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Frog - Wikipedia

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id="toc-Etymology_and_taxonomy-sublist" class="vector-toc-list"> <li id="toc-Etymology" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Etymology"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.1</span> <span>Etymology</span> </div> </a> <ul id="toc-Etymology-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Taxonomy" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Taxonomy"> <div class="vector-toc-text"> <span class="vector-toc-numb">1.2</span> <span>Taxonomy</span> </div> </a> <ul id="toc-Taxonomy-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Evolution" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#Evolution"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Evolution</span> </div> </a> <button aria-controls="toc-Evolution-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Evolution subsection</span> </button> <ul id="toc-Evolution-sublist" class="vector-toc-list"> <li id="toc-Phylogeny" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Phylogeny"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.1</span> <span>Phylogeny</span> </div> </a> <ul id="toc-Phylogeny-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Morphology_and_physiology" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#Morphology_and_physiology"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Morphology and physiology</span> </div> </a> <button aria-controls="toc-Morphology_and_physiology-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Morphology and physiology subsection</span> </button> <ul id="toc-Morphology_and_physiology-sublist" class="vector-toc-list"> <li id="toc-Feet_and_legs" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Feet_and_legs"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.1</span> <span>Feet and legs</span> </div> </a> <ul id="toc-Feet_and_legs-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Skin" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Skin"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.2</span> <span>Skin</span> </div> </a> <ul id="toc-Skin-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Respiration_and_circulation" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Respiration_and_circulation"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.3</span> <span>Respiration and circulation</span> </div> </a> <ul id="toc-Respiration_and_circulation-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Digestion_and_excretion" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Digestion_and_excretion"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.4</span> <span>Digestion and excretion</span> </div> </a> <ul id="toc-Digestion_and_excretion-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Reproductive_system" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Reproductive_system"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.5</span> <span>Reproductive system</span> </div> </a> <ul id="toc-Reproductive_system-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Nervous_system" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Nervous_system"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.6</span> <span>Nervous system</span> </div> </a> <ul id="toc-Nervous_system-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Sight" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Sight"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.7</span> <span>Sight</span> </div> </a> <ul id="toc-Sight-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Hearing" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Hearing"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.8</span> <span>Hearing</span> </div> </a> <ul id="toc-Hearing-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Call" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Call"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.9</span> <span>Call</span> </div> </a> <ul id="toc-Call-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Torpor" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Torpor"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.10</span> <span>Torpor</span> </div> </a> <ul id="toc-Torpor-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Locomotion" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#Locomotion"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Locomotion</span> </div> </a> <button aria-controls="toc-Locomotion-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Locomotion subsection</span> </button> <ul id="toc-Locomotion-sublist" class="vector-toc-list"> <li id="toc-Jumping" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Jumping"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.1</span> <span>Jumping</span> </div> </a> <ul id="toc-Jumping-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Walking_and_running" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Walking_and_running"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.2</span> <span>Walking and running</span> </div> </a> <ul id="toc-Walking_and_running-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Swimming" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Swimming"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.3</span> <span>Swimming</span> </div> </a> <ul id="toc-Swimming-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Burrowing" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Burrowing"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.4</span> <span>Burrowing</span> </div> </a> <ul id="toc-Burrowing-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Climbing" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Climbing"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.5</span> <span>Climbing</span> </div> </a> <ul id="toc-Climbing-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Gliding" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Gliding"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.6</span> <span>Gliding</span> </div> </a> <ul id="toc-Gliding-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Life_history" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#Life_history"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>Life history</span> </div> </a> <button aria-controls="toc-Life_history-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Life history subsection</span> </button> <ul id="toc-Life_history-sublist" class="vector-toc-list"> <li id="toc-Reproduction" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Reproduction"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.1</span> <span>Reproduction</span> </div> </a> <ul id="toc-Reproduction-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Life_cycle" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Life_cycle"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.2</span> <span>Life cycle</span> </div> </a> <ul id="toc-Life_cycle-sublist" class="vector-toc-list"> <li id="toc-Eggs_/_frogspawn" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Eggs_/_frogspawn"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.2.1</span> <span>Eggs / frogspawn</span> </div> </a> <ul id="toc-Eggs_/_frogspawn-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Tadpoles" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Tadpoles"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.2.2</span> <span>Tadpoles</span> </div> </a> <ul id="toc-Tadpoles-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Metamorphosis" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Metamorphosis"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.2.3</span> <span>Metamorphosis</span> </div> </a> <ul id="toc-Metamorphosis-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Adults" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Adults"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.2.4</span> <span>Adults</span> </div> </a> <ul id="toc-Adults-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Parental_care" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Parental_care"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.3</span> <span>Parental care</span> </div> </a> <ul id="toc-Parental_care-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Defence" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#Defence"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>Defence</span> </div> </a> <ul id="toc-Defence-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Distribution" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#Distribution"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>Distribution</span> </div> </a> <ul id="toc-Distribution-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Conservation" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#Conservation"> <div class="vector-toc-text"> <span class="vector-toc-numb">8</span> <span>Conservation</span> </div> </a> <ul id="toc-Conservation-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Human_uses" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#Human_uses"> <div class="vector-toc-text"> <span class="vector-toc-numb">9</span> <span>Human uses</span> </div> </a> <button aria-controls="toc-Human_uses-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Human uses subsection</span> </button> <ul id="toc-Human_uses-sublist" class="vector-toc-list"> <li id="toc-Culinary" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Culinary"> <div class="vector-toc-text"> <span class="vector-toc-numb">9.1</span> <span>Culinary</span> </div> </a> <ul id="toc-Culinary-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Scientific_research" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Scientific_research"> <div class="vector-toc-text"> <span class="vector-toc-numb">9.2</span> <span>Scientific research</span> </div> </a> <ul id="toc-Scientific_research-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Pharmaceutical" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Pharmaceutical"> <div class="vector-toc-text"> <span class="vector-toc-numb">9.3</span> <span>Pharmaceutical</span> </div> </a> <ul id="toc-Pharmaceutical-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Cultural_significance" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Cultural_significance"> <div class="vector-toc-text"> <span class="vector-toc-numb">9.4</span> <span>Cultural significance</span> </div> </a> <ul id="toc-Cultural_significance-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">10</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Further_reading" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#Further_reading"> <div class="vector-toc-text"> <span class="vector-toc-numb">11</span> <span>Further reading</span> </div> </a> <ul id="toc-Further_reading-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">12</span> <span>External links</span> </div> </a> <button aria-controls="toc-External_links-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle External links subsection</span> </button> <ul id="toc-External_links-sublist" class="vector-toc-list"> <li id="toc-Media" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Media"> <div class="vector-toc-text"> <span class="vector-toc-numb">12.1</span> <span>Media</span> </div> </a> <ul id="toc-Media-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" title="Table of Contents" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Toggle the table of contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Frog</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 171 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-171" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">171 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ace mw-list-item"><a href="https://ace.wikipedia.org/wiki/Canggu%C3%ABk" title="Cangguëk – Acehnese" lang="ace" hreflang="ace" data-title="Cangguëk" data-language-autonym="Acèh" data-language-local-name="Acehnese" class="interlanguage-link-target"><span>Acèh</span></a></li><li class="interlanguage-link interwiki-af mw-list-item"><a href="https://af.wikipedia.org/wiki/Padda" title="Padda – Afrikaans" lang="af" hreflang="af" data-title="Padda" data-language-autonym="Afrikaans" data-language-local-name="Afrikaans" class="interlanguage-link-target"><span>Afrikaans</span></a></li><li class="interlanguage-link interwiki-als mw-list-item"><a href="https://als.wikipedia.org/wiki/Froschlurche" title="Froschlurche – Alemannic" lang="gsw" hreflang="gsw" data-title="Froschlurche" data-language-autonym="Alemannisch" data-language-local-name="Alemannic" class="interlanguage-link-target"><span>Alemannisch</span></a></li><li class="interlanguage-link interwiki-am mw-list-item"><a href="https://am.wikipedia.org/wiki/%E1%8C%93%E1%8C%89%E1%8A%95%E1%89%B8%E1%88%AD" title="ጓጉንቸር – Amharic" lang="am" hreflang="am" data-title="ጓጉንቸር" data-language-autonym="አማርኛ" data-language-local-name="Amharic" class="interlanguage-link-target"><span>አማርኛ</span></a></li><li class="interlanguage-link interwiki-ang mw-list-item"><a href="https://ang.wikipedia.org/wiki/Frogga" title="Frogga – Old English" lang="ang" hreflang="ang" data-title="Frogga" data-language-autonym="Ænglisc" data-language-local-name="Old English" class="interlanguage-link-target"><span>Ænglisc</span></a></li><li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B6%D9%81%D8%AF%D8%B9" title="ضفدع – Arabic" lang="ar" hreflang="ar" data-title="ضفدع" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-an mw-list-item"><a href="https://an.wikipedia.org/wiki/Anura" title="Anura – Aragonese" lang="an" hreflang="an" data-title="Anura" data-language-autonym="Aragonés" data-language-local-name="Aragonese" class="interlanguage-link-target"><span>Aragonés</span></a></li><li class="interlanguage-link interwiki-roa-rup mw-list-item"><a href="https://roa-rup.wikipedia.org/wiki/Broaticu" title="Broaticu – Aromanian" lang="rup" hreflang="rup" data-title="Broaticu" data-language-autonym="Armãneashti" data-language-local-name="Aromanian" class="interlanguage-link-target"><span>Armãneashti</span></a></li><li class="interlanguage-link interwiki-frp mw-list-item"><a href="https://frp.wikipedia.org/wiki/Anuros" title="Anuros – Arpitan" lang="frp" hreflang="frp" data-title="Anuros" data-language-autonym="Arpetan" data-language-local-name="Arpitan" class="interlanguage-link-target"><span>Arpetan</span></a></li><li class="interlanguage-link interwiki-as mw-list-item"><a href="https://as.wikipedia.org/wiki/%E0%A6%AD%E0%A7%87%E0%A6%95%E0%A7%81%E0%A6%B2%E0%A7%80" title="ভেকুলী – Assamese" lang="as" hreflang="as" data-title="ভেকুলী" data-language-autonym="অসমীয়া" data-language-local-name="Assamese" class="interlanguage-link-target"><span>অসমীয়া</span></a></li><li class="interlanguage-link interwiki-ast mw-list-item"><a href="https://ast.wikipedia.org/wiki/Anura" title="Anura – Asturian" lang="ast" hreflang="ast" data-title="Anura" data-language-autonym="Asturianu" data-language-local-name="Asturian" class="interlanguage-link-target"><span>Asturianu</span></a></li><li class="interlanguage-link interwiki-gn mw-list-item"><a href="https://gn.wikipedia.org/wiki/Ju%E2%80%99i" title="Ju’i – Guarani" lang="gn" hreflang="gn" data-title="Ju’i" data-language-autonym="Avañe&#039;ẽ" data-language-local-name="Guarani" class="interlanguage-link-target"><span>Avañe&#039;ẽ</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Quyruqsuzlar" title="Quyruqsuzlar – Azerbaijani" lang="az" hreflang="az" data-title="Quyruqsuzlar" data-language-autonym="Azərbaycanca" data-language-local-name="Azerbaijani" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D9%82%D9%88%DB%8C%D8%B1%D9%88%D9%82%D8%B3%D9%88%D8%B2%D9%84%D8%A7%D8%B1" title="قویروقسوزلار – South Azerbaijani" lang="azb" hreflang="azb" data-title="قویروقسوزلار" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-ban mw-list-item"><a href="https://ban.wikipedia.org/wiki/Katak" title="Katak – Balinese" lang="ban" hreflang="ban" data-title="Katak" data-language-autonym="Basa Bali" data-language-local-name="Balinese" class="interlanguage-link-target"><span>Basa Bali</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%AC%E0%A7%8D%E0%A6%AF%E0%A6%BE%E0%A6%99" title="ব্যাঙ – Bangla" lang="bn" hreflang="bn" data-title="ব্যাঙ" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-zh-min-nan mw-list-item"><a href="https://zh-min-nan.wikipedia.org/wiki/Ch%C3%BAi-ke" title="Chúi-ke – Minnan" lang="nan" hreflang="nan" data-title="Chúi-ke" data-language-autonym="閩南語 / Bân-lâm-gú" data-language-local-name="Minnan" class="interlanguage-link-target"><span>閩南語 / Bân-lâm-gú</span></a></li><li class="interlanguage-link interwiki-ba mw-list-item"><a href="https://ba.wikipedia.org/wiki/%D2%A0%D0%BE%D0%B9%D1%80%D0%BE%D2%A1%D2%BB%D0%BE%D2%99%D2%99%D0%B0%D1%80" title="Ҡойроҡһоҙҙар – Bashkir" lang="ba" hreflang="ba" data-title="Ҡойроҡһоҙҙар" data-language-autonym="Башҡортса" data-language-local-name="Bashkir" class="interlanguage-link-target"><span>Башҡортса</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%91%D1%8F%D1%81%D1%85%D0%B2%D0%BE%D1%81%D1%82%D1%8B%D1%8F_%D0%B7%D0%B5%D0%BC%D0%BD%D0%B0%D0%B2%D0%BE%D0%B4%D0%BD%D1%8B%D1%8F" title="Бясхвостыя земнаводныя – Belarusian" lang="be" hreflang="be" data-title="Бясхвостыя земнаводныя" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-be-x-old mw-list-item"><a href="https://be-tarask.wikipedia.org/wiki/%D0%91%D1%8F%D1%81%D1%85%D0%B2%D0%BE%D1%81%D1%82%D1%8B%D1%8F" title="Бясхвостыя – Belarusian (Taraškievica orthography)" lang="be-tarask" hreflang="be-tarask" data-title="Бясхвостыя" data-language-autonym="Беларуская (тарашкевіца)" data-language-local-name="Belarusian (Taraškievica orthography)" class="interlanguage-link-target"><span>Беларуская (тарашкевіца)</span></a></li><li class="interlanguage-link interwiki-bh mw-list-item"><a href="https://bh.wikipedia.org/wiki/%E0%A4%AE%E0%A5%87%E0%A4%98%E0%A4%BE" title="मेघा – Bhojpuri" lang="bh" hreflang="bh" data-title="मेघा" data-language-autonym="भोजपुरी" data-language-local-name="Bhojpuri" class="interlanguage-link-target"><span>भोजपुरी</span></a></li><li class="interlanguage-link interwiki-bcl mw-list-item"><a href="https://bcl.wikipedia.org/wiki/Talapang" title="Talapang – Central Bikol" lang="bcl" hreflang="bcl" data-title="Talapang" data-language-autonym="Bikol Central" data-language-local-name="Central Bikol" class="interlanguage-link-target"><span>Bikol Central</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%91%D0%B5%D0%B7%D0%BE%D0%BF%D0%B0%D1%88%D0%B0%D1%82%D0%B8_%D0%B7%D0%B5%D0%BC%D0%BD%D0%BE%D0%B2%D0%BE%D0%B4%D0%BD%D0%B8" title="Безопашати земноводни – Bulgarian" lang="bg" hreflang="bg" data-title="Безопашати земноводни" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-bar mw-list-item"><a href="https://bar.wikipedia.org/wiki/Froschlurche" title="Froschlurche – Bavarian" lang="bar" hreflang="bar" data-title="Froschlurche" data-language-autonym="Boarisch" data-language-local-name="Bavarian" class="interlanguage-link-target"><span>Boarisch</span></a></li><li class="interlanguage-link interwiki-bs mw-list-item"><a href="https://bs.wikipedia.org/wiki/%C5%BDaba" title="Žaba – Bosnian" lang="bs" hreflang="bs" data-title="Žaba" data-language-autonym="Bosanski" data-language-local-name="Bosnian" class="interlanguage-link-target"><span>Bosanski</span></a></li><li class="interlanguage-link interwiki-br mw-list-item"><a href="https://br.wikipedia.org/wiki/Anura" title="Anura – Breton" lang="br" hreflang="br" data-title="Anura" data-language-autonym="Brezhoneg" data-language-local-name="Breton" class="interlanguage-link-target"><span>Brezhoneg</span></a></li><li class="interlanguage-link interwiki-ca badge-Q17437796 badge-featuredarticle mw-list-item" title="featured article badge"><a href="https://ca.wikipedia.org/wiki/Anurs" title="Anurs – Catalan" lang="ca" hreflang="ca" data-title="Anurs" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cv mw-list-item"><a href="https://cv.wikipedia.org/wiki/%D0%A5%D3%B3%D1%80%D0%B5%D1%81%C4%95%D1%80_%C3%A7%C4%95%D1%80%D1%82%D0%B8-%D1%88%D1%8B%D0%B2%D1%80%D0%B8%D1%81%D0%B5%D0%BC" title="Хӳресĕр çĕрти-шыврисем – Chuvash" lang="cv" hreflang="cv" data-title="Хӳресĕр çĕрти-шыврисем" data-language-autonym="Чӑвашла" data-language-local-name="Chuvash" class="interlanguage-link-target"><span>Чӑвашла</span></a></li><li class="interlanguage-link interwiki-ceb mw-list-item"><a href="https://ceb.wikipedia.org/wiki/Baki" title="Baki – Cebuano" lang="ceb" hreflang="ceb" data-title="Baki" data-language-autonym="Cebuano" data-language-local-name="Cebuano" class="interlanguage-link-target"><span>Cebuano</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/%C5%BD%C3%A1by" title="Žáby – Czech" lang="cs" hreflang="cs" data-title="Žáby" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-sn mw-list-item"><a href="https://sn.wikipedia.org/wiki/Datya" title="Datya – Shona" lang="sn" hreflang="sn" data-title="Datya" data-language-autonym="ChiShona" data-language-local-name="Shona" class="interlanguage-link-target"><span>ChiShona</span></a></li><li class="interlanguage-link interwiki-cy mw-list-item"><a href="https://cy.wikipedia.org/wiki/Llyffant" title="Llyffant – Welsh" lang="cy" hreflang="cy" data-title="Llyffant" data-language-autonym="Cymraeg" data-language-local-name="Welsh" class="interlanguage-link-target"><span>Cymraeg</span></a></li><li class="interlanguage-link interwiki-dag mw-list-item"><a href="https://dag.wikipedia.org/wiki/L%C9%94%C5%8B" title="Lɔŋ – Dagbani" lang="dag" hreflang="dag" data-title="Lɔŋ" data-language-autonym="Dagbanli" data-language-local-name="Dagbani" class="interlanguage-link-target"><span>Dagbanli</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Springpadder" title="Springpadder – Danish" lang="da" hreflang="da" data-title="Springpadder" data-language-autonym="Dansk" data-language-local-name="Danish" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-pdc mw-list-item"><a href="https://pdc.wikipedia.org/wiki/Frosch" title="Frosch – Pennsylvania German" lang="pdc" hreflang="pdc" data-title="Frosch" data-language-autonym="Deitsch" data-language-local-name="Pennsylvania German" class="interlanguage-link-target"><span>Deitsch</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Froschlurche" title="Froschlurche – German" lang="de" hreflang="de" data-title="Froschlurche" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-nv mw-list-item"><a href="https://nv.wikipedia.org/wiki/Ch%CA%BCa%C5%82" title="Chʼał – Navajo" lang="nv" hreflang="nv" data-title="Chʼał" data-language-autonym="Diné bizaad" data-language-local-name="Navajo" class="interlanguage-link-target"><span>Diné bizaad</span></a></li><li class="interlanguage-link interwiki-dty mw-list-item"><a href="https://dty.wikipedia.org/wiki/%E0%A4%AD%E0%A5%87%E0%A4%97%E0%A5%81%E0%A4%A4%E0%A5%8B" title="भेगुतो – Doteli" lang="dty" hreflang="dty" data-title="भेगुतो" data-language-autonym="डोटेली" data-language-local-name="Doteli" class="interlanguage-link-target"><span>डोटेली</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/P%C3%A4riskonnalised" title="Päriskonnalised – Estonian" lang="et" hreflang="et" data-title="Päriskonnalised" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%92%CE%AC%CF%84%CF%81%CE%B1%CF%87%CE%BF%CF%82" title="Βάτραχος – Greek" lang="el" hreflang="el" data-title="Βάτραχος" data-language-autonym="Ελληνικά" data-language-local-name="Greek" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Anura" title="Anura – Spanish" lang="es" hreflang="es" data-title="Anura" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Anuroj" title="Anuroj – Esperanto" lang="eo" hreflang="eo" data-title="Anuroj" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Anura" title="Anura – Basque" lang="eu" hreflang="eu" data-title="Anura" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D9%82%D9%88%D8%B1%D8%A8%D8%A7%D8%BA%D9%87" title="قورباغه – Persian" lang="fa" hreflang="fa" data-title="قورباغه" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-hif mw-list-item"><a href="https://hif.wikipedia.org/wiki/Medhak" title="Medhak – Fiji Hindi" lang="hif" hreflang="hif" data-title="Medhak" data-language-autonym="Fiji Hindi" data-language-local-name="Fiji Hindi" class="interlanguage-link-target"><span>Fiji Hindi</span></a></li><li class="interlanguage-link interwiki-fo mw-list-item"><a href="https://fo.wikipedia.org/wiki/Padda" title="Padda – Faroese" lang="fo" hreflang="fo" data-title="Padda" data-language-autonym="Føroyskt" data-language-local-name="Faroese" class="interlanguage-link-target"><span>Føroyskt</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Anoures" title="Anoures – French" lang="fr" hreflang="fr" data-title="Anoures" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Frog" title="Frog – Irish" lang="ga" hreflang="ga" data-title="Frog" data-language-autonym="Gaeilge" data-language-local-name="Irish" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-gv mw-list-item"><a href="https://gv.wikipedia.org/wiki/Rannag" title="Rannag – Manx" lang="gv" hreflang="gv" data-title="Rannag" data-language-autonym="Gaelg" data-language-local-name="Manx" class="interlanguage-link-target"><span>Gaelg</span></a></li><li class="interlanguage-link interwiki-gd mw-list-item"><a href="https://gd.wikipedia.org/wiki/Losgann" title="Losgann – Scottish Gaelic" lang="gd" hreflang="gd" data-title="Losgann" data-language-autonym="Gàidhlig" data-language-local-name="Scottish Gaelic" class="interlanguage-link-target"><span>Gàidhlig</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Anuro" title="Anuro – Galician" lang="gl" hreflang="gl" data-title="Anuro" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-inh mw-list-item"><a href="https://inh.wikipedia.org/wiki/%D0%9F%D1%85%D1%8C%D0%B8%D0%B4%D0%B0%D1%80%D1%87" title="Пхьидарч – Ingush" lang="inh" hreflang="inh" data-title="Пхьидарч" data-language-autonym="ГӀалгӀай" data-language-local-name="Ingush" class="interlanguage-link-target"><span>ГӀалгӀай</span></a></li><li class="interlanguage-link interwiki-gan mw-list-item"><a href="https://gan.wikipedia.org/wiki/%E8%9D%A6%E8%9F%86" title="蝦蟆 – Gan" lang="gan" hreflang="gan" data-title="蝦蟆" data-language-autonym="贛語" data-language-local-name="Gan" class="interlanguage-link-target"><span>贛語</span></a></li><li class="interlanguage-link interwiki-ki mw-list-item"><a href="https://ki.wikipedia.org/wiki/K%C4%A9%C5%A9ra" title="Kĩũra – Kikuyu" lang="ki" hreflang="ki" data-title="Kĩũra" data-language-autonym="Gĩkũyũ" data-language-local-name="Kikuyu" class="interlanguage-link-target"><span>Gĩkũyũ</span></a></li><li class="interlanguage-link interwiki-glk mw-list-item"><a href="https://glk.wikipedia.org/wiki/%D8%B4%D9%88%D8%A8%D9%88%DA%A9" title="شوبوک – Gilaki" lang="glk" hreflang="glk" data-title="شوبوک" data-language-autonym="گیلکی" data-language-local-name="Gilaki" class="interlanguage-link-target"><span>گیلکی</span></a></li><li class="interlanguage-link interwiki-hak mw-list-item"><a href="https://hak.wikipedia.org/wiki/Ko%C3%A1i-%C3%A9" title="Koái-é – Hakka Chinese" lang="hak" hreflang="hak" data-title="Koái-é" data-language-autonym="客家語 / Hak-kâ-ngî" data-language-local-name="Hakka Chinese" class="interlanguage-link-target"><span>客家語 / Hak-kâ-ngî</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EA%B0%9C%EA%B5%AC%EB%A6%AC%EB%AA%A9" title="개구리목 – Korean" lang="ko" hreflang="ko" data-title="개구리목" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-ha mw-list-item"><a href="https://ha.wikipedia.org/wiki/Kwa%C9%97o" title="Kwaɗo – Hausa" lang="ha" hreflang="ha" data-title="Kwaɗo" data-language-autonym="Hausa" data-language-local-name="Hausa" class="interlanguage-link-target"><span>Hausa</span></a></li><li class="interlanguage-link interwiki-hy mw-list-item"><a href="https://hy.wikipedia.org/wiki/%D4%B1%D5%B6%D5%BA%D5%B8%D5%B9_%D5%A5%D6%80%D5%AF%D5%AF%D5%A5%D5%B6%D6%81%D5%A1%D5%B2%D5%B6%D5%A5%D6%80" title="Անպոչ երկկենցաղներ – Armenian" lang="hy" hreflang="hy" data-title="Անպոչ երկկենցաղներ" data-language-autonym="Հայերեն" data-language-local-name="Armenian" class="interlanguage-link-target"><span>Հայերեն</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%AE%E0%A5%87%E0%A4%82%E0%A4%A2%E0%A4%95" title="मेंढक – Hindi" lang="hi" hreflang="hi" data-title="मेंढक" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/%C5%BDabe" title="Žabe – Croatian" lang="hr" hreflang="hr" data-title="Žabe" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-gor mw-list-item"><a href="https://gor.wikipedia.org/wiki/Tumbihe" title="Tumbihe – Gorontalo" lang="gor" hreflang="gor" data-title="Tumbihe" data-language-autonym="Bahasa Hulontalo" data-language-local-name="Gorontalo" class="interlanguage-link-target"><span>Bahasa Hulontalo</span></a></li><li class="interlanguage-link interwiki-ilo mw-list-item"><a href="https://ilo.wikipedia.org/wiki/Tukak" title="Tukak – Iloko" lang="ilo" hreflang="ilo" data-title="Tukak" data-language-autonym="Ilokano" data-language-local-name="Iloko" class="interlanguage-link-target"><span>Ilokano</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Anura" title="Anura – Indonesian" lang="id" hreflang="id" data-title="Anura" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-ia mw-list-item"><a href="https://ia.wikipedia.org/wiki/Anura" title="Anura – Interlingua" lang="ia" hreflang="ia" data-title="Anura" data-language-autonym="Interlingua" data-language-local-name="Interlingua" class="interlanguage-link-target"><span>Interlingua</span></a></li><li class="interlanguage-link interwiki-iu mw-list-item"><a href="https://iu.wikipedia.org/wiki/%E1%93%88%E1%95%8C%E1%94%A9%E1%96%85" title="ᓈᕌᔩᖅ – Inuktitut" lang="iu" hreflang="iu" data-title="ᓈᕌᔩᖅ" data-language-autonym="ᐃᓄᒃᑎᑐᑦ / inuktitut" data-language-local-name="Inuktitut" class="interlanguage-link-target"><span>ᐃᓄᒃᑎᑐᑦ / inuktitut</span></a></li><li class="interlanguage-link interwiki-ik mw-list-item"><a href="https://ik.wikipedia.org/wiki/Naa%C4%A1aayiq" title="Naaġaayiq – Inupiaq" lang="ik" hreflang="ik" data-title="Naaġaayiq" data-language-autonym="Iñupiatun" data-language-local-name="Inupiaq" class="interlanguage-link-target"><span>Iñupiatun</span></a></li><li class="interlanguage-link interwiki-os mw-list-item"><a href="https://os.wikipedia.org/wiki/%D0%A5%C3%A6%D1%84%D1%81" title="Хæфс – Ossetic" lang="os" hreflang="os" data-title="Хæфс" data-language-autonym="Ирон" data-language-local-name="Ossetic" class="interlanguage-link-target"><span>Ирон</span></a></li><li class="interlanguage-link interwiki-zu mw-list-item"><a href="https://zu.wikipedia.org/wiki/Iselesele" title="Iselesele – Zulu" lang="zu" hreflang="zu" data-title="Iselesele" data-language-autonym="IsiZulu" data-language-local-name="Zulu" class="interlanguage-link-target"><span>IsiZulu</span></a></li><li class="interlanguage-link interwiki-is mw-list-item"><a href="https://is.wikipedia.org/wiki/Froskar" title="Froskar – Icelandic" lang="is" hreflang="is" data-title="Froskar" data-language-autonym="Íslenska" data-language-local-name="Icelandic" class="interlanguage-link-target"><span>Íslenska</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Anura" title="Anura – Italian" lang="it" hreflang="it" data-title="Anura" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%97%D7%A1%D7%A8%D7%99_%D7%96%D7%A0%D7%91" title="חסרי זנב – Hebrew" lang="he" hreflang="he" data-title="חסרי זנב" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-jv mw-list-item"><a href="https://jv.wikipedia.org/wiki/Kodhok" title="Kodhok – Javanese" lang="jv" hreflang="jv" data-title="Kodhok" data-language-autonym="Jawa" data-language-local-name="Javanese" class="interlanguage-link-target"><span>Jawa</span></a></li><li class="interlanguage-link interwiki-kn mw-list-item"><a href="https://kn.wikipedia.org/wiki/%E0%B2%95%E0%B2%AA%E0%B3%8D%E0%B2%AA%E0%B3%86" title="ಕಪ್ಪೆ – Kannada" lang="kn" hreflang="kn" data-title="ಕಪ್ಪೆ" data-language-autonym="ಕನ್ನಡ" data-language-local-name="Kannada" class="interlanguage-link-target"><span>ಕನ್ನಡ</span></a></li><li class="interlanguage-link interwiki-pam mw-list-item"><a href="https://pam.wikipedia.org/wiki/Tugak" title="Tugak – Pampanga" lang="pam" hreflang="pam" data-title="Tugak" data-language-autonym="Kapampangan" data-language-local-name="Pampanga" class="interlanguage-link-target"><span>Kapampangan</span></a></li><li class="interlanguage-link interwiki-ka mw-list-item"><a href="https://ka.wikipedia.org/wiki/%E1%83%A3%E1%83%99%E1%83%A3%E1%83%93%E1%83%9D_%E1%83%90%E1%83%9B%E1%83%A4%E1%83%98%E1%83%91%E1%83%98%E1%83%94%E1%83%91%E1%83%98" title="უკუდო ამფიბიები – Georgian" lang="ka" hreflang="ka" data-title="უკუდო ამფიბიები" data-language-autonym="ქართული" data-language-local-name="Georgian" class="interlanguage-link-target"><span>ქართული</span></a></li><li class="interlanguage-link interwiki-ks mw-list-item"><a href="https://ks.wikipedia.org/wiki/%D9%85%D9%90%D9%86%DB%81%D9%90" title="مِنہِ – Kashmiri" lang="ks" hreflang="ks" data-title="مِنہِ" data-language-autonym="कॉशुर / کٲشُر" data-language-local-name="Kashmiri" class="interlanguage-link-target"><span>कॉशुर / کٲشُر</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D2%9A%D2%B1%D0%B9%D1%80%D1%8B%D2%9B%D1%81%D1%8B%D0%B7_%D2%9B%D0%BE%D1%81%D0%BC%D0%B5%D0%BA%D0%B5%D0%BD%D0%B4%D1%96%D0%BB%D0%B5%D1%80" title="Құйрықсыз қосмекенділер – Kazakh" lang="kk" hreflang="kk" data-title="Құйрықсыз қосмекенділер" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-kw mw-list-item"><a href="https://kw.wikipedia.org/wiki/Kwilkyn" title="Kwilkyn – Cornish" lang="kw" hreflang="kw" data-title="Kwilkyn" data-language-autonym="Kernowek" data-language-local-name="Cornish" class="interlanguage-link-target"><span>Kernowek</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Chura" title="Chura – Swahili" lang="sw" hreflang="sw" data-title="Chura" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-ht mw-list-item"><a href="https://ht.wikipedia.org/wiki/Krapo" title="Krapo – Haitian Creole" lang="ht" hreflang="ht" data-title="Krapo" data-language-autonym="Kreyòl ayisyen" data-language-local-name="Haitian Creole" class="interlanguage-link-target"><span>Kreyòl ayisyen</span></a></li><li class="interlanguage-link interwiki-ku mw-list-item"><a href="https://ku.wikipedia.org/wiki/Beq" title="Beq – Kurdish" lang="ku" hreflang="ku" data-title="Beq" data-language-autonym="Kurdî" data-language-local-name="Kurdish" class="interlanguage-link-target"><span>Kurdî</span></a></li><li class="interlanguage-link interwiki-ky mw-list-item"><a href="https://ky.wikipedia.org/wiki/%D0%91%D0%B0%D0%BA%D0%B0" title="Бака – Kyrgyz" lang="ky" hreflang="ky" data-title="Бака" data-language-autonym="Кыргызча" data-language-local-name="Kyrgyz" class="interlanguage-link-target"><span>Кыргызча</span></a></li><li class="interlanguage-link interwiki-mrj mw-list-item"><a href="https://mrj.wikipedia.org/wiki/%D0%9F%D0%B0%D1%87%D0%B4%D1%8B%D0%BC%D1%8B%D0%B2%D0%BB%D3%93" title="Пачдымывлӓ – Western Mari" lang="mrj" hreflang="mrj" data-title="Пачдымывлӓ" data-language-autonym="Кырык мары" data-language-local-name="Western Mari" class="interlanguage-link-target"><span>Кырык мары</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Anura" title="Anura – Latin" lang="la" hreflang="la" data-title="Anura" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-lv mw-list-item"><a href="https://lv.wikipedia.org/wiki/Bezastainie_abinieki" title="Bezastainie abinieki – Latvian" lang="lv" hreflang="lv" data-title="Bezastainie abinieki" data-language-autonym="Latviešu" data-language-local-name="Latvian" class="interlanguage-link-target"><span>Latviešu</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Beuodegiai_varliagyviai" title="Beuodegiai varliagyviai – Lithuanian" lang="lt" hreflang="lt" data-title="Beuodegiai varliagyviai" data-language-autonym="Lietuvių" data-language-local-name="Lithuanian" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-li mw-list-item"><a href="https://li.wikipedia.org/wiki/Kwakkers_en_kr%C3%B3ddele" title="Kwakkers en króddele – Limburgish" lang="li" hreflang="li" data-title="Kwakkers en króddele" data-language-autonym="Limburgs" data-language-local-name="Limburgish" class="interlanguage-link-target"><span>Limburgs</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/B%C3%A9k%C3%A1k" title="Békák – Hungarian" lang="hu" hreflang="hu" data-title="Békák" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-mk mw-list-item"><a href="https://mk.wikipedia.org/wiki/%D0%96%D0%B0%D0%B1%D0%B0" title="Жаба – Macedonian" lang="mk" hreflang="mk" data-title="Жаба" data-language-autonym="Македонски" data-language-local-name="Macedonian" class="interlanguage-link-target"><span>Македонски</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%A4%E0%B4%B5%E0%B4%B3" title="തവള – Malayalam" lang="ml" hreflang="ml" data-title="തവള" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-mr mw-list-item"><a href="https://mr.wikipedia.org/wiki/%E0%A4%AC%E0%A5%87%E0%A4%A1%E0%A5%82%E0%A4%95" title="बेडूक – Marathi" lang="mr" hreflang="mr" data-title="बेडूक" data-language-autonym="मराठी" data-language-local-name="Marathi" class="interlanguage-link-target"><span>मराठी</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%B6%D9%81%D8%AF%D8%B9" title="ضفدع – Egyptian Arabic" lang="arz" hreflang="arz" data-title="ضفدع" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D9%88%DA%A9" title="وک – Mazanderani" lang="mzn" hreflang="mzn" data-title="وک" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Katak" title="Katak – Malay" lang="ms" hreflang="ms" data-title="Katak" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-min mw-list-item"><a href="https://min.wikipedia.org/wiki/Koncek" title="Koncek – Minangkabau" lang="min" hreflang="min" data-title="Koncek" data-language-autonym="Minangkabau" data-language-local-name="Minangkabau" class="interlanguage-link-target"><span>Minangkabau</span></a></li><li class="interlanguage-link interwiki-cdo mw-list-item"><a href="https://cdo.wikipedia.org/wiki/U%C3%B2ng-b%C4%AD" title="Uòng-bĭ – Mindong" lang="cdo" hreflang="cdo" data-title="Uòng-bĭ" data-language-autonym="閩東語 / Mìng-dĕ̤ng-ngṳ̄" data-language-local-name="Mindong" class="interlanguage-link-target"><span>閩東語 / Mìng-dĕ̤ng-ngṳ̄</span></a></li><li class="interlanguage-link interwiki-mdf mw-list-item"><a href="https://mdf.wikipedia.org/wiki/%D0%92%D0%B0%D1%82%D1%80%D0%B0%D0%BA%D1%88" title="Ватракш – Moksha" lang="mdf" hreflang="mdf" data-title="Ватракш" data-language-autonym="Мокшень" data-language-local-name="Moksha" class="interlanguage-link-target"><span>Мокшень</span></a></li><li class="interlanguage-link interwiki-mn mw-list-item"><a href="https://mn.wikipedia.org/wiki/%D0%9C%D1%8D%D0%BB%D1%85%D0%B8%D0%B9" title="Мэлхий – Mongolian" lang="mn" hreflang="mn" data-title="Мэлхий" data-language-autonym="Монгол" data-language-local-name="Mongolian" class="interlanguage-link-target"><span>Монгол</span></a></li><li class="interlanguage-link interwiki-my mw-list-item"><a href="https://my.wikipedia.org/wiki/%E1%80%96%E1%80%AC%E1%80%B8" title="ဖား – Burmese" lang="my" hreflang="my" data-title="ဖား" data-language-autonym="မြန်မာဘာသာ" data-language-local-name="Burmese" class="interlanguage-link-target"><span>မြန်မာဘာသာ</span></a></li><li class="interlanguage-link interwiki-nl badge-Q17437796 badge-featuredarticle mw-list-item" title="featured article badge"><a href="https://nl.wikipedia.org/wiki/Kikkers" title="Kikkers – Dutch" lang="nl" hreflang="nl" data-title="Kikkers" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-nds-nl mw-list-item"><a href="https://nds-nl.wikipedia.org/wiki/Kikvors" title="Kikvors – Low Saxon" lang="nds-NL" hreflang="nds-NL" data-title="Kikvors" data-language-autonym="Nedersaksies" data-language-local-name="Low Saxon" class="interlanguage-link-target"><span>Nedersaksies</span></a></li><li class="interlanguage-link interwiki-ne mw-list-item"><a href="https://ne.wikipedia.org/wiki/%E0%A4%AD%E0%A5%8D%E0%A4%AF%E0%A4%BE%E0%A4%97%E0%A5%81%E0%A4%A4%E0%A5%8B" title="भ्यागुतो – Nepali" lang="ne" hreflang="ne" data-title="भ्यागुतो" data-language-autonym="नेपाली" data-language-local-name="Nepali" class="interlanguage-link-target"><span>नेपाली</span></a></li><li class="interlanguage-link interwiki-new mw-list-item"><a href="https://new.wikipedia.org/wiki/%E0%A4%AC%E0%A5%8D%E0%A4%AF%E0%A4%BE%E0%A4%82%E0%A4%9A%E0%A4%BE" title="ब्यांचा – Newari" lang="new" hreflang="new" data-title="ब्यांचा" data-language-autonym="नेपाल भाषा" data-language-local-name="Newari" class="interlanguage-link-target"><span>नेपाल भाषा</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%AB%E3%82%A8%E3%83%AB" title="カエル – Japanese" lang="ja" hreflang="ja" data-title="カエル" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-ce mw-list-item"><a href="https://ce.wikipedia.org/wiki/%D0%9F%D1%85%D1%8C%D0%B8%D0%B4%D0%B0%D1%80%D1%87%D0%B8%D0%B9" title="Пхьидарчий – Chechen" lang="ce" hreflang="ce" data-title="Пхьидарчий" data-language-autonym="Нохчийн" data-language-local-name="Chechen" class="interlanguage-link-target"><span>Нохчийн</span></a></li><li class="interlanguage-link interwiki-frr mw-list-item"><a href="https://frr.wikipedia.org/wiki/Hobelfasker_an_Poden" title="Hobelfasker an Poden – Northern Frisian" lang="frr" hreflang="frr" data-title="Hobelfasker an Poden" data-language-autonym="Nordfriisk" data-language-local-name="Northern