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Lystrosaurus - Wikipedia
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</button> <ul id="toc-History_of_discovery-sublist" class="vector-toc-list"> <li id="toc-Plate_tectonics" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Plate_tectonics"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.1</span> <span>Plate tectonics</span> </div> </a> <ul id="toc-Plate_tectonics-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Distribution_and_species" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Distribution_and_species"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.2</span> <span>Distribution and species</span> </div> </a> <ul id="toc-Distribution_and_species-sublist" class="vector-toc-list"> <li id="toc-Species_found_in_Africa" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Species_found_in_Africa"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.2.1</span> <span>Species found in Africa</span> </div> </a> <ul id="toc-Species_found_in_Africa-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Other_species" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Other_species"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.2.2</span> <span>Other species</span> </div> </a> <ul id="toc-Other_species-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> </ul> </li> <li id="toc-Description" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Description"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Description</span> </div> </a> <ul id="toc-Description-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Paleoecology" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Paleoecology"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Paleoecology</span> </div> </a> <button aria-controls="toc-Paleoecology-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Paleoecology subsection</span> </button> <ul id="toc-Paleoecology-sublist" class="vector-toc-list"> <li id="toc-Dominance_of_the_Early_Triassic" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Dominance_of_the_Early_Triassic"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.1</span> <span>Dominance of the Early Triassic</span> </div> </a> <ul id="toc-Dominance_of_the_Early_Triassic-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-See_also" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#See_also"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>See also</span> </div> </a> <ul id="toc-See_also-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Toggle the table of contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><i>Lystrosaurus</i></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 33 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-33" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">33 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-af mw-list-item"><a href="https://af.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – Afrikaans" lang="af" hreflang="af" data-title="Lystrosaurus" data-language-autonym="Afrikaans" data-language-local-name="Afrikaans" class="interlanguage-link-target"><span>Afrikaans</span></a></li><li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B3%D8%AD%D9%84%D9%8A%D8%A9_%D8%A7%D9%84%D9%85%D8%AC%D8%B1%D9%81%D8%A9" title="سحلية المجرفة – Arabic" lang="ar" hreflang="ar" data-title="سحلية المجرفة" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-br mw-list-item"><a href="https://br.wikipedia.org/wiki/Listrosaor" title="Listrosaor – Breton" lang="br" hreflang="br" data-title="Listrosaor" data-language-autonym="Brezhoneg" data-language-local-name="Breton" class="interlanguage-link-target"><span>Brezhoneg</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Listrosaure" title="Listrosaure – Catalan" lang="ca" hreflang="ca" data-title="Listrosaure" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – Czech" lang="cs" hreflang="cs" data-title="Lystrosaurus" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – German" lang="de" hreflang="de" data-title="Lystrosaurus" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%9B%CF%85%CF%83%CF%84%CF%81%CF%8C%CF%83%CE%B1%CF%85%CF%81%CE%BF%CF%82" title="Λυστρόσαυρος – Greek" lang="el" hreflang="el" data-title="Λυστρόσαυρος" data-language-autonym="Ελληνικά" data-language-local-name="Greek" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – Spanish" lang="es" hreflang="es" data-title="Lystrosaurus" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – Esperanto" lang="eo" hreflang="eo" data-title="Lystrosaurus" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%A8%DB%8C%D9%84%E2%80%8C%D8%AE%D8%B2%D9%86%D8%AF%D9%87%E2%80%8C%D9%87%D8%A7" title="بیلخزندهها – Persian" lang="fa" hreflang="fa" data-title="بیلخزندهها" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – French" lang="fr" hreflang="fr" data-title="Lystrosaurus" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EB%A6%AC%EC%8A%A4%ED%8A%B8%EB%A1%9C%EC%82%AC%EC%9A%B0%EB%A3%A8%EC%8A%A4" title="리스트로사우루스 – Korean" lang="ko" hreflang="ko" data-title="리스트로사우루스" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – Indonesian" lang="id" hreflang="id" data-title="Lystrosaurus" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – Italian" lang="it" hreflang="it" data-title="Lystrosaurus" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%9C%D7%99%D7%A1%D7%98%D7%A8%D7%95%D7%96%D7%90%D7%95%D7%A8%D7%95%D7%A1" title="ליסטרוזאורוס – Hebrew" lang="he" hreflang="he" data-title="ליסטרוזאורוס" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Listrozauras" title="Listrozauras – Lithuanian" lang="lt" hreflang="lt" data-title="Listrozauras" data-language-autonym="Lietuvių" data-language-local-name="Lithuanian" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-olo mw-list-item"><a href="https://olo.wikipedia.org/wiki/Listozauru" title="Listozauru – Livvi-Karelian" lang="olo" hreflang="olo" data-title="Listozauru" data-language-autonym="Livvinkarjala" data-language-local-name="Livvi-Karelian" class="interlanguage-link-target"><span>Livvinkarjala</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – Hungarian" lang="hu" hreflang="hu" data-title="Lystrosaurus" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – Dutch" lang="nl" hreflang="nl" data-title="Lystrosaurus" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%83%AA%E3%82%B9%E3%83%88%E3%83%AD%E3%82%B5%E3%82%A6%E3%83%AB%E3%82%B9" title="リストロサウルス – Japanese" lang="ja" hreflang="ja" data-title="リストロサウルス" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Lystrozaur" title="Lystrozaur – Polish" lang="pl" hreflang="pl" data-title="Lystrozaur" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – Portuguese" lang="pt" hreflang="pt" data-title="Lystrosaurus" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9B%D0%B8%D1%81%D1%82%D1%80%D0%BE%D0%B7%D0%B0%D0%B2%D1%80%D1%8B" title="Листрозавры – Russian" lang="ru" hreflang="ru" data-title="Листрозавры" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sco mw-list-item"><a href="https://sco.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – Scots" lang="sco" hreflang="sco" data-title="Lystrosaurus" data-language-autonym="Scots" data-language-local-name="Scots" class="interlanguage-link-target"><span>Scots</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – Simple English" lang="en-simple" hreflang="en-simple" data-title="Lystrosaurus" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – Slovak" lang="sk" hreflang="sk" data-title="Lystrosaurus" data-language-autonym="Slovenčina" data-language-local-name="Slovak" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%9B%D0%B8%D1%81%D1%82%D1%80%D0%BE%D1%81%D0%B0%D1%83%D1%80%D1%83%D1%81" title="Листросаурус – Serbian" lang="sr" hreflang="sr" data-title="Листросаурус" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – Serbo-Croatian" lang="sh" hreflang="sh" data-title="Lystrosaurus" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="Serbo-Croatian" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – Finnish" lang="fi" hreflang="fi" data-title="Lystrosaurus" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – Swedish" lang="sv" hreflang="sv" data-title="Lystrosaurus" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%9B%D1%96%D1%81%D1%82%D1%80%D0%BE%D0%B7%D0%B0%D0%B2%D1%80" title="Лістрозавр – Ukrainian" lang="uk" hreflang="uk" data-title="Лістрозавр" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Lystrosaurus" title="Lystrosaurus – Vietnamese" lang="vi" hreflang="vi" data-title="Lystrosaurus" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E6%B0%B4%E9%BE%8D%E7%8D%B8%E5%B1%AC" title="水龍獸屬 – Chinese" lang="zh" hreflang="zh" data-title="水龍獸屬" data-language-autonym="中文" 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id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Genus of Late Permian and Early Triassic dicynodont therapsids</div> <p class="mw-empty-elt"> </p> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><i>Lystrosaurus</i><br /><div style="font-size: 85%;">Temporal range: <a href="/wiki/Late_Permian" class="mw-redirect" title="Late Permian">Late Permian</a> (<a href="/wiki/Lopingian" title="Lopingian">Lopingian</a>)–<a href="/wiki/Early_Triassic" title="Early Triassic">Early Triassic</a> (<a href="/wiki/Olenekian" title="Olenekian">Olenekian</a>), <span class="noprint"><span style="display:inline-block;"></span><span style="display:inline-block;">255–248 <a href="/wiki/Megaannum" class="mw-redirect" title="Megaannum">Ma</a></span> <span style="display:inline-block;"></span><div id="Timeline-row" style="margin: 4px auto 0; clear:both; width:220px; padding:0px; height:18px; overflow:visible; white-space:nowrap; border:1px #666; border-style:solid none; position:relative; z-index:0; font-size:97%;"> <div style="position:absolute; height:100%; left:0px; width:207.23076923077px; padding-left:5px; text-align:left; background-color:rgb(254,217,106); background-image: linear-gradient(to right, rgba(255,255,255,1), rgba(254,217,106,1) 15%, rgba(254,217,106,1));"><a href="/wiki/Precambrian" title="Precambrian">PreꞒ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,160,86); left:37.636923076923px; width:18.073846153846px;"><a href="/wiki/Cambrian" title="Cambrian">Ꞓ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(0,146,112); left:55.710769230769px; width:14.08px;"><a href="/wiki/Ordovician" title="Ordovician">O</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(179,225,182); left:69.790769230769px; width:8.3261538461539px;"><a href="/wiki/Silurian" title="Silurian">S</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(203,140,55); left:78.116923076923px; width:20.409230769231px;"><a href="/wiki/Devonian" title="Devonian">D</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(103,165,153); left:98.526153846154px; width:20.307692307692px;"><a href="/wiki/Carboniferous" title="Carboniferous">C</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(240,64,40); left:118.83384615385px; width:15.907015384615px;"><a href="/wiki/Permian" title="Permian">P</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(129,43,146); left:134.74086153846px; width:17.092984615385px;"><a href="/wiki/Triassic" title="Triassic">T</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(52,178,201); left:151.83384615385px; width:19.089230769231px;"><a href="/wiki/Jurassic" title="Jurassic">J</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,198,78); left:170.92307692308px; width:26.738461538462px;"><a href="/wiki/Cretaceous" title="Cretaceous">K</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(253,154,82); left:197.66153846154px; width:14.543692307692px;"><a href="/wiki/Paleogene" title="Paleogene">Pg</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(255,230,25); left:212.20523076923px; width:6.9215384615385px;"><a href="/wiki/Neogene" title="Neogene">N</a></div> <div id="end-border" style="position:absolute; height:100%; background-color:#666; width:1px; left:219px"></div><div style="margin:0 auto; line-height:0; clear:both; width:220px; padding:0px; height:8px; overflow:visible; background-color:transparent; position:relative; top:-4px; z-index:100;"><div style="position:absolute; height:8px; left:133.69230769231px; width:2.3692307692308px; background-color:#360; opacity:0.42;"></div> <div style="position:absolute; height:8px; left:133.69230769231px; width:2.3692307692308px; background-color:#360; opacity:1;"></div> <div style="position:absolute; height:6px; top:1px; left:134.69230769231px; width:0.36923076923077px; background-color:#6c3;"></div> </div> </div></span></div> </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Lystrosaurus_hedini_IMG_4469.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/27/Lystrosaurus_hedini_IMG_4469.jpg/250px-Lystrosaurus_hedini_IMG_4469.jpg" decoding="async" width="250" height="167" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/27/Lystrosaurus_hedini_IMG_4469.jpg/375px-Lystrosaurus_hedini_IMG_4469.