Ensembl genome browser 113

<!DOCTYPE html> <html lang="en-gb"> <head> <title>Ensembl genome browser 113</title> <link rel="stylesheet" type="text/css" media="all" href="/minified/98adc89a8b4418c03a4c81d5bf76272d.css"/><link rel="stylesheet" type="text/css" media="all" href="/minified/9bafd8bb591fb39bcb766b0160434c45.image.css"/> <link rel="icon" type="image/png" href="/i/ensembl-favicon.png" /> <link href="/apple-touch-icon.png" type="image/png" rel="apple-touch-icon" /> <link rel="search" title="Ensembl (All)" href="//useast.ensembl.org/opensearch/all.xml" type="application/opensearchdescription+xml" /> <meta name="google-site-verification" content="tG3WW0bg9gNaIHH002gJl-fV2k0bzf-ZxC8pBqXTaNo" /> <meta name="viewport" content="target-densitydpi=device-dpi, width=device-width, initial-scale=1.0, maximum-scale=2.0, user-scalable=yes" /> <script>var prefetch = 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<div class="grid-col-4"> <div class="plain-box no-top-margin"> <div style="float:left;width:15%"> <div class="plain-box unbordered"> <h2>Tools</h2> <p><a href="/info/docs/tools/index.html">All tools</a></p> </div> </div> <div style="float:left;width:28%"> <div class="plain-box unbordered"> <h2><a href="/biomart/martview" class="nodeco">BioMart&nbsp ></a></h2> <p>Export custom datasets from Ensembl with this data-mining tool</p> </div> </div> <div style="float:left;width:28%"> <div class="plain-box unbordered"> <h2><a href="/Multi/Tools/Blast?db=core" class="nodeco">BLAST/BLAT&nbsp ></a></h2> <p>Search our genomes for your DNA or protein sequence</p> </div> </div> <div style="float:left;width:28%"> <div class="plain-box unbordered"> <h2><a href="/info/docs/tools/vep/" class="nodeco">Variant Effect Predictor  ></a></h2> <p>Analyse your own variants and predict the functional consequences of known and unknown variants</p> </div> </div></div> <div class="tinted-box no-top-margin home-search"> <div id="SpeciesSearch" class="js_panel"><input type="hidden" class="panel_type" value="SearchBox" /><form class="search-form homepage-search-form clear no-sel" action="/Multi/Psychic" method="get"><fieldset><input name="site" type="hidden" value="ensembl" /><div class="form-field"><div class="ff-right"><h3>Search</h3></div></div><div class="form-field"><div class="ff-right"><select name="species" class="input fselect" id="species"><option value="">All species</option><option value="help">Help and Documentation</option><option value="" disabled="disabled">---</option><optgroup label="Favourite species"><option value="Homo_sapiens">Human</option><option value="Mus_musculus">Mouse</option><option value="Danio_rerio">Zebrafish</option></optgroup><option value="" disabled="disabled">---</option><option value="Chelonoidis_abingdonii">Abingdon island giant tortoise</option><option value="Struthio_camelus_australis">African ostrich</option><option value="Gopherus_agassizii">Agassiz's desert tortoise</option><option value="Mus_spretus">Algerian mouse</option><option value="Vicugna_pacos">Alpaca</option><option value="Marmota_marmota_marmota">Alpine marmot</option><option value="Poecilia_formosa">Amazon molly</option><option value="Castor_canadensis">American beaver</option><option value="Bison_bison_bison">American bison</option><option value="Ursus_americanus">American black bear</option><option value="Neovison_vison">American mink</option><option value="Colobus_angolensis_palliatus">Angola colobus</option><option value="Camelus_dromedarius">Arabian camel</option><option value="Urocitellus_parryii">Arctic ground squirrel</option><option value="Salvator_merianae">Argentine black and white tegu</option><option value="Dasypus_novemcinctus">Armadillo</option><option value="Scleropages_formosus">Asian bonytongue</option><option value="Ursus_thibetanus_thibetanus">Asiatic black bear</option><option value="Gadus_morhua">Atlantic cod</option><option value="Clupea_harengus">Atlantic herring </option><option value="Salmo_salar">Atlantic salmon</option><option value="Crocodylus_porosus">Australian