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[o]" accesskey="o"><span class="vector-icon mw-ui-icon-logIn mw-ui-icon-wikimedia-logIn"></span> <span>Đăng nhập</span></a></li> </ul> </div> </div> <div id="p-user-menu-anon-editor" class="vector-menu mw-portlet mw-portlet-user-menu-anon-editor" > <div class="vector-menu-heading"> Trang dành cho người dùng chưa đăng nhập <a href="/wiki/Tr%E1%BB%A3_gi%C3%BAp:Gi%E1%BB%9Bi_thi%E1%BB%87u" aria-label="Tìm hiểu thêm về sửa đổi"><span>tìm hiểu thêm</span></a> </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-anoncontribs" class="mw-list-item"><a href="/wiki/%C4%90%E1%BA%B7c_bi%E1%BB%87t:%C4%90%C3%B3ng_g%C3%B3p_c%E1%BB%A7a_t%C3%B4i" title="Danh sách các sửa đổi được thực hiện qua địa chỉ IP này [y]" accesskey="y"><span>Đóng góp</span></a></li><li id="pt-anontalk" class="mw-list-item"><a href="/wiki/%C4%90%E1%BA%B7c_bi%E1%BB%87t:Th%E1%BA%A3o_lu%E1%BA%ADn_t%C3%B4i" title="Thảo luận với địa chỉ IP này [n]" accesskey="n"><span>Thảo luận cho địa chỉ IP này</span></a></li> </ul> </div> </div> </div> </div> </nav> </div> </header> </div> <div class="mw-page-container"> <div class="mw-page-container-inner"> <div class="vector-sitenotice-container"> <div id="siteNotice"></div> </div> <div class="vector-column-start"> <div class="vector-main-menu-container"> <div id="mw-navigation"> <nav id="mw-panel" class="vector-main-menu-landmark" aria-label="Trang Web"> <div id="vector-main-menu-pinned-container" class="vector-pinned-container"> </div> </nav> </div> </div> <div class="vector-sticky-pinned-container"> <nav id="mw-panel-toc" aria-label="Nội dung" data-event-name="ui.sidebar-toc" class="mw-table-of-contents-container vector-toc-landmark"> <div id="vector-toc-pinned-container" class="vector-pinned-container"> <div id="vector-toc" class="vector-toc vector-pinnable-element"> <div class="vector-pinnable-header vector-toc-pinnable-header vector-pinnable-header-pinned" data-feature-name="toc-pinned" data-pinnable-element-id="vector-toc" > <h2 class="vector-pinnable-header-label">Nội dung</h2> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-toc.pin">chuyển sang thanh bên</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-toc.unpin">ẩn</button> </div> <ul class="vector-toc-contents" id="mw-panel-toc-list"> <li id="toc-mw-content-text" class="vector-toc-list-item vector-toc-level-1"> <a href="#" class="vector-toc-link"> <div class="vector-toc-text">Đầu</div> </a> </li> <li id="toc-Chú_thích" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Chú_thích"> <div class="vector-toc-text"> <span class="vector-toc-numb">1</span> <span>Chú thích</span> </div> </a> <ul id="toc-Chú_thích-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Tham_khảo" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Tham_khảo"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Tham khảo</span> </div> </a> <ul id="toc-Tham_khảo-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Nội dung" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Đóng mở mục lục" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Đóng mở mục lục</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><i>Mycobacterium tuberculosis</i></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Xem bài viết trong ngôn ngữ khác. Bài có sẵn trong 48 ngôn ngữ" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-48" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">48 ngôn ngữ</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-am mw-list-item"><a href="https://am.wikipedia.org/wiki/%E1%88%9B%E1%8B%AD%E1%8A%AE%E1%89%A3%E1%8A%AD%E1%89%B4%E1%88%AA%E1%8B%A8%E1%88%9D_%E1%89%B1%E1%89%A0%E1%88%AD%E1%8A%AD%E1%88%8E%E1%88%B2%E1%88%B5" title="ማይኮባክቴሪየም ቱበርክሎሲስ – Tiếng Amharic" lang="am" hreflang="am" data-title="ማይኮባክቴሪየም ቱበርክሎሲስ" data-language-autonym="አማርኛ" data-language-local-name="Tiếng Amharic" class="interlanguage-link-target"><span>አማርኛ</span></a></li><li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D9%85%D8%AA%D9%81%D8%B7%D8%B1%D8%A9_%D8%B3%D9%84%D9%8A%D8%A9" title="متفطرة سلية – Tiếng Ả Rập" lang="ar" hreflang="ar" data-title="متفطرة سلية" data-language-autonym="العربية" data-language-local-name="Tiếng Ả Rập" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Indonesia" lang="id" hreflang="id" data-title="Mycobacterium tuberculosis" data-language-autonym="Bahasa Indonesia" data-language-local-name="Tiếng Indonesia" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Mã Lai" lang="ms" hreflang="ms" data-title="Mycobacterium tuberculosis" data-language-autonym="Bahasa Melayu" data-language-local-name="Tiếng Mã Lai" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%AE%E0%A6%BE%E0%A6%87%E0%A6%95%E0%A7%8B%E0%A6%AC%E0%A7%8D%E0%A6%AF%E0%A6%BE%E0%A6%95%E0%A7%8D%E0%A6%9F%E0%A7%87%E0%A6%B0%E0%A6%BF%E0%A6%AF%E0%A6%BC%E0%A6%BE%E0%A6%AE_%E0%A6%9F%E0%A6%BF%E0%A6%89%E0%A6%AC%E0%A6%BE%E0%A6%B0%E0%A6%95%E0%A6%BF%E0%A6%89%E0%A6%B2%E0%A7%8B%E0%A6%B8%E0%A6%BF%E0%A6%B8" title="মাইকোব্যাক্টেরিয়াম টিউবারকিউলোসিস – Tiếng Bangla" lang="bn" hreflang="bn" data-title="মাইকোব্যাক্টেরিয়াম