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Yarasuchus - Wikipedia

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vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button 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</div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Extinct genus of reptiles</div> <p class="mw-empty-elt"> </p> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><i>Yarasuchus</i><br /><div style="font-size: 85%;">Temporal range: <a href="/wiki/Middle_Triassic" title="Middle Triassic">Middle Triassic</a>, <span class="noprint"><span style="display:inline-block;"></span><span style="display:inline-block;">Anisian</span> <span style="display:inline-block;"></span><div id="Timeline-row" style="margin: 4px auto 0; clear:both; width:220px; padding:0px; height:18px; overflow:visible; white-space:nowrap; border:1px #666; border-style:solid none; position:relative; z-index:0; font-size:97%;"> <div style="position:absolute; height:100%; left:0px; width:207.23076923077px; padding-left:5px; text-align:left; background-color:rgb(254,217,106); background-image: linear-gradient(to right, rgba(255,255,255,1), rgba(254,217,106,1) 15%, rgba(254,217,106,1));"><a href="/wiki/Precambrian" title="Precambrian">PreꞒ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,160,86); left:37.636923076923px; width:18.073846153846px;"><a href="/wiki/Cambrian" title="Cambrian">Ꞓ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(0,146,112); left:55.710769230769px; width:14.08px;"><a href="/wiki/Ordovician" title="Ordovician">O</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(179,225,182); left:69.790769230769px; width:8.3261538461539px;"><a href="/wiki/Silurian" title="Silurian">S</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(203,140,55); left:78.116923076923px; width:20.409230769231px;"><a href="/wiki/Devonian" title="Devonian">D</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(103,165,153); left:98.526153846154px; width:20.307692307692px;"><a href="/wiki/Carboniferous" title="Carboniferous">C</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(240,64,40); left:118.83384615385px; width:15.907015384615px;"><a href="/wiki/Permian" title="Permian">P</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(129,43,146); left:134.74086153846px; width:17.092984615385px;"><a href="/wiki/Triassic" title="Triassic">T</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(52,178,201); left:151.83384615385px; width:19.089230769231px;"><a href="/wiki/Jurassic" title="Jurassic">J</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,198,78); left:170.92307692308px; width:26.738461538462px;"><a href="/wiki/Cretaceous" title="Cretaceous">K</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(253,154,82); left:197.66153846154px; width:14.543692307692px;"><a href="/wiki/Paleogene" title="Paleogene">Pg</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(255,230,25); left:212.20523076923px; width:6.9215384615385px;"><a href="/wiki/Neogene" title="Neogene">N</a></div> <div id="end-border" style="position:absolute; height:100%; background-color:#666; width:1px; left:219px"></div><div style="margin:0 auto; line-height:0; clear:both; width:220px; padding:0px; height:8px; overflow:visible; background-color:transparent; position:relative; top:-4px; z-index:100;"><div style="position:absolute; height:8px; left:136.33230769231px; width:1.76px; background-color:#360; opacity:0.42;"></div> <div style="position:absolute; height:8px; left:136.33230769231px; width:1.76px; background-color:#360; opacity:1;"></div> <div style="position:absolute; height:6px; top:1px; left:137.33230769231px; width:-0.24px; background-color:#6c3;"></div> </div> </div></span></div> </th></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Yarasuchus" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Chordate" title="Chordate">Chordata</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Archosaur" title="Archosaur">Archosauria</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Avemetatarsalia" title="Avemetatarsalia">Avemetatarsalia</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><span style="font-style:normal;font-weight:normal;">†</span><a href="/wiki/Aphanosauria" title="Aphanosauria">Aphanosauria</a> </td></tr> <tr> <td>Genus: </td> <td><span style="font-style:normal;font-weight:normal;">†</span><a class="mw-selflink selflink"><i>Yarasuchus</i></a><br /><small>Sen, <a href="/wiki/2005_in_paleontology" title="2005 in paleontology">2005</a></small> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Type_species" title="Type species">Type species</a> </th></tr> <tr> <td colspan="2" style="text-align: center"><i><b>Yarasuchus deccanensis</b></i><br /><div style="font-size: 85%;">Sen, 2005</div> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><i><b>Yarasuchus</b></i> (meaning "red crocodile") is an <a href="/wiki/Extinct" class="mw-redirect" title="Extinct">extinct</a> <a href="/wiki/Genus" title="Genus">genus</a> of <a href="/wiki/Avemetatarsalia" title="Avemetatarsalia">avemetatarsalian</a> <a href="/wiki/Archosaur" title="Archosaur">archosaur</a> that lived during the <a href="/wiki/Anisian" title="Anisian">Anisian</a> stage of the <a href="/wiki/Middle_Triassic" title="Middle Triassic">Middle Triassic</a> of <a href="/wiki/India" title="India">India</a>.<sup id="cite_ref-BS99_1-0" class="reference"><a href="#cite_note-BS99-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> The genus was named and described in 2005 from a collection of disarticulated but fairly complete fossil material found from the <a href="/wiki/Middle_Triassic" title="Middle Triassic">Middle Triassic</a> <a href="/wiki/Yerrapalli_Formation" title="Yerrapalli Formation">Yerrapalli Formation</a>. The material is thought to be from two individuals, possibly three, with one being much more complete and articulated than the other.