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class="vector-dropdown-checkbox " aria-label="個人用ツール" > <label id="vector-user-links-dropdown-label" for="vector-user-links-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-ellipsis mw-ui-icon-wikimedia-ellipsis"></span> <span class="vector-dropdown-label-text">個人用ツール</span> </label> <div class="vector-dropdown-content"> <div id="p-personal" class="vector-menu mw-portlet mw-portlet-personal user-links-collapsible-item" title="利用者メニュー" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-sitesupport" class="user-links-collapsible-item mw-list-item"><a href="//donate.wikimedia.org/wiki/Special:FundraiserRedirector?utm_source=donate&utm_medium=sidebar&utm_campaign=C13_ja.wikipedia.org&uselang=ja"><span>寄付</span></a></li><li id="pt-createaccount" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=%E7%89%B9%E5%88%A5:%E3%82%A2%E3%82%AB%E3%82%A6%E3%83%B3%E3%83%88%E4%BD%9C%E6%88%90&returnto=%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9" title="アカウントを作成してログインすることをお勧めしますが、必須ではありません"><span class="vector-icon mw-ui-icon-userAdd mw-ui-icon-wikimedia-userAdd"></span> <span>アカウント作成</span></a></li><li id="pt-login" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=%E7%89%B9%E5%88%A5:%E3%83%AD%E3%82%B0%E3%82%A4%E3%83%B3&returnto=%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9" title="ログインすることを推奨します。ただし、必須ではありません。 [o]" accesskey="o"><span class="vector-icon mw-ui-icon-logIn mw-ui-icon-wikimedia-logIn"></span> <span>ログイン</span></a></li> </ul> </div> </div> <div id="p-user-menu-anon-editor" class="vector-menu mw-portlet mw-portlet-user-menu-anon-editor" > <div class="vector-menu-heading"> ログアウトした編集者のページ <a href="/wiki/Wikipedia:%E3%82%A6%E3%82%A3%E3%82%AD%E3%83%9A%E3%83%87%E3%82%A3%E3%82%A2%E3%81%B8%E3%82%88%E3%81%86%E3%81%93%E3%81%9D" aria-label="編集の詳細"><span>もっと詳しく</span></a> </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-anoncontribs" class="mw-list-item"><a href="/wiki/%E7%89%B9%E5%88%A5:%E8%87%AA%E5%88%86%E3%81%AE%E6%8A%95%E7%A8%BF%E8%A8%98%E9%8C%B2" title="このIPアドレスからなされた編集の一覧 [y]" accesskey="y"><span>投稿記録</span></a></li><li id="pt-anontalk" class="mw-list-item"><a href="/wiki/%E7%89%B9%E5%88%A5:%E3%83%88%E3%83%BC%E3%82%AF%E3%83%9A%E3%83%BC%E3%82%B8" title="このIPアドレスからなされた編集についての議論 [n]" accesskey="n"><span>トーク</span></a></li> </ul> </div> </div> </div> </div> </nav> </div> </header> </div> <div class="mw-page-container"> <div class="mw-page-container-inner"> <div class="vector-sitenotice-container"> <div id="siteNotice"></div> </div> <div class="vector-column-start"> <div class="vector-main-menu-container"> <div id="mw-navigation"> <nav id="mw-panel" class="vector-main-menu-landmark" aria-label="サイト"> <div id="vector-main-menu-pinned-container" class="vector-pinned-container"> </div> </nav> </div> </div> <div class="vector-sticky-pinned-container"> <nav id="mw-panel-toc" aria-label="目次" data-event-name="ui.sidebar-toc" class="mw-table-of-contents-container vector-toc-landmark"> <div id="vector-toc-pinned-container" class="vector-pinned-container"> <div id="vector-toc" class="vector-toc vector-pinnable-element"> <div class="vector-pinnable-header vector-toc-pinnable-header vector-pinnable-header-pinned" data-feature-name="toc-pinned" data-pinnable-element-id="vector-toc" > <h2 class="vector-pinnable-header-label">目次</h2> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-toc.pin">サイドバーに移動</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-toc.unpin">非表示</button> </div> <ul class="vector-toc-contents" id="mw-panel-toc-list"> <li id="toc-mw-content-text" class="vector-toc-list-item vector-toc-level-1"> <a href="#" class="vector-toc-link"> <div class="vector-toc-text">ページ先頭</div> </a> </li> <li id="toc-命名" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#命名"> <div class="vector-toc-text"> <span class="vector-toc-numb">1</span> <span>命名</span> </div> </a> <ul id="toc-命名-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-特徴" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#特徴"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>特徴</span> </div> </a> <ul id="toc-特徴-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-系統発生" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#系統発生"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>系統発生</span> </div> </a> <ul id="toc-系統発生-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-出典" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#出典"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>出典</span> </div> </a> <ul id="toc-出典-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="目次" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="目次の表示・非表示を切り替え" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">目次の表示・非表示を切り替え</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">タナトテリステス</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="特定の記事の別の言語版に移動します。利用可能な言語12件" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-12" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">12の言語版</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Thanatotheristes" title="チェコ語: Thanatotheristes" lang="cs" hreflang="cs" data-title="Thanatotheristes" data-language-autonym="Čeština" data-language-local-name="チェコ語" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Thanatotheristes" title="英語: Thanatotheristes" lang="en" hreflang="en" data-title="Thanatotheristes" data-language-autonym="English" data-language-local-name="英語" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Thanatotheristes_degrootorum" title="スペイン語: Thanatotheristes degrootorum" lang="es" hreflang="es" data-title="Thanatotheristes degrootorum" data-language-autonym="Español" data-language-local-name="スペイン語" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Thanatotheristes" title="フランス語: Thanatotheristes" lang="fr" hreflang="fr" data-title="Thanatotheristes" data-language-autonym="Français" data-language-local-name="フランス語" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%AA%D7%A0%D7%98%D7%95%D7%AA%D7%A8%D7%99%D7%A1%D7%98%D7%A1" title="ヘブライ語: תנטותריסטס" lang="he" hreflang="he" data-title="תנטותריסטס" data-language-autonym="עברית" data-language-local-name="ヘブライ語" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Thanatotheristes" title="イタリア語: Thanatotheristes" lang="it" hreflang="it" data-title="Thanatotheristes" data-language-autonym="Italiano" data-language-local-name="イタリア語" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Thanatotheristes" title="オランダ語: Thanatotheristes" lang="nl" hreflang="nl" data-title="Thanatotheristes" data-language-autonym="Nederlands" data-language-local-name="オランダ語" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Thanatotheristes" title="ポルトガル語: Thanatotheristes" lang="pt" hreflang="pt" data-title="Thanatotheristes" data-language-autonym="Português" data-language-local-name="ポルトガル語" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Thanatotheristes" title="ルーマニア語: Thanatotheristes" lang="ro" hreflang="ro" data-title="Thanatotheristes" data-language-autonym="Română" data-language-local-name="ルーマニア語" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/Thanatotheristes" title="ロシア語: Thanatotheristes" lang="ru" hreflang="ru" data-title="Thanatotheristes" data-language-autonym="Русский" data-language-local-name="ロシア語" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%A2%D0%B0%D0%BD%D0%B0%D1%82%D0%BE%D1%82%D0%B5%D1%80%D0%B8%D1%81%D1%82" title="ウクライナ語: Танатотерист" lang="uk" hreflang="uk" data-title="Танатотерист" data-language-autonym="Українська" data-language-local-name="ウクライナ語" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E6%AD%BB%E6%8E%A0%E9%BE%99%E5%B1%9E" title="中国語: 死掠龙属" lang="zh" hreflang="zh" data-title="死掠龙属" data-language-autonym="中文" data-language-local-name="中国語" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q83546128#sitelinks-wikipedia" title="言語間リンクを編集" class="wbc-editpage">リンクを編集</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="名前空間"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9" title="本文を閲覧 [c]" accesskey="c"><span>ページ</span></a></li><li id="ca-talk" class="new vector-tab-noicon mw-list-item"><a href="/w/index.php?title=%E3%83%8E%E3%83%BC%E3%83%88:%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9&action=edit&redlink=1" rel="discussion" class="new" title="「本文ページについての議論」 (存在しないページ) [t]" accesskey="t"><span>ノート</span></a></li> </ul> </div> </div> <div id="vector-variants-dropdown" class="vector-dropdown emptyPortlet" > <input type="checkbox" id="vector-variants-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-variants-dropdown" class="vector-dropdown-checkbox " aria-label="別の言語に切り替える" > <label id="vector-variants-dropdown-label" for="vector-variants-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">日本語</span> </label> <div class="vector-dropdown-content"> <div id="p-variants" class="vector-menu mw-portlet mw-portlet-variants emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> </div> </div> </nav> </div> <div id="right-navigation" class="vector-collapsible"> <nav aria-label="表示"> <div id="p-views" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-views" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-view" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9"><span>閲覧</span></a></li><li id="ca-edit" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9&action=edit" title="このページのソースコードを編集する [e]" accesskey="e"><span>編集</span></a></li><li id="ca-history" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9&action=history" title="このページの過去の版 [h]" accesskey="h"><span>履歴表示</span></a></li> </ul> </div> </div> </nav> <nav class="vector-page-tools-landmark" aria-label="ページツール"> <div id="vector-page-tools-dropdown" class="vector-dropdown vector-page-tools-dropdown" > <input type="checkbox" id="vector-page-tools-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-tools-dropdown" class="vector-dropdown-checkbox " aria-label="ツール" > <label id="vector-page-tools-dropdown-label" for="vector-page-tools-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">ツール</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-tools-unpinned-container" class="vector-unpinned-container"> <div id="vector-page-tools" class="vector-page-tools vector-pinnable-element"> <div class="vector-pinnable-header vector-page-tools-pinnable-header vector-pinnable-header-unpinned" data-feature-name="page-tools-pinned" data-pinnable-element-id="vector-page-tools" data-pinned-container-id="vector-page-tools-pinned-container" data-unpinned-container-id="vector-page-tools-unpinned-container" > <div class="vector-pinnable-header-label">ツール</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-page-tools.pin">サイドバーに移動</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-page-tools.unpin">非表示</button> </div> <div id="p-cactions" class="vector-menu mw-portlet mw-portlet-cactions emptyPortlet vector-has-collapsible-items" title="その他の操作" > <div class="vector-menu-heading"> 操作 </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-more-view" class="selected vector-more-collapsible-item mw-list-item"><a href="/wiki/%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9"><span>閲覧</span></a></li><li id="ca-more-edit" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9&action=edit" title="このページのソースコードを編集する [e]" accesskey="e"><span>編集</span></a></li><li id="ca-more-history" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9&action=history"><span>履歴表示</span></a></li> </ul> </div> </div> <div id="p-tb" class="vector-menu mw-portlet mw-portlet-tb" > <div class="vector-menu-heading"> 全般 </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="t-whatlinkshere" class="mw-list-item"><a href="/wiki/%E7%89%B9%E5%88%A5:%E3%83%AA%E3%83%B3%E3%82%AF%E5%85%83/%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9" title="ここにリンクしている全ウィキページの一覧 [j]" accesskey="j"><span>リンク元</span></a></li><li id="t-recentchangeslinked" class="mw-list-item"><a href="/wiki/%E7%89%B9%E5%88%A5:%E9%96%A2%E9%80%A3%E3%83%9A%E3%83%BC%E3%82%B8%E3%81%AE%E6%9B%B4%E6%96%B0%E7%8A%B6%E6%B3%81/%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9" rel="nofollow" title="このページからリンクしているページの最近の更新 [k]" accesskey="k"><span>関連ページの更新状況</span></a></li><li id="t-upload" class="mw-list-item"><a href="/wiki/Wikipedia:%E3%83%95%E3%82%A1%E3%82%A4%E3%83%AB%E3%81%AE%E3%82%A2%E3%83%83%E3%83%97%E3%83%AD%E3%83%BC%E3%83%89" title="ファイルをアップロードする [u]" accesskey="u"><span>ファイルをアップロード</span></a></li><li id="t-specialpages" class="mw-list-item"><a href="/wiki/%E7%89%B9%E5%88%A5:%E7%89%B9%E5%88%A5%E3%83%9A%E3%83%BC%E3%82%B8%E4%B8%80%E8%A6%A7" title="特別ページの一覧 [q]" accesskey="q"><span>特別ページ</span></a></li><li id="t-permalink" class="mw-list-item"><a href="/w/index.php?title=%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9&oldid=100965312" title="このページのこの版への固定リンク"><span>この版への固定リンク</span></a></li><li id="t-info" class="mw-list-item"><a href="/w/index.php?title=%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9&action=info" title="このページについての詳細情報"><span>ページ情報</span></a></li><li id="t-cite" class="mw-list-item"><a href="/w/index.php?title=%E7%89%B9%E5%88%A5:%E3%81%93%E3%81%AE%E3%83%9A%E3%83%BC%E3%82%B8%E3%82%92%E5%BC%95%E7%94%A8&page=%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9&id=100965312&wpFormIdentifier=titleform" title="このページの引用方法"><span>このページを引用</span></a></li><li id="t-urlshortener" class="mw-list-item"><a href="/w/index.php?title=%E7%89%B9%E5%88%A5:UrlShortener&url=https%3A%2F%2Fja.wikipedia.org%2Fwiki%2F%25E3%2582%25BF%25E3%2583%258A%25E3%2583%2588%25E3%2583%2586%25E3%2583%25AA%25E3%2582%25B9%25E3%2583%2586%25E3%2582%25B9"><span>短縮URLを取得する</span></a></li><li id="t-urlshortener-qrcode" class="mw-list-item"><a