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/27/Lystrosaurus_hedini_IMG_4469.jpg/500px-Lystrosaurus_hedini_IMG_4469.jpg 2x" data-file-width="4416" data-file-height="2944" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%"><i>L. hedini</i> skeletal mount from the Natural history museum of <a href="/wiki/Z%C3%BCrich" class="mw-redirect" title="Zürich">Zürich</a> </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Lystrosaurus" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Chordate" title="Chordate">Chordata</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Synapsida" title="Synapsida">Synapsida</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Therapsida" title="Therapsida">Therapsida</a> </td></tr> <tr> <td>Suborder: </td> <td><span style="font-style:normal;font-weight:normal;">†</span><a href="/wiki/Anomodontia" title="Anomodontia">Anomodontia</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><span style="font-style:normal;font-weight:normal;">†</span><a href="/wiki/Dicynodontia" title="Dicynodontia">Dicynodontia</a> </td></tr> <tr> <td>Family: </td> <td><span style="font-style:normal;font-weight:normal;">†</span><a href="/wiki/Lystrosauridae" title="Lystrosauridae">Lystrosauridae</a> </td></tr> <tr> <td>Genus: </td> <td><span style="font-style:normal;font-weight:normal;">†</span><a class="mw-selflink selflink"><i>Lystrosaurus</i></a><br /><small><a href="/wiki/Edward_Drinker_Cope" title="Edward Drinker Cope">Cope</a>, <a href="/wiki/1870_in_paleontology" title="1870 in paleontology">1870</a></small> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">Species </th></tr> <tr> <td colspan="2" style="text-align: left"> <div class="collapsible-list mw-collapsible mw-collapsed" style="text-align: left;"> <div style="line-height: 1.6em; font-weight: bold;"><div><small>List</small></div></div> <ul class="mw-collapsible-content" style="margin-top: 0; margin-bottom: 0; line-height: inherit;"><li style="line-height: inherit; margin: 0"> <i>L. murrayi</i> <small>(<a href="/wiki/Thomas_Henry_Huxley" title="Thomas Henry Huxley">Huxley</a>, 1859) (<a href="/wiki/Type_species" title="Type species">type</a>)</small> </li><li style="line-height: inherit; margin: 0"> <i>L. declivis</i> <small>(<a href="/wiki/Richard_Owen" title="Richard Owen">Owen</a>, 1860)</small> </li><li style="line-height: inherit; margin: 0"> <i>L. curvatus</i> <small>(Owen, 1876)</small> </li><li style="line-height: inherit; margin: 0"> <i>L. maccaigi</i> <small><a href="/wiki/Harry_Seeley" title="Harry Seeley">Seeley</a>, 1898</small> </li></ul> </div> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><i><b>Lystrosaurus</b></i> (<span class="rt-commentedText nowrap"><span class="IPA nopopups noexcerpt" lang="en-fonipa"><a href="/wiki/Help:IPA/English" title="Help:IPA/English">/<span style="border-bottom:1px dotted"><span title="/ˌ/: secondary stress follows">ˌ</span><span title="'l' in 'lie'">l</span><span title="/ɪ/: 'i' in 'kit'">ɪ</span><span title="'s' in 'sigh'">s</span><span title="'t' in 'tie'">t</span><span title="'r' in 'rye'">r</span><span title="/oʊ/: 'o' in 'code'">oʊ</span><span title="/ˈ/: primary stress follows">ˈ</span><span title="'s' in 'sigh'">s</span><span title="/ɔːr/: 'ar' in 'war'">ɔːr</span><span title="/ə/: 'a' in 'about'">ə</span><span title="'s' in 'sigh'">s</span></span>/</a></span></span>; 'shovel lizard'; proper Greek is <span class="texhtml">λίστρον</span> <i>lístron</i> ‘tool for leveling or smoothing, shovel, spade, hoe’) is an extinct genus of <a href="/wiki/Herbivore" title="Herbivore">herbivorous</a> <a href="/wiki/Dicynodont" class="mw-redirect" title="Dicynodont">dicynodont</a> <a href="/wiki/Therapsid" class="mw-redirect" title="Therapsid">therapsids</a> from the <a href="/wiki/Lopingian" title="Lopingian">late Permian</a> and <a href="/wiki/Early_Triassic" title="Early Triassic">Early Triassic</a> <a href="/wiki/Epoch_(geology)" class="mw-redirect" title="Epoch (geology)">epochs</a> (around 248 million years ago). It lived in what is now <a href="/wiki/Antarctica" title="Antarctica">Antarctica</a>, <a href="/wiki/India" title="India">India</a>, <a href="/wiki/China" title="China">China</a>, <a href="/wiki/Mongolia" title="Mongolia">Mongolia</a>, <a href="/wiki/European_Russia" title="European Russia">European Russia</a> and <a href="/wiki/South_Africa" title="South Africa">South Africa</a>. Four to six species are currently recognized, although from the 1930s to 1970s the number of species was thought to be much higher. They ranged in size from that of a small dog to 8 feet (2.5 meters) long.<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p><p>As a dicynodont, <i>Lystrosaurus</i> had only two teeth (a pair of <a href="/wiki/Tusk" title="Tusk">tusk</a>-like <a href="/wiki/Canine_tooth" title="Canine tooth">canines</a>), and is thought to have had a horny beak that was used for biting off pieces of vegetation. <i>Lystrosaurus</i> was a heavily built, herbivorous animal. The structure of its shoulders and hip joints suggests that <i>Lystrosaurus</i> moved with a <a href="/wiki/Terrestrial_locomotion" title="Terrestrial locomotion">semi-sprawling gait</a>. The forelimbs were even more robust than the hindlimbs, and the animal is thought to have been a powerful digger that nested in burrows. </p><p><i>Lystrosaurus</i> survived the <a href="/wiki/Permian-Triassic_extinction" class="mw-redirect" title="Permian-Triassic extinction">Permian-Triassic extinction</a>, 252 million years ago. In the <a href="/wiki/Early_Triassic" title="Early Triassic">Early Triassic</a>, they were by far the most common terrestrial vertebrates, accounting for as many as 95% of the total individuals in some fossil beds.<sup id="cite_ref-Barendskraal_2-0" class="reference"><a href="#cite_note-Barendskraal-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> Researchers have offered various hypotheses for why <i>Lystrosaurus</i> survived the extinction event and thrived in the early Triassic. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="History_of_discovery">History of discovery</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Lystrosaurus&action=edit&section=1" title="Edit section: History of discovery"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure typeof="mw:File/Thumb"><a href="/wiki/File:Wegener_fossils-mapped.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/fc/Wegener_fossils-mapped.png/350px-Wegener_fossils-mapped.png" decoding="async" width="350" height="208" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/fc/Wegener_fossils-mapped.png/525px-Wegener_fossils-mapped.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/fc/Wegener_fossils-mapped.png/700px-Wegener_fossils-mapped.png 2x" data-file-width="2742" data-file-height="1628" /></a><figcaption>Map of <a href="/wiki/Pangea" class="mw-redirect" title="Pangea">Pangea</a> showing locations of <i>Lystrosaurus</i> remains as yellow disks. Distorted boundaries of modern continents shown as grey lines. (Distributions for lystrosaurs and three other Permian and Triassic fossil groups used as biogeographic evidence for <a href="/wiki/Continental_drift" title="Continental drift">continental drift</a> and certain <a href="/wiki/Land_bridge" title="Land bridge">land bridges</a>.)</figcaption></figure> <p>Dr. Elias Root Beadle, a Philadelphia missionary and avid fossil collector, discovered the first <i>Lystrosaurus</i> skull. Beadle wrote to the eminent paleontologist <a href="/wiki/Othniel_Charles_Marsh" title="Othniel Charles Marsh">Othniel Charles Marsh</a>, but received no reply. Marsh's rival, <a href="/wiki/Edward_Drinker_Cope" title="Edward Drinker Cope">Edward Drinker Cope</a>, was very interested in seeing the find, and described and named <i>Lystrosaurus</i> in the <i>Proceedings of the American Philosophical Society</i> in 1870.<sup id="cite_ref-wallace2000_3-0" class="reference"><a href="#cite_note-wallace2000-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Its name is derived from the <a href="/wiki/Ancient_Greek" title="Ancient Greek">Ancient Greek</a> words <i>listron</i>, "shovel", and <i>sauros</i>, "lizard".<sup id="cite_ref-Liddell1980_4-0" class="reference"><a href="#cite_note-Liddell1980-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Marsh belatedly purchased the skull in May 1871, although his interest in an already-described specimen was unclear; he may have wanted to carefully scrutinize Cope's description and illustration.<sup id="cite_ref-wallace2000_3-1" class="reference"><a href="#cite_note-wallace2000-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Plate_tectonics">Plate tectonics</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Lystrosaurus&action=edit&section=2" title="Edit section: Plate tectonics"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The discovery of <i>Lystrosaurus</i> fossils at Coalsack Bluff in the <a href="/wiki/Transantarctic_Mountains" title="Transantarctic Mountains">Transantarctic Mountains</a> by <a href="/wiki/Edwin_H._Colbert" title="Edwin H. Colbert">Edwin H. Colbert</a> and his team in 1969–1970 helped support the hypothesis of <a href="/wiki/Plate_tectonics" title="Plate tectonics">plate tectonics</a> and strengthen the theory, since <i>Lystrosaurus</i> had already been found in the lower Triassic of southern Africa as well as in India and China.<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Distribution_and_species">Distribution and species</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Lystrosaurus&action=edit&section=3" title="Edit section: Distribution and species"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><i>Lystrosaurus</i> fossils have been found in many <a href="/wiki/Late_Permian" class="mw-redirect" title="Late Permian">Late Permian</a> and <a href="/wiki/Early_Triassic" title="Early Triassic">Early Triassic</a> terrestrial <a href="/wiki/Bone_bed" title="Bone bed">bone beds</a>, most abundantly in Africa, and to a lesser extent in parts of what are now India, China, Mongolia, European Russia, and Antarctica (which was not over the South Pole at the time).<sup id="cite_ref-Surkov-Kalndze-Benton-2005_7-0" class="reference"><a href="#cite_note-Surkov-Kalndze-Benton-2005-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Species_found_in_Africa">Species found in Africa</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Lystrosaurus&action=edit&section=4" title="Edit section: Species found in Africa"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Lystrosaurus_murrayi_7.JPG" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/fe/Lystrosaurus_murrayi_7.JPG/220px-Lystrosaurus_murrayi_7.JPG" decoding="async" width="220" height="127" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/fe/Lystrosaurus_murrayi_7.JPG/330px-Lystrosaurus_murrayi_7.JPG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/fe/Lystrosaurus_murrayi_7.JPG/440px-Lystrosaurus_murrayi_7.JPG 2x" data-file-width="3152" data-file-height="1820" /></a><figcaption><i>L. murrayi</i> skeleton</figcaption></figure> <p>Most <i>Lystrosaurus</i> fossils have been found in the <a href="/wiki/Balfour_Formation" title="Balfour Formation">Balfour</a> and <a href="/wiki/Katberg_Formation" title="Katberg Formation">Katberg Formations</a> of the <a href="/wiki/Karoo_Basin" class="mw-redirect" title="Karoo Basin">Karoo basin</a> in <a href="/wiki/South_Africa" title="South Africa">South Africa</a>; these specimens offer the best prospects of identifying species because they are the most numerous and have been studied for the longest time. As so often with fossils, there is debate in the <a href="/wiki/Scientific_community" title="Scientific community">paleontological community</a> as to exactly how many species have been found in the Karoo basin. Studies from the 1930s to 1970s suggested a large number (23 in one case).<sup id="cite_ref-Botha-Smith-2007_8-0" class="reference"><a href="#cite_note-Botha-Smith-2007-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> However, by the 1980s and 1990s, only 6 species were recognized in the <a href="/wiki/Karoo" title="Karoo">Karoo</a>: <i>L. curvatus</i>, <i>L. platyceps</i>, <i>L. oviceps</i>, <i>L. maccaigi</i>, <i>L. murrayi</i>, and <i>L. declivis</i>. A study in 2011 reduced that number to four, treating the fossils previously labeled as <i>L. platyceps</i> and <i>L. oviceps</i> as members of <i>L. curvatus</i>.<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> </p><p><i>L. maccaigi</i> is the largest and apparently most specialized species, while <i>L. curvatus</i> was the least specialized. A <i>Lystrosaurus</i>-like fossil, <i>Kwazulusaurus shakai</i>, has also been found in South Africa. Although not assigned to the same <a href="/wiki/Genus" title="Genus">genus</a>, <i>K. shakai</i> is very similar to <i>L. curvatus</i>. Some paleontologists have therefore proposed that <i>K. shakai</i> was possibly an ancestor of or closely related to the ancestors of <i>L. curvatus</i>, while <i>L. maccaigi</i> arose from a different lineage.<sup id="cite_ref-Botha-Smith-2007_8-1" class="reference"><a href="#cite_note-Botha-Smith-2007-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> <i>L. maccaigi</i> is found only in sediments from the <a href="/wiki/Permian" title="Permian">Permian</a> period, and apparently did not survive the <a href="/wiki/Permian%E2%80%93Triassic_extinction_event" title="Permian–Triassic extinction event">Permian–Triassic extinction event</a>. Its specialized features and sudden appearance in the <a href="/wiki/Fossil_record" class="mw-redirect" title="Fossil record">fossil record</a> without an obvious ancestor may indicate that it immigrated into the Karoo from an area in which Late Permian sediments have not been found.