<sup id="cite_ref-Sen,_2005_2-0" class="reference"><a href="#cite_note-Sen,_2005-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> The <a href="/wiki/Type_species" title="Type species">type</a> and only <a href="/wiki/Species" title="Species">species</a> is <i><b>Y. deccanensis</b></i>. <i>Yarasuchus</i> was a quadruped roughly 2–2.5 metres (6.6–8.2&#160;ft) long, with an elongated neck and tall spines on its vertebrae. Unlike other quadrupedal Triassic reptiles, the limbs and shoulders of <i>Yarasuchus</i> were slender, and more like those of <a href="/wiki/Ornithodira" class="mw-redirect" title="Ornithodira">ornithodirans</a>. </p><p><i>Yarasuchus</i> has had a complicated taxonomic history, after originally being described as a "<a href="/wiki/Prestosuchidae" title="Prestosuchidae">prestosuchid</a> <a href="/wiki/Rauisuchia" title="Rauisuchia">rauisuchian</a>", it was later variously recovered as a <a href="/wiki/Poposauroidea" title="Poposauroidea">poposauroid</a> <a href="/wiki/Pseudosuchia" title="Pseudosuchia">pseudosuchian</a> and a non-archosaurian <a href="/wiki/Archosauriformes" title="Archosauriformes">archosauriform</a> of unstable position. In 2017 it was determined to be related to the similarly enigmatic Triassic reptiles <i><a href="/wiki/Teleocrater" title="Teleocrater">Teleocrater</a></i>, <i><a href="/wiki/Dongusuchus" title="Dongusuchus">Dongusuchus</a></i> and <i><a href="/wiki/Spondylosoma" title="Spondylosoma">Spondylosoma</a>.</i> Together, they belong to a group called <a href="/wiki/Aphanosauria" title="Aphanosauria">Aphanosauria</a> and are placed at the base of Avemetatarsalia, sister to Ornithodira, making <i>Yarasuchus</i> one of the earliest diverging bird-line archosaurs known. The relative completeness of <i>Yarasuchus</i> and its evolutionary position helps to shed light on the origins of later, well known bird-line archosaurs such as the <a href="/wiki/Dinosaur" title="Dinosaur">dinosaurs</a> and <a href="/wiki/Pterosaur" title="Pterosaur">pterosaurs</a>.<sup id="cite_ref-Teleo_3-0" class="reference"><a href="#cite_note-Teleo-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Description">Description</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yarasuchus&amp;action=edit&amp;section=1" title="Edit section: Description"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><i>Yarasuchus</i> was a relatively small, gracile, low-slung quadruped with a long neck and a small head, estimated to be around 2–2.5 metres (6.6–8.2&#160;ft) long.<sup id="cite_ref-Sen,_2005_2-1" class="reference"><a href="#cite_note-Sen,_2005-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-:0_4-0" class="reference"><a href="#cite_note-:0-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> Despite its appearance, <i>Yarasuchus</i> has a number of features in common with other avemetatarsalians, such as its elongated neck vertebrae. It has long, slender limb bones, unlike the more robust limbs of most pseudosuchians, but similar to other avemetatarsalians. Unlike later avemetatarsalians though, <i>Yarasuchus</i> has a "crocodile-normal" ankle configuration, more similar to pseudosuchians and some stem-archosaurs. <i>Yarasuchus</i> is very similar in appearance to its close relative <i>Teleocrater</i>, and differs in only a few minor anatomical features of the skeleton, such as having a less of a rear facing <a href="/wiki/Glenoid_cavity" class="mw-redirect" title="Glenoid cavity">glenoid</a> (the bony shoulder socket) than that of <i>Teleocrater</i>.<sup id="cite_ref-Teleo_3-1" class="reference"><a href="#cite_note-Teleo-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Skull">Skull</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yarasuchus&amp;action=edit&amp;section=2" title="Edit section: Skull"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The skull of <i>Yarasuchus</i> is poorly represented, known from only a few isolated pieces. A number of bones were initially identified by Sen (2005), including a <a href="/wiki/Jugal" class="mw-redirect" title="Jugal">jugal</a>, <a href="/wiki/Quadrate_bone" title="Quadrate bone">quadrate</a> and part of the <a href="/wiki/Quadratojugal" class="mw-redirect" title="Quadratojugal">quadratojugal</a>, <a href="/wiki/Squamosal" class="mw-redirect" title="Squamosal">squamosal</a>, both <a href="/wiki/Pterygoid_bone" title="Pterygoid bone">pterygoids</a> and two <a href="/wiki/Maxilla" title="Maxilla">maxilla</a> that included a portion of <a href="/wiki/Premaxilla" title="Premaxilla">premaxilla</a> attached to one of them, as well as an associated tooth.<sup id="cite_ref-Sen,_2005_2-2" class="reference"><a href="#cite_note-Sen,_2005-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> However both maxillae differ from the known maxilla in <i>Teleocrater</i> and instead more closely resemble those of an <a href="/wiki/Allokotosaur" class="mw-redirect" title="Allokotosaur">allokotosaur</a>. <a href="/wiki/Sterling_Nesbitt" title="Sterling Nesbitt">Nesbitt</a> and colleagues also regarded the jugal, quadrate and quadratojugal as indeterminate bones, and re-identified the squamosal as a <a href="/wiki/Postorbital_bone" title="Postorbital bone">postorbital</a> belonging to a larger individual than the holotype specimen. As of 2017, the right postorbital and both pterygoids are the only positively identified cranial material belonging to <i>Yarasuchus</i>.<sup id="cite_ref-Teleo_3-2" class="reference"><a href="#cite_note-Teleo-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Skeleton">Skeleton</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yarasuchus&amp;action=edit&amp;section=3" title="Edit section: Skeleton"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Yarasuchus.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/26/Yarasuchus.jpg/220px-Yarasuchus.jpg" decoding="async" width="220" height="96" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/26/Yarasuchus.jpg/330px-Yarasuchus.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/26/Yarasuchus.jpg/440px-Yarasuchus.jpg 2x" data-file-width="2698" data-file-height="1172" /></a><figcaption>Outdated life restoration depicting <i>Yarasuchus</i> as a "raisuchian" with osteoderms.