Frisian" class="interlanguage-link-target"><span>Nordfriisk</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Springpadder" title="Springpadder – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Springpadder" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-nn mw-list-item"><a href="https://nn.wikipedia.org/wiki/Halelause_padder" title="Halelause padder – Norwegian Nynorsk" lang="nn" hreflang="nn" data-title="Halelause padder" data-language-autonym="Norsk nynorsk" data-language-local-name="Norwegian Nynorsk" class="interlanguage-link-target"><span>Norsk nynorsk</span></a></li><li class="interlanguage-link interwiki-nrm mw-list-item"><a href="https://nrm.wikipedia.org/wiki/Ra%C3%AEnotte" title="Raînotte – Norman" lang="nrf" hreflang="nrf" data-title="Raînotte" data-language-autonym="Nouormand" data-language-local-name="Norman" class="interlanguage-link-target"><span>Nouormand</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Anura" title="Anura – Occitan" lang="oc" hreflang="oc" data-title="Anura" data-language-autonym="Occitan" data-language-local-name="Occitan" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-or mw-list-item"><a href="https://or.wikipedia.org/wiki/%E0%AC%AC%E0%AD%87%E0%AC%99%E0%AD%8D%E0%AC%97" title="ବେଙ୍ଗ – Odia" lang="or" hreflang="or" data-title="ବେଙ୍ଗ" data-language-autonym="ଓଡ଼ିଆ" data-language-local-name="Odia" class="interlanguage-link-target"><span>ଓଡ଼ିଆ</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/Qurbaqa" title="Qurbaqa – Uzbek" lang="uz" hreflang="uz" data-title="Qurbaqa" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Uzbek" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-pa mw-list-item"><a href="https://pa.wikipedia.org/wiki/%E0%A8%A1%E0%A9%B1%E0%A8%A1%E0%A9%82" title="ਡੱਡੂ – Punjabi" lang="pa" hreflang="pa" data-title="ਡੱਡੂ" data-language-autonym="ਪੰਜਾਬੀ" data-language-local-name="Punjabi" class="interlanguage-link-target"><span>ਪੰਜਾਬੀ</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%DA%88%DA%88%D9%88" title="ڈڈو – Western Punjabi" lang="pnb" hreflang="pnb" data-title="ڈڈو" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-ps mw-list-item"><a href="https://ps.wikipedia.org/wiki/%DA%86%D9%88%D9%86%DA%AF%DA%9A%D9%87" title="چونگښه – Pashto" lang="ps" hreflang="ps" data-title="چونگښه" data-language-autonym="پښتو" data-language-local-name="Pashto" class="interlanguage-link-target"><span>پښتو</span></a></li><li class="interlanguage-link interwiki-km mw-list-item"><a href="https://km.wikipedia.org/wiki/%E1%9E%80%E1%9E%84%E1%9F%92%E1%9E%80%E1%9F%82%E1%9E%94" title="កង្កែប – Khmer" lang="km" hreflang="km" data-title="កង្កែប" data-language-autonym="ភាសាខ្មែរ" data-language-local-name="Khmer" class="interlanguage-link-target"><span>ភាសាខ្មែរ</span></a></li><li class="interlanguage-link interwiki-pms mw-list-item"><a href="https://pms.wikipedia.org/wiki/Ran-a" title="Ran-a – Piedmontese" lang="pms" hreflang="pms" data-title="Ran-a" data-language-autonym="Piemontèis" data-language-local-name="Piedmontese" class="interlanguage-link-target"><span>Piemontèis</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/P%C5%82azy_bezogonowe" title="Płazy bezogonowe – Polish" lang="pl" hreflang="pl" data-title="Płazy bezogonowe" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Anura" title="Anura – Portuguese" lang="pt" hreflang="pt" data-title="Anura" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Broasc%C4%83" title="Broască – Romanian" lang="ro" hreflang="ro" data-title="Broască" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-qu mw-list-item"><a href="https://qu.wikipedia.org/wiki/K%27ayra" title="K&#039;ayra – Quechua" lang="qu" hreflang="qu" data-title="K&#039;ayra" data-language-autonym="Runa Simi" data-language-local-name="Quechua" class="interlanguage-link-target"><span>Runa Simi</span></a></li><li class="interlanguage-link interwiki-rue mw-list-item"><a href="https://rue.wikipedia.org/wiki/%D0%96%D0%B0%D0%B1%D0%B0" title="Жаба – Rusyn" lang="rue" hreflang="rue" data-title="Жаба" data-language-autonym="Русиньскый" data-language-local-name="Rusyn" class="interlanguage-link-target"><span>Русиньскый</span></a></li><li class="interlanguage-link interwiki-ru badge-Q17437796 badge-featuredarticle mw-list-item" title="featured article badge"><a href="https://ru.wikipedia.org/wiki/%D0%91%D0%B5%D1%81%D1%85%D0%B2%D0%BE%D1%81%D1%82%D1%8B%D0%B5_%D0%B7%D0%B5%D0%BC%D0%BD%D0%BE%D0%B2%D0%BE%D0%B4%D0%BD%D1%8B%D0%B5" title="Бесхвостые земноводные – Russian" lang="ru" hreflang="ru" data-title="Бесхвостые земноводные" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sah mw-list-item"><a href="https://sah.wikipedia.org/wiki/%D0%91%D0%B0%D2%95%D0%B0" title="Баҕа – Yakut" lang="sah" hreflang="sah" data-title="Баҕа" data-language-autonym="Саха тыла" data-language-local-name="Yakut" class="interlanguage-link-target"><span>Саха тыла</span></a></li><li class="interlanguage-link interwiki-szy mw-list-item"><a href="https://szy.wikipedia.org/wiki/apap" title="apap – Sakizaya" lang="szy" hreflang="szy" data-title="apap" data-language-autonym="Sakizaya" data-language-local-name="Sakizaya" class="interlanguage-link-target"><span>Sakizaya</span></a></li><li class="interlanguage-link interwiki-sat mw-list-item"><a href="https://sat.wikipedia.org/wiki/%E1%B1%A8%E1%B1%9A%E1%B1%B4%E1%B1%AE" title="ᱨᱚᱴᱮ – Santali" lang="sat" hreflang="sat" data-title="ᱨᱚᱴᱮ" data-language-autonym="ᱥᱟᱱᱛᱟᱲᱤ" data-language-local-name="Santali" class="interlanguage-link-target"><span>ᱥᱟᱱᱛᱟᱲᱤ</span></a></li><li class="interlanguage-link interwiki-sco mw-list-item"><a href="https://sco.wikipedia.org/wiki/Frog" title="Frog – Scots" lang="sco" hreflang="sco" data-title="Frog" data-language-autonym="Scots" data-language-local-name="Scots" class="interlanguage-link-target"><span>Scots</span></a></li><li class="interlanguage-link interwiki-nso mw-list-item"><a href="https://nso.wikipedia.org/wiki/Segwagwa" title="Segwagwa – Northern Sotho" lang="nso" hreflang="nso" data-title="Segwagwa" data-language-autonym="Sesotho sa Leboa" data-language-local-name="Northern Sotho" class="interlanguage-link-target"><span>Sesotho sa Leboa</span></a></li><li class="interlanguage-link interwiki-sq mw-list-item"><a href="https://sq.wikipedia.org/wiki/Bretkosa" title="Bretkosa – Albanian" lang="sq" hreflang="sq" data-title="Bretkosa" data-language-autonym="Shqip" data-language-local-name="Albanian" class="interlanguage-link-target"><span>Shqip</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Anura" title="Anura – Simple English" lang="en-simple" hreflang="en-simple" data-title="Anura" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sd mw-list-item"><a href="https://sd.wikipedia.org/wiki/%DA%8F%D9%8A%DA%8F%D8%B1" title="ڏيڏر – Sindhi" lang="sd" hreflang="sd" data-title="ڏيڏر" data-language-autonym="سنڌي" data-language-local-name="Sindhi" class="interlanguage-link-target"><span>سنڌي</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/%C5%BDabotvar%C3%A9" title="Žabotvaré – Slovak" lang="sk" hreflang="sk" data-title="Žabotvaré" data-language-autonym="Slovenčina" data-language-local-name="Slovak" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Brezrepe_dvo%C5%BEivke" title="Brezrepe dvoživke – Slovenian" lang="sl" hreflang="sl" data-title="Brezrepe dvoživke" data-language-autonym="Slovenščina" data-language-local-name="Slovenian" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-so mw-list-item"><a href="https://so.wikipedia.org/wiki/Rah" title="Rah – Somali" lang="so" hreflang="so" data-title="Rah" data-language-autonym="Soomaaliga" data-language-local-name="Somali" class="interlanguage-link-target"><span>Soomaaliga</span></a></li><li class="interlanguage-link interwiki-ckb mw-list-item"><a href="https://ckb.wikipedia.org/wiki/%D8%A8%DB%86%D9%82" title="بۆق – Central Kurdish" lang="ckb" hreflang="ckb" data-title="بۆق" data-language-autonym="کوردی" data-language-local-name="Central Kurdish" class="interlanguage-link-target"><span>کوردی</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%96%D0%B0%D0%B1%D0%B5" title="Жабе – Serbian" lang="sr" hreflang="sr" data-title="Жабе" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/%C5%BDaba" title="Žaba – Serbo-Croatian" lang="sh" hreflang="sh" data-title="Žaba" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="Serbo-Croatian" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-su mw-list-item"><a href="https://su.wikipedia.org/wiki/Bangkong" title="Bangkong – Sundanese" lang="su" hreflang="su" data-title="Bangkong" data-language-autonym="Sunda" data-language-local-name="Sundanese" class="interlanguage-link-target"><span>Sunda</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Sammakot" title="Sammakot – Finnish" lang="fi" hreflang="fi" data-title="Sammakot" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Stj%C3%A4rtl%C3%B6sa_groddjur" title="Stjärtlösa groddjur – Swedish" lang="sv" hreflang="sv" data-title="Stjärtlösa groddjur" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-tl mw-list-item"><a href="https://tl.wikipedia.org/wiki/Palaka" title="Palaka – Tagalog" lang="tl" hreflang="tl" data-title="Palaka" data-language-autonym="Tagalog" data-language-local-name="Tagalog" class="interlanguage-link-target"><span>Tagalog</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%A4%E0%AE%B5%E0%AE%B3%E0%AF%88" title="தவளை – Tamil" lang="ta" hreflang="ta" data-title="தவளை" data-language-autonym="தமிழ்" data-language-local-name="Tamil" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-kab mw-list-item"><a href="https://kab.wikipedia.org/wiki/Agru" title="Agru – Kabyle" lang="kab" hreflang="kab" data-title="Agru" data-language-autonym="Taqbaylit" data-language-local-name="Kabyle" class="interlanguage-link-target"><span>Taqbaylit</span></a></li><li class="interlanguage-link interwiki-roa-tara mw-list-item"><a href="https://roa-tara.wikipedia.org/wiki/Maraj%C3%B2ttole" title="Marajòttole – Tarantino" lang="nap-x-tara" hreflang="nap-x-tara" data-title="Marajòttole" data-language-autonym="Tarandíne" data-language-local-name="Tarantino" class="interlanguage-link-target"><span>Tarandíne</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://tt.wikipedia.org/wiki/%D0%9A%D0%BE%D0%B9%D1%80%D1%8B%D0%BA%D1%81%D1%8B%D0%B7%D0%BB%D0%B0%D1%80" title="Койрыксызлар – Tatar" lang="tt" hreflang="tt" data-title="Койрыксызлар" data-language-autonym="Татарча / tatarça" data-language-local-name="Tatar" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li><li class="interlanguage-link interwiki-tay mw-list-item"><a href="https://tay.wikipedia.org/wiki/Qpatung" title="Qpatung – Atayal" lang="tay" hreflang="tay" data-title="Qpatung" data-language-autonym="Tayal" data-language-local-name="Atayal" class="interlanguage-link-target"><span>Tayal</span></a></li><li class="interlanguage-link interwiki-te mw-list-item"><a href="https://te.wikipedia.org/wiki/%E0%B0%95%E0%B0%AA%E0%B1%8D%E0%B0%AA" title="కప్ప – Telugu" lang="te" hreflang="te" data-title="కప్ప" data-language-autonym="తెలుగు" data-language-local-name="Telugu" class="interlanguage-link-target"><span>తెలుగు</span></a></li><li class="interlanguage-link interwiki-th mw-list-item"><a href="https://th.wikipedia.org/wiki/%E0%B8%AD%E0%B8%B1%E0%B8%99%E0%B8%94%E0%B8%B1%E0%B8%9A%E0%B8%81%E0%B8%9A" title="อันดับกบ – Thai" lang="th" hreflang="th" data-title="อันดับกบ" data-language-autonym="ไทย" data-language-local-name="Thai" class="interlanguage-link-target"><span>ไทย</span></a></li><li class="interlanguage-link interwiki-din mw-list-item"><a href="https://din.wikipedia.org/wiki/Aguek" title="Aguek – Dinka" lang="din" hreflang="din" data-title="Aguek" data-language-autonym="Thuɔŋjäŋ" data-language-local-name="Dinka" class="interlanguage-link-target"><span>Thuɔŋjäŋ</span></a></li><li class="interlanguage-link interwiki-tg mw-list-item"><a href="https://tg.wikipedia.org/wiki/%D2%9A%D1%83%D1%80%D0%B1%D0%BE%D2%9B%D0%B0" title="Қурбоқа – Tajik" lang="tg" hreflang="tg" data-title="Қурбоқа" data-language-autonym="Тоҷикӣ" data-language-local-name="Tajik" class="interlanguage-link-target"><span>Тоҷикӣ</span></a></li><li class="interlanguage-link interwiki-chr mw-list-item"><a href="https://chr.wikipedia.org/wiki/%E1%8F%A9%E1%8E%B6%E1%8F%8F" title="ᏩᎶᏏ – Cherokee" lang="chr" hreflang="chr" data-title="ᏩᎶᏏ" data-language-autonym="ᏣᎳᎩ" data-language-local-name="Cherokee" class="interlanguage-link-target"><span>ᏣᎳᎩ</span></a></li><li class="interlanguage-link interwiki-tcy mw-list-item"><a href="https://tcy.wikipedia.org/wiki/%E0%B2%95%E0%B2%AA%E0%B3%8D%E0%B2%AA%E0%B3%86" title="ಕಪ್ಪೆ – Tulu" lang="tcy" hreflang="tcy" data-title="ಕಪ್ಪೆ" data-language-autonym="ತುಳು" data-language-local-name="Tulu" class="interlanguage-link-target"><span>ತುಳು</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Kurba%C4%9Fa" title="Kurbağa – Turkish" lang="tr" hreflang="tr" data-title="Kurbağa" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%91%D0%B5%D0%B7%D1%85%D0%B2%D0%BE%D1%81%D1%82%D1%96" title="Безхвості – Ukrainian" lang="uk" hreflang="uk" data-title="Безхвості" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D9%85%DB%8C%D9%86%DA%88%DA%A9" title="مینڈک – Urdu" lang="ur" hreflang="ur" data-title="مینڈک" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-ug mw-list-item"><a href="https://ug.wikipedia.org/wiki/%D9%BE%D8%A7%D9%82%D8%A7" title="پاقا – Uyghur" lang="ug" hreflang="ug" data-title="پاقا" data-language-autonym="ئۇيغۇرچە / Uyghurche" data-language-local-name="Uyghur" class="interlanguage-link-target"><span>ئۇيغۇرچە / Uyghurche</span></a></li><li class="interlanguage-link interwiki-za mw-list-item"><a href="https://za.wikipedia.org/wiki/Goep" title="Goep – Zhuang" lang="za" hreflang="za" data-title="Goep" data-language-autonym="Vahcuengh" data-language-local-name="Zhuang" class="interlanguage-link-target"><span>Vahcuengh</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/B%E1%BB%99_Kh%C3%B4ng_%C4%91u%C3%B4i" title="Bộ Không đuôi – Vietnamese" lang="vi" hreflang="vi" data-title="Bộ Không đuôi" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-fiu-vro mw-list-item"><a href="https://fiu-vro.wikipedia.org/wiki/Kunn" title="Kunn – Võro" lang="vro" hreflang="vro" data-title="Kunn" data-language-autonym="Võro" data-language-local-name="Võro" class="interlanguage-link-target"><span>Võro</span></a></li><li class="interlanguage-link interwiki-wa mw-list-item"><a href="https://wa.wikipedia.org/wiki/Rinne_(biesse)" title="Rinne (biesse) – Walloon" lang="wa" hreflang="wa" data-title="Rinne (biesse)" data-language-autonym="Walon" data-language-local-name="Walloon" class="interlanguage-link-target"><span>Walon</span></a></li><li class="interlanguage-link interwiki-guc mw-list-item"><a href="https://guc.wikipedia.org/wiki/Eper%C3%BC%27%C3%BCi" title="Eperü&#039;üi – Wayuu" lang="guc" hreflang="guc" data-title="Eperü&#039;üi" data-language-autonym="Wayuunaiki" data-language-local-name="Wayuu" class="interlanguage-link-target"><span>Wayuunaiki</span></a></li><li class="interlanguage-link interwiki-zh-classical mw-list-item"><a href="https://zh-classical.wikipedia.org/wiki/%E8%9B%A4" title="蛤 – Literary Chinese" lang="lzh" hreflang="lzh" data-title="蛤" data-language-autonym="文言" data-language-local-name="Literary Chinese" class="interlanguage-link-target"><span>文言</span></a></li><li class="interlanguage-link interwiki-vls mw-list-item"><a href="https://vls.wikipedia.org/wiki/Puutn" title="Puutn – West Flemish" lang="vls" hreflang="vls" data-title="Puutn" data-language-autonym="West-Vlams" data-language-local-name="West Flemish" class="interlanguage-link-target"><span>West-Vlams</span></a></li><li class="interlanguage-link interwiki-war mw-list-item"><a href="https://war.wikipedia.org/wiki/Pakla" title="Pakla – Waray" lang="war" hreflang="war" data-title="Pakla" data-language-autonym="Winaray" data-language-local-name="Waray" class="interlanguage-link-target"><span>Winaray</span></a></li><li class="interlanguage-link interwiki-wuu mw-list-item"><a href="https://wuu.wikipedia.org/wiki/%E7%94%B0%E9%B8%A1" title="田鸡 – Wu" lang="wuu" hreflang="wuu" data-title="田鸡" data-language-autonym="吴语" data-language-local-name="Wu" class="interlanguage-link-target"><span>吴语</span></a></li><li class="interlanguage-link interwiki-yi mw-list-item"><a href="https://yi.wikipedia.org/wiki/%D7%96%D7%A9%D7%90%D7%91%D7%A2" title="זשאבע – Yiddish" lang="yi" hreflang="yi" data-title="זשאבע" data-language-autonym="ייִדיש" data-language-local-name="Yiddish" class="interlanguage-link-target"><span>ייִדיש</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/K%E1%BB%8D%CC%80nk%E1%BB%8D%CC%80" title="Kọ̀nkọ̀ – Yoruba" lang="yo" hreflang="yo" data-title="Kọ̀nkọ̀" data-language-autonym="Yorùbá" data-language-local-name="Yoruba" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E8%9B%A4%F0%A7%8A%85" title="蛤𧊅 – Cantonese" lang="yue" hreflang="yue" data-title="蛤𧊅" data-language-autonym="粵語" data-language-local-name="Cantonese" class="interlanguage-link-target"><span>粵語</span></a></li><li class="interlanguage-link interwiki-zea mw-list-item"><a href="https://zea.wikipedia.org/wiki/Puut%27n" title="Puut&#039;n – Zeelandic" lang="zea" hreflang="zea" data-title="Puut&#039;n" data-language-autonym="Zeêuws" data-language-local-name="Zeelandic" class="interlanguage-link-target"><span>Zeêuws</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E6%97%A0%E5%B0%BE%E7%9B%AE" title="无尾目 – Chinese" lang="zh" hreflang="zh" data-title="无尾目" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q53636#sitelinks-wikipedia" title="Edit interlanguage links" class="wbc-editpage">Edit links</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Namespaces"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li 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For other uses, see <a href="/wiki/Frog_(disambiguation)" class="mw-disambig" title="Frog (disambiguation)">Frog (disambiguation)</a>.</div> <p class="mw-empty-elt"> </p> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">Frogs<br /><div style="font-size: 85%;">Temporal range: <a href="/wiki/Early_Jurassic" title="Early Jurassic">Early Jurassic</a> – <a href="/wiki/Holocene" title="Holocene">Present</a>, <span class="noprint"><span style="display:inline-block;"></span><span style="display:inline-block;">200–0&#160;<a href="/wiki/Megaannum" class="mw-redirect" title="Megaannum">Ma</a></span> <span style="display:inline-block;"></span><div id="Timeline-row" style="margin: 4px auto 0; clear:both; width:220px; padding:0px; height:18px; overflow:visible; white-space:nowrap; border:1px #666; border-style:solid none; position:relative; z-index:0; font-size:97%;"> <div style="position:absolute; height:100%; left:0px; width:207.23076923077px; padding-left:5px; text-align:left; background-color:rgb(254,217,106); background-image: linear-gradient(to right, rgba(255,255,255,1), rgba(254,217,106,1) 15%, rgba(254,217,106,1));"><a href="/wiki/Precambrian" title="Precambrian">PreꞒ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,160,86); left:37.636923076923px; width:17.583076923077px;"><a href="/wiki/Cambrian" title="Cambrian">Ꞓ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(0,146,112); left:55.22px; width:14.807692307692px;"><a href="/wiki/Ordovician" title="Ordovician">O</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(179,225,182); left:70.027692307692px; width:7.9470769230769px;"><a href="/wiki/Silurian" title="Silurian">S</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(203,140,55); left:77.974769230769px; width:20.564923076923px;"><a href="/wiki/Devonian" title="Devonian">D</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(103,165,153); left:98.539692307692px; width:20.294153846154px;"><a href="/wiki/Carboniferous" title="Carboniferous">C</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(240,64,40); left:118.83384615385px; width:15.907015384615px;"><a href="/wiki/Permian" title="Permian">P</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(129,43,146); left:134.74086153846px; width:17.092984615385px;"><a href="/wiki/Triassic" title="Triassic">T</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(52,178,201); left:151.83384615385px; width:19.732307692308px;"><a href="/wiki/Jurassic" title="Jurassic">J</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,198,78); left:171.56615384615px; width:26.095384615385px;"><a href="/wiki/Cretaceous" title="Cretaceous">K</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(253,154,82); left:197.66153846154px; width:14.540307692308px;"><a href="/wiki/Paleogene" title="Paleogene">Pg</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(255,230,25); left:212.20184615385px; width:6.9249230769231px;"><a href="/wiki/Neogene" title="Neogene">N</a></div> <div id="end-border" style="position:absolute; height:100%; background-color:#666; width:1px; left:219px"></div><div style="margin:0 auto; line-height:0; clear:both; width:220px; padding:0px; height:8px; overflow:visible; background-color:transparent; position:relative; top:-4px; z-index:100;"><div style="position:absolute; height:8px; left:152.30769230769px; width:67.692307692308px; background-color:#360; opacity:0.42;"></div> <div style="position:absolute; height:8px; left:152.30769230769px; width:67.692307692308px; background-color:#360; opacity:1;"></div> <div style="position:absolute; height:6px; top:1px; left:153.30769230769px; width:65.692307692308px; background-color:#6c3;"></div> </div> </div></span></div> </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Anoures.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/0/01/Anoures.jpg/220px-Anoures.jpg" decoding="async" width="220" height="319" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/01/Anoures.jpg/330px-Anoures.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/01/Anoures.jpg/440px-Anoures.jpg 2x" data-file-width="2066" data-file-height="3000" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%">Various types of frog </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Anura" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Chordate" title="Chordate">Chordata</a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Amphibian" title="Amphibian">Amphibia</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Salientia" title="Salientia">Salientia</a> </td></tr> <tr> <td>Order: </td> <td><a class="mw-selflink selflink">Anura</a><br /><small><a href="/wiki/Andr%C3%A9_Marie_Constant_Dum%C3%A9ril" title="André Marie Constant Duméril">Duméril</a>, 1806 (as Anoures)</small> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">Subgroups </th></tr> <tr> <td colspan="2" style="text-align: left"> <p>See text </p> </td></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Distribution.anura.1.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/fb/Distribution.anura.1.png/220px-Distribution.anura.1.png" decoding="async" width="220" height="102" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/fb/Distribution.anura.1.png/330px-Distribution.anura.1.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/fb/Distribution.anura.1.png/440px-Distribution.anura.1.png 2x" data-file-width="1357" data-file-height="628" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%">Native distribution of frogs (in green) </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p>A <b>frog</b> is any member of a diverse and largely <a href="/wiki/Carnivore" title="Carnivore">carnivorous</a> group of short-bodied, tailless <a href="/wiki/Amphibian" title="Amphibian">amphibians</a> composing the <a href="/wiki/Order_(biology)" title="Order (biology)">order</a> <b>Anura</b><sup id="cite_ref-AOTW_1-0" class="reference"><a href="#cite_note-AOTW-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> (coming from the <a href="/wiki/Ancient_Greek" title="Ancient Greek">Ancient Greek</a> <span title="Ancient Greek (to 1453)-language text"><span lang="grc">ἀνούρα</span></span>, literally 'without tail'). The oldest fossil "proto-frog" <i><a href="/wiki/Triadobatrachus" title="Triadobatrachus">Triadobatrachus</a></i> is known from the <a href="/wiki/Early_Triassic" title="Early Triassic">Early Triassic</a> of <a href="/wiki/Madagascar" title="Madagascar">Madagascar</a> (250<span class="nowrap">&#160;</span><a href="/wiki/Myr" class="mw-redirect" title="Myr">million years ago</a>), but <a href="/wiki/Molecular_clock" title="Molecular clock">molecular clock dating</a> suggests their split from other amphibians may extend further back to the <a href="/wiki/Permian" title="Permian">Permian</a>, 265<span class="nowrap">&#160;</span>million years ago. Frogs are widely distributed, ranging from the <a href="/wiki/Tropics" title="Tropics">tropics</a> to <a href="/wiki/Subarctic" title="Subarctic">subarctic</a> regions, but the greatest concentration of <a href="/wiki/Species_diversity" title="Species diversity">species diversity</a> is in <a href="/wiki/Tropical_rainforest" title="Tropical rainforest">tropical rainforest</a>. Frogs account for around 88% of extant amphibian species. They are also one of the five most diverse <a href="/wiki/Vertebrate" title="Vertebrate">vertebrate</a> orders. Warty frog species tend to be called <a href="/wiki/Toad" title="Toad">toads</a>, but the distinction between frogs and toads is informal, not from <a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">taxonomy</a> or evolutionary history. </p><p>An adult frog has a stout body, protruding <a href="/wiki/Eye" title="Eye">eyes</a>, anteriorly-attached <a href="/wiki/Tongue" title="Tongue">tongue</a>, limbs folded underneath, and no <a href="/wiki/Tail" title="Tail">tail</a> (the tail of <a href="/wiki/Tailed_frog" title="Tailed frog">tailed frogs</a> is an extension of the male cloaca). Frogs have <a href="/wiki/Gland" title="Gland">glandular</a> skin, with secretions ranging from distasteful to toxic. Their skin varies in colour from well-<a href="/wiki/Camouflage" title="Camouflage">camouflaged</a> dappled brown, grey and green to vivid patterns of bright red or yellow and black to show toxicity and <a href="/wiki/Aposematism" title="Aposematism">ward off predators</a>. Adult frogs live in fresh water and on dry land; some species are adapted for living underground or in trees. </p><p>Frogs typically lay their <a href="/wiki/Egg" title="Egg">eggs</a> in the water. The eggs hatch into aquatic <a href="/wiki/Larva" title="Larva">larvae</a> called <a href="/wiki/Tadpole" title="Tadpole">tadpoles</a> that have tails and internal <a href="/wiki/Gill" title="Gill">gills</a>. They have highly specialised rasping mouth parts suitable for <a href="/wiki/Herbivore" title="Herbivore">herbivorous</a>, <a href="/wiki/Omnivore" title="Omnivore">omnivorous</a> or <a href="/wiki/Planktivore" title="Planktivore">planktivorous</a> diets. The <a href="/wiki/Biological_life_cycle" title="Biological life cycle">life cycle</a> is completed when they <a href="/wiki/Metamorphosis" title="Metamorphosis">metamorphose</a> into adults. A few species deposit eggs on land or bypass the tadpole stage. Adult frogs generally have a carnivorous diet consisting of small <a href="/wiki/Invertebrates" class="mw-redirect" title="Invertebrates">invertebrates</a>, but omnivorous species exist and a few feed on plant matter. Frog skin has a rich <a href="/wiki/Skin_flora" title="Skin flora">microbiome</a> which is important to their health. Frogs are extremely efficient at converting what they eat into body mass. They are an important food source for <a href="/wiki/Predation" title="Predation">predators</a> and part of the <a href="/wiki/Food_web" title="Food web">food web</a> dynamics of many of the world's <a href="/wiki/Ecosystem" title="Ecosystem">ecosystems</a>. The skin is <a href="/wiki/Semi-permeable" class="mw-redirect" title="Semi-permeable">semi-permeable</a>, making them susceptible to dehydration, so they either live in moist places or have special adaptations to deal with dry habitats. Frogs produce a wide range of <a href="/wiki/Animal_communication" title="Animal communication">vocalisations</a>, particularly in their <a href="/wiki/Breeding_season" class="mw-redirect" title="Breeding season">breeding season</a>, and exhibit many different kinds of complex <a href="/wiki/Ethology" title="Ethology">behaviors</a> to attract mates, to fend off predators and to generally survive. </p><p>Frogs are valued as food by humans and also have many cultural roles in literature, symbolism and religion. They are also seen as <a href="/wiki/Bioindicator" title="Bioindicator">environmental bellwethers</a>, with declines in frog populations often viewed as early warning signs of environmental damage. Frog populations have declined significantly since the 1950s. More than one third of species are considered to be <a href="/wiki/Threatened_species" title="Threatened species">threatened</a> with <a href="/wiki/Extinct" class="mw-redirect" title="Extinct">extinction</a> and over 120 are believed to have become extinct since the 1980s. The number of malformations among frogs is on the rise and an emerging <a href="/wiki/Fungus" title="Fungus">fungal</a> disease, <a href="/wiki/Chytridiomycosis" title="Chytridiomycosis">chytridiomycosis</a>, has spread around the world. <a href="/wiki/Conservation_biology" title="Conservation biology">Conservation biologists</a> are working to understand the causes of these problems and to resolve them. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Etymology_and_taxonomy">Etymology and taxonomy</h2></div> <p>The use of the common names <i>frog</i> and <i><a href="/wiki/Toad" title="Toad">toad</a></i> has no taxonomic justification. From a classification perspective, all members of the order Anura are frogs, but only members of the family <a href="/wiki/Bufonidae" class="mw-redirect" title="Bufonidae">Bufonidae</a> are considered "true toads". The use of the term <i>frog</i> in common names usually refers to species that are aquatic or semi-aquatic and have smooth, moist skins; the term <i>toad</i> generally refers to species that are terrestrial with dry, warty skins.<sup id="cite_ref-Salientia_2-0" class="reference"><a href="#cite_note-Salientia-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> There are numerous exceptions to this rule. The <a href="/wiki/European_fire-bellied_toad" title="European fire-bellied toad">European fire-bellied toad</a> (<i>Bombina bombina</i>) has a slightly warty skin and prefers a watery habitat<sup id="cite_ref-Bb_4-0" class="reference"><a href="#cite_note-Bb-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> whereas the <a href="/wiki/Panamanian_golden_frog" title="Panamanian golden frog">Panamanian golden frog</a> (<i>Atelopus zeteki</i>) is in the toad family Bufonidae and has a smooth skin.<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">&#91;</span>5<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Etymology">Etymology</h3></div> <p>The origin of the order name <i>Anura</i>—and its original spelling <i>Anoures</i>—is the <a href="/wiki/Ancient_Greek" title="Ancient Greek">Ancient Greek</a> <a href="/wiki/Alpha_privative" title="Alpha privative">alpha privative</a> prefix <span lang="grc"><a href="https://en.wiktionary.org/wiki/%E1%BC%80%CE%BD-#Ancient_Greek" class="extiw" title="wikt:ἀν-">ἀν-</a></span> (<span title="Ancient Greek transliteration" lang="grc-Latn"><i>an-</i></span> from <span lang="grc"><a href="https://en.wiktionary.org/wiki/%E1%BC%80-#Ancient_Greek" class="extiw" title="wikt:ἀ-">ἀ-</a></span> before a vowel) 'without',<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup> and <span lang="grc"><a href="https://en.wiktionary.org/wiki/%CE%BF%E1%BD%90%CF%81%CE%AC#Ancient_Greek" class="extiw" title="wikt:οὐρά">οὐρά</a></span> (<span title="Ancient Greek transliteration" lang="grc-Latn"><i>ourá</i></span>) 'animal tail'.<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">&#91;</span>7<span class="cite-bracket">&#93;</span></a></sup> meaning "tailless". It refers to the tailless character of these amphibians.<sup id="cite_ref-BaillyBook_8-0" class="reference"><a href="#cite_note-BaillyBook-8"><span class="cite-bracket">&#91;</span>8<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-BaillyWeb_9-0" class="reference"><a href="#cite_note-BaillyWeb-9"><span class="cite-bracket">&#91;</span>9<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">&#91;</span>10<span class="cite-bracket">&#93;</span></a></sup> </p><p>The origins of the word <i>frog</i> are uncertain and debated.<sup id="cite_ref-:0_11-0" class="reference"><a href="#cite_note-:0-11"><span class="cite-bracket">&#91;</span>11<span class="cite-bracket">&#93;</span></a></sup> The word is first attested in <a href="/wiki/Old_English" title="Old English">Old English</a> as <span title="Old English (ca. 450-1100)-language text"><i lang="ang">frogga</i></span>, but the usual Old English word for the frog was <span title="Old English (ca. 450-1100)-language text"><i lang="ang">frosc</i></span> (with variants such as <span title="Old English (ca. 450-1100)-language text"><i lang="ang">frox</i></span> and <span title="Old English (ca. 450-1100)-language text"><i lang="ang">forsc</i></span>), and it is agreed that the word <i>frog</i> is somehow related to this. Old English <span title="Old English (ca. 450-1100)-language text"><i lang="ang">frosc</i></span> remained in dialectal use in English as <i>frosh</i> and <i>frosk</i> into the nineteenth century,<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">&#91;</span>12<span class="cite-bracket">&#93;</span></a></sup> and is paralleled widely in other <a href="/wiki/Germanic_languages" title="Germanic languages">Germanic languages</a>, with examples in the modern languages including <a href="/wiki/German_language" title="German language">German</a> <span title="German-language text"><i lang="de">Frosch</i></span>, <a href="/wiki/Norwegian_language" title="Norwegian language">Norwegian</a> <span title="Norwegian-language text"><i lang="no">frosk</i></span>, <a href="/wiki/Icelandic_language" title="Icelandic language">Icelandic</a> <span title="Icelandic-language text"><i lang="is">froskur</i></span>, and <a href="/wiki/Dutch_language" title="Dutch language">Dutch</a> <i>(kik)vors</i>.<sup id="cite_ref-:0_11-1" class="reference"><a href="#cite_note-:0-11"><span class="cite-bracket">&#91;</span>11<span class="cite-bracket">&#93;</span></a></sup> These words allow reconstruction of a <a href="/wiki/Proto-Germanic_language" title="Proto-Germanic language">Common Germanic</a> ancestor <span title="Proto-Germanic-language text">&#42;<i lang="gem">froskaz</i></span>.<sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">&#91;</span>13<span class="cite-bracket">&#93;</span></a></sup> The third edition of the <i><a href="/wiki/Oxford_English_Dictionary" title="Oxford English Dictionary">Oxford English Dictionary</a></i> finds that the etymology of <span title="Proto-Germanic-language text">&#42;<i lang="gem">froskaz</i></span> is uncertain, but agrees with arguments that it could plausibly derive from a <a href="/wiki/Proto-Indo-European_language" title="Proto-Indo-European language">Proto-Indo-European</a> base along the lines of <span title="Proto-Indo-European-language text">&#42;<i lang="ine">preu</i></span>, meaning 'jump'.<sup id="cite_ref-:0_11-2" class="reference"><a href="#cite_note-:0-11"><span class="cite-bracket">&#91;</span>11<span class="cite-bracket">&#93;</span></a></sup> </p><p>How Old English <span title="Old English (ca. 450-1100)-language text"><i lang="ang">frosc</i></span> gave rise to <span title="Old English (ca. 450-1100)-language text"><i lang="ang">frogga</i></span> is, however, uncertain, as the development does not involve a regular <a href="/wiki/Sound_change" title="Sound change">sound-change</a>. Instead, it seems that there was a trend in Old English to coin nicknames for animals ending in -<i>g</i>, with examples—themselves all of uncertain etymology—including <i>dog</i>, <i>hog</i>, <i>pig, stag</i>, and <i>(ear)wig</i>. <i>Frog</i> appears to have been adapted from <span title="Old English (ca. 450-1100)-language text"><i lang="ang">frosc</i></span> as part of this trend.<sup id="cite_ref-:0_11-3" class="reference"><a href="#cite_note-:0-11"><span class="cite-bracket">&#91;</span>11<span class="cite-bracket">&#93;</span></a></sup> </p><p>Meanwhile, the word <i>toad</i>, first attested as Old English <span title="Old English (ca. 450-1100)-language text"><i lang="ang">tādige</i></span>, is unique to English and is likewise of uncertain etymology.<sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">&#91;</span>14<span class="cite-bracket">&#93;</span></a></sup> It is the basis for the word <i>tadpole</i>, first attested as <a href="/wiki/Middle_English" title="Middle English">Middle English</a> <span title="Middle English (1100-1500)-language text"><i lang="enm">taddepol</i></span>, apparently meaning 'toad-head'.<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">&#91;</span>15<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Taxonomy">Taxonomy</h3></div> <p>About 88% of <a href="/wiki/Amphibian" title="Amphibian">amphibian</a> species are <a href="/wiki/Biological_classification" class="mw-redirect" title="Biological classification">classified</a> in the <a href="/wiki/Order_(biology)" title="Order (biology)">order</a> Anura.<sup id="cite_ref-Pou92_16-0" class="reference"><a href="#cite_note-Pou92-16"><span class="cite-bracket">&#91;</span>16<span class="cite-bracket">&#93;</span></a></sup> These include over 7,700 species<sup id="cite_ref-AOTW_1-1" class="reference"><a href="#cite_note-AOTW-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> in 59 <a href="/wiki/Family_(biology)" title="Family (biology)">families</a>, of which the <a href="/wiki/Hylidae" title="Hylidae">Hylidae</a> (1062 spp.), <a href="/wiki/Strabomantidae" title="Strabomantidae">Strabomantidae</a> (807 spp.), <a href="/wiki/Microhylidae" title="Microhylidae">Microhylidae</a> (758 spp.), and <a href="/wiki/True_toad" title="True toad">Bufonidae</a> (657 spp.) are the <a href="/wiki/Species_richness" title="Species richness">richest in species</a>.<sup id="cite_ref-amdb_17-0" class="reference"><a href="#cite_note-amdb-17"><span class="cite-bracket">&#91;</span>17<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Bombina_bombina_1_(Marek_Szczepanek)_tight_crop.jpg" class="mw-file-description"><img alt="Dark-coloured toad facing left" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4f/Bombina_bombina_1_%28Marek_Szczepanek%29_tight_crop.jpg/220px-Bombina_bombina_1_%28Marek_Szczepanek%29_tight_crop.jpg" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4f/Bombina_bombina_1_%28Marek_Szczepanek%29_tight_crop.jpg/330px-Bombina_bombina_1_%28Marek_Szczepanek%29_tight_crop.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4f/Bombina_bombina_1_%28Marek_Szczepanek%29_tight_crop.jpg/440px-Bombina_bombina_1_%28Marek_Szczepanek%29_tight_crop.jpg 2x" data-file-width="1905" data-file-height="1430" /></a><figcaption><a href="/wiki/European_fire-bellied_toad" title="European fire-bellied toad">European fire-bellied toad</a> (<i>Bombina&#160;bombina</i>)</figcaption></figure> <p>The Anura include all modern frogs and any <a href="/wiki/Fossil" title="Fossil">fossil</a> species that fit within the anuran definition. The characteristics of anuran adults include: 9 or fewer presacral vertebrae, the presence of a urostyle formed of fused vertebrae, no tail, a long and forward-sloping ilium, shorter fore limbs than hind limbs, <a href="/wiki/Radius_(bone)" title="Radius (bone)">radius</a> and <a href="/wiki/Ulna" title="Ulna">ulna</a> fused, <a href="/wiki/Tibia" title="Tibia">tibia</a> and <a href="/wiki/Fibula" title="Fibula">fibula</a> fused, elongated <a href="/wiki/Ankle_bone" class="mw-redirect" title="Ankle bone">ankle bones</a>, absence of a prefrontal bone, presence of a <a href="/wiki/Hyoid_bone" title="Hyoid bone">hyoid plate</a>, a <a href="/wiki/Mandible" title="Mandible">lower jaw</a> without teeth (with the exception of <i><a href="/wiki/Gastrotheca_guentheri" title="Gastrotheca guentheri">Gastrotheca guentheri</a></i>) consisting of three pairs of bones (angulosplenial, dentary, and mentomeckelian, with the last pair being absent in <a href="/wiki/Pipoidea" title="Pipoidea">Pipoidea</a>),<sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">&#91;</span>18<span class="cite-bracket">&#93;</span></a></sup> an unsupported tongue, <a href="/wiki/Lymph" title="Lymph">lymph</a> spaces underneath the skin, and a muscle, the protractor lentis, attached to the <a href="/wiki/Lens_(anatomy)" class="mw-redirect" title="Lens (anatomy)">lens of the eye</a>.<sup id="cite_ref-ToLAnura_19-0" class="reference"><a href="#cite_note-ToLAnura-19"><span class="cite-bracket">&#91;</span>19<span class="cite-bracket">&#93;</span></a></sup> The anuran larva or tadpole has a single central respiratory <a href="/wiki/Spiracle_(vertebrates)" title="Spiracle (vertebrates)">spiracle</a> and mouthparts consisting of <a href="/wiki/Keratin" title="Keratin">keratinous</a> beaks and <a href="/wiki/Denticle_(tooth_feature)" title="Denticle (tooth feature)">denticles</a>.<sup id="cite_ref-ToLAnura_19-1" class="reference"><a href="#cite_note-ToLAnura-19"><span class="cite-bracket">&#91;</span>19<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Atelopus_zeteki1.jpg" class="mw-file-description"><img alt="Panamanian golden frog" src="//upload.wikimedia.org/wikipedia/commons/thumb/5/55/Atelopus_zeteki1.jpg/250px-Atelopus_zeteki1.jpg" decoding="async" width="220" height="139" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/55/Atelopus_zeteki1.jpg/330px-Atelopus_zeteki1.