<sup id="cite_ref-Botha-Smith-2007_8-2" class="reference"><a href="#cite_note-Botha-Smith-2007-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> <i>L. curvatus</i> is found in a relatively narrow band of sediments from shortly before and after the extinction, and can be used as an approximate marker for the boundary between the <a href="/wiki/Permian" title="Permian">Permian</a> and <a href="/wiki/Triassic" title="Triassic">Triassic</a> periods. A skull identified as <i>L. curvatus</i> has been found in late Permian sediments from <a href="/wiki/Zambia" title="Zambia">Zambia</a>. For many years it had been thought that there were no Permian specimens of <i>L. curvatus</i> in the Karoo, which led to suggestions that <i>L. curvatus</i> immigrated from Zambia into the Karoo. However, a re-examination of Permian specimens in the Karoo has identified some as <i>L. curvatus</i>, and there is no need to assume immigration.<sup id="cite_ref-Botha-Smith-2007_8-3" class="reference"><a href="#cite_note-Botha-Smith-2007-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p><p><i>L. murrayi</i> and <i>L. declivis</i> are found only in Triassic sediments.<sup id="cite_ref-Botha-Smith-2007_8-4" class="reference"><a href="#cite_note-Botha-Smith-2007-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Other_species">Other species</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Lystrosaurus&action=edit&section=5" title="Edit section: Other species"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><i>Lystrosaurus georgi</i> fossils have been found in the Earliest Triassic sediments of the <a href="/wiki/Moscow" title="Moscow">Moscow</a> Basin in Russia. It was probably closely related to the African <i>Lystrosaurus curvatus</i>,<sup id="cite_ref-Surkov-Kalndze-Benton-2005_7-1" class="reference"><a href="#cite_note-Surkov-Kalndze-Benton-2005-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> which is regarded as one of the least specialized species and has been found in very Late Permian and very Early Triassic sediments.<sup id="cite_ref-Botha-Smith-2007_8-5" class="reference"><a href="#cite_note-Botha-Smith-2007-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p><p><i>L. murrayi</i>, in addition to two undescribed species presently assigned to <i>L. curvatus</i> and <i>L. declivis</i>, is known from the Early Triassic <a href="/wiki/Panchet_Formation" title="Panchet Formation">Panchet Formation</a> of the <a href="/wiki/Damodar_River" title="Damodar River">Damodar Valley</a> and the <a href="/w/index.php?title=Kamthi_Formation&action=edit&redlink=1" class="new" title="Kamthi Formation (page does not exist)">Kamthi Formation</a> of the <a href="/wiki/Pranhita%E2%80%93Godavari_Basin" title="Pranhita–Godavari Basin">Pranhita-Godavari Basin</a> in <a href="/wiki/India" title="India">India</a>.<sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> Seven <i>Lystrosaurus</i> species have been described from the Early Triassic <a href="/w/index.php?title=Jiucaiyuan_Formation&action=edit&redlink=1" class="new" title="Jiucaiyuan Formation (page does not exist)">Jiucaiyuan</a>, <a href="/w/index.php?title=Guodikeng_Formation&action=edit&redlink=1" class="new" title="Guodikeng Formation (page does not exist)">Guodikeng</a> and <a href="/w/index.php?title=Wutonggou_Formation&action=edit&redlink=1" class="new" title="Wutonggou Formation (page does not exist)">Wutonggou</a> formations of the <a href="/wiki/Bogda_Shan" title="Bogda Shan">Bogda Mountains</a> in <a href="/wiki/Xinjiang" title="Xinjiang">Xinjiang</a>, <a href="/wiki/China" title="China">China</a>, although it is possible that only two (<i>L. youngi</i> and <i>L. hedini</i>) are valid; unusually, no Chinese <i>Lystrosaurus</i> specimens are known below the Permian-Triassic boundary in this region.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> <i>L. curvatus</i>, <i>L. murrayi</i>, and <i>L. maccaigi</i> are known from the <a href="/wiki/Fremouw_Formation" title="Fremouw Formation">Fremouw Formation</a> in the <a href="/wiki/Transantarctic_Mountains" title="Transantarctic Mountains">Transantarctic Mountains</a> of <a href="/wiki/Antarctica" title="Antarctica">Antarctica</a>.<sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Description">Description</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Lystrosaurus&action=edit&section=6" title="Edit section: Description"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Lystrosaurus_murrayi_scale.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/9/9b/Lystrosaurus_murrayi_scale.svg/170px-Lystrosaurus_murrayi_scale.svg.png" decoding="async" width="170" height="159" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/9b/Lystrosaurus_murrayi_scale.svg/255px-Lystrosaurus_murrayi_scale.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/9b/Lystrosaurus_murrayi_scale.svg/340px-Lystrosaurus_murrayi_scale.svg.png 2x" data-file-width="400" data-file-height="375" /></a><figcaption>Size of <i>Lystrosaurus murrayi</i> relative to a human.</figcaption></figure> <p><i>Lystrosaurus</i> was a <a href="/wiki/Dicynodont" class="mw-redirect" title="Dicynodont">dicynodont</a> <a href="/wiki/Therapsid" class="mw-redirect" title="Therapsid">therapsid</a>, between 0.6 to 2.5 m (2 to 8 ft) long with an average of about 0.9 m (3 ft) depending upon the species.<sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> </p><p>Unlike other therapsids, dicynodonts had very short snouts and no teeth except for the <a href="/wiki/Tusk" title="Tusk">tusk</a>-like upper <a href="/wiki/Canine_teeth" class="mw-redirect" title="Canine teeth">canines</a>. Dicynodonts are generally thought to have had <a href="/wiki/Horn_(anatomy)" title="Horn (anatomy)">horny</a> <a href="/wiki/Beak" title="Beak">beaks</a> like those of turtles, for shearing off pieces of vegetation, which were then ground on a horny <a href="/wiki/Secondary_palate" title="Secondary palate">secondary palate</a> when the mouth was shut. The jaw joint was weak and moved backwards and forwards with a shearing action, instead of the more common sideways or up and down movements. It is thought that the jaw muscles were attached unusually far forward on the skull and took up a lot of space on the top and back of the skull. As a result, the eyes were set high and well forward on the skull, and the face was short.<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> </p><p>Features of the skeleton indicate that <i>Lystrosaurus</i> moved with a <a href="/wiki/Terrestrial_locomotion" title="Terrestrial locomotion">semi-sprawling gait</a>. The lower rear corner of the <a href="/wiki/Scapula" title="Scapula">scapula</a> (shoulder blade) was strongly ossified (built of strong bone), which suggests that movement of the scapula contributed to the stride length of the forelimbs and reduced the sideways flexing of the body.<sup id="cite_ref-Surkov-Kalndze-Benton-2005_7-2" class="reference"><a href="#cite_note-Surkov-Kalndze-Benton-2005-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> The five <a href="/wiki/Sacral_vertebrae" class="mw-redirect" title="Sacral vertebrae">sacral vertebrae</a> were massive but not fused to each other and to the <a href="/wiki/Pelvis" title="Pelvis">pelvis</a>, making the back more rigid and reducing sideways flexing while the animal was walking. <a href="/wiki/Therapsida" title="Therapsida">Therapsids</a> with fewer than five sacral vertebrae are thought to have had sprawling limbs, like those of modern lizards.<sup id="cite_ref-Surkov-Kalndze-Benton-2005_7-3" class="reference"><a href="#cite_note-Surkov-Kalndze-Benton-2005-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> In <a href="/wiki/Dinosaur" title="Dinosaur">dinosaurs</a> and <a href="/wiki/Mammal" title="Mammal">mammals</a>, which have erect limbs, the sacral vertebrae are fused to each other and to the <a href="/wiki/Pelvis" title="Pelvis">pelvis</a>.<sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> A buttress above each acetabulum (hip socket) is thought to have prevented <a href="/wiki/Joint_dislocation" title="Joint dislocation">dislocation</a> of the <a href="/wiki/Femur" title="Femur">femur</a> (thigh bone) while <i>Lystrosaurus</i> was walking with a semi-sprawling gait.<sup id="cite_ref-Surkov-Kalndze-Benton-2005_7-4" class="reference"><a href="#cite_note-Surkov-Kalndze-Benton-2005-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> The forelimbs of <i>Lystrosaurus</i> were massive,<sup id="cite_ref-Surkov-Kalndze-Benton-2005_7-5" class="reference"><a href="#cite_note-Surkov-Kalndze-Benton-2005-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> and <i>Lystrosaurus</i> is thought to have been a powerful burrower. </p><p>Mummified specimens recovered from the Karoo Basin and described in 2022 revealed that <i>Lystrosaurus</i> had dimpled, leathery, and hairless skin.<sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Paleoecology">Paleoecology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Lystrosaurus&action=edit&section=7" title="Edit section: Paleoecology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Dominance_of_the_Early_Triassic">Dominance of the Early Triassic</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Lystrosaurus&action=edit&section=8" title="Edit section: Dominance of the Early Triassic"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Lystr_georg1DB.jpg" class="mw-file-description"><img alt="A pink/grey four-footed animal. The head is facing slightly toward you, and has two big teeth. It is covered in fur and has sharp claws." src="//upload.wikimedia.org/wikipedia/commons/thumb/0/0b/Lystr_georg1DB.jpg/220px-Lystr_georg1DB.jpg" decoding="async" width="220" height="111" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/0b/Lystr_georg1DB.jpg/330px-Lystr_georg1DB.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/0b/Lystr_georg1DB.jpg/440px-Lystr_georg1DB.jpg 2x" data-file-width="1500" data-file-height="760" /></a><figcaption><i>Lystrosaurus georgi</i></figcaption></figure> <p><i>Lystrosaurus</i> is notable for dominating southern <a href="/wiki/Pangaea" title="Pangaea">Pangaea</a> for millions of years during the <a href="/wiki/Early_Triassic" title="Early Triassic">Early Triassic</a>. At least one unidentified species of this genus survived the <a href="/wiki/Permian%E2%80%93Triassic_extinction_event" title="Permian–Triassic extinction event">end-Permian mass extinction</a> and, in the absence of predators and herbivorous competitors, went on to thrive and <a href="/wiki/Evolutionary_radiation" title="Evolutionary radiation">re-radiate</a> into a number of species within the genus,<sup id="cite_ref-Benton-2006_19-0" class="reference"><a href="#cite_note-Benton-2006-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> becoming the most common group of terrestrial vertebrates during the Early Triassic; for a while, 95% of land vertebrates were <i>Lystrosaurus</i>.<sup id="cite_ref-Benton-2006_19-1" class="reference"><a href="#cite_note-Benton-2006-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Cokinos-2007_20-0" class="reference"><a href="#cite_note-Cokinos-2007-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> This is the only time that a single species or genus of land animal dominated the Earth to such a degree.<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> A few other Permian therapsid genera also survived the mass extinction and appear in Triassic rocks—the <a href="/wiki/Therocephalia" title="Therocephalia">therocephalians</a> <i><a href="/wiki/Tetracynodon" title="Tetracynodon">Tetracynodon</a></i>, <i><a href="/wiki/Moschorhinus" title="Moschorhinus">Moschorhinus</a></i>, <i><a href="/wiki/Ictidosuchoides" title="Ictidosuchoides">Ictidosuchoides</a></i> and <i><a href="/wiki/Promoschorhynchus" title="Promoschorhynchus">Promoschorhynchus</a></i>—but do not appear to have been abundant in the Triassic;<sup id="cite_ref-Botha-Smith-2007_8-6" class="reference"><a href="#cite_note-Botha-Smith-2007-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup> complete ecological recovery took 30 million years, spanning the <a href="/wiki/Early_Triassic" title="Early Triassic">Early</a> and <a href="/wiki/Middle_Triassic" title="Middle Triassic">Middle Triassic</a>.<sup id="cite_ref-23" class="reference"><a href="#cite_note-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup> </p><p>Several attempts have been made to explain why <i>Lystrosaurus</i> survived the <a href="/wiki/Permian%E2%80%93Triassic_extinction_event" title="Permian–Triassic extinction event">Permian–Triassic extinction event</a>, the "mother of all mass extinctions",<sup id="cite_ref-Erwin1993_24-0" class="reference"><a href="#cite_note-Erwin1993-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup> and why it dominated Early Triassic fauna to such an unprecedented extent: </p> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Lystr_murr1DB.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/3f/Lystr_murr1DB.jpg/220px-Lystr_murr1DB.jpg" decoding="async" width="220" height="109" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/3f/Lystr_murr1DB.jpg/330px-Lystr_murr1DB.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/3f/Lystr_murr1DB.jpg/440px-Lystr_murr1DB.jpg 2x" data-file-width="1500" data-file-height="745" /></a><figcaption><i>Lystrosaurus murrayi</i></figcaption></figure> <ul><li>Growth marks in fossilized tusks suggest that <i>Lystrosaurus</i> living in Antarctica <a href="/wiki/Early_Triassic" title="Early Triassic">~250 Mya</a> could enter a prolonged state of <a href="/wiki/Torpor" title="Torpor">torpor</a> analogous to <a href="/wiki/Hibernation" title="Hibernation">hibernation</a>. This could be the oldest evidence of a hibernation-like state in a vertebrate animal and indicates that torpor arose in vertebrates before mammals and dinosaurs evolved.