saltwater crocodile</option><option value="Labrus_bergylta">Ballan wrasse</option><option value="Lates_calcarifer">Barramundi perch</option><option value="Delphinapterus_leucas">Beluga whale</option><option value="Lonchura_striata_domestica">Bengalese finch</option><option value="Stegastes_partitus">Bicolor damselfish</option><option value="Rhinopithecus_bieti">Black snub-nosed monkey</option><option value="Sinocyclocheilus_anshuiensis">Blind barbel</option><option value="Oreochromis_aureus">Blue tilapia</option><option value="Cyanistes_caeruleus">Blue tit</option><option value="Balaenoptera_musculus">Blue whale</option><option value="Lepidothrix_coronata">Blue-crowned manakin</option><option value="Laticauda_laticaudata">Blue-ringed sea krait</option><option value="Gouania_willdenowi">Blunt-snouted clingfish</option><option value="Saimiri_boliviensis_boliviensis">Bolivian squirrel monkey</option><option value="Pan_paniscus">Bonobo</option><option value="Cavia_aperea">Brazilian guinea pig</option><option value="Salmo_trutta">Brown trout</option><option value="Melopsittacus_undulatus">Budgerigar</option><option value="Athene_cunicularia">Burrowing owl</option><option value="Haplochromis_burtoni">Burton's mouthbrooder</option><option value="Otolemur_garnettii">Bushbaby</option><option value="Ciona_intestinalis">C.intestinalis</option><option value="Ciona_savignyi">C.savignyi</option><option value="Caenorhabditis_elegans">Caenorhabditis elegans (Nematode, N2)</option><option value="Zalophus_californianus">California sea lion</option><option value="Lynx_canadensis">Canada lynx</option><option value="Felis_catus">Cat</option><option value="Pogona_vitticeps">Central bearded dragon</option><option value="Catagonus_wagneri">Chacoan peccary</option><option value="Cottoperca_gobio">Channel bull blenny</option><option value="Ictalurus_punctatus">Channel catfish</option><option value="Gallus_gallus">Chicken</option><option value="Nothoprocta_perdicaria">Chilean tinamou</option><option value="Pan_troglodytes">Chimpanzee</option><option value="Cricetulus_griseus_chok1gshd">Chinese hamster CHOK1GS</option><option value="Cricetulus_griseus_crigri">Chinese hamster CriGri</option><option value="Cricetulus_griseus_picr">Chinese hamster PICR</option><option value="Oryzias_sinensis">Chinese medaka</option><option value="Pelodiscus_sinensis">Chinese softshell turtle</option><option value="Oncorhynchus_tshawytscha">Chinook salmon</option><option value="Anabas_testudineus">Climbing perch</option><option value="Amphiprion_ocellaris">Clown anemonefish</option><option value="Latimeria_chalumnae">Coelacanth</option><option value="Oncorhynchus_kisutch">Coho salmon</option><option value="Ficedula_albicollis">Collared flycatcher</option><option value="Serinus_canaria">Common canary</option><option value="Cyprinus_carpio_carpio">Common carp</option><option value="Cyprinus_carpio_german_mirror">Common carp german mirror</option><option value="Cyprinus_carpio_hebao_red">Common carp hebao red</option><option value="Cyprinus_carpio_huanghe">Common carp huanghe</option><option value="Falco_tinnunculus">Common kestrel</option><option value="Chelydra_serpentina">Common snapping turtle</option><option value="Podarcis_muralis">Common wall lizard</option><option value="Vombatus_ursinus">Common wombat</option><option value="Propithecus_coquereli">Coquerel's sifaka</option><option value="Bos_taurus">Cow</option><option value="Macaca_fascicularis">Crab-eating macaque</option><option value="Fukomys_damarensis">Damara mole rat</option><option value="Junco_hyemalis">Dark-eyed junco</option><option value="Spermophilus_dauricus">Daurian ground squirrel</option><option value="Octodon_degus">Degu</option><option value="Denticeps_clupeoides">Denticle herring</option><option value="Canis_lupus_dingo">Dingo</option><option value="Canis_lupus_familiaris">Dog</option><option value="Tursiops_truncatus">Dolphin</option><option value="Bos_grunniens">Domestic yak</option><option value="Equus_asinus">Donkey</option><option