CINXE.COM

<!DOCTYPE html> <html class="client-nojs vector-feature-language-in-header-enabled vector-feature-language-in-main-page-header-disabled vector-feature-sticky-header-disabled vector-feature-page-tools-pinned-disabled vector-feature-toc-pinned-clientpref-1 vector-feature-main-menu-pinned-disabled vector-feature-limited-width-clientpref-1 vector-feature-limited-width-content-enabled vector-feature-custom-font-size-clientpref-1 vector-feature-appearance-pinned-clientpref-1 vector-feature-night-mode-enabled skin-theme-clientpref-day vector-toc-available" lang="en" dir="ltr"> <head> <meta charset="UTF-8"> <title>Dynastinae - Wikipedia</title> <script>(function(){var className="client-js vector-feature-language-in-header-enabled vector-feature-language-in-main-page-header-disabled vector-feature-sticky-header-disabled vector-feature-page-tools-pinned-disabled vector-feature-toc-pinned-clientpref-1 vector-feature-main-menu-pinned-disabled vector-feature-limited-width-clientpref-1 vector-feature-limited-width-content-enabled vector-feature-custom-font-size-clientpref-1 vector-feature-appearance-pinned-clientpref-1 vector-feature-night-mode-enabled skin-theme-clientpref-day vector-toc-available";var cookie=document.cookie.match(/(?:^|; )enwikimwclientpreferences=([^;]+)/);if(cookie){cookie[1].split('%2C').forEach(function(pref){className=className.replace(new RegExp('(^| )'+pref.replace(/-clientpref-\w+$|[^\w-]+/g,'')+'-clientpref-\\w+( |$)'),'$1'+pref+'$2');});}document.documentElement.className=className;}());RLCONF={"wgBreakFrames":false,"wgSeparatorTransformTable":["",""],"wgDigitTransformTable":["",""],"wgDefaultDateFormat":"dmy", "wgMonthNames":["","January","February","March","April","May","June","July","August","September","October","November","December"],"wgRequestId":"14a7334e-9437-427e-bdf4-f21552d4dadf","wgCanonicalNamespace":"","wgCanonicalSpecialPageName":false,"wgNamespaceNumber":0,"wgPageName":"Dynastinae","wgTitle":"Dynastinae","wgCurRevisionId":1246203154,"wgRevisionId":1246203154,"wgArticleId":2143758,"wgIsArticle":true,"wgIsRedirect":false,"wgAction":"view","wgUserName":null,"wgUserGroups":["*"],"wgCategories":["Articles with short description","Short description matches Wikidata","Articles lacking in-text citations from November 2015","All articles lacking in-text citations","Articles with 'species' microformats","Commons category link is on Wikidata","Dynastinae","Insects as food","Insects in culture","Extant Eocene first appearances"],"wgPageViewLanguage":"en","wgPageContentLanguage":"en","wgPageContentModel":"wikitext","wgRelevantPageName":"Dynastinae","wgRelevantArticleId":2143758, "wgIsProbablyEditable":true,"wgRelevantPageIsProbablyEditable":true,"wgRestrictionEdit":[],"wgRestrictionMove":[],"wgRedirectedFrom":"Unicorn_beetle","wgNoticeProject":"wikipedia","wgCiteReferencePreviewsActive":false,"wgFlaggedRevsParams":{"tags":{"status":{"levels":1}}},"wgMediaViewerOnClick":true,"wgMediaViewerEnabledByDefault":true,"wgPopupsFlags":0,"wgVisualEditor":{"pageLanguageCode":"en","pageLanguageDir":"ltr","pageVariantFallbacks":"en"},"wgMFDisplayWikibaseDescriptions":{"search":true,"watchlist":true,"tagline":false,"nearby":true},"wgWMESchemaEditAttemptStepOversample":false,"wgWMEPageLength":10000,"wgInternalRedirectTargetUrl":"/wiki/Dynastinae","wgRelatedArticlesCompat":[],"wgEditSubmitButtonLabelPublish":true,"wgULSPosition":"interlanguage","wgULSisCompactLinksEnabled":false,"wgVector2022LanguageInHeader":true,"wgULSisLanguageSelectorEmpty":false,"wgWikibaseItemId":"Q641142","wgCheckUserClientHintsHeadersJsApi":["brands","architecture","bitness","fullVersionList","mobile" ,"model","platform","platformVersion"],"GEHomepageSuggestedEditsEnableTopics":true,"wgGETopicsMatchModeEnabled":false,"wgGEStructuredTaskRejectionReasonTextInputEnabled":false,"wgGELevelingUpEnabledForUser":false};RLSTATE={"ext.globalCssJs.user.styles":"ready","site.styles":"ready","user.styles":"ready","ext.globalCssJs.user":"ready","user":"ready","user.options":"loading","ext.cite.styles":"ready","skins.vector.search.codex.styles":"ready","skins.vector.styles":"ready","skins.vector.icons":"ready","jquery.makeCollapsible.styles":"ready","ext.wikimediamessages.styles":"ready","ext.visualEditor.desktopArticleTarget.noscript":"ready","ext.uls.interlanguage":"ready","wikibase.client.init":"ready","ext.wikimediaBadges":"ready"};RLPAGEMODULES=["mediawiki.action.view.redirect","ext.cite.ux-enhancements","mediawiki.page.media","ext.scribunto.logs","site","mediawiki.page.ready","jquery.makeCollapsible","mediawiki.toc","skins.vector.js","ext.centralNotice.geoIP","ext.centralNotice.startUp", "ext.gadget.ReferenceTooltips","ext.gadget.switcher","ext.urlShortener.toolbar","ext.centralauth.centralautologin","mmv.bootstrap","ext.popups","ext.visualEditor.desktopArticleTarget.init","ext.visualEditor.targetLoader","ext.echo.centralauth","ext.eventLogging","ext.wikimediaEvents","ext.navigationTiming","ext.uls.interface","ext.cx.eventlogging.campaigns","ext.cx.uls.quick.actions","wikibase.client.vector-2022","ext.checkUser.clientHints","ext.growthExperiments.SuggestedEditSession","wikibase.sidebar.tracking"];</script> <script>(RLQ=window.RLQ||[]).push(function(){mw.loader.impl(function(){return["user.options@12s5i",function($,jQuery,require,module){mw.user.tokens.set({"patrolToken":"+\\","watchToken":"+\\","csrfToken":"+\\"}); }];});});</script> <link rel="stylesheet" href="/w/load.php?lang=en&modules=ext.cite.styles%7Cext.uls.interlanguage%7Cext.visualEditor.desktopArticleTarget.noscript%7Cext.wikimediaBadges%7Cext.wikimediamessages.styles%7Cjquery.makeCollapsible.styles%7Cskins.vector.icons%2Cstyles%7Cskins.vector.search.codex.styles%7Cwikibase.client.init&only=styles&skin=vector-2022"> <script async="" src="/w/load.php?lang=en&modules=startup&only=scripts&raw=1&skin=vector-2022"></script> <meta name="ResourceLoaderDynamicStyles" content=""> <link rel="stylesheet" href="/w/load.php?lang=en&modules=site.styles&only=styles&skin=vector-2022"> <meta name="generator" content="MediaWiki 1.44.0-wmf.5"> <meta name="referrer" content="origin"> <meta name="referrer" content="origin-when-cross-origin"> <meta name="robots" content="max-image-preview:standard"> <meta name="format-detection" content="telephone=no"> <meta property="og:image" content="https://upload.wikimedia.org/wikipedia/commons/thumb/e/ea/Dynastinae.jpg/1200px-Dynastinae.jpg"> <meta property="og:image:width" content="1200"> <meta property="og:image:height" content="1230"> <meta property="og:image" content="https://upload.wikimedia.org/wikipedia/commons/thumb/e/ea/Dynastinae.jpg/800px-Dynastinae.jpg"> <meta property="og:image:width" content="800"> <meta property="og:image:height" content="820"> <meta property="og:image" content="https://upload.wikimedia.org/wikipedia/commons/thumb/e/ea/Dynastinae.jpg/640px-Dynastinae.jpg"> <meta property="og:image:width" content="640"> <meta property="og:image:height" content="656"> <meta name="viewport" content="width=1120"> <meta property="og:title" content="Dynastinae - Wikipedia"> <meta property="og:type" content="website"> <link rel="preconnect" href="//upload.wikimedia.org"> <link rel="alternate" media="only screen and (max-width: 640px)" href="//en.m.wikipedia.org/wiki/Dynastinae"> <link rel="alternate" type="application/x-wiki" title="Edit this page" href="/w/index.php?title=Dynastinae&action=edit"> <link rel="apple-touch-icon" href="/static/apple-touch/wikipedia.png"> <link rel="icon" href="/static/favicon/wikipedia.ico"> <link rel="search" type="application/opensearchdescription+xml" href="/w/rest.php/v1/search" title="Wikipedia (en)"> <link rel="EditURI" type="application/rsd+xml" href="//en.wikipedia.org/w/api.php?action=rsd"> <link rel="canonical" href="https://en.wikipedia.org/wiki/Dynastinae"> <link rel="license" href="https://creativecommons.org/licenses/by-sa/4.0/deed.en"> <link rel="alternate" type="application/atom+xml" title="Wikipedia Atom feed" href="/w/index.php?title=Special:RecentChanges&feed=atom"> <link rel="dns-prefetch" href="//meta.wikimedia.org" /> <link rel="dns-prefetch" href="//login.wikimedia.org"> </head> <body class="skin--responsive skin-vector skin-vector-search-vue mediawiki ltr sitedir-ltr mw-hide-empty-elt ns-0 ns-subject mw-editable page-Dynastinae rootpage-Dynastinae skin-vector-2022 action-view"><a class="mw-jump-link" href="#bodyContent">Jump to content</a> <div class="vector-header-container"> <header class="vector-header mw-header"> <div class="vector-header-start"> <nav class="vector-main-menu-landmark" aria-label="Site"> <div id="vector-main-menu-dropdown" class="vector-dropdown vector-main-menu-dropdown vector-button-flush-left vector-button-flush-right" > <input type="checkbox" id="vector-main-menu-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-main-menu-dropdown" class="vector-dropdown-checkbox " aria-label="Main menu" > <label id="vector-main-menu-dropdown-label" for="vector-main-menu-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-menu mw-ui-icon-wikimedia-menu"></span> <span class="vector-dropdown-label-text">Main menu</span> </label> <div class="vector-dropdown-content"> <div id="vector-main-menu-unpinned-container" class="vector-unpinned-container"> <div id="vector-main-menu" class="vector-main-menu vector-pinnable-element"> <div class="vector-pinnable-header vector-main-menu-pinnable-header vector-pinnable-header-unpinned" data-feature-name="main-menu-pinned" data-pinnable-element-id="vector-main-menu" data-pinned-container-id="vector-main-menu-pinned-container" data-unpinned-container-id="vector-main-menu-unpinned-container" > <div class="vector-pinnable-header-label">Main menu</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-main-menu.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-main-menu.unpin">hide</button> </div> <div id="p-navigation" class="vector-menu mw-portlet mw-portlet-navigation" > <div class="vector-menu-heading"> Navigation </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="n-mainpage-description" class="mw-list-item"><a href="/wiki/Main_Page" title="Visit the main page [z]" accesskey="z"><span>Main page</span></a></li><li id="n-contents" class="mw-list-item"><a href="/wiki/Wikipedia:Contents" title="Guides to browsing Wikipedia"><span>Contents</span></a></li><li id="n-currentevents" class="mw-list-item"><a href="/wiki/Portal:Current_events" title="Articles related to current events"><span>Current events</span></a></li><li id="n-randompage" class="mw-list-item"><a href="/wiki/Special:Random" title="Visit a randomly selected article [x]" accesskey="x"><span>Random article</span></a></li><li id="n-aboutsite" class="mw-list-item"><a href="/wiki/Wikipedia:About" title="Learn about Wikipedia and how it works"><span>About Wikipedia</span></a></li><li id="n-contactpage" class="mw-list-item"><a href="//en.wikipedia.org/wiki/Wikipedia:Contact_us" title="How to contact Wikipedia"><span>Contact us</span></a></li> </ul> </div> </div> <div id="p-interaction" class="vector-menu mw-portlet mw-portlet-interaction" > <div class="vector-menu-heading"> Contribute </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="n-help" class="mw-list-item"><a href="/wiki/Help:Contents" title="Guidance on how to use and edit Wikipedia"><span>Help</span></a></li><li id="n-introduction" class="mw-list-item"><a href="/wiki/Help:Introduction" title="Learn how to edit Wikipedia"><span>Learn to edit</span></a></li><li id="n-portal" class="mw-list-item"><a href="/wiki/Wikipedia:Community_portal" title="The hub for editors"><span>Community portal</span></a></li><li id="n-recentchanges" class="mw-list-item"><a href="/wiki/Special:RecentChanges" title="A list of recent changes to Wikipedia [r]" accesskey="r"><span>Recent changes</span></a></li><li id="n-upload" class="mw-list-item"><a href="/wiki/Wikipedia:File_upload_wizard" title="Add images or other media for use on Wikipedia"><span>Upload file</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> <a href="/wiki/Main_Page" class="mw-logo"> <img class="mw-logo-icon" src="/static/images/icons/wikipedia.png" alt="" aria-hidden="true" height="50" width="50"> <span class="mw-logo-container skin-invert"> <img class="mw-logo-wordmark" alt="Wikipedia" src="/static/images/mobile/copyright/wikipedia-wordmark-en.svg" style="width: 7.5em; height: 1.125em;"> <img class="mw-logo-tagline" alt="The Free Encyclopedia" src="/static/images/mobile/copyright/wikipedia-tagline-en.svg" width="117" height="13" style="width: 7.3125em; height: 0.8125em;"> </span> </a> </div> <div class="vector-header-end"> <div id="p-search" role="search" class="vector-search-box-vue vector-search-box-collapses vector-search-box-show-thumbnail vector-search-box-auto-expand-width vector-search-box"> <a href="/wiki/Special:Search" class="cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only search-toggle" title="Search Wikipedia [f]" accesskey="f"><span class="vector-icon mw-ui-icon-search mw-ui-icon-wikimedia-search"></span> <span>Search</span> </a> <div class="vector-typeahead-search-container"> <div class="cdx-typeahead-search cdx-typeahead-search--show-thumbnail cdx-typeahead-search--auto-expand-width"> <form action="/w/index.php" id="searchform" class="cdx-search-input cdx-search-input--has-end-button"> <div id="simpleSearch" class="cdx-search-input__input-wrapper" data-search-loc="header-moved"> <div class="cdx-text-input cdx-text-input--has-start-icon"> <input class="cdx-text-input__input" type="search" name="search" placeholder="Search Wikipedia" aria-label="Search Wikipedia" autocapitalize="sentences" title="Search Wikipedia [f]" accesskey="f" id="searchInput" > <span class="cdx-text-input__icon cdx-text-input__start-icon"></span> </div> <input type="hidden" name="title" value="Special:Search"> </div> <button class="cdx-button cdx-search-input__end-button">Search</button> </form> </div> </div> </div> <nav class="vector-user-links vector-user-links-wide" aria-label="Personal tools"> <div class="vector-user-links-main"> <div id="p-vector-user-menu-preferences" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <div id="p-vector-user-menu-userpage" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <nav class="vector-appearance-landmark" aria-label="Appearance"> <div id="vector-appearance-dropdown" class="vector-dropdown " title="Change