টিউবারকিউলোসিস" data-language-autonym="বাংলা" data-language-local-name="Tiếng Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Belarus" lang="be" hreflang="be" data-title="Mycobacterium tuberculosis" data-language-autonym="Беларуская" data-language-local-name="Tiếng Belarus" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Bacil_de_Koch" title="Bacil de Koch – Tiếng Catalan" lang="ca" hreflang="ca" data-title="Bacil de Koch" data-language-autonym="Català" data-language-local-name="Tiếng Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Séc" lang="cs" hreflang="cs" data-title="Mycobacterium tuberculosis" data-language-autonym="Čeština" data-language-local-name="Tiếng Séc" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Đức" lang="de" hreflang="de" data-title="Mycobacterium tuberculosis" data-language-autonym="Deutsch" data-language-local-name="Tiếng Đức" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Tuberkuloosi_m%C3%BCkobakter" title="Tuberkuloosi mükobakter – Tiếng Estonia" lang="et" hreflang="et" data-title="Tuberkuloosi mükobakter" data-language-autonym="Eesti" data-language-local-name="Tiếng Estonia" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Anh" lang="en" hreflang="en" data-title="Mycobacterium tuberculosis" data-language-autonym="English" data-language-local-name="Tiếng Anh" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Tây Ban Nha" lang="es" hreflang="es" data-title="Mycobacterium tuberculosis" data-language-autonym="Español" data-language-local-name="Tiếng Tây Ban Nha" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Basque" lang="eu" hreflang="eu" data-title="Mycobacterium tuberculosis" data-language-autonym="Euskara" data-language-local-name="Tiếng Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D9%85%D8%A7%DB%8C%DA%A9%D9%88%D8%A8%D8%A7%DA%A9%D8%AA%D8%B1%DB%8C%D9%88%D9%85_%D8%AA%D9%88%D8%A8%D8%B1%DA%A9%D9%84%D9%88%D8%B2%DB%8C%D8%B3" title="مایکوباکتریوم توبرکلوزیس – Tiếng Ba Tư" lang="fa" hreflang="fa" data-title="مایکوباکتریوم توبرکلوزیس" data-language-autonym="فارسی" data-language-local-name="Tiếng Ba Tư" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Pháp" lang="fr" hreflang="fr" data-title="Mycobacterium tuberculosis" data-language-autonym="Français" data-language-local-name="Tiếng Pháp" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Galician" lang="gl" hreflang="gl" data-title="Mycobacterium tuberculosis" data-language-autonym="Galego" data-language-local-name="Tiếng Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EA%B2%B0%ED%95%B5%EA%B7%A0" title="결핵균 – Tiếng Hàn" lang="ko" hreflang="ko" data-title="결핵균" data-language-autonym="한국어" data-language-local-name="Tiếng Hàn" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%AE%E0%A4%BE%E0%A4%87%E0%A4%95%E0%A5%8B%E0%A4%AC%E0%A5%88%E0%A4%95%E0%A5%8D%E0%A4%9F%E0%A5%80%E0%A4%B0%E0%A4%BF%E0%A4%AF%E0%A4%AE_%E0%A4%9F%E0%A5%8D%E0%A4%AF%E0%A5%82%E0%A4%AC%E0%A4%B0%E0%A5%8D%E0%A4%95%E0%A5%8D%E0%A4%AF%E0%A5%81%E0%A4%B2%E0%A5%8B%E0%A4%B8%E0%A4%BF%E0%A4%B8" title="माइकोबैक्टीरियम ट्यूबर्क्युलोसिस – Tiếng Hindi" lang="hi" hreflang="hi" data-title="माइकोबैक्टीरियम ट्यूबर्क्युलोसिस" data-language-autonym="हिन्दी" data-language-local-name="Tiếng Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-is mw-list-item"><a href="https://is.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Iceland" lang="is" hreflang="is" data-title="Mycobacterium tuberculosis" data-language-autonym="Íslenska" data-language-local-name="Tiếng Iceland" class="interlanguage-link-target"><span>Íslenska</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Italy" lang="it" hreflang="it" data-title="Mycobacterium tuberculosis" data-language-autonym="Italiano" data-language-local-name="Tiếng Italy" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Do Thái" lang="he" hreflang="he" data-title="Mycobacterium tuberculosis" data-language-autonym="עברית" data-language-local-name="Tiếng Do Thái" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng La-tinh" lang="la" hreflang="la" data-title="Mycobacterium tuberculosis" data-language-autonym="Latina" data-language-local-name="Tiếng La-tinh" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-lv mw-list-item"><a href="https://lv.wikipedia.org/wiki/Tuberkulozes_mikobakt%C4%93rija" title="Tuberkulozes mikobaktērija – Tiếng Latvia" lang="lv" hreflang="lv" data-title="Tuberkulozes mikobaktērija" data-language-autonym="Latviešu" data-language-local-name="Tiếng Latvia" class="interlanguage-link-target"><span>Latviešu</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Hungary" lang="hu" hreflang="hu" data-title="Mycobacterium tuberculosis" data-language-autonym="Magyar" data-language-local-name="Tiếng Hungary" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-mk mw-list-item"><a href="https://mk.