<sup id="cite_ref-Lystrosaurus_25-0" class="reference"><a href="#cite_note-Lystrosaurus-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-26" class="reference"><a href="#cite_note-26"><span class="cite-bracket">[</span>26<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-27" class="reference"><a href="#cite_note-27"><span class="cite-bracket">[</span>27<span class="cite-bracket">]</span></a></sup></li> <li>One of the more recent theories is that the extinction event reduced the atmosphere's oxygen content and increased its <a href="/wiki/Carbon_dioxide" title="Carbon dioxide">carbon dioxide</a> content, so that many terrestrial species died out because they found breathing too difficult.<sup id="cite_ref-Cokinos-2007_20-1" class="reference"><a href="#cite_note-Cokinos-2007-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> It has therefore been suggested that <i>Lystrosaurus</i> survived and became dominant because its burrowing life-style made it able to cope with an atmosphere of "stale air", and that specific features of its anatomy were part of this adaptation: a barrel chest that accommodated large lungs, short internal <a href="/wiki/Nostril" title="Nostril">nostrils</a> that facilitated rapid breathing, and high <a href="/wiki/Neural_spine" class="mw-redirect" title="Neural spine">neural spines</a> (projections on the dorsal side of the <a href="/wiki/Vertebra" title="Vertebra">vertebrae</a>) that gave greater leverage to the muscles that expanded and contracted its chest. However, there are weaknesses in all these points: the chest of <i>Lystrosaurus</i> was not significantly larger in proportion to its size than in other <a href="/wiki/Dicynodont" class="mw-redirect" title="Dicynodont">dicynodonts</a> that became extinct; although Triassic dicynodonts appear to have had longer neural spines than their Permian counterparts, this feature may be related to posture, locomotion or even body size rather than respiratory efficiency; <i>L. murrayi</i> and <i>L. declivis</i> are much more abundant than other Early Triassic burrowers such as <i><a href="/wiki/Procolophon" title="Procolophon">Procolophon</a></i> or <i><a href="/wiki/Thrinaxodon" title="Thrinaxodon">Thrinaxodon</a></i>.<sup id="cite_ref-Botha-Smith-2007_8-7" class="reference"><a href="#cite_note-Botha-Smith-2007-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup></li></ul> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Lystrosaurus_skeleton.jpg" class="mw-file-description"><img alt="Full body view of the skeleton of a four footed animal." src="//upload.wikimedia.org/wikipedia/commons/thumb/d/d3/Lystrosaurus_skeleton.jpg/220px-Lystrosaurus_skeleton.jpg" decoding="async" width="220" height="112" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/d3/Lystrosaurus_skeleton.jpg/330px-Lystrosaurus_skeleton.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/d3/Lystrosaurus_skeleton.jpg/440px-Lystrosaurus_skeleton.jpg 2x" data-file-width="2035" data-file-height="1038" /></a><figcaption><i>Lystrosaurus</i> skeletal diagram</figcaption></figure> <ul><li>The suggestion that <i>Lystrosaurus</i> was helped to survive and dominate by being semi-aquatic has a similar weakness: although <a href="/wiki/Temnospondyl" class="mw-redirect" title="Temnospondyl">temnospondyls</a> become more abundant in the Karoo's Triassic sediments, they were much less numerous than <i>L. murrayi</i> and <i>L. declivis</i>.<sup id="cite_ref-Botha-Smith-2007_8-8" class="reference"><a href="#cite_note-Botha-Smith-2007-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup></li> <li>The most specialized and the largest animals are at higher risk in mass extinctions; this may explain why the unspecialized <i>L. curvatus</i> survived while the larger and more specialized <i>L. maccaigi</i> perished along with all the other large Permian herbivores and carnivores.<sup id="cite_ref-Botha-Smith-2007_8-9" class="reference"><a href="#cite_note-Botha-Smith-2007-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> Although <i>Lystrosaurus</i> generally looks adapted to feed on plants similar to <i><a href="/wiki/Dicroidium" title="Dicroidium">Dicroidium</a></i>, which dominated the Early Triassic, the larger size of <i>L. maccaigi</i> may have forced it to rely on the larger members of the <i><a href="/wiki/Glossopteris" title="Glossopteris">Glossopteris</a></i> flora, which did not survive the end-Permian extinction.<sup id="cite_ref-Botha-Smith-2007_8-10" class="reference"><a href="#cite_note-Botha-Smith-2007-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup></li> <li>Only the 1.5 m (5 ft)–long therocephalian <i><a href="/wiki/Moschorhinus" title="Moschorhinus">Moschorhinus</a></i> and the large <a href="/wiki/Archosauriform" class="mw-redirect" title="Archosauriform">archosauriform</a> <i><a href="/wiki/Proterosuchus" title="Proterosuchus">Proterosuchus</a></i> appear to be large enough to have preyed on the Triassic <i>Lystrosaurus</i> species, and this shortage of predators may have been responsible for a <i>Lystrosaurus</i> population boom in the Early Triassic.<sup id="cite_ref-Botha-Smith-2007_8-11" class="reference"><a href="#cite_note-Botha-Smith-2007-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup></li> <li>According to Benton, "Perhaps the survival of <i>Lystrosaurus</i> was simply a matter of luck".<sup id="cite_ref-Benton-2006_19-2" class="reference"><a href="#cite_note-Benton-2006-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup></li></ul> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Lystrosaurus&action=edit&section=9" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1259569809">.mw-parser-output .portalbox{padding:0;margin:0.5em 0;display:table;box-sizing:border-box;max-width:175px;list-style:none}.mw-parser-output .portalborder{border:1px solid var(--border-color-base,#a2a9b1);padding:0.1em;background:var(--background-color-neutral-subtle,#f8f9fa)}.mw-parser-output .portalbox-entry{display:table-row;font-size:85%;line-height:110%;height:1.9em;font-style:italic;font-weight:bold}.mw-parser-output .portalbox-image{display:table-cell;padding:0.2em;vertical-align:middle;text-align:center}.mw-parser-output .portalbox-link{display:table-cell;padding:0.2em 0.2em 0.2em 0.3em;vertical-align:middle}@media(min-width:720px){.mw-parser-output .portalleft{clear:left;float:left;margin:0.5em 1em 0.5em 0}.mw-parser-output .portalright{clear:right;float:right;margin:0.5em 0 0.5em 1em}}</style><ul role="navigation" aria-label="Portals" class="noprint portalbox portalborder portalright"> <li class="portalbox-entry"><span class="portalbox-image"><span class="noviewer" typeof="mw:File"><a href="/wiki/File:WikiProject_Geology.svg" class="mw-file-description"><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e7/WikiProject_Geology.svg/28px-WikiProject_Geology.svg.png" decoding="async" width="28" height="28" class="mw-file-element" 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.reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width" style="column-width: 25em;"> <ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFCluver1978" class="citation book cs1">Cluver, Michael Albert (1978). <i>Fossil reptiles of the South African Karoo</i>. The South African Museum. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/9780908407583" title="Special:BookSources/9780908407583"><bdi>9780908407583</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Fossil+reptiles+of+the+South+African+Karoo&rft.pub=The+South+African+Museum&rft.date=1978&rft.isbn=9780908407583&rft.aulast=Cluver&rft.aufirst=Michael+Albert&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALystrosaurus" class="Z3988"></span></span> </li> <li id="cite_note-Barendskraal-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-Barendskraal_2-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFDamianiNevelingModestoYates2004" class="citation journal cs1">Damiani, R.J.; Neveling, J.; Modesto, S.P. & Yates, A.M. (2004). "Barendskraal, a diverse amniote locality from the <i>Lystrosaurus</i> assemblage zone, Early Triassic of South Africa". <i>Palaeontologia Africana</i>. <b>39</b>: 53–62.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Palaeontologia+Africana&rft.atitle=Barendskraal%2C+a+diverse+amniote+locality+from+the+Lystrosaurus+assemblage+zone%2C+Early+Triassic+of+South+Africa&rft.volume=39&rft.pages=53-62&rft.date=2004&rft.aulast=Damiani&rft.aufirst=R.J.&rft.au=Neveling%2C+J.&rft.au=Modesto%2C+S.P.&rft.au=Yates%2C+A.M.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALystrosaurus" class="Z3988"></span></span> </li> <li id="cite_note-wallace2000-3"><span class="mw-cite-backlink">^ <a href="#cite_ref-wallace2000_3-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-wallace2000_3-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWallace,_David_Rains2000" class="citation book cs1">Wallace, David Rains (2000). <i>The Bonehunters' Revenge: Dinosaurs, Greed, and the Greatest Scientific Feud of the Gilded Age</i>. 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Retrieved <span class="nowrap">7 September</span> 2020</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=Fossil+suggests+animals+have+been+hibernating+for+250+million+years&rft_id=https%3A%2F%2Fwww.upi.com%2FScience_News%2F2020%2F08%2F27%2FFossil-suggests-animals-have-been-hibernating-for-250-million-years%2F8421598530111%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALystrosaurus" class="Z3988"></span></span> </li> <li id="cite_note-27"><span class="mw-cite-backlink"><b><a href="#cite_ref-27">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWhitneySidor2020" class="citation journal cs1">Whitney, Megan R.; Sidor, Christian A. (27 August 2020). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7453012">"Evidence of torpor in the tusks of Lystrosaurus from the Early Triassic of Antarctica"</a>. <i>Communications Biology</i>. <b>3</b> (1): 471. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2Fs42003-020-01207-6">10.1038/s42003-020-01207-6</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/2399-3642">2399-3642</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7453012">7453012</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/32855434">32855434</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Communications+Biology&rft.atitle=Evidence+of+torpor+in+the+tusks+of+Lystrosaurus+from+the+Early+Triassic+of+Antarctica&rft.volume=3&rft.issue=1&rft.pages=471&rft.date=2020-08-27&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC7453012%23id-name%3DPMC&rft.issn=2399-3642&rft_id=info%3Apmid%2F32855434&rft_id=info%3Adoi%2F10.1038%2Fs42003-020-01207-6&rft.aulast=Whitney&rft.aufirst=Megan+R.&rft.au=Sidor%2C+Christian+A.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC7453012&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALystrosaurus" class="Z3988"></span> <span typeof="mw:File"><a href="/wiki/File:CC_BY_icon.svg" class="mw-file-description"><img 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Archived from <a rel="nofollow" class="external text" href="http://www.palaeos.com/Vertebrates/Units/400Therapsida/400.725.html#Dicynodontia">the original</a> on 22 February 2006.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Palaeos.com&rft.atitle=Dicynodontia&rft_id=http%3A%2F%2Fwww.palaeos.com%2FVertebrates%2FUnits%2F400Therapsida%2F400.725.html%23Dicynodontia&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALystrosaurus" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFRance" class="citation web cs1">Rance, Hugh. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20060427060532/http://geowords.com/histbooknetscape/i07.htm">"Mammal-like reptiles of Pangea"</a>. <i>geowords.com</i>. The present is the key to the past. 