the appearance of the page's font size, width, and color" > <input type="checkbox" id="vector-appearance-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-appearance-dropdown" class="vector-dropdown-checkbox " aria-label="Appearance" > <label id="vector-appearance-dropdown-label" for="vector-appearance-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-appearance mw-ui-icon-wikimedia-appearance"></span> <span class="vector-dropdown-label-text">Appearance</span> </label> <div class="vector-dropdown-content"> <div id="vector-appearance-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <div id="p-vector-user-menu-notifications" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <div id="p-vector-user-menu-overflow" class="vector-menu mw-portlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-sitesupport-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="https://donate.wikimedia.org/wiki/Special:FundraiserRedirector?utm_source=donate&utm_medium=sidebar&utm_campaign=C13_en.wikipedia.org&uselang=en" class=""><span>Donate</span></a> </li> <li id="pt-createaccount-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="/w/index.php?title=Special:CreateAccount&returnto=Dynastinae" title="You are encouraged to create an account and log in; however, it is not mandatory" class=""><span>Create account</span></a> </li> <li id="pt-login-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="/w/index.php?title=Special:UserLogin&returnto=Dynastinae" title="You're encouraged to log in; however, it's not mandatory. [o]" accesskey="o" class=""><span>Log in</span></a> </li> </ul> </div> </div> </div> <div id="vector-user-links-dropdown" class="vector-dropdown vector-user-menu vector-button-flush-right vector-user-menu-logged-out" title="Log in and more options" > <input type="checkbox" id="vector-user-links-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-user-links-dropdown" class="vector-dropdown-checkbox " aria-label="Personal tools" > <label id="vector-user-links-dropdown-label" for="vector-user-links-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-ellipsis mw-ui-icon-wikimedia-ellipsis"></span> <span class="vector-dropdown-label-text">Personal tools</span> </label> <div class="vector-dropdown-content"> <div id="p-personal" class="vector-menu mw-portlet mw-portlet-personal user-links-collapsible-item" title="User menu" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-sitesupport" class="user-links-collapsible-item mw-list-item"><a href="https://donate.wikimedia.org/wiki/Special:FundraiserRedirector?utm_source=donate&utm_medium=sidebar&utm_campaign=C13_en.wikipedia.org&uselang=en"><span>Donate</span></a></li><li id="pt-createaccount" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=Special:CreateAccount&returnto=Dynastinae" title="You are encouraged to create an account and log in; however, it is not mandatory"><span class="vector-icon mw-ui-icon-userAdd mw-ui-icon-wikimedia-userAdd"></span> <span>Create account</span></a></li><li id="pt-login" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=Special:UserLogin&returnto=Dynastinae" title="You're encouraged to log in; however, it's not mandatory. [o]" accesskey="o"><span class="vector-icon mw-ui-icon-logIn mw-ui-icon-wikimedia-logIn"></span> <span>Log in</span></a></li> </ul> </div> </div> <div id="p-user-menu-anon-editor" class="vector-menu mw-portlet mw-portlet-user-menu-anon-editor" > <div class="vector-menu-heading"> Pages for logged out editors <a href="/wiki/Help:Introduction" aria-label="Learn more about editing"><span>learn more</span></a> </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-anoncontribs" class="mw-list-item"><a href="/wiki/Special:MyContributions" title="A list of edits made from this IP address [y]" accesskey="y"><span>Contributions</span></a></li><li id="pt-anontalk" class="mw-list-item"><a href="/wiki/Special:MyTalk" title="Discussion about edits from this IP address [n]" accesskey="n"><span>Talk</span></a></li> </ul> </div> </div> </div> </div> </nav> </div> </header> </div> <div class="mw-page-container"> <div class="mw-page-container-inner"> <div class="vector-sitenotice-container"> <div id="siteNotice"></div> </div> <div class="vector-column-start"> <div class="vector-main-menu-container"> <div id="mw-navigation"> <nav id="mw-panel" class="vector-main-menu-landmark" aria-label="Site"> <div id="vector-main-menu-pinned-container" class="vector-pinned-container"> </div> </nav> </div> </div> <div class="vector-sticky-pinned-container"> <nav id="mw-panel-toc" aria-label="Contents" data-event-name="ui.sidebar-toc" class="mw-table-of-contents-container vector-toc-landmark"> <div id="vector-toc-pinned-container" class="vector-pinned-container"> <div id="vector-toc" class="vector-toc vector-pinnable-element"> <div class="vector-pinnable-header vector-toc-pinnable-header vector-pinnable-header-pinned" data-feature-name="toc-pinned" data-pinnable-element-id="vector-toc" > <h2 class="vector-pinnable-header-label">Contents</h2> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-toc.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-toc.unpin">hide</button> </div> <ul class="vector-toc-contents" id="mw-panel-toc-list"> <li id="toc-mw-content-text" class="vector-toc-list-item vector-toc-level-1"> <a href="#" class="vector-toc-link"> <div class="vector-toc-text">(Top)</div> </a> </li> <li id="toc-Description" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Description"> <div class="vector-toc-text"> <span class="vector-toc-numb">1</span> <span>Description</span> </div> </a> <ul id="toc-Description-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Ecology" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Ecology"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Ecology</span> </div> </a> <button aria-controls="toc-Ecology-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Ecology subsection</span> </button> <ul id="toc-Ecology-sublist" class="vector-toc-list"> <li id="toc-Mating" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Mating"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.1</span> <span>Mating</span> </div> </a> <ul id="toc-Mating-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-As_pests" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#As_pests"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>As pests</span> </div> </a> <ul id="toc-As_pests-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Uses" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Uses"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Uses</span> </div> </a> <ul id="toc-Uses-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Tribes_with_selected_genera_and_species" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Tribes_with_selected_genera_and_species"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>Tribes with selected genera and species</span> </div> </a> <button aria-controls="toc-Tribes_with_selected_genera_and_species-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Tribes with selected genera and species subsection</span> </button> <ul id="toc-Tribes_with_selected_genera_and_species-sublist" class="vector-toc-list"> <li id="toc-Agaocephalini" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Agaocephalini"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.1</span> <span>Agaocephalini</span> </div> </a> <ul id="toc-Agaocephalini-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Cyclocephalini" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Cyclocephalini"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.2</span> <span>Cyclocephalini</span> </div> </a> <ul id="toc-Cyclocephalini-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Dynastini" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Dynastini"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.3</span> <span>Dynastini</span> </div> </a> <ul id="toc-Dynastini-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Hexodontini" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Hexodontini"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.4</span> <span>Hexodontini</span> </div> </a> <ul id="toc-Hexodontini-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Oryctini" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Oryctini"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.5</span> <span>Oryctini</span> </div> </a> <ul id="toc-Oryctini-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Oryctoderini" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Oryctoderini"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.6</span> <span>Oryctoderini</span> </div> </a> <ul id="toc-Oryctoderini-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Pentodontini" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Pentodontini"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.7</span> <span>Pentodontini</span> </div> </a> <ul id="toc-Pentodontini-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Phileurini" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Phileurini"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.8</span> <span>Phileurini</span> </div> </a> <ul id="toc-Phileurini-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Notes" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Notes"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>Notes</span> </div> </a> <ul id="toc-Notes-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Further_reading" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Further_reading"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>Further reading</span> </div> </a> <ul id="toc-Further_reading-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">8</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Toggle the table of contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Dynastinae</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 41 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-41" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">41 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-af mw-list-item"><a href="https://af.wikipedia.org/wiki/Renosterkewer" title="Renosterkewer – Afrikaans" lang="af" hreflang="af" data-title="Renosterkewer" data-language-autonym="Afrikaans" data-language-local-name="Afrikaans" class="interlanguage-link-target"><span>Afrikaans</span></a></li><li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B9%D8%B1%D9%8A%D9%82%D8%B7%D8%A9" title="عريقطة – Arabic" lang="ar" hreflang="ar" data-title="عريقطة" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-ast mw-list-item"><a href="https://ast.wikipedia.org/wiki/Dynastinae" title="Dynastinae – Asturian" lang="ast" hreflang="ast" data-title="Dynastinae" data-language-autonym="Asturianu" data-language-local-name="Asturian" class="interlanguage-link-target"><span>Asturianu</span></a></li><li class="interlanguage-link interwiki-zh-min-nan mw-list-item"><a href="https://zh-min-nan.wikipedia.org/wiki/Tau-th%C3%A2ng" title="Tau-thâng – Minnan" lang="nan" hreflang="nan" data-title="Tau-thâng" data-language-autonym="閩南語 / Bân-lâm-gú" data-language-local-name="Minnan" class="interlanguage-link-target"><span>閩南語 / Bân-lâm-gú</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Dinastins" title="Dinastins – Catalan" lang="ca" hreflang="ca" data-title="Dinastins" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-ceb mw-list-item"><a href="https://ceb.wikipedia.org/wiki/Dynastidae" title="Dynastidae – Cebuano" lang="ceb" hreflang="ceb" data-title="Dynastidae" data-language-autonym="Cebuano" data-language-local-name="Cebuano" class="interlanguage-link-target"><span>Cebuano</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Nosoro%C5%BE%C3%ADci" title="Nosorožíci – Czech" lang="cs" hreflang="cs" data-title="Nosorožíci" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/N%C3%A6sehornsbiller" title="Næsehornsbiller – Danish" lang="da" hreflang="da" data-title="Næsehornsbiller" data-language-autonym="Dansk" data-language-local-name="Danish" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Riesenk%C3%A4fer" title="Riesenkäfer – German" lang="de" hreflang="de" data-title="Riesenkäfer" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-nv mw-list-item"><a href="https://nv.wikipedia.org/wiki/Ch%C3%A9%C5%82chaa%CA%BCtsoh" title="Chéłchaaʼtsoh – Navajo" lang="nv" hreflang="nv" data-title="Chéłchaaʼtsoh" data-language-autonym="Diné bizaad" data-language-local-name="Navajo" class="interlanguage-link-target"><span>Diné bizaad</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Dynastinae" title="Dynastinae – Spanish" lang="es" hreflang="es" data-title="Dynastinae" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%B3%D9%88%D8%B3%DA%A9%E2%80%8C%D9%87%D8%A7%DB%8C_%DA%A9%D8%B1%DA%AF%D8%AF%D9%86%DB%8C" title="سوسک‌های کرگدنی – Persian" lang="fa" hreflang="fa" data-title="سوسک‌های کرگدنی" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Dynastinae" title="Dynastinae – French" lang="fr" hreflang="fr" data-title="Dynastinae" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Dynastinae" title="Dynastinae – Galician" lang="gl" hreflang="gl" data-title="Dynastinae" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%9E%A5%EC%88%98%ED%92%8D%EB%8E%85%EC%9D%B4%EC%95%84%EA%B3%BC" title="장수풍뎅이아과 – Korean" lang="ko" hreflang="ko" data-title="장수풍뎅이아과" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Rinocero-skarabeo" title="Rinocero-skarabeo – Ido" lang="io" hreflang="io" data-title="Rinocero-skarabeo" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Kumbang_badak" title="Kumbang badak – Indonesian" lang="id" hreflang="id" data-title="Kumbang badak" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Dynastinae" title="Dynastinae – Italian" lang="it" hreflang="it" data-title="Dynastinae" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-kn mw-list-item"><a href="https://kn.wikipedia.org/wiki/%E0%B2%96%E0%B2%A1%E0%B3%8D%E0%B2%97_%E0%B2%9C%E0%B3%80%E0%B2%B0%E0%B3%81%E0%B2%82%E0%B2%A1%E0%B3%86%E0%B2%97%E0%B2%B3%E0%B3%81" title="ಖಡ್ಗ ಜೀರುಂಡೆಗಳು – Kannada" lang="kn" hreflang="kn" data-title="ಖಡ್ಗ ಜೀರುಂಡೆಗಳು" data-language-autonym="ಕನ್ನಡ" data-language-local-name="Kannada" class="interlanguage-link-target"><span>ಕನ್ನಡ</span></a></li><li class="interlanguage-link interwiki-pam mw-list-item"><a