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/55/Atelopus_zeteki1.jpg/500px-Atelopus_zeteki1.jpg 2x" data-file-width="2720" data-file-height="1720" /></a><figcaption><a href="/wiki/Panamanian_golden_frog" title="Panamanian golden frog">Panamanian golden frog</a> (<i>Atelopus&#160;zeteki</i>)</figcaption></figure> <p>Frogs and toads are broadly classified into three suborders: <a href="/wiki/Archaeobatrachia" title="Archaeobatrachia">Archaeobatrachia</a>, which includes four families of primitive frogs; <a href="/wiki/Mesobatrachia" title="Mesobatrachia">Mesobatrachia</a>, which includes five families of more evolutionary intermediate frogs; and <a href="/wiki/Neobatrachia" title="Neobatrachia">Neobatrachia</a>, by far the largest group, which contains the remaining families of modern frogs, including most common species throughout the world. The suborder Neobatrachia is further divided into the two superfamilies <a href="/wiki/Hyloidea" title="Hyloidea">Hyloidea</a> and <a href="/wiki/Ranoidea" title="Ranoidea">Ranoidea</a>.<sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">&#91;</span>20<span class="cite-bracket">&#93;</span></a></sup> This classification is based on such <a href="/wiki/Morphology_(biology)" title="Morphology (biology)">morphological</a> features as the number of vertebrae, the structure of the <a href="/wiki/Pectoral_girdle" class="mw-redirect" title="Pectoral girdle">pectoral girdle</a>, and the morphology of tadpoles. While this classification is largely accepted, relationships among families of frogs are still debated.<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">&#91;</span>21<span class="cite-bracket">&#93;</span></a></sup> </p><p>Some species of anurans <a href="/wiki/Hybrid_(biology)" title="Hybrid (biology)">hybridise</a> readily. For instance, the <a href="/wiki/Edible_frog" title="Edible frog">edible frog</a> (<i>Pelophylax esculentus</i>) is a hybrid between the <a href="/wiki/Pool_frog" title="Pool frog">pool frog</a> (<i>P. lessonae</i>) and the <a href="/wiki/Marsh_frog" title="Marsh frog">marsh frog</a> (<i>P. ridibundus</i>).<sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">&#91;</span>22<span class="cite-bracket">&#93;</span></a></sup> The fire-bellied toads <i>Bombina bombina</i> and <i><a href="/wiki/Yellow-bellied_toad" title="Yellow-bellied toad">B. variegata</a></i> are similar in forming hybrids. These are less fertile than their parents, giving rise to a <a href="/wiki/Hybrid_zone" title="Hybrid zone">hybrid zone</a> where the hybrids are prevalent.<sup id="cite_ref-23" class="reference"><a href="#cite_note-23"><span class="cite-bracket">&#91;</span>23<span class="cite-bracket">&#93;</span></a></sup> </p> <div style="clear:left;" class=""></div> <div class="mw-heading mw-heading2"><h2 id="Evolution">Evolution</h2></div> <p>The origins and evolutionary relationships between the three main groups of amphibians are hotly debated. A <a href="/wiki/Molecular_phylogeny" class="mw-redirect" title="Molecular phylogeny">molecular phylogeny</a> based on <a href="/wiki/Ribosomal_DNA" title="Ribosomal DNA">rDNA</a> analysis dating from 2005 suggests that <a href="/wiki/Salamanders" class="mw-redirect" title="Salamanders">salamanders</a> and <a href="/wiki/Caecilian" title="Caecilian">caecilians</a> are more closely related to each other than they are to frogs and the <a href="/wiki/Genetic_divergence" title="Genetic divergence">divergence</a> of the three groups took place in the <a href="/wiki/Paleozoic" title="Paleozoic">Paleozoic</a> or early <a href="/wiki/Mesozoic" title="Mesozoic">Mesozoic</a> before the break-up of the supercontinent <a href="/wiki/Pangaea" title="Pangaea">Pangaea</a> and soon after their divergence from the <a href="/wiki/Sarcopterygii" title="Sarcopterygii">lobe-finned fishes</a>. This would help account for the relative scarcity of amphibian fossils from the period before the groups split.<sup id="cite_ref-24" class="reference"><a href="#cite_note-24"><span class="cite-bracket">&#91;</span>24<span class="cite-bracket">&#93;</span></a></sup> Another molecular phylogenetic analysis conducted about the same time concluded that <a href="/wiki/Lissamphibian" class="mw-redirect" title="Lissamphibian">lissamphibians</a> first appeared about 330 million years ago and that the <a href="/wiki/Temnospondyli" title="Temnospondyli">temnospondyl-origin</a> hypothesis is more credible than other theories. The <a href="/wiki/Neobatrachia" title="Neobatrachia">neobatrachians</a> seemed to have originated in Africa/India, the salamanders in East Asia and the caecilians in tropical Pangaea.<sup id="cite_ref-25" class="reference"><a href="#cite_note-25"><span class="cite-bracket">&#91;</span>25<span class="cite-bracket">&#93;</span></a></sup> Other researchers, while agreeing with the main thrust of this study, questioned the choice of calibration points used to synchronise the data. They proposed that the date of lissamphibian diversification should be placed in the <a href="/wiki/Permian" title="Permian">Permian</a>, rather less than 300 million years ago, a date in better agreement with the palaeontological data.<sup id="cite_ref-26" class="reference"><a href="#cite_note-26"><span class="cite-bracket">&#91;</span>26<span class="cite-bracket">&#93;</span></a></sup> A further study in 2011 using both extinct and living taxa sampled for morphological, as well as molecular data, came to the conclusion that Lissamphibia is <a href="/wiki/Monophyly" title="Monophyly">monophyletic</a> and that it should be nested within <a href="/wiki/Lepospondyli" title="Lepospondyli">Lepospondyli</a> rather than within <a href="/wiki/Temnospondyli" title="Temnospondyli">Temnospondyli</a>. The study postulated that Lissamphibia originated no earlier than the late <a href="/wiki/Carboniferous" title="Carboniferous">Carboniferous</a>, some 290 to 305 million years ago. The split between Anura and <a href="/wiki/Caudata" title="Caudata">Caudata</a> was estimated as taking place 292 million years ago, rather later than most molecular studies suggest, with the caecilians splitting off 239 million years ago.<sup id="cite_ref-27" class="reference"><a href="#cite_note-27"><span class="cite-bracket">&#91;</span>27<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Fossilised_frog.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/1/17/Fossilised_frog.jpg/250px-Fossilised_frog.jpg" decoding="async" width="220" height="189" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/17/Fossilised_frog.jpg/330px-Fossilised_frog.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/17/Fossilised_frog.jpg/500px-Fossilised_frog.jpg 2x" data-file-width="2016" data-file-height="1732" /></a><figcaption>A fossilised frog from the <a href="/wiki/Czech_Republic" title="Czech Republic">Czech Republic</a>, possibly <i><a href="/wiki/Palaeobatrachus_gigas" class="mw-redirect" title="Palaeobatrachus gigas">Palaeobatrachus gigas</a></i></figcaption></figure> <p>In 2008, <i><a href="/wiki/Gerobatrachus_hottoni" class="mw-redirect" title="Gerobatrachus hottoni">Gerobatrachus hottoni</a></i>, a <a href="/wiki/Temnospondyli" title="Temnospondyli">temnospondyl</a> with many frog- and salamander-like characteristics, was discovered in <a href="/wiki/Texas" title="Texas">Texas</a>. It dated back 290 million years and was hailed as a <a href="/wiki/Transitional_fossil" title="Transitional fossil">missing link</a>, a <a href="/wiki/Stem_group" class="mw-redirect" title="Stem group">stem</a> batrachian close to the <a href="/wiki/Common_ancestor" class="mw-redirect" title="Common ancestor">common ancestor</a> of frogs and salamanders, consistent with the widely accepted hypothesis that frogs and salamanders are more closely related to each other (forming a <a href="/wiki/Clade" title="Clade">clade</a> called Batrachia) than they are to caecilians.<sup id="cite_ref-28" class="reference"><a href="#cite_note-28"><span class="cite-bracket">&#91;</span>28<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-29" class="reference"><a href="#cite_note-29"><span class="cite-bracket">&#91;</span>29<span class="cite-bracket">&#93;</span></a></sup> However, others have suggested that <i>Gerobatrachus hottoni</i> was only a <a href="/wiki/Dissorophoidea" title="Dissorophoidea">dissorophoid</a> temnospondyl unrelated to extant amphibians.<sup id="cite_ref-M&amp;L09_30-0" class="reference"><a href="#cite_note-M&amp;L09-30"><span class="cite-bracket">&#91;</span>30<span class="cite-bracket">&#93;</span></a></sup> </p><p><a href="/wiki/Salientia" title="Salientia">Salientia</a> (Latin <i>salire</i> (<i>salio</i>), "to jump") is the name of the total group that includes modern frogs in the order Anura as well as their close fossil relatives, the "proto-frogs" or "stem-frogs". The common features possessed by these proto-frogs include 14 <a href="/wiki/Vertebral_column" class="mw-redirect" title="Vertebral column">presacral vertebrae</a> (modern frogs have eight or 9), a long and forward-sloping <a href="/wiki/Ilium_(bone)" title="Ilium (bone)">ilium</a> in the <a href="/wiki/Pelvis" title="Pelvis">pelvis</a>, the presence of a <a href="/wiki/Parietal_bone" title="Parietal bone">frontoparietal bone</a>, and a <a href="/wiki/Mandible" title="Mandible">lower jaw</a> without teeth. The earliest known amphibians that were more closely related to frogs than to salamanders are <i><a href="/wiki/Triadobatrachus_massinoti" class="mw-redirect" title="Triadobatrachus massinoti">Triadobatrachus massinoti</a></i>, from the early <a href="/wiki/Triassic" title="Triassic">Triassic</a> period of <a href="/wiki/Madagascar" title="Madagascar">Madagascar</a> (about 250 million years ago), and <i><a href="/wiki/Czatkobatrachus_polonicus" class="mw-redirect" title="Czatkobatrachus polonicus">Czatkobatrachus polonicus</a></i>, from the Early Triassic of <a href="/wiki/Poland" title="Poland">Poland</a> (about the same age as <i>Triadobatrachus</i>).<sup id="cite_ref-TOLweb_31-0" class="reference"><a href="#cite_note-TOLweb-31"><span class="cite-bracket">&#91;</span>31<span class="cite-bracket">&#93;</span></a></sup> The skull of <i>Triadobatrachus</i> is frog-like, being broad with large eye sockets, but the fossil has features diverging from modern frogs. These include a longer body with more <a href="/wiki/Vertebrae" class="mw-redirect" title="Vertebrae">vertebrae</a>. The tail has separate vertebrae unlike the fused urostyle or coccyx in modern frogs. The tibia and fibula bones are also separate, making it probable that <i>Triadobatrachus</i> was not an efficient leaper.<sup id="cite_ref-TOLweb_31-1" class="reference"><a href="#cite_note-TOLweb-31"><span class="cite-bracket">&#91;</span>31<span class="cite-bracket">&#93;</span></a></sup> A 2019 study has noted the presence of Salientia from the <a href="/wiki/Chinle_Formation" title="Chinle Formation">Chinle Formation</a>, and suggested that anurans might have first appeared during the <a href="/wiki/Late_Triassic" title="Late Triassic">Late Triassic</a>.<sup id="cite_ref-32" class="reference"><a href="#cite_note-32"><span class="cite-bracket">&#91;</span>32<span class="cite-bracket">&#93;</span></a></sup> </p><p>On the basis of fossil evidence, the earliest known "true frogs" that fall into the anuran lineage proper all lived in the early <a href="/wiki/Jurassic" title="Jurassic">Jurassic</a> period.<sup id="cite_ref-Salientia_2-1" class="reference"><a href="#cite_note-Salientia-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Heatwole00_33-0" class="reference"><a href="#cite_note-Heatwole00-33"><span class="cite-bracket">&#91;</span>33<span class="cite-bracket">&#93;</span></a></sup> One such early frog species, <i><a href="/wiki/Prosalirus_bitis" class="mw-redirect" title="Prosalirus bitis">Prosalirus bitis</a></i>, was discovered in <a href="/wiki/1995_in_paleontology" title="1995 in paleontology">1995</a> in the <a href="/wiki/Kayenta_Formation" title="Kayenta Formation">Kayenta Formation</a> of <a href="/wiki/Arizona" title="Arizona">Arizona</a> and dates back to the <a href="/wiki/Early_Jurassic" title="Early Jurassic">Early Jurassic</a> epoch (199.6 to 175 million years ago), making <i>Prosalirus</i> somewhat more recent than <i>Triadobatrachus</i>.<sup id="cite_ref-jurassicdistribution_34-0" class="reference"><a href="#cite_note-jurassicdistribution-34"><span class="cite-bracket">&#91;</span>34<span class="cite-bracket">&#93;</span></a></sup> Like the latter, <i>Prosalirus</i> did not have greatly enlarged legs, but had the typical three-pronged <a href="/wiki/Pelvic" class="mw-redirect" title="Pelvic">pelvic</a> structure of modern frogs. Unlike <i>Triadobatrachus</i>, <i>Prosalirus</i> had already lost nearly all of its tail<sup id="cite_ref-35" class="reference"><a href="#cite_note-35"><span class="cite-bracket">&#91;</span>35<span class="cite-bracket">&#93;</span></a></sup> and was well adapted for jumping.<sup id="cite_ref-foster-anura_36-0" class="reference"><a href="#cite_note-foster-anura-36"><span class="cite-bracket">&#91;</span>36<span class="cite-bracket">&#93;</span></a></sup> Another Early Jurassic frog is <i><a href="/wiki/Vieraella_herbsti" class="mw-redirect" title="Vieraella herbsti">Vieraella herbsti</a></i>, which is known only from <a href="/wiki/Dorsum_(biology)" class="mw-redirect" title="Dorsum (biology)">dorsal</a> and <a href="/wiki/Ventral" class="mw-redirect" title="Ventral">ventral</a> impressions of a single animal and was estimated to be <style data-mw-deduplicate="TemplateStyles:r1154941027">.mw-parser-output .frac{white-space:nowrap}.mw-parser-output .frac .num,.mw-parser-output .frac .den{font-size:80%;line-height:0;vertical-align:super}.mw-parser-output .frac .den{vertical-align:sub}.mw-parser-output .sr-only{border:0;clip:rect(0,0,0,0);clip-path:polygon(0px 0px,0px 0px,0px 0px);height:1px;margin:-1px;overflow:hidden;padding:0;position:absolute;width:1px}</style>33&#160;mm (<span class="frac">1<span class="sr-only">+</span><span class="num">1</span>&#8260;<span class="den">4</span></span>&#160;in) from snout to vent. <i><a href="/wiki/Notobatrachus_degiustoi" class="mw-redirect" title="Notobatrachus degiustoi">Notobatrachus degiustoi</a></i> from the <a href="/wiki/Middle_Jurassic" title="Middle Jurassic">middle Jurassic</a> is slightly younger, about 155–170 million years old. The main evolutionary changes in this species involved the shortening of the body and the loss of the tail. Tadpoles of <i>N. degiustoi</i> constitute the oldest tadpoles found as of 2024, dating back to 168–161 million years ago. These tadpoles also showed adaptations for <a href="/wiki/Filter-feeding" class="mw-redirect" title="Filter-feeding">filter-feeding</a>, implying residence in temporary pools by filter-feeding larvae was already commonplace.<sup id="cite_ref-37" class="reference"><a href="#cite_note-37"><span class="cite-bracket">&#91;</span>37<span class="cite-bracket">&#93;</span></a></sup> The evolution of modern Anura likely was complete by the Jurassic period. Since then, evolutionary changes in <a href="/wiki/Ploidy" title="Ploidy">chromosome numbers</a> have taken place about 20 times faster in mammals than in frogs, which means <a href="/wiki/Speciation" title="Speciation">speciation</a> is occurring more rapidly in mammals.<sup id="cite_ref-38" class="reference"><a href="#cite_note-38"><span class="cite-bracket">&#91;</span>38<span class="cite-bracket">&#93;</span></a></sup> </p><p>According to genetic studies, the families <a href="/wiki/Hyloidea" title="Hyloidea">Hyloidea</a>, <a href="/wiki/Microhylidae" title="Microhylidae">Microhylidae</a>, and the clade <a href="/w/index.php?title=Natatanura&amp;action=edit&amp;redlink=1" class="new" title="Natatanura (page does not exist)">Natatanura</a> (comprising about 88% of living frogs) diversified simultaneously some 66 million years ago, soon after the <a href="/wiki/Cretaceous%E2%80%93Paleogene_extinction_event" title="Cretaceous–Paleogene extinction event">Cretaceous–Paleogene extinction event</a> associated with the <a href="/wiki/Chicxulub_impactor" class="mw-redirect" title="Chicxulub impactor">Chicxulub impactor</a>. All origins of arboreality (e.g. in Hyloidea and Natatanura) follow from that time and the resurgence of forest that occurred afterwards.<sup id="cite_ref-39" class="reference"><a href="#cite_note-39"><span class="cite-bracket">&#91;</span>39<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-40" class="reference"><a href="#cite_note-40"><span class="cite-bracket">&#91;</span>40<span class="cite-bracket">&#93;</span></a></sup> </p><p>Frog fossils have been found on all of the Earth's continents.<sup id="cite_ref-41" class="reference"><a href="#cite_note-41"><span class="cite-bracket">&#91;</span>41<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-42" class="reference"><a href="#cite_note-42"><span class="cite-bracket">&#91;</span>42<span class="cite-bracket">&#93;</span></a></sup> In 2020, it was announced that 40 million year old <a href="/wiki/Calyptocephalellidae" title="Calyptocephalellidae">helmeted frog</a> fossils had been discovered by a team of vertebrate palaeontologists in <a href="/wiki/Seymour_Island" title="Seymour Island">Seymour Island</a> on the <a href="/wiki/Antarctic_Peninsula" title="Antarctic Peninsula">Antarctic Peninsula</a>, indicating that this region was once home to frogs related to those now living in South American <a href="/wiki/Valdivian_temperate_rain_forest" class="mw-redirect" title="Valdivian temperate rain forest"><i>Nothofagus</i> forest</a>.<sup id="cite_ref-43" class="reference"><a href="#cite_note-43"><span class="cite-bracket">&#91;</span>43<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Phylogeny">Phylogeny</h3></div> <p>A <a href="/wiki/Cladogram" title="Cladogram">cladogram</a> showing the relationships of the different <a href="/wiki/Family_(biology)" title="Family (biology)">families</a> of frogs in the clade Anura can be seen in the table below. This diagram, in the form of a <a href="/wiki/Phylogenetic_tree" title="Phylogenetic tree">tree</a>, shows how each frog family is related to other families, with each node representing a point of common ancestry. It is based on Frost <i>et al.</i> (2006),<sup id="cite_ref-44" class="reference"><a href="#cite_note-44"><span class="cite-bracket">&#91;</span>44<span class="cite-bracket">&#93;</span></a></sup> Heinicke <i>et al.</i> (2009)<sup id="cite_ref-45" class="reference"><a href="#cite_note-45"><span class="cite-bracket">&#91;</span>45<span class="cite-bracket">&#93;</span></a></sup> and Pyron and Wiens (2011).<sup id="cite_ref-46" class="reference"><a href="#cite_note-46"><span class="cite-bracket">&#91;</span>46<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="clade"><style data-mw-deduplicate="TemplateStyles:r1261294616">body.skin-vector-2022 .mw-parser-output div.clade,body.skin-minerva .mw-parser-output div.clade{overflow-x:auto;overflow-y:hidden}body.skin-minerva .mw-parser-output div.clade p{font-size:inherit}.mw-parser-output table.clade{border-spacing:0;margin:0;font-size:100%;line-height:100%;border-collapse:separate;width:auto;display:table}.mw-parser-output table.clade table.clade{width:100%;line-height:inherit}.mw-parser-output table.clade td.clade-label{min-width:0.2em;width:0.2em;padding:0.1em 0.25em;vertical-align:bottom;text-align:center;border-left:1px solid;border-bottom:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-label::before,.mw-parser-output table.clade td.clade-slabel::before{content:"\2060 "}.mw-parser-output table.clade td.clade-fixed-width{overflow:hidden;text-overflow:ellipsis}.mw-parser-output table.clade td.clade-fixed-width:hover{overflow:visible}.mw-parser-output table.clade td.clade-label.first{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-label.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-slabel{padding:0.1em 0.25em;vertical-align:top;text-align:center;border-left:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-slabel:hover{overflow:visible}.mw-parser-output table.clade td.clade-slabel.last{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-slabel.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-bar{vertical-align:middle;text-align:left;padding:0 0.5em;position:relative}.mw-parser-output table.clade td.clade-bar.reverse{text-align:right;position:relative}.mw-parser-output table.clade td.clade-leaf{border:0;padding:0;text-align:left}.mw-parser-output table.clade td.clade-leaf p{padding-right:5px;padding-left:2px}.mw-parser-output table.clade td.clade-leafR{border:0;padding:0;text-align:right}.mw-parser-output table.clade td.clade-leafR p{padding-left:5px;padding-right:2px}.mw-parser-output table.clade td.clade-leaf.reverse{text-align:right}.mw-parser-output table.clade td.clade-leaf.reverse p{padding-left:5px;padding-right:2px}.mw-parser-output table.clade:hover span.linkA{background-color:yellow}.mw-parser-output table.clade:hover span.linkB{background-color:green}</style> <table class="clade" style="font-size:80%;line-height:100%"> <tbody><tr> <td class="clade-label first"><b>Anura</b> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Leiopelmatidae" class="mw-redirect" title="Leiopelmatidae">Leiopelmatidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Ascaphidae" class="mw-redirect" title="Ascaphidae">Ascaphidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/w/index.php?title=Bombianura&amp;action=edit&amp;redlink=1" class="new" title="Bombianura (page does not exist)">Bombianura</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/w/index.php?title=Costata&amp;action=edit&amp;redlink=1" class="new" title="Costata (page does not exist)">Costata</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Bombinatoridae" title="Bombinatoridae">Bombinatoridae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Alytidae" title="Alytidae">Alytidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Discoglossidae" class="mw-redirect" title="Discoglossidae">Discoglossidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/w/index.php?title=Pipanura&amp;action=edit&amp;redlink=1" class="new" title="Pipanura (page does not exist)">Pipanura</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Xenoanura" class="mw-redirect" title="Xenoanura">Xenoanura</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Pipidae" title="Pipidae">Pipidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Rhinophrynidae" title="Rhinophrynidae">Rhinophrynidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/w/index.php?title=Acosmanura&amp;action=edit&amp;redlink=1" class="new" title="Acosmanura (page does not exist)">Acosmanura</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/w/index.php?title=Anomocoela&amp;action=edit&amp;redlink=1" class="new" title="Anomocoela (page does not exist)">Anomocoela</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Scaphiopodidae" class="mw-redirect" title="Scaphiopodidae">Scaphiopodidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Pelodytidae" title="Pelodytidae">Pelodytidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Pelobatoidea" title="Pelobatoidea">Pelobatoidea</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Pelobatidae" class="mw-redirect" title="Pelobatidae">Pelobatidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Megophryidae" title="Megophryidae">Megophryidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Neobatrachia" title="Neobatrachia">Neobatrachia</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Heleophrynidae" class="mw-redirect" title="Heleophrynidae">Heleophrynidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/w/index.php?title=Phthanobatrachia&amp;action=edit&amp;redlink=1" class="new" title="Phthanobatrachia (page does not exist)">Phthanobatrachia</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Sooglossoidea" title="Sooglossoidea">Sooglossoidea</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Sooglossidae" title="Sooglossidae">Sooglossidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Nasikabatrachidae" class="mw-redirect" title="Nasikabatrachidae">Nasikabatrachidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/w/index.php?title=Notogaeanura&amp;action=edit&amp;redlink=1" class="new" title="Notogaeanura (page does not exist)">Notogaeanura</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Australobatrachia" title="Australobatrachia">Australobatrachia</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Calyptocephalellidae" title="Calyptocephalellidae">Calyptocephalellidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Myobatrachidae" title="Myobatrachidae">Myobatrachidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Limnodynastidae" title="Limnodynastidae">Limnodynastidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/w/index.php?title=Nobleoanura&amp;action=edit&amp;redlink=1" class="new" title="Nobleoanura (page does not exist)">Nobleoanura</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Brachycephaloidea" title="Brachycephaloidea">Brachycephaloidea</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Ceuthomantidae" class="mw-redirect" title="Ceuthomantidae">Ceuthomantidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Brachycephalidae" title="Brachycephalidae">Brachycephalidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Eleutherodactylidae" title="Eleutherodactylidae">Eleutherodactylidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Craugastoridae" title="Craugastoridae">Craugastoridae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last">(<a href="/wiki/Terrarana" class="mw-redirect" title="Terrarana">Terrarana</a>) </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Hemiphractidae" title="Hemiphractidae">Hemiphractidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/w/index.php?title=Athesphatanura&amp;action=edit&amp;redlink=1" class="new" title="Athesphatanura (page does not exist)">Athesphatanura</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Hylidae" title="Hylidae">Hylidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/w/index.php?title=Leptodactyliformes&amp;action=edit&amp;redlink=1" class="new" title="Leptodactyliformes (page does not exist)">Leptodactyliformes</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/w/index.php?title=Agastorophrynia&amp;action=edit&amp;redlink=1" class="new" title="Agastorophrynia (page does not exist)">Agastorophrynia</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Bufonidae" class="mw-redirect" title="Bufonidae">Bufonidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Aromobatidae" title="Aromobatidae">Aromobatidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Dendrobatidae" class="mw-redirect" title="Dendrobatidae">Dendrobatidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/w/index.php?title=Diphyabatracea&amp;action=edit&amp;redlink=1" class="new" title="Diphyabatracea (page does not exist)">Diphyabatracea</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Leptodactylidae" title="Leptodactylidae">Leptodactylidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Allophrynidae" class="mw-redirect" title="Allophrynidae">Allophrynidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Centrolenidae" class="mw-redirect" title="Centrolenidae">Centrolenidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Ceratophryidae" title="Ceratophryidae">Ceratophryidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Odontophrynidae" title="Odontophrynidae">Odontophrynidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Cycloramphidae" title="Cycloramphidae">Cycloramphidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Alsodidae" title="Alsodidae">Alsodidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Hylodidae" title="Hylodidae">Hylodidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Telmatobiidae" class="mw-redirect" title="Telmatobiidae">Telmatobiidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Batrachylidae" title="Batrachylidae">Batrachylidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Rhinodermatidae" title="Rhinodermatidae">Rhinodermatidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Ranoidea" title="Ranoidea">Ranoides</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/w/index.php?title=Allodapanura&amp;action=edit&amp;redlink=1" class="new" title="Allodapanura (page does not exist)">Allodapanura</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Microhylidae" title="Microhylidae">Microhylidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label">Afrobatrachia </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/w/index.php?title=Xenosyneunitanura&amp;action=edit&amp;redlink=1" class="new" title="Xenosyneunitanura (page does not exist)">Xenosyneunitanura</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Brevicipitidae" title="Brevicipitidae">Brevicipitidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Hemisotidae" class="mw-redirect" title="Hemisotidae">Hemisotidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/w/index.php?title=Laurentobatrachia&amp;action=edit&amp;redlink=1" class="new" title="Laurentobatrachia (page does not exist)">Laurentobatrachia</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Hyperoliidae" title="Hyperoliidae">Hyperoliidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Arthroleptidae" title="Arthroleptidae">Arthroleptidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/w/index.php?title=Natatanura&amp;action=edit&amp;redlink=1" class="new" title="Natatanura (page does not exist)">Natatanura</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Ptychadenidae" title="Ptychadenidae">Ptychadenidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/w/index.php?title=Victoranura&amp;action=edit&amp;redlink=1" class="new" title="Victoranura (page does not exist)">Victoranura</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Micrixalidae" class="mw-redirect" title="Micrixalidae">Micrixalidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Phrynobatrachidae" class="mw-redirect" title="Phrynobatrachidae">Phrynobatrachidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Conrauidae" class="mw-redirect" title="Conrauidae">Conrauidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/w/index.php?title=Pyxicephaloidea&amp;action=edit&amp;redlink=1" class="new" title="Pyxicephaloidea (page does not exist)">Pyxicephaloidea</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Petropedetidae" title="Petropedetidae">Petropedetidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Pyxicephalidae" title="Pyxicephalidae">Pyxicephalidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Nyctibatrachidae" title="Nyctibatrachidae">Nyctibatrachidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Ceratobatrachidae" title="Ceratobatrachidae">Ceratobatrachidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/w/index.php?title=Saukrobatrachia&amp;action=edit&amp;redlink=1" class="new" title="Saukrobatrachia (page does not exist)">Saukrobatrachia</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Ranixalidae" title="Ranixalidae">Ranixalidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Dicroglossidae" title="Dicroglossidae">Dicroglossidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/w/index.php?title=Aglaioanura&amp;action=edit&amp;redlink=1" class="new" title="Aglaioanura (page does not exist)">Aglaioanura</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Rhacophoridae" title="Rhacophoridae">Rhacophoridae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Mantellidae" title="Mantellidae">Mantellidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Ranidae" class="mw-redirect" title="Ranidae">Ranidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> <div class="mw-heading mw-heading2"><h2 id="Morphology_and_physiology">Morphology and physiology</h2></div> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Frog_limbs.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/3a/Frog_limbs.jpg/250px-Frog_limbs.jpg" decoding="async" width="220" height="199" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/3a/Frog_limbs.jpg/330px-Frog_limbs.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/3a/Frog_limbs.jpg/500px-Frog_limbs.jpg 2x" data-file-width="800" data-file-height="722" /></a><figcaption>A <a href="/wiki/American_bullfrog" title="American bullfrog">bullfrog</a> skeleton, showing elongated limb bones and extra joints. Red marks indicate bones which have been substantially elongated in frogs and joints which have become mobile. Blue indicates joints and bones which have not been modified or only somewhat elongated.</figcaption></figure> <p>Frogs have no tail, except as larvae, and most have long hind legs, elongated ankle bones, webbed toes, no claws, large eyes, and a smooth or warty skin. They have short vertebral columns, with no more than 10 free vertebrae and fused tailbones (urostyle or coccyx).<sup id="cite_ref-Flam_47-0" class="reference"><a href="#cite_note-Flam-47"><span class="cite-bracket">&#91;</span>47<span class="cite-bracket">&#93;</span></a></sup> Frogs range in size from <i><a href="/wiki/Paedophryne_amauensis" title="Paedophryne amauensis">Paedophryne amauensis</a></i> of <a href="/wiki/Papua_New_Guinea" title="Papua New Guinea">Papua New Guinea</a> that is 7.7&#160;mm (0.30&#160;in) in <a href="/wiki/Snout%E2%80%93vent_length" title="Snout–vent length">snout–vent length</a><sup id="cite_ref-48" class="reference"><a href="#cite_note-48"><span class="cite-bracket">&#91;</span>48<span class="cite-bracket">&#93;</span></a></sup> to the up to about 35&#160;cm (14&#160;in) and 3.3&#160;kg (7.3&#160;lb) <a href="/wiki/Goliath_frog" title="Goliath frog">goliath frog</a> (<i>Conraua goliath</i>) of central Africa.<sup id="cite_ref-Colgan1982_49-0" class="reference"><a href="#cite_note-Colgan1982-49"><span class="cite-bracket">&#91;</span>49<span class="cite-bracket">&#93;</span></a></sup> There are prehistoric, extinct species that reached even larger sizes.<sup id="cite_ref-50" class="reference"><a href="#cite_note-50"><span class="cite-bracket">&#91;</span>50<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Feet_and_legs">Feet and legs</h3></div> <p>The structure of the feet and legs varies greatly among frog species, depending in part on whether they live primarily on the ground, in water, in trees, or in burrows. Adult anurans have four fingers on the hands and five toes on the feet,<sup id="cite_ref-51" class="reference"><a href="#cite_note-51"><span class="cite-bracket">&#91;</span>51<span class="cite-bracket">&#93;</span></a></sup> but the smallest species often have hands and feet where some of the digits are vestigial.<sup id="cite_ref-52" class="reference"><a href="#cite_note-52"><span class="cite-bracket">&#91;</span>52<span class="cite-bracket">&#93;</span></a></sup> Frogs must be able to move quickly through their environment to catch prey and escape predators, and numerous adaptations help them to do so. Most frogs are either proficient at jumping or are descended from ancestors that were, with much of the <a href="/wiki/Musculoskeletal" class="mw-redirect" title="Musculoskeletal">musculoskeletal</a> <a href="/wiki/Morphology_(biology)" title="Morphology (biology)">morphology</a> modified for this purpose. The tibia, fibula, and <a href="/wiki/Tarsus_(skeleton)" title="Tarsus (skeleton)">tarsals</a> have been fused into a single strong <a href="/wiki/Bone" title="Bone">bone</a>, as have the radius and ulna in the fore limbs (which must absorb the impact on landing). The <a href="/wiki/Metatarsals" class="mw-redirect" title="Metatarsals">metatarsals</a> have become elongated to add to the leg length and allow frogs to push against the ground for a longer period on take-off. The <a href="/wiki/Ilium_(bone)" title="Ilium (bone)">ilium</a> has elongated and formed a mobile joint with the <a href="/wiki/Sacrum" title="Sacrum">sacrum</a> which, in specialist jumpers such as ranids and hylids, functions as an additional limb joint to further power the leaps. The tail vertebrae have fused into a urostyle which is retracted inside the pelvis. This enables the force to be transferred from the legs to the body during a leap.<sup id="cite_ref-Flam_47-1" class="reference"><a href="#cite_note-Flam-47"><span class="cite-bracket">&#91;</span>47<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Rana_temporaria_04_by-dpc.jpg" class="mw-file-description"><img alt="Webbed foot" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/9c/Rana_temporaria_04_by-dpc.jpg/220px-Rana_temporaria_04_by-dpc.jpg" decoding="async" width="220" height="182" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/9c/Rana_temporaria_04_by-dpc.jpg/330px-Rana_temporaria_04_by-dpc.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/9c/Rana_temporaria_04_by-dpc.jpg/440px-Rana_temporaria_04_by-dpc.jpg 2x" data-file-width="1926" data-file-height="1589" /></a><figcaption>Webbed hind foot of <a href="/wiki/Common_frog" title="Common frog">common frog</a> <br /> (<i>Rana temporaria</i>)</figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Litoria_tyleri.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/9/96/Litoria_tyleri.jpg/250px-Litoria_tyleri.jpg" decoding="async" width="220" height="196" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/96/Litoria_tyleri.jpg/330px-Litoria_tyleri.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/96/Litoria_tyleri.jpg/500px-Litoria_tyleri.jpg 2x" data-file-width="1131" data-file-height="1008" /></a><figcaption><a href="/wiki/Tyler%27s_tree_frog" title="Tyler&#39;s tree frog">Tyler's tree frog</a> (<i>Litoria tyleri</i>) has large toe pads and webbed feet.</figcaption></figure> <p>The muscular system has been similarly modified. The hind limbs of ancestral frogs presumably contained pairs of muscles which would act in opposition (one muscle to flex the knee, a different muscle to extend it), as is seen in most other limbed animals. However, in modern frogs, almost all muscles have been modified to contribute to the action of jumping, with only a few small muscles remaining to bring the limb back to the starting position and maintain posture. The muscles have also been greatly enlarged, with the main leg muscles accounting for over 17% of the total mass of frogs.<sup id="cite_ref-Frogjump_53-0" class="reference"><a href="#cite_note-Frogjump-53"><span class="cite-bracket">&#91;</span>53<span class="cite-bracket">&#93;</span></a></sup> </p><p>Many frogs have webbed feet and the degree of webbing is directly proportional to the amount of time the species spends in the water.<sup id="cite_ref-Exploratorium_54-0" class="reference"><a href="#cite_note-Exploratorium-54"><span class="cite-bracket">&#91;</span>54<span class="cite-bracket">&#93;</span></a></sup> The completely aquatic <a href="/wiki/African_dwarf_frog" title="African dwarf frog">African dwarf frog</a> (<i>Hymenochirus</i> sp.) has fully webbed toes, whereas those of <a href="/wiki/White%27s_tree_frog" class="mw-redirect" title="White&#39;s tree frog">White's tree frog</a> (<i>Litoria caerulea</i>), an arboreal species, are only a quarter or half webbed.<sup id="cite_ref-55" class="reference"><a href="#cite_note-55"><span class="cite-bracket">&#91;</span>55<span class="cite-bracket">&#93;</span></a></sup> Exceptions include <a href="/wiki/Flying_frog" title="Flying frog">flying frogs</a> in the <a href="/wiki/Hylidae" title="Hylidae">Hylidae</a> and <a href="/wiki/Rhacophoridae" title="Rhacophoridae">Rhacophoridae</a>, which also have fully webbed toes used in gliding. </p><p><a href="/wiki/Tree_frog" title="Tree frog">Arboreal frogs</a> have pads located on the ends of their toes to help grip vertical surfaces. These are not suction pads, the surface consisting instead of columnar cells with flat tops with small gaps between them lubricated by mucous glands. When the frog applies pressure, the cells adhere to irregularities on the surface and the grip is maintained through <a href="/wiki/Capillarity" class="mw-redirect" title="Capillarity">surface tension</a>. This allows the frog to climb on smooth surfaces, but the system does not function efficiently when the pads are excessively wet.<sup id="cite_ref-56" class="reference"><a href="#cite_note-56"><span class="cite-bracket">&#91;</span>56<span class="cite-bracket">&#93;</span></a></sup> </p><p>In many arboreal frogs, a small "intercalary structure" on each toe increases the surface area touching the <a href="/wiki/Substrate_(biology)" title="Substrate (biology)">substrate</a>. Furthermore, many arboreal frogs have hip joints that allow both hopping and walking. Some frogs that live high in trees even possess an elaborate degree of webbing between their toes. This allows the frogs to "parachute" or make a controlled glide from one position in the canopy to another.<sup id="cite_ref-57" class="reference"><a href="#cite_note-57"><span class="cite-bracket">&#91;</span>57<span class="cite-bracket">&#93;</span></a></sup> </p><p>Ground-dwelling frogs generally lack the adaptations of aquatic and arboreal frogs. Most have smaller toe pads, if any, and little webbing. Some burrowing frogs such as <a href="/wiki/Couch%27s_Spadefoot_Toad" class="mw-redirect" title="Couch&#39;s Spadefoot Toad">Couch's spadefoot</a> (<i>Scaphiopus couchii</i>) have a flap-like toe extension on the hind feet, a <a href="/wiki/Keratin" title="Keratin">keratinised</a> <a href="/wiki/Tubercle" title="Tubercle">tubercle</a> often referred to as a spade, that helps them to burrow.