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.navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Anomodontia" title="Template:Anomodontia"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Anomodontia" title="Template talk:Anomodontia"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Anomodontia" title="Special:EditPage/Template:Anomodontia"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Anomodontia" style="font-size:114%;margin:0 4em"><a href="/wiki/Anomodontia" title="Anomodontia">Anomodontia</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="*_Kingdom:_Animalia_*_Phylum:_Chordata_*_Clade:_Synapsida_*_Clade:_Therapsida" style="font-size:114%;margin:0 4em"><span style="font-weight:normal; font-style:normal"><span style="font-size:85%;"> <ul><li>Kingdom: <a href="/wiki/Animal" title="Animal">Animalia</a></li> <li>Phylum: <a href="/wiki/Chordate" title="Chordate">Chordata</a></li> <li><i>Clade</i>: <a href="/wiki/Synapsida" title="Synapsida">Synapsida</a></li> <li><i>Clade</i>: <a href="/wiki/Therapsida" title="Therapsida">Therapsida</a></li></ul> </span></span></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Synapsida" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Synapsida" title="Synapsida">Synapsida</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Synapsida" title="Template:Synapsida">Synapsida</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Anomodontia" title="Anomodontia">Anomodontia</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><div id="Anomodontia" style="font-size:114%;margin:0 4em"><a href="/wiki/Anomodontia" title="Anomodontia">Anomodontia</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Anomodontia" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute; transform: translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Anomodontia" title="Anomodontia">Anomodontia</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Biseridens" title="Biseridens">Biseridens</a></i></li> <li><i><a href="/wiki/Novocynodon" title="Novocynodon">Novocynodon</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Anomocephaloidea" title="Anomocephaloidea">Anomocephaloidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Anomocephalus" title="Anomocephalus">Anomocephalus</a></i></li> <li><i><a href="/wiki/Tiarajudens" title="Tiarajudens">Tiarajudens</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Venyukovioidea" title="Venyukovioidea">Venyukovioidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Otsheria" title="Otsheria">Otsheria</a></i></li> <li><i><a href="/wiki/Parasuminia" title="Parasuminia">Parasuminia</a></i></li> <li><i><a href="/wiki/Suminia" title="Suminia">Suminia</a></i></li> <li><i><a href="/wiki/Ulemica" title="Ulemica">Ulemica</a></i></li> <li><i><a href="/wiki/Venyukovia" title="Venyukovia">Venyukovia</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Chainosauria" title="Chainosauria">Chainosauria</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Galechirus" title="Galechirus">Galechirus</a></i></li> <li><i><a href="/wiki/Galeops" title="Galeops">Galeops</a></i></li> <li><i><a href="/wiki/Galepus" title="Galepus">Galepus</a></i></li> <li><i><a href="/wiki/Patranomodon" title="Patranomodon">Patranomodon</a></i></li> <li><b><a href="/wiki/Anomocephaloidea" title="Anomocephaloidea">Anomocephaloidea</a></b>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Dicynodontia" title="Dicynodontia">Dicynodontia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Galepus.jpg" class="mw-file-description" title="Galepus jouberti"><img alt="Galepus jouberti" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/34/Galepus.jpg/100px-Galepus.jpg" decoding="async" width="100" height="123" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/34/Galepus.jpg/150px-Galepus.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/34/Galepus.jpg/200px-Galepus.jpg 2x" data-file-width="691" data-file-height="852" /></a></span></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><div id="Dicynodontia" style="font-size:114%;margin:0 4em"><a href="/wiki/Dicynodontia" title="Dicynodontia">Dicynodontia</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Dicynodontia" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute; transform: translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Dicynodontia" title="Dicynodontia">Dicynodontia</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Brachyprosopus" title="Brachyprosopus">Brachyprosopus</a></i></li> <li><i><a href="/wiki/Colobodectes" title="Colobodectes">Colobodectes</a></i></li> <li><i><a href="/wiki/Eodicynodon" title="Eodicynodon">Eodicynodon</a></i></li> <li><i><a href="/wiki/Lanthanostegus" title="Lanthanostegus">Lanthanostegus</a></i></li> <li><i><a href="/wiki/Nyaphulia" title="Nyaphulia">Nyaphulia</a></i></li> <li><i><a href="/wiki/Pristerodon" title="Pristerodon">Pristerodon</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Pylaecephalidae" title="Pylaecephalidae">Pylaecephalidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Diictodon" title="Diictodon">Diictodon</a></i></li> <li><i><a href="/wiki/Eosimops" title="Eosimops">Eosimops</a></i></li> <li><i><a href="/wiki/Prosictodon" title="Prosictodon">Prosictodon</a></i></li> <li><i><a href="/wiki/Robertia" title="Robertia">Robertia</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Endothiodontia" title="Endothiodontia">Endothiodontia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Abajudon&action=edit&redlink=1" class="new" title="Abajudon (page does not exist)">Abajudon</a></i></li> <li><i><a href="/wiki/Endothiodon" title="Endothiodon">Endothiodon</a></i></li> <li><i><a href="/wiki/Niassodon" title="Niassodon">Niassodon</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Therochelonia" title="Therochelonia">Therochelonia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Emydopoidea" title="Emydopoidea">Emydopoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Digalodon" title="Digalodon">Digalodon</a></i></li> <li><i><a href="/wiki/Emydorhinus" title="Emydorhinus">Emydorhinus</a></i>?</li> <li><i><a href="/wiki/Palemydops" title="Palemydops">Palemydops</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Emydopidae" title="Emydopidae">Emydopidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Compsodon" title="Compsodon">Compsodon</a></i></li> <li><i><a href="/wiki/Emydops" title="Emydops">Emydops</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Myosauridae" title="Myosauridae">Myosauridae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Myosauroides" title="Myosauroides">Myosauroides</a></i></li> <li><i><a href="/wiki/Myosaurus" title="Myosaurus">Myosaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Kingoriidae" title="Kingoriidae">Kingoriidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Dicynodontoides" title="Dicynodontoides">Dicynodontoides</a></i></li> <li><i><a href="/wiki/Kombuisia" title="Kombuisia">Kombuisia</a></i></li> <li><i><a href="/wiki/Thliptosaurus" title="Thliptosaurus">Thliptosaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cistecephalidae" title="Cistecephalidae">Cistecephalidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Cistecephaloides" title="Cistecephaloides">Cistecephaloides</a></i></li> <li><i><a href="/wiki/Cistecephalus" title="Cistecephalus">Cistecephalus</a></i></li> <li><i><a href="/wiki/Kawingasaurus" title="Kawingasaurus">Kawingasaurus</a></i></li> <li><i><a href="/wiki/Kembawacela" title="Kembawacela">Kembawacela</a></i></li> <li><i><a href="/wiki/Sauroscaptor" title="Sauroscaptor">Sauroscaptor</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Bidentalia" title="Bidentalia">Bidentalia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Diictodon_EF.jpg" class="mw-file-description" title="Diictodon feliceps"><img alt="Diictodon feliceps" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/87/Diictodon_EF.jpg/140px-Diictodon_EF.jpg" decoding="async" width="140" height="79" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/87/Diictodon_EF.jpg/210px-Diictodon_EF.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/87/Diictodon_EF.jpg/280px-Diictodon_EF.jpg 2x" data-file-width="2000" data-file-height="1125" /></a></span> <span typeof="mw:File"><a href="/wiki/File:KingoriaDB.jpg" class="mw-file-description" title="Dicynodontoides recurvidens"><img alt="Dicynodontoides recurvidens" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/93/KingoriaDB.jpg/140px-KingoriaDB.jpg" decoding="async" width="140" height="58" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/93/KingoriaDB.jpg/210px-KingoriaDB.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/93/KingoriaDB.jpg/280px-KingoriaDB.jpg 2x" data-file-width="1000" data-file-height="415" /></a></span></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible uncollapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><div id="Bidentalia" style="font-size:114%;margin:0 4em"><a href="/wiki/Bidentalia" title="Bidentalia">Bidentalia</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Bidentalia" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute; transform: translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Bidentalia" title="Bidentalia">Bidentalia</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Daqingshanodon" title="Daqingshanodon">Daqingshanodon</a></i></li> <li><i><a href="/wiki/Idelesaurus" title="Idelesaurus">Idelesaurus</a></i></li> <li><i><a href="/wiki/Keyseria" title="Keyseria">Keyseria</a></i></li> <li><i><a href="/wiki/Odontocyclops" title="Odontocyclops">Odontocyclops</a></i></li> <li><i><a href="/wiki/Rastodon" title="Rastodon">Rastodon</a></i></li> <li><i><a href="/wiki/Syops" title="Syops">Syops</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Elphidae" title="Elphidae">Elphidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Elph_(therapsid)" title="Elph (therapsid)">Elph</a></i></li> <li><i><a href="/wiki/Interpresosaurus" title="Interpresosaurus">Interpresosaurus</a></i></li> <li><i><a href="/wiki/Katumbia" title="Katumbia">Katumbia</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Geikiidae" title="Geikiidae">Geikiidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Aulacephalodon" title="Aulacephalodon">Aulacephalodon</a></i></li> <li><i><a href="/wiki/Bulbasaurus" title="Bulbasaurus">Bulbasaurus</a></i></li> <li><i><a href="/wiki/Geikia" title="Geikia">Geikia</a></i></li> <li><i><a href="/wiki/Pelanomodon" title="Pelanomodon">Pelanomodon</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Oudenodontidae" title="Oudenodontidae">Oudenodontidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Australobarbarus" title="Australobarbarus">Australobarbarus</a></i></li> <li><i><a href="/wiki/Oudenodon" title="Oudenodon">Oudenodon</a></i></li> <li><i><a href="/wiki/Tropidostoma" title="Tropidostoma">Tropidostoma</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Rhachiocephalidae" title="Rhachiocephalidae">Rhachiocephalidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Kitchinganomodon" title="Kitchinganomodon">Kitchinganomodon</a></i></li> <li><i><a href="/wiki/Rhachiocephalus" title="Rhachiocephalus">Rhachiocephalus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Dicynodontoidea" title="Dicynodontoidea">Dicynodontoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Counillonia" title="Counillonia">Counillonia</a></i></li> <li><i><a href="/wiki/Daptocephalus" title="Daptocephalus">Daptocephalus</a></i></li> <li><i><a href="/wiki/Delectosaurus" title="Delectosaurus">Delectosaurus</a></i></li> <li><i><a href="/wiki/Dicynodon" title="Dicynodon">Dicynodon</a></i></li> <li><i><a href="/wiki/Dinanomodon" title="Dinanomodon">Dinanomodon</a></i></li> <li><i><a href="/wiki/Gordonia_(synapsid)" title="Gordonia (synapsid)">Gordonia</a></i></li> <li><i><a href="/wiki/Kunpania" title="Kunpania">Kunpania</a></i></li> <li><i><a href="/wiki/Peramodon" title="Peramodon">Peramodon</a></i></li> <li><i><a href="/wiki/Repelinosaurus" title="Repelinosaurus">Repelinosaurus</a></i></li> <li><i><a href="/wiki/Taoheodon" title="Taoheodon">Taoheodon</a></i></li> <li><i><a href="/wiki/Turfanodon" title="Turfanodon">Turfanodon</a></i></li> <li><i><a href="/wiki/Vivaxosaurus" title="Vivaxosaurus">Vivaxosaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Lystrosauridae" title="Lystrosauridae">Lystrosauridae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Basilodon" title="Basilodon">Basilodon</a></i>?</li> <li><i><a href="/wiki/Euptychognathus" title="Euptychognathus">Euptychognathus</a></i>?</li> <li><i><a href="/wiki/Jimusaria" title="Jimusaria">Jimusaria</a></i>?</li> <li><i><a href="/wiki/Kwazulusaurus" title="Kwazulusaurus">Kwazulusaurus</a></i></li> <li><i><a class="mw-selflink selflink">Lystrosaurus</a></i></li> <li><i><a href="/wiki/Sintocephalus" title="Sintocephalus">Sintocephalus</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Kannemeyeriiformes" title="Kannemeyeriiformes">Kannemeyeriiformes</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:GeikiaDB17.jpg" class="mw-file-description" title="Geikia elginensis"><img alt="Geikia elginensis" src="//upload.wikimedia.org/wikipedia/commons/thumb/5/51/GeikiaDB17.jpg/140px-GeikiaDB17.jpg" decoding="async" width="140" height="67" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/51/GeikiaDB17.jpg/210px-GeikiaDB17.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/51/GeikiaDB17.jpg/280px-GeikiaDB17.jpg 2x" data-file-width="2284" data-file-height="1092" /></a></span> <span typeof="mw:File"><a href="/wiki/File:Lystrosaurus_BW.jpg" class="mw-file-description" title="Lystrosaurus murrayi"><img alt="Lystrosaurus murrayi" src="//upload.wikimedia.org/wikipedia/commons/thumb/1/17/Lystrosaurus_BW.jpg/140px-Lystrosaurus_BW.jpg" decoding="async" width="140" height="72" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/17/Lystrosaurus_BW.jpg/210px-Lystrosaurus_BW.