href="https://pam.wikipedia.org/wiki/Uang" title="Uang – Pampanga" lang="pam" hreflang="pam" data-title="Uang" data-language-autonym="Kapampangan" data-language-local-name="Pampanga" class="interlanguage-link-target"><span>Kapampangan</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Chonga" title="Chonga – Swahili" lang="sw" hreflang="sw" data-title="Chonga" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Ragvabaliai" title="Ragvabaliai – Lithuanian" lang="lt" hreflang="lt" data-title="Ragvabaliai" data-language-autonym="Lietuvių" data-language-local-name="Lithuanian" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/%C3%93ri%C3%A1sbog%C3%A1rform%C3%A1k" title="Óriásbogárformák – Hungarian" lang="hu" hreflang="hu" data-title="Óriásbogárformák" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%AE%D9%86%D9%81%D8%B3%D8%A7%D8%A1_%D9%88%D8%AD%D9%8A%D8%AF_%D8%A7%D9%84%D9%82%D8%B1%D9%86" title="خنفساء وحيد القرن – Egyptian Arabic" lang="arz" hreflang="arz" data-title="خنفساء وحيد القرن" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Kumbang_badak" title="Kumbang badak – Malay" lang="ms" hreflang="ms" data-title="Kumbang badak" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Dynastinae" title="Dynastinae – Dutch" lang="nl" hreflang="nl" data-title="Dynastinae" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%AB%E3%83%96%E3%83%88%E3%83%A0%E3%82%B7%E4%BA%9C%E7%A7%91" title="カブトムシ亜科 – Japanese" lang="ja" hreflang="ja" data-title="カブトムシ亜科" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Hornbiller" title="Hornbiller – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Hornbiller" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Rohaty%C5%84cowate" title="Rohatyńcowate – Polish" lang="pl" hreflang="pl" data-title="Rohatyńcowate" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Dynastinae" title="Dynastinae – Portuguese" lang="pt" hreflang="pt" data-title="Dynastinae" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-qu mw-list-item"><a href="https://qu.wikipedia.org/wiki/Waqra_suntu" title="Waqra suntu – Quechua" lang="qu" hreflang="qu" data-title="Waqra suntu" data-language-autonym="Runa Simi" data-language-local-name="Quechua" class="interlanguage-link-target"><span>Runa Simi</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%94%D1%83%D0%BF%D0%BB%D1%8F%D0%BA%D0%B8" title="Дупляки – Russian" lang="ru" hreflang="ru" data-title="Дупляки" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Dynastinae" title="Dynastinae – Finnish" lang="fi" hreflang="fi" data-title="Dynastinae" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Noshornsbaggar" title="Noshornsbaggar – Swedish" lang="sv" hreflang="sv" data-title="Noshornsbaggar" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%95%E0%AE%BE%E0%AE%A3%E0%AF%8D%E0%AE%9F%E0%AE%BE%E0%AE%AE%E0%AE%BF%E0%AE%B0%E0%AF%81%E0%AE%95_%E0%AE%B5%E0%AE%A3%E0%AF%8D%E0%AE%9F%E0%AF%81" title="காண்டாமிருக வண்டு – Tamil" lang="ta" hreflang="ta" data-title="காண்டாமிருக வண்டு" data-language-autonym="தமிழ்" data-language-local-name="Tamil" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-th mw-list-item"><a href="https://th.wikipedia.org/wiki/%E0%B8%94%E0%B9%89%E0%B8%A7%E0%B8%87%E0%B8%81%E0%B8%A7%E0%B9%88%E0%B8%B2%E0%B8%87" title="ด้วงกว่าง – Thai" lang="th" hreflang="th" data-title="ด้วงกว่าง" data-language-autonym="ไทย" data-language-local-name="Thai" class="interlanguage-link-target"><span>ไทย</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/Dynastinae" title="Dynastinae – Ukrainian" lang="uk" hreflang="uk" data-title="Dynastinae" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Ki%E1%BA%BFn_v%C6%B0%C6%A1ng" title="Kiến vương – Vietnamese" lang="vi" hreflang="vi" data-title="Kiến vương" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-war mw-list-item"><a href="https://war.wikipedia.org/wiki/Dynastidae" title="Dynastidae – Waray" lang="war" hreflang="war" data-title="Dynastidae" data-language-autonym="Winaray" data-language-local-name="Waray" class="interlanguage-link-target"><span>Winaray</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E5%85%9C%E8%9F%B2" title="兜蟲 – Cantonese" lang="yue" hreflang="yue" data-title="兜蟲" data-language-autonym="粵語" data-language-local-name="Cantonese" class="interlanguage-link-target"><span>粵語</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E5%85%9C%E8%9F%B2%E4%BA%9E%E7%A7%91" title="兜蟲亞科 – Chinese" lang="zh" hreflang="zh" data-title="兜蟲亞科" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q641142#sitelinks-wikipedia" title="Edit interlanguage links" class="wbc-editpage">Edit links</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Namespaces"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Dynastinae" title="View the content page [c]" accesskey="c"><span>Article</span></a></li><li id="ca-talk" class="vector-tab-noicon mw-list-item"><a href="/wiki/Talk:Dynastinae" rel="discussion" title="Discuss improvements to the content page [t]" accesskey="t"><span>Talk</span></a></li> </ul> </div> </div> <div id="vector-variants-dropdown" class="vector-dropdown emptyPortlet" > <input type="checkbox" id="vector-variants-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-variants-dropdown" class="vector-dropdown-checkbox " aria-label="Change language variant" > <label id="vector-variants-dropdown-label" for="vector-variants-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">English</span> </label> <div class="vector-dropdown-content"> <div id="p-variants" class="vector-menu mw-portlet mw-portlet-variants emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> </div> </div> </nav> </div> <div id="right-navigation" class="vector-collapsible"> <nav aria-label="Views"> <div id="p-views" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-views" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-view" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Dynastinae"><span>Read</span></a></li><li id="ca-edit" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Dynastinae&action=edit" title="Edit this page [e]" accesskey="e"><span>Edit</span></a></li><li id="ca-history" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Dynastinae&action=history" title="Past revisions of this page [h]" accesskey="h"><span>View history</span></a></li> </ul> </div> </div> </nav> <nav class="vector-page-tools-landmark" aria-label="Page tools"> <div id="vector-page-tools-dropdown" class="vector-dropdown vector-page-tools-dropdown" > <input type="checkbox" id="vector-page-tools-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-tools-dropdown" class="vector-dropdown-checkbox " aria-label="Tools" > <label id="vector-page-tools-dropdown-label" for="vector-page-tools-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">Tools</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-tools-unpinned-container" class="vector-unpinned-container"> <div id="vector-page-tools" class="vector-page-tools vector-pinnable-element"> <div class="vector-pinnable-header vector-page-tools-pinnable-header vector-pinnable-header-unpinned" data-feature-name="page-tools-pinned" data-pinnable-element-id="vector-page-tools" data-pinned-container-id="vector-page-tools-pinned-container" data-unpinned-container-id="vector-page-tools-unpinned-container" > <div class="vector-pinnable-header-label">Tools</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-page-tools.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-page-tools.unpin">hide</button> </div> <div id="p-cactions" class="vector-menu mw-portlet mw-portlet-cactions emptyPortlet vector-has-collapsible-items" title="More options" > <div class="vector-menu-heading"> Actions </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-more-view" class="selected vector-more-collapsible-item mw-list-item"><a href="/wiki/Dynastinae"><span>Read</span></a></li><li id="ca-more-edit" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Dynastinae&action=edit" title="Edit this page [e]" accesskey="e"><span>Edit</span></a></li><li id="ca-more-history" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Dynastinae&action=history"><span>View history</span></a></li> </ul> </div> </div> <div id="p-tb" class="vector-menu mw-portlet mw-portlet-tb" > <div class="vector-menu-heading"> General </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="t-whatlinkshere" class="mw-list-item"><a href="/wiki/Special:WhatLinksHere/Dynastinae" title="List of all English Wikipedia pages containing links to this page [j]" accesskey="j"><span>What links here</span></a></li><li id="t-recentchangeslinked" class="mw-list-item"><a href="/wiki/Special:RecentChangesLinked/Dynastinae" rel="nofollow" title="Recent changes in pages linked from this page [k]" accesskey="k"><span>Related changes</span></a></li><li id="t-upload" class="mw-list-item"><a href="/wiki/Wikipedia:File_Upload_Wizard" title="Upload files [u]" accesskey="u"><span>Upload file</span></a></li><li id="t-specialpages" class="mw-list-item"><a href="/wiki/Special:SpecialPages" title="A list of all special pages [q]" accesskey="q"><span>Special pages</span></a></li><li id="t-permalink" class="mw-list-item"><a href="/w/index.php?title=Dynastinae&oldid=1246203154" title="Permanent link to this revision of this page"><span>Permanent link</span></a></li><li id="t-info" class="mw-list-item"><a href="/w/index.php?title=Dynastinae&action=info" title="More information about this page"><span>Page information</span></a></li><li id="t-cite" class="mw-list-item"><a href="/w/index.php?title=Special:CiteThisPage&page=Dynastinae&id=1246203154&wpFormIdentifier=titleform" title="Information on how to cite this page"><span>Cite this page</span></a></li><li id="t-urlshortener" class="mw-list-item"><a href="/w/index.php?title=Special:UrlShortener&url=https%3A%2F%2Fen.wikipedia.org%2Fwiki%2FDynastinae"><span>Get shortened URL</span></a></li><li id="t-urlshortener-qrcode" class="mw-list-item"><a href="/w/index.php?title=Special:QrCode&url=https%3A%2F%2Fen.wikipedia.org%2Fwiki%2FDynastinae"><span>Download QR code</span></a></li> </ul> </div> </div> <div id="p-coll-print_export" class="vector-menu mw-portlet mw-portlet-coll-print_export" > <div class="vector-menu-heading"> Print/export </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="coll-download-as-rl" class="mw-list-item"><a href="/w/index.php?title=Special:DownloadAsPdf&page=Dynastinae&action=show-download-screen" title="Download this page as a PDF file"><span>Download as PDF</span></a></li><li id="t-print" class="mw-list-item"><a href="/w/index.php?title=Dynastinae&printable=yes" title="Printable version of this page [p]" accesskey="p"><span>Printable version</span></a></li> </ul> </div> </div> <div id="p-wikibase-otherprojects" class="vector-menu mw-portlet mw-portlet-wikibase-otherprojects" > <div class="vector-menu-heading"> In other projects </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="wb-otherproject-link wb-otherproject-commons mw-list-item"><a href="https://commons.wikimedia.org/wiki/Category:Dynastinae" hreflang="en"><span>Wikimedia Commons</span></a></li><li class="wb-otherproject-link wb-otherproject-species mw-list-item"><a href="https://species.wikimedia.org/wiki/Dynastinae" hreflang="en"><span>Wikispecies</span></a></li><li id="t-wikibase" class="wb-otherproject-link wb-otherproject-wikibase-dataitem mw-list-item"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q641142" title="Structured data on this page hosted by Wikidata [g]" accesskey="g"><span>Wikidata item</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> </div> </div> </div> <div class="vector-column-end"> <div class="vector-sticky-pinned-container"> <nav class="vector-page-tools-landmark" aria-label="Page tools"> <div id="vector-page-tools-pinned-container" class="vector-pinned-container"> </div> </nav> <nav class="vector-appearance-landmark" aria-label="Appearance"> <div id="vector-appearance-pinned-container" class="vector-pinned-container"> <div id="vector-appearance" class="vector-appearance vector-pinnable-element"> <div class="vector-pinnable-header vector-appearance-pinnable-header vector-pinnable-header-pinned" data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">Appearance</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">hide</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"><span class="mw-redirectedfrom">(Redirected from <a href="/w/index.php?title=Unicorn_beetle&redirect=no" class="mw-redirect" title="Unicorn beetle">Unicorn beetle</a>)</span></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Subfamily of beetles</div> <style data-mw-deduplicate="TemplateStyles:r1251242444">.mw-parser-output .ambox{border:1px solid #a2a9b1;border-left:10px solid #36c;background-color:#fbfbfb;box-sizing:border-box}.mw-parser-output .ambox+link+.ambox,.mw-parser-output .ambox+link+style+.ambox,.mw-parser-output .ambox+link+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+style+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+link+.ambox{margin-top:-1px}html body.mediawiki .mw-parser-output .ambox.mbox-small-left{margin:4px 1em 4px 0;overflow:hidden;width:238px;border-collapse:collapse;font-size:88%;line-height:1.25em}.mw-parser-output .ambox-speedy{border-left:10px solid #b32424;background-color:#fee7e6}.mw-parser-output .ambox-delete{border-left:10px solid #b32424}.mw-parser-output .ambox-content{border-left:10px solid #f28500}.mw-parser-output .ambox-style{border-left:10px solid #fc3}.mw-parser-output .ambox-move{border-left:10px solid #9932cc}.mw-parser-output .ambox-protection{border-left:10px solid #a2a9b1}.mw-parser-output .ambox .mbox-text{border:none;padding:0.25em 0.5em;width:100%}.mw-parser-output .ambox .mbox-image{border:none;padding:2px 0 2px 0.5em;text-align:center}.mw-parser-output .ambox .mbox-imageright{border:none;padding:2px 0.5em 2px 0;text-align:center}.mw-parser-output .ambox .mbox-empty-cell{border:none;padding:0;width:1px}.mw-parser-output .ambox .mbox-image-div{width:52px}@media(min-width:720px){.mw-parser-output .ambox{margin:0 10%}}@media print{body.ns-0 .mw-parser-output .ambox{display:none!important}}</style><table class="box-More_footnotes_needed plainlinks metadata ambox ambox-style ambox-More_footnotes_needed" role="presentation"><tbody><tr><td class="mbox-image"><div class="mbox-image-div"><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a4/Text_document_with_red_question_mark.svg/40px-Text_document_with_red_question_mark.svg.png" decoding="async" width="40" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/a4/Text_document_with_red_question_mark.svg/60px-Text_document_with_red_question_mark.