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Macedonia" lang="mk" hreflang="mk" data-title="Mycobacterium tuberculosis" data-language-autonym="Македонски" data-language-local-name="Tiếng Macedonia" class="interlanguage-link-target"><span>Македонски</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%AE%E0%B5%88%E0%B4%95%E0%B5%8D%E0%B4%95%E0%B5%8B%E0%B4%AC%E0%B4%BE%E0%B4%95%E0%B5%8D%E0%B4%B1%E0%B5%8D%E0%B4%B1%E0%B5%80%E0%B4%B0%E0%B4%BF%E0%B4%AF%E0%B4%82_%E0%B4%9F%E0%B5%8D%E0%B4%AF%E0%B5%82%E0%B4%AC%E0%B5%BC%E0%B4%95%E0%B5%81%E0%B4%B2%E0%B5%8B%E0%B4%B8%E0%B4%BF%E0%B4%B8%E0%B5%8D" title="മൈക്കോബാക്റ്റീരിയം ട്യൂബർകുലോസിസ് – Tiếng Malayalam" lang="ml" hreflang="ml" data-title="മൈക്കോബാക്റ്റീരിയം ട്യൂബർകുലോസിസ്" data-language-autonym="മലയാളം" data-language-local-name="Tiếng Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D9%85%D8%AA%D9%81%D8%B7%D8%B1%D9%87_%D8%B3%D9%84%D9%8A%D9%87" title="متفطره سليه – Tiếng Ả Rập Ai Cập" lang="arz" hreflang="arz" data-title="متفطره سليه" data-language-autonym="مصرى" data-language-local-name="Tiếng Ả Rập Ai Cập" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Hà Lan" lang="nl" hreflang="nl" data-title="Mycobacterium tuberculosis" data-language-autonym="Nederlands" data-language-local-name="Tiếng Hà Lan" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E7%B5%90%E6%A0%B8%E8%8F%8C" title="結核菌 – Tiếng Nhật" lang="ja" hreflang="ja" data-title="結核菌" data-language-autonym="日本語" data-language-local-name="Tiếng Nhật" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Na Uy (Bokmål)" lang="nb" hreflang="nb" data-title="Mycobacterium tuberculosis" data-language-autonym="Norsk bokmål" data-language-local-name="Tiếng Na Uy (Bokmål)" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-nn mw-list-item"><a href="https://nn.wikipedia.org/wiki/Tuberkelbakterie" title="Tuberkelbakterie – Tiếng Na Uy (Nynorsk)" lang="nn" hreflang="nn" data-title="Tuberkelbakterie" data-language-autonym="Norsk nynorsk" data-language-local-name="Tiếng Na Uy (Nynorsk)" class="interlanguage-link-target"><span>Norsk nynorsk</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Uzbek" lang="uz" hreflang="uz" data-title="Mycobacterium tuberculosis" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Tiếng Uzbek" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D9%85%D8%A7%D8%A6%DA%A9%D8%B1%D9%88%D8%A8%DB%8C%DA%A9%D9%B9%DB%8C%D8%B1%DB%8C%D9%85_%D9%B9%DB%8C%D9%88%D8%A8%D8%B1%DA%A9%D9%84%D9%88%D8%B2%D8%B2" title="مائکروبیکٹیریم ٹیوبرکلوزز – Western Punjabi" lang="pnb" hreflang="pnb" data-title="مائکروبیکٹیریم ٹیوبرکلوزز" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-km mw-list-item"><a href="https://km.wikipedia.org/wiki/%E1%9E%87%E1%9F%86%E1%9E%84%E1%9E%BA%E1%9E%9A%E1%9E%94%E1%9F%81%E1%9E%84" title="ជំងឺរបេង – Tiếng Khmer" lang="km" hreflang="km" data-title="ជំងឺរបេង" data-language-autonym="ភាសាខ្មែរ" data-language-local-name="Tiếng Khmer" class="interlanguage-link-target"><span>ភាសាខ្មែរ</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Pr%C4%85tek_gru%C5%BAlicy" title="Prątek gruźlicy – Tiếng Ba Lan" lang="pl" hreflang="pl" data-title="Prątek gruźlicy" data-language-autonym="Polski" data-language-local-name="Tiếng Ba Lan" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Bồ Đào Nha" lang="pt" hreflang="pt" data-title="Mycobacterium tuberculosis" data-language-autonym="Português" data-language-local-name="Tiếng Bồ Đào Nha" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Romania" lang="ro" hreflang="ro" data-title="Mycobacterium tuberculosis" data-language-autonym="Română" data-language-local-name="Tiếng Romania" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Nga" lang="ru" hreflang="ru" data-title="Mycobacterium tuberculosis" data-language-autonym="Русский" data-language-local-name="Tiếng Nga" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sq mw-list-item"><a href="https://sq.wikipedia.org/wiki/Bacili_i_tuberkulozit" title="Bacili i tuberkulozit – Tiếng Albania" lang="sq" hreflang="sq" data-title="Bacili i tuberkulozit" data-language-autonym="Shqip" data-language-local-name="Tiếng Albania" class="interlanguage-link-target"><span>Shqip</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Simple English" lang="en-simple" hreflang="en-simple" data-title="Mycobacterium tuberculosis" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Slovak" lang="sk" hreflang="sk" data-title="Mycobacterium tuberculosis" data-language-autonym="Slovenčina" data-language-local-name="Tiếng Slovak" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Slovenia" lang="sl" hreflang="sl" data-title="Mycobacterium tuberculosis" data-language-autonym="Slovenščina" data-language-local-name="Tiếng Slovenia" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Phần Lan" lang="fi" hreflang="fi" data-title="Mycobacterium tuberculosis" data-language-autonym="Suomi" data-language-local-name="Tiếng Phần Lan" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv badge-Q17559452 badge-recommendedarticle mw-list-item" title="bài chất lượng cao"><a href="https://sv.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Thụy Điển" lang="sv" hreflang="sv" data-title="Mycobacterium tuberculosis" data-language-autonym="Svenska" data-language-local-name="Tiếng Thụy Điển" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-th mw-list-item"><a href="https://th.