value="Mandrillus_leucophaeus">Drill</option><option value="Drosophila_melanogaster">Drosophila melanogaster (Fruit fly)</option><option value="Anas_platyrhynchos_platyrhynchos">Duck</option><option value="Pseudonaja_textilis">Eastern brown snake</option><option value="Buteo_japonicus">Eastern buzzard</option><option value="Astatotilapia_calliptera">Eastern happy</option><option value="Anas_zonorhyncha">Eastern spot-billed duck</option><option value="Electrophorus_electricus">Electric eel</option><option value="Loxodonta_africana">Elephant</option><option value="Callorhinchus_milii">Elephant shark</option><option value="Dromaius_novaehollandiae">Emu</option><option value="Bubo_bubo">Eurasian eagle-owl</option><option value="Sciurus_vulgaris">Eurasian red squirrel</option><option value="Accipiter_nisus">Eurasian sparrowhawk</option><option value="Dicentrarchus_labrax">European seabass</option><option value="Mustela_putorius_furo">Ferret</option><option value="Takifugu_rubripes">Fugu</option><option value="Theropithecus_gelada">Gelada</option><option value="Ailuropoda_melanoleuca">Giant panda</option><option value="Nomascus_leucogenys">Gibbon</option><option value="Sparus_aurata">Gilthead seabream</option><option value="Capra_hircus">Goat</option><option value="Aquila_chrysaetos_chrysaetos">Golden eagle</option><option value="Mesocricetus_auratus">Golden Hamster</option><option value="Chrysolophus_pictus">Golden pheasant</option><option value="Rhinopithecus_roxellana">Golden snub-nosed monkey</option><option value="Manacus_vitellinus">Golden-collared manakin</option><option value="Sinocyclocheilus_grahami">Golden-line barbel</option><option value="Carassius_auratus">Goldfish</option><option value="Gopherus_evgoodei">Goodes thornscrub tortoise</option><option value="Gorilla_gorilla">Gorilla</option><option value="Erythrura_gouldiae">Gouldian finch</option><option value="Apteryx_haastii">Great spotted kiwi</option><option value="Parus_major">Great Tit</option><option value="Seriola_dumerili">Greater amberjack</option><option value="Prolemur_simus">Greater bamboo lemur</option><option value="Rhinolophus_ferrumequinum">Greater horseshoe bat</option><option value="Anolis_carolinensis">Green anole</option><option value="Cavia_porcellus">Guinea Pig</option><option value="Poecilia_reticulata">Guppy</option><option value="Eptatretus_burgeri">Hagfish</option><option value="Erinaceus_europaeus">Hedgehog</option><option value="Numida_meleagris">Helmeted guineafowl</option><option value="Sinocyclocheilus_rhinocerous">Horned golden-line barbel</option><option value="Equus_caballus">Horse</option><option value="Hucho_hucho">Huchen</option><option value="Homo_sapiens">Human</option><option value="Bos_indicus_hybrid">Hybrid - Bos Indicus</option><option value="Bos_taurus_hybrid">Hybrid - Bos Taurus</option><option value="Procavia_capensis">Hyrax</option><option value="Naja_naja">Indian cobra</option><option value="Parambassis_ranga">Indian glassy fish</option><option value="Oryzias_melastigma">Indian medaka</option><option value="Pavo_cristatus">Indian peafowl</option><option value="Oryzias_latipes">Japanese medaka HdrR</option><option value="Oryzias_latipes_hni">Japanese medaka HNI</option><option value="Oryzias_latipes_hsok">Japanese medaka HSOK</option><option value="Coturnix_japonica">Japanese quail</option><option value="Oryzias_javanicus">Javanese ricefish</option><option value="Salarias_fasciatus">Jewelled blenny</option><option value="Strigops_habroptila">Kakapo</option><option value="Dipodomys_ordii">Kangaroo rat</option><option value="Phascolarctos_cinereus">Koala</option><option value="Varanus_komodoensis">Komodo dragon</option><option value="Petromyzon_marinus">Lamprey</option><option value="Larimichthys_crocea">Large yellow croaker</option><option value="Leptobrachium_leishanense">Leishan spiny toad</option><option value="Panthera_pardus">Leopard</option><option value="Jaculus_jaculus">Lesser Egyptian jerboa</option><option value="Echinops_telfairi">Lesser