<sup id="cite_ref-58" class="reference"><a href="#cite_note-58"><span class="cite-bracket">&#91;</span>58<span class="cite-bracket">&#93;</span></a></sup> </p><p>Sometimes during the tadpole stage, one of the developing rear legs is eaten by a predator such as a <a href="/wiki/Dragonfly_nymph" class="mw-redirect" title="Dragonfly nymph">dragonfly nymph</a>. In some cases, the full leg still grows, but in others it does not, although the frog may still live out its normal lifespan with only three limbs. Occasionally, a parasitic <a href="/wiki/Flatworm" title="Flatworm">flatworm</a> (<i><a href="/wiki/Ribeiroia_ondatrae" title="Ribeiroia ondatrae">Ribeiroia ondatrae</a></i>) digs into the rear of a tadpole, causing a rearrangement of the limb bud cells and the frog develops one or more extra legs.<sup id="cite_ref-59" class="reference"><a href="#cite_note-59"><span class="cite-bracket">&#91;</span>59<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Ranapipiensmoulting.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/f1/Ranapipiensmoulting.jpg/250px-Ranapipiensmoulting.jpg" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/f1/Ranapipiensmoulting.jpg/330px-Ranapipiensmoulting.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/f1/Ranapipiensmoulting.jpg/500px-Ranapipiensmoulting.jpg 2x" data-file-width="3648" data-file-height="2736" /></a><figcaption><a href="/wiki/Northern_leopard_frog" title="Northern leopard frog">Northern leopard frog</a> (<i>Rana pipiens</i>) moulting and eating its skin</figcaption></figure> <div class="mw-heading mw-heading3"><h3 id="Skin">Skin</h3></div> <p>A frog's skin is protective, has a respiratory function, can absorb water, and helps control body temperature. It has many glands, particularly on the head and back, which often exude distasteful and toxic substances (<a href="/wiki/Skin" title="Skin">granular glands</a>). The secretion is often sticky and helps keep the skin moist, protects against the entry of moulds and bacteria, and makes the animal slippery and more able to escape from predators.<sup id="cite_ref-60" class="reference"><a href="#cite_note-60"><span class="cite-bracket">&#91;</span>60<span class="cite-bracket">&#93;</span></a></sup> The skin is shed every few weeks. It usually splits down the middle of the back and across the belly, and the frog pulls its arms and legs free. The sloughed skin is then worked towards the head where it is quickly eaten.<sup id="cite_ref-61" class="reference"><a href="#cite_note-61"><span class="cite-bracket">&#91;</span>61<span class="cite-bracket">&#93;</span></a></sup> </p><p>Being cold-blooded, frogs have to adopt suitable behaviour patterns to regulate their temperature. To warm up, they can move into the sun or onto a warm surface; if they overheat, they can move into the shade or adopt a stance that exposes the minimum area of skin to the air. This posture is also used to prevent water loss and involves the frog squatting close to the substrate with its hands and feet tucked under its chin and body.<sup id="cite_ref-BandN_62-0" class="reference"><a href="#cite_note-BandN-62"><span class="cite-bracket">&#91;</span>62<span class="cite-bracket">&#93;</span></a></sup> The colour of a frog's skin is used for thermoregulation. In cool damp conditions, the colour will be darker than on a hot dry day. The <a href="/wiki/Grey_Foam-nest_Treefrog" class="mw-redirect" title="Grey Foam-nest Treefrog">grey foam-nest tree frog</a> (<i>Chiromantis xerampelina</i>) is even able to turn white to minimise the chance of overheating.<sup id="cite_ref-63" class="reference"><a href="#cite_note-63"><span class="cite-bracket">&#91;</span>63<span class="cite-bracket">&#93;</span></a></sup> </p><p>Many frogs are able to absorb water and oxygen directly through the skin, especially around the pelvic area, but the permeability of a frog's skin can also result in water loss. Glands located all over the body exude mucus which helps keep the skin moist and reduces evaporation. Some glands on the hands and chest of males are specialised to produce sticky secretions to aid in <a href="/wiki/Amplexus" title="Amplexus">amplexus</a>. Similar glands in tree frogs produce a glue-like substance on the adhesive discs of the feet. Some arboreal frogs reduce water loss by having a waterproof layer of skin, and several South American species coat their skin with a waxy secretion. Other frogs have adopted behaviours to conserve water, including becoming <a href="/wiki/Nocturnal" class="mw-redirect" title="Nocturnal">nocturnal</a> and resting in a water-conserving position. Some frogs may also rest in large groups with each frog pressed against its neighbours. This reduces the amount of skin exposed to the air or a dry surface, and thus reduces water loss.<sup id="cite_ref-BandN_62-1" class="reference"><a href="#cite_note-BandN-62"><span class="cite-bracket">&#91;</span>62<span class="cite-bracket">&#93;</span></a></sup> <a href="/wiki/Woodhouse%27s_toad" title="Woodhouse&#39;s toad">Woodhouse's toad</a> (<i>Bufo woodhousii</i>), if given access to water after confinement in a dry location, sits in the shallows to rehydrate.<sup id="cite_ref-64" class="reference"><a href="#cite_note-64"><span class="cite-bracket">&#91;</span>64<span class="cite-bracket">&#93;</span></a></sup> The male <a href="/wiki/Hairy_frog" title="Hairy frog">hairy frog</a> (<i>Trichobatrachus robustus</i>) has <a href="/wiki/Dermal_papillae" class="mw-redirect" title="Dermal papillae">dermal papillae</a> projecting from its lower back and thighs, giving it a bristly appearance. These contain blood vessels and are thought to increase the area of the skin available for respiration.<sup id="cite_ref-65" class="reference"><a href="#cite_note-65"><span class="cite-bracket">&#91;</span>65<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Hip-pocket_Frog_-_Assa_darlingtoni.jpg" class="mw-file-description"><img alt="Frog barely recognisable against brown decaying leaf litter." src="//upload.wikimedia.org/wikipedia/commons/thumb/1/11/Hip-pocket_Frog_-_Assa_darlingtoni.jpg/250px-Hip-pocket_Frog_-_Assa_darlingtoni.jpg" decoding="async" width="220" height="126" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/11/Hip-pocket_Frog_-_Assa_darlingtoni.jpg/330px-Hip-pocket_Frog_-_Assa_darlingtoni.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/11/Hip-pocket_Frog_-_Assa_darlingtoni.jpg/500px-Hip-pocket_Frog_-_Assa_darlingtoni.jpg 2x" data-file-width="2136" data-file-height="1220" /></a><figcaption><a href="/wiki/Pouched_frog" title="Pouched frog">Pouched frog</a> (<i>Assa darlingtoni</i>) camouflaged against leaf litter</figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:WoodFrog_DarienLakesStatePark_2020-06-16_(02).jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/f6/WoodFrog_DarienLakesStatePark_2020-06-16_%2802%29.jpg/250px-WoodFrog_DarienLakesStatePark_2020-06-16_%2802%29.jpg" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/f6/WoodFrog_DarienLakesStatePark_2020-06-16_%2802%29.jpg/330px-WoodFrog_DarienLakesStatePark_2020-06-16_%2802%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/f6/WoodFrog_DarienLakesStatePark_2020-06-16_%2802%29.jpg/500px-WoodFrog_DarienLakesStatePark_2020-06-16_%2802%29.jpg 2x" data-file-width="2520" data-file-height="1889" /></a><figcaption><a href="/wiki/Wood_frog" title="Wood frog">Wood frog</a> (<i>Lithobates sylvaticus</i>) uses <a href="/wiki/Disruptive_coloration" title="Disruptive coloration">disruptive coloration</a>.</figcaption></figure> <p>Some species have <a href="/wiki/Osteoderm" title="Osteoderm">bony plates</a> embedded in the skin, a trait that appears to have evolved independently several times.<sup id="cite_ref-66" class="reference"><a href="#cite_note-66"><span class="cite-bracket">&#91;</span>66<span class="cite-bracket">&#93;</span></a></sup> In certain other species, the skin at the top of the head is compacted and the connective tissue of the dermis is co-ossified with the bones of the skull (<a href="/wiki/Exostosis" title="Exostosis">exostosis</a>).<sup id="cite_ref-67" class="reference"><a href="#cite_note-67"><span class="cite-bracket">&#91;</span>67<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-68" class="reference"><a href="#cite_note-68"><span class="cite-bracket">&#91;</span>68<span class="cite-bracket">&#93;</span></a></sup> </p><p><a href="/wiki/Camouflage" title="Camouflage">Camouflage</a> is a common defensive mechanism in frogs. Features such as warts and <a href="/wiki/Skin_fold" title="Skin fold">skin folds</a> are usually on ground-dwelling frogs, for whom smooth skin would not provide such effective camouflage. Certain frogs change colour between night and day, as light and moisture stimulate the pigment cells and cause them to expand or contract.<sup id="cite_ref-Observer_69-0" class="reference"><a href="#cite_note-Observer-69"><span class="cite-bracket">&#91;</span>69<span class="cite-bracket">&#93;</span></a></sup> Some are even able to control their skin texture.<sup id="cite_ref-70" class="reference"><a href="#cite_note-70"><span class="cite-bracket">&#91;</span>70<span class="cite-bracket">&#93;</span></a></sup> The <a href="/wiki/Pacific_tree_frog" title="Pacific tree frog">Pacific tree frog</a> (<i>Pseudacris regilla</i>) has green and brown morphs, plain or spotted, and changes colour depending on the time of year and general background colour.<sup id="cite_ref-71" class="reference"><a href="#cite_note-71"><span class="cite-bracket">&#91;</span>71<span class="cite-bracket">&#93;</span></a></sup> The <a href="/wiki/Wood_frog" title="Wood frog">Wood frog</a> (<i>Lithobates sylvaticus</i>) uses <a href="/wiki/Disruptive_coloration" title="Disruptive coloration">disruptive coloration</a> including black eye markings similar to voids between leaves, bands of the dorsal skin (dorsolateral dermal plica) similar to a leaf <a href="/wiki/Midrib" class="mw-redirect" title="Midrib">midrib</a> as well as stains, spots and leg stripes similar to fallen leaf features. </p> <div class="mw-heading mw-heading3"><h3 id="Respiration_and_circulation">Respiration and circulation</h3></div> <p>Like other amphibians, <a href="/wiki/Oxygen" title="Oxygen">oxygen</a> can pass through their highly permeable skins. This unique feature allows them to remain in places without access to the air, respiring through their skins. Ribs are generally absent, so the lungs are filled by <a href="/wiki/Buccal_pumping" title="Buccal pumping">buccal pumping</a> and a frog deprived of its lungs can maintain its body functions without them.<sup id="cite_ref-Observer_69-1" class="reference"><a href="#cite_note-Observer-69"><span class="cite-bracket">&#91;</span>69<span class="cite-bracket">&#93;</span></a></sup> The fully aquatic <a href="/wiki/Bornean_flat-headed_frog" title="Bornean flat-headed frog">Bornean flat-headed frog</a> (<i>Barbourula kalimantanensis</i>) is the first frog known to lack lungs entirely.<sup id="cite_ref-72" class="reference"><a href="#cite_note-72"><span class="cite-bracket">&#91;</span>72<span class="cite-bracket">&#93;</span></a></sup> </p><p>Frogs have three-chambered <a href="/wiki/Heart" title="Heart">hearts</a>, a feature they share with <a href="/wiki/Lizard" title="Lizard">lizards</a>. Oxygenated blood from the lungs and de-oxygenated blood from the <a href="/wiki/Respiration_(physiology)" title="Respiration (physiology)">respiring</a> tissues enter the heart through separate <a href="/wiki/Atrium_(anatomy)" class="mw-redirect" title="Atrium (anatomy)">atria</a>. When these chambers contract, the two blood streams pass into a common <a href="/wiki/Ventricle_(heart)" title="Ventricle (heart)">ventricle</a> before being pumped via a spiral valve to the appropriate vessel, the <a href="/wiki/Aorta" title="Aorta">aorta</a> for oxygenated blood and <a href="/wiki/Pulmonary_artery" title="Pulmonary artery">pulmonary artery</a> for deoxygenated blood.<sup id="cite_ref-Kimball_73-0" class="reference"><a href="#cite_note-Kimball-73"><span class="cite-bracket">&#91;</span>73<span class="cite-bracket">&#93;</span></a></sup> </p><p>Some species of frog have adaptations that allow them to survive in oxygen deficient water. The <a href="/wiki/Titicaca_water_frog" class="mw-redirect" title="Titicaca water frog">Titicaca water frog</a> (<i>Telmatobius culeus</i>) is one such species and has wrinkly skin that increases its surface area to enhance gas exchange. It normally makes no use of its rudimentary lungs but will sometimes raise and lower its body rhythmically while on the lake bed to increase the flow of water around it.<sup id="cite_ref-74" class="reference"><a href="#cite_note-74"><span class="cite-bracket">&#91;</span>74<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Frog_anatomy_tags.PNG" class="mw-file-description"><img alt="Dissected frog" src="//upload.wikimedia.org/wikipedia/commons/thumb/2/28/Frog_anatomy_tags.PNG/220px-Frog_anatomy_tags.PNG" decoding="async" width="220" height="294" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/28/Frog_anatomy_tags.PNG/330px-Frog_anatomy_tags.PNG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/28/Frog_anatomy_tags.PNG/440px-Frog_anatomy_tags.PNG 2x" data-file-width="1248" data-file-height="1668" /></a><figcaption>Anatomical model of a dissected frog: 1&#160;Right&#160;atrium, 2&#160;Lungs, 3&#160;Aorta, 4&#160;Egg mass, 5&#160;Colon, 6&#160;Left&#160;atrium, 7&#160;Ventricle, 8&#160;Stomach, 9&#160;Liver, 10&#160;Gallbladder, 11&#160;Small&#160;intestine, 12&#160;Cloaca</figcaption></figure> <div class="mw-heading mw-heading3"><h3 id="Digestion_and_excretion">Digestion and excretion</h3></div> <p>Frogs have maxillary teeth along their upper jaw which are used to hold food before it is swallowed. These teeth are very weak, and cannot be used to chew or catch and harm agile prey. Instead, the frog uses its sticky, cleft tongue to catch insects and other small moving prey. The tongue normally lies coiled in the mouth, free at the back and attached to the mandible at the front. It can be shot out and retracted at great speed.<sup id="cite_ref-Exploratorium_54-1" class="reference"><a href="#cite_note-Exploratorium-54"><span class="cite-bracket">&#91;</span>54<span class="cite-bracket">&#93;</span></a></sup> In amphibians there are salvary glands on the tongue, which in frogs produce what is called a two-phase viscoelastic fluid. When exposed to pressure, like when the tongue is wrapping around a prey, it becomes runny and covers the prey's body. As the pressure drops, it returns to a thick and elastic state, which gives the tongue an extra grip.<sup id="cite_ref-75" class="reference"><a href="#cite_note-75"><span class="cite-bracket">&#91;</span>75<span class="cite-bracket">&#93;</span></a></sup> Some frogs have no tongue and just stuff food into their mouths with their hands.<sup id="cite_ref-Exploratorium_54-2" class="reference"><a href="#cite_note-Exploratorium-54"><span class="cite-bracket">&#91;</span>54<span class="cite-bracket">&#93;</span></a></sup> The African bullfrog (<i><a href="/wiki/Pyxicephalus" title="Pyxicephalus">Pyxicephalus</a></i>), which preys on relatively large animals such as mice and other frogs, has cone shaped bony projections called odontoid processes at the front of the lower jaw which function like teeth.<sup id="cite_ref-Pou92_16-1" class="reference"><a href="#cite_note-Pou92-16"><span class="cite-bracket">&#91;</span>16<span class="cite-bracket">&#93;</span></a></sup> The eyes assist in the swallowing of food as they can be retracted through holes in the skull and help push food down the throat.<sup id="cite_ref-Exploratorium_54-3" class="reference"><a href="#cite_note-Exploratorium-54"><span class="cite-bracket">&#91;</span>54<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-76" class="reference"><a href="#cite_note-76"><span class="cite-bracket">&#91;</span>76<span class="cite-bracket">&#93;</span></a></sup> </p><p>The food then moves through the oesophagus into the stomach where digestive enzymes are added and it is churned up. It then proceeds to the small intestine (duodenum and ileum) where most digestion occurs. Pancreatic juice from the pancreas, and bile, produced by the liver and stored in the gallbladder, are secreted into the small intestine, where the fluids digest the food and the nutrients are absorbed. The food residue passes into the large intestine where excess water is removed and the wastes are passed out through the <a href="/wiki/Cloaca" title="Cloaca">cloaca</a>.<sup id="cite_ref-77" class="reference"><a href="#cite_note-77"><span class="cite-bracket">&#91;</span>77<span class="cite-bracket">&#93;</span></a></sup> </p><p>Although adapted to terrestrial life, frogs resemble freshwater fish in their inability to conserve body water effectively. When they are on land, much water is lost by evaporation from the skin. The excretory system is similar to that of mammals and there are two <a href="/wiki/Kidney" title="Kidney">kidneys</a> that remove nitrogenous products from the blood. Frogs produce large quantities of dilute urine in order to flush out toxic products from the kidney tubules.<sup id="cite_ref-Doritexcretion_78-0" class="reference"><a href="#cite_note-Doritexcretion-78"><span class="cite-bracket">&#91;</span>78<span class="cite-bracket">&#93;</span></a></sup> The nitrogen is excreted as <a href="/wiki/Ammonia" title="Ammonia">ammonia</a> by tadpoles and aquatic frogs but mainly as <a href="/wiki/Urea" title="Urea">urea</a>, a less toxic product, by most terrestrial adults. A few species of tree frog with little access to water excrete the even less toxic <a href="/wiki/Uric_acid" title="Uric acid">uric acid</a>.<sup id="cite_ref-Doritexcretion_78-1" class="reference"><a href="#cite_note-Doritexcretion-78"><span class="cite-bracket">&#91;</span>78<span class="cite-bracket">&#93;</span></a></sup> The urine passes along paired <a href="/wiki/Ureter" title="Ureter">ureters</a> to the <a href="/wiki/Urinary_bladder" class="mw-redirect" title="Urinary bladder">urinary bladder</a> from which it is vented periodically into the cloaca. All bodily wastes exit the body through the cloaca which terminates in a cloacal vent.<sup id="cite_ref-TutorVista_79-0" class="reference"><a href="#cite_note-TutorVista-79"><span class="cite-bracket">&#91;</span>79<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Reproductive_system">Reproductive system</h3></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951" /><div role="note" class="hatnote navigation-not-searchable">See also: <a href="/wiki/Sexual_selection_in_amphibians" title="Sexual selection in amphibians">Sexual selection in amphibians</a></div> <p>In the male frog, the two <a href="/wiki/Testis" class="mw-redirect" title="Testis">testes</a> are attached to the kidneys and <a href="/wiki/Semen" title="Semen">semen</a> passes into the kidneys through fine tubes called <a href="/wiki/Efferent_ducts" title="Efferent ducts">efferent ducts</a>. It then travels on through the ureters, which are consequently known as urinogenital ducts. There is no penis, and sperm is ejected from the cloaca directly onto the eggs as the female lays them. The ovaries of the female frog are beside the kidneys and the eggs pass down a pair of oviducts and through the cloaca to the exterior.<sup id="cite_ref-TutorVista_79-1" class="reference"><a href="#cite_note-TutorVista-79"><span class="cite-bracket">&#91;</span>79<span class="cite-bracket">&#93;</span></a></sup> </p><p>When frogs mate, the male climbs on the back of the female and wraps his fore limbs round her body, either behind the front legs or just in front of the hind legs. This position is called <a href="/wiki/Amplexus" title="Amplexus">amplexus</a> and may be held for several days.<sup id="cite_ref-80" class="reference"><a href="#cite_note-80"><span class="cite-bracket">&#91;</span>80<span class="cite-bracket">&#93;</span></a></sup> The male frog has certain hormone-dependent <a href="/wiki/Secondary_sexual_characteristic" class="mw-redirect" title="Secondary sexual characteristic">secondary sexual characteristics</a>. These include the development of special pads on his thumbs in the breeding season, to give him a firm hold.<sup id="cite_ref-81" class="reference"><a href="#cite_note-81"><span class="cite-bracket">&#91;</span>81<span class="cite-bracket">&#93;</span></a></sup> The grip of the male frog during amplexus stimulates the female to release eggs, usually wrapped in jelly, as spawn. In many species the male is smaller and slimmer than the female. Males have vocal cords and make a range of croaks, particularly in the breeding season, and in some species they also have <a href="/wiki/Vocal_sac" title="Vocal sac">vocal sacs</a> to amplify the sound.<sup id="cite_ref-TutorVista_79-2" class="reference"><a href="#cite_note-TutorVista-79"><span class="cite-bracket">&#91;</span>79<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Nervous_system">Nervous system</h3></div> <p>Frogs have a highly developed nervous system that consists of a brain, spinal cord and nerves. Many parts of frog brains correspond with those of humans. It consists of two olfactory lobes, two cerebral hemispheres, a pineal body, two optic lobes, a cerebellum and a medulla oblongata. Muscular coordination and posture are controlled by the <a href="/wiki/Cerebellum" title="Cerebellum">cerebellum</a>, and the <a href="/wiki/Medulla_oblongata" title="Medulla oblongata">medulla oblongata</a> regulates respiration, digestion and other automatic functions. The relative size of the <a href="/wiki/Cerebrum" title="Cerebrum">cerebrum</a> in frogs is much smaller than it is in humans. Frogs have ten pairs of <a href="/wiki/Cranial_nerves" title="Cranial nerves">cranial nerves</a> which pass information from the outside directly to the brain, and ten pairs of <a href="/wiki/Spinal_nerves" class="mw-redirect" title="Spinal nerves">spinal nerves</a> which pass information from the extremities to the brain through the spinal cord.<sup id="cite_ref-TutorVista_79-3" class="reference"><a href="#cite_note-TutorVista-79"><span class="cite-bracket">&#91;</span>79<span class="cite-bracket">&#93;</span></a></sup> By contrast, all <a href="/wiki/Amniotes" class="mw-redirect" title="Amniotes">amniotes</a> (mammals, birds and reptiles) have twelve pairs of cranial nerves.<sup id="cite_ref-82" class="reference"><a href="#cite_note-82"><span class="cite-bracket">&#91;</span>82<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Groene_kikker_achter_Bekaert-draad-detail_oog.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/6/6f/Groene_kikker_achter_Bekaert-draad-detail_oog.jpg/220px-Groene_kikker_achter_Bekaert-draad-detail_oog.jpg" decoding="async" width="220" height="139" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/6f/Groene_kikker_achter_Bekaert-draad-detail_oog.jpg/330px-Groene_kikker_achter_Bekaert-draad-detail_oog.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/6f/Groene_kikker_achter_Bekaert-draad-detail_oog.jpg/440px-Groene_kikker_achter_Bekaert-draad-detail_oog.jpg 2x" data-file-width="771" data-file-height="488" /></a><figcaption>Close-up of frog's head showing eye, nostril, mouth, and tympanum</figcaption></figure> <div class="mw-heading mw-heading3"><h3 id="Sight">Sight</h3></div> <p>The eyes of most frogs are located on either side of the head near the top and project outwards as hemispherical bulges. They provide <a href="/wiki/Binocular_vision" title="Binocular vision">binocular vision</a> over a field of 100° to the front and a total visual field of almost 360°.<sup id="cite_ref-Howard_83-0" class="reference"><a href="#cite_note-Howard-83"><span class="cite-bracket">&#91;</span>83<span class="cite-bracket">&#93;</span></a></sup> They may be the only part of an otherwise submerged frog to protrude from the water. Each eye has closable upper and lower lids and a <a href="/wiki/Nictitating_membrane" title="Nictitating membrane">nictitating membrane</a> which provides further protection, especially when the frog is swimming.<sup id="cite_ref-Badger_84-0" class="reference"><a href="#cite_note-Badger-84"><span class="cite-bracket">&#91;</span>84<span class="cite-bracket">&#93;</span></a></sup> Members of the aquatic family <a href="/wiki/Pipidae" title="Pipidae">Pipidae</a> have the eyes located at the top of the head, a position better suited for detecting prey in the water above.<sup id="cite_ref-Howard_83-1" class="reference"><a href="#cite_note-Howard-83"><span class="cite-bracket">&#91;</span>83<span class="cite-bracket">&#93;</span></a></sup> The <a href="/wiki/Iris_(anatomy)" title="Iris (anatomy)">irises</a> come in a range of colours and the pupils in a range of shapes. The <a href="/wiki/Common_toad" title="Common toad">common toad</a> (<i>Bufo bufo</i>) has golden irises and horizontal slit-like pupils, the <a href="/wiki/Agalychnis_callidryas" title="Agalychnis callidryas">red-eyed tree frog</a> (<i>Agalychnis callidryas</i>) has vertical slit pupils, the <a href="/wiki/Poison_dart_frog" title="Poison dart frog">poison dart frog</a> has dark irises, the <a href="/wiki/Fire-bellied_toad" title="Fire-bellied toad">fire-bellied toad</a> (<i>Bombina spp.</i>) has triangular pupils and the <a href="/wiki/Dyscophus_antongilii" title="Dyscophus antongilii">tomato frog</a> (<i>Dyscophus spp.</i>) has circular ones. The irises of the <a href="/wiki/Southern_toad" title="Southern toad">southern toad</a> (<i>Anaxyrus terrestris</i>) are patterned so as to blend in with the surrounding camouflaged skin.<sup id="cite_ref-Badger_84-1" class="reference"><a href="#cite_note-Badger-84"><span class="cite-bracket">&#91;</span>84<span class="cite-bracket">&#93;</span></a></sup> </p><p>The distant vision of a frog is better than its near vision. Calling frogs will quickly become silent when they see an intruder or even a moving shadow but the closer an object is, the less well it is seen.<sup id="cite_ref-Badger_84-2" class="reference"><a href="#cite_note-Badger-84"><span class="cite-bracket">&#91;</span>84<span class="cite-bracket">&#93;</span></a></sup> When a frog shoots out its tongue to catch an insect it is reacting to a small moving object that it cannot see well and must line it up precisely beforehand because it shuts its eyes as the tongue is extended.<sup id="cite_ref-Exploratorium_54-4" class="reference"><a href="#cite_note-Exploratorium-54"><span class="cite-bracket">&#91;</span>54<span class="cite-bracket">&#93;</span></a></sup> Although it was formerly debated,<sup id="cite_ref-85" class="reference"><a href="#cite_note-85"><span class="cite-bracket">&#91;</span>85<span class="cite-bracket">&#93;</span></a></sup> more recent research has shown that frogs can see in colour, even in very low light.<sup id="cite_ref-86" class="reference"><a href="#cite_note-86"><span class="cite-bracket">&#91;</span>86<span class="cite-bracket">&#93;</span></a></sup> </p> <div style="clear:both;" class=""></div> <div class="mw-heading mw-heading3"><h3 id="Hearing">Hearing</h3></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><span><video id="mwe_player_0" poster="//upload.wikimedia.org/wikipedia/commons/thumb/d/d4/Synchrotron_microtomography_of_Atelopus_franciscus_head_-_pone.0022080.s003.ogv/220px--Synchrotron_microtomography_of_Atelopus_franciscus_head_-_pone.0022080.s003.ogv.jpg" controls="" preload="none" data-mw-tmh="" class="mw-file-element" width="220" height="139" data-durationhint="41" data-mwtitle="Synchrotron_microtomography_of_Atelopus_franciscus_head_-_pone.0022080.s003.ogv" data-mwprovider="wikimediacommons" resource="/wiki/File:Synchrotron_microtomography_of_Atelopus_franciscus_head_-_pone.0022080.s003.ogv"><source src="//upload.wikimedia.org/wikipedia/commons/d/d4/Synchrotron_microtomography_of_Atelopus_franciscus_head_-_pone.0022080.s003.ogv" type="video/ogg; codecs=&quot;theora&quot;" data-width="696" data-height="440" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/d/d4/Synchrotron_microtomography_of_Atelopus_franciscus_head_-_pone.0022080.s003.ogv/Synchrotron_microtomography_of_Atelopus_franciscus_head_-_pone.0022080.s003.ogv.240p.vp9.webm" type="video/webm; codecs=&quot;vp9, opus&quot;" data-transcodekey="240p.vp9.webm" data-width="380" data-height="240" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/d/d4/Synchrotron_microtomography_of_Atelopus_franciscus_head_-_pone.0022080.s003.ogv/Synchrotron_microtomography_of_Atelopus_franciscus_head_-_pone.0022080.s003.ogv.360p.vp9.webm" type="video/webm; codecs=&quot;vp9, opus&quot;" data-transcodekey="360p.vp9.webm" data-width="570" data-height="360" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/d/d4/Synchrotron_microtomography_of_Atelopus_franciscus_head_-_pone.0022080.s003.ogv/Synchrotron_microtomography_of_Atelopus_franciscus_head_-_pone.0022080.s003.ogv.360p.webm" type="video/webm; codecs=&quot;vp8, vorbis&quot;" data-transcodekey="360p.webm" data-width="570" data-height="360" /></video></span><figcaption>Surface rendering of the head of the frog <i><a href="/wiki/Atelopus_franciscus" title="Atelopus franciscus">Atelopus franciscus</a></i>, with ear parts highlighted</figcaption></figure> <p>Frogs can hear both in the air and below water. They do not have <a href="/wiki/Pinna_(anatomy)" class="mw-redirect" title="Pinna (anatomy)">external ears</a>; the eardrums (<a href="/wiki/Tympanum_(anatomy)" title="Tympanum (anatomy)">tympanic membranes</a>) are directly exposed or may be covered by a layer of skin and are visible as a circular area just behind the eye. The size and distance apart of the eardrums is related to the frequency and wavelength at which the frog calls. In some species such as the bullfrog, the size of the tympanum indicates the sex of the frog; males have tympani that are larger than their eyes while in females, the eyes and tympani are much the same size.<sup id="cite_ref-Neth38_87-0" class="reference"><a href="#cite_note-Neth38-87"><span class="cite-bracket">&#91;</span>87<span class="cite-bracket">&#93;</span></a></sup> A noise causes the tympanum to vibrate and the sound is transmitted to the middle and inner ear. The middle ear contains semicircular canals which help control balance and orientation. In the inner ear, the auditory hair cells are arranged in two areas of the cochlea, the basilar papilla and the amphibian papilla. The former detects high frequencies and the latter low frequencies.<sup id="cite_ref-Cohen_88-0" class="reference"><a href="#cite_note-Cohen-88"><span class="cite-bracket">&#91;</span>88<span class="cite-bracket">&#93;</span></a></sup> Because the cochlea is short, frogs use <a href="/wiki/Electrical_tuning" title="Electrical tuning">electrical tuning</a> to extend their range of audible frequencies and help discriminate different sounds.<sup id="cite_ref-89" class="reference"><a href="#cite_note-89"><span class="cite-bracket">&#91;</span>89<span class="cite-bracket">&#93;</span></a></sup> This arrangement enables detection of the territorial and breeding calls of their <a href="/wiki/Conspecificity" class="mw-redirect" title="Conspecificity">conspecifics</a>. In some species that inhabit arid regions, the sound of thunder or heavy rain may arouse them from a dormant state.<sup id="cite_ref-Cohen_88-1" class="reference"><a href="#cite_note-Cohen-88"><span class="cite-bracket">&#91;</span>88<span class="cite-bracket">&#93;</span></a></sup> A frog may be startled by an unexpected noise but it will not usually take any action until it has located the source of the sound by sight.<sup id="cite_ref-Neth38_87-1" class="reference"><a href="#cite_note-Neth38-87"><span class="cite-bracket">&#91;</span>87<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Call">Call</h3></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951" /><div role="note" class="hatnote navigation-not-searchable">See also: <a href="/wiki/Sexual_selection_in_frogs" class="mw-redirect" title="Sexual selection in frogs">Sexual selection in frogs</a></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Dendropsophus_microcephalus_-_calling_male_(Cope,_1886).jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8b/Dendropsophus_microcephalus_-_calling_male_%28Cope%2C_1886%29.jpg/250px-Dendropsophus_microcephalus_-_calling_male_%28Cope%2C_1886%29.jpg" decoding="async" width="220" height="147" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8b/Dendropsophus_microcephalus_-_calling_male_%28Cope%2C_1886%29.jpg/330px-Dendropsophus_microcephalus_-_calling_male_%28Cope%2C_1886%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8b/Dendropsophus_microcephalus_-_calling_male_%28Cope%2C_1886%29.jpg/500px-Dendropsophus_microcephalus_-_calling_male_%28Cope%2C_1886%29.jpg 2x" data-file-width="3000" data-file-height="2000" /></a><figcaption>A male <i><a href="/wiki/Dendropsophus_microcephalus" title="Dendropsophus microcephalus">Dendropsophus microcephalus</a></i> displaying its vocal sac during its call</figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><span><audio id="mwe_player_1" controls="" preload="none" data-mw-tmh="" class="mw-file-element" width="220" style="width:220px;" data-durationhint="95" data-mwtitle="Atelopus_franciscus_male_territorial_call_-_pone.0022080.s002.oga" data-mwprovider="wikimediacommons" resource="/wiki/File:Atelopus_franciscus_male_territorial_call_-_pone.0022080.s002.oga"><source src="//upload.wikimedia.org/wikipedia/commons/7/7e/Atelopus_franciscus_male_territorial_call_-_pone.0022080.s002.oga" type="audio/ogg; codecs=&quot;vorbis&quot;" data-width="0" data-height="0" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/7/7e/Atelopus_franciscus_male_territorial_call_-_pone.0022080.s002.oga/Atelopus_franciscus_male_territorial_call_-_pone.0022080.s002.oga.mp3" type="audio/mpeg" data-transcodekey="mp3" data-width="0" data-height="0" /></audio></span><figcaption>Advertisement call of male <i><a href="/wiki/Atelopus_franciscus" title="Atelopus franciscus">Atelopus franciscus</a></i></figcaption></figure> <p>The call or croak of a frog is unique to its species. Frogs create this sound by passing air through the <a href="/wiki/Larynx" title="Larynx">larynx</a> in the throat. In most calling frogs, the sound is amplified by one or more vocal sacs, membranes of skin under the throat or on the corner of the mouth, that distend during the amplification of the call. Some frog calls are so loud that they can be heard up to a mile (1.6<span class="nowrap">&#160;</span>km) away.<sup id="cite_ref-90" class="reference"><a href="#cite_note-90"><span class="cite-bracket">&#91;</span>90<span class="cite-bracket">&#93;</span></a></sup> Additionally, some species have been found to use man-made structures such as drain pipes for artificial amplification of their call.<sup id="cite_ref-91" class="reference"><a href="#cite_note-91"><span class="cite-bracket">&#91;</span>91<span class="cite-bracket">&#93;</span></a></sup> The <a href="/wiki/Ascaphus_truei" class="mw-redirect" title="Ascaphus truei">coastal tailed frog</a> (<i>Ascaphus truei</i>) lives in mountain streams in North America and does not vocalise.<sup id="cite_ref-92" class="reference"><a href="#cite_note-92"><span class="cite-bracket">&#91;</span>92<span class="cite-bracket">&#93;</span></a></sup> </p><p>The main function of calling is for male frogs to attract mates. Males may call individually or there may be a chorus of sound where numerous males have converged on breeding sites. In many frog species, such as the <a href="/wiki/Common_Tree_Frog" class="mw-redirect" title="Common Tree Frog">common tree frog</a> (<i>Polypedates leucomystax</i>), females reply to males' calls, which acts to reinforce reproductive activity in a breeding colony.<sup id="cite_ref-93" class="reference"><a href="#cite_note-93"><span class="cite-bracket">&#91;</span>93<span class="cite-bracket">&#93;</span></a></sup> Female frogs prefer males that produce sounds of greater intensity and lower frequency, attributes that stand out in a crowd. The rationale for this is thought to be that by demonstrating his prowess, the male shows his fitness to produce superior offspring.<sup id="cite_ref-94" class="reference"><a href="#cite_note-94"><span class="cite-bracket">&#91;</span>94<span class="cite-bracket">&#93;</span></a></sup> </p><p>A different call is emitted by a male frog or unreceptive female when mounted by another male. This is a distinct chirruping sound and is accompanied by a vibration of the body.<sup id="cite_ref-Badger3_95-0" class="reference"><a href="#cite_note-Badger3-95"><span class="cite-bracket">&#91;</span>95<span class="cite-bracket">&#93;</span></a></sup> Tree frogs and some non-aquatic species have a rain call that they make on the basis of humidity cues prior to a shower.<sup id="cite_ref-Badger3_95-1" class="reference"><a href="#cite_note-Badger3-95"><span class="cite-bracket">&#91;</span>95<span class="cite-bracket">&#93;</span></a></sup> Many species also have a territorial call that is used to drive away other males. All of these calls are emitted with the mouth of the frog closed.<sup id="cite_ref-Badger3_95-2" class="reference"><a href="#cite_note-Badger3-95"><span class="cite-bracket">&#91;</span>95<span class="cite-bracket">&#93;</span></a></sup> A distress call, emitted by some frogs when they are in danger, is produced with the mouth open resulting in a higher-pitched call. It is typically used when the frog has been grabbed by a predator and may serve to distract or disorient the attacker so that it releases the frog.<sup id="cite_ref-Badger3_95-3" class="reference"><a href="#cite_note-Badger3-95"><span class="cite-bracket">&#91;</span>95<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><span><audio id="mwe_player_2" controls="" preload="none" data-mw-tmh="" class="mw-file-element" width="220" style="width:220px;" data-durationhint="45" data-mwtitle="Banded_Bull_Frog_Call.ogg" data-mwprovider="wikimediacommons" resource="/wiki/File:Banded_Bull_Frog_Call.ogg"><source src="//upload.wikimedia.org/wikipedia/commons/c/c5/Banded_Bull_Frog_Call.ogg" type="audio/ogg; codecs=&quot;vorbis&quot;" data-width="0" data-height="0" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/c/c5/Banded_Bull_Frog_Call.ogg/Banded_Bull_Frog_Call.ogg.mp3" type="audio/mpeg" data-transcodekey="mp3" data-width="0" data-height="0" /></audio></span><figcaption>Distinctive low "jug-o-rum" sound of <a href="/wiki/Banded_bullfrog" title="Banded bullfrog">banded bullfrog</a></figcaption></figure> <p>Many species of frog have deep calls. The croak of the <a href="/wiki/American_bullfrog" title="American bullfrog">American bullfrog</a> (<i>Rana catesbiana</i>) is sometimes written as "jug o' rum".<sup id="cite_ref-96" class="reference"><a href="#cite_note-96"><span class="cite-bracket">&#91;</span>96<span class="cite-bracket">&#93;</span></a></sup> The <a href="/wiki/Pacific_Tree_Frog" class="mw-redirect" title="Pacific Tree Frog">Pacific tree frog</a> (<i>Pseudacris regilla</i>) produces the <a href="/wiki/Onomatopoeia" title="Onomatopoeia">onomatopoeic</a> "ribbit" often heard in films.<sup id="cite_ref-97" class="reference"><a href="#cite_note-97"><span class="cite-bracket">&#91;</span>97<span class="cite-bracket">&#93;</span></a></sup> Other renderings of frog calls into speech include "brekekekex koax koax", the call of the marsh frog (<i><a href="/wiki/Pelophylax_ridibundus" class="mw-redirect" title="Pelophylax ridibundus">Pelophylax ridibundus</a></i>) in <i>The Frogs</i>, an Ancient Greek comic drama by <a href="/wiki/Aristophanes" title="Aristophanes">Aristophanes</a>.<sup id="cite_ref-98" class="reference"><a href="#cite_note-98"><span class="cite-bracket">&#91;</span>98<span class="cite-bracket">&#93;</span></a></sup> The calls of the <a href="/wiki/Concave-eared_torrent_frog" title="Concave-eared torrent frog">Concave-eared torrent frog</a> (<i>Amolops tormotus</i>) are unusual in many aspects. The males are notable for their varieties of calls where upward and downward frequency modulations take place. When they communicate, they produce calls that fall in the <a href="/wiki/Ultrasound" title="Ultrasound">ultrasound</a> frequency range. The last aspect that makes this species of frog's calls unusual is that nonlinear acoustic phenomena are important components in their acoustic signals.<sup id="cite_ref-ultrasonic_99-0" class="reference"><a href="#cite_note-ultrasonic-99"><span class="cite-bracket">&#91;</span>99<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Torpor">Torpor</h3></div> <p>During extreme conditions, some frogs enter a state of <a href="/wiki/Torpor" title="Torpor">torpor</a> and remain inactive for months. In colder regions, many species of frog <a href="/wiki/Hibernation" title="Hibernation">hibernate</a> in winter. Those that live on land such as the <a href="/wiki/American_toad" title="American toad">American toad</a> (<i>Bufo americanus</i>) dig a burrow and make a <a href="/wiki/Hibernaculum_(zoology)" title="Hibernaculum (zoology)">hibernaculum</a> in which to lie <a href="/wiki/Dormancy" title="Dormancy">dormant</a>. Others, less proficient at digging, find a crevice or bury themselves in dead leaves. Aquatic species such as the <a href="/wiki/American_bullfrog" title="American bullfrog">American bullfrog</a> (<i>Rana catesbeiana</i>) normally sink to the bottom of the pond where they lie, semi-immersed in mud but still able to access the oxygen dissolved in the water. Their metabolism slows down and they live on their energy reserves. Some frogs such as the <a href="/wiki/Wood_frog" title="Wood frog">wood frog</a>, <a href="/wiki/Moor_frog" title="Moor frog">moor frog</a>, or <a href="/wiki/Spring_peeper" title="Spring peeper">spring peeper</a> can even survive being frozen. Ice crystals form under the skin and in the body cavity but the essential organs are protected from freezing by a high concentration of glucose. An apparently lifeless, frozen frog can resume respiration and its heartbeat can restart when conditions warm up.