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/a4/Text_document_with_red_question_mark.svg/80px-Text_document_with_red_question_mark.svg.png 2x" data-file-width="48" data-file-height="48" /></span></span></div></td><td class="mbox-text"><div class="mbox-text-span">This article includes a list of <a href="/wiki/Wikipedia:Citing_sources#General_references" title="Wikipedia:Citing sources">general references</a>, but <b>it lacks sufficient corresponding <a href="/wiki/Wikipedia:Citing_sources#Inline_citations" title="Wikipedia:Citing sources">inline citations</a></b>.<span class="hide-when-compact"> Please help to <a href="/wiki/Wikipedia:WikiProject_Reliability" title="Wikipedia:WikiProject Reliability">improve</a> this article by <a href="/wiki/Wikipedia:When_to_cite" title="Wikipedia:When to cite">introducing</a> more precise citations.</span> <span class="date-container"><i>(<span class="date">November 2015</span>)</i></span><span class="hide-when-compact"><i> (<small><a href="/wiki/Help:Maintenance_template_removal" title="Help:Maintenance template removal">Learn how and when to remove this message</a></small>)</i></span></div></td></tr></tbody></table> <p class="mw-empty-elt"> </p> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">Dynastinae<br /><div style="font-size: 85%;">Temporal range: <span class="noprint"><span style="display:inline-block;"></span><span style="display:inline-block;">Eocene–Recent</span> <span style="display:inline-block;"></span><div id="Timeline-row" style="margin: 4px auto 0; clear:both; width:220px; padding:0px; height:18px; overflow:visible; white-space:nowrap; border:1px #666; border-style:solid none; position:relative; z-index:0; font-size:97%;"> <div style="position:absolute; height:100%; left:0px; width:207.23076923077px; padding-left:5px; text-align:left; background-color:rgb(254,217,106); background-image: linear-gradient(to right, rgba(255,255,255,1), rgba(254,217,106,1) 15%, rgba(254,217,106,1));"><a href="/wiki/Precambrian" title="Precambrian">PreꞒ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,160,86); left:37.636923076923px; width:18.073846153846px;"><a href="/wiki/Cambrian" title="Cambrian">Ꞓ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(0,146,112); left:55.710769230769px; width:14.08px;"><a href="/wiki/Ordovician" title="Ordovician">O</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(179,225,182); left:69.790769230769px; width:8.3261538461539px;"><a href="/wiki/Silurian" title="Silurian">S</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(203,140,55); left:78.116923076923px; width:20.409230769231px;"><a href="/wiki/Devonian" title="Devonian">D</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(103,165,153); left:98.526153846154px; width:20.307692307692px;"><a href="/wiki/Carboniferous" title="Carboniferous">C</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(240,64,40); left:118.83384615385px; width:15.907015384615px;"><a href="/wiki/Permian" title="Permian">P</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(129,43,146); left:134.74086153846px; width:17.092984615385px;"><a href="/wiki/Triassic" title="Triassic">T</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(52,178,201); left:151.83384615385px; width:19.089230769231px;"><a href="/wiki/Jurassic" title="Jurassic">J</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,198,78); left:170.92307692308px; width:26.738461538462px;"><a href="/wiki/Cretaceous" title="Cretaceous">K</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(253,154,82); left:197.66153846154px; width:14.543692307692px;"><a href="/wiki/Paleogene" title="Paleogene">Pg</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(255,230,25); left:212.20523076923px; width:6.9215384615385px;"><a href="/wiki/Neogene" title="Neogene">N</a></div> <div id="end-border" style="position:absolute; height:100%; background-color:#666; width:1px; left:219px"></div><div style="margin:0 auto; line-height:0; clear:both; width:220px; padding:0px; height:8px; overflow:visible; background-color:transparent; position:relative; top:-4px; z-index:100;"><div style="position:absolute; height:8px; left:201.04615384615px; width:18.953846153846px; background-color:#360; opacity:0.42;"></div> <div style="position:absolute; height:8px; left:201.04615384615px; width:18.953846153846px; background-color:#360; opacity:1;"></div> <div style="position:absolute; height:6px; top:1px; left:202.04615384615px; width:16.953846153846px; background-color:#6c3;"></div> </div> </div></span></div> </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Dynastinae.jpg" class="mw-file-description"><img alt="Rhinoceros Beetle" src="//upload.wikimedia.org/wikipedia/commons/thumb/e/ea/Dynastinae.jpg/220px-Dynastinae.jpg" decoding="async" width="220" height="226" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/ea/Dynastinae.jpg/330px-Dynastinae.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/ea/Dynastinae.jpg/440px-Dynastinae.jpg 2x" data-file-width="2142" data-file-height="2196" /></a></span> </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Dynastinae" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Arthropod" title="Arthropod">Arthropoda</a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Insect" title="Insect">Insecta</a> </td></tr> <tr> <td>Order: </td> <td><a href="/wiki/Beetle" title="Beetle">Coleoptera</a> </td></tr> <tr> <td>Family: </td> <td><a href="/wiki/Scarabaeidae" title="Scarabaeidae">Scarabaeidae</a> </td></tr> <tr> <td>Subfamily: </td> <td><a class="mw-selflink selflink">Dynastinae</a><br /><small><a href="/wiki/William_Sharp_MacLeay" class="mw-redirect" title="William Sharp MacLeay">MacLeay</a>, 1819</small> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">Tribes </th></tr> <tr> <td colspan="2" style="text-align: left"> <p>See text<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><b>Dynastinae</b> or <b>rhinoceros beetles</b> are a <a href="/wiki/Subfamily" title="Subfamily">subfamily</a> of the <a href="/wiki/Scarabaeidae" title="Scarabaeidae">scarab beetle</a> <a href="/wiki/Family_(biology)" title="Family (biology)">family</a> (Scarabaeidae). Other common names – some for particular groups of rhinoceros beetles – include <b><a href="/wiki/Hercules_beetle" title="Hercules beetle">Hercules beetles</a></b>, <b>unicorn beetles</b> or <b>horn beetles</b>. Over 1,500 <a href="/wiki/Species" title="Species">species</a> and 225 <a href="/wiki/Genera" class="mw-redirect" title="Genera">genera</a> of rhinoceros beetles are known.<sup id="cite_ref-handbook_coleoptera_vol2_2-0" class="reference"><a href="#cite_note-handbook_coleoptera_vol2-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p><p>Many rhinoceros beetles are well known for their unique shapes and large sizes. Some famous species are, for example, the <a href="/wiki/Atlas_beetle" title="Atlas beetle">Atlas beetle</a> (<i>Chalcosoma atlas</i>), <a href="/wiki/Common_rhinoceros_beetle" class="mw-redirect" title="Common rhinoceros beetle">common rhinoceros beetle</a> (<i>Xylotrupes ulysses</i>), <a href="/wiki/Elephant_beetle" title="Elephant beetle">elephant beetle</a> (<i>Megasoma elephas</i>), <a href="/wiki/European_rhinoceros_beetle" title="European rhinoceros beetle">European rhinoceros beetle</a> (<i>Oryctes nasicornis</i>), <a href="/wiki/Hercules_beetle" title="Hercules beetle">Hercules beetle</a> (<i>Dynastes hercules</i>), <a href="/wiki/Japanese_rhinoceros_beetle" title="Japanese rhinoceros beetle">Japanese rhinoceros beetle</a> or <i>kabutomushi</i> (<i>Allomyrina dichotoma</i>), <a href="/wiki/Ox_beetle" class="mw-redirect" title="Ox beetle">ox beetle</a> (<i>Strategus aloeus</i>) and the <a href="/wiki/Eastern_Hercules_beetle" class="mw-redirect" title="Eastern Hercules beetle">Eastern Hercules beetle</a> (<i>Dynastes tityus</i>). </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Description">Description</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dynastinae&action=edit&section=1" title="Edit section: Description"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Oryctes_nasicornis_male_2012_G1.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Oryctes_nasicornis_male_2012_G1.jpg/220px-Oryctes_nasicornis_male_2012_G1.jpg" decoding="async" width="220" height="147" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Oryctes_nasicornis_male_2012_G1.jpg/330px-Oryctes_nasicornis_male_2012_G1.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Oryctes_nasicornis_male_2012_G1.jpg/440px-Oryctes_nasicornis_male_2012_G1.jpg 2x" data-file-width="4500" data-file-height="3000" /></a><figcaption><a href="/wiki/European_rhinoceros_beetle" title="European rhinoceros beetle">European rhinoceros beetle</a></figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Three_stages_rhinoceros_beetle.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/0/00/Three_stages_rhinoceros_beetle.jpg/220px-Three_stages_rhinoceros_beetle.jpg" decoding="async" width="220" height="185" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/0/00/Three_stages_rhinoceros_beetle.jpg 1.5x" data-file-width="289" data-file-height="243" /></a><figcaption>European rhinoceros beetle – three stages from larva to adult: <a href="/wiki/Larva" title="Larva">larva</a> (back), <a href="/wiki/Pupa" title="Pupa">pupa</a> (center), and <a href="/wiki/Imago" title="Imago">imago</a> (front)</figcaption></figure> <p>The Dynastinae are among the largest of beetles, reaching more than 15 centimetres (6 inches) in length, but are completely harmless to humans because they cannot bite or sting. Some species have been anecdotally claimed to lift up to 850 times their own weight.<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> An extinct <a href="/wiki/Eocene" title="Eocene">Eocene</a> <i><a href="/wiki/Oryctoantiquus" title="Oryctoantiquus">Oryctoantiquus borealis</a></i> was the largest fossil scarabeid, with a length of 5 centimetres (2.0 in).<sup id="cite_ref-Brett2009_4-0" class="reference"><a href="#cite_note-Brett2009-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Some modern <a href="/wiki/Oryctini" title="Oryctini">Oryctini</a> grew up to 7 cm (3 in).<sup id="cite_ref-Brett2009_4-1" class="reference"><a href="#cite_note-Brett2009-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Common names of the Dynastinae refer to the characteristic horns borne only by the males of most species in the group. Each has a horn on the head and another horn pointing forward from the center of the thorax. The horns are used in fighting other males during mating season, and for digging. The size of the horn is a good indicator of nutrition and physical health.<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p><p>The body of an adult rhinoceros beetle is covered by a thick <a href="/wiki/Exoskeleton" title="Exoskeleton">exoskeleton</a>. A pair of thick wings lie atop another set of membranous wings underneath, allowing the rhinoceros beetle to fly, although not very efficiently owing to its large size. Their best protection from <a href="/wiki/Predators" class="mw-redirect" title="Predators">predators</a> is their size and stature, also avoiding many due to being <a href="/wiki/Nocturnal" class="mw-redirect" title="Nocturnal">nocturnal</a>. When the sun is out, they hide under logs or in vegetation to <a href="/wiki/Camouflage" title="Camouflage">camouflage</a> themselves from the few predators big enough to eat them. If rhinoceros beetles are disturbed, some can release very loud, hissing squeaks created by rubbing their abdomens against the ends of their wing covers. </p> <div class="mw-heading mw-heading2"><h2 id="Ecology">Ecology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dynastinae&action=edit&section=2" title="Edit section: Ecology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>These beetles' <a href="/wiki/Larva" title="Larva">larval</a> stages can be several years long. The larvae feed on rotten wood and the adults feed on <a href="/wiki/Nectar" title="Nectar">nectar</a>, <a href="/wiki/Plant_sap" class="mw-redirect" title="Plant sap">plant sap</a>, and <a href="/wiki/Fruit" title="Fruit">fruit</a>. First, the larvae hatch from eggs and later develop into <a href="/wiki/Pupae" class="mw-redirect" title="Pupae">pupae</a> before they reach <a href="/wiki/Adult" title="Adult">adulthood</a>. </p> <div class="mw-heading mw-heading3"><h3 id="Mating">Mating</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dynastinae&action=edit&section=3" title="Edit section: Mating"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Male <a href="/wiki/Japanese_rhinoceros_beetle" title="Japanese rhinoceros beetle">Japanese rhinoceros beetles</a> (<i>Allomyrina dichotoma</i>) fight to dominate sap sites. Males use their horns to pry rival males off the area, which also may give them the chance to mate with a female. In this and other species that defend mating sites, larger males with larger horns mate more frequently, as they win more contests. Small males often avoid larger males and exhibit alternative strategies to gain access to females.<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="As_pests">As pests</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dynastinae&action=edit&section=4" title="Edit section: As pests"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Some species, such as the coconut rhinoceros beetle (<i><a href="/wiki/Oryctes_rhinoceros" title="Oryctes rhinoceros">Oryctes rhinoceros</a></i>), can become major <a href="/wiki/Pest_(organism)" title="Pest (organism)">pests</a>, e.g., in tree plantations.