hedgehog tenrec</option><option value="Panthera_leo">Lion</option><option value="Apteryx_owenii">Little spotted kiwi</option><option value="Echeneis_naucrates">Live sharksucker</option><option value="Chinchilla_lanigera">Long-tailed chinchilla</option><option value="Cyclopterus_lumpus">Lumpfish</option><option value="Neolamprologus_brichardi">Lyretail cichlid</option><option value="Aotus_nancymaae">Ma's night monkey</option><option value="Macaca_mulatta">Macaque</option><option value="Notechis_scutatus">Mainland tiger snake</option><option value="Pundamilia_nyererei">Makobe Island cichlid</option><option value="Anas_platyrhynchos">Mallard</option><option value="Kryptolebias_marmoratus">Mangrove rivulus</option><option value="Geospiza_fortis">Medium ground-finch</option><option value="Suricata_suricatta">Meerkat</option><option value="Pteropus_vampyrus">Megabat</option><option value="Astyanax_mexicanus">Mexican tetra</option><option value="Myotis_lucifugus">Microbat</option><option value="Amphilophus_citrinellus">Midas cichlid</option><option value="Meriones_unguiculatus">Mongolian gerbil</option><option value="Xiphophorus_couchianus">Monterrey platyfish</option><option value="Mus_musculus">Mouse</option><option value="Microcebus_murinus">Mouse Lemur</option><option value="Fundulus_heteroclitus">Mummichog</option><option value="Cairina_moschata_domestica">Muscovy Duck (domestic type)</option><option value="Heterocephalus_glaber_female">Naked mole-rat female</option><option value="Heterocephalus_glaber_male">Naked mole-rat male</option><option value="Monodon_monoceros">Narwhal</option><option value="Corvus_moneduloides">New Caledonian crow</option><option value="Oreochromis_niloticus">Nile tilapia</option><option value="Peromyscus_maniculatus_bairdii">Northern American deer mouse</option><option value="Esox_lucius">Northern pike</option><option value="Strix_occidentalis_caurina">Northern spotted owl</option><option value="Mola_mola">Ocean sunfish</option><option value="Apteryx_rowi">Okarito brown kiwi</option><option value="Papio_anubis">Olive baboon</option><option value="Monodelphis_domestica">Opossum</option><option value="Amphiprion_percula">Orange clownfish</option><option value="Sphaeramia_orbicularis">Orbiculate cardinalfish</option><option value="Otus_sunia">Oriental scops-owl</option><option value="Astyanax_mexicanus_pachon">Pachon cavefish</option><option value="Chrysemys_picta_bellii">Painted turtle</option><option value="Cebus_imitator">Panamanian white-faced capuchin</option><option value="Paramormyrops_kingsleyae">Paramormyrops kingsleyae</option><option value="Periophthalmus_magnuspinnatus">Periophthalmus magnuspinnatus</option><option value="Sus_scrofa">Pig</option><option value="Macaca_nemestrina">Pig-tailed macaque</option><option value="Ochotona_princeps">Pika</option><option value="Sander_lucioperca">Pike-perch</option><option value="Myripristis_murdjan">Pinecone soldierfish</option><option value="Anser_brachyrhynchus">Pink-footed goose</option><option value="Xiphophorus_maculatus">Platyfish</option><option value="Ornithorhynchus_anatinus">Platypus</option><option value="Ursus_maritimus">Polar bear</option><option value="Microtus_ochrogaster">Prairie vole</option><option value="Oryctolagus_cuniculus">Rabbit</option><option value="Oncorhynchus_mykiss">Rainbow trout</option><option value="Rattus_norvegicus">Rat</option><option value="Vulpes_vulpes">Red fox</option><option value="Pygocentrus_nattereri">Red-bellied piranha</option><option value="Erpetoichthys_calabaricus">Reedfish</option><option value="Phasianus_colchicus">Ring-necked pheasant</option><option value="Neogobius_melanostomus">Round goby</option><option value="Calidris_pugnax">Ruff</option><option value="Stachyris_ruficeps">Rufous-capped babbler</option><option value="Mus_caroli">Ryukyu mouse</option><option value="Saccharomyces_cerevisiae">Saccharomyces cerevisiae</option><option value="Poecilia_latipinna">Sailfin molly</option><option value="Ovis_aries_rambouillet">Sheep</option><option