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class="mw-page-title-main">Substantia gelatinosa of Rolando</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. 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class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Sostanza_gelatinosa_di_Rolando" title="Sostanza gelatinosa di Rolando – Italian" lang="it" hreflang="it" data-title="Sostanza gelatinosa di Rolando" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Subst%C3%A2ncia_gelatinosa_de_Rolando" title="Substância gelatinosa de Rolando – Portuguese" lang="pt" hreflang="pt" data-title="Substância gelatinosa de Rolando" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/Rolandova_%C5%BEelatinozna_supstanca" title="Rolandova želatinozna supstanca – Serbian" lang="sr" hreflang="sr" data-title="Rolandova želatinozna supstanca" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q17137094#sitelinks-wikipedia" title="Edit interlanguage links" class="wbc-editpage">Edit links</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Namespaces"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Substantia_gelatinosa_of_Rolando" title="View the content 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</div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">"Substantia gelatinosa" redirects here. For the substantia gelatinosa centralis, see <a href="/wiki/Central_gelatinous_substance_of_spinal_cord" class="mw-redirect" title="Central gelatinous substance of spinal cord">Central gelatinous substance of spinal cord</a>.</div> <style data-mw-deduplicate="TemplateStyles:r1257001546">.mw-parser-output .infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox-full-data:not(.notheme)>div:not(.notheme)[style]{background:#1f1f23!important;color:#f8f9fa}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox-full-data:not(.notheme) div:not(.notheme){background:#1f1f23!important;color:#f8f9fa}}@media(min-width:640px){body.skin--responsive .mw-parser-output .infobox-table{display:table!important}body.skin--responsive .mw-parser-output .infobox-table>caption{display:table-caption!important}body.skin--responsive .mw-parser-output .infobox-table>tbody{display:table-row-group}body.skin--responsive .mw-parser-output .infobox-table tr{display:table-row!important}body.skin--responsive .mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox"><tbody><tr><th colspan="2" class="infobox-above" style="background-color:dimgray; color: white">Substantia gelatinosa of Rolando</th></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Medulla_spinalis_-_Substantia_grisea_-_English.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c0/Medulla_spinalis_-_Substantia_grisea_-_English.svg/250px-Medulla_spinalis_-_Substantia_grisea_-_English.svg.png" decoding="async" width="250" height="184" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c0/Medulla_spinalis_-_Substantia_grisea_-_English.svg/375px-Medulla_spinalis_-_Substantia_grisea_-_English.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c0/Medulla_spinalis_-_Substantia_grisea_-_English.svg/500px-Medulla_spinalis_-_Substantia_grisea_-_English.svg.png 2x" data-file-width="700" data-file-height="514" /></a></span><div class="infobox-caption">Substantia gelatinosa of Rolando is <a href="/wiki/Rexed_lamina" class="mw-redirect" title="Rexed lamina">Rexed lamina</a> II, labeled at upper left.</div></td></tr><tr><th colspan="2" class="infobox-header" style="background-color: #efefef">Details</th></tr><tr><th colspan="2" class="infobox-header" style="background-color: #efefef">Identifiers</th></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/Latin" title="Latin">Latin</a></th><td class="infobox-data"><i>substantia gelatinosa cornu posterioris medullae spinalis</i></td></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/Medical_Subject_Headings" title="Medical Subject Headings">MeSH</a></th><td class="infobox-data"><a rel="nofollow" class="external text" href="https://meshb.nlm.nih.gov/record/ui?ui=D013376">D013376</a></td></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/Terminologia_Anatomica" title="Terminologia Anatomica">TA98</a></th><td class="infobox-data"><a rel="nofollow" class="external