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/17/Lystrosaurus_BW.jpg/280px-Lystrosaurus_BW.jpg 2x" data-file-width="800" data-file-height="410" /></a></span></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><div id="Kannemeyeriiformes" style="font-size:114%;margin:0 4em"><a href="/wiki/Kannemeyeriiformes" title="Kannemeyeriiformes">Kannemeyeriiformes</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Kannemeyeriiformes" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute; transform: translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Kannemeyeriiformes" title="Kannemeyeriiformes">Kannemeyeriiformes</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Angonisaurus" title="Angonisaurus">Angonisaurus</a></i></li> <li><i><a href="/wiki/Dinodontosaurus" title="Dinodontosaurus">Dinodontosaurus</a></i></li> <li><i><a href="/wiki/Elephantosaurus" title="Elephantosaurus">Elephantosaurus</a></i></li> <li><i><a href="/wiki/Woznikella" title="Woznikella">Woznikella</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Shansiodontidae" title="Shansiodontidae">Shansiodontidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Shansiodon" title="Shansiodon">Shansiodon</a></i></li> <li><i><a href="/wiki/Tetragonias" title="Tetragonias">Tetragonias</a></i></li> <li><i><a href="/wiki/Vinceria" title="Vinceria">Vinceria</a></i></li> <li><i><a href="/wiki/Rhinodicynodon" title="Rhinodicynodon">Rhinodicynodon</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Kannemeyeriidae" class="mw-redirect" title="Kannemeyeriidae">Kannemeyeriidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Acratophorus_argentinensis" title="Acratophorus argentinensis">Acratophorus</a></i></li> <li><i><a href="/wiki/Dolichuranus" title="Dolichuranus">Dolichuranus</a></i></li> <li><i><a href="/wiki/Kannemeyeria" title="Kannemeyeria">Kannemeyeria</a></i></li> <li><i><a href="/wiki/Parakannemeyeria" title="Parakannemeyeria">Parakannemeyeria</a></i></li> <li><i><a href="/wiki/Rabidosaurus" title="Rabidosaurus">Rabidosaurus</a></i></li> <li><i><a href="/wiki/Rechnisaurus" title="Rechnisaurus">Rechnisaurus</a></i></li> <li><i><a href="/wiki/Rhadiodromus" title="Rhadiodromus">Rhadiodromus</a></i></li> <li><i><a href="/wiki/Shaanbeikannemeyeria" title="Shaanbeikannemeyeria">Shaanbeikannemeyeria</a></i></li> <li><i><a href="/wiki/Sinokannemeyeria" title="Sinokannemeyeria">Sinokannemeyeria</a></i></li> <li><i><a href="/wiki/Uralokannemeyeria" title="Uralokannemeyeria">Uralokannemeyeria</a></i></li> <li><i><a href="/wiki/Wadiasaurus" title="Wadiasaurus">Wadiasaurus</a></i></li> <li><i><a href="/wiki/Xiyukannemeyeria" title="Xiyukannemeyeria">Xiyukannemeyeria</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Stahleckeriidae" title="Stahleckeriidae">Stahleckeriidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Sungeodon" title="Sungeodon">Sungeodon</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Stahleckeriidae" title="Stahleckeriidae">Placeriinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Argodicynodon" title="Argodicynodon">Argodicynodon</a></i></li> <li><i><a href="/wiki/Lisowicia" title="Lisowicia">Lisowicia</a></i></li> <li><i><a href="/wiki/Moghreberia" title="Moghreberia">Moghreberia</a></i></li> <li><i><a href="/wiki/Pentasaurus" title="Pentasaurus">Pentasaurus</a></i></li> <li><i><a href="/wiki/Placerias" title="Placerias">Placerias</a></i></li> <li><i><a href="/wiki/Zambiasaurus" title="Zambiasaurus">Zambiasaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Stahleckeriidae" title="Stahleckeriidae">Stahleckeriinae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Eubrachiosaurus" title="Eubrachiosaurus">Eubrachiosaurus</a></i></li> <li><i><a href="/wiki/Ischigualastia" title="Ischigualastia">Ischigualastia</a></i></li> <li><i><a href="/wiki/Jachaleria" title="Jachaleria">Jachaleria</a></i></li> <li><i><a href="/wiki/Sangusaurus" title="Sangusaurus">Sangusaurus</a></i></li> <li><i><a href="/wiki/Stahleckeria" title="Stahleckeria">Stahleckeria</a></i></li> <li><i><a href="/wiki/Ufudocyclops" title="Ufudocyclops">Ufudocyclops</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Lisowicia_bojani_J..png" class="mw-file-description" title="Lisowicia bojani"><img alt="Lisowicia bojani" src="//upload.wikimedia.org/wikipedia/commons/thumb/2/25/Lisowicia_bojani_J..png/140px-Lisowicia_bojani_J..png" decoding="async" width="140" height="81" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/25/Lisowicia_bojani_J..png/210px-Lisowicia_bojani_J..png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/25/Lisowicia_bojani_J..png/280px-Lisowicia_bojani_J..png 2x" data-file-width="2844" data-file-height="1651" /></a></span></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Other_articles" style="font-size:114%;margin:0 4em">Other articles</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Nomen_dubium" title="Nomen dubium">Nomina dubia</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Azarifeneria" class="mw-redirect" title="Azarifeneria">Azarifeneria</a></i></li> <li><i><a href="/w/index.php?title=Baiopsis&action=edit&redlink=1" class="new" title="Baiopsis (page does not exist)">Baiopsis</a></i></li> <li><i><a href="/wiki/Calleonasus" title="Calleonasus">Calleonasus</a></i></li> <li><i><a href="/w/index.php?title=Chelyoposaurus&action=edit&redlink=1" class="new" title="Chelyoposaurus (page does not exist)">Chelyoposaurus</a></i></li> <li><i><a href="/w/index.php?title=Cristonasus&action=edit&redlink=1" class="new" title="Cristonasus (page does not exist)">Cristonasus</a></i></li> <li><i><a href="/wiki/Diaelurodon" class="mw-redirect" title="Diaelurodon">Diaelurodon</a></i></li> <li><i><a href="/w/index.php?title=Dicranozygoma&action=edit&redlink=1" class="new" title="Dicranozygoma (page does not exist)">Dicranozygoma</a></i></li> <li><i><a href="/wiki/Elatosaurus" title="Elatosaurus">Elatosaurus</a></i></li> <li><i><a href="/wiki/Eurycarpus" class="mw-redirect" title="Eurycarpus">Eurycarpus</a></i></li> <li><i><a href="/wiki/Haughtoniana" title="Haughtoniana">Haughtoniana</a></i></li> <li><i><a href="/w/index.php?title=Keirognathus&action=edit&redlink=1" class="new" title="Keirognathus (page does not exist)">Keirognathus</a></i></li> <li><i><a href="/wiki/Koupia" title="Koupia">Koupia</a></i></li> <li><i><a href="/w/index.php?title=Nasoplanites&action=edit&redlink=1" class="new" title="Nasoplanites (page does not exist)">Nasoplanites</a></i></li> <li><i><a href="/w/index.php?title=Opisthoctenodon&action=edit&redlink=1" class="new" title="Opisthoctenodon (page does not exist)">Opisthoctenodon</a></i></li> <li><i><a href="/w/index.php?title=Parvobestiola&action=edit&redlink=1" class="new" title="Parvobestiola (page does not exist)">Parvobestiola</a></i></li> <li><i><a href="/w/index.php?title=Planitorostris&action=edit&redlink=1" class="new" title="Planitorostris (page does not exist)">Planitorostris</a></i></li> <li><i><a href="/w/index.php?title=Platypodosaurus&action=edit&redlink=1" class="new" title="Platypodosaurus (page does not exist)">Platypodosaurus</a></i></li> <li><i><a href="/wiki/Putillosaurus" title="Putillosaurus">Putillosaurus</a></i></li> <li><i><a href="/wiki/Taognathus" title="Taognathus">Taognathus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Paraphyly" title="Paraphyly">Paraphyletic groups</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cryptodontia" title="Cryptodontia">Cryptodontia</a>?</li> <li><a href="/wiki/Dromasauria" title="Dromasauria">Dromasauria</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Geology_of_South_Africa" style="padding:3px"><table class="nowraplinks hlist mw-collapsible mw-collapsed navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Geology_of_South_Africa" title="Template:Geology of South Africa"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Geology_of_South_Africa" title="Template talk:Geology of South Africa"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Geology_of_South_Africa" title="Special:EditPage/Template:Geology of South Africa"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Geology_of_South_Africa" style="font-size:114%;margin:0 4em"><a href="/wiki/Geology_of_South_Africa" title="Geology of South Africa">Geology of South Africa</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Geological_formations" style="font-size:114%;margin:0 4em">Geological formations</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/List_of_fossiliferous_stratigraphic_units_in_South_Africa" title="List of fossiliferous stratigraphic units in South Africa">List of fossiliferous stratigraphic units in South Africa</a></li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Palaeoarchaean</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bushveld_Igneous_Complex" title="Bushveld Igneous Complex">Bushveld Igneous Complex</a></li> <li><a href="/wiki/Fig_Tree_Formation" title="Fig Tree Formation">Fig Tree Formation</a></li> <li><a href="/wiki/Komatii_Formation" title="Komatii Formation">Komatii Formation</a></li> <li><a href="/wiki/Moodies_Group" title="Moodies Group">Moodies Group</a></li> <li><a href="/wiki/Onverwacht_Group" title="Onverwacht Group">Onverwacht Group</a></li> <li><a href="/wiki/Transvaal_Supergroup" title="Transvaal Supergroup">Transvaal Supergroup</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Paleozoic</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Carboniferous <ul><li><a href="/wiki/Dwyka_Group" title="Dwyka Group">Dwyka Group</a></li> <li><a href="/wiki/Ganigobis_Formation" title="Ganigobis Formation">Ganigobis Formation</a></li></ul></li> <li>Devonian <ul><li><a href="/wiki/Bokkeveld_Group" title="Bokkeveld Group">Bokkeveld Group</a></li> <li><a href="/wiki/Table_Mountain_Sandstone" title="Table Mountain Sandstone">Table Mountain Sandstone</a></li></ul></li> <li>Permian <ul><li><a href="/wiki/Abrahamskraal_Formation" title="Abrahamskraal Formation">Abrahamskraal Formation</a></li> <li><a href="/wiki/Balfour_Formation" title="Balfour Formation">Balfour Formation</a></li> <li><a href="/wiki/Dwyka_Group" title="Dwyka Group">Dwyka Group</a></li> <li><a href="/wiki/Ganigobis_Formation" title="Ganigobis Formation">Ganigobis Formation</a></li> <li><a href="/wiki/Teekloof_Formation" title="Teekloof Formation">Teekloof Formation</a></li> <li><a href="/wiki/Whitehill_Formation" title="Whitehill Formation">Whitehill Formation</a></li></ul></li> <li>Ordovician <ul><li><a href="/wiki/Soom_Shale" title="Soom Shale">Soom Shale</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Mesozoic</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Cretaceous <ul><li><a href="/wiki/Uitenhage_Group" title="Uitenhage Group">Uitenhage Group</a> <ul><li><a href="/wiki/Buffelskloof_Formation" title="Buffelskloof Formation">Buffelskloof Formation</a></li> <li><a href="/wiki/Enon_Formation" title="Enon Formation">Enon Formation</a></li> <li><a href="/wiki/Kirkwood_Formation" title="Kirkwood Formation">Kirkwood Formation</a></li> <li><a href="/wiki/Sundays_River_Formation" title="Sundays River Formation">Sundays River Formation</a></li></ul></li> <li><a href="/wiki/Kalahari_Deposits" title="Kalahari Deposits">Kalahari Deposits</a></li></ul></li></ul> <ul><li>Jurassic <ul><li><a href="/wiki/Drakensberg_Group" title="Drakensberg Group">Drakensberg Group</a></li> <li><a href="/wiki/Enon_Formation" title="Enon Formation">Enon Formation</a></li> <li><a href="/wiki/Uitenhage_Group" title="Uitenhage Group">Uitenhage Group</a></li></ul></li> <li><a href="/wiki/Karoo_Supergroup" title="Karoo Supergroup">Karoo Supergroup</a> <ul><li><a href="/wiki/Beaufort_Group" title="Beaufort Group">Beaufort Group</a> <ul><li><a href="/wiki/Abrahamskraal_Formation" title="Abrahamskraal Formation">Abrahamskraal Formation</a></li> <li><a href="/wiki/Balfour_Formation" title="Balfour Formation">Balfour Formation</a></li> <li><a href="/wiki/Katberg_Formation" title="Katberg Formation">Katberg Formation</a></li> <li><a href="/wiki/Middleton_Formation" title="Middleton Formation">Middleton Formation</a></li> <li><a href="/wiki/Teekloof_Formation" title="Teekloof Formation">Teekloof Formation</a></li></ul></li> <li><a href="/wiki/Dwyka_Group" title="Dwyka Group">Dwyka Group</a></li> <li><a href="/wiki/Ecca_Group" title="Ecca Group">Ecca Group</a> <ul><li><a href="/wiki/Whitehill_Formation" title="Whitehill Formation">Whitehill Formation</a></li></ul></li> <li><a href="/wiki/Stormberg_Group" title="Stormberg Group">Stormberg Group</a> <ul><li><a href="/wiki/Bushveld_Sandstone" title="Bushveld Sandstone">Bushveld Sandstone</a></li> <li><a href="/wiki/Clarens_Formation" title="Clarens Formation">Clarens Formation</a></li> <li><a href="/wiki/Elliot_Formation" title="Elliot Formation">Elliot Formation</a></li></ul></li></ul></li> <li>Triassic <ul><li><a href="/wiki/Elliot_Formation" title="Elliot Formation">Elliot Formation</a></li> <li><a href="/wiki/Katberg_Formation" title="Katberg Formation">Katberg Formation</a></li> <li><a href="/wiki/Molteno_Formation" title="Molteno Formation">Molteno Formation</a></li> <li><a href="/wiki/Normandien_Formation" title="Normandien Formation">Normandien Formation</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Cenozoic</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Algoa_Group" title="Algoa Group">Algoa Group</a></li></ul> </div></td></tr></tbody></table><div> </div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Tectonics_and_orogeny" style="font-size:114%;margin:0 4em">Tectonics and orogeny</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Algoa_Basin" title="Algoa Basin">Algoa Basin</a></li> <li><a href="/wiki/Cape_Fold_Belt" title="Cape Fold Belt">Cape Fold Belt</a></li> <li><a href="/wiki/Ellisras_Basin" title="Ellisras Basin">Ellisras Basin</a></li> <li><a href="/wiki/Gondwanide_orogeny" title="Gondwanide orogeny">Gondwanide orogeny</a></li> <li><a href="/wiki/Kaapvaal_Craton" title="Kaapvaal Craton">Kaapvaal Craton</a></li> <li><a href="/wiki/Kalahari_Craton" title="Kalahari Craton">Kalahari Craton</a></li> <li><a href="/wiki/Makhonjwa_Mountains" title="Makhonjwa Mountains">Makhonjwa Mountains</a></li> <li><a href="/wiki/Tectonic_evolution_of_the_Barberton_greenstone_belt" title="Tectonic evolution of the Barberton greenstone belt">Tectonic evolution of the Barberton greenstone belt</a></li> <li><a href="/wiki/Terra_Australis_Orogen" title="Terra Australis Orogen">Terra Australis Orogen</a></li> <li><a href="/wiki/Transvaal_Basin" title="Transvaal Basin">Transvaal Basin</a></li> <li><a href="/wiki/Witteberg" title="Witteberg">Witteberg</a></li> <li><a href="/wiki/Witwatersrand_Basin" class="mw-redirect" title="Witwatersrand Basin">Witwatersrand Basin</a></li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Volcanism</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/List_of_volcanoes_in_South_Africa" title="List