<sup id="cite_ref-100" class="reference"><a href="#cite_note-100"><span class="cite-bracket">&#91;</span>100<span class="cite-bracket">&#93;</span></a></sup> </p><p>At the other extreme, the <a href="/wiki/Striped_burrowing_frog" title="Striped burrowing frog">striped burrowing frog</a> (<i>Cyclorana alboguttata</i>) regularly <a href="/wiki/Aestivation" title="Aestivation">aestivates</a> during the hot, dry season in Australia, surviving in a dormant state without access to food and water for nine or ten months of the year. It burrows underground and curls up inside a protective <a href="/wiki/Cocoon_(silk)" class="mw-redirect" title="Cocoon (silk)">cocoon</a> formed by its shed skin. Researchers at the <a href="/wiki/University_of_Queensland" title="University of Queensland">University of Queensland</a> have found that during aestivation, the <a href="/wiki/Metabolism" title="Metabolism">metabolism</a> of the frog is altered and the operational efficiency of the <a href="/wiki/Mitochondria" class="mw-redirect" title="Mitochondria">mitochondria</a> is increased. This means that the limited amount of energy available to the comatose frog is used in a more efficient manner. This survival mechanism is only useful to animals that remain completely unconscious for an extended period of time and whose energy requirements are low because they are cold-blooded and have no need to generate heat.<sup id="cite_ref-101" class="reference"><a href="#cite_note-101"><span class="cite-bracket">&#91;</span>101<span class="cite-bracket">&#93;</span></a></sup> Other research showed that, to provide these energy requirements, muscles atrophy, but hind limb muscles are preferentially unaffected.<sup id="cite_ref-102" class="reference"><a href="#cite_note-102"><span class="cite-bracket">&#91;</span>102<span class="cite-bracket">&#93;</span></a></sup> Frogs have been found to have upper critical temperatures of around 41 degrees Celsius.<sup id="cite_ref-103" class="reference"><a href="#cite_note-103"><span class="cite-bracket">&#91;</span>103<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Locomotion">Locomotion</h2></div> <p>Different species of frog use a number of methods of moving around including <a href="/wiki/Jumping" title="Jumping">jumping</a>, <a href="/wiki/Running" title="Running">running</a>, <a href="/wiki/Walking" title="Walking">walking</a>, <a href="/wiki/Aquatic_locomotion" title="Aquatic locomotion">swimming</a>, <a href="/wiki/Burrow" title="Burrow">burrowing</a>, <a href="/wiki/Climbing" title="Climbing">climbing</a> and <a href="/wiki/Flying_frog" title="Flying frog">gliding</a>. </p> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Colostethus_flotator_jumping.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/85/Colostethus_flotator_jumping.jpg/250px-Colostethus_flotator_jumping.jpg" decoding="async" width="220" height="138" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/85/Colostethus_flotator_jumping.jpg/330px-Colostethus_flotator_jumping.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/85/Colostethus_flotator_jumping.jpg/500px-Colostethus_flotator_jumping.jpg 2x" data-file-width="3000" data-file-height="1887" /></a><figcaption><a href="/wiki/Rainforest_rocket_frog" class="mw-redirect" title="Rainforest rocket frog">Rainforest rocket frog</a> jumping</figcaption></figure> <div class="mw-heading mw-heading3"><h3 id="Jumping">Jumping</h3></div> <p>Frogs are generally recognised as exceptional jumpers and, relative to their size, the best jumpers of all vertebrates.<sup id="cite_ref-104" class="reference"><a href="#cite_note-104"><span class="cite-bracket">&#91;</span>104<span class="cite-bracket">&#93;</span></a></sup> The <a href="/wiki/Striped_rocket_frog" title="Striped rocket frog">striped rocket frog</a>, <i>Litoria nasuta</i>, can leap over <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1154941027" />two metres (<span class="frac">6<span class="sr-only">+</span><span class="num">1</span>&#8260;<span class="den">2</span></span> feet), a distance that is more than fifty times its body length of <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1154941027" />55&#160;mm (<span class="frac">2<span class="sr-only">+</span><span class="num">1</span>&#8260;<span class="den">4</span></span>&#160;in).<sup id="cite_ref-105" class="reference"><a href="#cite_note-105"><span class="cite-bracket">&#91;</span>105<span class="cite-bracket">&#93;</span></a></sup> There are tremendous differences between species in jumping capability. Within a species, jump distance increases with increasing size, but relative jumping distance (body-lengths jumped) decreases. The <a href="/wiki/Euphlyctis_cyanophlyctis" title="Euphlyctis cyanophlyctis">Indian skipper frog</a> (<i>Euphlyctis cyanophlyctis</i>) has the ability to leap out of the water from a position floating on the surface.<sup id="cite_ref-106" class="reference"><a href="#cite_note-106"><span class="cite-bracket">&#91;</span>106<span class="cite-bracket">&#93;</span></a></sup> The tiny <a href="/wiki/Northern_cricket_frog" title="Northern cricket frog">northern cricket frog</a> (<i>Acris crepitans</i>) can "skitter" across the surface of a pond with a series of short rapid jumps.<sup id="cite_ref-Badger2_107-0" class="reference"><a href="#cite_note-Badger2-107"><span class="cite-bracket">&#91;</span>107<span class="cite-bracket">&#93;</span></a></sup> </p><p>Slow-motion photography shows that the muscles have passive flexibility. They are first stretched while the frog is still in the crouched position, then they are contracted before being stretched again to launch the frog into the air. The fore legs are folded against the chest and the hind legs remain in the extended, streamlined position for the duration of the jump.<sup id="cite_ref-Frogjump_53-1" class="reference"><a href="#cite_note-Frogjump-53"><span class="cite-bracket">&#91;</span>53<span class="cite-bracket">&#93;</span></a></sup> In some extremely capable jumpers, such as the <a href="/wiki/Cuban_tree_frog" title="Cuban tree frog">Cuban tree frog</a> (<i>Osteopilus septentrionalis</i>) and the <a href="/wiki/Northern_leopard_frog" title="Northern leopard frog">northern leopard frog</a> (<i>Lithobates pipiens</i>), the peak power exerted during a jump can exceed that which the muscle is theoretically capable of producing. When the muscles contract, the energy is first transferred into the stretched tendon which is wrapped around the ankle bone. Then the muscles stretch again at the same time as the tendon releases its energy like a <a href="/wiki/Catapult" title="Catapult">catapult</a> to produce a powerful acceleration beyond the limits of muscle-powered acceleration.<sup id="cite_ref-108" class="reference"><a href="#cite_note-108"><span class="cite-bracket">&#91;</span>108<span class="cite-bracket">&#93;</span></a></sup> A similar mechanism has been documented in <a href="/wiki/Locust" title="Locust">locusts</a> and <a href="/wiki/Grasshopper" title="Grasshopper">grasshoppers</a>.<sup id="cite_ref-109" class="reference"><a href="#cite_note-109"><span class="cite-bracket">&#91;</span>109<span class="cite-bracket">&#93;</span></a></sup> </p><p>Early hatching of froglets can have negative effects on frog jumping performance and overall locomotion.<sup id="cite_ref-:1_110-0" class="reference"><a href="#cite_note-:1-110"><span class="cite-bracket">&#91;</span>110<span class="cite-bracket">&#93;</span></a></sup> The hindlimbs are unable to completely form, which results in them being shorter and much weaker relative to a normal hatching froglet.<sup id="cite_ref-:1_110-1" class="reference"><a href="#cite_note-:1-110"><span class="cite-bracket">&#91;</span>110<span class="cite-bracket">&#93;</span></a></sup> Early hatching froglets may tend to depend on other forms of locomotion more often, such as swimming and walking.<sup id="cite_ref-:1_110-2" class="reference"><a href="#cite_note-:1-110"><span class="cite-bracket">&#91;</span>110<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Walking_and_running">Walking and running</h3></div> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Phrynosequence_bodyFoR2.gif" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/f3/Phrynosequence_bodyFoR2.gif/250px-Phrynosequence_bodyFoR2.gif" decoding="async" width="220" height="142" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/f3/Phrynosequence_bodyFoR2.gif/330px-Phrynosequence_bodyFoR2.gif 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/f3/Phrynosequence_bodyFoR2.gif/500px-Phrynosequence_bodyFoR2.gif 2x" data-file-width="930" data-file-height="600" /></a><figcaption><i><a href="/wiki/Phrynomantis_bifasciatus" class="mw-redirect" title="Phrynomantis bifasciatus">Phrynomantis bifasciatus</a></i> walking on a level surface</figcaption></figure> <p>Frogs in the families Bufonidae, <a href="/wiki/Rhinophrynidae" title="Rhinophrynidae">Rhinophrynidae</a>, and <a href="/wiki/Microhylidae" title="Microhylidae">Microhylidae</a> have short back legs and tend to walk rather than jump.<sup id="cite_ref-Britannica_111-0" class="reference"><a href="#cite_note-Britannica-111"><span class="cite-bracket">&#91;</span>111<span class="cite-bracket">&#93;</span></a></sup> When they try to move rapidly, they speed up the rate of movement of their limbs or resort to an ungainly hopping gait. The <a href="/wiki/Great_Plains_narrow-mouthed_toad" class="mw-redirect" title="Great Plains narrow-mouthed toad">Great Plains narrow-mouthed toad</a> (<i>Gastrophryne olivacea</i>) has been described as having a gait that is "a combination of running and short hops that are usually only an inch or two in length".<sup id="cite_ref-112" class="reference"><a href="#cite_note-112"><span class="cite-bracket">&#91;</span>112<span class="cite-bracket">&#93;</span></a></sup> In an experiment, <a href="/wiki/Bufo_fowleri" class="mw-redirect" title="Bufo fowleri">Fowler's toad</a> (<i>Anaxyrus</i> <i>fowleri</i>) was placed on a treadmill which was turned at varying speeds. By measuring the toad's uptake of oxygen it was found that hopping was an inefficient use of resources during sustained locomotion but was a useful strategy during short bursts of high-intensity activity.<sup id="cite_ref-113" class="reference"><a href="#cite_note-113"><span class="cite-bracket">&#91;</span>113<span class="cite-bracket">&#93;</span></a></sup> </p><p>The <a href="/wiki/Kassina_maculata" class="mw-redirect" title="Kassina maculata">red-legged running frog</a> (<i>Kassina maculata</i>) has short, slim hind limbs unsuited to jumping. It can move fast by using a running gait in which the two hind legs are used alternately. Slow-motion photography shows, unlike a horse that can trot or gallop, the frog's gait remained similar at slow, medium, and fast speeds.<sup id="cite_ref-114" class="reference"><a href="#cite_note-114"><span class="cite-bracket">&#91;</span>114<span class="cite-bracket">&#93;</span></a></sup> This species can also climb trees and shrubs, and does so at night to catch insects.<sup id="cite_ref-115" class="reference"><a href="#cite_note-115"><span class="cite-bracket">&#91;</span>115<span class="cite-bracket">&#93;</span></a></sup> The Indian skipper frog (<i>Euphlyctis cyanophlyctis</i>) has broad feet and can run across the surface of the water for several metres (yards).<sup id="cite_ref-Badger2_107-1" class="reference"><a href="#cite_note-Badger2-107"><span class="cite-bracket">&#91;</span>107<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Swimming">Swimming</h3></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Frog_%C5%BE%C3%A1ba.gif" class="mw-file-description"><img alt="Common toad swimming" src="//upload.wikimedia.org/wikipedia/commons/thumb/b/be/Frog_%C5%BE%C3%A1ba.gif/250px-Frog_%C5%BE%C3%A1ba.gif" decoding="async" width="220" height="179" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/b/be/Frog_%C5%BE%C3%A1ba.gif 1.5x" data-file-width="291" data-file-height="237" /></a><figcaption><a href="/wiki/Common_toad" title="Common toad">Common toad</a> (<i>Bufo bufo</i>) swimming</figcaption></figure> <p>Frogs that live in or visit water have adaptations that improve their swimming abilities. The hind limbs are heavily muscled and strong. The webbing between the toes of the hind feet increases the area of the foot and helps propel the frog powerfully through the water. Members of the family <a href="/wiki/Pipidae" title="Pipidae">Pipidae</a> are wholly aquatic and show the most marked specialisation. They have inflexible vertebral columns, flattened, streamlined bodies, <a href="/wiki/Lateral_line" title="Lateral line">lateral line</a> systems, and powerful hind limbs with large webbed feet.<sup id="cite_ref-Pipidae_116-0" class="reference"><a href="#cite_note-Pipidae-116"><span class="cite-bracket">&#91;</span>116<span class="cite-bracket">&#93;</span></a></sup> Tadpoles mostly have large tail fins which provide thrust when the tail is moved from side to side.<sup id="cite_ref-Duellman_117-0" class="reference"><a href="#cite_note-Duellman-117"><span class="cite-bracket">&#91;</span>117<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Burrowing">Burrowing</h3></div> <p>Some frogs have become adapted for burrowing and a life underground. They tend to have rounded bodies, short limbs, small heads with bulging eyes, and hind feet adapted for excavation. An extreme example of this is the <a href="/wiki/Purple_frog" title="Purple frog">purple frog</a> (<i>Nasikabatrachus sahyadrensis</i>) from southern India which feeds on <a href="/wiki/Termite" title="Termite">termites</a> and spends almost its whole life underground. It emerges briefly during the <a href="/wiki/Monsoon" title="Monsoon">monsoon</a> to mate and breed in temporary pools. It has a tiny head with a pointed snout and a plump, rounded body. Because of this <a href="/wiki/Fossorial" title="Fossorial">fossorial</a> existence, it was <a href="/wiki/Species_description" title="Species description">first described</a> in 2003, being new to the scientific community at that time, although previously known to local people.<sup id="cite_ref-118" class="reference"><a href="#cite_note-118"><span class="cite-bracket">&#91;</span>118<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Nasikabatrachus_sahyadrensis.jpg" class="mw-file-description"><img alt="Purple frog" src="//upload.wikimedia.org/wikipedia/commons/thumb/0/08/Nasikabatrachus_sahyadrensis.jpg/250px-Nasikabatrachus_sahyadrensis.jpg" decoding="async" width="220" height="167" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/08/Nasikabatrachus_sahyadrensis.jpg/330px-Nasikabatrachus_sahyadrensis.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/08/Nasikabatrachus_sahyadrensis.jpg/500px-Nasikabatrachus_sahyadrensis.jpg 2x" data-file-width="1699" data-file-height="1288" /></a><figcaption><a href="/wiki/Purple_frog" title="Purple frog">Purple frog</a> (<i>Nasikabatrachus sahyadrensis</i>)</figcaption></figure> <p>The spadefoot toads of North America are also adapted to underground life. The <a href="/wiki/Plains_spadefoot_toad" title="Plains spadefoot toad">Plains spadefoot toad</a> (<i>Spea bombifrons</i>) is typical and has a flap of keratinised bone attached to one of the <a href="/wiki/Metatarsus" class="mw-redirect" title="Metatarsus">metatarsals</a> of the hind feet which it uses to dig itself backwards into the ground. As it digs, the toad wriggles its hips from side to side to sink into the loose soil. It has a shallow burrow in the summer from which it emerges at night to forage. In winter, it digs much deeper and has been recorded at a depth of 4.5&#160;m (14&#160;ft 9&#160;in).<sup id="cite_ref-Farrar_119-0" class="reference"><a href="#cite_note-Farrar-119"><span class="cite-bracket">&#91;</span>119<span class="cite-bracket">&#93;</span></a></sup> The tunnel is filled with soil and the toad hibernates in a small chamber at the end. During this time, urea accumulates in its tissues and water is drawn in from the surrounding damp soil by <a href="/wiki/Osmosis" title="Osmosis">osmosis</a> to supply the toad's needs.<sup id="cite_ref-Farrar_119-1" class="reference"><a href="#cite_note-Farrar-119"><span class="cite-bracket">&#91;</span>119<span class="cite-bracket">&#93;</span></a></sup> Spadefoot toads are "explosive breeders", all emerging from their burrows at the same time and converging on temporary pools, attracted to one of these by the calling of the first male to find a suitable breeding location.<sup id="cite_ref-120" class="reference"><a href="#cite_note-120"><span class="cite-bracket">&#91;</span>120<span class="cite-bracket">&#93;</span></a></sup> </p><p>The burrowing frogs of Australia have a rather different lifestyle. The <a href="/wiki/Western_spotted_frog" title="Western spotted frog">western spotted frog</a> (<i>Heleioporus albopunctatus</i>) digs a burrow beside a river or in the bed of an ephemeral stream and regularly emerges to forage. Mating takes place and eggs are laid in a foam nest inside the burrow. The eggs partially develop there, but do not hatch until they are submerged following heavy rainfall. The tadpoles then swim out into the open water and rapidly complete their development.<sup id="cite_ref-121" class="reference"><a href="#cite_note-121"><span class="cite-bracket">&#91;</span>121<span class="cite-bracket">&#93;</span></a></sup> Madagascan burrowing frogs are less fossorial and mostly bury themselves in leaf litter. One of these, the <a href="/wiki/Scaphiophryne_marmorata" title="Scaphiophryne marmorata">green burrowing frog</a> (<i>Scaphiophryne marmorata</i>), has a flattened head with a short snout and well-developed metatarsal tubercles on its hind feet to help with excavation. It also has greatly enlarged terminal discs on its fore feet that help it to clamber around in bushes.<sup id="cite_ref-122" class="reference"><a href="#cite_note-122"><span class="cite-bracket">&#91;</span>122<span class="cite-bracket">&#93;</span></a></sup> It breeds in temporary pools that form after rains.<sup id="cite_ref-123" class="reference"><a href="#cite_note-123"><span class="cite-bracket">&#91;</span>123<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Climbing">Climbing</h3></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Phyllomedusa_burmeisteri01.jpg" class="mw-file-description"><img alt="Frog climbing" src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a9/Phyllomedusa_burmeisteri01.jpg/220px-Phyllomedusa_burmeisteri01.jpg" decoding="async" width="220" height="145" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/a9/Phyllomedusa_burmeisteri01.jpg/330px-Phyllomedusa_burmeisteri01.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/a9/Phyllomedusa_burmeisteri01.jpg/440px-Phyllomedusa_burmeisteri01.jpg 2x" data-file-width="769" data-file-height="507" /></a><figcaption><a href="/wiki/Phyllomedusa_burmeisteri" title="Phyllomedusa burmeisteri">Burmeister's leaf frog</a></figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Glass_frogs.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/6/62/Glass_frogs.jpg/220px-Glass_frogs.jpg" decoding="async" width="220" height="147" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/62/Glass_frogs.jpg/330px-Glass_frogs.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/62/Glass_frogs.jpg/440px-Glass_frogs.jpg 2x" data-file-width="4397" data-file-height="2931" /></a><figcaption>Group of <a href="/wiki/Glass_frog" title="Glass frog">glass frogs</a></figcaption></figure> <p>Tree frogs live high in the <a href="/wiki/Canopy_(biology)" title="Canopy (biology)">canopy</a>, where they scramble around on the branches, twigs, and leaves, sometimes never coming down to earth. The "true" tree frogs belong to the family Hylidae, but members of other frog families have independently adopted an arboreal habit, a case of <a href="/wiki/Convergent_evolution" title="Convergent evolution">convergent evolution</a>. These include the <a href="/wiki/Glass_frogs" class="mw-redirect" title="Glass frogs">glass frogs</a> (Centrolenidae), the <a href="/wiki/Hyperoliidae" title="Hyperoliidae">bush frogs</a> (Hyperoliidae), some of the narrow-mouthed frogs (Microhylidae), and the <a href="/wiki/Rhacophoridae" title="Rhacophoridae">shrub frogs</a> (Rhacophoridae).<sup id="cite_ref-Britannica_111-1" class="reference"><a href="#cite_note-Britannica-111"><span class="cite-bracket">&#91;</span>111<span class="cite-bracket">&#93;</span></a></sup> Most tree frogs are under 10&#160;cm (4&#160;in) in length, with long legs and long toes with adhesive pads on the tips. The surface of the toe pads is formed from a closely packed layer of flat-topped, hexagonal <a href="/wiki/Epidermis_(skin)" class="mw-redirect" title="Epidermis (skin)">epidermal</a> cells separated by grooves into which glands secrete <a href="/wiki/Mucus" title="Mucus">mucus</a>. These toe pads, moistened by the mucus, provide the grip on any wet or dry surface, including glass. The forces involved include <a href="/wiki/Boundary_friction" title="Boundary friction">boundary friction</a> of the toe pad epidermis on the surface and also <a href="/wiki/Surface_tension" title="Surface tension">surface tension</a> and <a href="/wiki/Viscosity" title="Viscosity">viscosity</a>.<sup id="cite_ref-124" class="reference"><a href="#cite_note-124"><span class="cite-bracket">&#91;</span>124<span class="cite-bracket">&#93;</span></a></sup> Tree frogs are very acrobatic and can catch insects while hanging by one toe from a twig or clutching onto the blade of a windswept reed.<sup id="cite_ref-125" class="reference"><a href="#cite_note-125"><span class="cite-bracket">&#91;</span>125<span class="cite-bracket">&#93;</span></a></sup> Some members of the subfamily <a href="/wiki/Phyllomedusinae" title="Phyllomedusinae">Phyllomedusinae</a> have <a href="/wiki/Opposable_thumb#Opposition_and_apposition" class="mw-redirect" title="Opposable thumb">opposable toes</a> on their feet. The <a href="/wiki/Phyllomedusa_ayeaye" class="mw-redirect" title="Phyllomedusa ayeaye">reticulated leaf frog</a> (<i>Phyllomedusa ayeaye</i>) has a single opposed <a href="/wiki/Digit_(anatomy)" title="Digit (anatomy)">digit</a> on each fore foot and two opposed digits on its hind feet. This allows it to grasp the stems of bushes as it clambers around in its riverside habitat.<sup id="cite_ref-126" class="reference"><a href="#cite_note-126"><span class="cite-bracket">&#91;</span>126<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Gliding">Gliding</h3></div> <p>During the evolutionary history of frogs, several different groups have independently taken to the air.<sup id="cite_ref-127" class="reference"><a href="#cite_note-127"><span class="cite-bracket">&#91;</span>127<span class="cite-bracket">&#93;</span></a></sup> Some frogs in the tropical rainforest are specially adapted for gliding from tree to tree or parachuting to the forest floor. Typical of them is <a href="/wiki/Rhacophorus_nigropalmatus" class="mw-redirect" title="Rhacophorus nigropalmatus">Wallace's flying frog</a> (<i>Rhacophorus nigropalmatus</i>) from Malaysia and Borneo. It has large feet with the fingertips expanded into flat adhesive discs and the digits fully webbed. Flaps of skin occur on the lateral margins of the limbs and across the tail region. With the digits splayed, the limbs outstretched, and these flaps spread, it can glide considerable distances, but is unable to undertake powered flight.<sup id="cite_ref-128" class="reference"><a href="#cite_note-128"><span class="cite-bracket">&#91;</span>128<span class="cite-bracket">&#93;</span></a></sup> It can alter its direction of travel and navigate distances of up to 15&#160;m (50&#160;ft) between trees.<sup id="cite_ref-129" class="reference"><a href="#cite_note-129"><span class="cite-bracket">&#91;</span>129<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Life_history">Life history</h2></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Greenfrog_life_stages.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/0/09/Greenfrog_life_stages.svg/250px-Greenfrog_life_stages.svg.png" decoding="async" width="220" height="164" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/09/Greenfrog_life_stages.svg/330px-Greenfrog_life_stages.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/09/Greenfrog_life_stages.svg/500px-Greenfrog_life_stages.svg.png 2x" data-file-width="1207" data-file-height="902" /></a><figcaption>Life cycle of the <a href="/wiki/Rana_clamitans" class="mw-redirect" title="Rana clamitans">green frog</a> <br /> (<i>Rana clamitans</i>)</figcaption></figure> <div class="mw-heading mw-heading3"><h3 id="Reproduction">Reproduction</h3></div> <p>Two main types of reproduction occur in frogs, prolonged breeding and explosive breeding. In the former, adopted by the majority of species, adult frogs at certain times of year assemble at a pond, lake or stream to breed. Many frogs return to the bodies of water in which they developed as larvae. This often results in annual migrations involving thousands of individuals. In explosive breeders, mature adult frogs arrive at breeding sites in response to certain trigger factors such as rainfall occurring in an arid area. In these frogs, mating and spawning take place promptly and the speed of larval growth is rapid in order to make use of the ephemeral pools before they dry up.<sup id="cite_ref-Stebbins_130-0" class="reference"><a href="#cite_note-Stebbins-130"><span class="cite-bracket">&#91;</span>130<span class="cite-bracket">&#93;</span></a></sup> </p><p>Among prolonged breeders, males usually arrive at the breeding site first and remain there for some time whereas females tend to arrive later and depart soon after they have spawned. This means that males outnumber females at the water's edge and defend territories from which they expel other males. They advertise their presence by calling, often alternating their croaks with neighbouring frogs. Larger, stronger males tend to have deeper calls and maintain higher quality territories. Females select their mates at least partly on the basis of the depth of their voice.<sup id="cite_ref-131" class="reference"><a href="#cite_note-131"><span class="cite-bracket">&#91;</span>131<span class="cite-bracket">&#93;</span></a></sup> In some species there are satellite males who have no territory and do not call. They may intercept females that are approaching a calling male or take over a vacated territory. Calling is an energy-sapping activity. Sometimes the two roles are reversed and a calling male gives up its territory and becomes a satellite.<sup id="cite_ref-Stebbins_130-1" class="reference"><a href="#cite_note-Stebbins-130"><span class="cite-bracket">&#91;</span>130<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Bufo_bufo_couple_during_migration(2005).jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/f7/Bufo_bufo_couple_during_migration%282005%29.jpg/250px-Bufo_bufo_couple_during_migration%282005%29.jpg" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/f7/Bufo_bufo_couple_during_migration%282005%29.jpg/330px-Bufo_bufo_couple_during_migration%282005%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/f7/Bufo_bufo_couple_during_migration%282005%29.jpg/500px-Bufo_bufo_couple_during_migration%282005%29.jpg 2x" data-file-width="2048" data-file-height="1536" /></a><figcaption>Male and female <a href="/wiki/Common_toad" title="Common toad">common toads</a> (<i>Bufo bufo</i>) in <a href="/wiki/Amplexus" title="Amplexus">amplexus</a></figcaption></figure> <p>In explosive breeders, the first male that finds a suitable breeding location, such as a temporary pool, calls loudly and other frogs of both sexes converge on the pool. Explosive breeders tend to call in unison creating a chorus that can be heard from far away. The spadefoot toads (<i><a href="/wiki/Scaphiopus" title="Scaphiopus">Scaphiopus</a> spp.</i>) of North America fall into this category. Mate selection and courtship is not as important as speed in reproduction. In some years, suitable conditions may not occur and the frogs may go for two or more years without breeding.<sup id="cite_ref-Stebbins_130-2" class="reference"><a href="#cite_note-Stebbins-130"><span class="cite-bracket">&#91;</span>130<span class="cite-bracket">&#93;</span></a></sup> Some female <a href="/wiki/New_Mexico_Spadefoot_Toad" class="mw-redirect" title="New Mexico Spadefoot Toad">New Mexico spadefoot toads</a> (<i>Spea multiplicata</i>) only spawn half of the available eggs at a time, perhaps retaining some in case a better reproductive opportunity arises later.<sup id="cite_ref-132" class="reference"><a href="#cite_note-132"><span class="cite-bracket">&#91;</span>132<span class="cite-bracket">&#93;</span></a></sup> </p><p>At the breeding site, the male mounts the female and grips her tightly round the body. Typically, <a href="/wiki/Amplexus" title="Amplexus">amplexus</a> takes place in the water, the female releases her eggs and the male covers them with sperm; fertilisation is <a href="/wiki/External_fertilization" title="External fertilization">external</a>. In many species such as the <a href="/wiki/Great_Plains_toad" title="Great Plains toad">Great Plains toad</a> (<i>Bufo cognatus</i>), the male restrains the eggs with his back feet, holding them in place for about three minutes.<sup id="cite_ref-Stebbins_130-3" class="reference"><a href="#cite_note-Stebbins-130"><span class="cite-bracket">&#91;</span>130<span class="cite-bracket">&#93;</span></a></sup> Members of the West African genus <i><a href="/wiki/Nimbaphrynoides" title="Nimbaphrynoides">Nimbaphrynoides</a></i> are unique among frogs in that they are <a href="/wiki/Viviparity" title="Viviparity">viviparous</a>; <i><a href="/wiki/Limnonectes_larvaepartus" title="Limnonectes larvaepartus">Limnonectes larvaepartus</a></i>, <i><a href="/wiki/Eleutherodactylus_jasperi" class="mw-redirect" title="Eleutherodactylus jasperi">Eleutherodactylus jasperi</a></i> and members of the <a href="/wiki/Tanzania" title="Tanzania">Tanzanian</a> genus <i><a href="/wiki/Nectophrynoides" title="Nectophrynoides">Nectophrynoides</a></i> are the only frogs known to be <a href="/wiki/Ovoviviparity" title="Ovoviviparity">ovoviviparous</a>. In these species, fertilisation is <a href="/wiki/Internal_fertilization" title="Internal fertilization">internal</a> and females give birth to fully developed juvenile frogs, except <i>L. larvaepartus</i>, which give birth to tadpoles.<sup id="cite_ref-Iskandar2014_133-0" class="reference"><a href="#cite_note-Iskandar2014-133"><span class="cite-bracket">&#91;</span>133<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-134" class="reference"><a href="#cite_note-134"><span class="cite-bracket">&#91;</span>134<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-135" class="reference"><a href="#cite_note-135"><span class="cite-bracket">&#91;</span>135<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Life_cycle">Life cycle</h3></div> <div class="mw-heading mw-heading4"><h4 id="Eggs_/_frogspawn"><span id="Eggs_.2F_frogspawn"></span>Eggs / frogspawn <span class="anchor" id="Frogspawn"></span><span class="anchor" id="Frog_spawn"></span></h4></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Frogspawn_closeup.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/7d/Frogspawn_closeup.jpg/250px-Frogspawn_closeup.jpg" decoding="async" width="220" height="168" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/7d/Frogspawn_closeup.jpg/330px-Frogspawn_closeup.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/7d/Frogspawn_closeup.jpg/500px-Frogspawn_closeup.jpg 2x" data-file-width="666" data-file-height="510" /></a><figcaption>Frogspawn</figcaption></figure> <p>Frogs may lay their eggs as clumps, surface films, strings, or individually. Around half of species deposit eggs in water, others lay eggs in vegetation, on the ground or in excavations.<sup id="cite_ref-136" class="reference"><a href="#cite_note-136"><span class="cite-bracket">&#91;</span>136<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-137" class="reference"><a href="#cite_note-137"><span class="cite-bracket">&#91;</span>137<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-138" class="reference"><a href="#cite_note-138"><span class="cite-bracket">&#91;</span>138<span class="cite-bracket">&#93;</span></a></sup> The tiny <a href="/wiki/Yellow-Striped_Pygmy_Eleuth" class="mw-redirect" title="Yellow-Striped Pygmy Eleuth">yellow-striped pygmy eleuth</a> (<i>Eleutherodactylus limbatus</i>) lays eggs singly, burying them in moist soil.<sup id="cite_ref-139" class="reference"><a href="#cite_note-139"><span class="cite-bracket">&#91;</span>139<span class="cite-bracket">&#93;</span></a></sup> The <a href="/wiki/Smoky_Jungle_Frog" class="mw-redirect" title="Smoky Jungle Frog">smoky jungle frog</a> (<i>Leptodactylus pentadactylus</i>) makes a nest of foam in a hollow. The eggs hatch when the nest is flooded, or the tadpoles may complete their development in the foam if flooding does not occur.<sup id="cite_ref-140" class="reference"><a href="#cite_note-140"><span class="cite-bracket">&#91;</span>140<span class="cite-bracket">&#93;</span></a></sup> The red-eyed treefrog (<i>Agalychnis callidryas</i>) deposits its eggs on a leaf above a pool and when they hatch, the larvae fall into the water below.<sup id="cite_ref-141" class="reference"><a href="#cite_note-141"><span class="cite-bracket">&#91;</span>141<span class="cite-bracket">&#93;</span></a></sup> </p><p>In certain species, such as the <a href="/wiki/Wood_frog" title="Wood frog">wood frog</a> (<i>Rana sylvatica</i>), <a href="/wiki/Symbiosis" title="Symbiosis">symbiotic</a> unicellular green algae are present in the gelatinous material. It is thought that these may benefit the developing larvae by providing them with extra oxygen through <a href="/wiki/Photosynthesis" title="Photosynthesis">photosynthesis</a>.<sup id="cite_ref-142" class="reference"><a href="#cite_note-142"><span class="cite-bracket">&#91;</span>142<span class="cite-bracket">&#93;</span></a></sup> The interior of globular egg clusters of the <a href="/wiki/Wood_Frog" class="mw-redirect" title="Wood Frog">wood frog</a> has also been found to be up to 6&#160;°C (11&#160;°F) warmer than the surrounding water and this speeds up the development of the larvae.<sup id="cite_ref-143" class="reference"><a href="#cite_note-143"><span class="cite-bracket">&#91;</span>143<span class="cite-bracket">&#93;</span></a></sup> The larvae developing in the eggs can detect vibrations caused by nearby predatory wasps or snakes, and will hatch early to avoid being eaten.<sup id="cite_ref-144" class="reference"><a href="#cite_note-144"><span class="cite-bracket">&#91;</span>144<span class="cite-bracket">&#93;</span></a></sup> In general, the length of the egg stage depends on the species and the environmental conditions. Aquatic eggs normally hatch within one week when the capsule splits as a result of <a href="/wiki/Enzyme" title="Enzyme">enzymes</a> released by the developing larvae.<sup id="cite_ref-Stebbins2_145-0" class="reference"><a href="#cite_note-Stebbins2-145"><span class="cite-bracket">&#91;</span>145<span class="cite-bracket">&#93;</span></a></sup> </p><p><b>Direct development</b>, where eggs hatch into juveniles like small adults, is also known in many frogs, for example, <i><a href="/wiki/Ischnocnema_henselii" title="Ischnocnema henselii">Ischnocnema henselii</a>,</i><sup id="cite_ref-146" class="reference"><a href="#cite_note-146"><span class="cite-bracket">&#91;</span>146<span class="cite-bracket">&#93;</span></a></sup> <i><a href="/wiki/Common_coqu%C3%AD" title="Common coquí">Eleutherodactylus coqui</a></i>,<sup id="cite_ref-147" class="reference"><a href="#cite_note-147"><span class="cite-bracket">&#91;</span>147<span class="cite-bracket">&#93;</span></a></sup> and <i><a href="/wiki/Raorchestes_ochlandrae" title="Raorchestes ochlandrae">Raorchestes ochlandrae</a></i> and <i><a href="/wiki/Raorchestes_chalazodes" title="Raorchestes chalazodes">Raorchestes chalazodes</a>.</i><sup id="cite_ref-148" class="reference"><a href="#cite_note-148"><span class="cite-bracket">&#91;</span>148<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Tadpoles">Tadpoles</h4></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Frog_spawn_time-lapse.gif" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8e/Frog_spawn_time-lapse.gif/220px-Frog_spawn_time-lapse.gif" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8e/Frog_spawn_time-lapse.gif/330px-Frog_spawn_time-lapse.gif 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8e/Frog_spawn_time-lapse.gif/440px-Frog_spawn_time-lapse.gif 2x" data-file-width="480" data-file-height="360" /></a><figcaption>Frogspawn development</figcaption></figure> <p>The larvae that emerge from the eggs are known as tadpoles (or occasionally polliwogs). Tadpoles lack eyelids and limbs, and have cartilaginous skeletons, gills for respiration (external gills at first, internal gills later), and tails they use for swimming.<sup id="cite_ref-Duellman_117-1" class="reference"><a href="#cite_note-Duellman-117"><span class="cite-bracket">&#91;</span>117<span class="cite-bracket">&#93;</span></a></sup> As a general rule, free-living larvae are fully aquatic, but at least one species (<i><a href="/wiki/Nannophrys_ceylonensis" title="Nannophrys ceylonensis">Nannophrys ceylonensis</a></i>) has semiterrestrial tadpoles which live among wet rocks.<sup id="cite_ref-149" class="reference"><a href="#cite_note-149"><span class="cite-bracket">&#91;</span>149<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-150" class="reference"><a href="#cite_note-150"><span class="cite-bracket">&#91;</span>150<span class="cite-bracket">&#93;</span></a></sup> </p><p>From early in its development, a gill pouch covers the tadpole's gills and front legs. The lungs soon start to develop and are used as an accessory breathing organ. Some species go through metamorphosis while still inside the egg and hatch directly into small frogs. Tadpoles lack true teeth, but the jaws in most species have two elongated, parallel rows of small, <a href="/wiki/Keratin" title="Keratin">keratinized</a> structures called keradonts in their upper jaws. Their lower jaws usually have three rows of keradonts surrounded by a horny beak, but the number of rows can vary and the exact arrangements of mouth parts provide a means for species identification.<sup id="cite_ref-Stebbins2_145-1" class="reference"><a href="#cite_note-Stebbins2-145"><span class="cite-bracket">&#91;</span>145<span class="cite-bracket">&#93;</span></a></sup> In the Pipidae, with the exception of <i><a href="/wiki/Hymenochirus" class="mw-redirect" title="Hymenochirus">Hymenochirus</a></i>, the tadpoles have paired anterior barbels, which make them resemble small <a href="/wiki/Catfish" title="Catfish">catfish</a>.<sup id="cite_ref-Pipidae_116-1" class="reference"><a href="#cite_note-Pipidae-116"><span class="cite-bracket">&#91;</span>116<span class="cite-bracket">&#93;</span></a></sup> Their tails are stiffened by a <a href="/wiki/Notochord" title="Notochord">notochord</a>, but does not contain any bony or cartilaginous elements except for a few vertebrae at the base which forms the urostyle during metamorphosis. This has been suggested as an adaptation to their lifestyles; because the transformation into frogs happens very fast, the tail is made of soft tissue only, as bone and cartilage take a much longer time to be broken down and absorbed. The tail fin and tip is fragile and will easily tear, which is seen as an adaptation to escape from predators which try to grasp them by the tail.<sup id="cite_ref-151" class="reference"><a href="#cite_note-151"><span class="cite-bracket">&#91;</span>151<span class="cite-bracket">&#93;</span></a></sup> </p><p>Tadpoles are typically <a href="/wiki/Herbivore" title="Herbivore">herbivorous</a>, feeding mostly on <a href="/wiki/Alga" class="mw-redirect" title="Alga">algae</a>, including <a href="/wiki/Diatom" title="Diatom">diatoms</a> filtered from the water through the <a href="/wiki/Gill" title="Gill">gills</a>. Some species are carnivorous at the tadpole stage, eating insects, smaller tadpoles, and fish. The Cuban tree frog (<i>Osteopilus septentrionalis</i>) is one of a number of species in which the tadpoles can be <a href="/wiki/Cannibalism" title="Cannibalism">cannibalistic</a>. Tadpoles that develop legs early may be eaten by the others, so late developers may have better long-term survival prospects.