text" href="https://ifaa.unifr.ch/Public/EntryPage/TA98%20Tree/Entity%20TA98%20EN/14.1.02.119%20Entity%20TA98%20EN.htm">A14.1.02.119</a></td></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/Terminologia_Anatomica" title="Terminologia Anatomica">TA2</a></th><td class="infobox-data"><a rel="nofollow" class="external text" href="https://ta2viewer.openanatomy.org/?id=6067">6067</a></td></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/Foundational_Model_of_Anatomy" title="Foundational Model of Anatomy">FMA</a></th><td class="infobox-data"><a rel="nofollow" class="external text" href="https://bioportal.bioontology.org/ontologies/FMA/?p=classes&conceptid=http%3A%2F%2Fpurl.org%2Fsig%2Font%2Ffma%2Ffma74019">74019</a></td></tr><tr><td colspan="2" class="infobox-below"><a href="/wiki/Anatomical_terminology" title="Anatomical terminology">Anatomical terminology</a><div style="text-align: right;"><small class="noprint">[<a href="https://www.wikidata.org/wiki/Q17137094" class="extiw" title="d:Q17137094">edit on Wikidata</a>]</small></div></td></tr></tbody></table> <p>The apex of the <a href="/wiki/Posterior_grey_column" class="mw-redirect" title="Posterior grey column">posterior grey column</a>, one of the three <a href="/wiki/Grey_column" class="mw-redirect" title="Grey column">grey columns</a> of the <a href="/wiki/Spinal_cord" title="Spinal cord">spinal cord</a>, is capped by a V-shaped or crescentic mass of translucent, gelatinous neuroglia, termed the <b>substantia gelatinosa of Rolando</b> (or <b>SGR</b>) (or <b>gelatinous substance of posterior horn of spinal cord</b>), which contains both <a href="/wiki/Neuroglia" class="mw-redirect" title="Neuroglia">neuroglia</a> cells, and small <a href="/wiki/Neuron" title="Neuron">neurons</a>. The gelatinous appearance is due to an abundance of <a href="/wiki/Neuropil" title="Neuropil">neuropil</a> with a very low concentration of myelinated fibers.<sup id="cite_ref-Sheikh_1-0" class="reference"><a href="#cite_note-Sheikh-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:0_2-0" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> It extends the entire length of the spinal cord and into the <a href="/wiki/Medulla_oblongata" title="Medulla oblongata">medulla oblongata</a> where it becomes the <a href="/wiki/Spinal_trigeminal_nucleus" title="Spinal trigeminal nucleus">spinal trigeminal nucleus</a>. </p><p>It is named after <a href="/wiki/Luigi_Rolando" title="Luigi Rolando">Luigi Rolando</a>. </p><p>It corresponds to <a href="/wiki/Rexed_lamina" class="mw-redirect" title="Rexed lamina">Rexed lamina</a> II.<sup id="cite_ref-pmid10691235_3-0" class="reference"><a href="#cite_note-pmid10691235-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-urlSpringerLink_-_Journal_Article_4-0" class="reference"><a href="#cite_note-urlSpringerLink_-_Journal_Article-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Structure">Structure</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Substantia_gelatinosa_of_Rolando&action=edit&section=1" title="Edit section: Structure"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The SGR, or <a href="/wiki/Rexed_laminae" title="Rexed laminae">lamina II</a>, is composed of an outer lamina II and an inner lamina II.<sup id="cite_ref-:0_2-1" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> In rodents, the inner lamina II is divided into a dorsal and ventral inner lamina II. The distinction between these laminae lies in the areas of the spinal cord that send information to and from the laminae (input and output projections).<sup id="cite_ref-:0_2-2" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p><p>The cell types within the SGR include islet cells, central cells, stalked or large vertical cells, small vertical cells, and radial cells. The islet cells and small vertical cells are primarily <a href="/wiki/GABAergic" title="GABAergic">GABAergic</a>, while the large vertical cells and radial cells are primarily <a href="/wiki/Glutamatergic" title="Glutamatergic">glutamatergic</a>. The descriptors GABAergic and glutamatergic refer to the neurotransmitter (GABA and glutamate, respectively) that the cell releases. Typically, the release of GABA from one cell causes the next cell to stop firing. The release of glutamate typically causes the next cell to depolarize and fire. Central cells can be either glutamatergic or GABAergic. These cells synapse on each other to modulate pain signaling through the release of these different neurotransmitters and various neuropeptides.