value="Cyprinodon_variegatus">Sheepshead minnow</option><option value="Poecilia_mexicana">Shortfin molly</option><option value="Sorex_araneus">Shrew</option><option value="Mus_pahari">Shrew mouse</option><option value="Betta_splendens">Siamese fighting fish</option><option value="Moschus_moschiferus">Siberian musk deer</option><option value="Zosterops_lateralis_melanops">Silver-eye</option><option value="Choloepus_hoffmanni">Sloth</option><option value="Camarhynchus_parvulus">Small tree finch</option><option value="Cercocebus_atys">Sooty mangabey</option><option value="Physeter_catodon">Sperm whale</option><option value="Acanthochromis_polyacanthus">Spiny chromis</option><option value="Calidris_pygmaea">Spoon-billed sandpiper</option><option value="Lepisosteus_oculatus">Spotted gar</option><option value="Ictidomys_tridecemlineatus">Squirrel</option><option value="Mus_spicilegus">Steppe mouse</option><option value="Gasterosteus_aculeatus">Stickleback</option><option value="Pongo_abelii">Sumatran orangutan</option><option value="Malurus_cyaneus_samueli">Superb fairywren</option><option value="Catharus_ustulatus">Swainson's thrush</option><option value="Monopterus_albus">Swamp eel</option><option value="Anser_cygnoides">Swan goose</option><option value="Carlito_syrichta">Tarsier</option><option value="Sarcophilus_harrisii">Tasmanian devil</option><option value="Tetraodon_nigroviridis">Tetraodon</option><option value="Terrapene_carolina_triunguis">Three-toed box turtle</option><option value="Panthera_tigris_altaica">Tiger</option><option value="Hippocampus_comes">Tiger tail seahorse</option><option value="Cynoglossus_semilaevis">Tongue sole</option><option value="Tupaia_belangeri">Tree Shrew</option><option value="Xenopus_tropicalis">Tropical clawed frog</option><option value="Sphenodon_punctatus">Tuatara</option><option value="Scophthalmus_maximus">Turbot</option><option value="Meleagris_gallopavo">Turkey</option><option value="Nothobranchius_furzeri">Turquoise killifish</option><option value="Piliocolobus_tephrosceles">Ugandan red Colobus</option><option value="Nannospalax_galili">Upper Galilee mountains blind mole rat</option><option value="Phocoena_sinus">Vaquita</option><option value="Chlorocebus_sabaeus">Vervet-AGM</option><option value="Notamacropus_eugenii">Wallaby</option><option value="Pelusios_castaneus">West African mud turtle</option><option value="Gambusia_affinis">Western mosquitofish</option><option value="Zonotrichia_albicollis">White-throated sparrow</option><option value="Callithrix_jacchus">White-tufted-ear marmoset</option><option value="Bos_mutus">Wild yak</option><option value="Cervus_hanglu_yarkandensis">Yarkand deer</option><option value="Amazona_collaria">Yellow-billed parrot</option><option value="Seriola_lalandi_dorsalis">Yellowtail amberjack</option><option value="Taeniopygia_guttata">Zebra finch</option><option value="Maylandia_zebra">Zebra mbuna</option><option value="Danio_rerio">Zebrafish</option><option value="Mastacembelus_armatus">Zig-zag eel</option></select><label for="q"> for</label></div></div><div class="ff-multi form-field"><div class="ff-inline ff-right"><input id="q" type="text" class="input query ftext _string inactive optional" size="50" name="q" value=" " /><input value="Go" class="fbutton" type="submit" /></div><div class="ff-notes"><p class="search-example">e.g. <a class="nowrap" href="/Multi/Psychic?q=BRCA2">BRCA2</a> or <a class="nowrap" href="/Multi/Psychic?q=rat 5:62797383-63627669">rat 5:62797383-63627669</a> or <a class="nowrap" href="/Multi/Psychic?q=rs699">rs699</a> or <a class="nowrap" href="/Multi/Psychic?q=coronary heart disease">coronary heart disease</a></p></div></div></fieldset></form></div> </div> <div class="plain-box no-top-margin"> <div class="js_panel"> <input type="hidden" class="panel_type" value="SpeciesList" /> <div class="column_wrapper"><div class="static_all_species column-two"><h3>All genomes</h3> <p><select class="fselect _all_species"><option value="">-- Select a species --</option></select></p> <h3 class="space-above"></h3> <div class="species-box-outer"> <div class="species-box"> <a href="Sus_scrofa/Info/Breeds/"><img src="/i/species/Sus_scrofa.png" alt="Pig breeds" title="Browse Pig breeds" class="badge-48"/></a> <span class="species-name">Pig breeds</span><div class="assembly"><a href="/Sus_scrofa/" class="nodeco">Pig reference genome</a> and <a href="Sus_scrofa/Info/Breeds/" class="nodeco">20 additional breeds</a></div> </div> </div> <p><a href="/info/about/species.html">View full list of all species</a></p> </div><div class="column-two fave-genomes"><h3>Favourite genomes <a href="/Multi/Account/Login" class="_list_edit modal_link"><img src="/i/16/pencil.png" class="left-half-margin" title="Edit your favourites"></a></h3><div class="hidden clear reorder_species _species_sort_container"><p>For easy access to commonly used genomes, drag from the bottom list to the top one</p> <p><strong>Favourites</strong></p> <ul class="_favourites"></ul> <p><a href="#Done" class="button _list_done">Done</a> <a href="#Reset" class="button _list_reset">Restore default list</a></p> <p><strong>Other available species</strong></p> <ul class="_species"></ul> </div><div class="species-list _species_fav_container"> <div class="species-box-outer"> <div class="species-box"> <a href="/Homo_sapiens/Info/Index"><img src="/i/species/Homo_sapiens.png" alt="Homo_sapiens" title="Browse Homo_sapiens" class="badge-48"/></a> <a href="/Homo_sapiens/Info/Index" class="species-name">Human</a> <div class="assembly">GRCh38.p14</div> </div> <a href="/info/website/tutorials/grch37.html" class="species-extra">Still using GRCh37?</a> </div> <div class="species-box-outer"> <div class="species-box"> <a href="/Mus_musculus/Info/Index"><img src="/i/species/Mus_musculus.png" alt="Mus_musculus" title="Browse Mus_musculus" class="badge-48"/></a> <a href="/Mus_musculus/Info/Index" class="species-name">Mouse</a> <div class="assembly">GRCm39</div> </div> </div> <div class="species-box-outer"> <div class="species-box"> <a href="/Danio_rerio/Info/Index"><img src="/i/species/Danio_rerio.png" alt="Danio_rerio" title="Browse Danio_rerio" class="badge-48"/></a> <a href="/Danio_rerio/Info/Index" class="species-name">Zebrafish</a> <div class="assembly">GRCz11</div> </div> </div></div><input value=" <div class="species-box-outer"> <div class="species-box"> <a href="{{species.homepage}}"><img src="{{species.img}}" alt="{{species.name}}" title="Browse {{species.name}}" class="badge-48"/></a> <a href="{{species.homepage}}" class="species-name">{{species.common}}</a> <div class="assembly">{{species.assembly}}</div> </div> {{species.extra}} </div>" name="fav_template" type="hidden" class="js_param" /><input class="js_param" type="hidden" name="list_template" value="<li id="species-{{species.key}}">{{species.common}} (<em>{{species.name}}</em>)</li>" /><input name="species_list" type="hidden" class="json js_param" value="[{"name":"Homo_sapiens","extra":"<a href=\"/info/website/tutorials/grch37.html\" class=\"species-extra\">Still using 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breeds","name":"Ovis_aries_rambouillet"}]" type="hidden" class="js_param json" name="strains_list" /><input type="hidden" class="js_param" name="ajax_refresh_url" value="Ajax/html_doc?db=core;module=GenomeList" /><input value="/Multi/Account/Favourites/Save" type="hidden" class="js_param" name="ajax_save_url" /><input name="display_limit" class="js_param" type="hidden" value="6" /><input value="Favourite" name="fave_text" class="js_param" type="hidden" /><input value="{"Actinopterygii":"Fish","Amphibia":"Amphibians","Aves":"Birds","Sauropsida":"Reptiles","Chordata":"Other chordates","LOWCOVERAGE":"Low-coverage species","Euarchontoglires":"Rodents etc.","Vertebrata":"Other vertebrates","Eukaryota":"Other eukaryotes","Mammalia":"Other mammals"}" name="taxon_labels" type="hidden" class="json js_param" /><input name="taxon_order" type="hidden" class="js_param json" 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