<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> Usually though, beetle <a href="/wiki/Population_densities" class="mw-redirect" title="Population densities">population densities</a> are not as high as in some other pest insects, and they typically prefer food trees which are already sick or dying from some other cause. Some species' larvae, however, will attack healthy trees or even <a href="/wiki/Root_vegetable" title="Root vegetable">root vegetables</a>, and when they occur in large numbers, can cause economically significant damage. The fungus <i><a href="/wiki/Metarhizium_majus" title="Metarhizium majus">Metarhizium majus</a></i> is a proven biocontrol agent for beetle infestation in crops. </p> <div class="mw-heading mw-heading2"><h2 id="Uses">Uses</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dynastinae&action=edit&section=5" title="Edit section: Uses"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Rhinoceros beetles have become popular pets in parts of <a href="/wiki/Asia" title="Asia">Asia</a>,<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> due to being relatively clean, easy to maintain, and safe to handle. Also in Asia, male beetles are used for <a href="/wiki/Insect_fighting" title="Insect fighting">gambling fights</a>.<sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> Since males naturally have the tendency to fight each other for the attention of females, they are the ones used for battle. To get the two male beetles to lock in combat, a female beetle is used, or a small noisemaker duplicating the female's <a href="/wiki/Mating_call" title="Mating call">mating call</a>. </p><p>Entomologist <a href="/w/index.php?title=S%C3%A9verin_Tchibozo&action=edit&redlink=1" class="new" title="Séverin Tchibozo (page does not exist)">Séverin Tchibozo</a> suggests the larvae contain much more <a href="/wiki/Protein" title="Protein">protein</a> (40%) than <a href="/wiki/Chicken" title="Chicken">chicken</a> (20%) and <a href="/wiki/Beef" title="Beef">beef</a> (approximately 18%), and they could become a <a href="/wiki/Entomophagy" title="Entomophagy">protein source for a large human population</a>.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> </p><p>Dr. MinJun Kim, leading a team of engineers in <a href="/wiki/National_Science_Foundation" title="National Science Foundation">National Science Foundation</a>-funded research, examined the function and aerodynamics of <i><a href="/wiki/Allomyrina_dichotoma" class="mw-redirect" title="Allomyrina dichotoma">Allomyrina dichotoma</a></i> with the help of researchers in <a href="/wiki/Drexel_University" title="Drexel University">Drexel University</a>'s Mechanical Engineering Department and in collaboration with <a href="/wiki/Konkuk_University" title="Konkuk University">Konkuk University</a> in South Korea. Rhinoceros beetles could play a big part in the next generation of aircraft design.<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Tribes_with_selected_genera_and_species">Tribes with selected genera and species</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dynastinae&action=edit&section=6" title="Edit section: Tribes with selected genera and species"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Agaocephalini">Agaocephalini</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dynastinae&action=edit&section=7" title="Edit section: Agaocephalini"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Antodon_goryi_NMNS.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/1/10/Antodon_goryi_NMNS.jpg/220px-Antodon_goryi_NMNS.jpg" decoding="async" width="220" height="337" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/10/Antodon_goryi_NMNS.jpg/330px-Antodon_goryi_NMNS.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/10/Antodon_goryi_NMNS.jpg/440px-Antodon_goryi_NMNS.jpg 2x" data-file-width="1115" data-file-height="1710" /></a><figcaption><i>Antodon goryi</i></figcaption></figure> <p>Auth: Burmeister, 1847. all genera: </p> <ol><li><i><a href="/w/index.php?title=Aegopsis&action=edit&redlink=1" class="new" title="Aegopsis (page does not exist)">Aegopsis</a></i> <small>Burmeister, 1847</small></li> <li><i><a href="/wiki/Agaocephala" title="Agaocephala">Agaocephala</a></i> <small>Lepeletier & Audinet-Serville, 1828</small></li> <li><i><a href="/wiki/Antodon" title="Antodon">Antodon</a></i> <small>Brême, 1845</small></li> <li><i><a href="/w/index.php?title=Brachysiderus&action=edit&redlink=1" class="new" title="Brachysiderus (page does not exist)">Brachysiderus</a></i> <small>Waterhouse, 1881</small></li> <li><i><a href="/w/index.php?title=Colacus&action=edit&redlink=1" class="new" title="Colacus (page does not exist)">Colacus</a></i> <small>Ohaus, 1910</small></li> <li><a href="/w/index.php?title=Democrates_(beetle)&action=edit&redlink=1" class="new" title="Democrates (beetle) (page does not exist)"><i>Democrates</i> (beetle)</a> <small>Burmeister, 1847</small></li> <li><i><a href="/w/index.php?title=Gnathogolofa&action=edit&redlink=1" class="new" title="Gnathogolofa (page does not exist)">Gnathogolofa</a></i> <small>Arrow, 1914</small></li> <li><i><a href="/w/index.php?title=Horridocalia&action=edit&redlink=1" class="new" title="Horridocalia (page does not exist)">Horridocalia</a></i> <small>Endrödi, 1974</small></li> <li><a href="/w/index.php?title=Lycomedes_(beetle)&action=edit&redlink=1" class="new" title="Lycomedes (beetle) (page does not exist)"><i>Lycomedes</i> (beetle)</a> <small>Breme, 1844</small></li> <li><i><a href="/w/index.php?title=Mitracephala&action=edit&redlink=1" class="new" title="Mitracephala (page does not exist)">Mitracephala</a></i> <small>Thomson, 1859</small></li> <li><i><a href="/wiki/Spodistes" title="Spodistes">Spodistes</a></i> <small>Burmeister, 1847</small></li></ol> <div class="mw-heading mw-heading3"><h3 id="Cyclocephalini"><a href="/wiki/Cyclocephalini" title="Cyclocephalini">Cyclocephalini</a></h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dynastinae&action=edit&section=8" title="Edit section: Cyclocephalini"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Cyclocephal_borealis_Adult.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Cyclocephal_borealis_Adult.jpg/170px-Cyclocephal_borealis_Adult.jpg" decoding="async" width="170" height="131" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Cyclocephal_borealis_Adult.jpg/255px-Cyclocephal_borealis_Adult.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Cyclocephal_borealis_Adult.jpg/340px-Cyclocephal_borealis_Adult.jpg 2x" data-file-width="768" data-file-height="594" /></a><figcaption><i><a href="/wiki/Cyclocephala_borealis" title="Cyclocephala borealis">Cyclocephala borealis</a></i> (<a href="/wiki/Cyclocephalini" title="Cyclocephalini">Cyclocephalini</a>)</figcaption></figure> <p>Auth: Laporte, 1840. Selected genera: </p> <ul><li><i><a href="/wiki/Ancognatha" title="Ancognatha">Ancognatha</a></i> <small>Erichson, 1847</small></li> <li><i><a href="/wiki/Cyclocephala" title="Cyclocephala">Cyclocephala</a></i> <small>Dejean, 1821</small> (masked chafers)</li> <li><i><a href="/wiki/Dyscinetus" title="Dyscinetus">Dyscinetus</a></i> <small>Harold, 1869</small> (rice beetles)</li></ul> <div class="mw-heading mw-heading3"><h3 id="Dynastini"><a href="/wiki/Dynastini" title="Dynastini">Dynastini</a></h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dynastinae&action=edit&section=9" title="Edit section: Dynastini"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Eupatorus_gracilicornis_Vol.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/45/Eupatorus_gracilicornis_Vol.jpg/170px-Eupatorus_gracilicornis_Vol.jpg" decoding="async" width="170" height="137" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/45/Eupatorus_gracilicornis_Vol.jpg/255px-Eupatorus_gracilicornis_Vol.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/45/Eupatorus_gracilicornis_Vol.jpg/340px-Eupatorus_gracilicornis_Vol.jpg 2x" data-file-width="6821" data-file-height="5481" /></a><figcaption><i><a href="/wiki/Eupatorus_gracilicornis" title="Eupatorus gracilicornis">Eupatorus gracilicornis</a></i> (<a href="/wiki/Dynastini" title="Dynastini">Dynastini</a>)</figcaption></figure> <p>Auth: MacLeay, 1819. Selected genera: </p> <ul><li><i><a href="/wiki/Allomyrina" title="Allomyrina">Allomyrina</a></i> <small>Arrow, 1911</small> (including <i>Trypoxylus</i>) <ul><li><i>Allomyrina dichotoma</i> – <a href="/wiki/Japanese_rhinoceros_beetle" title="Japanese rhinoceros beetle">Japanese rhinoceros beetle</a></li></ul></li> <li><i><a href="/wiki/Chalcosoma" title="Chalcosoma">Chalcosoma</a></i> <small>Hope, 1837</small> <ul><li><i>Chalcosoma atlas</i> – <a href="/wiki/Atlas_beetle" title="Atlas beetle">Atlas beetle</a></li> <li><i><a href="/wiki/Chalcosoma_moellenkampi" class="mw-redirect" title="Chalcosoma moellenkampi">Chalcosoma moellenkampi</a></i> – Moellenkampi beetle</li> <li><i><a href="/wiki/Chalcosoma_caucasus" class="mw-redirect" title="Chalcosoma caucasus">Chalcosoma caucasus</a></i> – Caucasus beetle</li></ul></li> <li><i><a href="/wiki/Dynastes" title="Dynastes">Dynastes</a></i> <small>MacLeay, 1819</small> <ul><li><i><a href="/wiki/Dynastes_hercules" class="mw-redirect" title="Dynastes hercules">Dynastes hercules</a></i> – Hercules beetle</li> <li><i><a href="/wiki/Dynastes_neptunus" title="Dynastes neptunus">Dynastes neptunus</a></i> – Neptune beetle</li></ul></li> <li><i><a href="/wiki/Eupatorus" title="Eupatorus">Eupatorus</a></i> <small>Burmeister, 1847</small> <ul><li><i><a href="/wiki/Eupatorus_gracilicornis" title="Eupatorus gracilicornis">Eupatorus gracilicornis</a></i> – Five-horned rhinoceros beetle</li> <li><i><a href="/wiki/Eupatorus_siamensis" title="Eupatorus siamensis">Eupatorus siamensis</a></i> – Siamese beetle</li> <li><i><a href="/wiki/Eupatorus_birmanicus" title="Eupatorus birmanicus">Eupatorus birmanicus</a></i> – Rabbit beetle</li></ul></li> <li><i><a href="/wiki/Megasoma" title="Megasoma">Megasoma</a></i> <small>Kirby, 1825</small> <ul><li><i><a href="/wiki/Megasoma_mars" title="Megasoma mars">Megasoma mars</a></i> - Mars beetle</li> <li><i><a href="/wiki/Megasoma_elephas" class="mw-redirect" title="Megasoma elephas">Megasoma elephas</a></i> - Elephant beetle</li> <li><i><a href="/wiki/Megasoma_actaeon" class="mw-redirect" title="Megasoma actaeon">Megasoma actaeon</a></i> - Actaeon beetle</li></ul></li> <li><i><a href="/wiki/Xylotrupes" title="Xylotrupes">Xylotrupes</a></i> <small>Hope, 1837</small> <ul><li><i><a href="/wiki/Xylotrupes_gideon" title="Xylotrupes gideon">Xylotrupes gideon</a></i> – Siamese rhinoceros beetle</li> <li><i><a href="/wiki/Xylotrupes_ulysses" title="Xylotrupes ulysses">Xylotrupes ulysses</a></i></li></ul></li></ul> <div class="mw-heading mw-heading3"><h3 id="Hexodontini"><a href="/wiki/Hexodontini" title="Hexodontini">Hexodontini</a></h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dynastinae&action=edit&section=10" title="Edit section: Hexodontini"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Auth. Lacordaire, 1856; all genera from Madagascar: </p> <ol><li><i><a href="/w/index.php?title=Hemicyrthus&action=edit&redlink=1" class="new" title="Hemicyrthus (page does not exist)">Hemicyrthus</a></i> <small>Reiche, 1860</small></li> <li><i><a href="/w/index.php?title=Hexodon&action=edit&redlink=1" class="new" title="Hexodon (page does not exist)">Hexodon</a></i> <small>Olivier, 1789</small></li> <li><i><a href="/w/index.php?title=Hyboschema&action=edit&redlink=1" class="new" title="Hyboschema (page does not exist)">Hyboschema</a></i> <small>Péringuey, 1901</small></li></ol> <div class="mw-heading mw-heading3"><h3 id="Oryctini"><a href="/wiki/Oryctini" title="Oryctini">Oryctini</a></h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dynastinae&action=edit&section=11" title="Edit section: Oryctini"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Trichogomphus_mongol_(Arrow,1908)_60_mm_Rhinoceros_Beetle_male_(7941206664).jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/f5/Trichogomphus_mongol_%28Arrow%2C1908%29_60_mm_Rhinoceros_Beetle_male_%287941206664%29.jpg/170px-Trichogomphus_mongol_%28Arrow%2C1908%29_60_mm_Rhinoceros_Beetle_male_%287941206664%29.jpg" decoding="async" width="170" height="128" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/f5/Trichogomphus_mongol_%28Arrow%2C1908%29_60_mm_Rhinoceros_Beetle_male_%287941206664%29.jpg/255px-Trichogomphus_mongol_%28Arrow%2C1908%29_60_mm_Rhinoceros_Beetle_male_%287941206664%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/f5/Trichogomphus_mongol_%28Arrow%2C1908%29_60_mm_Rhinoceros_Beetle_male_%287941206664%29.jpg/340px-Trichogomphus_mongol_%28Arrow%2C1908%29_60_mm_Rhinoceros_Beetle_male_%287941206664%29.jpg 2x" data-file-width="3264" data-file-height="2448" /></a><figcaption><i><a href="/w/index.php?title=Trichogomphus_mongol&action=edit&redlink=1" class="new" title="Trichogomphus mongol (page does not exist)">Trichogomphus mongol</a></i> (<a href="/wiki/Oryctini" title="Oryctini">Oryctini</a>)</figcaption></figure> <p>Auth: Mulsant, 1842. Selected genera: </p> <ul><li><i><a href="/w/index.php?title=Enema_(beetle)&action=edit&redlink=1" class="new" title="Enema (beetle) (page does not exist)">Enema</a></i> <small>Hope, 1837</small></li> <li><i><a href="/w/index.php?title=Megaceras&action=edit&redlink=1" class="new" title="Megaceras (page does not exist)">Megaceras</a></i> <small>Hope, 1837</small> <ul><li><i><a href="/wiki/Megaceras_briansaltini" title="Megaceras briansaltini">Megaceras briansaltini</a></i></li></ul></li> <li><i><a href="/wiki/Oryctes" title="Oryctes">Oryctes</a></i> <small>Hellwig, 1798</small> <ul><li><i>Oryctes nasicornis</i> – <a href="/wiki/European_rhinoceros_beetle" title="European rhinoceros beetle">European rhinoceros beetle</a></li> <li><i><a href="/wiki/Oryctes_rhinoceros" title="Oryctes rhinoceros">Oryctes rhinoceros</a></i> – coconut rhinoceros beetle</li></ul></li> <li><i><a href="/wiki/Strategus_(beetle)" title="Strategus (beetle)">Strategus</a></i> <small>Kirby, 1828</small> <ul><li><i><a href="/wiki/Strategus_aloeus" title="Strategus aloeus">Strategus aloeus</a></i> – ox beetle</li></ul></li> <li><i><a href="/wiki/Trichogomphus" title="Trichogomphus">Trichogomphus</a></i> <small>Burmeister, 1847</small></li> <li><i><a href="/wiki/Xyloryctes" title="Xyloryctes">Xyloryctes</a></i></li></ul> <div class="mw-heading mw-heading3"><h3 id="Oryctoderini"><a href="/w/index.php?title=Oryctoderini&action=edit&redlink=1" class="new" title="Oryctoderini (page does not exist)">Oryctoderini</a></h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dynastinae&action=edit&section=12" title="Edit section: Oryctoderini"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Auth. Endrödi, 1966; all genera: </p> <ol><li><i><a