<sup id="cite_ref-:0_2-3" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p><p>The cells in the SGR receive input from each other and primary afferent neurons and project outwards to other cells within the lamina. Complex circuits of excitation and inhibition lead to transmission and inhibition of pain signals through the spinal cord to the thalamus.<sup id="cite_ref-:0_2-4" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Function">Function</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Substantia_gelatinosa_of_Rolando&action=edit&section=2" title="Edit section: Function"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">See also: <a href="/wiki/Gate_control_theory" title="Gate control theory">Gate control theory</a></div> <p>The substantia gelatinosa is one point (the <a href="/wiki/Nucleus_proprius" class="mw-redirect" title="Nucleus proprius">nucleus proprius</a> being the other) where first order neurons of the <a href="/wiki/Spinothalamic_tract" title="Spinothalamic tract">spinothalamic tract</a> synapse. </p><p>Many <a href="/wiki/%CE%9C-opioid_receptor" title="Μ-opioid receptor">μ</a> and <a href="/wiki/%CE%9A-opioid_receptor" title="Κ-opioid receptor">κ-opioid receptors</a>, <a href="/wiki/Presynaptic" class="mw-redirect" title="Presynaptic">presynaptic</a> and <a href="/wiki/Postsynaptic" class="mw-redirect" title="Postsynaptic">postsynaptic</a>, are found on these nerve cells; they can be targeted to manage pain of distal origin. For instance, neuraxial administration of opioids results in analgesia primarily by action in the dorsal horn of the spinal cord in the substantia gelatinosa where they inhibit release of excitatory neurotransmitters such as substance P and glutamate and inhibit afferent neural transmission to the brain from incoming peripheral pain neurons via hyperpolarization of postsynaptic neurons. </p><p><a href="/wiki/C_fiber" class="mw-redirect" title="C fiber">C fibers</a> terminate at this layer. Thus, the cell bodies located here are part of the neural pathway conveying slowly conducting, poorly localized <a href="/wiki/Pain" title="Pain">pain</a> sensation. However, some <a href="/wiki/A_delta_fiber" class="mw-redirect" title="A delta fiber">A delta fibers</a> (carrying fast, localized pain sensation) also terminate in the substantia gelatinosa, mostly via axons passing through this area to the <a href="/wiki/Nucleus_proprius" class="mw-redirect" title="Nucleus proprius">nucleus proprius</a>. Thus, there is cross talk between the two pain pathways. </p><p>C fibers carrying information about pain and temperature synapse in outer lamina II and dorsal inner lamina II and release glutamate to excite neurons in these regions. Some C fibers also release <a href="/wiki/Brain-derived_neurotrophic_factor" title="Brain-derived neurotrophic factor">BDNF</a>, which can be either excitatory or inhibitory, sometimes depending on the characteristics of the post-synaptic neuron. These fibers are part of a pathway which may be implicated in central sensitization in chronic pain conditions. Fibers synapsing on these laminae that release peptides <a href="/wiki/Somatostatin" title="Somatostatin">SST</a> and <a href="/wiki/Glial_cell_line-derived_neurotrophic_factor" title="Glial cell line-derived neurotrophic factor">GDNF</a> may be part of a pathway that inhibits pain signaling.<sup id="cite_ref-:0_2-5" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p><p>Some of the SGR projects to the <a href="/wiki/Marginal_nucleus_of_the_spinal_cord" class="mw-redirect" title="Marginal nucleus of the spinal cord">marginal nucleus of the spinal cord</a>, or lamina I, and laminae III-V. Most of these projections are excitatory.<sup id="cite_ref-:0_2-6" class="reference"><a href="#cite_note-:0-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Substantia_gelatinosa_of_Rolando&action=edit&section=3" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><span class="noviewer" typeof="mw:File"><span><img alt="Public domain" src="//upload.wikimedia.org/wikipedia/en/thumb/6/62/PD-icon.svg/12px-PD-icon.svg.png" decoding="async" width="12" height="12" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/6/62/PD-icon.svg/18px-PD-icon.