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Thái" lang="th" hreflang="th" data-title="Mycobacterium tuberculosis" data-language-autonym="ไทย" data-language-local-name="Tiếng Thái" class="interlanguage-link-target"><span>ไทย</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Thổ Nhĩ Kỳ" lang="tr" hreflang="tr" data-title="Mycobacterium tuberculosis" data-language-autonym="Türkçe" data-language-local-name="Tiếng Thổ Nhĩ Kỳ" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/Mycobacterium_tuberculosis" title="Mycobacterium tuberculosis – Tiếng Ukraina" lang="uk" hreflang="uk" data-title="Mycobacterium tuberculosis" data-language-autonym="Українська" data-language-local-name="Tiếng Ukraina" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E7%B5%90%E6%A0%B8%E5%88%86%E6%9E%9D%E6%A1%BF%E8%8F%8C" title="結核分枝桿菌 – Tiếng Trung" lang="zh" hreflang="zh" data-title="結核分枝桿菌" data-language-autonym="中文" data-language-local-name="Tiếng Trung" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q130971#sitelinks-wikipedia" title="Sửa liên kết giữa ngôn ngữ" class="wbc-editpage">Sửa liên kết</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Không gian tên"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Mycobacterium_tuberculosis" title="Xem bài viết [c]" accesskey="c"><span>Bài viết</span></a></li><li id="ca-talk" class="new vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Th%E1%BA%A3o_lu%E1%BA%ADn:Mycobacterium_tuberculosis&action=edit&redlink=1" rel="discussion" class="new" title="Thảo luận về trang này (trang không tồn tại) [t]" accesskey="t"><span>Thảo luận</span></a></li> </ul> </div> </div> <div id="vector-variants-dropdown" class="vector-dropdown emptyPortlet" > <input type="checkbox" id="vector-variants-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-variants-dropdown" class="vector-dropdown-checkbox " aria-label="Thay đổi biến thể ngôn ngữ" > <label id="vector-variants-dropdown-label" for="vector-variants-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">Tiếng Việt</span> </label> <div class="vector-dropdown-content"> <div id="p-variants" class="vector-menu mw-portlet mw-portlet-variants emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> </div> </div> </nav> </div> <div id="right-navigation" class="vector-collapsible"> <nav aria-label="Giao diện"> <div id="p-views" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-views" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-view" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Mycobacterium_tuberculosis"><span>Đọc</span></a></li><li id="ca-ve-edit" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Mycobacterium_tuberculosis&veaction=edit" title="Sửa đổi trang này [v]" accesskey="v"><span>Sửa đổi</span></a></li><li id="ca-edit" class="collapsible vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Mycobacterium_tuberculosis&action=edit" title="Sửa đổi mã nguồn của trang này [e]" accesskey="e"><span>Sửa mã nguồn</span></a></li><li id="ca-history" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Mycobacterium_tuberculosis&action=history" title="Các phiên bản cũ của trang này [h]" accesskey="h"><span>Xem lịch sử</span></a></li> </ul> </div> </div> </nav> <nav class="vector-page-tools-landmark" aria-label="Công cụ trang"> <div id="vector-page-tools-dropdown" class="vector-dropdown vector-page-tools-dropdown" > <input type="checkbox" id="vector-page-tools-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-tools-dropdown" class="vector-dropdown-checkbox " aria-label="Công cụ" > <label id="vector-page-tools-dropdown-label" for="vector-page-tools-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">Công cụ</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-tools-unpinned-container" class="vector-unpinned-container"> <div id="vector-page-tools" class="vector-page-tools vector-pinnable-element"> <div class="vector-pinnable-header vector-page-tools-pinnable-header vector-pinnable-header-unpinned" data-feature-name="page-tools-pinned" data-pinnable-element-id="vector-page-tools" data-pinned-container-id="vector-page-tools-pinned-container" data-unpinned-container-id="vector-page-tools-unpinned-container" > <div class="vector-pinnable-header-label">Công cụ</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-page-tools.pin">chuyển sang thanh bên</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-page-tools.unpin">ẩn</button> </div> <div id="p-cactions" class="vector-menu mw-portlet mw-portlet-cactions emptyPortlet vector-has-collapsible-items" title="Thêm tùy chọn" > <div class="vector-menu-heading"> Tác vụ </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-more-view" class="selected vector-more-collapsible-item mw-list-item"><a href="/wiki/Mycobacterium_tuberculosis"><span>Đọc</span></a></li><li id="ca-more-ve-edit" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Mycobacterium_tuberculosis&veaction=edit" title="Sửa đổi trang này [v]" accesskey="v"><span>Sửa đổi</span></a></li><li id="ca-more-edit" class="collapsible vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Mycobacterium_tuberculosis&action=edit" title="Sửa đổi mã nguồn của trang này [e]" accesskey="e"><span>Sửa mã nguồn</span></a></li><li