href="/w/index.php?title=Chalcasthenes&action=edit&redlink=1" class="new" title="Chalcasthenes (page does not exist)">Chalcasthenes</a></i> <small>Arrow, 1937</small></li> <li><i><a href="/w/index.php?title=Chalcocrates&action=edit&redlink=1" class="new" title="Chalcocrates (page does not exist)">Chalcocrates</a></i> <small>Heller, 1903</small></li> <li><i><a href="/w/index.php?title=Coenoryctoderus&action=edit&redlink=1" class="new" title="Coenoryctoderus (page does not exist)">Coenoryctoderus</a></i> <small>Prell, 1933</small></li> <li><i><a href="/w/index.php?title=Hatamus&action=edit&redlink=1" class="new" title="Hatamus (page does not exist)">Hatamus</a></i> <small>Sharp, 1877</small></li> <li><i><a href="/w/index.php?title=Melanhyphus&action=edit&redlink=1" class="new" title="Melanhyphus (page does not exist)">Melanhyphus</a></i> <small>Fairmaire, 1881</small></li> <li><i><a href="/w/index.php?title=Neohyphus&action=edit&redlink=1" class="new" title="Neohyphus (page does not exist)">Neohyphus</a></i> <small>Heller, 1896</small></li> <li><i><a href="/w/index.php?title=Onychionyx&action=edit&redlink=1" class="new" title="Onychionyx (page does not exist)">Onychionyx</a></i> <small>Arrow, 1914</small></li> <li><i><a href="/w/index.php?title=Oryctoderinus&action=edit&redlink=1" class="new" title="Oryctoderinus (page does not exist)">Oryctoderinus</a></i> <small>Endrödi, 1978</small></li> <li><i><a href="/w/index.php?title=Oryctoderus&action=edit&redlink=1" class="new" title="Oryctoderus (page does not exist)">Oryctoderus</a></i> <small>Boisduval, 1835</small></li> <li><i><a href="/w/index.php?title=Paroryctoderus&action=edit&redlink=1" class="new" title="Paroryctoderus (page does not exist)">Paroryctoderus</a></i> <small>Dechambre, 1994</small></li></ol> <div class="mw-heading mw-heading3"><h3 id="Pentodontini"><a href="/wiki/Pentodontini" title="Pentodontini">Pentodontini</a></h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dynastinae&action=edit&section=13" title="Edit section: Pentodontini"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Pentodon_idiota_-_%D0%9D%D0%B0%D0%B2%D0%BE%D0%B7%D0%BD%D0%B8%D0%BA_(%D0%94%D1%83%D0%BF%D0%BB%D1%8F%D0%BA)_%D0%BA%D1%83%D0%BA%D1%83%D1%80%D1%83%D0%B7%D0%BD%D1%8B%D0%B9.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/27/Pentodon_idiota_-_%D0%9D%D0%B0%D0%B2%D0%BE%D0%B7%D0%BD%D0%B8%D0%BA_%28%D0%94%D1%83%D0%BF%D0%BB%D1%8F%D0%BA%29_%D0%BA%D1%83%D0%BA%D1%83%D1%80%D1%83%D0%B7%D0%BD%D1%8B%D0%B9.jpg/170px-Pentodon_idiota_-_%D0%9D%D0%B0%D0%B2%D0%BE%D0%B7%D0%BD%D0%B8%D0%BA_%28%D0%94%D1%83%D0%BF%D0%BB%D1%8F%D0%BA%29_%D0%BA%D1%83%D0%BA%D1%83%D1%80%D1%83%D0%B7%D0%BD%D1%8B%D0%B9.jpg" decoding="async" width="170" height="128" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/27/Pentodon_idiota_-_%D0%9D%D0%B0%D0%B2%D0%BE%D0%B7%D0%BD%D0%B8%D0%BA_%28%D0%94%D1%83%D0%BF%D0%BB%D1%8F%D0%BA%29_%D0%BA%D1%83%D0%BA%D1%83%D1%80%D1%83%D0%B7%D0%BD%D1%8B%D0%B9.jpg/255px-Pentodon_idiota_-_%D0%9D%D0%B0%D0%B2%D0%BE%D0%B7%D0%BD%D0%B8%D0%BA_%28%D0%94%D1%83%D0%BF%D0%BB%D1%8F%D0%BA%29_%D0%BA%D1%83%D0%BA%D1%83%D1%80%D1%83%D0%B7%D0%BD%D1%8B%D0%B9.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/27/Pentodon_idiota_-_%D0%9D%D0%B0%D0%B2%D0%BE%D0%B7%D0%BD%D0%B8%D0%BA_%28%D0%94%D1%83%D0%BF%D0%BB%D1%8F%D0%BA%29_%D0%BA%D1%83%D0%BA%D1%83%D1%80%D1%83%D0%B7%D0%BD%D1%8B%D0%B9.jpg/340px-Pentodon_idiota_-_%D0%9D%D0%B0%D0%B2%D0%BE%D0%B7%D0%BD%D0%B8%D0%BA_%28%D0%94%D1%83%D0%BF%D0%BB%D1%8F%D0%BA%29_%D0%BA%D1%83%D0%BA%D1%83%D1%80%D1%83%D0%B7%D0%BD%D1%8B%D0%B9.jpg 2x" data-file-width="1024" data-file-height="768" /></a><figcaption><i><a href="/w/index.php?title=Pentodon_idiota&action=edit&redlink=1" class="new" title="Pentodon idiota (page does not exist)">Pentodon idiota</a></i> (<a href="/wiki/Pentodontini" title="Pentodontini">Pentodontini</a>)</figcaption></figure> <p>Auth: Mulsant, 1842. Selected genera: </p> <ul><li><i><a href="/w/index.php?title=Diloboderus&action=edit&redlink=1" class="new" title="Diloboderus (page does not exist)">Diloboderus</a></i> <small>Sturm, 1826</small> monotypic <i><a href="https://es.wikipedia.org/wiki/Diloboderus_abderus" class="extiw" title="es:Diloboderus abderus">Diloboderus abderus</a></i></li> <li><i><a href="/wiki/Ligyrus" title="Ligyrus">Ligyrus</a></i> <span style="font-size:85%;">Burmeister, 1847</span></li> <li><i><a href="/wiki/Pentodon_(beetle)" title="Pentodon (beetle)">Pentodon</a></i> <small>Hope, 1837</small></li> <li><i><a href="/wiki/Pericoptus" title="Pericoptus">Pericoptus</a></i> <small>Burmeister, 1847</small></li> <li><i><a href="/wiki/Tomarus" title="Tomarus">Tomarus</a></i> <small>Erichson, 1847</small></li></ul> <div class="mw-heading mw-heading3"><h3 id="Phileurini"><a href="/wiki/Phileurini" title="Phileurini">Phileurini</a></h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dynastinae&action=edit&section=14" title="Edit section: Phileurini"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Phileurus_valgus_P1620883a.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/49/Phileurus_valgus_P1620883a.jpg/170px-Phileurus_valgus_P1620883a.jpg" decoding="async" width="170" height="130" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/49/Phileurus_valgus_P1620883a.jpg/255px-Phileurus_valgus_P1620883a.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/49/Phileurus_valgus_P1620883a.jpg/340px-Phileurus_valgus_P1620883a.jpg 2x" data-file-width="1900" data-file-height="1453" /></a><figcaption><i><a href="/wiki/Phileurus_valgus" title="Phileurus valgus">Phileurus valgus</a></i> (<a href="/wiki/Phileurini" title="Phileurini">Phileurini</a>)</figcaption></figure> <p>Auth: Burmeister, 1847; selected genera: </p> <ul><li><i><a href="/w/index.php?title=Cryptodus&action=edit&redlink=1" class="new" title="Cryptodus (page does not exist)">Cryptodus</a></i> <small>MacLeay, 1819</small></li> <li><i><a href="/wiki/Phileurus" title="Phileurus">Phileurus</a></i> <small>Latreille, 1807</small></li></ul> <div class="mw-heading mw-heading2"><h2 id="Notes">Notes</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dynastinae&action=edit&section=15" title="Edit section: Notes"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width reflist-columns-3"> <ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text">Bouchard, P., Y. Bousquet, A. Davies, M. Alonso-Zarazaga, J. Lawrence, C. Lyal, A. Newton, et al. (2011). "Family-group names in Coleoptera (Insecta)". <i>ZooKeys, vol. 88</i>, 1-972.</span> </li> <li id="cite_note-handbook_coleoptera_vol2-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-handbook_coleoptera_vol2_2-0">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFBeutelLeschen2016" class="citation book cs1">Beutel, Rolf G.; Leschen, Richard A.B., eds. (2016-03-21). <i>Coleoptera, Beetles, Volume 1, Morphology and Systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim)</i>. De Gruyter. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1515%2F9783110373929">10.1515/9783110373929</a>. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-3-11-037392-9" title="Special:BookSources/978-3-11-037392-9"><bdi>978-3-11-037392-9</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Coleoptera%2C+Beetles%2C+Volume+1%2C+Morphology+and+Systematics+%28Archostemata%2C+Adephaga%2C+Myxophaga%2C+Polyphaga+partim%29&rft.pub=De+Gruyter&rft.date=2016-03-21&rft_id=info%3Adoi%2F10.1515%2F9783110373929&rft.isbn=978-3-11-037392-9&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADynastinae" class="Z3988"></span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-3">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFKram1996" class="citation journal cs1">Kram, Rodger (1 March 1996). <a rel="nofollow" class="external text" href="https://journals.biologists.com/jeb/article/199/3/609/7375/Inexpensive-Load-Carrying-by-Rhinoceros-Beetles">"Inexpensive load carrying by rhinoceros beetles"</a>. <i>Journal of Experimental Biology</i>. <b>199</b> (3): 609–612. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1242%2Fjeb.199.3.609">10.1242/jeb.199.3.609</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/9318326">9318326</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Journal+of+Experimental+Biology&rft.atitle=Inexpensive+load+carrying+by+rhinoceros+beetles&rft.volume=199&rft.issue=3&rft.pages=609-612&rft.date=1996-03-01&rft_id=info%3Adoi%2F10.1242%2Fjeb.199.3.609&rft_id=info%3Apmid%2F9318326&rft.aulast=Kram&rft.aufirst=Rodger&rft_id=https%3A%2F%2Fjournals.biologists.com%2Fjeb%2Farticle%2F199%2F3%2F609%2F7375%2FInexpensive-Load-Carrying-by-Rhinoceros-Beetles&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADynastinae" class="Z3988"></span></span> </li> <li id="cite_note-Brett2009-4"><span class="mw-cite-backlink">^ <a href="#cite_ref-Brett2009_4-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Brett2009_4-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text">Brett Ratcliffe, Dena M. Smith, Diane Marie Erwin. "<a rel="nofollow" class="external text" href="https://www.researchgate.net/publication/232682009_Oryctoantiquus_borealis_New_Genus_and_Species_from_the_Eocene_of_Oregon_USA_the_World's_Oldest_Fossil_Dynastine_and_Largest_Fossil_Scarabaeid_Coleoptera_Scarabaeidae_Dynastinae">Oryctoantiquus borealis, New Genus and Species from the Eocene of Oregon, U.S.A., the World's Oldest Fossil Dynastine and Largest Fossil Scarabaeid (Coleoptera: Scarabaeidae: Dynastinae)</a>". 2009. The Coleopterists Bulletin 59(Mar 2005):127-135 DOI:10.1649/0010-065X(2005)059[0127:OBNGAS]2.0.CO;2</span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-5">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.newscientist.com/blogs/nstv/2012/07/why-horn-size-matters-when-picking-a-mate.html?DCMP=OTC-rss&nsref=online-news">"Why horn size matters when picking a mate"</a>. <i>New Scientist</i>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=New+Scientist&rft.atitle=Why+horn+size+matters+when+picking+a+mate&rft_id=https%3A%2F%2Fwww.newscientist.com%2Fblogs%2Fnstv%2F2012%2F07%2Fwhy-horn-size-matters-when-picking-a-mate.html%3FDCMP%3DOTC-rss%26nsref%3Donline-news&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADynastinae" class="Z3988"></span></span> </li> <li id="cite_note-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-6">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBuchalskiGutierrezEmlenLavine2019" class="citation journal cs1">Buchalski, Benjamin; Gutierrez, Eric; Emlen, Douglas; Lavine, Laura; Swanson, Brook (15 October 2019). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6835817">"Variation in an Extreme Weapon: Horn Performance Differences across Rhinoceros Beetle (Trypoxylus dichotomus) Populations"</a>. <i>Insects</i>. <b>10</b> (10): 346. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.3390%2Finsects10100346">10.3390/insects10100346</a></span>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6835817">6835817</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/31618906">31618906</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Insects&rft.atitle=Variation+in+an+Extreme+Weapon%3A+Horn+Performance+Differences+across+Rhinoceros+Beetle+%28Trypoxylus+dichotomus%29+Populations&rft.volume=10&rft.issue=10&rft.pages=346&rft.date=2019-10-15&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC6835817%23id-name%3DPMC&rft_id=info%3Apmid%2F31618906&rft_id=info%3Adoi%2F10.3390%2Finsects10100346&rft.aulast=Buchalski&rft.aufirst=Benjamin&rft.au=Gutierrez%2C+Eric&rft.au=Emlen%2C+Douglas&rft.au=Lavine%2C+Laura&rft.au=Swanson%2C+Brook&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC6835817&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADynastinae" class="Z3988"></span></span> </li> <li id="cite_note-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-7">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFGoczałRossaTofilski2019" class="citation journal cs1">Goczał, Jakub; Rossa, Robert; Tofilski, Adam (17 April 2019). "Intersexual and intrasexual patterns of horn size and shape variation in the European rhinoceros beetle: quantifying the shape of weapons". <i>Biological Journal of the Linnean Society</i>. <b>127</b> (1): 34–43. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fbiolinnean%2Fblz026">10.1093/biolinnean/blz026</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Biological+Journal+of+the+Linnean+Society&rft.atitle=Intersexual+and+intrasexual+patterns+of+horn+size+and+shape+variation+in+the+European+rhinoceros+beetle%3A+quantifying+the+shape+of+weapons&rft.volume=127&rft.issue=1&rft.pages=34-43&rft.date=2019-04-17&rft_id=info%3Adoi%2F10.1093%2Fbiolinnean%2Fblz026&rft.aulast=Gocza%C5%82&rft.aufirst=Jakub&rft.au=Rossa%2C+Robert&rft.au=Tofilski%2C+Adam&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADynastinae" class="Z3988"></span></span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-8">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://entnemdept.ufl.edu/creatures/orn/palms/Oryctes_rhinoceros.htm">Featured Creatures: Coconut rhinoceros beetle</a></span> </li> <li id="cite_note-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-9">^</a></b></span> <span class="reference-text">"WHO? KNEW" (May 6, 2005) <i>Current Science</i> Vol.90 No.16</span> </li> <li id="cite_note-10"><span class="mw-cite-backlink"><b><a href="#cite_ref-10">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20120412084046/http://www.articlesextra.com/beetle-fight-xylotrupes-thailand.htm">"Rhinoceros beetle gambling in Thailand"</a>. Archived from <a rel="nofollow" class="external text" href="http://www.articlesextra.com/beetle-fight-xylotrupes-thailand.htm">the original</a> on 2012-04-12<span class="reference-accessdate">. Retrieved <span class="nowrap">2006-11-03</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Rhinoceros+beetle+gambling+in+Thailand&rft_id=http%3A%2F%2Fwww.articlesextra.com%2Fbeetle-fight-xylotrupes-thailand.htm&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADynastinae" class="Z3988"></span></span> </li> <li id="cite_note-11"><span class="mw-cite-backlink"><b><a href="#cite_ref-11">^</a></b></span> <span class="reference-text"><i><a href="/wiki/Global_Steak" title="Global Steak">Global Steak</a> - Demain nos enfants mangeront des criquets</i> (2010 French documentary)</span> </li> <li id="cite_note-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-12">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation news cs1"><a rel="nofollow" class="external text" href="https://www.sciencedaily.com/releases/2012/04/120410163531.htm">"Engineers Unlock Secrets of Beetle Flight"</a> <span class="cs1-format">(news story)</span>. <i>ScienceDaily</i>. April 11, 2012. <q><i>ScienceDaily (Apr. 10, 2012)</i> — Rhinoceros beetles could play a big part....