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/6/62/PD-icon.svg/24px-PD-icon.svg.png 2x" data-file-width="196" data-file-height="196" /></span></span> <i>This article incorporates text in the <a href="/wiki/Public_domain" title="Public domain">public domain</a> from <a rel="nofollow" class="external text" href="https://archive.org/stream/anatomyofhumanbo1918gray#page/753/mode/2up">page 753</a> of the 20th edition of</i> <a href="/wiki/Gray%27s_Anatomy" title="Gray's Anatomy">Gray's Anatomy</a> <i>(1918)</i> </p> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-Sheikh-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-Sheikh_1-0">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFSheikhDua2024" class="citation web cs1">Sheikh, Nafiz K.; Dua, Anterpreet (2024). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/books/NBK551522/">"Neuroanatomy, Substantia Gelatinosa"</a>. <i>StatPearls</i>. 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a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Spinal_cord" title="Template:Spinal cord"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Spinal_cord" title="Template talk:Spinal cord"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a 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title="Meninges">Meninges</a></li> <li><a href="/wiki/Central_canal" title="Central canal">Central canal</a> <ul><li><a href="/wiki/Terminal_ventricle" class="mw-redirect" title="Terminal ventricle">Terminal ventricle</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Grey_column" class="mw-redirect" title="Grey column">Grey columns</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Posterior_grey_column" class="mw-redirect" title="Posterior grey column">Posterior grey column</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Marginal_nucleus_of_spinal_cord" title="Marginal nucleus of spinal cord">Marginal nucleus</a></li> <li><a class="mw-selflink selflink">Substantia gelatinosa of Rolando</a></li> <li><a href="/wiki/Nucleus_proprius_of_spinal_cord" title="Nucleus proprius of spinal cord">Nucleus proprius</a></li> <li><a href="/wiki/Rexed_laminae" title="Rexed laminae">Rexed lamina V</a></li> <li><a href="/wiki/Rexed_laminae" title="Rexed laminae">Rexed lamina VI</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Lateral_grey_column" title="Lateral grey column">Lateral grey column</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Intermediolateral_nucleus" title="Intermediolateral nucleus">Intermediolateral nucleus</a></li> <li><a href="/wiki/Posterior_thoracic_nucleus" title="Posterior thoracic nucleus">Posterior thoracic nucleus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Anterior_grey_column" class="mw-redirect" title="Anterior grey column">Anterior grey column</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Interneuron" title="Interneuron">Interneuron</a></li> <li><a href="/wiki/Alpha_motor_neuron" title="Alpha motor neuron">Alpha motor neuron</a> <ul><li><a href="/wiki/Onuf%27s_nucleus" title="Onuf's nucleus">Onuf's nucleus</a></li></ul></li> <li><a href="/wiki/Gamma_motor_neuron" title="Gamma motor neuron">Gamma motor neuron</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Rexed_laminae" title="Rexed laminae">Rexed laminae</a></li> <li><a href="/wiki/Central_gelatinous_substance_of_spinal_cord" class="mw-redirect" title="Central gelatinous substance of spinal cord">Central gelatinous substance</a></li> <li><a href="/wiki/Gray_commissure" class="mw-redirect" title="Gray commissure">Gray commissure</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/White_matter" title="White matter">White matter</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Sensory</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Posterior_funiculus" class="mw-redirect" title="Posterior funiculus">Posterior</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Posterior_column-medial_lemniscus_pathway" class="mw-redirect" title="Posterior column-medial lemniscus pathway">Posterior column-medial lemniscus pathway</a></li> <li><a href="/wiki/Gracile_fasciculus" class="mw-redirect" title="Gracile fasciculus">Gracile</a></li> <li><a href="/wiki/Cuneate_fasciculus" class="mw-redirect" title="Cuneate