</figcaption></figure> <p>Almost the whole <a href="/wiki/Vertebral_column" title="Vertebral column">vertebral column</a> is represented in <i>Yarasuchus</i>, including at least 8 <a href="/wiki/Cervical_vertebrae" title="Cervical vertebrae">cervicals</a> (including the <a href="/wiki/Atlas_(anatomy)" title="Atlas (anatomy)">atlas</a> and <a href="/wiki/Axis_(anatomy)" title="Axis (anatomy)">axis</a>), 17 <a href="/wiki/Thoracic_vertebrae" title="Thoracic vertebrae">dorsals</a>, 2 <a href="/wiki/Sacral_vertebra" class="mw-redirect" title="Sacral vertebra">sacral</a>, and at least 11 proximal <a href="/wiki/Caudal_vertebra" class="mw-redirect" title="Caudal vertebra">caudal</a> vertebrae. The long neck of <i>Yarasuchus</i> is made up of a series of characteristically elongated cervical (neck) vertebrae, in contrast to its proportionately small skull. The articulating surfaces of the <a href="/wiki/Zygapophyses" class="mw-redirect" title="Zygapophyses">zygapophyses</a> that connect between each vertebra are inclined and the <a href="/wiki/Vertebral_centrum" class="mw-redirect" title="Vertebral centrum">centra</a> are strongly curved along the bottom margin with offset faces, indicating that the neck was held raised up from the body and arched along its length. The cervicals are uniquely characterised by a prominent midline keel that runs along the front half of the underside of each centrum, followed by two separate keels that continue to the rear edge.<sup id="cite_ref-NJKP07_5-0" class="reference"><a href="#cite_note-NJKP07-5"><span class="cite-bracket">&#91;</span>5<span class="cite-bracket">&#93;</span></a></sup> The posterior <a href="/wiki/Cervical_rib" title="Cervical rib">cervical ribs</a> have three heads, an unusual condition in archosaurs, and the corresponding cervicals possess an accessory articular facet to accommodate this.<sup id="cite_ref-Sen,_2005_2-3" class="reference"><a href="#cite_note-Sen,_2005-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Teleo_3-3" class="reference"><a href="#cite_note-Teleo-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-:0_4-1" class="reference"><a href="#cite_note-:0-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Teleo2_6-0" class="reference"><a href="#cite_note-Teleo2-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup> </p><p>Unlike the cervicals, the dorsal (back) vertebra are short and compact, almost only half the length of the cervicals, as are the two sacral (hip) vertebra. The front-most two or three dorsal vertebra possess similar keels to the cervicals, unlike the condition in <i>Teleocrater</i> which has no keeled dorsals. The preserved proximal caudal (tail) vertebra are consistently elongate, however the distal caudals are missing and so the end of the tail is unknown. The vertebrae all have tall <a href="/wiki/Neural_spines" class="mw-redirect" title="Neural spines">neural spines</a> that run down the back, the tallest of which are over the hip where the spines are two times taller than the corresponding centra. The neck has similarly tall neural spines, however they are longer than tall and inclined forward so as to overhang the vertebra before it. The cervical neural spines are also unusually thickened and roughly textured at the top. The caudal neural spines are also tall and narrow, unlike the elongated centra, and are inclined backwards, decreasing in height further down the tail. The tall, broad neural spines were initially believed to be associated with rows of paramedian <a href="/wiki/Osteoderm" title="Osteoderm">osteoderms</a> (although see below).<sup id="cite_ref-Sen,_2005_2-4" class="reference"><a href="#cite_note-Sen,_2005-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-:0_4-2" class="reference"><a href="#cite_note-:0-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> </p><p>The <a href="/wiki/Pectoral_girdle" class="mw-redirect" title="Pectoral girdle">pectoral (shoulder) girdle</a> is delicately built, with a tall and slender <a href="/wiki/Scapula" title="Scapula">scapula</a> that is expanded distally and constricted near the glenoid. The scapula has a continuous <a href="/wiki/Acromion_process" class="mw-redirect" title="Acromion process">acromion process</a>, as well as an unusual sharp, thin ridge of bone running down its posterior margin, a feature only found in aphanosaurs and <a href="/wiki/Silesauridae" title="Silesauridae">silesaurids</a>.<sup id="cite_ref-Teleo_3-4" class="reference"><a href="#cite_note-Teleo-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> The <a href="/wiki/Coracoid" title="Coracoid">coracoid</a> is small and rounded, and forms a down and rearward facing glenoid fossa with the scapula. The <a href="/wiki/Ilium_(bone)" title="Ilium (bone)">ilium</a> of the <a href="/wiki/Pelvic_girdle" class="mw-redirect" title="Pelvic girdle">pelvic (hip) girdle</a> has a prominent posterior process and supraacetabular crest—a ridge of bone over the <a href="/wiki/Acetabulum" title="Acetabulum">acetabulum</a>. The acetabulum itself is relatively large and almost completely closed, unlike the fully open acetabulum of dinosaurs. The <a href="/wiki/Pubis_(bone)" title="Pubis (bone)">pubis</a> is short and points downwards, with a thickened, flattened end. The <a href="/wiki/Ischium" title="Ischium">ischium</a> is very similar to that of <i>Teleocrater</i>, and is directed down and back from the hips, with a tapered shaft and slightly expanded distal end.<sup id="cite_ref-Teleo2_6-1" class="reference"><a href="#cite_note-Teleo2-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup> The pelvic girdle articulates with the spine via the sacral ribs, of which the first is larger and more robust. The nature of the articulation between the ilium and the sacral ribs suggests the hip was held sub-horizontally and faced ventrolaterally, causing the legs to be positioned down and outwards from the body.