href="/w/index.php?title=%E7%89%B9%E5%88%A5:QrCode&url=https%3A%2F%2Fja.wikipedia.org%2Fwiki%2F%25E3%2582%25BF%25E3%2583%258A%25E3%2583%2588%25E3%2583%2586%25E3%2583%25AA%25E3%2582%25B9%25E3%2583%2586%25E3%2582%25B9"><span>QRコードをダウンロード</span></a></li> </ul> </div> </div> <div id="p-coll-print_export" class="vector-menu mw-portlet mw-portlet-coll-print_export" > <div class="vector-menu-heading"> 印刷/書き出し </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="coll-create_a_book" class="mw-list-item"><a href="/w/index.php?title=%E7%89%B9%E5%88%A5:%E3%83%96%E3%83%83%E3%82%AF&bookcmd=book_creator&referer=%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9"><span>ブックの新規作成</span></a></li><li id="coll-download-as-rl" class="mw-list-item"><a href="/w/index.php?title=%E7%89%B9%E5%88%A5:DownloadAsPdf&page=%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9&action=show-download-screen"><span>PDF 形式でダウンロード</span></a></li><li id="t-print" class="mw-list-item"><a href="/w/index.php?title=%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9&printable=yes" title="このページの印刷用ページ [p]" accesskey="p"><span>印刷用バージョン</span></a></li> </ul> </div> </div> <div id="p-wikibase-otherprojects" class="vector-menu mw-portlet mw-portlet-wikibase-otherprojects" > <div class="vector-menu-heading"> 他のプロジェクト </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="wb-otherproject-link wb-otherproject-commons mw-list-item"><a href="https://commons.wikimedia.org/wiki/Category:Thanatotheristes" hreflang="en"><span>コモンズ</span></a></li><li id="t-wikibase" class="wb-otherproject-link wb-otherproject-wikibase-dataitem mw-list-item"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q83546128" title="関連付けられたデータリポジトリ項目へのリンク [g]" accesskey="g"><span>ウィキデータ項目</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> </div> </div> </div> <div class="vector-column-end"> <div class="vector-sticky-pinned-container"> <nav class="vector-page-tools-landmark" aria-label="ページツール"> <div id="vector-page-tools-pinned-container" class="vector-pinned-container"> </div> </nav> <nav class="vector-appearance-landmark" aria-label="表示"> <div id="vector-appearance-pinned-container" class="vector-pinned-container"> <div id="vector-appearance" class="vector-appearance vector-pinnable-element"> <div class="vector-pinnable-header vector-appearance-pinnable-header vector-pinnable-header-pinned" data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">表示</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">サイドバーに移動</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">非表示</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">出典: フリー百科事典『ウィキペディア（Wikipedia）』</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="ja" dir="ltr"><table class="infobox bordered" style="width:18em;"> <tbody><tr> <th style="font-size:110%;background:transparent;background:rgb(211,211,164);text-align:center;">タナトテリステス </th></tr> <tr> <td style="background:transparent;background:rgb(211,211,164);text-align:center;"><div style="text-align:center; font-size:smaller;">生息年代: <a href="/wiki/%E4%B8%AD%E7%94%9F%E4%BB%A3" title="中生代">中生代</a><a href="/wiki/%E5%BE%8C%E6%9C%9F%E7%99%BD%E4%BA%9C%E7%B4%80" title="後期白亜紀">後期白亜紀</a>、80.1–79.5 Ma <div id="Timeline-row" style="margin: 4px auto 0; clear:both; width:200px; padding:0px; height:18px; overflow:visible; border:1px #666; border-style:solid none; position:relative; z-index:0; font-size:smaller; font-weight:bolder; white-space:nowrap;"> <div id="PreЄ_marker" style="z-index:1000; white-space:nowrap; position:absolute; left:5px; overflow:visible; width:0; height:100%;"><a href="/wiki/%E5%85%88%E3%82%AB%E3%83%B3%E3%83%96%E3%83%AA%E3%82%A2%E6%99%82%E4%BB%A3" title="先カンブリア時代">PreЄ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.05; filter:alpha(opacity=5); width:1px; left:0px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.1; filter:alpha(opacity=10); width:1px; left:1px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.15; filter:alpha(opacity=15); width:1px; left:2px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.2; filter:alpha(opacity=20); width:1px; left:3px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.25; filter:alpha(opacity=25); width:1px; left:4px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.3; filter:alpha(opacity=30); width:1px; left:5px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.35; filter:alpha(opacity=30); width:1px; left:6px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.4; filter:alpha(opacity=40); width:1px; left:7px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.45; filter:alpha(opacity=45); width:1px; left:8px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.5; filter:alpha(opacity=50); width:1px; left:9px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.55; filter:alpha(opacity=55); width:1px; left:10px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.6; filter:alpha(opacity=60); width:1px; left:11px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.65; filter:alpha(opacity=65); width:1px; left:12px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.7; filter:alpha(opacity=70); width:1px; left:13px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.75; filter:alpha(opacity=75); width:1px; left:14px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.8; filter:alpha(opacity=80); width:1px; left:15px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.85; filter:alpha(opacity=85); width:1px; left:16px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.9; filter:alpha(opacity=90); width:1px; left:17px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); opacity:0.95; filter:alpha(opacity=95); width:1px; left:18px"></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(254,217,106); left:19px; width:14.230769230769px;"></div> <div id="bar" style="position:absolute; height:100%; text-align:center; background-color:rgb(127,160,86); left:33.538461538462px; width:17.107692307692px;"><a href="/wiki/%E3%82%AB%E3%83%B3%E3%83%96%E3%83%AA%E3%82%A2%E7%B4%80" title="カンブリア紀">Є</a></div> <div id="bar" style="position:absolute; height:100%; text-align:center; background-color:rgb(0,146,112); left:50.646153846154px; width:12.8px;"><a href="/wiki/%E3%82%AA%E3%83%AB%E3%83%89%E3%83%93%E3%82%B9%E7%B4%80" title="オルドビス紀">O</a></div> <div id="bar" style="position:absolute; height:100%; text-align:center; background-color:rgb(179,225,182); left:63.446153846154px; width:7.5692307692308px;"><a href="/wiki/%E3%82%B7%E3%83%AB%E3%83%AB%E7%B4%80" title="シルル紀">S</a></div> <div id="bar" style="position:absolute; height:100%; text-align:center; background-color:rgb(203,140,55); left:71.015384615385px; width:18.553846153846px;"><a href="/wiki/%E3%83%87%E3%83%9C%E3%83%B3%E7%B4%80" title="デボン紀">D</a></div> <div id="bar" style="position:absolute; height:100%; text-align:center; background-color:rgb(103,165,153); left:89.569230769231px; width:18.461538461538px;"><a href="/wiki/%E7%9F%B3%E7%82%AD%E7%B4%80" title="石炭紀">C</a></div> <div