fasciculus">Cuneate</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Lateral_funiculus" title="Lateral funiculus">Lateral</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Spinocerebellar_tract" class="mw-redirect" title="Spinocerebellar tract">Spinocerebellar</a> <ul><li><a href="/wiki/Dorsal_spinocerebellar_tract" class="mw-redirect" title="Dorsal spinocerebellar tract">dorsal</a></li> <li><a href="/wiki/Ventral_spinocerebellar_tract" class="mw-redirect" title="Ventral spinocerebellar tract">ventral</a></li></ul></li> <li><a href="/wiki/Spinothalamic_tract" title="Spinothalamic tract">Spinothalamic</a> <ul><li><a href="/wiki/Lateral_spinothalamic_tract" class="mw-redirect" title="Lateral spinothalamic tract">lateral</a></li> <li><a href="/wiki/Anterior_spinothalamic_tract" class="mw-redirect" title="Anterior spinothalamic tract">anterior</a></li></ul></li> <li><a href="/wiki/Posterolateral_tract" title="Posterolateral tract">Posterolateral</a></li> <li><a href="/wiki/Spinotectal_tract" class="mw-redirect" title="Spinotectal tract">Spinotectal</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Anterior_funiculus" title="Anterior funiculus">Anterior</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Spinoreticular_tract" title="Spinoreticular tract">Spinoreticular tract</a></li> <li><a href="/wiki/Spino-olivary_tract" title="Spino-olivary tract">Spino-olivary tract</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Motor</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Lateral_funiculus" title="Lateral funiculus">Lateral</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pyramidal_tracts" title="Pyramidal tracts">Corticospinal</a> <ul><li><a href="/wiki/Lateral_corticospinal_tract" title="Lateral corticospinal tract">Lateral</a></li></ul></li> <li><a href="/wiki/Extrapyramidal_system" title="Extrapyramidal system">Extrapyramidal</a> <ul><li><a href="/wiki/Rubrospinal_tract" title="Rubrospinal tract">Rubrospinal</a></li> <li><a href="/wiki/Lateral_raphespinal_tract" class="mw-redirect" title="Lateral raphespinal tract">Raphespinal</a></li> <li><a href="/wiki/Reticular_formation#Descending_reticulospinal_tracts" title="Reticular formation">Reticulospinal</a></li> <li><a href="/wiki/Hypothalamospinal_tract" title="Hypothalamospinal tract">Hypothalamospinal</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Anterior_funiculus" title="Anterior funiculus">Anterior</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Corticospinal <ul><li><a href="/wiki/Anterior_corticospinal_tract" title="Anterior corticospinal tract">Anterior</a></li></ul></li> <li>Extrapyramidal <ul><li><a href="/wiki/Vestibulospinal_tract" title="Vestibulospinal tract">Vestibulospinal</a></li> <li><a href="/wiki/Reticular_formation#Descending_reticulospinal_tracts" title="Reticular formation">Reticulospinal</a></li> <li><a href="/wiki/Tectospinal_tract" title="Tectospinal tract">Tectospinal</a></li> <li><a href="/wiki/Anterior_raphespinal_tract" class="mw-redirect" title="Anterior raphespinal tract">Raphespinal</a></li> <li><a href="/wiki/Olivospinal_tract" title="Olivospinal tract">Olivospinal</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Both</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Anterior_white_commissure" title="Anterior white commissure">Anterior white commissure</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">External features</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Ventral <ul><li><a href="/wiki/Anterior_median_fissure_of_spinal_cord" title="Anterior median fissure of spinal cord">Anterior median fissure</a></li> <li><a href="/wiki/Anterolateral_sulcus_of_spinal_cord" title="Anterolateral sulcus of spinal cord">Anterolateral sulcus</a></li></ul></li> <li>Dorsal <ul><li><a href="/wiki/Posterior_median_sulcus_of_spinal_cord" title="Posterior median sulcus of spinal cord">Posterior median sulcus</a></li> <li><a href="/wiki/Posterolateral_sulcus_of_spinal_cord" title="Posterolateral sulcus of spinal cord">Posterolateral sulcus</a></li></ul></li></ul> </div></td></tr></tbody></table></div> <style data-mw-deduplicate="TemplateStyles:r1130092004">.mw-parser-output .portal-bar{font-size:88%;font-weight:bold;display:flex;justify-content:center;align-items:baseline}.mw-parser-output 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