<sup id="cite_ref-Sen,_2005_2-5" class="reference"><a href="#cite_note-Sen,_2005-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-:0_4-3" class="reference"><a href="#cite_note-:0-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> </p><p>The limb bones of <i>Yarasuchus</i> are characteristically slender. The <a href="/wiki/Humerus" title="Humerus">humerus</a> is long and cylindrical, with a moderately developed elongated deltopectoral crest that occupies roughly 30% of the length of the bone, similar to the condition in dinosaurs. The <a href="/wiki/Ulna" title="Ulna">ulna</a> is arched somewhat, and is roughly equal in length to the humerus. The <a href="/wiki/Femur" title="Femur">femur</a> is similarly gracile, and has a sigmoidal (s-shaped) curve along its length. The head of the femur is not turned in. A moderately developed fourth trochanter is placed proximally on the femur, unlike the well developed fourth trochanters of dinosaurs. The manus and pes are almost entirely missing, except for the calcaneus of the ankle joint. The calcaneus indicates that <i>Yarasuchus</i> had a "crocodile-normal" ankle, which allowed for more rotation of the foot than the derived hinge-like avemetatarsalian ankle. The forelimb to hindlimb length ratio is roughly 3:5, with much longer back legs than the front. Although the manus is missing, the overall similarity of <i>Teleocrater</i> suggests it would have had a relatively small hand.<sup id="cite_ref-Sen,_2005_2-6" class="reference"><a href="#cite_note-Sen,_2005-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Teleo_3-5" class="reference"><a href="#cite_note-Teleo-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-:0_4-4" class="reference"><a href="#cite_note-:0-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> </p><p>The osteoderms attributed to <i>Yarasuchus</i> are greatly sculptured with a ridge running anteroposteriorly along each one.<sup id="cite_ref-Sen,_2005_2-7" class="reference"><a href="#cite_note-Sen,_2005-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> This is unusual in comparison to <i>Teleocrater,</i> which has been interpreted as lacking any form of similar dermal armour. However, these osteoderms are larger in proportion to the presacral vertebra than would be expected and bear a close similarity to osteoderms of an <a href="/wiki/Erythrosuchidae" title="Erythrosuchidae">erythrosuchid</a> also found in the same formation. Nesbitt and colleagues considered it probable that the osteoderms do not belong <i>Yarasuchus</i> and so it may have also lacked osteoderms like other aphanosaurs and avemetatarsalians.<sup id="cite_ref-Teleo_3-6" class="reference"><a href="#cite_note-Teleo-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Discovery_and_naming">Discovery and naming</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yarasuchus&amp;action=edit&amp;section=4" title="Edit section: Discovery and naming"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><i>Yarasuchus</i> is known from at least two individuals collected from a single 1 square metre (11&#160;sq&#160;ft) assemblage in the <a href="/wiki/Yerrapalli_Formation" title="Yerrapalli Formation">Yerrapalli Formation</a>, located near the Bhimaram village in the <a href="/wiki/Adilabad_district" title="Adilabad district">Adilabad district</a> of <a href="/wiki/India" title="India">India</a>, in a layer of fine red mudstone. The material was found disarticulated, however it represents the majority of the skeleton, missing only the distal caudal vertebrae, <a href="/wiki/Radius_(bone)" title="Radius (bone)">radius</a>, <a href="/wiki/Fibula" title="Fibula">fibula</a>, manus, and most of the pes and skull. The name is derived from 'Yara', meaning red in the local dialect, and the Greek <i>suchos</i> ("crocodile"), referring to the red mudstones the fossils were discovered in. The specific name refers to the <a href="/wiki/Deccan_(disambiguation)" class="mw-disambig" title="Deccan (disambiguation)">Deccan region</a> of India, where the Yerrapalli Formation and surrounding <a href="/wiki/Pranhita-Godavari_Basin" class="mw-redirect" title="Pranhita-Godavari Basin">Pranhita-Godavari Basin</a> are situated.<sup id="cite_ref-Sen,_2005_2-8" class="reference"><a href="#cite_note-Sen,_2005-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> The fossils were also found in association with two specimens of the allokotosaur <i><a href="/wiki/Pamelaria" title="Pamelaria">Pamelaria</a>.</i><sup id="cite_ref-Sen,_2005_2-9" class="reference"><a href="#cite_note-Sen,_2005-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-:0_4-5" class="reference"><a href="#cite_note-:0-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-SK03_7-0" class="reference"><a href="#cite_note-SK03-7"><span class="cite-bracket">&#91;</span>7<span class="cite-bracket">&#93;</span></a></sup> It was initially reported on in a 1993 thesis by Dasgupta, who considered it an indeterminate <a href="/wiki/Rauisuchidae" title="Rauisuchidae">rauisuchid</a>,<sup id="cite_ref-:0_4-6" class="reference"><a href="#cite_note-:0-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> before being officially described and named as a new taxon by Sen in 2005.<sup id="cite_ref-Sen,_2005_2-10" class="reference"><a href="#cite_note-Sen,_2005-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> All the material is held at the Geology Museum of the <a href="/wiki/Indian_Statistical_Institute" title="Indian Statistical Institute">Indian Statistical Institute</a> in <a href="/wiki/Kolkata" title="Kolkata">Kolkata</a>, India.<sup id="cite_ref-Sen,_2005_2-11" class="reference"><a href="#cite_note-Sen,_2005-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Teleo_3-7" class="reference"><a href="#cite_note-Teleo-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-:0_4-7" class="reference"><a href="#cite_note-:0-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> </p><p>In 2016, the material was re-examined by Nesbitt and colleagues in their description of <i>Teleocrater</i>, which revealed a number of previously unrecognised anatomical characteristics. This re-evaluation prompted the inclusion of <i>Yarasuchus</i> within the newly recognised avemetatarsalian clade Aphanosauria. In their examination, Nesbitt and colleagues were able to refer a number of previously undescribed <a href="/wiki/Calcaneus" title="Calcaneus">calcanea</a> collected at the site to the hypodigm of <i>Yarasuchus</i> based on their similarity to <i>Teleocrater</i>, and identified ischia that were originally reported as missing. They also removed a number of erroneously referred skull bones and osteoderms from the hypodigm. The relatively complete skeleton of <i>Yarasuchus</i> allowed Nesbitt and colleagues to confidently refer a number of isolated bones as all belonging to specimens of <i>Teleocrater</i> because of their close similarity to those of <i>Yarasuchus</i>, which provided a key reference point for understanding the anatomy of the previously enigmatic taxon.