id="bar" style="position:absolute; height:100%; text-align:center; background-color:rgb(240,64,40); left:108.03076923077px; width:14.460923076923px;"><a href="/wiki/%E3%83%9A%E3%83%AB%E3%83%A0%E7%B4%80" title="ペルム紀">P</a></div> <div id="bar" style="position:absolute; height:100%; text-align:center; background-color:rgb(129,43,146); left:122.49169230769px; width:15.569846153846px;"><a href="/wiki/%E4%B8%89%E7%95%B3%E7%B4%80" title="三畳紀">T</a></div> <div id="bar" style="position:absolute; height:100%; text-align:center; background-color:rgb(52,178,201); left:138.06153846154px; width:17.323076923077px;"><a href="/wiki/%E3%82%B8%E3%83%A5%E3%83%A9%E7%B4%80" title="ジュラ紀">J</a></div> <div id="bar" style="position:absolute; height:100%; text-align:center; background-color:rgb(127,198,78); left:155.38461538462px; width:24.307692307692px;"><a href="/wiki/%E7%99%BD%E4%BA%9C%E7%B4%80" title="白亜紀">K</a></div> <div id="bar" style="position:absolute; height:100%; text-align:center; background-color:rgb(253,154,82); left:179.69230769231px; width:13.221538461538px;"><a href="/wiki/%E5%8F%A4%E7%AC%AC%E4%B8%89%E7%B4%80" title="古第三紀"><small>Pg</small></a></div> <div id="bar" style="position:absolute; height:100%; text-align:center; background-color:rgb(255,230,25); left:192.91384615385px; width:6.2923076923077px;"><a href="/wiki/%E6%96%B0%E7%AC%AC%E4%B8%89%E7%B4%80" title="新第三紀"><small>N</small></a></div> <div id="end-border" style="position:absolute; height:100%; background-color:#666; width:1px; left:199px"></div></div><div style="margin:0 auto; line-height:0; clear:both; width:200px; padding:0px; height:8px; overflow:visible; background-color:transparent; position:relative; top:-4px; z-index:100;"> <div style="position:absolute; left:175.35384615385px; font-size:50%"><div style="position:relative; left:-0.42em">↓</div></div> </div></div> </td></tr> <tr style="text-align:center; font-size:smaller;"> <td><span typeof="mw:File"><a href="/wiki/%E3%83%95%E3%82%A1%E3%82%A4%E3%83%AB:Thanatotheristes.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/41/Thanatotheristes.jpg/250px-Thanatotheristes.jpg" decoding="async" width="250" height="97" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/41/Thanatotheristes.jpg/375px-Thanatotheristes.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/41/Thanatotheristes.jpg/500px-Thanatotheristes.jpg 2x" data-file-width="5096" data-file-height="1972" /></a></span> <div>タナトテリステスの復元図</div> </td></tr> <tr> <th style="background:transparent;background:rgb(211,211,164);text-align:center;"><a href="/wiki/%E5%9C%B0%E8%B3%AA%E6%99%82%E4%BB%A3" title="地質時代">地質時代</a> </th></tr> <tr> <td style="text-align:center"><a href="/wiki/%E4%B8%AD%E7%94%9F%E4%BB%A3" title="中生代">中生代</a><a href="/wiki/%E5%BE%8C%E6%9C%9F%E7%99%BD%E4%BA%9C%E7%B4%80" title="後期白亜紀">後期白亜紀</a><br />（約8010万~7950万年前） </td></tr> <tr> <th style="background:transparent;background:rgb(211,211,164);text-align:center;"><a href="/wiki/%E7%94%9F%E7%89%A9%E3%81%AE%E5%88%86%E9%A1%9E" title="生物の分類">分類</a> </th></tr> <tr> <td> <table class="borderless" style="margin:auto;"> <tbody><tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><small><a href="/wiki/%E3%83%89%E3%83%A1%E3%82%A4%E3%83%B3_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="ドメイン (分類学)">ドメイン</a></small> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E7%9C%9F%E6%A0%B8%E7%94%9F%E7%89%A9" title="真核生物">真核生物</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/en:Eukaryote" class="extiw" title="w:en:Eukaryote">Eukaryota</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><a href="/wiki/%E7%95%8C_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="界 (分類学)">界</a> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E5%8B%95%E7%89%A9" title="動物">動物界</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/en:Animal" class="extiw" title="w:en:Animal">Animalia</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><a href="/wiki/%E9%96%80_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="門 (分類学)">門</a> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E8%84%8A%E7%B4%A2%E5%8B%95%E7%89%A9" title="脊索動物">脊索動物門</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/en:Chordate" class="extiw" title="w:en:Chordate">Chordata</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><a href="/wiki/%E9%96%80_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="門 (分類学)">亜門</a> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E8%84%8A%E6%A4%8E%E5%8B%95%E7%89%A9" title="脊椎動物">脊椎動物亜門</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/en:Vertebrate" class="extiw" title="w:en:Vertebrate">Vertebrata</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><a href="/wiki/%E7%B6%B1_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="綱 (分類学)">綱</a> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E7%88%AC%E8%99%AB%E9%A1%9E" title="爬虫類">爬虫綱</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/Reptilia" class="extiw" title="w:Reptilia">Reptilia</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><a href="/wiki/%E7%B6%B1_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="綱 (分類学)">亜綱</a> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E5%8F%8C%E5%BC%93%E9%A1%9E" title="双弓類">双弓亜綱</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/en:Diapsid" class="extiw" title="w:en:Diapsid">Diapsida</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><a href="/wiki/%E7%B6%B1_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="綱 (分類学)">下綱</a> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E4%B8%BB%E7%AB%9C%E5%BD%A2%E9%A1%9E" title="主竜形類">主竜形下綱</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/Archosauromorpha" class="extiw" title="w:Archosauromorpha">Archosauromorpha</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><a href="/wiki/%E7%9B%AE_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="目 (分類学)">上目</a> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E6%81%90%E7%AB%9C" title="恐竜">恐竜上目</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/en:Dinosaur" class="extiw" title="w:en:Dinosaur">Dinosauria</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><a href="/wiki/%E7%9B%AE_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="目 (分類学)">目</a> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E7%AB%9C%E7%9B%A4%E9%A1%9E" title="竜盤類">竜盤目</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/Saurischia" class="extiw" title="w:Saurischia">Saurischia</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><small>階級なし</small> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E7%9C%9F%E7%AB%9C%E7%9B%A4%E9%A1%9E" class="mw-redirect" title="真竜盤類">真竜盤類</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/Eusaurischia" class="extiw" title="w:Eusaurischia">Eusaurischia</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><a href="/wiki/%E7%9B%AE_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="目 (分類学)">亜目</a> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E7%8D%A3%E8%84%9A%E9%A1%9E" title="獣脚類">獣脚亜目</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/Theropoda" class="extiw" title="w:Theropoda">Theropoda</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><small>階級なし</small> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E3%82%A2%E3%83%B4%E3%82%A7%E3%83%AD%E3%82%B9%E3%83%88%E3%83%A9" class="mw-redirect" title="アヴェロストラ">鳥吻類</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/Averostra" class="extiw" title="w:Averostra">Averostra</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><a href="/wiki/%E7%9B%AE_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="目 (分類学)">下目</a> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E3%83%86%E3%82%BF%E3%83%8C%E3%83%A9%E9%A1%9E" title="テタヌラ類">テタヌラ下目</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/Tetanurae" class="extiw" title="w:Tetanurae">Tetanurae</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><small>階級なし</small> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E3%82%B3%E3%82%A8%E3%83%AB%E3%83%AD%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9%E9%A1%9E" title="コエルロサウルス類">コエルロサウルス類</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/Coelurosauria" class="extiw" title="w:Coelurosauria">Coelurosauria</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><a href="/wiki/%E7%A7%91_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="科 (分類学)">上科</a> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E3%83%86%E3%82%A3%E3%83%A9%E3%83%8E%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9%E4%B8%8A%E7%A7%91" title="ティラノサウルス上科">ティラノサウルス上科</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/Tyrannosauroidea" class="extiw" title="w:Tyrannosauroidea">Tyrannosauroidea</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><a href="/wiki/%E7%A7%91_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="科 (分類学)">科</a> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E3%83%86%E3%82%A3%E3%83%A9%E3%83%8E%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9%E7%A7%91" title="ティラノサウルス科">ティラノサウルス科</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/Tyrannosauridae" class="extiw" title="w:Tyrannosauridae">Tyrannosauridae</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><a href="/wiki/%E7%A7%91_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="科 (分類学)">亜科</a> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E3%83%86%E3%82%A3%E3%83%A9%E3%83%8E%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9%E4%BA%9C%E7%A7%91" title="ティラノサウルス亜科">ティラノサウルス亜科</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/Tyrannosaurinae" class="extiw" title="w:Tyrannosaurinae">Tyrannosaurinae</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><a href="/wiki/%E6%97%8F_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="族 (分類学)">族</a> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><a href="/wiki/%E3%83%80%E3%82%B9%E3%83%97%E3%83%AC%E3%83%88%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9%E6%97%8F" title="ダスプレトサウルス族">ダスプレトサウルス族</a> <span lang="la"><a href="https://en.wikipedia.org/wiki/Daspletosaurini" class="extiw" title="w:Daspletosaurini">Daspletosaurini</a></span> </td></tr> <tr> <td style="white-space:nowrap; vertical-align:middle; text-align:right;"><a href="/wiki/%E5%B1%9E_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="属 (分類学)">属</a> </td> <td style="vertical-align:middle;">: </td> <td style="vertical-align:middle; text-align:left;"><b>タナトテリステス属</b> <i><span lang="la"><a href="https://en.wikipedia.org/wiki/Thanatotheristes" class="extiw" title="w:Thanatotheristes">Thanatotheristes</a></span></i> </td></tr> </tbody></table> </td></tr> <tr> <th style="background:transparent;background:rgb(211,211,164);text-align:center;"><a href="/wiki/%E5%AD%A6%E5%90%8D" title="学名">学名</a> </th></tr> <tr> <td lang="la" style="text-align:center"><i><b>Thanatotheristes</b></i><br /><span lang="la">Voris <i>et al.</i>, <a href="/wiki/2020%E5%B9%B4" title="2020年">2020</a></span> </td></tr> <tr> <th style="background:transparent;background:rgb(211,211,164);text-align:center;"><a href="/wiki/%E7%A8%AE_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="種 (分類学)">種</a> </th></tr> <tr> <td> <ul><li><i>T. degrootorum</i> <span lang="la">Voris <i>et al.</i>, <a href="/wiki/2020%E5%B9%B4" title="2020年">2020</a></span>（タイプ種）</li></ul> </td></tr> </tbody></table> <p><b>タナトテリステス</b>(<i>Thanatotheristes</i>)（「刈り取る<a href="/wiki/%E6%AD%BB%E7%A5%9E" title="死神">死神</a>」の意）<sup id="cite_ref-Voris2020_1-0" class="reference"><a href="#cite_note-Voris2020-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup>は、<a href="/wiki/%E5%BE%8C%E6%9C%9F%E7%99%BD%E4%BA%9C%E7%B4%80" title="後期白亜紀">後期白亜紀</a>（約7950万年前）の<a href="/wiki/%E3%83%A9%E3%83%A9%E3%83%9F%E3%83%87%E3%82%A3%E3%82%A2%E5%A4%A7%E9%99%B8" title="ララミディア大陸">ララミディア大陸</a>に生息していた<a href="/wiki/%E3%83%86%E3%82%A3%E3%83%A9%E3%83%8E%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9%E7%A7%91" title="ティラノサウルス科">ティラノサウルス科</a>の<a href="/wiki/%E7%8D%A3%E8%84%9A%E9%A1%9E" title="獣脚類">獣脚類</a>の<a href="/wiki/%E5%B1%9E_(%E5%88%86%E9%A1%9E%E5%AD%A6)" title="属 (分類学)">属</a>の一つ。<a href="/wiki/%E3%82%A2%E3%83%AB%E3%83%90%E3%83%BC%E3%82%BF%E5%B7%9E" title="アルバータ州">アルバータ州</a>の<a href="/wiki/%E3%83%95%E3%82%A9%E3%82%A2%E3%83%A2%E3%82%B9%E3%83%88%E5%B1%A4" class="mw-redirect" title="フォアモスト層">フォアモスト層</a>で<a href="/wiki/%E5%8C%96%E7%9F%B3" title="化石">化石</a>が発見された。中型<a href="/wiki/%E3%82%B1%E3%83%A9%E3%83%88%E3%83%97%E3%82%B9%E9%A1%9E" class="mw-redirect" title="ケラトプス類">ケラトプス類</a>の<a href="/wiki/%E3%82%BC%E3%83%8E%E3%82%B1%E3%83%A9%E3%83%88%E3%83%97%E3%82%B9" title="ゼノケラトプス">ゼノケラトプス</a>や小型<a href="/wiki/%E3%83%91%E3%82%AD%E3%82%B1%E3%83%95%E3%82%A1%E3%83%AD%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9%E9%A1%9E" class="mw-redirect" title="パキケファロサウルス類">パキケファロサウルス類</a>の<a href="/wiki/%E3%82%B3%E3%83%AC%E3%83%94%E3%82%AA%E3%82%B1%E3%83%95%E3%82%A1%E3%83%AC" title="コレピオケファレ">コレピオケファレ</a>などと共存した<sup id="cite_ref-講談社_3-0" class="reference"><a href="#cite_note-講談社-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup>。<a href="/wiki/%E3%83%80%E3%82%B9%E3%83%97%E3%83%AC%E3%83%88%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9" title="ダスプレトサウルス">ダスプレトサウルス</a>の可能性がある。 </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="命名"><span id=".E5.91.BD.E5.90.8D"></span>命名</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9&action=edit&section=1" title="節を編集: 命名"><span>編集</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>2020年に模式種ナトテリステス・デグロオトルムが命名された。属名は<a href="/wiki/%E6%AD%BB" title="死">死</a>を神格化した<a href="/wiki/%E3%82%AE%E3%83%AA%E3%82%B7%E3%83%A3%E7%A5%9E%E8%A9%B1" class="mw-redirect" title="ギリシャ神話">ギリシャ神話</a>の神<a href="/wiki/%E3%82%BF%E3%83%8A%E3%83%88%E3%82%B9" title="タナトス">タナトス</a>に由来し、種小名は「刈り取る者」を意味し、併せて「死を刈り取る神」といった意になる<sup id="cite_ref-講談社_3-1" class="reference"><a href="#cite_note-講談社-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup>。 </p> <div class="mw-heading mw-heading2"><h2 id="特徴"><span id=".E7.89.B9.E5.BE.B4"></span>特徴</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9&action=edit&section=2" title="節を編集: 特徴"><span>編集</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/%E3%83%95%E3%82%A1%E3%82%A4%E3%83%AB:Thanatotheristes_Size_Comparison.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/89/Thanatotheristes_Size_Comparison.svg/220px-Thanatotheristes_Size_Comparison.svg.png" decoding="async" width="220" height="81" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/89/Thanatotheristes_Size_Comparison.svg/330px-Thanatotheristes_Size_Comparison.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/89/Thanatotheristes_Size_Comparison.svg/440px-Thanatotheristes_Size_Comparison.svg.png 2x" data-file-width="512" data-file-height="189" /></a><figcaption>タナトテリステスと<a href="/wiki/%E3%83%92%E3%83%88" title="ヒト">ヒト</a>との大きさの比較</figcaption></figure> <p>タナトテリステス・デグロオトルムの<a href="/wiki/%E3%83%9B%E3%83%AD%E3%82%BF%E3%82%A4%E3%83%97" class="mw-redirect" title="ホロタイプ">ホロタイプ</a>(TMP 2010.5.7) は、右<a href="/wiki/%E4%B8%8A%E9%A1%8E%E9%AA%A8" title="上顎骨">上顎骨</a>、右<a href="/wiki/%E9%A0%AC%E9%AA%A8" title="頬骨">頬骨</a>、右後眼窩骨、右上角骨、右方形骨、右の側蝶形骨、左<a href="/wiki/%E5%89%8D%E9%A0%AD%E9%AA%A8" title="前頭骨">前頭骨</a>、両方の<a href="/wiki/%E6%AD%AF%E9%AA%A8" class="mw-redirect" title="歯骨">歯骨</a>で構成される。頭骨長は約80センチメートル。近縁属である<a href="/wiki/%E3%83%80%E3%82%B9%E3%83%97%E3%83%AC%E3%83%88%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9" title="ダスプレトサウルス">ダスプレトサウルス</a>よりも小さいが、ホロタイプの個体は骨格的には未成熟である。参考標本TMP 2018.016.0001 は、亜成体の断片的な右上顎骨のみに基づく。これはフォアモスト層のヘロントン砂岩上部のTwelve Mile Coulee で発見された。<a href="/wiki/%E7%A8%AE%E5%B0%8F%E5%90%8D" class="mw-redirect" title="種小名">種小名</a>は模式標本を発見したジョン・デ・グルートとサンドラ・デ・グルートへの献名である<sup id="cite_ref-Voris2020_1-1" class="reference"><a href="#cite_note-Voris2020-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>。 </p><p>T・デグロオトルムは以下の形質によって特徴付けられる<sup id="cite_ref-Voris2020_1-2" class="reference"><a href="#cite_note-Voris2020-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>。 </p> <ul><li>頭骨の表面における、上顎骨の腹側と前眼窩窓に等間隔で並んだ正中線方向の一列の隆起。</li> <li>顎骨の丸みを帯びた膨らみのある眼窩縁。</li> <li>前頭骨側矢状突起は上顎骨稜の前方に広く丸みを帯びた隆起として延びる。</li> <li>前頭骨と涙骨の接触面が前頭骨間縫合線に対して60°以下で前方に延びる。</li> <li>頭頂骨の腹側表面に前頭骨と関節を形成する2つの後方に突出した枝を持つ。</li></ul> <p>このタナトテリステスの模式種の全長は、頭骨要素と近縁種からの比較情報に基づき約8メートルと推定されている<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup>。 </p> <div class="mw-heading mw-heading2"><h2 id="系統発生"><span id=".E7.B3.BB.E7.B5.B1.E7.99.BA.E7.94.9F"></span>系統発生</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9&action=edit&section=3" title="節を編集: 系統発生"><span>編集</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>タナトテリステスの正式記載により、ティラノサウルス科の中にティラノサウルス亜科の姉妹群である<a href="/wiki/%E3%83%80%E3%82%B9%E3%83%97%E3%83%AC%E3%83%88%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9%E6%97%8F" title="ダスプレトサウルス族">ダスプレトサウルス族</a>という新分類群が誕生した。この分類群は1種のタナトテリステスと2種の<a href="/wiki/%E3%83%80%E3%82%B9%E3%83%97%E3%83%AC%E3%83%88%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9" title="ダスプレトサウルス">ダスプレトサウルス</a>、そして未命名の<a href="/wiki/%E3%83%80%E3%82%A4%E3%83%8A%E3%82%BD%E3%83%BC%E3%83%91%E3%83%BC%E3%82%AF%E5%B1%A4" class="mw-redirect" title="ダイナソーパーク層">ダイナソーパーク層</a>のティラノサウルス科 (FMNH PR308)を内包する。ダスプレトサウルス亜科の存在はティラノサウルス科の中でも地理的な棲み分けがあることを示しており、ララミディアの南部には<a href="/wiki/%E3%83%93%E3%82%B9%E3%82%BF%E3%83%92%E3%82%A8%E3%83%B4%E3%82%A7%E3%83%AB%E3%82%BD%E3%83%AB" title="ビスタヒエヴェルソル">ビスタヒエヴェルソル</a>、<a href="/wiki/%E3%83%AA%E3%83%88%E3%83%AD%E3%83%8A%E3%82%AF%E3%82%B9" title="リトロナクス">リトロナクス</a>、<a href="/wiki/%E3%83%86%E3%83%A9%E3%83%88%E3%83%95%E3%82%A9%E3%83%8D%E3%82%A6%E3%82%B9" title="テラトフォネウス">テラトフォネウス</a>によって形成された<a href="/wiki/%E3%82%AF%E3%83%AC%E3%83%BC%E3%83%89" class="mw-redirect" title="クレード">クレード</a>が、カナダおよびアメリカ北部には<a href="/wiki/%E3%82%A2%E3%83%AB%E3%83%90%E3%83%BC%E3%83%88%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9%E4%BA%9C%E7%A7%91" title="アルバートサウルス亜科">アルバートサウルス亜科</a>とダスプレトサウルス族が棲息していた<sup id="cite_ref-Voris2020_1-3" class="reference"><a href="#cite_note-Voris2020-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>。 </p><p>以下の<a href="/wiki/%E3%82%AF%E3%83%A9%E3%83%89%E3%82%B0%E3%83%A9%E3%83%A0" title="クラドグラム">クラドグラム</a>は Voris <i>et al.</i> (2020) の系統解析に基づく<sup id="cite_ref-Voris2020_1-4" class="reference"><a href="#cite_note-Voris2020-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>。 <style data-mw-deduplicate="TemplateStyles:r89142105">.mw-parser-output table.clade{border-spacing:0;margin:0;font-size:100%;line-height:100%;border-collapse:separate;width:auto}.mw-parser-output table.clade table.clade{width:100%}.mw-parser-output table.clade td.clade-label{width:0.7em;padding:0 0.15em;vertical-align:bottom;text-align:center;border-left:1px solid;border-bottom:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-fixed-width{overflow:hidden;text-overflow:ellipsis}.mw-parser-output table.clade td.clade-fixed-width:hover{overflow:visible}.mw-parser-output table.clade td.clade-label.first{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-label.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-slabel{padding:0 0.15em;vertical-align:top;text-align:center;border-left:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-slabel:hover{overflow:visible}.mw-parser-output table.clade td.clade-slabel.last{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-slabel.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-bar{vertical-align:middle;text-align:left;padding:0 0.5em;position:relative}.mw-parser-output table.clade td.clade-bar.reverse{text-align:right;position:relative}.mw-parser-output table.clade td.clade-leaf{border:0;padding:0;text-align:left}.mw-parser-output table.clade td.clade-leafR{border:0;padding:0;text-align:right}.mw-parser-output table.clade td.clade-leaf.reverse{text-align:right}.mw-parser-output table.clade:hover span.linkA{background-color:yellow}.mw-parser-output table.clade:hover span.linkB{background-color:green}</style> </p> <table class="clade" style="font-size:100%; line-height:100%; width:65%"> <tbody><tr> <td class="clade-label first"><a href="/wiki/%E3%82%A8%E3%82%A6%E3%83%86%E3%82%A3%E3%83%A9%E3%83%8E%E3%82%B5%E3%82%A6%E3%83%AA%E3%82%A2" class="mw-redirect" title="エウティラノサウリア">エウティラノサウリア</a> </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first">  </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first">  </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/%E3%83%89%E3%83%AA%E3%83%97%E3%83%88%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9" title="ドリプトサウルス">Dryptosaurus aquilunguis</a></i> </p> </td></tr> <tr> <td class="clade-slabel">  </td></tr> <tr> <td class="clade-label">  </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first">  </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/%E3%82%A2%E3%83%91%E3%83%A9%E3%83%81%E3%82%AA%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9" title="アパラチオサウルス">Appalachiosaurus montgomeriensis</a></i> </p> </td></tr> <tr> <td class="clade-slabel">  </td></tr> <tr> <td class="clade-label">  </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first">  </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/%E3%83%93%E3%82%B9%E3%82%BF%E3%83%92%E3%82%A8%E3%83%B4%E3%82%A7%E3%83%AB%E3%82%BD%E3%83%AB" title="ビスタヒエヴェルソル">Bistahieversor sealeyi</a></i> </p> </td></tr> <tr> <td class="clade-slabel">  </td></tr> <tr> <td class="clade-label">  </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/%E3%83%86%E3%82%A3%E3%83%A9%E3%83%8E%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9%E7%A7%91" title="ティラノサウルス科">ティラノサウルス科</a> </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/%E3%82%A2%E3%83%AB%E3%83%90%E3%83%BC%E3%83%88%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9%E4%BA%9C%E7%A7%91" title="アルバートサウルス亜科">アルバートサウルス亜科</a> </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first">  </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/%E3%82%B4%E3%83%AB%E3%82%B4%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9" title="ゴルゴサウルス">Gorgosaurus libratus</a></i> </p> </td></tr> <tr> <td class="clade-slabel">  </td></tr> <tr> <td class="clade-label">  </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/%E3%82%A2%E3%83%AB%E3%83%90%E3%83%BC%E3%83%88%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9" title="アルバートサウルス">Albertosaurus sarcophagus</a></i> </p> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel">  </td></tr> <tr> <td class="clade-label"><a href="/wiki/%E3%83%86%E3%82%A3%E3%83%A9%E3%83%8E%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9%E4%BA%9C%E7%A7%91" title="ティラノサウルス亜科">ティラノサウルス亜科</a> </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/%E3%82%A2%E3%83%AA%E3%82%AA%E3%83%A9%E3%83%A0%E3%82%B9%E6%97%8F" title="アリオラムス族">アリオラムス族</a> </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first">  </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/%E3%82%AD%E3%82%A2%E3%83%B3%E3%82%BE%E3%82%A6%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9" title="キアンゾウサウルス">Qianzhousaurus sinensis</a></i> </p> </td></tr> <tr> <td class="clade-slabel">  </td></tr> <tr> <td class="clade-label">  </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first">  </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/%E3%82%A2%E3%83%AA%E3%82%AA%E3%83%A9%E3%83%A0%E3%82%B9" title="アリオラムス">Alioramus remotus</a></i> </p> </td></tr> <tr> <td class="clade-slabel">  </td></tr> <tr> <td class="clade-label">  </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/%E3%82%A2%E3%83%AA%E3%82%AA%E3%83%A9%E3%83%A0%E3%82%B9" title="アリオラムス">Alioramus altai</a></i> </p> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel">  </td></tr> <tr> <td class="clade-label">  </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first">  </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first">  </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/%E3%83%86%E3%83%A9%E3%83%88%E3%83%95%E3%82%A9%E3%83%8D%E3%82%A6%E3%82%B9" title="テラトフォネウス">Teratophoneus curriei</a></i> </p> </td></tr> <tr> <td class="clade-slabel">  </td></tr> <tr> <td class="clade-label">  </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/%E3%83%87%E3%82%A3%E3%83%8A%E3%83%A2%E3%83%86%E3%83%A9%E3%83%BC" title="ディナモテラー">Dynamoterror dynastes</a></i> </p> </td></tr> <tr> <td class="clade-slabel">  </td></tr> <tr> <td class="clade-label">  </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/%E3%83%AA%E3%83%88%E3%83%AD%E3%83%8A%E3%82%AF%E3%82%B9" title="リトロナクス">Lythronax argestes</a></i> </p> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel">  </td></tr> <tr> <td class="clade-label">  </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first">  </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/%E3%83%8A%E3%83%8C%E3%83%BC%E3%82%AF%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9" title="ナヌークサウルス">Nanuqsaurus hoglundi</a></i> </p> </td></tr> <tr> <td class="clade-slabel">  </td></tr> <tr> <td class="clade-label">  </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first">  </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/%E3%83%80%E3%82%B9%E3%83%97%E3%83%AC%E3%83%88%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9%E6%97%8F" title="ダスプレトサウルス族">ダスプレトサウルス族</a> </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first">  </td> <td rowspan="2" class="clade-leaf"> <p><i><b>Thanatotheristes degrootorum</b></i> </p> </td></tr> <tr> <td class="clade-slabel">  </td></tr> <tr> <td class="clade-label">  </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first">  </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/%E3%83%80%E3%82%B9%E3%83%97%E3%83%AC%E3%83%88%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9" title="ダスプレトサウルス">Daspletosaurus torosus</a></i> </p> </td></tr> <tr> <td class="clade-slabel">  </td></tr> <tr> <td class="clade-label">  </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/%E3%83%80%E3%82%B9%E3%83%97%E3%83%AC%E3%83%88%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9" title="ダスプレトサウルス">Daspletosaurus horneri</a></i> </p> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel">  </td></tr> <tr> <td class="clade-label">  </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first">  </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/%E3%82%BA%E3%82%B1%E3%83%B3%E3%83%86%E3%82%A3%E3%83%A9%E3%83%8C%E3%82%B9" title="ズケンティラヌス">Zhuchengtyrannus magnus</a></i> </p> </td></tr> <tr> <td class="clade-slabel">  </td></tr> <tr> <td class="clade-label">  </td> <td rowspan="2" class="clade-leaf"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r89142105"> <table class="clade"> <tbody><tr> <td class="clade-label first">  </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/%E3%82%BF%E3%83%AB%E3%83%9C%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9" title="タルボサウルス">Tarbosaurus bataar</a></i> </p> </td></tr> <tr> <td class="clade-slabel">  </td></tr> <tr> <td class="clade-label">  </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/%E3%83%86%E3%82%A3%E3%83%A9%E3%83%8E%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9" title="ティラノサウルス">Tyrannosaurus rex</a></i> </p> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> </td></tr> <tr> <td class="clade-slabel last">  </td></tr></tbody></table> <p>なお、Yun (2020) による亜成体のダスプレトサウルス・トロススの<a href="/wiki/%E5%89%8D%E9%A0%AD%E9%AA%A8" title="前頭骨">前頭骨</a>の解析では、前頭骨の形態の差が大きいことから、ティラノサウルス科の前頭骨の<a href="/wiki/%E5%9B%BA%E6%9C%89%E6%B4%BE%E7%94%9F%E5%BD%A2%E8%B3%AA" title="固有派生形質">固有派生形質</a>のいくつかに不備があるとされた。