of volcanoes in South Africa">List of volcanoes in South Africa</a></li> <li><a href="/wiki/Pilanesberg" title="Pilanesberg">Pilanesberg</a></li> <li><a href="/wiki/Pilanesberg_Game_Reserve" class="mw-redirect" title="Pilanesberg Game Reserve">Pilanesberg Game Reserve</a></li> <li><a href="/wiki/Prince_Edward_Islands" title="Prince Edward Islands">Prince Edward Islands</a></li> <li>Diatremes <ul><li><a href="/wiki/Big_Hole" title="Big Hole">Big Hole</a></li> <li><a href="/wiki/Finsch_diamond_mine" title="Finsch diamond mine">Finsch diamond mine</a></li> <li><a href="/wiki/Jagersfontein_Mine" title="Jagersfontein Mine">Jagersfontein Mine</a></li> <li><a href="/wiki/Koffiefontein_mine" title="Koffiefontein mine">Koffiefontein mine</a></li></ul></li> <li><a href="/wiki/Barberton_Greenstone_Belt" title="Barberton Greenstone Belt">Barberton Greenstone Belt</a></li> <li><a href="/wiki/Drakensberg" title="Drakensberg">Drakensberg</a></li> <li><a href="/wiki/Karoo-Ferrar" title="Karoo-Ferrar">Karoo-Ferrar</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/List_of_earthquakes_in_South_Africa" title="List of earthquakes in South Africa">Earthquakes</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/1620_Robben_Island_earthquake" title="1620 Robben Island earthquake">1620 Robben Island earthquake</a></li> <li><a href="/wiki/1969_Tulbagh_earthquake" title="1969 Tulbagh earthquake">1969 Tulbagh earthquake</a></li> <li><a href="/wiki/2006_Mozambique_earthquake" title="2006 Mozambique earthquake">2006 Mozambique earthquake</a></li> <li><a href="/wiki/2014_Orkney_earthquake" title="2014 Orkney earthquake">2014 Orkney earthquake</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Impact craters</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Kalkkop_crater" title="Kalkkop crater">Kalkkop crater</a></li> <li><a href="/wiki/Morokweng_impact_structure" title="Morokweng impact structure">Morokweng impact structure</a></li> <li><a href="/wiki/Tswaing_crater" title="Tswaing crater">Tswaing crater</a></li> <li><a href="/wiki/Vredefort_impact_structure" title="Vredefort impact structure">Vredefort impact structure</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Meteorites</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bellsbank_meteorite" title="Bellsbank meteorite">Bellsbank meteorite</a></li></ul> </div></td></tr></tbody></table><div> </div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Mineral_deposits" style="font-size:114%;margin:0 4em">Mineral deposits</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Kimberlite" title="Kimberlite">Kimberlite</a></li> <li><a href="/wiki/Merensky_Reef" title="Merensky Reef">Merensky Reef</a></li> <li><a href="/wiki/Witwatersrand" title="Witwatersrand">Witwatersrand</a> <ul><li><a href="/wiki/Banket_(mining_term)" title="Banket (mining term)">Banket (mining term)</a></li></ul></li></ul> <ul><li><a href="/wiki/Coal_in_South_Africa" title="Coal in South Africa">Coal in South Africa</a></li> <li>Iron ore in Africa</li> <li><a href="/wiki/Platinum_in_Africa" title="Platinum in Africa">Platinum in Africa</a></li> <li><a href="/wiki/Titanium_in_Africa" title="Titanium in Africa">Titanium in Africa</a></li> <li><a href="/wiki/Uranium_in_Africa" title="Uranium in Africa">Uranium in Africa</a></li></ul> <ul><li><a href="/wiki/Minerals_Council_South_Africa" title="Minerals Council South Africa">Minerals Council South Africa</a></li> <li><a href="/wiki/Department_of_Mineral_Resources_(South_Africa)" title="Department of Mineral Resources (South Africa)">Department of Mineral Resources (South Africa)</a></li> <li><a href="/wiki/Department_of_Mineral_Resources_and_Energy" title="Department of Mineral Resources and Energy">Department of Mineral Resources and Energy</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible uncollapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Paleontology" style="font-size:114%;margin:0 4em">Paleontology</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/List_of_fossiliferous_stratigraphic_units_in_South_Africa" title="List of fossiliferous stratigraphic units in South Africa">List of fossiliferous stratigraphic units in South Africa</a></li> <li><a href="/wiki/Archean_life_in_the_Barberton_Greenstone_Belt" title="Archean life in the Barberton Greenstone Belt">Archean life in the Barberton Greenstone Belt</a></li> <li><a href="/wiki/Waterloo_Farm_lagerst%C3%A4tte" title="Waterloo Farm lagerstätte">Waterloo Farm lagerstätte</a></li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Ordovician</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Notiodella" title="Notiodella">Notiodella</a></li> <li><a href="/wiki/Onychopterella" title="Onychopterella">Onychopterella</a></li> <li><a href="/wiki/Promissum" title="Promissum">Promissum</a></li> <li><a href="/wiki/Soomaspis" title="Soomaspis">Soomaspis</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Devonian</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Africanaspis" title="Africanaspis">Africanaspis</a></li> <li><a href="/wiki/Antarctilamna" title="Antarctilamna">Antarctilamna</a></li> <li><a href="/wiki/Bainella" title="Bainella">Bainella</a></li> <li><a href="/wiki/Deltacephalaspis" title="Deltacephalaspis">Deltacephalaspis</a></li> <li><a href="/wiki/Diplacanthus" title="Diplacanthus">Diplacanthus</a></li> <li><a href="/wiki/Gondwanascorpio" title="Gondwanascorpio">Gondwanascorpio</a></li> <li><a href="/wiki/Priscomyzon" title="Priscomyzon">Priscomyzon</a></li> <li><a href="/wiki/Serenichthys" title="Serenichthys">Serenichthys</a></li> <li><a href="/wiki/Tutusius" title="Tutusius">Tutusius</a></li> <li><a href="/wiki/Umzantsia" title="Umzantsia">Umzantsia</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Carboniferous</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Dadoxylon" class="mw-redirect" title="Dadoxylon">Dadoxylon</a></li> <li><a href="/wiki/Lepidodendron" title="Lepidodendron">Lepidodendron</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Permian</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Abdalodon" title="Abdalodon">Abdalodon</a></li> <li><a href="/wiki/Alopecognathus" title="Alopecognathus">Alopecognathus</a></li> <li><a href="/wiki/Anomocephalus" title="Anomocephalus">Anomocephalus</a></li> <li><a href="/wiki/Basilodon" title="Basilodon">Basilodon</a></li> <li><a href="/wiki/Bulbasaurus" title="Bulbasaurus">Bulbasaurus</a></li> <li><a href="/wiki/Charassognathus" title="Charassognathus">Charassognathus</a></li> <li><a href="/wiki/Cistecephaloides" title="Cistecephaloides">Cistecephaloides</a></li> <li><a href="/wiki/Crapartinella" class="mw-redirect" title="Crapartinella">Crapartinella</a></li> <li><a href="/wiki/Cynosaurus" title="Cynosaurus">Cynosaurus</a></li> <li><a href="/wiki/Dicynodontoides" title="Dicynodontoides">Dicynodontoides</a></li> <li><a href="/wiki/Diictodon" title="Diictodon">Diictodon</a></li> <li><a href="/wiki/Dinanomodon" title="Dinanomodon">Dinanomodon</a></li> <li><a href="/wiki/Dinogorgon" title="Dinogorgon">Dinogorgon</a></li> <li><a href="/wiki/Dwykaselachus" title="Dwykaselachus">Dwykaselachus</a></li> <li><a href="/wiki/Endothiodon" title="Endothiodon">Endothiodon</a></li> <li><a href="/wiki/Eunotosaurus" title="Eunotosaurus">Eunotosaurus</a></li> <li><a href="/wiki/Galechirus" title="Galechirus">Galechirus</a></li> <li><a href="/wiki/Glanosuchus" title="Glanosuchus">Glanosuchus</a></li> <li><a href="/wiki/Glossopteris" title="Glossopteris">Glossopteris</a></li> <li><a href="/wiki/Hofmeyria" title="Hofmeyria">Hofmeyria</a></li> <li><a href="/wiki/Ictidognathus" title="Ictidognathus">Ictidognathus</a></li> <li><a href="/wiki/Ictidosuchoides" title="Ictidosuchoides">Ictidosuchoides</a></li> <li><a href="/wiki/Lepidodendron" title="Lepidodendron">Lepidodendron</a></li> <li><a href="/wiki/Lycosuchus" title="Lycosuchus">Lycosuchus</a></li> <li><a href="/wiki/Mesosaurus" title="Mesosaurus">Mesosaurus</a></li> <li><a href="/wiki/Moschops" title="Moschops">Moschops</a></li> <li><a href="/wiki/Odontocyclops" title="Odontocyclops">Odontocyclops</a></li> <li><a href="/wiki/Procynosuchus" title="Procynosuchus">Procynosuchus</a></li> <li><a href="/wiki/Scylacosaurus" title="Scylacosaurus">Scylacosaurus</a></li> <li><a href="/wiki/Scymnosaurus" title="Scymnosaurus">Scymnosaurus</a></li> <li><a href="/wiki/Simorhinella" title="Simorhinella">Simorhinella</a></li> <li><a href="/wiki/Stereosternum" title="Stereosternum">Stereosternum</a></li> <li><a href="/wiki/Theriognathus" title="Theriognathus">Theriognathus</a></li> <li><a href="/wiki/Thliptosaurus" title="Thliptosaurus">Thliptosaurus</a></li> <li><a href="/wiki/Trochosaurus" title="Trochosaurus">Trochosaurus</a></li> <li><a href="/wiki/Youngina" title="Youngina">Youngina</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Beaufort Group</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Aelurosaurus" title="Aelurosaurus">Aelurosaurus</a></li> <li><a href="/wiki/Anomocephalus" title="Anomocephalus">Anomocephalus</a></li> <li><a href="/wiki/Basilodon" title="Basilodon">Basilodon</a></li> <li><a href="/wiki/Blattoidealestes" title="Blattoidealestes">Blattoidealestes</a></li> <li><a href="/wiki/Brachyprosopus" title="Brachyprosopus">Brachyprosopus</a></li> <li><a href="/wiki/Broomistega" title="Broomistega">Broomistega</a></li> <li><a href="/wiki/Cistecephaloides" title="Cistecephaloides">Cistecephaloides</a></li> <li><a href="/wiki/Choerosaurus" title="Choerosaurus">Choerosaurus</a></li> <li><a href="/wiki/Clelandina" title="Clelandina">Clelandina</a></li> <li><a href="/wiki/Colobodectes" title="Colobodectes">Colobodectes</a></li> <li><a href="/wiki/Compsodon" title="Compsodon">Compsodon</a></li> <li><a href="/wiki/Cryptocynodon" class="mw-redirect" title="Cryptocynodon">Cryptocynodon</a></li> <li><a href="/wiki/Cynariognathus" class="mw-redirect" title="Cynariognathus">Cynariognathus</a></li> <li><a href="/wiki/Cynosaurus" title="Cynosaurus">Cynosaurus</a></li> <li><a href="/wiki/Diictodon" title="Diictodon">Diictodon</a></li> <li><a href="/wiki/Dinanomodon" title="Dinanomodon">Dinanomodon</a></li> <li><a href="/wiki/Eodicynodon" title="Eodicynodon">Eodicynodon</a></li> <li><a href="/wiki/Eohyosaurus" title="Eohyosaurus">Eohyosaurus</a></li> <li><a href="/wiki/Emyduranus" class="mw-redirect" title="Emyduranus">Emyduranus</a></li> <li><a href="/wiki/Euchambersia" title="Euchambersia">Euchambersia</a></li> <li><a href="/wiki/Homodontosaurus" title="Homodontosaurus">Homodontosaurus</a></li> <li><a href="/wiki/Icticephalus" title="Icticephalus">Icticephalus</a></li> <li><a href="/wiki/Ictidochampsa" title="Ictidochampsa">Ictidochampsa</a></li> <li><a href="/wiki/Ictidodon" title="Ictidodon">Ictidodon</a></li> <li><a href="/wiki/Ictidodraco" title="Ictidodraco">Ictidodraco</a></li> <li><a href="/wiki/Ictidostoma" title="Ictidostoma">Ictidostoma</a></li> <li><a href="/wiki/Lanthanostegus" title="Lanthanostegus">Lanthanostegus</a></li> <li><a href="/wiki/Leontosaurus" title="Leontosaurus">Leontosaurus</a></li> <li><a href="/wiki/Lepidodendron" title="Lepidodendron">Lepidodendron</a></li> <li><a href="/wiki/Lycideops" title="Lycideops">Lycideops</a></li> <li><a href="/wiki/Macroscelesaurus" title="Macroscelesaurus">Macroscelesaurus</a></li> <li><a href="/wiki/Moschorhinus" title="Moschorhinus">Moschorhinus</a></li> <li><a href="/wiki/Paliguana" title="Paliguana">Paliguana</a></li> <li><a href="/wiki/Pelictosuchus" title="Pelictosuchus">Pelictosuchus</a></li> <li><a href="/wiki/Polycynodon" title="Polycynodon">Polycynodon</a></li> <li><a href="/wiki/Pristerodon" title="Pristerodon">Pristerodon</a></li> <li><a href="/wiki/Pristerognathoides" class="mw-redirect" title="Pristerognathoides">Pristerognathoides</a></li> <li><a href="/wiki/Pristerognathus" title="Pristerognathus">Pristerognathus</a></li> <li><a href="/wiki/Procolophon" title="Procolophon">Procolophon</a></li> <li><a href="/wiki/Promoschorhynchus" title="Promoschorhynchus">Promoschorhynchus</a></li> <li><a href="/wiki/Scalopocephalus" title="Scalopocephalus">Scalopocephalus</a></li> <li><a href="/wiki/Silpholestes" title="Silpholestes">Silpholestes</a></li> <li><a href="/wiki/Tamboeria" title="Tamboeria">Tamboeria</a></li> <li><a href="/wiki/Uranocentrodon" title="Uranocentrodon">Uranocentrodon</a></li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Assemblages" scope="row" class="navbox-group" style="width:1%">Assemblages</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cistecephalus_Assemblage_Zone" title="Cistecephalus Assemblage Zone">Cistecephalus Assemblage Zone</a></li> <li><a href="/wiki/Daptocephalus_Assemblage_Zone" title="Daptocephalus Assemblage Zone">Daptocephalus Assemblage Zone</a></li> <li><a href="/wiki/Eodicynodon_Assemblage_Zone" title="Eodicynodon Assemblage Zone">Eodicynodon Assemblage Zone</a></li> <li><a href="/wiki/Pristerognathus_Assemblage_Zone" title="Pristerognathus Assemblage Zone">Pristerognathus Assemblage Zone</a></li> <li><a href="/wiki/Tapinocephalus_Assemblage_Zone" title="Tapinocephalus Assemblage Zone">Tapinocephalus Assemblage Zone</a></li> <li><a href="/wiki/Tropidostoma_Assemblage_Zone" title="Tropidostoma Assemblage Zone">Tropidostoma Assemblage Zone</a></li></ul> </div></td></tr></tbody></table><div> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Triassic</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Batrachosuchus" title="Batrachosuchus">Batrachosuchus</a></li> <li><a