</q></cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=ScienceDaily&rft.atitle=Engineers+Unlock+Secrets+of+Beetle+Flight&rft.date=2012-04-11&rft_id=https%3A%2F%2Fwww.sciencedaily.com%2Freleases%2F2012%2F04%2F120410163531.htm&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADynastinae" class="Z3988"></span></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="Further_reading">Further reading</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dynastinae&action=edit&section=16" title="Edit section: Further reading"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFEndrödi1985" class="citation book cs1"><a href="/wiki/Seb%C3%B6_Endr%C3%B6di" title="Sebö Endrödi">Endrödi, S.</a> (1985). <i>The Dynastinae of the World</i>. Series Entomologica. Vol. 28. Dr. W. Junk Publishers. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-9061931386" title="Special:BookSources/978-9061931386"><bdi>978-9061931386</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=The+Dynastinae+of+the+World&rft.series=Series+Entomologica&rft.pub=Dr.+W.+Junk+Publishers&rft.date=1985&rft.isbn=978-9061931386&rft.aulast=Endr%C3%B6di&rft.aufirst=S.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ADynastinae" class="Z3988"></span></li> <li>Dechambre (R.-P.) & Lachaume (G.) <i><a href="/wiki/The_Beetles_of_the_World" title="The Beetles of the World">The Beetles of the World</a></i>, volume 27, The genus <i>Oryctes</i> (Dynastidae), <a href="/wiki/Hillside_Books,_Canterbury" title="Hillside Books, Canterbury">Hillside Books, Canterbury</a> <a rel="nofollow" class="external autonumber" href="https://web.archive.org/web/20100806171758/http://www.insects.demon.co.uk/cm/cm-27-english.html">[1]</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Dynastinae&action=edit&section=17" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 0;box-sizing:border-box;border:1px solid #aaa;font-size:88%;line-height:1.25em;background-color:var(--background-color-interactive-subtle,#f8f9fa);display:flow-root}.mw-parser-output .side-box-abovebelow,.mw-parser-output .side-box-text{padding:0.25em 0.9em}.mw-parser-output .side-box-image{padding:2px 0 2px 0.9em;text-align:center}.mw-parser-output .side-box-imageright{padding:2px 0.9em 2px 0;text-align:center}@media(min-width:500px){.mw-parser-output .side-box-flex{display:flex;align-items:center}.mw-parser-output .side-box-text{flex:1;min-width:0}}@media(min-width:720px){.mw-parser-output .side-box{width:238px}.mw-parser-output .side-box-right{clear:right;float:right;margin-left:1em}.mw-parser-output .side-box-left{margin-right:1em}}</style><style data-mw-deduplicate="TemplateStyles:r1237033735">@media print{body.ns-0 .mw-parser-output .sistersitebox{display:none!important}}@media screen{html.skin-theme-clientpref-night .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}</style><div class="side-box side-box-right plainlinks sistersitebox"><style data-mw-deduplicate="TemplateStyles:r1126788409">.mw-parser-output .plainlist ol,.mw-parser-output .plainlist ul{line-height:inherit;list-style:none;margin:0;padding:0}.mw-parser-output .plainlist ol li,.mw-parser-output .plainlist ul li{margin-bottom:0}</style> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/30px-Commons-logo.svg.png" decoding="async" width="30" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/45px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/59px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></span></span></div> <div class="side-box-text plainlist">Wikimedia Commons has media related to <span style="font-weight: bold; font-style: italic;"><a href="https://commons.wikimedia.org/wiki/Category:Dynastinae" class="extiw" title="commons:Category:Dynastinae">Dynastinae</a></span>.</div></div> </div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1235681985"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1237033735"><div class="side-box side-box-right plainlinks sistersitebox"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/34px-Wikispecies-logo.svg.png" decoding="async" width="34" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/51px-Wikispecies-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/68px-Wikispecies-logo.svg.png 2x" data-file-width="941" data-file-height="1103" /></span></span></div> <div class="side-box-text plainlist"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a> has information related to <i><b><a href="https://species.wikimedia.org/wiki/Dynastinae" class="extiw" title="wikispecies:Dynastinae">Dynastinae</a></b></i>.</div></div> </div> <ul><li><a rel="nofollow" class="external text" href="http://www.naturalworlds.org/scarabaeidae/index.htm">Family SCARABAEIDAE</a></li> <li><a rel="nofollow" class="external text" href="https://www.biolib.cz/en/taxonsubtaxa/id8133/">Subfamily Dynastinae</a></li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol ol,.mw-parser-output .hlist ol ul,.mw-parser-output .hlist ul dl,.mw-parser-output .hlist ul ol,.mw-parser-output .hlist ul ul{display:inline}.mw-parser-output .hlist .mw-empty-li{display:none}.mw-parser-output .hlist dt::after{content:": "}.mw-parser-output .hlist dd::after,.mw-parser-output .hlist li::after{content:" · ";font-weight:bold}.mw-parser-output .hlist dd:last-child::after,.mw-parser-output .hlist dt:last-child::after,.mw-parser-output .hlist li:last-child::after{content:none}.mw-parser-output .hlist dd dd:first-child::before,.mw-parser-output .hlist dd dt:first-child::before,.mw-parser-output .hlist dd li:first-child::before,.mw-parser-output .hlist dt dd:first-child::before,.mw-parser-output .hlist dt dt:first-child::before,.mw-parser-output .hlist dt li:first-child::before,.mw-parser-output .hlist li dd:first-child::before,.mw-parser-output .hlist li dt:first-child::before,.mw-parser-output .hlist li li:first-child::before{content:" (";font-weight:normal}.mw-parser-output .hlist dd dd:last-child::after,.mw-parser-output .hlist dd dt:last-child::after,.mw-parser-output .hlist dd li:last-child::after,.mw-parser-output .hlist dt dd:last-child::after,.mw-parser-output .hlist dt dt:last-child::after,.mw-parser-output .hlist dt li:last-child::after,.mw-parser-output .hlist li dd:last-child::after,.mw-parser-output .hlist li dt:last-child::after,.mw-parser-output .hlist li li:last-child::after{content:")";font-weight:normal}.mw-parser-output .hlist ol{counter-reset:listitem}.mw-parser-output .hlist ol>li{counter-increment:listitem}.mw-parser-output .hlist ol>li::before{content:" "counter(listitem)"\a0 "}.mw-parser-output .hlist dd ol>li:first-child::before,.mw-parser-output .hlist dt ol>li:first-child::before,.mw-parser-output .hlist li ol>li:first-child::before{content:" ("counter(listitem)"\a0 "}</style><style data-mw-deduplicate="TemplateStyles:r1236075235">.mw-parser-output .navbox{box-sizing:border-box;border:1px solid #a2a9b1;width:100%;clear:both;font-size:88%;text-align:center;padding:1px;margin:1em auto 0}.mw-parser-output .navbox .navbox{margin-top:0}.mw-parser-output .navbox+.navbox,.mw-parser-output .navbox+.navbox-styles+.navbox{margin-top:-1px}.mw-parser-output .navbox-inner,.mw-parser-output .navbox-subgroup{width:100%}.mw-parser-output .navbox-group,.mw-parser-output .navbox-title,.mw-parser-output .navbox-abovebelow{padding:0.25em 1em;line-height:1.5em;text-align:center}.mw-parser-output .navbox-group{white-space:nowrap;text-align:right}.mw-parser-output .navbox,.mw-parser-output .navbox-subgroup{background-color:#fdfdfd}.mw-parser-output .navbox-list{line-height:1.5em;border-color:#fdfdfd}.mw-parser-output .navbox-list-with-group{text-align:left;border-left-width:2px;border-left-style:solid}.mw-parser-output tr+tr>.navbox-abovebelow,.mw-parser-output tr+tr>.navbox-group,.mw-parser-output tr+tr>.navbox-image,.mw-parser-output tr+tr>.navbox-list{border-top:2px solid #fdfdfd}.mw-parser-output .navbox-title{background-color:#ccf}.mw-parser-output .navbox-abovebelow,.mw-parser-output .navbox-group,.mw-parser-output .navbox-subgroup .navbox-title{background-color:#ddf}.mw-parser-output .navbox-subgroup .navbox-group,.mw-parser-output .navbox-subgroup .navbox-abovebelow{background-color:#e6e6ff}.mw-parser-output .navbox-even{background-color:#f7f7f7}.mw-parser-output .navbox-odd{background-color:transparent}.mw-parser-output .navbox .hlist td dl,.mw-parser-output .navbox .hlist td ol,.mw-parser-output .navbox .hlist td ul,.mw-parser-output .navbox td.hlist dl,.mw-parser-output .navbox td.hlist ol,.mw-parser-output .navbox td.hlist ul{padding:0.125em 0}.mw-parser-output .navbox .navbar{display:block;font-size:100%}.mw-parser-output .navbox-title .navbar{float:left;text-align:left;margin-right:0.5em}body.skin--responsive .mw-parser-output .navbox-image img{max-width:none!important}@media print{body.ns-0 .mw-parser-output .navbox{display:none!important}}</style></div><div role="navigation" class="navbox" aria-labelledby="Human_interactions_with_insects" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Insects_in_culture" title="Template:Insects in culture"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Insects_in_culture" title="Template talk:Insects in culture"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Insects_in_culture" title="Special:EditPage/Template:Insects in culture"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Human_interactions_with_insects" style="font-size:114%;margin:0 4em"><a href="/wiki/Human_interactions_with_insects" title="Human interactions with insects">Human interactions with insects</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">Aspects<br />of <a href="/wiki/Insect" title="Insect">insects</a><br />in culture</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">In the arts</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Insects_in_art" title="Insects in art">Insects in art</a> <ul><li><a href="/wiki/Beetlewing" title="Beetlewing">Beetlewing</a></li> <li><i><a href="/wiki/Musca_depicta" title="Musca depicta">Musca depicta</a></i></li></ul></li> <li><a href="/wiki/Arthropods_in_film" title="Arthropods in film">Arthropods in film</a></li> <li><a href="/wiki/Insects_in_literature" title="Insects in literature">Insects in literature</a></li> <li><a href="/wiki/Insects_in_music" title="Insects in music">Insects in music</a> <ul><li><a href="/wiki/List_of_insect-inspired_songs" title="List of insect-inspired songs">List of insect-inspired songs</a></li></ul></li> <li><a href="/wiki/Insects_on_stamps" title="Insects on stamps">Insects on stamps</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">In fishing</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Fishing_bait" title="Fishing bait">Fishing bait</a></li> <li><a href="/wiki/Fly_fishing" title="Fly fishing">Fly fishing</a></li> <li><a href="/wiki/Artificial_fly" title="Artificial fly">Artificial fly</a></li> <li><a href="/wiki/Fly_tying" title="Fly tying">Fly tying</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Insects_in_medicine" title="Insects in medicine">In medicine</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Apitherapy" title="Apitherapy">Apitherapy</a> <ul><li><a href="/wiki/Apitoxin" title="Apitoxin">Apitoxin</a></li> <li><a href="/wiki/Melittin" title="Melittin">Melittin</a></li></ul></li> <li><a href="/wiki/Lytta_vesicatoria" title="Lytta vesicatoria">Spanish fly</a> <ul><li><a href="/wiki/Cantharidin" title="Cantharidin">Cantharidin</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Insects_in_mythology" title="Insects in mythology">In mythology</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bees_in_mythology" title="Bees in mythology">Bees in mythology</a></li> <li><a href="/wiki/Cicadas_in_mythology" title="Cicadas in mythology">Cicadas in mythology</a></li> <li><a href="/wiki/Scarab_(artifact)" title="Scarab (artifact)">Scarab (artifact)</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other aspects</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biomimetics" title="Biomimetics">Biomimicry</a></li> <li><a href="/wiki/Cockroach_racing" title="Cockroach racing">Cockroach racing</a></li> <li><a href="/wiki/Cockroaches_in_popular_culture" title="Cockroaches in popular culture">Cockroaches in popular culture</a></li> <li><a href="/wiki/Cricket_fighting" title="Cricket fighting">Cricket fighting</a></li> <li><a href="/wiki/Entomological_warfare" title="Entomological warfare">Entomological warfare</a></li> <li><a href="/wiki/Entomophagy_in_humans" title="Entomophagy in humans">Entomophagy</a></li> <li><a href="/wiki/Insect_farming" title="Insect farming">Insect farming</a></li> <li><a href="/wiki/Flea_circus" title="Flea circus">Flea circus</a></li> <li><a href="/wiki/Insects_in_ethics" title="Insects in ethics">Insects in ethics</a></li> <li><a href="/wiki/Insects_in_religion" title="Insects in religion">Insects in religion</a></li> <li><a href="/wiki/Jingzhe" title="Jingzhe">Jingzhe</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Economic_entomology" title="Economic entomology">Economic<br />entomology</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Beneficial_insect" title="Beneficial insect">Beneficial insect</a></li> <li><a href="/wiki/Biological_pest_control" title="Biological pest control">Biological pest control</a></li> <li><a href="/wiki/Beekeeping" title="Beekeeping">Beekeeping</a> <ul><li><a href="/wiki/Bee_pollen" title="Bee pollen">Bee pollen</a></li> <li><a href="/wiki/List_of_crop_plants_pollinated_by_bees" title="List of crop plants pollinated by bees">List of crop plants pollinated by bees</a></li> <li><a href="/wiki/Beeswax" title="Beeswax">Beeswax</a></li> <li><a href="/wiki/Honey" title="Honey">Honey</a></li> <li><a href="/wiki/Propolis" title="Propolis">Propolis</a></li> <li><a href="/wiki/Royal_jelly" title="Royal jelly">Royal jelly</a></li></ul></li> <li><a href="/wiki/Carmine" title="Carmine">Carmine/Cochineal</a> <ul><li><a href="/wiki/Polish_cochineal" title="Polish