すなわち、同研究ではタナトテリステス・デグロオトルムを独立した属と見なすための形質が不十分とされ、本種はダスプレトサウルスの種として扱われている<sup id="cite_ref-yun2020_5-0" class="reference"><a href="#cite_note-yun2020-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup>。 </p> <div class="mw-heading mw-heading2"><h2 id="出典"><span id=".E5.87.BA.E5.85.B8"></span>出典</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9&action=edit&section=4" title="節を編集: 出典"><span>編集</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="reflist" style="list-style-type: decimal;"> <ol class="references"> <li id="cite_note-Voris2020-1">^ <a href="#cite_ref-Voris2020_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Voris2020_1-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Voris2020_1-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-Voris2020_1-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-Voris2020_1-4"><sup><i><b>e</b></i></sup></a> <span class="reference-text"><cite style="font-style:normal" class="citation journal">Voris, Jared T.; Therrien, Francois; Zelenitzky, Darla K.; Brown, Caleb M. (2020). “A new tyrannosaurine (Theropoda:Tyrannosauridae) from the Campanian Foremost Formation of Alberta, Canada, provides insight into the evolution and biogeography of tyrannosaurids”. <i><a href="https://en.wikipedia.org/wiki/Cretaceous_Research" class="extiw" title="w:Cretaceous Research">Cretaceous Research</a></i> <b>110</b>: 104388. <a href="/wiki/Doi_(%E8%AD%98%E5%88%A5%E5%AD%90)" class="mw-redirect" title="Doi (識別子)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.cretres.2020.104388">10.1016/j.cretres.2020.104388</a>.</cite><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=A+new+tyrannosaurine+%28Theropoda%3ATyrannosauridae%29+from+the+Campanian+Foremost+Formation+of+Alberta%2C+Canada%2C+provides+insight+into+the+evolution+and+biogeography+of+tyrannosaurids&rft.jtitle=%5B%5Bw%3ACretaceous+Research%7CCretaceous+Research%5D%5D&rft.aulast=Voris&rft.aufirst=Jared+T.&rft.au=Voris%2C%26%2332%3BJared+T.&rft.au=Therrien%2C%26%2332%3BFrancois&rft.au=Zelenitzky%2C%26%2332%3BDarla+K.&rft.au=Brown%2C%26%2332%3BCaleb+M.&rft.date=2020&rft.volume=110&rft.pages=104388&rft_id=info:doi/10.1016%2Fj.cretres.2020.104388&rfr_id=info:sid/ja.wikipedia.org:%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9"><span style="display: none;"> </span></span></span> </li> <li id="cite_note-2"><b><a href="#cite_ref-2">^</a></b> <span class="reference-text"><cite class="citation web" style="font-style:normal">“<a rel="nofollow" class="external text" href="http://www.oed.com/view/Entry/200131">Thanatos, n.</a>”. <i>OED Online</i>.  Oxford University Press (September 2014). <span title="">18 November 2014</span>閲覧。</cite></span> </li> <li id="cite_note-講談社-3">^ <a href="#cite_ref-講談社_3-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-講談社_3-1"><sup><i><b>b</b></i></sup></a> <span class="reference-text"><cite class="citation web" style="font-style:normal"><a href="/wiki/%E5%B0%8F%E6%9E%97%E5%BF%AB%E6%AC%A1" title="小林快次">小林快次</a> (2021年3月10日). “<a rel="nofollow" class="external text" href="https://gendai.media/articles/-/80961?page=2">50年ぶりの新種発見…カナダにはいつからティラノ軍団が棲息していたのか？</a>”.  <a href="/wiki/%E8%AC%9B%E8%AB%87%E7%A4%BE" title="講談社">講談社</a>. <span title="">2021年3月10日</span>閲覧。</cite></span> </li> <li id="cite_note-4"><b><a href="#cite_ref-4">^</a></b> <span class="reference-text"><cite style="font-style:normal" class="citation news">Black, Riley (February 10, 2020). <a rel="nofollow" class="external text" href="https://www.smithsonianmag.com/science-nature/newly-discovered-tyrannosaur-was-key-rise-giant-meat-eaters-180974156/">“Newly Discovered Tyrannosaur Was Key to the Rise of Giant Meat-Eaters”</a> (英語). <i><a href="https://en.wikipedia.org/wiki/Smithsonian_(magazine)" class="extiw" title="w:Smithsonian (magazine)">Smithsonian</a></i><span style="display:none;">. <a rel="nofollow" class="external free" href="https://www.smithsonianmag.com/science-nature/newly-discovered-tyrannosaur-was-key-rise-giant-meat-eaters-180974156/">https://www.smithsonianmag.com/science-nature/newly-discovered-tyrannosaur-was-key-rise-giant-meat-eaters-180974156/</a></span> <span class="reference-accessdate"><span title="">2020年2月11日</span>閲覧。</span></cite><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=Newly+Discovered+Tyrannosaur+Was+Key+to+the+Rise+of+Giant+Meat-Eaters&rft.jtitle=%5B%5Bw%3ASmithsonian+%28magazine%29%7CSmithsonian%5D%5D&rft.aulast=Black&rft.aufirst=Riley&rft.au=Black%2C%26%2332%3BRiley&rft.date=February+10%2C+2020&rft_id=https%3A%2F%2Fwww.smithsonianmag.com%2Fscience-nature%2Fnewly-discovered-tyrannosaur-was-key-rise-giant-meat-eaters-180974156%2F&rfr_id=info:sid/ja.wikipedia.org:%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9"><span style="display: none;"> </span></span></span> </li> <li id="cite_note-yun2020-5"><b><a href="#cite_ref-yun2020_5-0">^</a></b> <span class="reference-text"><cite style="font-style:normal" class="citation journal">Chan-gyu Yun (2020). <a rel="nofollow" class="external text" href="https://www.researchgate.net/publication/344187485_A_Subadult_Frontal_of_Daspletosaurus_torosus_Theropoda_Tyrannosauridae_from_the_Late_Cretaceous_of_Alberta_Canada_with_Implications_for_Tyrannosaurid_Ontogeny_and_Taxonomy">“A Subadult Frontal of <i>Daspletosaurus torosus</i> (Theropoda: Tyrannosauridae) from the Late Cretaceous of Alberta, Canada with Implications for Tyrannosaurid Ontogeny and Taxonomy”</a>. <i>PalArch's Journal of Vertebrate Palaeontology</i> <b>17</b>: 1–13<span style="display:none;">. <a rel="nofollow" class="external free" href="https://www.researchgate.net/publication/344187485_A_Subadult_Frontal_of_Daspletosaurus_torosus_Theropoda_Tyrannosauridae_from_the_Late_Cretaceous_of_Alberta_Canada_with_Implications_for_Tyrannosaurid_Ontogeny_and_Taxonomy">https://www.researchgate.net/publication/344187485_A_Subadult_Frontal_of_Daspletosaurus_torosus_Theropoda_Tyrannosauridae_from_the_Late_Cretaceous_of_Alberta_Canada_with_Implications_for_Tyrannosaurid_Ontogeny_and_Taxonomy</a></span>.</cite><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=A+Subadult+Frontal+of+%27%27Daspletosaurus+torosus%27%27+%28Theropoda%3A+Tyrannosauridae%29+from+the+Late+Cretaceous+of+Alberta%2C+Canada+with+Implications+for+Tyrannosaurid+Ontogeny+and+Taxonomy&rft.jtitle=PalArch%27s+Journal+of+Vertebrate+Palaeontology&rft.aulast=Chan-gyu+Yun&rft.au=Chan-gyu+Yun&rft.date=2020&rft.volume=17&rft.pages=1%E2%80%9313&rft_id=https%3A%2F%2Fwww.researchgate.net%2Fpublication%2F344187485_A_Subadult_Frontal_of_Daspletosaurus_torosus_Theropoda_Tyrannosauridae_from_the_Late_Cretaceous_of_Alberta_Canada_with_Implications_for_Tyrannosaurid_Ontogeny_and_Taxonomy&rfr_id=info:sid/ja.wikipedia.org:%E3%82%BF%E3%83%8A%E3%83%88%E3%83%86%E3%83%AA%E3%82%B9%E3%83%86%E3%82%B9"><span style="display: none;"> </span></span></span> </li> </ol></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" 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