<sup id="cite_ref-Teleo_3-8" class="reference"><a href="#cite_note-Teleo-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Teleo2_6-2" class="reference"><a href="#cite_note-Teleo2-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Classification">Classification</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yarasuchus&amp;action=edit&amp;section=5" title="Edit section: Classification"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><i>Yarasuchus</i> was originally described as a "<a href="/wiki/Prestosuchid" class="mw-redirect" title="Prestosuchid">prestosuchid</a>", similar to <i><a href="/wiki/Prestosuchus" title="Prestosuchus">Prestosuchus</a></i>, <i><a href="/wiki/Ticinosuchus" title="Ticinosuchus">Ticinosuchus</a>,</i> and <i><a href="/wiki/Mandasuchus" title="Mandasuchus">Mandasuchus</a></i> by Sen, however no <a href="/wiki/Phylogenetic_analysis" class="mw-redirect" title="Phylogenetic analysis">phylogenetic analysis</a> was performed and the referral was made based on general morphological similarities, despite the already debated validity of "Prestosuchidae".<sup id="cite_ref-Sen,_2005_2-12" class="reference"><a href="#cite_note-Sen,_2005-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> In 2010, <a href="/wiki/Stephen_L._Brusatte" title="Stephen L. Brusatte">Brusatte</a> and colleagues conclusively demonstrated that "Prestosuchidae" was a <a href="/wiki/Paraphyletic" class="mw-redirect" title="Paraphyletic">paraphyletic</a> <a href="/wiki/Evolutionary_grade" title="Evolutionary grade">grade</a> of <a href="/wiki/Paracrocodylomorpha" title="Paracrocodylomorpha">paracrocodylomorphs</a>, and that the supposed shared characteristics of the group were in fact found throughout <a href="/wiki/Pseudosuchia" title="Pseudosuchia">Pseudosuchia</a>. They also performed a detailed <a href="/wiki/Cladistics" title="Cladistics">cladistic</a> analysis of fossil Triassic archosaurs, which instead found <i>Yarasuchus</i> to be a basal member of <a href="/wiki/Poposauroidea" title="Poposauroidea">Poposauroidea</a>, although support for this position was weak.<sup id="cite_ref-Betal10_8-0" class="reference"><a href="#cite_note-Betal10-8"><span class="cite-bracket">&#91;</span>8<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Qianosuchus_BW.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8d/Qianosuchus_BW.jpg/220px-Qianosuchus_BW.jpg" decoding="async" width="220" height="60" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8d/Qianosuchus_BW.jpg/330px-Qianosuchus_BW.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8d/Qianosuchus_BW.jpg/440px-Qianosuchus_BW.jpg 2x" data-file-width="759" data-file-height="207" /></a><figcaption>Restoration of <i><a href="/wiki/Qianosuchus" title="Qianosuchus">Qianosuchus</a></i>, a basal <a href="/wiki/Poposauroid" class="mw-redirect" title="Poposauroid">poposauroid</a> formerly considered to be a close relative of <i>Yarasuchus</i>.</figcaption></figure> <p>The position of <i>Yarasuchus</i> in subsequent studies has been similarly unstable,<sup id="cite_ref-Butleretal11_9-0" class="reference"><a href="#cite_note-Butleretal11-9"><span class="cite-bracket">&#91;</span>9<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Decuriasuchus_10-0" class="reference"><a href="#cite_note-Decuriasuchus-10"><span class="cite-bracket">&#91;</span>10<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Nundasuchus_11-0" class="reference"><a href="#cite_note-Nundasuchus-11"><span class="cite-bracket">&#91;</span>11<span class="cite-bracket">&#93;</span></a></sup> though it has nonetheless typically been regarded as a pseudosuchian.<sup id="cite_ref-Desojo09_12-0" class="reference"><a href="#cite_note-Desojo09-12"><span class="cite-bracket">&#91;</span>12<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Rauisuchus_13-0" class="reference"><a href="#cite_note-Rauisuchus-13"><span class="cite-bracket">&#91;</span>13<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Chatterjee2017_14-0" class="reference"><a href="#cite_note-Chatterjee2017-14"><span class="cite-bracket">&#91;</span>14<span class="cite-bracket">&#93;</span></a></sup> A study by Ezcurra on archosauromorph phylogeny in 2016 found <i>Yarasuchus</i> in a position outside of Archosauria, clading together with <i>Dongusuchus</i> in a <a href="/wiki/Polytomy" title="Polytomy">polytomy</a> with <i><a href="/wiki/Euparkeria" title="Euparkeria">Euparkeria</a></i> and a clade made up of <a href="/wiki/Proterochampsia" title="Proterochampsia">Proterochampsia</a> and archosaurs.<sup id="cite_ref-Ezcurra2016_15-0" class="reference"><a href="#cite_note-Ezcurra2016-15"><span class="cite-bracket">&#91;</span>15<span class="cite-bracket">&#93;</span></a></sup> The left cladogram depicts the results of Brusatte and colleagues in 2010,<sup id="cite_ref-Betal10_8-1" class="reference"><a href="#cite_note-Betal10-8"><span class="cite-bracket">&#91;</span>8<span class="cite-bracket">&#93;</span></a></sup> while the right cladogram depicts that of Ezcurra in 2016:<sup id="cite_ref-Ezcurra2016_15-1" class="reference"><a href="#cite_note-Ezcurra2016-15"><span class="cite-bracket">&#91;</span>15<span class="cite-bracket">&#93;</span></a></sup> </p> <table width="75%"> <tbody><tr valign="top"> <td align="left" width="50%"> <div class="clade"><style data-mw-deduplicate="TemplateStyles:r1258728058">body.skin-vector-2022 .mw-parser-output div.clade,body.skin-minerva .mw-parser-output div.clade{overflow-x:auto;overflow-y:hidden}body.skin-minerva .mw-parser-output div.clade p{font-size:inherit}.mw-parser-output table.clade{border-spacing:0;margin:0;font-size:100%;line-height:100%;border-collapse:separate;width:auto}.mw-parser-output table.clade table.clade{width:100%;line-height:inherit}.mw-parser-output table.clade td.clade-label{min-width:0.2em;width:0.2em;padding:0.1em 0.25em;vertical-align:bottom;text-align:center;border-left:1px solid;border-bottom:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-label::before,.mw-parser-output table.clade td.clade-slabel::before{content:"\2060 "}.mw-parser-output table.clade td.clade-fixed-width{overflow:hidden;text-overflow:ellipsis}.mw-parser-output table.clade td.clade-fixed-width:hover{overflow:visible}.mw-parser-output table.clade td.clade-label.first{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-label.