<sup id="cite_ref-152" class="reference"><a href="#cite_note-152"><span class="cite-bracket">&#91;</span>152<span class="cite-bracket">&#93;</span></a></sup> </p><p>Tadpoles are highly vulnerable to being eaten by fish, <a href="/wiki/Newt" title="Newt">newts</a>, predatory <a href="/wiki/Diving_beetle" class="mw-redirect" title="Diving beetle">diving beetles</a>, and birds, particularly <a href="/wiki/Water_bird" title="Water bird">water birds</a>, such as <a href="/wiki/Stork" title="Stork">storks</a> and <a href="/wiki/Heron" title="Heron">herons</a> and <a href="/wiki/Domestic_duck" title="Domestic duck">domestic ducks</a>. Some tadpoles, including those of the <a href="/wiki/Cane_toad" title="Cane toad">cane toad</a> (<i>Rhinella marina</i>), are poisonous. The tadpole stage may be as short as a week in explosive breeders or it may last through one or more winters followed by metamorphosis in the spring.<sup id="cite_ref-153" class="reference"><a href="#cite_note-153"><span class="cite-bracket">&#91;</span>153<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Metamorphosis">Metamorphosis</h4></div> <p>At the end of the tadpole stage, a frog undergoes metamorphosis in which its body makes a sudden transition into the adult form. This metamorphosis typically lasts only 24 hours, and is initiated by production of the <a href="/wiki/Hormone" title="Hormone">hormone</a> <a href="/wiki/Thyroxine" title="Thyroxine">thyroxine</a>. This causes different tissues to develop in different ways. The principal changes that take place include the development of the lungs and the disappearance of the gills and gill pouch, making the front legs visible. The lower jaw transforms into the big mandible of the carnivorous adult, and the long, spiral gut of the herbivorous tadpole is replaced by the typical short gut of a predator.<sup id="cite_ref-Stebbins2_145-2" class="reference"><a href="#cite_note-Stebbins2-145"><span class="cite-bracket">&#91;</span>145<span class="cite-bracket">&#93;</span></a></sup> Homeostatic feedback control of food intake is largely absent, making tadpoles eat constantly when food is present. But shortly before and during metamorphosis the sensation of hunger is suppressed, and they stop eating while their gut and internal organs are reorganised and prepared for a different diet.<sup id="cite_ref-154" class="reference"><a href="#cite_note-154"><span class="cite-bracket">&#91;</span>154<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-155" class="reference"><a href="#cite_note-155"><span class="cite-bracket">&#91;</span>155<span class="cite-bracket">&#93;</span></a></sup> Also the <a href="/wiki/Gut_microbiota" title="Gut microbiota">gut microbiota</a> changes, from being similar to that of fish to resembling that of amniotes.<sup id="cite_ref-156" class="reference"><a href="#cite_note-156"><span class="cite-bracket">&#91;</span>156<span class="cite-bracket">&#93;</span></a></sup> Exceptions are carnivorous tadpoles like <a href="/wiki/Lepidobatrachus_laevis" title="Lepidobatrachus laevis">Lepidobatrachus laevis</a>, which has a gut already adapted to a diet similar to that of adults. These continue to eat during metamorphosis.<sup id="cite_ref-157" class="reference"><a href="#cite_note-157"><span class="cite-bracket">&#91;</span>157<span class="cite-bracket">&#93;</span></a></sup> The nervous system becomes adapted for hearing and stereoscopic vision, and for new methods of locomotion and feeding.<sup id="cite_ref-Stebbins2_145-3" class="reference"><a href="#cite_note-Stebbins2-145"><span class="cite-bracket">&#91;</span>145<span class="cite-bracket">&#93;</span></a></sup> The eyes are repositioned higher up on the head and the eyelids and associated glands are formed. The eardrum, middle ear, and inner ear are developed. The skin becomes thicker and tougher, the lateral line system is lost, and skin glands are developed.<sup id="cite_ref-Stebbins2_145-4" class="reference"><a href="#cite_note-Stebbins2-145"><span class="cite-bracket">&#91;</span>145<span class="cite-bracket">&#93;</span></a></sup> The final stage is the disappearance of the tail, but this takes place rather later, the tissue being used to produce a spurt of growth in the limbs.<sup id="cite_ref-158" class="reference"><a href="#cite_note-158"><span class="cite-bracket">&#91;</span>158<span class="cite-bracket">&#93;</span></a></sup> Frogs are at their most vulnerable to predators when they are undergoing metamorphosis. At this time, the tail is being lost and locomotion by means of limbs is only just becoming established.<sup id="cite_ref-Britannica_111-2" class="reference"><a href="#cite_note-Britannica-111"><span class="cite-bracket">&#91;</span>111<span class="cite-bracket">&#93;</span></a></sup> </p> <style data-mw-deduplicate="TemplateStyles:r1275594942">@media all and (max-width:720px){.mw-parser-output .mod-gallery{width:100%!important}}.mw-parser-output .mod-gallery{display:table}.mw-parser-output .mod-gallery-default{background:transparent;margin-top:4px}.mw-parser-output .mod-gallery-center{margin-left:auto;margin-right:auto}.mw-parser-output .mod-gallery-left{float:left}.mw-parser-output .mod-gallery-right{float:right}.mw-parser-output .mod-gallery-none{float:none}.mw-parser-output .mod-gallery-center .gallery{justify-content:center}.mw-parser-output .mod-gallery-left .gallery{justify-content:left}.mw-parser-output .mod-gallery-right .gallery{justify-content:right}.mw-parser-output .mod-gallery-collapsible{width:100%}.mw-parser-output .mod-gallery .title,.mw-parser-output .mod-gallery .main,.mw-parser-output .mod-gallery .footer{display:table-row}.mw-parser-output .mod-gallery .title>div{display:table-cell;padding:0 4px 4px;text-align:center;font-weight:bold}.mw-parser-output .mod-gallery .main>div{display:table-cell}.mw-parser-output .mod-gallery .gallery.gallery.gallery{line-height:1.35em;display:flex;flex-wrap:wrap;column-gap:4px}.mw-parser-output .mod-gallery .footer>div{display:table-cell;padding:4px;text-align:right;font-size:85%;line-height:1em}.mw-parser-output .mod-gallery .title>div *,.mw-parser-output .mod-gallery .footer>div *{overflow:visible}.mw-parser-output .mod-gallery .gallerybox img{background:none!important}.mw-parser-output .mod-gallery .bordered-images .thumb img{outline:solid var(--background-color-neutral,#eaecf0)1px}.mw-parser-output .mod-gallery .whitebg .thumb{background:var(--background-color-base,#fff)!important}@media screen{html.skin-theme-clientpref-night .mw-parser-output .mod-gallery .bordered-images .thumb img[alt*="\200b \200b \200b "],html.skin-theme-clientpref-night .mw-parser-output .skin-invert-image .mod-gallery .whitebg .thumb.thumb.thumb img{outline:solid #d7d7d7 1px}html.skin-theme-clientpref-night .mw-parser-output .skin-invert-image .mod-gallery .whitebg .thumb.thumb.thumb img{background:none!important}html.skin-theme-clientpref-night .mw-parser-output .mod-gallery .whitebg .thumb img:not([alt*="\200b \200b \200b "]):not([alt*="\200b \200b \200c "]){background:white!important}html.skin-theme-clientpref-night .mw-parser-output .mod-gallery img[alt*="\200b \200b \200b "]{filter:invert(1)hue-rotate(180deg)}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .mod-gallery .bordered-images .thumb img[alt*="\200b \200b \200b "],html.skin-theme-clientpref-os .mw-parser-output .skin-invert-image .mod-gallery .whitebg .thumb.thumb.thumb img{outline:solid #d7d7d7 1px}html.skin-theme-clientpref-os .mw-parser-output .skin-invert-image .mod-gallery .whitebg .thumb.thumb.thumb img{background:none!important}html.skin-theme-clientpref-os .mw-parser-output .mod-gallery .whitebg .thumb img:not([alt*="\200b \200b \200b "]):not([alt*="\200b \200b \200c "]){background:white!important}html.skin-theme-clientpref-os .mw-parser-output .mod-gallery img[alt*="\200b \200b \200b "]{filter:invert(1)hue-rotate(180deg)}}</style><div class="mod-gallery mod-gallery-default mod-gallery-center"><div class="main"><div><ul class="gallery mw-gallery-traditional nochecker bordered-images whitebg"> <li class="gallerybox" style="width: 255px"> <div class="thumb" style="width: 250px; height: 210px;"><span typeof="mw:File"><a href="/wiki/File:RanaTemporariaLarva2.jpg" class="mw-file-description" title="Larva of the common frog Rana temporaria a day before metamorphosis"><img alt="Larva of the common frog Rana temporaria a day before metamorphosis" src="//upload.wikimedia.org/wikipedia/commons/thumb/0/0e/RanaTemporariaLarva2.jpg/250px-RanaTemporariaLarva2.jpg" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/0e/RanaTemporariaLarva2.jpg/330px-RanaTemporariaLarva2.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/0e/RanaTemporariaLarva2.jpg/500px-RanaTemporariaLarva2.jpg 2x" data-file-width="2272" data-file-height="1704" /></a></span></div> <div class="gallerytext">Larva of the common frog <i>Rana temporaria</i> a day before metamorphosis</div> </li> <li class="gallerybox" style="width: 255px"> <div class="thumb" style="width: 250px; height: 210px;"><span typeof="mw:File"><a href="/wiki/File:Rana_Temporaria_-_Larva_Final_Stage.jpg" class="mw-file-description" title="Metamorphosis stage with deforming jaws, large eyes, and remains of gill pouch"><img alt="Metamorphosis stage with deforming jaws, large eyes, and remains of gill pouch" src="//upload.wikimedia.org/wikipedia/commons/thumb/5/5a/Rana_Temporaria_-_Larva_Final_Stage.jpg/250px-Rana_Temporaria_-_Larva_Final_Stage.jpg" decoding="async" width="220" height="178" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/5a/Rana_Temporaria_-_Larva_Final_Stage.jpg/330px-Rana_Temporaria_-_Larva_Final_Stage.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/5a/Rana_Temporaria_-_Larva_Final_Stage.jpg/500px-Rana_Temporaria_-_Larva_Final_Stage.jpg 2x" data-file-width="1443" data-file-height="1168" /></a></span></div> <div class="gallerytext">Metamorphosis stage with deforming jaws, large eyes, and remains of gill pouch</div> </li> <li class="gallerybox" style="width: 255px"> <div class="thumb" style="width: 250px; height: 210px;"><span typeof="mw:File"><a href="/wiki/File:Juvenile_Frog_with_tail_top_view_(1).JPG" class="mw-file-description" title="Young frog with a stumpy tail, metamorphosis nearly complete"><img alt="Young frog with a stumpy tail, metamorphosis nearly complete" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c8/Juvenile_Frog_with_tail_top_view_%281%29.JPG/220px-Juvenile_Frog_with_tail_top_view_%281%29.JPG" decoding="async" width="220" height="162" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c8/Juvenile_Frog_with_tail_top_view_%281%29.JPG/330px-Juvenile_Frog_with_tail_top_view_%281%29.JPG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c8/Juvenile_Frog_with_tail_top_view_%281%29.JPG/440px-Juvenile_Frog_with_tail_top_view_%281%29.JPG 2x" data-file-width="744" data-file-height="547" /></a></span></div> <div class="gallerytext">Young frog with a stumpy tail, metamorphosis nearly complete</div> </li> </ul></div></div></div> <div class="mw-heading mw-heading4"><h4 id="Adults">Adults</h4></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><span><video id="mwe_player_3" poster="//upload.wikimedia.org/wikipedia/commons/thumb/c/ca/Xenopus_laevis_froglet_swimming_-1749-8104-7-13-S10.ogv/220px--Xenopus_laevis_froglet_swimming_-1749-8104-7-13-S10.ogv.jpg" controls="" preload="none" data-mw-tmh="" class="mw-file-element" width="220" height="165" data-durationhint="5" data-mwtitle="Xenopus_laevis_froglet_swimming_-1749-8104-7-13-S10.ogv" data-mwprovider="wikimediacommons" resource="/wiki/File:Xenopus_laevis_froglet_swimming_-1749-8104-7-13-S10.ogv"><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/c/ca/Xenopus_laevis_froglet_swimming_-1749-8104-7-13-S10.ogv/Xenopus_laevis_froglet_swimming_-1749-8104-7-13-S10.ogv.240p.vp9.webm" type="video/webm; codecs=&quot;vp9, opus&quot;" data-transcodekey="240p.vp9.webm" data-width="320" data-height="240" /><source src="//upload.wikimedia.org/wikipedia/commons/c/ca/Xenopus_laevis_froglet_swimming_-1749-8104-7-13-S10.ogv" type="video/ogg; codecs=&quot;theora, vorbis&quot;" data-width="320" data-height="240" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/c/ca/Xenopus_laevis_froglet_swimming_-1749-8104-7-13-S10.ogv/Xenopus_laevis_froglet_swimming_-1749-8104-7-13-S10.ogv.360p.webm" type="video/webm; codecs=&quot;vp8, vorbis&quot;" data-transcodekey="360p.webm" data-width="320" data-height="240" /></video></span><figcaption>A <i><a href="/wiki/Xenopus_laevis" class="mw-redirect" title="Xenopus laevis">Xenopus laevis</a></i> froglet after metamorphosis</figcaption></figure> <p>Adult frogs may live in or near water, but few are fully aquatic.<sup id="cite_ref-159" class="reference"><a href="#cite_note-159"><span class="cite-bracket">&#91;</span>159<span class="cite-bracket">&#93;</span></a></sup> Almost all frog species are <a href="/wiki/Carnivore" title="Carnivore">carnivorous</a> as adults, preying on invertebrates, including <a href="/wiki/Insect" title="Insect">insects</a>, <a href="/wiki/Crab" title="Crab">crabs</a>, <a href="/wiki/Spider" title="Spider">spiders</a>, <a href="/wiki/Mite" title="Mite">mites</a>, <a href="/wiki/Annelid" title="Annelid">worms</a>, <a href="/wiki/Snails" class="mw-redirect" title="Snails">snails</a>, and <a href="/wiki/Slug" title="Slug">slugs</a>. A few of the larger ones may eat other frogs, small <a href="/wiki/Mammal" title="Mammal">mammals</a> and reptiles, and <a href="/wiki/Fish" title="Fish">fish</a>.<sup id="cite_ref-160" class="reference"><a href="#cite_note-160"><span class="cite-bracket">&#91;</span>160<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-161" class="reference"><a href="#cite_note-161"><span class="cite-bracket">&#91;</span>161<span class="cite-bracket">&#93;</span></a></sup> A few species also eat plant matter; the tree frog <i><a href="/wiki/Xenohyla_truncata" title="Xenohyla truncata">Xenohyla truncata</a></i> is partly herbivorous, its diet including a large proportion of fruit, floral structures and nectar.<sup id="cite_ref-162" class="reference"><a href="#cite_note-162"><span class="cite-bracket">&#91;</span>162<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-163" class="reference"><a href="#cite_note-163"><span class="cite-bracket">&#91;</span>163<span class="cite-bracket">&#93;</span></a></sup> <i><a href="/wiki/Leptodactylus_mystaceus" title="Leptodactylus mystaceus">Leptodactylus mystaceus</a></i> has been found to eat plants,<sup id="cite_ref-164" class="reference"><a href="#cite_note-164"><span class="cite-bracket">&#91;</span>164<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-165" class="reference"><a href="#cite_note-165"><span class="cite-bracket">&#91;</span>165<span class="cite-bracket">&#93;</span></a></sup> and <a href="/wiki/Folivory" class="mw-redirect" title="Folivory">folivory</a> occurs in <i><a href="/wiki/Euphlyctis_hexadactylus" title="Euphlyctis hexadactylus">Euphlyctis hexadactylus</a></i>, with plants constituting 79.5% of its diet by volume.<sup id="cite_ref-166" class="reference"><a href="#cite_note-166"><span class="cite-bracket">&#91;</span>166<span class="cite-bracket">&#93;</span></a></sup> Many frogs use their sticky tongues to catch prey, while others simply grab them with their mouths.<sup id="cite_ref-167" class="reference"><a href="#cite_note-167"><span class="cite-bracket">&#91;</span>167<span class="cite-bracket">&#93;</span></a></sup> Adult frogs are themselves attacked by many predators. The <a href="/wiki/Northern_leopard_frog" title="Northern leopard frog">northern leopard frog</a> (<i>Rana pipiens</i>) is eaten by <a href="/wiki/Heron" title="Heron">herons</a>, <a href="/wiki/Hawk" title="Hawk">hawks</a>, fish, large <a href="/wiki/Salamanders" class="mw-redirect" title="Salamanders">salamanders</a>, <a href="/wiki/Snake" title="Snake">snakes</a>, <a href="/wiki/Raccoon" title="Raccoon">raccoons</a>, <a href="/wiki/Skunk" title="Skunk">skunks</a>, <a href="/wiki/Mink" title="Mink">mink</a>, bullfrogs, and other animals.<sup id="cite_ref-168" class="reference"><a href="#cite_note-168"><span class="cite-bracket">&#91;</span>168<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:TrophicWeb.jpg" class="mw-file-description"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/1/13/TrophicWeb.jpg/250px-TrophicWeb.jpg" decoding="async" width="220" height="91" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/13/TrophicWeb.jpg/330px-TrophicWeb.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/13/TrophicWeb.jpg/500px-TrophicWeb.jpg 2x" data-file-width="3260" data-file-height="1350" /></a><figcaption>A trophic pyramid showing frogs as primary predators</figcaption></figure> <p>Frogs are primary predators and an important part of the <a href="/wiki/Food_web" title="Food web">food web</a>. Being <a href="/wiki/Ectotherm" title="Ectotherm">cold-blooded</a>, they make efficient use of the food they eat with little energy being used for metabolic processes, while the rest is transformed into <a href="/wiki/Biomass_(ecology)" title="Biomass (ecology)">biomass</a>. They are themselves eaten by secondary predators and are the primary terrestrial consumers of invertebrates, most of which feed on plants. By reducing herbivory, they play a part in increasing the growth of plants and are thus part of a delicately balanced ecosystem.<sup id="cite_ref-169" class="reference"><a href="#cite_note-169"><span class="cite-bracket">&#91;</span>169<span class="cite-bracket">&#93;</span></a></sup> </p><p>Little is known about the longevity of frogs and toads in the wild, but some can live for many years. <a href="/wiki/Skeletochronology" title="Skeletochronology">Skeletochronology</a> is a method of examining bones to determine age. Using this method, the ages of <a href="/wiki/Mountain_yellow-legged_frog" title="Mountain yellow-legged frog">mountain yellow-legged frogs</a> (<i>Rana muscosa</i>) were studied, the phalanges of the toes showing seasonal lines where growth slows in winter. The oldest frogs had ten bands, so their age was believed to be 14 years, including the four-year tadpole stage.<sup id="cite_ref-170" class="reference"><a href="#cite_note-170"><span class="cite-bracket">&#91;</span>170<span class="cite-bracket">&#93;</span></a></sup> Captive frogs and toads have been recorded as living for up to 40 years, an age achieved by a European common toad (<i>Bufo bufo</i>). The cane toad (<i>Rhinella marina</i>) has been known to survive 24 years in captivity, and the American bullfrog (<i>Rana catesbeiana</i>) 14 years.<sup id="cite_ref-171" class="reference"><a href="#cite_note-171"><span class="cite-bracket">&#91;</span>171<span class="cite-bracket">&#93;</span></a></sup> Frogs from temperate climates hibernate during the winter, and four species are known to be able to withstand freezing during this time, including the wood frog (<i>Rana sylvatica</i>).<sup id="cite_ref-172" class="reference"><a href="#cite_note-172"><span class="cite-bracket">&#91;</span>172<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Parental_care">Parental care</h3></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:AlytesObstetricansMaleWithEggs.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/45/AlytesObstetricansMaleWithEggs.jpg/250px-AlytesObstetricansMaleWithEggs.jpg" decoding="async" width="220" height="169" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/45/AlytesObstetricansMaleWithEggs.jpg/330px-AlytesObstetricansMaleWithEggs.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/45/AlytesObstetricansMaleWithEggs.jpg/500px-AlytesObstetricansMaleWithEggs.jpg 2x" data-file-width="1713" data-file-height="1314" /></a><figcaption>Male <a href="/wiki/Common_midwife_toad" title="Common midwife toad">common midwife toad</a> (<i>Alytes obstetricans</i>) with eggs</figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Assa_darlingtoni.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/ca/Assa_darlingtoni.jpg/250px-Assa_darlingtoni.jpg" decoding="async" width="220" height="133" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/ca/Assa_darlingtoni.jpg/330px-Assa_darlingtoni.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/ca/Assa_darlingtoni.jpg/500px-Assa_darlingtoni.jpg 2x" data-file-width="2284" data-file-height="1384" /></a><figcaption><a href="/wiki/Pouched_frog" title="Pouched frog">Pouched frog</a> (<i>Assa darlingtoni</i>)</figcaption></figure> <p>Although care of offspring is poorly understood in frogs, up to an estimated 20% of amphibian species may care for their young in some way.<sup id="cite_ref-173" class="reference"><a href="#cite_note-173"><span class="cite-bracket">&#91;</span>173<span class="cite-bracket">&#93;</span></a></sup> The <a href="/wiki/Evolution_of_biparental_care_in_tropical_frogs" title="Evolution of biparental care in tropical frogs">evolution of parental care</a> in frogs is driven primarily by the size of the water body in which they breed. Those that breed in smaller water bodies tend to have greater and more complex parental care behaviour.<sup id="cite_ref-Brown1_174-0" class="reference"><a href="#cite_note-Brown1-174"><span class="cite-bracket">&#91;</span>174<span class="cite-bracket">&#93;</span></a></sup> Because predation of eggs and larvae is high in large water bodies, some frog species started to lay their eggs on land. Once this happened, the desiccating terrestrial environment demands that one or both parents keep them moist to ensure their survival.<sup id="cite_ref-175" class="reference"><a href="#cite_note-175"><span class="cite-bracket">&#91;</span>175<span class="cite-bracket">&#93;</span></a></sup> The subsequent need to transport hatched tadpoles to a water body required an even more intense form of parental care.<sup id="cite_ref-Brown1_174-1" class="reference"><a href="#cite_note-Brown1-174"><span class="cite-bracket">&#91;</span>174<span class="cite-bracket">&#93;</span></a></sup> </p><p>In small pools, predators are mostly absent and competition between tadpoles becomes the variable that constrains their survival. Certain frog species avoid this competition by making use of smaller <a href="/wiki/Phytotelma" title="Phytotelma">phytotelmata</a> (water-filled leaf <a href="https://en.wiktionary.org/wiki/axil" class="extiw" title="wikt:axil">axils</a> or small woody cavities) as sites for depositing a few tadpoles.<sup id="cite_ref-176" class="reference"><a href="#cite_note-176"><span class="cite-bracket">&#91;</span>176<span class="cite-bracket">&#93;</span></a></sup> While these smaller rearing sites are free from competition, they also lack sufficient nutrients to support a tadpole without parental assistance. Frog species that changed from the use of larger to smaller phytotelmata have evolved a strategy of providing their offspring with nutritive but unfertilised eggs.<sup id="cite_ref-Brown1_174-2" class="reference"><a href="#cite_note-Brown1-174"><span class="cite-bracket">&#91;</span>174<span class="cite-bracket">&#93;</span></a></sup> The female <a href="/wiki/Strawberry_poison-dart_frog" title="Strawberry poison-dart frog">strawberry poison-dart frog</a> (<i>Oophaga pumilio</i>) lays her eggs on the forest floor. The male frog guards them from predation and carries water in his cloaca to keep them moist. When they hatch, the female moves the tadpoles on her back to a water-holding <a href="/wiki/Bromeliad" class="mw-redirect" title="Bromeliad">bromeliad</a> or other similar water body, depositing just one in each location. She visits them regularly and feeds them by laying one or two unfertilised eggs in the phytotelma, continuing to do this until the young are large enough to undergo metamorphosis.<sup id="cite_ref-177" class="reference"><a href="#cite_note-177"><span class="cite-bracket">&#91;</span>177<span class="cite-bracket">&#93;</span></a></sup> The <a href="/wiki/Granular_poison_frog" title="Granular poison frog">granular poison frog</a> (<i>Oophaga granulifera</i>) looks after its tadpoles in a similar way.<sup id="cite_ref-178" class="reference"><a href="#cite_note-178"><span class="cite-bracket">&#91;</span>178<span class="cite-bracket">&#93;</span></a></sup> </p><p>Many other diverse forms of parental care are seen in frogs. The tiny male <i><a href="/wiki/Colostethus_subpunctatus" class="mw-redirect" title="Colostethus subpunctatus">Colostethus subpunctatus</a></i> stands guard over his egg cluster, laid under a stone or log. When the eggs hatch, he transports the tadpoles on his back to a temporary pool, where he partially immerses himself in the water and one or more tadpoles drop off. He then moves on to another pool.<sup id="cite_ref-179" class="reference"><a href="#cite_note-179"><span class="cite-bracket">&#91;</span>179<span class="cite-bracket">&#93;</span></a></sup> The male <a href="/wiki/Common_midwife_toad" title="Common midwife toad">common midwife toad</a> (<i>Alytes obstetricans</i>) carries the eggs around with him attached to his hind legs. He keeps them damp in dry weather by immersing himself in a pond, and prevents them from getting too wet in soggy vegetation by raising his hindquarters. After three to six weeks, he travels to a pond and the eggs hatch into tadpoles.<sup id="cite_ref-180" class="reference"><a href="#cite_note-180"><span class="cite-bracket">&#91;</span>180<span class="cite-bracket">&#93;</span></a></sup> The <a href="/wiki/Tungara_frog" class="mw-redirect" title="Tungara frog">tungara frog</a> (<i>Physalaemus pustulosus</i>) builds a floating nest from foam to protect its eggs from predation. The foam is made from <a href="/wiki/Protein" title="Protein">proteins</a> and <a href="/wiki/Lectin" title="Lectin">lectins</a>, and seems to have antimicrobial properties.<sup id="cite_ref-181" class="reference"><a href="#cite_note-181"><span class="cite-bracket">&#91;</span>181<span class="cite-bracket">&#93;</span></a></sup> Several pairs of frogs may form a colonial nest on a previously built raft. The eggs are laid in the centre, followed by alternate layers of foam and eggs, finishing with a foam capping.<sup id="cite_ref-182" class="reference"><a href="#cite_note-182"><span class="cite-bracket">&#91;</span>182<span class="cite-bracket">&#93;</span></a></sup> </p><p>Some frogs protect their offspring inside their own bodies. Both male and female <a href="/wiki/Pouched_frog" title="Pouched frog">pouched frogs</a> (<i>Assa darlingtoni</i>) guard their eggs, which are laid on the ground. When the eggs hatch, the male lubricates his body with the jelly surrounding them and immerses himself in the egg mass. The tadpoles wriggle into skin pouches on his side, where they develop until they metamorphose into juvenile frogs.<sup id="cite_ref-183" class="reference"><a href="#cite_note-183"><span class="cite-bracket">&#91;</span>183<span class="cite-bracket">&#93;</span></a></sup> The female <a href="/wiki/Gastric-brooding_frog" title="Gastric-brooding frog">gastric-brooding frog</a> (<i>Rheobatrachus</i> sp.) from <a href="/wiki/Australia" title="Australia">Australia</a>, now probably extinct, swallows her fertilised eggs, which then develop inside her stomach. She ceases to feed and stops secreting <a href="/wiki/Stomach_acid" class="mw-redirect" title="Stomach acid">stomach acid</a>. The tadpoles rely on the yolks of the eggs for nourishment. After six or seven weeks, they are ready for metamorphosis. The mother regurgitates the tiny frogs, which hop away from her mouth.<sup id="cite_ref-184" class="reference"><a href="#cite_note-184"><span class="cite-bracket">&#91;</span>184<span class="cite-bracket">&#93;</span></a></sup> The female <a href="/wiki/Darwin%27s_frog" title="Darwin&#39;s frog">Darwin's frog</a> (<i>Rhinoderma darwinii</i>) from <a href="/wiki/Chile" title="Chile">Chile</a> lays up to 40 eggs on the ground, where they are guarded by the male. When the tadpoles are about to hatch, they are engulfed by the male, which carries them around inside his much-enlarged vocal sac. Here they are immersed in a frothy, viscous liquid that contains some nourishment to supplement what they obtain from the yolks of the eggs. They remain in the sac for seven to ten weeks before undergoing metamorphosis, after which they move into the male's mouth and emerge.<sup id="cite_ref-185" class="reference"><a href="#cite_note-185"><span class="cite-bracket">&#91;</span>185<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Defence">Defence</h2></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:R._imitator_Chazuta.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/6/6e/R._imitator_Chazuta.jpg/220px-R._imitator_Chazuta.jpg" decoding="async" width="220" height="156" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/6e/R._imitator_Chazuta.jpg/330px-R._imitator_Chazuta.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/6e/R._imitator_Chazuta.jpg/440px-R._imitator_Chazuta.jpg 2x" data-file-width="1754" data-file-height="1240" /></a><figcaption>The mildly toxic <i><a href="/wiki/Ranitomeya_imitator" class="mw-redirect" title="Ranitomeya imitator">Ranitomeya imitator</a></i></figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Dendrobates_pumilio.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/ab/Dendrobates_pumilio.jpg/250px-Dendrobates_pumilio.jpg" decoding="async" width="220" height="156" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/ab/Dendrobates_pumilio.jpg/330px-Dendrobates_pumilio.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/ab/Dendrobates_pumilio.jpg/500px-Dendrobates_pumilio.jpg 2x" data-file-width="1000" data-file-height="709" /></a><figcaption><a href="/wiki/Strawberry_poison-dart_frog" title="Strawberry poison-dart frog">Strawberry poison-dart frog</a> contains numerous alkaloids which deter predators.</figcaption></figure> <p>At first sight, frogs seem rather defenceless because of their small size, slow movement, thin skin, and lack of defensive structures, such as spines, claws or teeth. Many use camouflage to avoid detection, the skin often being spotted or streaked in neutral colours that allow a stationary frog to merge into its surroundings. Some can make prodigious leaps, often into water, that help them to evade potential attackers, while many have other defensive adaptations and strategies.<sup id="cite_ref-Stebbins_130-4" class="reference"><a href="#cite_note-Stebbins-130"><span class="cite-bracket">&#91;</span>130<span class="cite-bracket">&#93;</span></a></sup> </p><p>The skin of many frogs contains mild toxic substances called <a href="/wiki/Bufotoxin" title="Bufotoxin">bufotoxins</a> to make them unpalatable to potential predators. Most toads and some frogs have large poison glands, the <a href="/wiki/Parotoid_gland" title="Parotoid gland">parotoid glands</a>, located on the sides of their heads behind the eyes and other glands elsewhere on their bodies. These glands secrete mucus and a range of toxins that make frogs slippery to hold and distasteful or poisonous. If the noxious effect is immediate, the predator may cease its action and the frog may escape. If the effect develops more slowly, the predator may learn to avoid that species in future.<sup id="cite_ref-186" class="reference"><a href="#cite_note-186"><span class="cite-bracket">&#91;</span>186<span class="cite-bracket">&#93;</span></a></sup> Poisonous frogs tend to advertise their toxicity with bright colours, an adaptive strategy known as <a href="/wiki/Aposematism" title="Aposematism">aposematism</a>. The <a href="/wiki/Poison_dart_frog" title="Poison dart frog">poison dart frogs</a> in the family Dendrobatidae do this. They are typically red, orange, or yellow, often with contrasting black markings on their bodies. <i><a href="/wiki/Allobates_zaparo" title="Allobates zaparo">Allobates zaparo</a></i> is not poisonous, but mimics the appearance of two different toxic species with which it shares a common range in an effort to deceive predators.<sup id="cite_ref-187" class="reference"><a href="#cite_note-187"><span class="cite-bracket">&#91;</span>187<span class="cite-bracket">&#93;</span></a></sup> Other species, such as the <a href="/wiki/European_Fire-bellied_Toad" class="mw-redirect" title="European Fire-bellied Toad">European fire-bellied toad</a> (<i>Bombina bombina</i>), have their warning colour underneath. They "flash" this when attacked, adopting a pose that exposes the vivid colouring on their bellies.<sup id="cite_ref-Bb_4-1" class="reference"><a href="#cite_note-Bb-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Bufo_bufo-defensive_reaction1.JPG" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/6/6b/Bufo_bufo-defensive_reaction1.JPG/250px-Bufo_bufo-defensive_reaction1.JPG" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/6b/Bufo_bufo-defensive_reaction1.JPG/330px-Bufo_bufo-defensive_reaction1.JPG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/6b/Bufo_bufo-defensive_reaction1.JPG/500px-Bufo_bufo-defensive_reaction1.JPG 2x" data-file-width="1600" data-file-height="1200" /></a><figcaption>A common toad adopting a defensive stance</figcaption></figure> <p>Some frogs, such as the <a href="/wiki/Poison_dart_frog" title="Poison dart frog">poison dart frogs</a>, are especially toxic. The native peoples of South America extract poison from these frogs to apply to their <a href="/wiki/Dart_(missile)" title="Dart (missile)">weapons</a> for hunting,<sup id="cite_ref-188" class="reference"><a href="#cite_note-188"><span class="cite-bracket">&#91;</span>188<span class="cite-bracket">&#93;</span></a></sup> although few species are toxic enough to be used for this purpose. At least two non-poisonous frog species in tropical America (<i><a href="/wiki/Eleutherodactylus" title="Eleutherodactylus">Eleutherodactylus gaigei</a></i> and <i><a href="/wiki/Lithodytes_lineatus" class="mw-redirect" title="Lithodytes lineatus">Lithodytes lineatus</a></i>) <a href="/wiki/Batesian_mimicry" title="Batesian mimicry">mimic</a> the colouration of dart poison frogs for self-protection.<sup id="cite_ref-189" class="reference"><a href="#cite_note-189"><span class="cite-bracket">&#91;</span>189<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-190" class="reference"><a href="#cite_note-190"><span class="cite-bracket">&#91;</span>190<span class="cite-bracket">&#93;</span></a></sup> Some frogs obtain poisons from the ants and other arthropods they eat.<sup id="cite_ref-191" class="reference"><a href="#cite_note-191"><span class="cite-bracket">&#91;</span>191<span class="cite-bracket">&#93;</span></a></sup> Others, such as the Australian <a href="/wiki/Corroboree_frog" title="Corroboree frog">corroboree frogs</a> (<i>Pseudophryne corroboree</i> and <i>Pseudophryne pengilleyi</i>), can synthesize the <a href="/wiki/Alkaloid" title="Alkaloid">alkaloids</a> themselves.<sup id="cite_ref-192" class="reference"><a href="#cite_note-192"><span class="cite-bracket">&#91;</span>192<span class="cite-bracket">&#93;</span></a></sup> The chemicals involved may be irritants, <a href="/wiki/Hallucinogen" title="Hallucinogen">hallucinogens</a>, <a href="/wiki/Seizure" title="Seizure">convulsants</a>, <a href="/wiki/Neurotoxin" title="Neurotoxin">nerve poisons</a> or <a href="/wiki/Vasoconstrictor" class="mw-redirect" title="Vasoconstrictor">vasoconstrictors</a>. Many predators of frogs have become adapted to tolerate high levels of these poisons, but other creatures, including humans who handle the frogs, may be severely affected.<sup id="cite_ref-193" class="reference"><a href="#cite_note-193"><span class="cite-bracket">&#91;</span>193<span class="cite-bracket">&#93;</span></a></sup> </p><p>Some frogs use bluff or deception. The European common toad (<i>Bufo bufo</i>) adopts a characteristic stance when attacked, inflating its body and standing with its hindquarters raised and its head lowered.<sup id="cite_ref-194" class="reference"><a href="#cite_note-194"><span class="cite-bracket">&#91;</span>194<span class="cite-bracket">&#93;</span></a></sup> The bullfrog (<i>Rana catesbeiana</i>) crouches down with eyes closed and head tipped forward when threatened. This places the parotoid glands in the most effective position, the other glands on its back begin to ooze noxious secretions and the most vulnerable parts of its body are protected.<sup id="cite_ref-Stebbins_130-5" class="reference"><a href="#cite_note-Stebbins-130"><span class="cite-bracket">&#91;</span>130<span class="cite-bracket">&#93;</span></a></sup> Another tactic used by some frogs is to "scream", the sudden loud noise tending to startle the predator. The grey tree frog (<i><a href="/wiki/Hyla_versicolor" class="mw-redirect" title="Hyla versicolor">Hyla versicolor</a></i>) makes an explosive sound that sometimes repels the shrew <i><a href="/wiki/Blarina_brevicauda" class="mw-redirect" title="Blarina brevicauda">Blarina brevicauda</a></i>.<sup id="cite_ref-Stebbins_130-6" class="reference"><a href="#cite_note-Stebbins-130"><span class="cite-bracket">&#91;</span>130<span class="cite-bracket">&#93;</span></a></sup> Although toads are avoided by many predators, the <a href="/wiki/Common_Garter_Snake" class="mw-redirect" title="Common Garter Snake">common garter snake</a> (<i>Thamnophis sirtalis</i>) regularly feeds on them. The strategy employed by juvenile American toads (<i><a href="/wiki/Bufo_americanus" class="mw-redirect" title="Bufo americanus">Bufo americanus</a></i>) on being approached by a snake is to crouch down and remain immobile. This is usually successful, with the snake passing by and the toad remaining undetected. If it is encountered by the snake's head, however, the toad hops away before crouching defensively.<sup id="cite_ref-195" class="reference"><a href="#cite_note-195"><span class="cite-bracket">&#91;</span>195<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Distribution">Distribution</h2></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Wood_Frog_(Rana_sylvatica)_(25234151669).jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/6/68/Wood_Frog_%28Rana_sylvatica%29_%2825234151669%29.jpg/220px-Wood_Frog_%28Rana_sylvatica%29_%2825234151669%29.jpg" decoding="async" width="220" height="148" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/68/Wood_Frog_%28Rana_sylvatica%29_%2825234151669%29.jpg/330px-Wood_Frog_%28Rana_sylvatica%29_%2825234151669%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/68/Wood_Frog_%28Rana_sylvatica%29_%2825234151669%29.jpg/440px-Wood_Frog_%28Rana_sylvatica%29_%2825234151669%29.jpg 2x" data-file-width="4200" data-file-height="2820" /></a><figcaption>Although frogs are most diverse in warm regions, a few species like the <a href="/wiki/Wood_frog" title="Wood frog">wood frog</a> live at the <a href="/wiki/Arctic_Circle" title="Arctic Circle">Arctic Circle</a>.</figcaption></figure> <p>Frogs live on every continent except Antarctica, but they are not present on certain islands, especially those far away from continental land masses.<sup id="cite_ref-196" class="reference"><a href="#cite_note-196"><span class="cite-bracket">&#91;</span>196<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-197" class="reference"><a href="#cite_note-197"><span class="cite-bracket">&#91;</span>197<span class="cite-bracket">&#93;</span></a></sup> Many species are isolated in restricted ranges by changes of climate or inhospitable territory, such as stretches of sea, mountain ridges, deserts, forest clearance, road construction, or other human-made barriers.<sup id="cite_ref-198" class="reference"><a href="#cite_note-198"><span class="cite-bracket">&#91;</span>198<span class="cite-bracket">&#93;</span></a></sup> Usually, a greater diversity of frogs occurs in tropical areas than in temperate regions, such as Europe.<sup id="cite_ref-199" class="reference"><a href="#cite_note-199"><span class="cite-bracket">&#91;</span>199<span class="cite-bracket">&#93;</span></a></sup> Some frogs inhabit arid areas, such as deserts, and rely on specific adaptations to survive. Members of the Australian genus <i><a href="/wiki/Cyclorana" class="mw-redirect" title="Cyclorana">Cyclorana</a></i> bury themselves underground where they create a water-impervious cocoon in which to <a href="/wiki/Aestivation" title="Aestivation">aestivate</a> during dry periods. Once it rains, they emerge, find a temporary pool, and breed. Egg and tadpole development is very fast compared with those of most other frogs, so breeding can be completed before the pond dries up.<sup id="cite_ref-200" class="reference"><a href="#cite_note-200"><span class="cite-bracket">&#91;</span>200<span class="cite-bracket">&#93;</span></a></sup> Some frog species are adapted to a cold environment. The <a href="/wiki/Wood_frog" title="Wood frog">wood frog</a> (<i>Rana sylvatica</i>), whose habitat extends into the <a href="/wiki/Arctic_Circle" title="Arctic Circle">Arctic Circle</a>, buries itself in the ground during winter. Although much of its body freezes during this time, it maintains a high concentration of glucose in its vital organs, which protects them from damage.<sup id="cite_ref-Exploratorium_54-5" class="reference"><a href="#cite_note-Exploratorium-54"><span class="cite-bracket">&#91;</span>54<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Conservation">Conservation</h2></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951" /><div role="note" class="hatnote navigation-not-searchable">See also: <a href="/wiki/Decline_in_amphibian_populations" title="Decline in amphibian populations">Decline in amphibian populations</a></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Bufo_periglenes2.