id="ca-more-history" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Mycobacterium_tuberculosis&action=history"><span>Xem lịch sử</span></a></li> </ul> </div> </div> <div id="p-tb" class="vector-menu mw-portlet mw-portlet-tb" > <div class="vector-menu-heading"> Chung </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="t-whatlinkshere" class="mw-list-item"><a href="/wiki/%C4%90%E1%BA%B7c_bi%E1%BB%87t:Li%C3%AAn_k%E1%BA%BFt_%C4%91%E1%BA%BFn_%C4%91%C3%A2y/Mycobacterium_tuberculosis" title="Các trang liên kết đến đây [j]" accesskey="j"><span>Các liên kết đến đây</span></a></li><li id="t-recentchangeslinked" class="mw-list-item"><a href="/wiki/%C4%90%E1%BA%B7c_bi%E1%BB%87t:Thay_%C4%91%E1%BB%95i_li%C3%AAn_quan/Mycobacterium_tuberculosis" rel="nofollow" title="Thay đổi gần đây của các trang liên kết đến đây [k]" accesskey="k"><span>Thay đổi liên quan</span></a></li><li id="t-specialpages" class="mw-list-item"><a href="/wiki/%C4%90%E1%BA%B7c_bi%E1%BB%87t:Trang_%C4%91%E1%BA%B7c_bi%E1%BB%87t" title="Một danh sách chứa tất cả trang đặc biệt [q]" accesskey="q"><span>Trang đặc biệt</span></a></li><li id="t-permalink" class="mw-list-item"><a href="/w/index.php?title=Mycobacterium_tuberculosis&oldid=71032768" title="Liên kết thường trực đến phiên bản này của trang"><span>Liên kết thường trực</span></a></li><li id="t-info" class="mw-list-item"><a href="/w/index.php?title=Mycobacterium_tuberculosis&action=info" title="Thêm chi tiết về trang này"><span>Thông tin trang</span></a></li><li id="t-cite" class="mw-list-item"><a href="/w/index.php?title=%C4%90%E1%BA%B7c_bi%E1%BB%87t:Tr%C3%ADch_d%E1%BA%ABn&page=Mycobacterium_tuberculosis&id=71032768&wpFormIdentifier=titleform" title="Hướng dẫn cách trích dẫn trang này"><span>Trích dẫn trang này</span></a></li><li id="t-urlshortener" class="mw-list-item"><a href="/w/index.php?title=%C4%90%E1%BA%B7c_bi%E1%BB%87t:UrlShortener&url=https%3A%2F%2Fvi.wikipedia.org%2Fwiki%2FMycobacterium_tuberculosis"><span>Lấy URL ngắn gọn</span></a></li><li id="t-urlshortener-qrcode" class="mw-list-item"><a href="/w/index.php?title=%C4%90%E1%BA%B7c_bi%E1%BB%87t:QrCode&url=https%3A%2F%2Fvi.wikipedia.org%2Fwiki%2FMycobacterium_tuberculosis"><span>Tải mã QR</span></a></li> </ul> </div> </div> <div id="p-coll-print_export" class="vector-menu mw-portlet mw-portlet-coll-print_export" > <div class="vector-menu-heading"> In và xuất </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="coll-create_a_book" class="mw-list-item"><a href="/w/index.php?title=%C4%90%E1%BA%B7c_bi%E1%BB%87t:S%C3%A1ch&bookcmd=book_creator&referer=Mycobacterium+tuberculosis"><span>Tạo một quyển sách</span></a></li><li id="coll-download-as-rl" class="mw-list-item"><a href="/w/index.php?title=%C4%90%E1%BA%B7c_bi%E1%BB%87t:DownloadAsPdf&page=Mycobacterium_tuberculosis&action=show-download-screen"><span>Tải dưới dạng PDF</span></a></li><li id="t-print" class="mw-list-item"><a href="/w/index.php?title=Mycobacterium_tuberculosis&printable=yes" title="Bản để in ra của trang [p]" accesskey="p"><span>Bản để in ra</span></a></li> </ul> </div> </div> <div id="p-wikibase-otherprojects" class="vector-menu mw-portlet mw-portlet-wikibase-otherprojects" > <div class="vector-menu-heading"> Tại dự án khác </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="wb-otherproject-link wb-otherproject-commons mw-list-item"><a href="https://commons.wikimedia.org/wiki/Category:Mycobacterium_tuberculosis" hreflang="en"><span>Wikimedia Commons</span></a></li><li class="wb-otherproject-link wb-otherproject-species mw-list-item"><a href="https://species.wikimedia.org/wiki/Mycobacterium_tuberculosis" hreflang="en"><span>Wikispecies</span></a></li><li id="t-wikibase" class="wb-otherproject-link wb-otherproject-wikibase-dataitem mw-list-item"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q130971" title="Liên kết đến khoản mục kết nối trong kho dữ liệu [g]" accesskey="g"><span>Khoản mục Wikidata</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> </div> </div> </div> <div class="vector-column-end"> <div class="vector-sticky-pinned-container"> <nav class="vector-page-tools-landmark" aria-label="Công cụ trang"> <div id="vector-page-tools-pinned-container" class="vector-pinned-container"> </div> </nav> <nav class="vector-appearance-landmark" aria-label="Giao diện"> <div id="vector-appearance-pinned-container" class="vector-pinned-container"> <div id="vector-appearance" class="vector-appearance vector-pinnable-element"> <div class="vector-pinnable-header vector-appearance-pinnable-header vector-pinnable-header-pinned" data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">Giao diện</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">chuyển sang thanh bên</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" 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src="//upload.wikimedia.org/wikipedia/commons/thumb/0/0a/TB_Culture.jpg/280px-TB_Culture.jpg" decoding="async" width="280" height="186" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/0a/TB_Culture.jpg/420px-TB_Culture.