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-slabel{padding:0.1em 0.25em;vertical-align:top;text-align:center;border-left:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-slabel:hover{overflow:visible}.mw-parser-output table.clade td.clade-slabel.last{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-slabel.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-bar{vertical-align:middle;text-align:left;padding:0 0.5em;position:relative}.mw-parser-output table.clade td.clade-bar.reverse{text-align:right;position:relative}.mw-parser-output table.clade td.clade-leaf{border:0;padding:0;text-align:left}.mw-parser-output table.clade td.clade-leafR{border:0;padding:0;text-align:right}.mw-parser-output table.clade td.clade-leaf.reverse{text-align:right}.mw-parser-output table.clade:hover span.linkA{background-color:yellow}.mw-parser-output table.clade:hover span.linkB{background-color:green}</style> <table class="clade" style="font-size:85%;line-height:85%;"> <tbody><tr> <td class="clade-label first">&#160;<a href="/wiki/Poposauroidea" title="Poposauroidea">Poposauroidea</a>&#160; </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><b>Yarasuchus</b></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Qianosuchus" title="Qianosuchus">Qianosuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Arizonasaurus" title="Arizonasaurus">Arizonasaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Bromsgroveia" title="Bromsgroveia">Bromsgroveia</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Lotosaurus" title="Lotosaurus">Lotosaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Poposaurus" title="Poposaurus">Poposaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Sillosuchus" title="Sillosuchus">Sillosuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label">&#160;<a href="/wiki/Shuvosauridae" title="Shuvosauridae">Shuvosauridae</a>&#160; </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Effigia" title="Effigia">Effigia</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Shuvosaurus" title="Shuvosaurus">Shuvosaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td> <td align="left" width="50%"> <div class="clade"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade" style="font-size:85%;line-height:85%;"> <tbody><tr> <td class="clade-label first">&#160;<a href="/wiki/Eucrocopoda" class="mw-redirect" title="Eucrocopoda">Eucrocopoda</a>&#160; </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Dorosuchus" title="Dorosuchus">Dorosuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Euparkeria" title="Euparkeria">Euparkeria</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Dongusuchus" title="Dongusuchus">Dongusuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><b>Yarasuchus</b></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Proterochampsia" title="Proterochampsia">Proterochampsia</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Doswelliidae" title="Doswelliidae">Doswelliidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Proterochampsidae" title="Proterochampsidae">Proterochampsidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Archosauria" class="mw-redirect" title="Archosauria">Archosauria</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Ornithodira" class="mw-redirect" title="Ornithodira">Ornithodira</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Pseudosuchia" title="Pseudosuchia">Pseudosuchia</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr></tbody></table> <p>Further complicating the issue were suggestions that the hypodigm of <i>Yarasuchus</i> was a chimeric assemblage of material from both a "rauisuchian" archosaur and a <a href="/wiki/Prolacertiformes" class="mw-redirect" title="Prolacertiformes">prolacertiform</a> archosauromorph, if not including material from yet other archosauromorphs.<sup id="cite_ref-Chimera_16-0" class="reference"><a href="#cite_note-Chimera-16"><span class="cite-bracket">&#91;</span>16<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Rauisuchia_17-0" class="reference"><a href="#cite_note-Rauisuchia-17"><span class="cite-bracket">&#91;</span>17<span class="cite-bracket">&#93;</span></a></sup> However, later examination of the material by Ezcurra found that none of it could be assigned to either a "rauisuchian" or prolacertiform, and that they all likely pertained to a single non-<a href="/wiki/Suchia" title="Suchia">suchian</a> archosauriform taxon.<sup id="cite_ref-Ezcurra2016_15-2" class="reference"><a href="#cite_note-Ezcurra2016-15"><span class="cite-bracket">&#91;</span>15<span class="cite-bracket">&#93;</span></a></sup> </p><p>In 2017, <i>Yarasuchus</i> was included in an updated analysis of Triassic archosauromorphs by Nesbitt and colleagues in their official description of <i>Teleocrater</i>, utilising two modified datasets, those of Nesbitt (2011)<sup id="cite_ref-NSJ11_18-0" class="reference"><a href="#cite_note-NSJ11-18"><span class="cite-bracket">&#91;</span>18<span class="cite-bracket">&#93;</span></a></sup> and Ezcurra (2016).<sup id="cite_ref-Ezcurra2016_15-3" class="reference"><a href="#cite_note-Ezcurra2016-15"><span class="cite-bracket">&#91;</span>15<span class="cite-bracket">&#93;</span></a></sup> Both analyses recovered <i>Yarasuchus</i> in a newly recognised clade they named Aphanosauria, which included <i>Yarasuchus</i> in a polytomy with <i>Teleocrater</i> and <i>Dongusuchus</i>, as well as <i>Spondylosoma</i>. Indeed, the clade Aphanosauria was cladistically defined as "the most inclusive clade containing <i>Teleocrater rhadinus</i> and <i>Yarasuchus deccanensis</i> but not <i><a href="/wiki/House_sparrow" title="House sparrow">Passer domesticus</a></i> or <i><a href="/wiki/Nile_crocodile" title="Nile crocodile">Crocodylus niloticus</a>"</i>. Aphanosaurs were found to be the earliest diverging clade of <a href="/wiki/Avemetatarsalia" title="Avemetatarsalia">Avemetatarsalia</a>, sister taxon to the clade <a href="/wiki/Ornithodira" class="mw-redirect" title="Ornithodira">Ornithodira</a> that includes the <a href="/wiki/Pterosauria" class="mw-redirect" title="Pterosauria">Pterosauria</a> and <a href="/wiki/Dinosauromorpha" title="Dinosauromorpha">Dinosauromorpha</a>.