cochineal">Polish</a></li></ul></li> <li><a href="/wiki/Chitin" title="Chitin">Chitin</a></li> <li><a href="/wiki/Kermes_(dye)" title="Kermes (dye)">Kermes</a></li> <li><a href="/wiki/Sericulture" title="Sericulture">Sericulture</a> <ul><li><a href="/wiki/Silk" title="Silk">Silk</a></li></ul></li> <li><a href="/wiki/Shellac" title="Shellac">Shellac</a></li> <li><a href="/wiki/Model_organism" title="Model organism">Model organism</a> <ul><li><i><a href="/wiki/Drosophila_melanogaster" title="Drosophila melanogaster">Drosophila melanogaster</a></i></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Pest_(organism)#Insects" title="Pest (organism)">Harmful<br />insects</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Insect_bites_and_stings" class="mw-redirect" title="Insect bites and stings">Insect bites and stings</a></li> <li><a href="/wiki/Insect_sting_allergy" title="Insect sting allergy">Insect sting allergy</a></li> <li><a href="/wiki/Bed_bug" title="Bed bug">Bed bug</a></li> <li><a href="/wiki/Woodworm" title="Woodworm">Woodworm</a></li> <li><a href="/wiki/Home-stored_product_entomology" title="Home-stored product entomology">Home-stored product entomology</a></li> <li><a href="/wiki/Clothes_moth" title="Clothes moth">Clothes moth</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Pioneers</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Jan_Swammerdam" title="Jan Swammerdam">Jan Swammerdam</a></li> <li><a href="/wiki/Alfred_Russel_Wallace" title="Alfred Russel Wallace">Alfred Russel Wallace</a></li> <li><a href="/wiki/Jean-Henri_Fabre" title="Jean-Henri Fabre">Jean-Henri Fabre</a></li> <li><a href="/wiki/Hans_Zinsser" title="Hans Zinsser">Hans Zinsser</a> (<i><a href="/wiki/Rats,_Lice_and_History" title="Rats, Lice and History">Rats, Lice and History</a></i>)</li> <li><a href="/wiki/Lafcadio_Hearn" title="Lafcadio Hearn">Lafcadio Hearn</a> (<i>Insect Literature</i>)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Concerns</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bees_and_toxic_chemicals" title="Bees and toxic chemicals">Bees and toxic chemicals</a></li> <li><a href="/wiki/Colony_collapse_disorder" title="Colony collapse disorder">Colony collapse disorder</a></li> <li><a href="/wiki/Decline_in_insect_populations" title="Decline in insect populations">Decline in insect populations</a></li> <li><a href="/wiki/Habitat_destruction" title="Habitat destruction">Habitat destruction</a></li> <li><a href="/wiki/List_of_endangered_insects" title="List of endangered insects">List of endangered insects</a></li> <li><a href="/wiki/Pesticide" title="Pesticide">Pesticide</a> <ul><li><a href="/wiki/Insecticide" title="Insecticide">Insecticide</a></li> <li><a href="/wiki/Neonicotinoid" title="Neonicotinoid">Neonicotinoid</a></li> <li><a href="/wiki/Pesticide_toxicity_to_bees" title="Pesticide toxicity to bees">Pesticide toxicity to bees</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Categories,<br />templates</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Category:Insects_and_humans" title="Category:Insects and humans">Insects and humans</a></li> <li><a href="/wiki/Template:Insecticides" title="Template:Insecticides">Insecticides</a></li> <li><a href="/wiki/Template:Pesticides" title="Template:Pesticides">Pesticides</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2" style="font-weight:bold;"><div><span class="nowrap"><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Beetle_icon.svg" class="mw-file-description"><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/6c/Beetle_icon.svg/25px-Beetle_icon.svg.png" decoding="async" width="25" height="28" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/6c/Beetle_icon.svg/38px-Beetle_icon.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/6c/Beetle_icon.svg/50px-Beetle_icon.svg.png 2x" data-file-width="143" data-file-height="159" /></a></span> </span><a href="/wiki/Portal:Insects" title="Portal:Insects">Insects portal</a></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Dynastinae</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q641142" class="extiw" title="wikidata:Q641142">Q641142</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Dynastinae" class="extiw" title="wikispecies:Dynastinae">Dynastinae</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Australian_Faunal_Directory" title="Australian Faunal Directory">AFD</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://biodiversity.org.au/afd/taxa/Dynastinae">Dynastinae</a></span></span></li> <li><span style="white-space:nowrap;">BioLib: <span class="uid"><a rel="nofollow" class="external text" href="https://www.biolib.cz/en/taxon/id8133">8133</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/BugGuide" title="BugGuide">BugGuide</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://bugguide.net/node/view/12431">12431</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Catalogue_of_Life" title="Catalogue of Life">CoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.catalogueoflife.org/data/taxon/7NLQQ">7NLQQ</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Fauna_Europaea" title="Fauna Europaea">Fauna Europaea</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://www.eu-nomen.eu/portal/taxon.php?GUID=urn:lsid:faunaeur.org:taxname:246911">246911</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Fauna_Europaea" title="Fauna Europaea">Fauna Europaea (new)</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://fauna-eu.org/cdm_dataportal/taxon/9aa5cb31-5738-4393-ab61-abf9d2b0af1f">9aa5cb31-5738-4393-ab61-abf9d2b0af1f</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/INaturalist" title="INaturalist">iNaturalist</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://inaturalist.org/taxa/136361">136361</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Integrated_Taxonomic_Information_System" title="Integrated Taxonomic Information System">ITIS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=678510">678510</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=50519">50519</a></span></span></li> <li><span style="white-space:nowrap;">NZOR: <span class="uid"><a rel="nofollow" class="external text" href="https://www.nzor.org.nz/names/b353af6b-5c28-4965-b432-3b1cb2b0ddf7">b353af6b-5c28-4965-b432-3b1cb2b0ddf7</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=70053">70053</a></span></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><style data-mw-deduplicate="TemplateStyles:r1038841319">.mw-parser-output .tooltip-dotted{border-bottom:1px dotted;cursor:help}</style></div><div role="navigation" class="navbox authority-control" aria-label="Navbox" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Help:Authority_control" title="Help:Authority control">Authority control databases</a>: National <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q641142#identifiers" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><span class="rt-commentedText tooltip tooltip-dotted" title="nosorožíci"><a rel="nofollow" class="external text" href="https://aleph.nkp.cz/F/?func=find-c&local_base=aut&ccl_term=ica=ph471364&CON_LNG=ENG">Czech Republic</a></span></span></li></ul></div></td></tr></tbody></table></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1&useformat=desktop" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Retrieved from "<a dir="ltr" href="https://en.wikipedia.org/w/index.php?title=Dynastinae&oldid=1246203154">https://en.wikipedia.org/w/index.php?title=Dynastinae&oldid=1246203154</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Help:Category" title="Help:Category">Categories</a>: <ul><li><a href="/wiki/Category:Dynastinae" title="Category:Dynastinae">Dynastinae</a></li><li><a href="/wiki/Category:Insects_as_food" title="Category:Insects as food">Insects as food</a></li><li><a href="/wiki/Category:Insects_in_culture" title="Category:Insects in culture">Insects in culture</a></li><li><a href="/wiki/Category:Extant_Eocene_first_appearances" title="Category:Extant Eocene first appearances">Extant Eocene first appearances</a></li></ul></div><div id="mw-hidden-catlinks" class="mw-hidden-catlinks mw-hidden-cats-hidden">Hidden categories: <ul><li><a href="/wiki/Category:Articles_with_short_description" title="Category:Articles with short description">Articles with short description</a></li><li><a href="/wiki/Category:Short_description_matches_Wikidata" title="Category:Short description matches Wikidata">Short description matches Wikidata</a></li><li><a href="/wiki/Category:Articles_lacking_in-text_citations_from_November_2015" title="Category:Articles lacking in-text citations from November 2015">Articles lacking in-text citations from November 2015</a></li><li><a href="/wiki/Category:All_articles_lacking_in-text_citations" title="Category:All articles lacking in-text citations">All articles lacking in-text citations</a></li><li><a href="/wiki/Category:Articles_with_%27species%27_microformats" title="Category:Articles with 'species' microformats">Articles with 'species' microformats</a></li><li><a href="/wiki/Category:Commons_category_link_is_on_Wikidata" title="Category:Commons category link is on Wikidata">Commons category link is on Wikidata</a></li></ul></div></div> </div> </main> </div> <div class="mw-footer-container"> <footer id="footer" class="mw-footer" > <ul id="footer-info"> <li id="footer-info-lastmod"> This page was last edited on 17 September 2024, at 15:00<span class="anonymous-show"> (UTC)</span>.</li> <li id="footer-info-copyright">Text is available under the <a href="/wiki/Wikipedia:Text_of_the_Creative_Commons_Attribution-ShareAlike_4.0_International_License" title="Wikipedia:Text of the Creative Commons Attribution-ShareAlike 4.0 International License">Creative Commons Attribution-ShareAlike 4.0 License</a>; additional terms may apply. By using this site, you agree to the <a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Terms_of_Use" class="extiw" title="foundation:Special:MyLanguage/Policy:Terms of Use">Terms of Use</a> and <a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Privacy_policy" class="extiw" title="foundation:Special:MyLanguage/Policy:Privacy policy">Privacy Policy</a>. Wikipedia® is a registered trademark of the <a rel="nofollow" class="external text" href="https://wikimediafoundation.org/">Wikimedia Foundation, Inc.</a>, a non-profit organization.</li> </ul> <ul id="footer-places"> <li id="footer-places-privacy"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Privacy_policy">Privacy policy</a></li> <li id="footer-places-about"><a href="/wiki/Wikipedia:About">About Wikipedia</a></li> <li id="footer-places-disclaimers"><a href="/wiki/Wikipedia:General_disclaimer">Disclaimers</a></li> <li id="footer-places-contact"><a href="//en.wikipedia.org/wiki/Wikipedia:Contact_us">Contact Wikipedia</a></li> <li id="footer-places-wm-codeofconduct"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Universal_Code_of_Conduct">Code of Conduct</a></li> <li id="footer-places-developers"><a href="https://developer.wikimedia.org">Developers</a></li> <li id="footer-places-statslink"><a href="https://stats.wikimedia.org/#/en.wikipedia.org">Statistics</a></li> <li id="footer-places-cookiestatement"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Cookie_statement">Cookie statement</a></li> <li id="footer-places-mobileview"><a href="//en.m.wikipedia.org/w/index.php?title=Dynastinae&mobileaction=toggle_view_mobile" class="noprint stopMobileRedirectToggle">Mobile view</a></li> </ul> <ul id="footer-icons" class="noprint"> <li id="footer-copyrightico"><a href="https://wikimediafoundation.org/" class="cdx-button cdx-button--fake-button cdx-button--size-large cdx-button--fake-button--enabled"><img src="/static/images/footer/wikimedia-button.svg" width="84" height="29" alt="Wikimedia Foundation" loading="lazy"></a></li> <li id="footer-poweredbyico"><a href="https://www.mediawiki.org/" class="cdx-button cdx-button--fake-button cdx-button--size-large cdx-button--fake-button--enabled"><img src="/w/resources/assets/poweredby_mediawiki.svg" alt="Powered by MediaWiki" width="88" height="31" loading="lazy"></a></li> </ul> </footer> </div> </div> </div> <div class="vector-settings" id="p-dock-bottom"> <ul></ul> </div><script>(RLQ=window.RLQ||[]).push(function(){mw.config.set({"wgHostname":"mw-web.codfw.main-5c59558b9d-j9zbk","wgBackendResponseTime":173,"wgPageParseReport":{"limitreport":{"cputime":"0.849","walltime":"1.107","ppvisitednodes":{"value":18524,"limit":1000000},"postexpandincludesize":{"value":108431,"limit":2097152},"templateargumentsize":{"value":22960,"limit":2097152},"expansiondepth":{"value":24,"limit":100},"expensivefunctioncount":{"value":14,"limit":500},"unstrip-depth":{"value":1,"limit":20},"unstrip-size":{"value":54846,"limit":5000000},"entityaccesscount":{"value":12,"limit":400},"timingprofile":["100.00% 989.367 1 -total"," 38.07% 376.646 1 Template:Automatic_taxobox"," 13.62% 134.770 1 Template:Reflist"," 11.52% 113.942 1 Template:Fossil_range"," 10.78% 106.616 1 Template:Taxonbar"," 8.97% 88.759 2 Template:Navbox"," 8.96% 88.674 1 Template:Insects_in_culture"," 8.94% 88.415 2 Template:Cite_book"," 8.74% 86.493 1 Template:Phanerozoic_220px"," 8.02% 79.331 38 Template:Period_start"]},"scribunto":{"limitreport-timeusage":{"value":"0.578","limit":"10.000"},"limitreport-memusage":{"value":8659907,"limit":52428800},"limitreport-logs":"table#1 {\n [\"size\"] = \"small\",\n}\n"},"cachereport":{"origin":"mw-web.codfw.main-f69cdc8f6-f2wr6","timestamp":"20241124113123","ttl":2592000,"transientcontent":false}}});});</script> <script type="application/ld+json">{"@context":"https:\/\/schema.org","@type":"Article","name":"Dynastinae","url":"https:\/\/en.wikipedia.org\/wiki\/Dynastinae","sameAs":"http:\/\/www.wikidata.org\/entity\/Q641142","mainEntity":"http:\/\/www.wikidata.org\/entity\/Q641142","author":{"@type":"Organization","name":"Contributors to Wikimedia projects"},"publisher":{"@type":"Organization","name":"Wikimedia Foundation, Inc.","logo":{"@type":"ImageObject","url":"https:\/\/www.wikimedia.org\/static\/images\/wmf-hor-googpub.png"}},"datePublished":"2005-06-30T14:51:11Z","dateModified":"2024-09-17T15:00:13Z","image":"https:\/\/upload.wikimedia.org\/wikipedia\/commons\/e\/ea\/Dynastinae.jpg","headline":"subfamily of beetles"}</script> </body> </html>