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/0a/TB_Culture.jpg/560px-TB_Culture.jpg 2x" data-file-width="700" data-file-height="466" /></a></span><div style="padding:5px">Vi khuẩn <i>M. tuberculosis</i> cụm</div></td></tr><tr><th colspan="2" style="text-align:center;padding:5px;background: lightgrey;"><a href="/wiki/Ph%C3%A2n_lo%E1%BA%A1i_sinh_h%E1%BB%8Dc" title="Phân loại sinh học">Phân loại khoa học</a></th></tr><tr><th scope="row"><a href="/wiki/Gi%E1%BB%9Bi_(sinh_h%E1%BB%8Dc)" title="Giới (sinh học)">Giới</a> <small>(<i>regnum</i>)</small></th><td><a href="/wiki/Vi_khu%E1%BA%A9n" title="Vi khuẩn">Vi khuẩn</a></td></tr><tr><th scope="row"><a href="/wiki/Ng%C3%A0nh_(sinh_h%E1%BB%8Dc)" title="Ngành (sinh học)">Ngành</a> <small>(<i>phylum</i>)</small></th><td><a href="/wiki/X%E1%BA%A1_khu%E1%BA%A9n" title="Xạ khuẩn">Xạ khuẩn</a></td></tr><tr><th scope="row"><a href="/wiki/B%E1%BB%99_(sinh_h%E1%BB%8Dc)" title="Bộ (sinh học)">Bộ</a> <small>(<i>ordo</i>)</small></th><td><a href="/w/index.php?title=Actinomycetales&action=edit&redlink=1" class="new" title="Actinomycetales (trang không tồn tại)">Actinomycetales</a></td></tr><tr><th scope="row"><a href="/wiki/Ph%C3%A2n_b%E1%BB%99" class="mw-redirect" title="Phân bộ">Phân bộ</a> <small>(<i>subordo</i>)</small></th><td><a href="/w/index.php?title=Corynebacterineae&action=edit&redlink=1" class="new" title="Corynebacterineae (trang không tồn tại)">Corynebacterineae</a></td></tr><tr><th scope="row"><a href="/wiki/H%E1%BB%8D_(sinh_h%E1%BB%8Dc)" title="Họ (sinh học)">Họ</a> <small>(<i>familia</i>)</small></th><td><a href="/wiki/Mycobacterium" title="Mycobacterium">Mycobacteriaceae</a></td></tr><tr><th scope="row"><a href="/wiki/Chi_(sinh_h%E1%BB%8Dc)" title="Chi (sinh học)">Chi</a> <small>(<i>genus</i>)</small></th><td><i><a href="/wiki/Mycobacterium" title="Mycobacterium">Mycobacterium</a></i></td></tr><tr><th scope="row"><a href="/wiki/Lo%C3%A0i" title="Loài">Loài</a> <small>(<i>species</i>)</small></th><td><i><b>M. tuberculosis</b></i></td></tr><tr><th colspan="2" style="text-align:center;padding:5px;background: lightgrey;"><a href="/wiki/Danh_ph%C3%A1p_hai_ph%E1%BA%A7n" title="Danh pháp hai phần">Danh pháp hai phần</a></th></tr><tr><td colspan="2" style="text-align:center"><span style="font-weight: bold"><i>Mycobacterium tuberculosis</i></span><br /><span style="font-variant: small-caps;">Zopf 1883 </span></td></tr><tr><th colspan="2" style="text-align:center;padding:5px;background: lightgrey;"><a href="/wiki/Danh_ph%C3%A1p_%C4%91%E1%BB%93ng_ngh%C4%A9a_(ph%C3%A2n_lo%E1%BA%A1i_h%E1%BB%8Dc)" class="mw-redirect" title="Danh pháp đồng nghĩa (phân loại học)">Danh pháp đồng nghĩa</a></th></tr><tr><td colspan="2" style="text-align:center"><div align="left"> <i>Tubercle bacillus</i> <small><a href="/wiki/Robert_Koch" title="Robert Koch">Koch</a> 1882</small></div></td></tr></tbody></table> <p><b>Mycobacterium tuberculosis</b> là một loài <a href="/wiki/Vi_khu%E1%BA%A9n" title="Vi khuẩn">vi khuẩn</a> gây bệnh trong chi <i><a href="/wiki/Mycobacterium" title="Mycobacterium">Mycobacterium</a></i> và là tác nhân nhân gây bệnh của hầu hết các ca <a href="/wiki/Lao" title="Lao">bệnh lao</a> (nên còn gọi là "<i>vi khuẩn lao</i>")<sup id="cite_ref-Sherris_1-0" class="reference"><a href="#cite_note-Sherris-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>. Lần đầu được phát hiện ra vào năm 1882 bởi <a href="/wiki/Robert_Koch" title="Robert Koch">Robert Koch</a>, <i>M. tuberculosis</i> có một lớp phủ sáp bất thường trên bề mặt tế bào (chủ yếu là <a href="/w/index.php?title=Mycolic_acid&action=edit&redlink=1" class="new" title="Mycolic acid (trang không tồn tại)">mycolic acid</a>), mà làm cho các tế bào không thấm nước để nhuộm gram kỹ thuật để phát hiện, acid nhanh được sử dụng thay thế.Trực khuẩn lao được xác định dưới kính hiển vi bằng đặc tính nhuộm của nó: nó vẫn giữ màu nhuộm sau khi bị xử lý với dung dịch acid, vì vậy nó được phân loại là "trực khuẩn kháng acid" (acid-fast bacillus, viết tắt là AFB). Loài này được phân loại là gram dương do thành tế bào có chứa peptidoglycan, nhưng vì chúng có đặc tính kháng acid, kháng cồn nên được gọi là "trực khuẩn kháng acid". Đây là vi khuẩn hiếu khí và đòi hỏi mức độ cao của oxy. Trực khuẩn lao có hình dạng giống que nhỏ, có thể chịu đựng được chất sát khuẩn yếu và sống sót trong trạng thái khô trong nhiều tuần nhưng, trong điều kiện tự nhiên, chỉ có thể phát triển trong sinh vật ký chủ Bộ <a href="/wiki/Gen" title="Gen">gen</a> đã được giải mã vào năm 1998<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup>. <i>M. tuberculosis</i> phân chia sau mỗi 15-20 giờ, rất chậm so với các vi khuẩn khác có thời gian phân chia tính bằng phút (Escherichia coli (E. coli) có thể phân chia khoảng mỗi 20 phút). Nó là một trực khuẩn nhỏ, có thể chịu được thuốc sát khuẩn yếu và có thể tồn tại trong trạng thái khô trong nhiều tuần. Nó có thành tế bào bất thường, giàu chất béo (ví dụ, acid mycolic), có khả năng chịu mang đến khả năng tính kháng này và là một yếu tố độc lực quan trọng<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup>. </p><p>Khi ở trong <a href="/wiki/Ph%E1%BB%95i" title="Phổi">phổi</a>, bệnh lao M. được đưa lên bởi các <a href="/wiki/%C4%90%E1%BA%A1i_th%E1%BB%B1c_b%C3%A0o" title="Đại