<sup id="cite_ref-Teleo_3-9" class="reference"><a href="#cite_note-Teleo-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> The results of their analyses are reproduced and simplified below, combining the general topology of the Ezcurra dataset with the Nesbitt dataset results for Avemetatarsalia. </p> <div class="clade"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade" style="font-size:85%; line-height:85%"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Archosauriformes" title="Archosauriformes">Archosauriformes</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Proterosuchidae" title="Proterosuchidae">Proterosuchidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Erythrosuchidae" title="Erythrosuchidae">Erythrosuchidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Archosauriformes" title="Archosauriformes">Eucrocopoda</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Proterochampsia" title="Proterochampsia">Proterochampsia</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Archosauria" class="mw-redirect" title="Archosauria">Archosauria</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Pseudosuchia" title="Pseudosuchia">Pseudosuchia</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Avemetatarsalia" title="Avemetatarsalia">Avemetatarsalia</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Aphanosauria" title="Aphanosauria">Aphanosauria</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Spondylosoma" title="Spondylosoma">Spondylosoma</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><b>Yarasuchus</b></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Dongusuchus" title="Dongusuchus">Dongusuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Teleocrater" title="Teleocrater">Teleocrater</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Avemetatarsalia" title="Avemetatarsalia">Ornithodira</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Pterosauria" class="mw-redirect" title="Pterosauria">Pterosauria</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Lagerpetidae" title="Lagerpetidae">Lagerpetidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Dinosauriformes" class="mw-redirect" title="Dinosauriformes">Dinosauriformes</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Marasuchus" title="Marasuchus">Marasuchus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Silesauridae" title="Silesauridae">Silesauridae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Dinosauria" class="mw-redirect" title="Dinosauria">Dinosauria</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> <div class="mw-heading mw-heading3"><h3 id="Evolutionary_significance">Evolutionary significance</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yarasuchus&amp;action=edit&amp;section=6" title="Edit section: Evolutionary significance"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Teleocrater_v1.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a6/Teleocrater_v1.png/220px-Teleocrater_v1.png" decoding="async" width="220" height="59" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/a6/Teleocrater_v1.png/330px-Teleocrater_v1.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/a6/Teleocrater_v1.png/440px-Teleocrater_v1.png 2x" data-file-width="1800" data-file-height="481" /></a><figcaption>Life restoration of the closely related aphanosaur <i><a href="/wiki/Teleocrater" title="Teleocrater">Teleocrater</a></i>.</figcaption></figure> <p>The recognition of a close relationship between <i>Yarasuchus</i>, <i>Teleocrater</i> and other aphanosaurs and their relation to other avemetatarsalians settled a number of the unusual anatomical features of <i>Yarasuchus</i>. Many of the previously unique features of <i>Yarasuchus</i> unite it with other aphanosaurs, including the elongated neck, high neural spines, three-headed cervical ribs and slender <a href="/wiki/Appendicular_skeleton" title="Appendicular skeleton">appendicular skeleton</a>. <i>Yarasuchus</i> and the other aphanosaurs play a significant role in our understanding of early avemetatarsalian evolution, exemplified in the relatively completely known anatomy of <i>Yarasuchus</i>. The structure of the foot, particularly the bones of the ankle (such as the calcaneus), demonstrate that avemetatarsalians evolved from ancestors with 'crocodile-normal' ankles, unlike the simple hinge-like ankles characteristic of derived ornithodirans. The 'crocodile-normal' ankle was once thought to be unique to pseudosuchians, but its presence in aphanosaurs like <i>Yarasuchus</i> imply that the evolution of the avemetatarsalian ankle was a more complicated process than initially believed.<sup id="cite_ref-Teleo_3-10" class="reference"><a href="#cite_note-Teleo-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> The anatomy of <i>Yarasuchus</i> also demonstrates that other typical avemetatarsalian features, such as slender limb girdles, had evolved prior to the eponymous 'advanced mesotarsal' ankles.<sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">&#91;</span>19<span class="cite-bracket">&#93;</span></a></sup> </p><p>Unusually, <i>Yarasuchus</i> and other aphanosaurs share a number of features convergently evolved with poposauroids. Aphanosaurs and poposauroids share only one unique trait (the presence of an accessory articulation facet just above the parapophysis of the cervicals for the three-headed cervical rib), however they have also convergently acquired a similar set of traits that are found throughout archosaurs. The previous phylogenetic position of <i>Yarasuchus</i> as a poposauroid by Brusatte and colleagues was likely due to this convergence.<sup id="cite_ref-Teleo2_6-3" class="reference"><a href="#cite_note-Teleo2-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup> The convergence between <i>Yarasuchus</i> and poposauroids could be attributed to the broader trend of poposauroids converging on <a href="/wiki/Coelurosaur" class="mw-redirect" title="Coelurosaur">coelurosaurian</a> <a href="/wiki/Theropods" class="mw-redirect" title="Theropods">theropods</a>, a derived clade of avemetatarsalians.