href="/wiki/Blikanasaurus" title="Blikanasaurus">Blikanasaurus</a></li> <li><a href="/wiki/Bolotridon" title="Bolotridon">Bolotridon</a></li> <li><a href="/wiki/Cynognathus" title="Cynognathus">Cynognathus</a></li> <li><a href="/wiki/Diademodon" title="Diademodon">Diademodon</a></li> <li><a href="/wiki/Dinanomodon" title="Dinanomodon">Dinanomodon</a></li> <li><a href="/wiki/Elliotherium" title="Elliotherium">Elliotherium</a></li> <li><a href="/wiki/Ericiolacerta" title="Ericiolacerta">Ericiolacerta</a></li> <li><a href="/wiki/Erythrosuchus" title="Erythrosuchus">Erythrosuchus</a></li> <li><a href="/wiki/Erythrotherium" title="Erythrotherium">Erythrotherium</a></li> <li><a href="/wiki/Eucnemesaurus" title="Eucnemesaurus">Eucnemesaurus</a></li> <li><a href="/wiki/Euskelosaurus" title="Euskelosaurus">Euskelosaurus</a></li> <li><a href="/wiki/Galesaurus" title="Galesaurus">Galesaurus</a></li> <li><a href="/wiki/Howesia" title="Howesia">Howesia</a></li> <li><a href="/wiki/Ictidosuchoides" title="Ictidosuchoides">Ictidosuchoides</a></li> <li><a href="/wiki/Kannemeyeria" title="Kannemeyeria">Kannemeyeria</a></li> <li><a href="/wiki/Lepidodendron" title="Lepidodendron">Lepidodendron</a></li> <li><a href="/wiki/Lydekkerina" title="Lydekkerina">Lydekkerina</a></li> <li><a class="mw-selflink selflink">Lystrosaurus</a></li> <li><a href="/wiki/Melanorosaurus" title="Melanorosaurus">Melanorosaurus</a></li> <li><a href="/wiki/Mesosuchus" title="Mesosuchus">Mesosuchus</a></li> <li><a href="/wiki/Microgomphodon" title="Microgomphodon">Microgomphodon</a></li> <li><a href="/wiki/Micropholis_(amphibian)" title="Micropholis (amphibian)">Micropholis</a></li> <li><a href="/wiki/Myosaurus" title="Myosaurus">Myosaurus</a></li> <li><a href="/wiki/Noteosuchus" title="Noteosuchus">Noteosuchus</a></li> <li><a href="/wiki/Olivierosuchus" title="Olivierosuchus">Olivierosuchus</a></li> <li><a href="/wiki/Orosaurus" title="Orosaurus">Orosaurus</a></li> <li><a href="/wiki/Pentasaurus" title="Pentasaurus">Pentasaurus</a></li> <li><a href="/wiki/Plateosauravus" title="Plateosauravus">Plateosauravus</a></li> <li><a href="/wiki/Platycraniellus" title="Platycraniellus">Platycraniellus</a></li> <li><a href="/wiki/Procolophon" title="Procolophon">Procolophon</a></li> <li><a href="/wiki/Progalesaurus" title="Progalesaurus">Progalesaurus</a></li> <li><a href="/wiki/Prolacerta" title="Prolacerta">Prolacerta</a></li> <li><a href="/wiki/Proterosuchus" title="Proterosuchus">Proterosuchus</a></li> <li><a href="/wiki/Ptychoceratodus" title="Ptychoceratodus">Ptychoceratodus</a></li> <li><a href="/wiki/Regisaurus" title="Regisaurus">Regisaurus</a></li> <li><a href="/wiki/Scaloposaurus" title="Scaloposaurus">Scaloposaurus</a></li> <li><a href="/wiki/Scymnosaurus" title="Scymnosaurus">Scymnosaurus</a></li> <li><a href="/wiki/Shansiodon" title="Shansiodon">Shansiodon</a></li> <li><a href="/wiki/Tetracynodon" title="Tetracynodon">Tetracynodon</a></li> <li><a href="/wiki/Thrinaxodon" title="Thrinaxodon">Thrinaxodon</a></li> <li><a href="/wiki/Trirachodon" title="Trirachodon">Trirachodon</a></li> <li><a href="/wiki/Ufudocyclops" title="Ufudocyclops">Ufudocyclops</a></li> <li><a href="/wiki/Zorillodontops" title="Zorillodontops">Zorillodontops</a></li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Assemblages" scope="row" class="navbox-group" style="width:1%">Assemblages</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cynognathus_Assemblage_Zone" title="Cynognathus Assemblage Zone">Cynognathus Assemblage Zone</a></li> <li><a href="/wiki/Lystrosaurus_Assemblage_Zone" title="Lystrosaurus Assemblage Zone">Lystrosaurus Assemblage Zone</a></li></ul> </div></td></tr></tbody></table><div> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Jurassic</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Aardonyx" title="Aardonyx">Aardonyx</a></li> <li><a href="/wiki/Abrictosaurus" title="Abrictosaurus">Abrictosaurus</a></li> <li><a href="/wiki/Antetonitrus" title="Antetonitrus">Antetonitrus</a></li> <li><a href="/wiki/Arcusaurus" title="Arcusaurus">Arcusaurus</a></li> <li><a href="/wiki/Coelophysis" title="Coelophysis">Coelophysis</a></li> <li><a href="/wiki/Coelophysis_rhodesiensis" class="mw-redirect" title="Coelophysis rhodesiensis">Coelophysis rhodesiensis</a></li> <li><a href="/wiki/Dracovenator" title="Dracovenator">Dracovenator</a></li> <li><a href="/wiki/Eocursor" title="Eocursor">Eocursor</a></li> <li><a href="/wiki/Erythrotherium" title="Erythrotherium">Erythrotherium</a></li> <li><a href="/wiki/Fabrosaurus" title="Fabrosaurus">Fabrosaurus</a></li> <li><a href="/wiki/Geranosaurus" title="Geranosaurus">Geranosaurus</a></li> <li><a href="/wiki/Gigantoscelus" title="Gigantoscelus">Gigantoscelus</a></li> <li><a href="/wiki/Gryponyx" title="Gryponyx">Gryponyx</a></li> <li><a href="/wiki/Gyposaurus" title="Gyposaurus">Gyposaurus</a></li> <li><a href="/wiki/Heterodontosaurus" title="Heterodontosaurus">Heterodontosaurus</a></li> <li><a href="/wiki/Ledumahadi" title="Ledumahadi">Ledumahadi</a></li> <li><a href="/wiki/Lesothosaurus" title="Lesothosaurus">Lesothosaurus</a></li> <li><a href="/wiki/Litargosuchus" title="Litargosuchus">Litargosuchus</a></li> <li><a href="/wiki/Lycorhinus" title="Lycorhinus">Lycorhinus</a></li> <li><a href="/wiki/Massospondylus" title="Massospondylus">Massospondylus</a></li> <li><a href="/wiki/Ngwevu" title="Ngwevu">Ngwevu</a></li> <li><a href="/wiki/Pegomastax" title="Pegomastax">Pegomastax</a></li> <li><a href="/wiki/Ptychoceratodus" title="Ptychoceratodus">Ptychoceratodus</a></li> <li><a href="/wiki/Pulanesaura" title="Pulanesaura">Pulanesaura</a></li> <li><a href="/wiki/Sefapanosaurus" title="Sefapanosaurus">Sefapanosaurus</a></li> <li><a href="/wiki/Sphenosuchus" title="Sphenosuchus">Sphenosuchus</a></li> <li><a href="/wiki/Tritheledon" title="Tritheledon">Tritheledon</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Cretaceous</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Acanthohoplites" title="Acanthohoplites">Acanthohoplites</a></li> <li><a href="/wiki/Algoasaurus" title="Algoasaurus">Algoasaurus</a></li> <li><a href="/wiki/Baculites" title="Baculites">Baculites</a></li> <li><a href="/wiki/Kangnasaurus" title="Kangnasaurus">Kangnasaurus</a></li> <li><a href="/wiki/Macroscaphites" title="Macroscaphites">Macroscaphites</a></li> <li><a href="/wiki/Nqwebasaurus" title="Nqwebasaurus">Nqwebasaurus</a></li> <li><a href="/wiki/Paranthodon" title="Paranthodon">Paranthodon</a></li> <li><a href="/wiki/Ptychoceratodus" title="Ptychoceratodus">Ptychoceratodus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Paleogene</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Nummulite" title="Nummulite">Nummulite</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Neogene</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Africanacetus" title="Africanacetus">Africanacetus</a></li> <li><a href="/wiki/Nummulite" title="Nummulite">Nummulite</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Pleistocene</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Boskop_Man" title="Boskop Man">Boskop Man</a></li> <li><a href="/wiki/Homo_naledi" title="Homo naledi">Homo naledi</a></li> <li><a href="/wiki/Panthera_shawi" title="Panthera shawi">Panthera shawi</a></li> <li><a href="/wiki/Vulpes_skinneri" title="Vulpes skinneri">Vulpes skinneri</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cradle_of_Humankind" title="Cradle of Humankind">Cradle of Humankind</a></li> <li><a href="/wiki/Planolites" title="Planolites">Planolites</a></li> <li><a href="/wiki/Scylacorhinus" class="mw-redirect" title="Scylacorhinus">Scylacorhinus</a></li> <li><a href="/wiki/West_Coast_Fossil_Park" title="West Coast Fossil Park">West Coast Fossil Park</a></li> <li><a href="/wiki/Waterloo_Farm_lagerst%C3%A4tte" title="Waterloo Farm lagerstätte">Waterloo Farm lagerstätte</a></li></ul> </div></td></tr></tbody></table><div> </div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Research_and_administration" style="font-size:114%;margin:0 4em">Research and administration</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Council_for_Geoscience" title="Council for Geoscience">Council for Geoscience</a></li> <li><a href="/wiki/Evolutionary_Studies_Institute" title="Evolutionary Studies Institute">Evolutionary Studies Institute</a></li> <li><a href="/wiki/Geological_Commission_of_the_Cape_of_Good_Hope" title="Geological Commission of the Cape of Good Hope">Geological Commission of the Cape of Good Hope</a></li> <li><a href="/wiki/Geological_Society_of_South_Africa" title="Geological Society of South Africa">Geological Society of South Africa</a></li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Geologists</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Hans_Merensky" title="Hans Merensky">Hans Merensky</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Paleontologists</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Andrew_Geddes_Bain" title="Andrew Geddes Bain">Andrew Geddes Bain</a></li> <li><a href="/wiki/Lieuwe_Dirk_Boonstra" title="Lieuwe Dirk Boonstra">Lieuwe Dirk Boonstra</a></li> <li><a href="/wiki/Charles_Kimberlin_Brain" title="Charles Kimberlin Brain">Charles Kimberlin Brain</a></li> <li><a href="/wiki/Robert_Broom" title="Robert Broom">Robert Broom</a></li> <li><a href="/wiki/Alfred_Brown_(palaeontologist)" title="Alfred Brown (palaeontologist)">Alfred Brown</a></li> <li><a href="/wiki/Anusuya_Chinsamy-Turan" title="Anusuya Chinsamy-Turan">Anusuya Chinsamy-Turan</a></li> <li><a href="/wiki/H._Basil_S._Cooke" title="H. Basil S. Cooke">H. Basil S. Cooke</a></li> <li><a href="/wiki/Alfred_W._Crompton" title="Alfred W. Crompton">Alfred W. Crompton</a></li> <li><a href="/wiki/Daniel_Rossouw_Kannemeyer" title="Daniel Rossouw Kannemeyer">Daniel Rossouw Kannemeyer</a></li> <li><a href="/wiki/Andre_Keyser" title="Andre Keyser">Andre Keyser</a></li> <li><a href="/wiki/James_Kitching" title="James Kitching">James Kitching</a></li> <li><a href="/wiki/John_Talbot_Robinson" title="John Talbot Robinson">John Talbot Robinson</a></li> <li><a href="/wiki/George_William_Stow" title="George William Stow">George William Stow</a></li> <li><a href="/wiki/Eduard_Meine_van_Zinderen-Bakker" title="Eduard Meine van Zinderen-Bakker">Eduard Meine van Zinderen-Bakker</a></li></ul> </div></td></tr></tbody></table><div> </div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Regional_articles" style="font-size:114%;margin:0 4em">Regional articles</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Geology_of_Cape_Town" title="Geology of Cape Town">Geology of Cape Town</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Other" style="font-size:114%;margin:0 4em">Other</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pothole_(geology)" title="Pothole (geology)">Pothole (geology)</a></li> <li><a href="/wiki/Steep_structure" title="Steep structure">Steep structure</a></li> <li><a href="/wiki/Swazian" title="Swazian">Swazian</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2" style="font-weight: bold;"><div> <ul><li><b><span class="nowrap"><span class="noviewer" typeof="mw:File"><a href="/wiki/File:WikiProject_Geology.svg" class="mw-file-description"><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e7/WikiProject_Geology.svg/16px-WikiProject_Geology.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/e7/WikiProject_Geology.svg/24px-WikiProject_Geology.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/e7/WikiProject_Geology.svg/32px-WikiProject_Geology.svg.png 2x" data-file-width="327" data-file-height="328" /></a></span> </span><a href="/wiki/Portal:Geology" title="Portal:Geology">Geology portal</a></b></li> <li><b><span class="nowrap"><span class="mw-image-border noviewer" 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1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <b><a href="/wiki/Category:Geology_of_South_Africa" title="Category:Geology of South Africa">Category</a></b></li> <li><span class="noviewer" typeof="mw:File"><span title="List-Class article"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/d/db/Symbol_list_class.svg/16px-Symbol_list_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/d/db/Symbol_list_class.svg/23px-Symbol_list_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/d/db/Symbol_list_class.svg/31px-Symbol_list_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span></li> <li><b><a href="/wiki/Glossary_of_geology" title="Glossary of geology">Glossary</a></b></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Lystrosaurus</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q131680" class="extiw" title="wikidata:Q131680">Q131680</a></span></span></span></li> <li><span style="white-space:nowrap;">BioLib: <span class="uid"><a rel="nofollow" class="external text" href="https://www.biolib.cz/en/taxon/id417226">417226</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/4529455">4529455</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/4817808">4817808</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&id=1195743">1195743</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=4126421">4126421</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=39049">39049</a></span></span></li></ul> </div></td></tr></tbody></table></div> <!-- NewPP limit report Parsed by mw‐web.codfw.main‐7fc47fc68d‐229k2 Cached time: 20241128193911 Cache expiry: 2592000 Reduced expiry: false Complications: [vary‐revision‐sha1, show‐toc] CPU time usage: 1.214 seconds Real time usage: 1.500 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