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/5/5b/Bufo_periglenes2.jpg/250px-Bufo_periglenes2.jpg" decoding="async" width="220" height="147" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/5b/Bufo_periglenes2.jpg/330px-Bufo_periglenes2.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/5b/Bufo_periglenes2.jpg/500px-Bufo_periglenes2.jpg 2x" data-file-width="3576" data-file-height="2385" /></a><figcaption><a href="/wiki/Golden_toad" title="Golden toad">Golden toad</a> (<i>Bufo periglenes</i>)&#160;– last seen in 1989</figcaption></figure> <p>In 2006, of 4,035 species of amphibians that depend on water during some lifecycle stage, 1,356 (33.6%) were considered to be threatened. This is likely to be an underestimate because it excludes 1,427 species for which evidence was insufficient to assess their status.<sup id="cite_ref-201" class="reference"><a href="#cite_note-201"><span class="cite-bracket">&#91;</span>201<span class="cite-bracket">&#93;</span></a></sup> Frog populations have declined dramatically since the 1950s. More than one-third of frog species are considered to be threatened with <a href="/wiki/Extinction" title="Extinction">extinction</a>, and more than 120 species are believed to have become extinct since the 1980s.<sup id="cite_ref-Stuart_202-0" class="reference"><a href="#cite_note-Stuart-202"><span class="cite-bracket">&#91;</span>202<span class="cite-bracket">&#93;</span></a></sup> Among these species are the gastric-brooding frogs of Australia and the <a href="/wiki/Golden_toad" title="Golden toad">golden toad</a> of Costa Rica. The latter is of particular concern to scientists because it inhabited the pristine <a href="/wiki/Monteverde_Cloud_Forest_Reserve" title="Monteverde Cloud Forest Reserve">Monteverde Cloud Forest Reserve</a> and its population crashed in 1987, along with about 20 other frog species in the area. This could not be linked directly to human activities, such as deforestation, and was outside the range of normal fluctuations in population size.<sup id="cite_ref-203" class="reference"><a href="#cite_note-203"><span class="cite-bracket">&#91;</span>203<span class="cite-bracket">&#93;</span></a></sup> Elsewhere, habitat loss is a significant cause of frog population decline, as are pollutants, climate change, increased <a href="/wiki/Ultraviolet" title="Ultraviolet">UVB</a> radiation, and the introduction of <a href="/wiki/Indigenous_(ecology)" class="mw-redirect" title="Indigenous (ecology)">non-native</a> predators and competitors.<sup id="cite_ref-204" class="reference"><a href="#cite_note-204"><span class="cite-bracket">&#91;</span>204<span class="cite-bracket">&#93;</span></a></sup> A Canadian study conducted in 2006 suggested heavy traffic in their environment was a larger threat to frog populations than was habitat loss.<sup id="cite_ref-205" class="reference"><a href="#cite_note-205"><span class="cite-bracket">&#91;</span>205<span class="cite-bracket">&#93;</span></a></sup> Emerging infectious diseases, including <a href="/wiki/Chytridiomycosis" title="Chytridiomycosis">chytridiomycosis</a> and <a href="/wiki/Ranavirus" title="Ranavirus">ranavirus</a>, are also devastating populations.<sup id="cite_ref-206" class="reference"><a href="#cite_note-206"><span class="cite-bracket">&#91;</span>206<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-207" class="reference"><a href="#cite_note-207"><span class="cite-bracket">&#91;</span>207<span class="cite-bracket">&#93;</span></a></sup> </p><p>Many environmental scientists believe amphibians, including frogs, are good biological <a href="/wiki/Indicator_species" class="mw-redirect" title="Indicator species">indicators</a> of broader <a href="/wiki/Ecosystem" title="Ecosystem">ecosystem</a> health because of their intermediate positions in food chains, their permeable skins, and typically biphasic lives (aquatic larvae and terrestrial adults).<sup id="cite_ref-208" class="reference"><a href="#cite_note-208"><span class="cite-bracket">&#91;</span>208<span class="cite-bracket">&#93;</span></a></sup> It appears that species with both aquatic eggs and larvae are most affected by the decline, while those with direct development are the most resistant.<sup id="cite_ref-209" class="reference"><a href="#cite_note-209"><span class="cite-bracket">&#91;</span>209<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Deformed_Frog.gif" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/af/Deformed_Frog.gif/250px-Deformed_Frog.gif" decoding="async" width="220" height="210" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/a/af/Deformed_Frog.gif 1.5x" data-file-width="288" data-file-height="275" /></a><figcaption>Deformed <a href="/wiki/Mink_frog" title="Mink frog">mink frog</a> with an extra left leg</figcaption></figure> <p><a href="/wiki/Frog_mutations_and_genetic_defects" class="mw-redirect" title="Frog mutations and genetic defects">Frog mutations and genetic defects</a> have increased since the 1990s. These often include missing legs or extra legs. Various causes have been identified or hypothesized, including an increase in <a href="/wiki/Ultraviolet" title="Ultraviolet">ultraviolet radiation</a> affecting the spawn on the surface of ponds, chemical contamination from pesticides and fertilizers, and parasites such as the <a href="/wiki/Trematode" class="mw-redirect" title="Trematode">trematode</a> <i><a href="/wiki/Ribeiroia_ondatrae" title="Ribeiroia ondatrae">Ribeiroia ondatrae</a></i>. Probably all these are involved in a complex way as <a href="/wiki/Stressor" title="Stressor">stressors</a>, environmental factors contributing to rates of disease, and vulnerability to attack by parasites. Malformations impair mobility and the individuals may not survive to adulthood. An increase in the number of frogs eaten by birds may actually increase the likelihood of parasitism of other frogs, because the trematode's complex lifecycle includes the <a href="/wiki/Planorbidae" title="Planorbidae">ramshorn snail</a> and several intermediate hosts such as birds.<sup id="cite_ref-Blaustein_210-0" class="reference"><a href="#cite_note-Blaustein-210"><span class="cite-bracket">&#91;</span>210<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-211" class="reference"><a href="#cite_note-211"><span class="cite-bracket">&#91;</span>211<span class="cite-bracket">&#93;</span></a></sup> </p><p>In a few cases, captive breeding programs have been established and have largely been successful.<sup id="cite_ref-212" class="reference"><a href="#cite_note-212"><span class="cite-bracket">&#91;</span>212<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-213" class="reference"><a href="#cite_note-213"><span class="cite-bracket">&#91;</span>213<span class="cite-bracket">&#93;</span></a></sup> The <a href="/wiki/World_Association_of_Zoos_and_Aquariums" title="World Association of Zoos and Aquariums">World Association of Zoos and Aquariums</a> named 2008 as the "Year of the Frog" in order to draw attention to the conservation issues faced by them.<sup id="cite_ref-214" class="reference"><a href="#cite_note-214"><span class="cite-bracket">&#91;</span>214<span class="cite-bracket">&#93;</span></a></sup> </p><p>The <a href="/wiki/Cane_toad" title="Cane toad">cane toad</a> (<i>Rhinella marina</i>) is a very adaptable species native to South and Central America. In the 1930s, it was introduced into Puerto Rico, and later various other islands in the Pacific and Caribbean region, as a <a href="/wiki/Biological_pest_control" title="Biological pest control">biological pest control</a> agent.<sup id="cite_ref-215" class="reference"><a href="#cite_note-215"><span class="cite-bracket">&#91;</span>215<span class="cite-bracket">&#93;</span></a></sup> In 1935, 3000 toads were liberated in the <a href="/wiki/Sugar_cane" class="mw-redirect" title="Sugar cane">sugar cane</a> fields of Queensland, Australia, in an attempt to control <a href="/wiki/Cane_beetle" class="mw-redirect" title="Cane beetle">cane beetles</a> such as <i>Dermolepida albohirtum</i>, the larvae of which damage and kill the canes. Initial results in many of these countries were positive, but it later became apparent that the toads upset the ecological balance in their new environments. They bred freely, competed with native frog species, ate bees and other harmless native invertebrates, had few predators in their adopted habitats, and poisoned pets, carnivorous birds, and mammals. In many of these countries, they are now regarded both as pests and <a href="/wiki/Invasive_species" title="Invasive species">invasive species</a>, and scientists are looking for a biological method to control them.<sup id="cite_ref-216" class="reference"><a href="#cite_note-216"><span class="cite-bracket">&#91;</span>216<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Human_uses">Human uses</h2></div> <div class="mw-heading mw-heading3"><h3 id="Culinary">Culinary</h3></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 0;box-sizing:border-box;border:1px solid #aaa;font-size:88%;line-height:1.25em;background-color:var(--background-color-interactive-subtle,#f8f9fa);display:flow-root}.mw-parser-output .side-box-abovebelow,.mw-parser-output .side-box-text{padding:0.25em 0.9em}.mw-parser-output .side-box-image{padding:2px 0 2px 0.9em;text-align:center}.mw-parser-output .side-box-imageright{padding:2px 0.9em 2px 0;text-align:center}@media(min-width:500px){.mw-parser-output .side-box-flex{display:flex;align-items:center}.mw-parser-output .side-box-text{flex:1;min-width:0}}@media(min-width:720px){.mw-parser-output .side-box{width:238px}.mw-parser-output .side-box-right{clear:right;float:right;margin-left:1em}.mw-parser-output .side-box-left{margin-right:1em}}</style><style data-mw-deduplicate="TemplateStyles:r1237033735">@media print{body.ns-0 .mw-parser-output .sistersitebox{display:none!important}}@media screen{html.skin-theme-clientpref-night .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}</style><div class="side-box side-box-right plainlinks sistersitebox"><style data-mw-deduplicate="TemplateStyles:r1126788409">.mw-parser-output .plainlist ol,.mw-parser-output .plainlist ul{line-height:inherit;list-style:none;margin:0;padding:0}.mw-parser-output .plainlist ol li,.mw-parser-output .plainlist ul li{margin-bottom:0}</style> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Wikibooks-logo-en-noslogan.svg" class="mw-file-description"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikibooks-logo-en-noslogan.svg/40px-Wikibooks-logo-en-noslogan.svg.png" decoding="async" width="40" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikibooks-logo-en-noslogan.svg/60px-Wikibooks-logo-en-noslogan.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikibooks-logo-en-noslogan.svg/120px-Wikibooks-logo-en-noslogan.svg.png 2x" data-file-width="400" data-file-height="400" /></a></span></div> <div class="side-box-text plainlist">Wikibooks <a href="https://en.wikibooks.org/wiki/Cookbook" class="extiw" title="wikibooks:Cookbook">Cookbook</a> has a recipe/module on <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409" /><div class="plainlist" style="margin-left: 1.6em;"> <ul><li><i><b><a href="https://en.wikibooks.org/wiki/Cookbook:Frog" class="extiw" title="wikibooks:Cookbook:Frog"> Frog</a></b></i></li></ul> </div></div></div> </div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951" /><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Frog_legs" title="Frog legs">Frog legs</a></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:2012_Froschschenkel_anagoria.JPG" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e4/2012_Froschschenkel_anagoria.JPG/220px-2012_Froschschenkel_anagoria.JPG" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/e4/2012_Froschschenkel_anagoria.JPG/330px-2012_Froschschenkel_anagoria.JPG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/e4/2012_Froschschenkel_anagoria.JPG/440px-2012_Froschschenkel_anagoria.JPG 2x" data-file-width="3648" data-file-height="2736" /></a><figcaption>French <i>cuisses de grenouille</i></figcaption></figure> <p><a href="/wiki/Frog_legs" title="Frog legs">Frog legs</a> are eaten by humans in many parts of the world. Indonesia is the world's largest exporter of frog meat, exporting more than 5,000 tonnes of frog meat each year, mostly to France, Belgium and Luxembourg.<sup id="cite_ref-abc_news_217-0" class="reference"><a href="#cite_note-abc_news-217"><span class="cite-bracket">&#91;</span>217<span class="cite-bracket">&#93;</span></a></sup> Originally, they were supplied from local wild populations, but overexploitation led to a diminution in the supply. This resulted in the development of <a href="/wiki/Aquaculture" title="Aquaculture">frog farming</a> and a global trade in frogs. The main importing countries are France, Belgium, Luxembourg, and the United States, while the chief exporting nations are Indonesia and China.<sup id="cite_ref-Warkentin_218-0" class="reference"><a href="#cite_note-Warkentin-218"><span class="cite-bracket">&#91;</span>218<span class="cite-bracket">&#93;</span></a></sup> The annual global trade in the <a href="/wiki/American_bullfrog" title="American bullfrog">American bullfrog</a> (<i>Rana catesbeiana</i>), mostly farmed in China, varies between 1200 and 2400 tonnes.<sup id="cite_ref-219" class="reference"><a href="#cite_note-219"><span class="cite-bracket">&#91;</span>219<span class="cite-bracket">&#93;</span></a></sup> </p><p>The <a href="/wiki/Leptodactylus_fallax" title="Leptodactylus fallax">mountain chicken frog</a>, so-called as it tastes of chicken, is now endangered, in part due to human consumption, and was a major food choice of the <a href="/wiki/Dominica" title="Dominica">Dominicans</a>.<sup id="cite_ref-220" class="reference"><a href="#cite_note-220"><span class="cite-bracket">&#91;</span>220<span class="cite-bracket">&#93;</span></a></sup> <a href="/wiki/Raccoon" title="Raccoon">Raccoon</a>, <a href="/wiki/Virginia_opossum" title="Virginia opossum">opossum</a>, <a href="/wiki/Partridges" class="mw-redirect" title="Partridges">partridges</a>, <a href="/wiki/Greater_prairie_chicken" class="mw-redirect" title="Greater prairie chicken">prairie chicken</a>, and frogs were among the fare <a href="/wiki/Mark_Twain" title="Mark Twain">Mark Twain</a> recorded as part of American cuisine.<sup id="cite_ref-TwainWarner1904_221-0" class="reference"><a href="#cite_note-TwainWarner1904-221"><span class="cite-bracket">&#91;</span>221<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Scientific_research">Scientific research</h3></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951" /><div role="note" class="hatnote navigation-not-searchable">See also: <a href="/wiki/Animal_testing_on_frogs" title="Animal testing on frogs">Animal testing on frogs</a> and <a href="/wiki/Frogs_in_captivity" title="Frogs in captivity">Frogs in captivity</a></div> <p>In November 1970, <a href="/wiki/NASA" title="NASA">NASA</a> sent two bullfrogs into space for six days during the <a href="/wiki/Orbiting_Frog_Otolith" title="Orbiting Frog Otolith">Orbiting Frog Otolith</a> mission to test weightlessness. </p><p>Frogs are used for <a href="/wiki/Dissection" title="Dissection">dissections</a> in high school and university anatomy classes, often first being injected with coloured substances to enhance contrasts among the <a href="/wiki/Biological_system" title="Biological system">biological systems</a>. This practice is declining due to <a href="/wiki/Animal_welfare" title="Animal welfare">animal welfare</a> concerns, and "digital frogs" are now available for virtual dissection.<sup id="cite_ref-222" class="reference"><a href="#cite_note-222"><span class="cite-bracket">&#91;</span>222<span class="cite-bracket">&#93;</span></a></sup> </p><p>Frogs have served as <a href="/wiki/Animal_testing" title="Animal testing">experimental animals</a> throughout the history of science. Eighteenth-century biologist <a href="/wiki/Luigi_Galvani" title="Luigi Galvani">Luigi Galvani</a> discovered the link between <a href="/wiki/Electricity" title="Electricity">electricity</a> and the <a href="/wiki/Nervous_system" title="Nervous system">nervous system</a> by studying frogs. He created <a href="/wiki/Frog_galvanometer" class="mw-redirect" title="Frog galvanometer">one of the first tools</a> for measuring <a href="/wiki/Electric_current" title="Electric current">electric current</a> out of a frog leg.<sup id="cite_ref-223" class="reference"><a href="#cite_note-223"><span class="cite-bracket">&#91;</span>223<span class="cite-bracket">&#93;</span></a></sup> In 1852, H. F. Stannius used a frog's heart in a procedure called a <a href="/wiki/Stannius_ligature" title="Stannius ligature">Stannius ligature</a> to demonstrate the ventricle and atria beat independently of each other and at different rates.<sup id="cite_ref-224" class="reference"><a href="#cite_note-224"><span class="cite-bracket">&#91;</span>224<span class="cite-bracket">&#93;</span></a></sup> The <a href="/wiki/African_clawed_frog" title="African clawed frog">African clawed frog</a> or platanna (<i>Xenopus laevis</i>) was first widely used in laboratories in pregnancy tests in the first half of the 20th century. A sample of urine from a pregnant woman injected into a female frog induces it to lay <a href="/wiki/Egg" title="Egg">eggs</a>, a discovery made by English zoologist <a href="/wiki/Lancelot_Hogben" title="Lancelot Hogben">Lancelot Hogben</a>. This is because a hormone, <a href="/wiki/Human_chorionic_gonadotropin" title="Human chorionic gonadotropin">human chorionic gonadotropin</a>, is present in substantial quantities in the urine of women during pregnancy.<sup id="cite_ref-225" class="reference"><a href="#cite_note-225"><span class="cite-bracket">&#91;</span>225<span class="cite-bracket">&#93;</span></a></sup> In 1952, <a href="/wiki/Robert_William_Briggs" class="mw-redirect" title="Robert William Briggs">Robert Briggs</a> and <a href="/wiki/Thomas_J._King" class="mw-redirect" title="Thomas J. King">Thomas J. King</a> cloned a frog by <a href="/wiki/Somatic_cell_nuclear_transfer" title="Somatic cell nuclear transfer">somatic cell nuclear transfer</a>. This same technique was later used to create <a href="/wiki/Dolly_(sheep)" title="Dolly (sheep)">Dolly the sheep</a>, and their experiment was the first time a successful nuclear transplantation had been accomplished in higher animals.<sup id="cite_ref-226" class="reference"><a href="#cite_note-226"><span class="cite-bracket">&#91;</span>226<span class="cite-bracket">&#93;</span></a></sup> </p><p>Frogs are used in cloning research and other branches of <a href="/wiki/Embryology" title="Embryology">embryology</a>. Although alternative pregnancy tests have been developed, biologists continue to use <i>Xenopus</i> as a <a href="/wiki/Model_organism" title="Model organism">model organism</a> in <a href="/wiki/Developmental_biology" title="Developmental biology">developmental biology</a> because their embryos are large and easy to manipulate, they are readily obtainable, and can easily be kept in the laboratory.<sup id="cite_ref-227" class="reference"><a href="#cite_note-227"><span class="cite-bracket">&#91;</span>227<span class="cite-bracket">&#93;</span></a></sup> <i>Xenopus laevis</i> is increasingly being displaced by its smaller relative, <i><a href="/wiki/Xenopus_tropicalis" class="mw-redirect" title="Xenopus tropicalis">Xenopus tropicalis</a></i>, which reaches its reproductive age in five months rather than the one to two years for <i>X. laevis</i>,<sup id="cite_ref-228" class="reference"><a href="#cite_note-228"><span class="cite-bracket">&#91;</span>228<span class="cite-bracket">&#93;</span></a></sup> thus facilitating faster studies across generations. </p><p>Genomes of <i>Xenopus laevis</i>, <i>X. tropicalis</i>, <i>Rana catesbeiana</i>, <i>Rhinella marina</i>, and <i>Nanorana parkeri</i> have been sequenced and deposited in the <a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a> Genome database.<sup id="cite_ref-229" class="reference"><a href="#cite_note-229"><span class="cite-bracket">&#91;</span>229<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Pharmaceutical">Pharmaceutical</h3></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Schrecklicherpfeilgiftfrosch-01.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/6/6e/Schrecklicherpfeilgiftfrosch-01.jpg/250px-Schrecklicherpfeilgiftfrosch-01.jpg" decoding="async" width="170" height="130" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/6e/Schrecklicherpfeilgiftfrosch-01.jpg/330px-Schrecklicherpfeilgiftfrosch-01.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/6e/Schrecklicherpfeilgiftfrosch-01.jpg/500px-Schrecklicherpfeilgiftfrosch-01.jpg 2x" data-file-width="895" data-file-height="683" /></a><figcaption><a href="/wiki/Golden_poison_frog" title="Golden poison frog">Golden poison frog</a> (<i>Phyllobates terribilis</i>)</figcaption></figure> <p>Because frog toxins are extraordinarily diverse, they have raised the interest of biochemists as a "natural pharmacy". The alkaloid <a href="/wiki/Epibatidine" title="Epibatidine">epibatidine</a>, a painkiller 200 times more potent than <a href="/wiki/Morphine" title="Morphine">morphine</a>, is made by some species of <a href="/wiki/Poison_dart_frog" title="Poison dart frog">poison dart frogs</a>. Other chemicals isolated from the skins of frogs may offer resistance to <a href="/wiki/HIV" title="HIV">HIV</a> infection.<sup id="cite_ref-230" class="reference"><a href="#cite_note-230"><span class="cite-bracket">&#91;</span>230<span class="cite-bracket">&#93;</span></a></sup> Dart poisons are under active investigation for their potential as therapeutic drugs.<sup id="cite_ref-231" class="reference"><a href="#cite_note-231"><span class="cite-bracket">&#91;</span>231<span class="cite-bracket">&#93;</span></a></sup> </p><p>It has long been suspected that pre-Columbian <a href="/wiki/Mesoamerica" title="Mesoamerica">Mesoamericans</a> used a toxic secretion produced by the cane toad as a <a href="/wiki/Hallucinogen" title="Hallucinogen">hallucinogen</a>, but more likely they used substances secreted by the <a href="/wiki/Colorado_River_toad" title="Colorado River toad">Colorado River toad</a> (<i>Bufo alvarius</i>). These contain <a href="/wiki/Bufotenin" title="Bufotenin">bufotenin</a> (5-MeO-DMT), a <a href="/wiki/Psychoactive_drug" title="Psychoactive drug">psychoactive compound</a> that has been used in modern times as a <a href="/wiki/Recreational_drug" class="mw-redirect" title="Recreational drug">recreational drug</a>. Typically, the skin secretions are dried and then smoked.<sup id="cite_ref-232" class="reference"><a href="#cite_note-232"><span class="cite-bracket">&#91;</span>232<span class="cite-bracket">&#93;</span></a></sup> Illicit drug use by licking the skin of a toad has been reported in the media, but this may be an <a href="/wiki/Urban_myth" class="mw-redirect" title="Urban myth">urban myth</a>.<sup id="cite_ref-233" class="reference"><a href="#cite_note-233"><span class="cite-bracket">&#91;</span>233<span class="cite-bracket">&#93;</span></a></sup> </p><p>Exudations from the skin of the <a href="/wiki/Golden_poison_frog" title="Golden poison frog">golden poison frog</a> (<i>Phyllobates terribilis</i>) are traditionally used by native Colombians to poison the darts they use for hunting. The tip of the projectile is rubbed over the back of the frog and the dart is launched from a <a href="/wiki/Blowgun" title="Blowgun">blowgun</a>. The combination of the two alkaloid toxins <a href="/wiki/Batrachotoxin" title="Batrachotoxin">batrachotoxin</a> and <a href="/wiki/Homobatrachotoxin" class="mw-redirect" title="Homobatrachotoxin">homobatrachotoxin</a> is so powerful, one frog contains enough poison to kill an estimated 22,000 mice.<sup id="cite_ref-dartpoison_234-0" class="reference"><a href="#cite_note-dartpoison-234"><span class="cite-bracket">&#91;</span>234<span class="cite-bracket">&#93;</span></a></sup> Two other species, the <a href="/wiki/Kokoe_poison_dart_frog" class="mw-redirect" title="Kokoe poison dart frog">Kokoe poison dart frog</a> (<i>Phyllobates aurotaenia</i>) and the <a href="/wiki/Black-legged_dart_frog" class="mw-redirect" title="Black-legged dart frog">black-legged dart frog</a> (<i>Phyllobates bicolor</i>) are also used for this purpose. These are less toxic and less abundant than the golden poison frog. They are impaled on pointed sticks and may be heated over a fire to maximise the quantity of poison that can be transferred to the dart.<sup id="cite_ref-dartpoison_234-1" class="reference"><a href="#cite_note-dartpoison-234"><span class="cite-bracket">&#91;</span>234<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Toadlarcomuseum.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/75/Toadlarcomuseum.jpg/170px-Toadlarcomuseum.jpg" decoding="async" width="170" height="175" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/7/75/Toadlarcomuseum.jpg 1.5x" data-file-width="243" data-file-height="250" /></a><figcaption><a href="/wiki/Moche_culture" title="Moche culture">Moche</a> frog sculpture</figcaption></figure> <div class="mw-heading mw-heading3"><h3 id="Cultural_significance">Cultural significance</h3></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951" /><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Frogs_in_culture" title="Frogs in culture">Frogs in culture</a></div> <p>Frogs have been featured in mythology, <a href="/wiki/Fairy_tale" title="Fairy tale">fairy tales</a> and popular culture. In traditional Chinese myths, the world rests on a giant frog, who would try to swallow the moon, causing the <a href="/wiki/Lunar_eclipse" title="Lunar eclipse">lunar eclipse</a>. Frogs have been featured in religion, folklore, and popular culture. The <a href="/wiki/Ancient_Egyptian" class="mw-redirect" title="Ancient Egyptian">ancient Egyptians</a> depicted the god <a href="/wiki/Heqet" title="Heqet">Heqet</a>, protector of newborns, with the head of a frog. For the <a href="/wiki/Mayan_civilization" class="mw-redirect" title="Mayan civilization">Mayans</a>, frogs represented water, crops, fertility and birth and were associated with the god <a href="/wiki/Chaac" title="Chaac">Chaac</a>. In the <a href="/wiki/Bible" title="Bible">Bible</a>, <a href="/wiki/Moses" title="Moses">Moses</a> unleashes a <a href="/wiki/Plagues_of_Egypt" title="Plagues of Egypt">plague of frogs</a> on the Egyptians. Medieval Europeans associated frogs and toads with evil and <a href="/wiki/Witchcraft" title="Witchcraft">witchcraft</a>.<sup id="cite_ref-culture_235-0" class="reference"><a href="#cite_note-culture-235"><span class="cite-bracket">&#91;</span>235<span class="cite-bracket">&#93;</span></a></sup> The <a href="/wiki/Brothers_Grimm" title="Brothers Grimm">Brothers Grimm</a> fairy tale <i><a href="/wiki/The_Frog_Prince" title="The Frog Prince">The Frog Prince</a></i> features a princess taking in a frog and it turning into a handsome prince.<sup id="cite_ref-236" class="reference"><a href="#cite_note-236"><span class="cite-bracket">&#91;</span>236<span class="cite-bracket">&#93;</span></a></sup> In modern culture, frogs may take a comedic or hapless role, such as <a href="/wiki/Mr._Toad" title="Mr. Toad">Mr. Toad</a> of the 1908 novel <i><a href="/wiki/The_Wind_in_the_Willows" title="The Wind in the Willows">The Wind in the Willows</a></i>, <a href="/wiki/Michigan_J._Frog" title="Michigan J. Frog">Michigan J. Frog</a> of <a href="/wiki/Warner_Bros._Cartoons" title="Warner Bros. Cartoons">Warner Bros. Cartoons</a>, the <a href="/wiki/Muppet" class="mw-redirect" title="Muppet">Muppet</a> <a href="/wiki/Kermit_the_Frog" title="Kermit the Frog">Kermit the Frog</a> and in the game <i><a href="/wiki/Frogger" title="Frogger">Frogger</a></i>.<sup id="cite_ref-237" class="reference"><a href="#cite_note-237"><span class="cite-bracket">&#91;</span>237<span class="cite-bracket">&#93;</span></a></sup> </p> <div style="clear:both;" class=""></div> <div class="mw-heading mw-heading2"><h2 id="References">References</h2></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width" style="column-width: 30em;"> <ol class="references"> <li id="cite_note-AOTW-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-AOTW_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-AOTW_1-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFFrost2021" class="citation web cs1">Frost, Darrel R. (2021) [1998]. <a rel="nofollow" class="external text" href="https://amphibiansoftheworld.amnh.org/Amphibia/Anura">"Anura"</a>. <i>Amphibian Species of the World</i>. American Museum of Natural History<span class="reference-accessdate">. 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(1997). <a rel="nofollow" class="external text" href="http://tolweb.org/Salientia/">"Salientia"</a>. <i>Tree of Life Web Project</i><span class="reference-accessdate">. Retrieved <span class="nowrap">August 7,</span> 2012</span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=unknown&amp;rft.jtitle=Tree+of+Life+Web+Project&amp;rft.atitle=Salientia&amp;rft.date=1997&amp;rft.au=Cannatella%2C+David+C.&amp;rft_id=http%3A%2F%2Ftolweb.org%2FSalientia%2F&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AFrog" class="Z3988"></span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-3">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFBadgerNetherton,_J.1995" class="citation book cs1">Badger, D.; Netherton, J. (1995). <i>Frogs</i>. 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font-size:95%; color: var( --color-subtle, #555 )">(Subscription or <a rel="nofollow" class="external text" href="https://www.oed.com/public/login/loggingin#withyourlibrary">participating institution membership</a> required.)</span></span> </li> <li id="cite_note-:0-11"><span class="mw-cite-backlink">^ <a href="#cite_ref-:0_11-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-:0_11-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-:0_11-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-:0_11-3"><sup><i><b>d</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFReference-OED-frog,_n.1_and_adj." class="citation encyclopaedia cs1"><span class="id-lock-subscription" title="Paid subscription required"><a rel="nofollow" class="external text" href="https://www.oed.com/search/dictionary/?q=frog%2C+n.1+and+adj.">"frog, n.1 and adj."</a></span>. <i><a href="/wiki/Oxford_English_Dictionary" title="Oxford English Dictionary">Oxford English Dictionary</a></i> (Online&#160;ed.). <a href="/wiki/Oxford_University_Press" title="Oxford University Press">Oxford University Press</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=bookitem&amp;rft.atitle=frog%2C+n.1+and+adj.&amp;rft.btitle=Oxford+English+Dictionary&amp;rft.edition=Online&amp;rft.pub=Oxford+University+Press&amp;rft_id=https%3A%2F%2Fwww.oed.com%2Fsearch%2Fdictionary%2F%3Fq%3Dfrog%252C%2Bn.1%2Band%2Badj.&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AFrog" class="Z3988"></span>&#32;<span style="font-size:0.95em; font-size:95%; color: var( --color-subtle, #555 )">(Subscription or <a rel="nofollow" class="external text" href="https://www.oed.com/public/login/loggingin#withyourlibrary">participating institution membership</a> required.)</span></span> </li> <li id="cite_note-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-12">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFReference-OED-frosh_&#124;_frosk,_n.1." class="citation encyclopaedia cs1"><span class="id-lock-subscription" title="Paid subscription required"><a rel="nofollow" class="external text" href="https://www.oed.com/search/dictionary/?q=frosh+%7C+frosk%2C+n.1.">"frosh | frosk, n.1."</a></span>. <i><a href="/wiki/Oxford_English_Dictionary" title="Oxford English Dictionary">Oxford English Dictionary</a></i> (Online&#160;ed.). <a href="/wiki/Oxford_University_Press" title="Oxford University Press">Oxford University Press</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=bookitem&amp;rft.atitle=frosh+%7C+frosk%2C+n.1.&amp;rft.btitle=Oxford+English+Dictionary&amp;rft.edition=Online&amp;rft.pub=Oxford+University+Press&amp;rft_id=https%3A%2F%2Fwww.oed.com%2Fsearch%2Fdictionary%2F%3Fq%3Dfrosh%2B%257C%2Bfrosk%252C%2Bn.1.&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AFrog" class="Z3988"></span>&#32;<span style="font-size:0.95em; font-size:95%; color: var( --color-subtle, #555 )">(Subscription or <a rel="nofollow" class="external text" href="https://www.oed.com/public/login/loggingin#withyourlibrary">participating institution membership</a> required.)</span></span> </li> <li id="cite_note-13"><span class="mw-cite-backlink"><b><a href="#cite_ref-13">^</a></b></span> <span class="reference-text">Jerzy Wełna, '<a rel="nofollow" class="external text" href="https://repozytorium.amu.edu.pl/bitstream/10593/18454/1/31welna.pdf">Metathetic and Non-Metathetic Form Selection in Middle English</a>', <i>Studia Anglica Posnaniensia</i>, 30 (2002), 501–18 (p. 504).</span> </li> <li id="cite_note-14"><span class="mw-cite-backlink"><b><a href="#cite_ref-14">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFReference-OED-toad,_n." class="citation encyclopaedia cs1"><span class="id-lock-subscription" title="Paid subscription required"><a rel="nofollow" class="external text" href="https://www.oed.com/search/dictionary/?q=toad%2C+n.">"toad, n."</a></span>. <i><a href="/wiki/Oxford_English_Dictionary" title="Oxford English Dictionary">Oxford English Dictionary</a></i> (Online&#160;ed.). <a href="/wiki/Oxford_University_Press" title="Oxford University Press">Oxford University Press</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=bookitem&amp;rft.atitle=toad%2C+n.&amp;rft.btitle=Oxford+English+Dictionary&amp;rft.edition=Online&amp;rft.pub=Oxford+University+Press&amp;rft_id=https%3A%2F%2Fwww.oed.com%2Fsearch%2Fdictionary%2F%3Fq%3Dtoad%252C%2Bn.&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AFrog" class="Z3988"></span>&#32;<span style="font-size:0.95em; 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(2010). <a rel="nofollow" class="external text" href="http://www.nimbadarwin.org/resources/nimbatoad.pdf">"Rediscovery of the Liberian Nimba toad, <i>Nimbaphrynoides liberiensis</i> (Xavier, 1978) (Amphibia: Anura: Bufonidae), and reassessment of its taxonomic status"</a> <span class="cs1-format">(PDF)</span>. <i>Zootaxa</i>. <b>4355</b>: <span class="nowrap">56–</span>68. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.11646%2Fzootaxa.2355.1.3">10.11646/zootaxa.2355.1.3</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a>&#160;<a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/1175-5334">1175-5334</a>. <a rel="nofollow" class="external text" href="https://ghostarchive.org/archive/20221009/http://www.nimbadarwin.org/resources/nimbatoad.pdf">Archived</a> <span class="cs1-format">(PDF)</span> from the original on October 9, 2022.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Zootaxa&amp;rft.atitle=Rediscovery+of+the+Liberian+Nimba+toad%2C+Nimbaphrynoides+liberiensis+%28Xavier%2C+1978%29+%28Amphibia%3A+Anura%3A+Bufonidae%29%2C+and+reassessment+of+its+taxonomic+status&amp;rft.volume=4355&amp;rft.pages=%3Cspan+class%3D%22nowrap%22%3E56-%3C%2Fspan%3E68&amp;rft.date=2010&amp;rft_id=info%3Adoi%2F10.11646%2Fzootaxa.2355.1.3&amp;rft.issn=1175-5334&amp;rft.au=Sandberger%2C+L.&amp;rft.au=Hillers%2C+A.&amp;rft.au=Doumbia%2C+J.&amp;rft.au=Loua%2C+N-S.&amp;rft.au=Brede+C.&amp;rft.au=R%C3%B6del%2C+M-O.&amp;rft_id=http%3A%2F%2Fwww.nimbadarwin.org%2Fresources%2Fnimbatoad.pdf&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AFrog" class="Z3988"></span></span> </li> <li id="cite_note-136"><span class="mw-cite-backlink"><b><a href="#cite_ref-136">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFMattison,_Chris2011" class="citation book cs1">Mattison, Chris (2011). <i>Frogs and Toads of the Worlds</i>. Princeton University Press. p.&#160;91. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/978-0-691-14968-4" title="Special:BookSources/978-0-691-14968-4"><bdi>978-0-691-14968-4</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=book&amp;rft.btitle=Frogs+and+Toads+of+the+Worlds&amp;rft.pages=91&amp;rft.pub=Princeton+University+Press&amp;rft.date=2011&amp;rft.isbn=978-0-691-14968-4&amp;rft.au=Mattison%2C+Chris&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AFrog" class="Z3988"></span></span> </li> <li id="cite_note-137"><span class="mw-cite-backlink"><b><a href="#cite_ref-137">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFDuellman,_W._E.Zug,_G._R." class="citation encyclopaedia cs1">Duellman, W. E.; Zug, G. 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"Misuse and legend in the "toad licking" phenomenon". <i>International Journal of the Addictions</i>. <b>28</b> (6): <span class="nowrap">521–</span>538. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.3109%2F10826089309039645">10.3109/10826089309039645</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/8486435">8486435</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=International+Journal+of+the+Addictions&amp;rft.atitle=Misuse+and+legend+in+the+%22toad+licking%22+phenomenon&amp;rft.volume=28&amp;rft.issue=6&amp;rft.pages=%3Cspan+class%3D%22nowrap%22%3E521-%3C%2Fspan%3E538&amp;rft.date=1993&amp;rft_id=info%3Adoi%2F10.3109%2F10826089309039645&amp;rft_id=info%3Apmid%2F8486435&amp;rft.au=Lyttle%2C+T.&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AFrog" class="Z3988"></span></span> </li> <li id="cite_note-dartpoison-234"><span class="mw-cite-backlink">^ <a href="#cite_ref-dartpoison_234-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-dartpoison_234-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFMyers,_Charles_W.Daly,_John_W.Malkin,_Borys1978" class="citation journal cs1">Myers, Charles W.; Daly, John W.; Malkin, Borys (1978). "A dangerously toxic new frog (<i>Phyllobates</i>) used by Emberá Indians of western Colombia, with discussion of blowgun fabrication and dart poisoning". <i>Bulletin of the American Museum of Natural History</i>. <b>161</b>: <span class="nowrap">307–</span>366. <a href="/wiki/Hdl_(identifier)" class="mw-redirect" title="Hdl (identifier)">hdl</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://hdl.handle.net/2246%2F1286">2246/1286</a></span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Bulletin+of+the+American+Museum+of+Natural+History&amp;rft.atitle=A+dangerously+toxic+new+frog+%28Phyllobates%29+used+by+Ember%C3%A1+Indians+of+western+Colombia%2C+with+discussion+of+blowgun+fabrication+and+dart+poisoning&amp;rft.volume=161&amp;rft.pages=%3Cspan+class%3D%22nowrap%22%3E307-%3C%2Fspan%3E366&amp;rft.date=1978&amp;rft_id=info%3Ahdl%2F2246%2F1286&amp;rft.au=Myers%2C+Charles+W.&amp;rft.au=Daly%2C+John+W.&amp;rft.au=Malkin%2C+Borys&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AFrog" class="Z3988"></span></span> </li> <li id="cite_note-culture-235"><span class="mw-cite-backlink"><b><a href="#cite_ref-culture_235-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFMattison,_Chris2011" class="citation book cs1">Mattison, Chris (2011). <i>Frogs and Toads of the Worlds</i>. Princeton University Press. pp.&#160;<span class="nowrap">91–</span>92. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/978-0-691-14968-4" title="Special:BookSources/978-0-691-14968-4"><bdi>978-0-691-14968-4</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=book&amp;rft.btitle=Frogs+and+Toads+of+the+Worlds&amp;rft.pages=%3Cspan+class%3D%22nowrap%22%3E91-%3C%2Fspan%3E92&amp;rft.pub=Princeton+University+Press&amp;rft.date=2011&amp;rft.isbn=978-0-691-14968-4&amp;rft.au=Mattison%2C+Chris&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AFrog" class="Z3988"></span></span> </li> <li id="cite_note-236"><span class="mw-cite-backlink"><b><a href="#cite_ref-236">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFSleigh,_Charlotee2012" class="citation book cs1">Sleigh, Charlotee (2012). <i>Frog</i>. Reaktion Books. pp.&#160;<span class="nowrap">40–</span>42. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/978-1-86189-920-0" title="Special:BookSources/978-1-86189-920-0"><bdi>978-1-86189-920-0</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=book&amp;rft.btitle=Frog&amp;rft.pages=%3Cspan+class%3D%22nowrap%22%3E40-%3C%2Fspan%3E42&amp;rft.pub=Reaktion+Books&amp;rft.date=2012&amp;rft.isbn=978-1-86189-920-0&amp;rft.au=Sleigh%2C+Charlotee&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AFrog" class="Z3988"></span></span> </li> <li id="cite_note-237"><span class="mw-cite-backlink"><b><a href="#cite_ref-237">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFSleigh,_Charlotee2012" class="citation book cs1">Sleigh, Charlotee (2012). <i>Frog</i>. Reaktion Books. pp.&#160;<span class="nowrap">167–</span>175. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/978-1-86189-920-0" title="Special:BookSources/978-1-86189-920-0"><bdi>978-1-86189-920-0</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=book&amp;rft.btitle=Frog&amp;rft.pages=%3Cspan+class%3D%22nowrap%22%3E167-%3C%2Fspan%3E175&amp;rft.pub=Reaktion+Books&amp;rft.date=2012&amp;rft.isbn=978-1-86189-920-0&amp;rft.au=Sleigh%2C+Charlotee&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AFrog" class="Z3988"></span></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="Further_reading">Further reading</h2></div> <ul><li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFBeltz2005" class="citation book cs1">Beltz, Ellin (2005). <a rel="nofollow" class="external text" href="https://archive.org/details/frogs00elli"><i>Frogs: Inside their Remarkable World</i></a>. Firefly Books. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/978-1-55297-869-6" title="Special:BookSources/978-1-55297-869-6"><bdi>978-1-55297-869-6</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=book&amp;rft.btitle=Frogs%3A+Inside+their+Remarkable+World&amp;rft.pub=Firefly+Books&amp;rft.date=2005&amp;rft.isbn=978-1-55297-869-6&amp;rft.aulast=Beltz&amp;rft.aufirst=Ellin&amp;rft_id=https%3A%2F%2Farchive.org%2Fdetails%2Ffrogs00elli&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AFrog" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFCoggerZweifelKirschner2004" class="citation book cs1">Cogger, H. G.; Zweifel, R. G.; Kirschner, D. (2004). <i>Encyclopedia of Reptiles &amp; Amphibians</i> (2nd&#160;ed.). Fog City Press. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/978-1-877019-69-2" title="Special:BookSources/978-1-877019-69-2"><bdi>978-1-877019-69-2</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=book&amp;rft.btitle=Encyclopedia+of+Reptiles+%26+Amphibians&amp;rft.edition=2nd&amp;rft.pub=Fog+City+Press&amp;rft.date=2004&amp;rft.isbn=978-1-877019-69-2&amp;rft.aulast=Cogger&amp;rft.aufirst=H.+G.&amp;rft.au=Zweifel%2C+R.+G.&amp;rft.au=Kirschner%2C+D.&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AFrog" class="Z3988"></span></li> <li>Estes, R., and O. A. Reig. (1973). "The early fossil record of frogs: a review of the evidence." pp.&#160;11–63 In J. L. Vial (Ed.), <i>Evolutionary Biology of the Anurans: Contemporary Research on Major Problems</i>. University of Missouri Press, Columbia.</li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFGissiSan_MauroPesoleZardoya2006" class="citation journal cs1">Gissi, Carmela; San Mauro, Diego; Pesole, Graziano; Zardoya, Rafael (February 2006). "Mitochondrial phylogeny of Anura (Amphibia): A case study of congruent phylogenetic reconstruction using amino acid and nucleotide characters". <i>Gene</i>. <b>366</b> (2): <span class="nowrap">228–</span>237. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.gene.2005.07.034">10.1016/j.gene.2005.07.034</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/16307849">16307849</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Gene&amp;rft.atitle=Mitochondrial+phylogeny+of+Anura+%28Amphibia%29%3A+A+case+study+of+congruent+phylogenetic+reconstruction+using+amino+acid+and+nucleotide+characters&amp;rft.volume=366&amp;rft.issue=2&amp;rft.pages=%3Cspan+class%3D%22nowrap%22%3E228-%3C%2Fspan%3E237&amp;rft.date=2006-02&amp;rft_id=info%3Adoi%2F10.1016%2Fj.gene.2005.07.034&amp;rft_id=info%3Apmid%2F16307849&amp;rft.aulast=Gissi&amp;rft.aufirst=Carmela&amp;rft.au=San+Mauro%2C+Diego&amp;rft.au=Pesole%2C+Graziano&amp;rft.au=Zardoya%2C+Rafael&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AFrog" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFHolman2004" class="citation book cs1">Holman, J. A. (2004). <i>Fossil Frogs and Toads of North America</i>. Indiana University Press. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/978-0-253-34280-5" title="Special:BookSources/978-0-253-34280-5"><bdi>978-0-253-34280-5</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=book&amp;rft.btitle=Fossil+Frogs+and+Toads+of+North+America&amp;rft.pub=Indiana+University+Press&amp;rft.date=2004&amp;rft.isbn=978-0-253-34280-5&amp;rft.aulast=Holman&amp;rft.aufirst=J.+A.&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AFrog" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFSan_MauroVencesAlcobendasZardoya2005" class="citation journal cs1">San Mauro, Diego; Vences, Miguel; Alcobendas, Marina; Zardoya, Rafael; Meyer, Axel (May 2005). <a rel="nofollow" class="external text" href="http://nbn-resolving.de/urn:nbn:de:bsz:352-opus-33053">"Initial diversification of living amphibians predated the breakup of Pangaea"</a>. <i>American Naturalist</i>. <b>165</b> (5): <span class="nowrap">590–</span>599. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1086%2F429523">10.1086/429523</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/15795855">15795855</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a>&#160;<a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:17021360">17021360</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=American+Naturalist&amp;rft.atitle=Initial+diversification+of+living+amphibians+predated+the+breakup+of+Pangaea&amp;rft.volume=165&amp;rft.issue=5&amp;rft.pages=%3Cspan+class%3D%22nowrap%22%3E590-%3C%2Fspan%3E599&amp;rft.date=2005-05&amp;rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A17021360%23id-name%3DS2CID&amp;rft_id=info%3Apmid%2F15795855&amp;rft_id=info%3Adoi%2F10.1086%2F429523&amp;rft.aulast=San+Mauro&amp;rft.aufirst=Diego&amp;rft.au=Vences%2C+Miguel&amp;rft.au=Alcobendas%2C+Marina&amp;rft.au=Zardoya%2C+Rafael&amp;rft.au=Meyer%2C+Axel&amp;rft_id=http%3A%2F%2Fnbn-resolving.de%2Furn%3Anbn%3Ade%3Absz%3A352-opus-33053&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AFrog" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFTyler1994" class="citation book cs1">Tyler, M. J. (1994). <i>Australian Frogs A Natural History</i>. Reed Books. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/978-0-7301-0468-1" title="Special:BookSources/978-0-7301-0468-1"><bdi>978-0-7301-0468-1</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=book&amp;rft.btitle=Australian+Frogs+A+Natural+History&amp;rft.pub=Reed+Books&amp;rft.date=1994&amp;rft.isbn=978-0-7301-0468-1&amp;rft.aulast=Tyler&amp;rft.aufirst=M.+J.&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AFrog" class="Z3988"></span></li></ul> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1235681985" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1237033735" /><div class="side-box side-box-right plainlinks sistersitebox"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409" /> 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title="q:Frogs">Frogs</a></b></i>.</div></div> </div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1235681985" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1237033735" /><div class="side-box side-box-right plainlinks sistersitebox"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409" /> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Wikispecies-logo.svg" class="mw-file-description"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/40px-Wikispecies-logo.svg.png" decoding="async" width="34" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/60px-Wikispecies-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/120px-Wikispecies-logo.svg.png 2x" data-file-width="941" data-file-height="1103" /></a></span></div> <div class="side-box-text plainlist"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a> has information related to <i><b><a href="https://species.wikimedia.org/wiki/Anura" class="extiw" title="wikispecies:Anura">Anura</a></b></i>.</div></div> </div> <ul><li><a rel="nofollow" class="external text" href="http://amphibiaweb.org/">AmphibiaWeb</a></li> <li><a rel="nofollow" class="external text" href="http://www.theinformationarchives.com/frogs/">Gallery of Frogs</a>&#160;– Photography and images of various frog species</li> <li><a rel="nofollow" class="external text" href="http://froggy.lbl.gov/">The Whole Frog Project</a>&#160;– Virtual frog dissection and anatomy</li> <li><a rel="nofollow" class="external text" href="http://raysweb.net/specialplaces/pages/frogsdecline.html">"Disappearance of toads, frogs has some scientists worried"</a> <i>San Francisco Chronicle</i>, April 20, 1992</li> <li><a rel="nofollow" class="external text" href="http://calphotos.berkeley.edu/browse_imgs/amphibian_sci_1.html">Amphibian photo gallery by scientific name</a>&#160;– Features many unusual frogs</li> <li><a rel="nofollow" class="external text" href="http://www.scientificamerican.com/article/researchers-pinpoint-sour/"><i>Scientific American</i>: Researchers Pinpoint Source of Poison Frogs' Deadly Defenses</a></li></ul> <div class="mw-heading mw-heading3"><h3 id="Media">Media</h3></div> <ul><li><a rel="nofollow" class="external text" href="http://www.thinkoholic.com/2005/03/31/time-lapse-part-5-frog-spawn/">Time-lapse video showing the egg's development until hatching</a></li> <li><a rel="nofollow" class="external text" href="https://sounds.bl.uk/Environment/Amphibians">Frog vocalisations from around the world</a>&#160;– From the <a href="/wiki/British_Library_Sound_Archive" title="British Library Sound Archive">British Library Sound Archive</a></li> <li><a rel="nofollow" class="external text" href="http://www.naturenorth.com/spring/sound/shfrsnd.html">Frog calls</a>&#160;– From <a href="/wiki/Manitoba" title="Manitoba">Manitoba</a>, Canada</li></ul> <p><br /> </p> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol ol,.mw-parser-output .hlist ol ul,.mw-parser-output .hlist ul dl,.mw-parser-output .hlist ul ol,.mw-parser-output .hlist ul ul{display:inline}.mw-parser-output .hlist .mw-empty-li{display:none}.mw-parser-output .hlist dt::after{content:": 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4em"><a href="/wiki/Lissamphibia" title="Lissamphibia">Lissamphibia</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="*Kingdom:_Animalia_*Phylum:_Chordata_*Clade:_Osteichthyes_*Clade:_Sarcopterygii_*Clade:_Tetrapoda291" style="font-size:114%;margin:0 4em"><span style="font-weight:normal; font-style:normal"><span style="font-size: 85%;"> <ul><li>Kingdom: <a href="/wiki/Animal" title="Animal">Animalia</a></li> <li>Phylum: <a href="/wiki/Chordate" title="Chordate">Chordata</a></li> <li><i>Clade</i>: <a href="/wiki/Osteichthyes" title="Osteichthyes">Osteichthyes</a></li> <li><i>Clade</i>: <a href="/wiki/Sarcopterygii" title="Sarcopterygii">Sarcopterygii</a></li> <li><i>Clade</i>: <a href="/wiki/Tetrapod" title="Tetrapod">Tetrapoda</a></li></ul> </span></span></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Tetrapodomorpha41" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Tetrapodomorpha" title="Tetrapodomorpha">Tetrapodomorpha</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Tetrapodomorpha" title="Template:Tetrapodomorpha">Tetrapodomorpha</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Batrachomorpha" title="Batrachomorpha">Batrachomorpha</a> / <a href="/wiki/Temnospondyli" title="Temnospondyli">Temnospondyli</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Temnospondyli" title="Template:Temnospondyli">Temnospondyli</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Dissorophoidea" title="Dissorophoidea">Dissorophoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Dissorophoidea" title="Template:Dissorophoidea">Dissorophoidea</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Lissamphibia" title="Lissamphibia">Lissamphibia</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2" style="background:#ddddff;font-size:115%"><div><b><a href="/wiki/Lissamphibia" title="Lissamphibia">Lissamphibia</a></b></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="†Albanerpetontidae?178" scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Albanerpetontidae" title="Albanerpetontidae">Albanerpetontidae</a>?</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Albanerpeton" title="Albanerpeton">Albanerpeton</a></i></li> <li><i><a href="/wiki/Anoualerpeton" title="Anoualerpeton">Anoualerpeton</a></i></li> <li><i><a href="/wiki/Celtedens" title="Celtedens">Celtedens</a></i></li> <li><i><a href="/wiki/Shirerpeton" title="Shirerpeton">Shirerpeton</a></i></li> <li><i><a href="/wiki/Wesserpeton" title="Wesserpeton">Wesserpeton</a></i></li> <li><i><a href="/wiki/Yaksha_perettii" title="Yaksha perettii">Yaksha</a></i></li></ul> </div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="3" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Albanerpeton_inexpectatum.jpg" class="mw-file-description" title="Albanerpeton inexpectatum"><img alt="Albanerpeton inexpectatum" src="//upload.wikimedia.org/wikipedia/commons/thumb/f/f5/Albanerpeton_inexpectatum.jpg/250px-Albanerpeton_inexpectatum.jpg" decoding="async" width="140" height="53" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/f5/Albanerpeton_inexpectatum.jpg/330px-Albanerpeton_inexpectatum.jpg 2x" data-file-width="1600" data-file-height="603" /></a></span> <p><span typeof="mw:File"><a href="/wiki/File:Eocaecilia_BW.jpg" class="mw-file-description" title="Eocaecilia micropodia"><img alt="Eocaecilia micropodia" src="//upload.wikimedia.org/wikipedia/commons/thumb/2/27/Eocaecilia_BW.jpg/250px-Eocaecilia_BW.jpg" decoding="async" width="140" height="128" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/27/Eocaecilia_BW.jpg/330px-Eocaecilia_BW.jpg 2x" data-file-width="800" data-file-height="732" /></a></span> <span typeof="mw:File"><a href="/wiki/File:Triassurus_NT.png" class="mw-file-description" title="Triassurus sixtelae"><img alt="Triassurus sixtelae" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/78/Triassurus_NT.png/140px-Triassurus_NT.png" decoding="async" width="140" height="114" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/78/Triassurus_NT.png/210px-Triassurus_NT.png 1.5x, //upload.wikimedia.org/wikipedia/commons/7/78/Triassurus_NT.png 2x" data-file-width="229" data-file-height="187" /></a></span> </p> <span style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Triadobatrachus_BW.jpg" class="mw-file-description" title="Triadobatrachus massinoti"><img alt="Triadobatrachus massinoti" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/90/Triadobatrachus_BW.jpg/140px-Triadobatrachus_BW.jpg" decoding="async" width="140" height="105" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/90/Triadobatrachus_BW.jpg/210px-Triadobatrachus_BW.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/90/Triadobatrachus_BW.jpg/280px-Triadobatrachus_BW.jpg 2x" data-file-width="800" data-file-height="600" /></a></span></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;transform: translate(-50%,-50%) rotate(270deg)"><a href="/wiki/Gymnophionomorpha" class="mw-redirect" title="Gymnophionomorpha">Gymnophionomorpha <small><span style="font-weight:normal; font-style:normal">(caecilians total group)</span></small></a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Eocaecilia" title="Eocaecilia">Eocaecilia</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Funcusvermis" title="Funcusvermis">Funcusvermis</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Rubricacaecilia" title="Rubricacaecilia">Rubricacaecilia</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Chinlestegophis" title="Chinlestegophis">Chinlestegophis</a></i>?</li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><b><a href="/wiki/Albanerpetontidae" title="Albanerpetontidae">Albanerpetontidae</a></b>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;transform: translate(-50%,-50%) rotate(270deg)"><a href="/wiki/Caecilian" title="Caecilian">Gymnophiona <small><span style="font-weight:normal; font-style:normal">(caecilians crown group)</span></small></a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Rhinatrematidae_(American_tailed_caecilians)138" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Rhinatrematidae" title="Rhinatrematidae">Rhinatrematidae <small><span style="font-weight:normal; font-style:normal">(American tailed caecilians)</span></small></a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Amazops" title="Amazops">Amazops</a></i></li> <li><i><a href="/wiki/Epicrionops" title="Epicrionops">Epicrionops</a></i></li> <li><i><a href="/wiki/Rhinatrema" title="Rhinatrema">Rhinatrema</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;transform: translate(-50%,-50%) rotate(270deg)"><a href="/wiki/Caecilian" title="Caecilian">Stegokrotaphia</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Ichthyophiidae_(Asian_tailed_caecilians)133" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ichthyophiidae" title="Ichthyophiidae">Ichthyophiidae <small><span style="font-weight:normal; font-style:normal">(Asian tailed caecilians)</span></small></a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Ichthyophis" title="Ichthyophis">Ichthyophis</a></i></li> <li><i><a href="/wiki/Uraeotyphlus" title="Uraeotyphlus">Uraeotyphlus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;transform: translate(-50%,-50%) rotate(270deg)"><a href="/wiki/Caecilian" title="Caecilian">Teresomata</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Scolecomorphidae" title="Scolecomorphidae">Scolecomorphidae <small><span style="font-weight:normal; font-style:normal">(buried-eyed caecilians)</span></small></a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Scolecomorphidae" title="Scolecomorphidae">Crotaphatrema</a></i></li> <li><i><a href="/wiki/Scolecomorphidae" title="Scolecomorphidae">Scolecomorphus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Chikilidae" title="Chikilidae">Chikilidae <small><span style="font-weight:normal; font-style:normal">(Northeast Indian caecilians)</span></small></a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Chikila" title="Chikila">Chikila</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Herpelidae" title="Herpelidae">Herpelidae <small><span style="font-weight:normal; font-style:normal">(African caecilians)</span></small></a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Boulengerula" title="Boulengerula">Boulengerula</a></i></li> <li><i><a href="/wiki/Herpele" title="Herpele">Herpele</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Typhlonectidae" title="Typhlonectidae">Typhlonectidae <small><span style="font-weight:normal; font-style:normal">(aquatic caecilians)</span></small></a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Atretochoana" title="Atretochoana">Atretochoana</a></i></li> <li><i><a href="/wiki/Chthonerpeton" title="Chthonerpeton">Chthonerpeton</a></i></li> <li><i><a href="/wiki/Nectocaecilia" title="Nectocaecilia">Nectocaecilia</a></i></li> <li><i><a href="/wiki/Potamotyphlus_kaupii" title="Potamotyphlus kaupii">Potamotyphlus</a></i></li> <li><i><a href="/wiki/Typhlonectes" title="Typhlonectes">Typhlonectes</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Ymboirana" title="Ymboirana">Ymboirana</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Caeciliidae" title="Caeciliidae">Caeciliidae <small><span style="font-weight:normal; font-style:normal">(common caecilians)</span></small></a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Apodops" title="Apodops">Apodops</a></i></li> <li><i><a href="/wiki/Caecilia" title="Caecilia">Caecilia</a></i></li> <li><i><a href="/wiki/Oscaecilia" title="Oscaecilia">Oscaecilia</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Grandisoniidae" title="Grandisoniidae">Grandisoniidae <small><span style="font-weight:normal; font-style:normal">(Indo-African caecilians)</span></small></a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Gegeneophis" title="Gegeneophis">Gegeneophis</a></i></li> <li><i><a href="/wiki/Grandisonia" title="Grandisonia">Grandisonia</a></i></li> <li><i><a href="/wiki/Hypogeophis" title="Hypogeophis">Hypogeophis</a></i></li> <li><i><a href="/wiki/Makumuno_Assumbo_caecilian" title="Makumuno Assumbo caecilian">Idiocranium</a></i></li> <li><i><a href="/wiki/Indotyphlus" title="Indotyphlus">Indotyphlus</a></i></li> <li><i><a href="/wiki/Praslinia" title="Praslinia">Praslinia</a></i></li> <li><i><a href="/wiki/Sylvacaecilia" title="Sylvacaecilia">Sylvacaecilia</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Dermophiidae" title="Dermophiidae">Dermophiidae <small><span style="font-weight:normal; font-style:normal">(Neotropical caecilians)</span></small></a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Dermophis" title="Dermophis">Dermophis</a></i></li> <li><i><a href="/wiki/Geotrypetes" title="Geotrypetes">Geotrypetes</a></i></li> <li><i><a href="/wiki/Gymnopis" title="Gymnopis">Gymnopis</a></i></li> <li><i><a href="/wiki/Schistometopum" title="Schistometopum">Schistometopum</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Siphonopidae" title="Siphonopidae">Siphonopidae <small><span style="font-weight:normal; font-style:normal">(South American caecilians)</span></small></a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Brasilotyphlus" title="Brasilotyphlus">Brasilotyphlus</a></i></li> <li><i><a href="/wiki/Luetkenotyphlus" title="Luetkenotyphlus">Luetkenotyphlus</a></i></li> <li><i><a href="/wiki/Microcaecilia" title="Microcaecilia">Microcaecilia</a></i></li> <li><i><a href="/wiki/Mimosiphonops" title="Mimosiphonops">Mimosiphonops</a></i></li> <li><i><a href="/wiki/Siphonops" title="Siphonops">Siphonops</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:2em;line-height:1em"><div style="position:absolute;transform: translate(-50%,-50%) rotate(270deg)"><br /><br /><a href="/wiki/Batrachia" title="Batrachia">Batrachia<br /><small><span style="font-weight:normal; font-style:normal">(salamanders + frogs<br />crown group)</span></small></a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;line-height:1em"><a href="/wiki/Caudata" title="Caudata">Caudata<br /><small><span style="font-weight:normal; font-style:normal">(salamanders<br />total group)</span></small></a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/w/index.php?title=Egoria&amp;action=edit&amp;redlink=1" class="new" title="Egoria (page does not exist)">Egoria</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Kulgeriherpeton" title="Kulgeriherpeton">Kulgeriherpeton</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Triassurus" title="Triassurus">Triassurus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Urupia" title="Urupia">Urupia</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;line-height:1em"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Karauridae" title="Karauridae">Karauridae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Karaurus" title="Karaurus">Karaurus</a></i></li> <li><i><a href="/wiki/Kokartus" title="Kokartus">Kokartus</a></i></li> <li><i><a href="/w/index.php?title=Kuzbassia&amp;action=edit&amp;redlink=1" class="new" title="Kuzbassia (page does not exist)">Kuzbassia</a></i></li> <li><i><a href="/wiki/Marmorerpeton" title="Marmorerpeton">Marmorerpeton</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;line-height:1em"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Batrachosauroididae" title="Batrachosauroididae">Batrachosauroididae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Batrachosauroides" title="Batrachosauroides">Batrachosauroides</a></i></li> <li><i><a href="/wiki/Opisthotriton" title="Opisthotriton">Opisthotriton</a></i></li> <li><i><a href="/wiki/Palaeoproteus" title="Palaeoproteus">Palaeoproteus</a></i></li> <li><i><a href="/wiki/Parrisia" title="Parrisia">Parrisia</a></i></li> <li><i><a href="/wiki/Peratosauroides" title="Peratosauroides">Peratosauroides</a></i></li> <li><i><a href="/wiki/Prodesmodon" title="Prodesmodon">Prodesmodon</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;line-height:1em"><a href="/wiki/Salamander" title="Salamander">Urodela <small><span style="font-weight:normal; font-style:normal">(salamanders crown group)</span></small></a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>115 genera</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;line-height:1em"><a href="/wiki/Salientia" title="Salientia">Salientia<br /><small><span style="font-weight:normal; font-style:normal">(frogs<br />total group)</span></small></a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Czatkobatrachus" title="Czatkobatrachus">Czatkobatrachus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Notobatrachus" title="Notobatrachus">Notobatrachus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Prosalirus" title="Prosalirus">Prosalirus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Triadobatrachus" title="Triadobatrachus">Triadobatrachus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Vieraella" title="Vieraella">Vieraella</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;line-height:1em"><a class="mw-selflink selflink">Anura <small><span style="font-weight:normal; font-style:normal">(frogs crown group)</span></small></a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>several hundred genera</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li>See also: <a href="/wiki/List_of_amphibian_genera" title="List of amphibian genera">List of amphibian genera</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235" /></div><div role="navigation" class="navbox" aria-labelledby="Extant_amphibian_positions_by_subclass172" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231" /><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Amphibians" title="Template:Amphibians"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Amphibians" title="Template talk:Amphibians"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Amphibians" title="Special:EditPage/Template:Amphibians"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Extant_amphibian_positions_by_subclass172" style="font-size:114%;margin:0 4em">Extant <a href="/wiki/Amphibian" title="Amphibian">amphibian</a> positions by subclass</div></th></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li>Kingdom <a href="/wiki/Animal" title="Animal">Animalia</a></li> <li>Phylum <a href="/wiki/Chordate" title="Chordate">Chordata</a></li> <li>Subphylum <a href="/wiki/Craniate" title="Craniate">Craniata</a></li> <li>Superclass <a href="/wiki/Tetrapod" title="Tetrapod">Tetrapoda</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Lissamphibia" title="Lissamphibia">Lissamphibia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a class="mw-selflink selflink">Anura (frogs)</a></li> <li><a href="/wiki/Salamander" title="Salamander">Urodela (salamanders)</a></li> <li><a href="/wiki/Caecilian" title="Caecilian">Gymnophiona (caecilians)</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235" /></div><div role="navigation" class="navbox" aria-labelledby="Extant_anuran_families_by_suborder401" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231" /><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Anura" title="Template:Anura"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Anura" title="Template talk:Anura"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Anura" title="Special:EditPage/Template:Anura"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Extant_anuran_families_by_suborder401" style="font-size:114%;margin:0 4em">Extant <a class="mw-selflink selflink">anuran</a> families by suborder</div></th></tr><tr><td class="navbox-abovebelow" colspan="3"><div> <ul><li>Kingdom <a href="/wiki/Animal" title="Animal">Animalia</a></li> <li>Phylum <a href="/wiki/Chordate" title="Chordate">Chordata</a></li> <li>Subphylum <a href="/wiki/Craniate" title="Craniate">Craniata</a></li> <li>Superclass <a href="/wiki/Tetrapod" title="Tetrapod">Tetrapoda</a></li> <li>Class <a href="/wiki/Amphibian" title="Amphibian">Amphibia</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Archaeobatrachia" title="Archaeobatrachia">Archaeobatrachia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Tailed_frog" title="Tailed frog">Ascaphidae</a></li> <li><a href="/wiki/Bombinatoridae" title="Bombinatoridae">Bombinatoridae</a></li> <li><a href="/wiki/Discoglossidae" class="mw-redirect" title="Discoglossidae">Discoglossidae</a></li> <li><a href="/wiki/Leiopelma" title="Leiopelma">Leiopelmatidae</a></li></ul> </div></td><td class="noviewer navbox-image" rowspan="3" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:White-Lipped-small.jpg" class="mw-file-description" title="White-lipped Tree Frog"><img alt="White-lipped Tree Frog" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/9e/White-Lipped-small.jpg/50px-White-Lipped-small.jpg" decoding="async" width="50" height="37" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/9e/White-Lipped-small.jpg/75px-White-Lipped-small.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/9/9e/White-Lipped-small.jpg 2x" data-file-width="100" data-file-height="73" /></a></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Mesobatrachia" title="Mesobatrachia">Mesobatrachia</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Megophryidae" title="Megophryidae">Megophryidae</a></li> <li><a href="/wiki/European_spadefoot_toad" title="European spadefoot toad">Pelobatidae</a></li> <li><a href="/wiki/Parsley_frog" title="Parsley frog">Pelodytidae</a></li> <li><a href="/wiki/Pipidae" title="Pipidae">Pipidae</a></li> <li><a href="/wiki/American_spadefoot_toad" title="American spadefoot toad">Scaphiopodidae</a></li> <li><a href="/wiki/Mexican_burrowing_toad" title="Mexican burrowing toad">Rhinophrynidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Neobatrachia" title="Neobatrachia">Neobatrachia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Amphignathodontidae" title="Amphignathodontidae">Amphignathodontidae</a></li> <li><a href="/wiki/Aromobatidae" title="Aromobatidae">Aromobatidae</a></li> <li><a href="/wiki/Arthroleptidae" title="Arthroleptidae">Arthroleptidae</a></li> <li><a href="/wiki/Brachycephalidae" title="Brachycephalidae">Brachycephalidae</a></li> <li><a href="/wiki/Brevicipitidae" title="Brevicipitidae">Brevicipitidae</a></li> <li><a href="/wiki/True_toad" title="True toad">Bufonidae</a></li> <li><a href="/wiki/Glass_frog" title="Glass frog">Centrolenidae</a></li> <li><a href="/wiki/Craugastoridae" title="Craugastoridae">Craugastoridae</a></li> <li><a href="/wiki/Poison_dart_frog" title="Poison dart frog">Dendrobatidae</a></li> <li><a href="/wiki/Ghost_frog" title="Ghost frog">Heleophrynidae</a></li> <li><a href="/wiki/Hemiphractidae" title="Hemiphractidae">Hemiphractidae</a></li> <li><a href="/wiki/Shovelnose_frog" title="Shovelnose frog">Hemisotidae</a></li> <li><a href="/wiki/Hylidae" title="Hylidae">Hylidae</a></li> <li><a href="/wiki/Hyperoliidae" title="Hyperoliidae">Hyperoliidae</a></li> <li><a href="/wiki/Leptodactylidae" title="Leptodactylidae">Leptodactylidae</a></li> <li><a href="/wiki/Mantellidae" title="Mantellidae">Mantellidae</a></li> <li><a href="/wiki/Microhylidae" title="Microhylidae">Microhylidae</a></li> <li><a href="/wiki/Myobatrachidae" title="Myobatrachidae">Myobatrachidae</a></li> <li><a href="/wiki/Petropedetidae" title="Petropedetidae">Petropedetidae</a></li> <li><a href="/wiki/Pyxicephalidae" title="Pyxicephalidae">Pyxicephalidae</a></li> <li><a href="/wiki/True_frog" title="True frog">Ranidae</a></li> <li><a href="/wiki/Rhacophoridae" title="Rhacophoridae">Rhacophoridae</a></li> <li><a href="/wiki/Rhinodermatidae" title="Rhinodermatidae">Rhinodermatidae</a></li> <li><a href="/wiki/Sooglossidae" title="Sooglossidae">Sooglossidae</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="3"><div><span class="nowrap"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c1/Variegated_golden_frog_%28Mantella_baroni%29_Ranomafana.jpg/21px-Variegated_golden_frog_%28Mantella_baroni%29_Ranomafana.jpg" decoding="async" width="21" height="14" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c1/Variegated_golden_frog_%28Mantella_baroni%29_Ranomafana.jpg/32px-Variegated_golden_frog_%28Mantella_baroni%29_Ranomafana.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c1/Variegated_golden_frog_%28Mantella_baroni%29_Ranomafana.jpg/42px-Variegated_golden_frog_%28Mantella_baroni%29_Ranomafana.jpg 2x" data-file-width="4302" data-file-height="2869" /></span></span> </span><a href="/wiki/Portal:Frogs" title="Portal:Frogs">Frogs&#32;portal</a></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235" /></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers3767" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers3767" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Anura</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q53636" class="extiw" title="wikidata:Q53636">Q53636</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Anura" class="extiw" title="wikispecies:Anura">Anura</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Animal_Diversity_Web" title="Animal Diversity Web">ADW</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://animaldiversity.org/accounts/Anura/">Anura</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Australian_Faunal_Directory" title="Australian Faunal Directory">AFD</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://biodiversity.org.au/afd/taxa/Anura">Anura</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Amphibian_Species_of_the_World" title="Amphibian Species of the World">ASW</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://amphibiansoftheworld.amnh.org/Amphibia/Anura">Anura</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Barcode_of_Life_Data_System" title="Barcode of Life Data System">BOLD</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://bench.boldsystems.org/index.php/TaxBrowser_TaxonPage?taxid=219">219</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Catalogue_of_Life" title="Catalogue of Life">CoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.catalogueoflife.org/data/taxon/PW">PW</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/1553">1553</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/EPPO_Code" title="EPPO Code">EPPO</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://gd.eppo.int/taxon/1ANURO">1ANURO</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Fauna_Europaea" title="Fauna Europaea">Fauna Europaea</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://www.eu-nomen.eu/portal/taxon.php?GUID=urn:lsid:faunaeur.org:taxname:15534">15534</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Fauna_Europaea" title="Fauna Europaea">Fauna Europaea (new)</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://fauna-eu.org/cdm_dataportal/taxon/8675db52-461d-4ab4-8574-dbc5007b80c3">8675db52-461d-4ab4-8574-dbc5007b80c3</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/952">952</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/INaturalist" title="INaturalist">iNaturalist</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://inaturalist.org/taxa/20979">20979</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&amp;id=10211">10211</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Integrated_Taxonomic_Information_System" title="Integrated Taxonomic Information System">ITIS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&amp;search_value=173423">173423</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Biodiversity_Network" title="National Biodiversity Network">NBN</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://species.nbnatlas.org/species/NHMSYS0000375308">NHMSYS0000375308</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&amp;id=8342">8342</a></span></span></li> <li><span style="white-space:nowrap;">NZOR: <span class="uid"><a rel="nofollow" class="external text" href="https://www.nzor.org.nz/names/a330081c-9f45-47fb-80dd-ba8d870e5bd2">a330081c-9f45-47fb-80dd-ba8d870e5bd2</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=991547">991547</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=150288">150288</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Plazi" title="Plazi">Plazi</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://treatment.plazi.org/id/CB2D87E1-2E34-FFC6-FC3B-41D7FC23FC69">CB2D87E1-2E34-FFC6-FC3B-41D7FC23FC69</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/World_Register_of_Marine_Species" title="World Register of Marine Species">WoRMS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.marinespecies.org/aphia.php?p=taxdetails&amp;id=448306">448306</a></span></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235" /><style data-mw-deduplicate="TemplateStyles:r1038841319">.mw-parser-output .tooltip-dotted{border-bottom:1px dotted;cursor:help}</style><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1038841319" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1038841319" /></div><div role="navigation" class="navbox authority-control" aria-label="Navbox1338" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Help:Authority_control" title="Help:Authority control">Authority control databases</a>: National <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q53636#identifiers" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://d-nb.info/gnd/4018698-2">Germany</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/sh85052057">United States</a></span></li><li><span class="uid"><span class="rt-commentedText tooltip tooltip-dotted" title="Anoures"><a rel="nofollow" class="external text" href="https://catalogue.bnf.fr/ark:/12148/cb11974461k">France</a></span></span></li><li><span class="uid"><span class="rt-commentedText tooltip tooltip-dotted" title="Anoures"><a rel="nofollow" class="external text" href="https://data.bnf.fr/ark:/12148/cb11974461k">BnF data</a></span></span></li><li><span class="uid"><span class="rt-commentedText tooltip tooltip-dotted" title="žáby"><a rel="nofollow" class="external text" href="https://aleph.nkp.cz/F/?func=find-c&amp;local_base=aut&amp;ccl_term=ica=ph128054&amp;CON_LNG=ENG">Czech Republic</a></span></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.nli.org.il/en/authorities/987007553171905171">Israel</a></span></li></ul></div></td></tr></tbody></table></div> <style data-mw-deduplicate="TemplateStyles:r1130092004">.mw-parser-output .portal-bar{font-size:88%;font-weight:bold;display:flex;justify-content:center;align-items:baseline}.mw-parser-output .portal-bar-bordered{padding:0 2em;background-color:#fdfdfd;border:1px solid #a2a9b1;clear:both;margin:1em auto 0}.mw-parser-output .portal-bar-related{font-size:100%;justify-content:flex-start}.mw-parser-output .portal-bar-unbordered{padding:0 1.7em;margin-left:0}.mw-parser-output .portal-bar-header{margin:0 1em 0 0.5em;flex:0 0 auto;min-height:24px}.mw-parser-output .portal-bar-content{display:flex;flex-flow:row wrap;flex:0 1 auto;padding:0.15em 0;column-gap:1em;align-items:baseline;margin:0;list-style:none}.mw-parser-output .portal-bar-content-related{margin:0;list-style:none}.mw-parser-output .portal-bar-item{display:inline-block;margin:0.15em 0.2em;min-height:24px;line-height:24px}@media screen and (max-width:768px){.mw-parser-output .portal-bar{font-size:88%;font-weight:bold;display:flex;flex-flow:column wrap;align-items:baseline}.mw-parser-output .portal-bar-header{text-align:center;flex:0;padding-left:0.5em;margin:0 auto}.mw-parser-output .portal-bar-related{font-size:100%;align-items:flex-start}.mw-parser-output .portal-bar-content{display:flex;flex-flow:row wrap;align-items:center;flex:0;column-gap:1em;border-top:1px solid #a2a9b1;margin:0 auto;list-style:none}.mw-parser-output .portal-bar-content-related{border-top:none;margin:0;list-style:none}}.mw-parser-output .navbox+link+.portal-bar,.mw-parser-output .navbox+style+.portal-bar,.mw-parser-output .navbox+link+.portal-bar-bordered,.mw-parser-output .navbox+style+.portal-bar-bordered,.mw-parser-output .sister-bar+link+.portal-bar,.mw-parser-output .sister-bar+style+.portal-bar,.mw-parser-output .portal-bar+.navbox-styles+.navbox,.mw-parser-output .portal-bar+.navbox-styles+.sister-bar{margin-top:-1px}</style><div class="portal-bar noprint metadata noviewer portal-bar-bordered" role="navigation" aria-label="Portals"><span class="portal-bar-header"><a href="/wiki/Wikipedia:Contents/Portals" title="Wikipedia:Contents/Portals">Portals</a>:</span><ul class="portal-bar-content"><li class="portal-bar-item"><span class="nowrap"><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c1/Variegated_golden_frog_%28Mantella_baroni%29_Ranomafana.jpg/21px-Variegated_golden_frog_%28Mantella_baroni%29_Ranomafana.jpg" decoding="async" width="21" height="14" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c1/Variegated_golden_frog_%28Mantella_baroni%29_Ranomafana.jpg/32px-Variegated_golden_frog_%28Mantella_baroni%29_Ranomafana.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c1/Variegated_golden_frog_%28Mantella_baroni%29_Ranomafana.jpg/42px-Variegated_golden_frog_%28Mantella_baroni%29_Ranomafana.jpg 2x" data-file-width="4302" data-file-height="2869" /></span></span> </span><a href="/wiki/Portal:Frogs" title="Portal:Frogs">Frogs</a></li><li class="portal-bar-item"><span class="nowrap"><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/5/50/Frog_%282546%29_-_The_Noun_Project.svg/20px-Frog_%282546%29_-_The_Noun_Project.svg.png" decoding="async" width="19" height="19" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/50/Frog_%282546%29_-_The_Noun_Project.svg/40px-Frog_%282546%29_-_The_Noun_Project.svg.png 1.5x" data-file-width="512" data-file-height="512" /></span></span> </span><a href="/wiki/Portal:Amphibians" title="Portal:Amphibians">Amphibians</a></li><li class="portal-bar-item"><span class="nowrap"><span typeof="mw:File"><a href="/wiki/File:Wapiti_from_Wagon_Trails.jpg" class="mw-file-description"><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/44/Wapiti_from_Wagon_Trails.jpg/40px-Wapiti_from_Wagon_Trails.jpg" decoding="async" width="21" height="14" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/44/Wapiti_from_Wagon_Trails.jpg/60px-Wapiti_from_Wagon_Trails.jpg 2x" data-file-width="2064" data-file-height="1413" /></a></span> </span><a href="/wiki/Portal:Animals" title="Portal:Animals">Animals</a></li></ul></div> <p class="mw-empty-elt"> </p> <!-- NewPP limit report Parsed by mw‐web.codfw.main‐67db76d8d7‐r58c5 Cached time: 20250403183408 Cache expiry: 2592000 Reduced expiry: false Complications: [vary‐revision‐sha1, show‐toc] CPU time usage: 2.627 seconds 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