thực bào">đại thực bào</a> <a href="/wiki/Ph%E1%BB%95i" title="Phổi">phế nang</a>, nhưng chúng không thể tiêu hóa vi khuẩn. Thành tế bào của nó ngăn cản sự hợp nhất của các thể thực bào với một lizozimt. Cụ thể, M. tuberculosis ngăn cản phân tử bắc cầu, early endosomal autoantigen 1 (EEA1), tuy nhiên, phong tỏa này không ngăn cản sự hợp nhất của các túi chứa đầy chất dinh dưỡng. Do đó, các vi khuẩn phân chia không được kiểm soát trong các đại thực bào. Các vi khuẩn cũng mang gen UreC, ngăn ngừa quá trình axit hóa của các thể thực bào<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup>. Các vi khuẩn cũng né tránh sự tiêu diệt của đại thực bào bằng cách trung hòa các chất trung gian nitơ phản ứng. </p> <div class="mw-heading mw-heading2"><h2 id="Chú_thích"><span id="Ch.C3.BA_th.C3.ADch"></span>Chú thích</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Mycobacterium_tuberculosis&veaction=edit&section=1" title="Sửa đổi phần “Chú thích”" class="mw-editsection-visualeditor"><span>sửa</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Mycobacterium_tuberculosis&action=edit&section=1" title="Sửa mã nguồn tại đề mục: Chú thích"><span>sửa mã nguồn</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r71728118">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist columns references-column-width" style="-moz-column-width: 30em; -webkit-column-width: 30em; column-width: 30em; list-style-type: decimal;"> <ol class="references"> <li id="cite_note-Sherris-1"><b><a href="#cite_ref-Sherris_1-0">^</a></b> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r67233549">.mw-parser-output cite.citation{font-style:inherit}.mw-parser-output .citation q{quotes:"“""”""‘""’"}.mw-parser-output .id-lock-free a,.mw-parser-output .citation .cs1-lock-free a{background:linear-gradient(transparent,transparent),url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited a,.mw-parser-output .id-lock-registration a,.mw-parser-output .citation .cs1-lock-limited a,.mw-parser-output .citation .cs1-lock-registration a{background:linear-gradient(transparent,transparent),url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription a,.mw-parser-output .citation .cs1-lock-subscription a{background:linear-gradient(transparent,transparent),url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-subscription,.mw-parser-output .cs1-registration{color:#555}.mw-parser-output .cs1-subscription span,.mw-parser-output .cs1-registration span{border-bottom:1px dotted;cursor:help}.mw-parser-output .cs1-ws-icon a{background:linear-gradient(transparent,transparent),url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}.mw-parser-output code.cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;font-size:100%}.mw-parser-output .cs1-visible-error{font-size:100%}.mw-parser-output .cs1-maint{display:none;color:#33aa33;margin-left:0.3em}.mw-parser-output .cs1-format{font-size:95%}.mw-parser-output .cs1-kern-left,.mw-parser-output .cs1-kern-wl-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right,.mw-parser-output .cs1-kern-wl-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}</style><cite id="CITEREFRyan_KJ,_Ray_CG_(editors)2004" class="citation book cs1">Ryan KJ, Ray CG (editors) (2004). <a rel="nofollow" class="external text" href="https://archive.org/details/sherrismedicalmi0000unse_q1i3"><i>Sherris Medical Microbiology</i></a> (ấn bản thứ 4). McGraw Hill. <a href="/wiki/ISBN" title="ISBN">ISBN</a> <a href="/wiki/%C4%90%E1%BA%B7c_bi%E1%BB%87t:Ngu%E1%BB%93n_s%C3%A1ch/0-8385-8529-9" title="Đặc biệt:Nguồn sách/0-8385-8529-9"><bdi>0-8385-8529-9</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Sherris+Medical+Microbiology&rft.edition=4th&rft.pub=McGraw+Hill&rft.date=2004&rft.isbn=0-8385-8529-9&rft.au=Ryan+KJ%2C+Ray+CG+%28editors%29&rft_id=https%3A%2F%2Farchive.org%2Fdetails%2Fsherrismedicalmi0000unse_q1i3&rfr_id=info%3Asid%2Fvi.wikipedia.org%3AMycobacterium+tuberculosis" class="Z3988"></span><span class="cs1-maint citation-comment">Quản lý CS1: văn bản dư: danh sách tác giả (<a href="/wiki/Th%E1%BB%83_lo%E1%BA%A1i:Qu%E1%BA%A3n_l%C3%BD_CS1:_v%C4%83n_b%E1%BA%A3n_d%C6%B0:_danh_s%C3%A1ch_t%C3%A1c_gi%E1%BA%A3" title="Thể loại:Quản lý CS1: văn bản dư: danh sách tác giả">liên kết</a>)</span></span> </li> <li id="cite_note-2"><b><a href="#cite_ref-2">^</a></b> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67233549"><cite id="CITEREFCole_ST,_Brosch_R,_Parkhill_J1998" class="citation journal cs1">Cole ST, Brosch R, Parkhill J (1998). “Deciphering the biology of Mycobacterium tuberculosis from the complete genome sequence”. <i>Nature</i>. <b>393</b> (6685): 537–44. <a href="/wiki/%C4%90%E1%BB%8Bnh_danh_%C4%91%E1%BB%91i_t%C6%B0%E1%BB%A3ng_s%E1%BB%91" class="mw-redirect" title="Định danh đối tượng số">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2F31159">10.1038/31159</a>. <a href="/wiki/PMID" class="mw-redirect" title="PMID">PMID</a> <a rel="nofollow" class="external text" href="//pubmed.ncbi.nlm.nih.gov/9634230">9634230</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nature&rft.atitle=Deciphering+the+biology+of+Mycobacterium+tuberculosis+from+the+complete+genome+sequence&rft.volume=393&rft.issue=6685&rft.pages=537-44&rft.date=1998&rft_id=info%3Adoi%2F10.1038%2F31159&rft_id=info%3Apmid%2F9634230&rft.au=Cole+ST%2C+Brosch+R%2C+Parkhill+J&rfr_id=info%3Asid%2Fvi.wikipedia.org%3AMycobacterium+tuberculosis" class="Z3988"></span><span class="cs1-maint citation-comment">Quản lý CS1: nhiều tên: danh sách tác giả (<a href="/wiki/Th%E1%BB%83_lo%E1%BA%A1i:Qu%E1%BA%A3n_l%C3%BD_CS1:_nhi%E1%BB%81u_t%C3%AAn:_danh_s%C3%A1ch_t%C3%A1c_gi%E1%BA%A3" title="Thể loại:Quản lý CS1: nhiều tên: danh sách tác giả">liên kết</a>)</span></span> </li> <li id="cite_note-3"><b><a href="#cite_ref-3">^</a></b> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67233549"><cite id="CITEREFCamus_JC,_Pryor_MJ,_Médigue_C,_Cole_ST2002" class="citation journal cs1">Camus JC, Pryor MJ, Médigue C, Cole ST (2002). <a rel="nofollow" class="external text" href="http://mic.sgmjournals.org/cgi/pmidlookup?view=long&pmid=12368430">“Re-annotation of the genome sequence of Mycobacterium tuberculosis H37Rv”</a>. <i>Microbiology (Reading, Engl.)</i>. <b>148</b> (Pt 10): 2967–73. <a href="/wiki/PMID" class="mw-redirect" title="PMID">PMID</a> <a rel="nofollow" class="external text" href="//pubmed.ncbi.nlm.nih.gov/12368430">12368430</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Microbiology+%28Reading%2C+Engl.%29&rft.atitle=Re-annotation+of+the+genome+sequence+of+Mycobacterium+tuberculosis+H37Rv&rft.volume=148&rft.issue=Pt+10&rft.pages=2967-73&rft.date=2002&rft_id=info%3Apmid%2F12368430&rft.au=Camus+JC%2C+Pryor+MJ%2C+M%C3%A9digue+C%2C+Cole+ST&rft_id=http%3A%2F%2Fmic.sgmjournals.org%2Fcgi%2Fpmidlookup%3Fview%3Dlong%26pmid%3D12368430&rfr_id=info%3Asid%2Fvi.wikipedia.org%3AMycobacterium+tuberculosis" class="Z3988"></span><span class="cs1-maint citation-comment">Quản lý CS1: nhiều tên: danh sách tác giả (<a href="/wiki/Th%E1%BB%83_lo%E1%BA%A1i:Qu%E1%BA%A3n_l%C3%BD_CS1:_nhi%E1%BB%81u_t%C3%AAn:_danh_s%C3%A1ch_t%C3%A1c_gi%E1%BA%A3" title="Thể loại:Quản lý CS1: nhiều tên: danh sách tác giả">liên kết</a>)</span></span> </li> <li id="cite_note-4"><b><a href="#cite_ref-4">^</a></b> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67233549"><cite id="CITEREFMurray_PR,_Rosenthal_KS,_Pfaller_MA2005" class="citation book cs1">Murray PR, Rosenthal KS, Pfaller MA (2005). <i>Medical Microbiology</i>. Elsevier Mosby.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Medical+Microbiology&rft.pub=Elsevier+Mosby&rft.date=2005&rft.au=Murray+PR%2C+Rosenthal+KS%2C+Pfaller+MA&rfr_id=info%3Asid%2Fvi.wikipedia.org%3AMycobacterium+tuberculosis" class="Z3988"></span><span class="cs1-maint citation-comment">Quản lý CS1: nhiều tên: danh sách tác giả (<a href="/wiki/Th%E1%BB%83_lo%E1%BA%A1i:Qu%E1%BA%A3n_l%C3%BD_CS1:_nhi%E1%BB%81u_t%C3%AAn:_danh_s%C3%A1ch_t%C3%A1c_gi%E1%BA%A3" title="Thể loại:Quản lý CS1: nhiều tên: danh sách tác giả">liên kết</a>)</span></span> </li> <li id="cite_note-5"><b><a href="#cite_ref-5">^</a></b> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67233549"><cite id="CITEREFBell_E2005" class="citation journal cs1">Bell E (2005). “Vaccines: A souped-up version of BCG”. <i>Nature Reviews Immunology</i>. <b>5</b> (10): 746. <a href="/wiki/%C4%90%E1%BB%8Bnh_danh_%C4%91%E1%BB%91i_t%C6%B0%E1%BB%A3ng_s%E1%BB%91" class="mw-redirect" title="Định danh đối tượng số">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2Fnri1720">10.1038/nri1720</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nature+Reviews+Immunology&rft.atitle=Vaccines%3A+A+souped-up+version+of+BCG&rft.volume=5&rft.issue=10&rft.pages=746&rft.date=2005&rft_id=info%3Adoi%2F10.1038%2Fnri1720&rft.au=Bell+E&rfr_id=info%3Asid%2Fvi.wikipedia.org%3AMycobacterium+tuberculosis" class="Z3988"></span></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="Tham_khảo"><span id="Tham_kh.E1.BA.A3o"></span>Tham khảo</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Mycobacterium_tuberculosis&veaction=edit&section=2" title="Sửa đổi phần “Tham khảo”" class="mw-editsection-visualeditor"><span>sửa</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Mycobacterium_tuberculosis&action=edit&section=2" title="Sửa mã nguồn tại đề mục: Tham khảo"><span>sửa mã nguồn</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><span class="noviewer" typeof="mw:File"><a href="/wiki/T%E1%BA%ADp_tin:Wikispecies-logo.svg" class="mw-file-description"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/14px-Wikispecies-logo.svg.png" decoding="async" width="14" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/20px-Wikispecies-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/27px-Wikispecies-logo.svg.png 2x" data-file-width="941" data-file-height="1103" /></a></span> Dữ liệu liên quan tới <a href="https://species.wikimedia.org/wiki/Special:Search/Mycobacterium_tuberculosis" class="extiw" title="wikispecies:Special:Search/Mycobacterium tuberculosis">Mycobacterium tuberculosis</a> tại <a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a></li></ul>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; 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