<sup id="cite_ref-Nesbitt2007_20-0" class="reference"><a href="#cite_note-Nesbitt2007-20"><span class="cite-bracket">&#91;</span>20<span class="cite-bracket">&#93;</span></a></sup> The similarity between poposauroids and aphanosaurs like <i>Yarasuchus</i> means it is difficult to determine the identity of isolated archosaur material that has features present in both groups, particularly as aphanosaurs are the earliest diverging avemetatarsalians while poposauroids are the oldest known pseudosuchians and so their <a href="/wiki/Stratigraphic" class="mw-redirect" title="Stratigraphic">stratigraphic</a> ranges broadly overlap with each other.<sup id="cite_ref-Teleo2_6-4" class="reference"><a href="#cite_note-Teleo2-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Palaeobiology">Palaeobiology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yarasuchus&amp;action=edit&amp;section=7" title="Edit section: Palaeobiology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><i>Yarasuchus</i> was proposed to be facultatively bipedal by Dasgupta in 1993 on the basis of its gracile body, slender shoulder girdle and proportionately short forelimbs, among other features, and this suggestion was repeated by Sen in its official description in 2005.<sup id="cite_ref-Sen,_2005_2-13" class="reference"><a href="#cite_note-Sen,_2005-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-:0_4-8" class="reference"><a href="#cite_note-:0-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> The closely related <i>Teleocrater</i> has since been interpreted to have been a quadruped, and as <i>Yarasuchus</i> has similar limb proportions, it likely was as well.<sup id="cite_ref-Teleo_3-11" class="reference"><a href="#cite_note-Teleo-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> The lack of definitive jaw material leaves the diet of <i>Yarasuchus</i> ambiguous, however the teeth of <i>Teleocrater</i> imply aphanosaurs were carnivorous, as with other early avemetatarsalians. Furthermore, <a href="/wiki/Histology" title="Histology">histological</a> samples from <i>Teleocrater</i> show that its growth rates were more similar to those of other avemetatarsalians compared to pseudosuchians and stem-archosaurs, and so <i>Yarasuchus</i> may have also had a similarly higher growth rate.<sup id="cite_ref-Teleo_3-12" class="reference"><a href="#cite_note-Teleo-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Palaeoecology">Palaeoecology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yarasuchus&amp;action=edit&amp;section=8" title="Edit section: Palaeoecology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Pamelaria_dolichotrachela.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/dc/Pamelaria_dolichotrachela.jpg/220px-Pamelaria_dolichotrachela.jpg" decoding="async" width="220" height="111" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/dc/Pamelaria_dolichotrachela.jpg/330px-Pamelaria_dolichotrachela.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/dc/Pamelaria_dolichotrachela.jpg/440px-Pamelaria_dolichotrachela.jpg 2x" data-file-width="2814" data-file-height="1414" /></a><figcaption>Restoration of the contemporary allokotosaur <i><a href="/wiki/Pamelaria" title="Pamelaria">Pamelaria</a></i>.</figcaption></figure> <p>Many other vertebrate remains have been found from the Yerrapalli Formation alongside those of <i>Yarasuchus</i>, and would have coexisted with it during the Middle Triassic. Remains of the allokotosaur <i>Pamelaria</i> in particular have been found in close proximity to those of <i>Yarasuchus</i>.<sup id="cite_ref-SK03_7-1" class="reference"><a href="#cite_note-SK03-7"><span class="cite-bracket">&#91;</span>7<span class="cite-bracket">&#93;</span></a></sup> Other vertebrate remains include those of the <a href="/wiki/Lungfish" title="Lungfish">lungfish</a> <i><a href="/wiki/Ceratodus_(genus)" class="mw-redirect" title="Ceratodus (genus)">Ceratodus</a></i>, the <a href="/wiki/Actinopterygii" title="Actinopterygii">actinopterygian</a> fish <i><a href="/wiki/Saurichthys" title="Saurichthys">Saurichthys</a></i>, the <a href="/wiki/Temnospondyl" class="mw-redirect" title="Temnospondyl">temnospondyl</a> <i><a href="/wiki/Parotosuchus" title="Parotosuchus">Parotosuchus</a></i>, the <a href="/wiki/Dicynodont" class="mw-redirect" title="Dicynodont">dicynodonts</a> <i><a href="/wiki/Rechnisaurus" title="Rechnisaurus">Rechnisaurus</a></i> and <i><a href="/wiki/Wadiasaurus" title="Wadiasaurus">Wadiasaurus</a></i>, the <a href="/wiki/Rhynchosaur" title="Rhynchosaur">rhynchosaur</a> <i><a href="/wiki/Mesodapedon" title="Mesodapedon">Mesodapedon</a></i>, and a large undescribed erythrosuchid.<sup id="cite_ref-BS99_1-1" class="reference"><a href="#cite_note-BS99-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-:0_4-9" class="reference"><a href="#cite_note-:0-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Chatterjee2017_14-1" class="reference"><a href="#cite_note-Chatterjee2017-14"><span class="cite-bracket">&#91;</span>14<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Wadiasaurus1DB.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/9/93/Wadiasaurus1DB.jpg/220px-Wadiasaurus1DB.jpg" decoding="async" width="220" height="132" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/93/Wadiasaurus1DB.jpg/330px-Wadiasaurus1DB.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/93/Wadiasaurus1DB.jpg/440px-Wadiasaurus1DB.jpg 2x" data-file-width="1400" data-file-height="839" /></a><figcaption>Restoration of the contemporary dicynodont <i><a href="/wiki/Wadiasaurus" title="Wadiasaurus">Wadiasaurus</a></i>.</figcaption></figure> <p>The anatomical similarity between <i>Yarasuchus</i> and <i>Teleocrater</i> is mirrored by the similarities in fauna and environment shared between the Yerrapalli Formation and the <a href="/wiki/Manda_Formation" title="Manda Formation">Manda Formation</a> in <a href="/wiki/Tanzania" title="Tanzania">Tanzania</a>, and both are estimated to be Anisian in age.<sup id="cite_ref-BS99_1-2" class="reference"><a href="#cite_note-BS99-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Sen,_2005_2-14" class="reference"><a href="#cite_note-Sen,_2005-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> At the time, India was still a part of the supercontinent <a href="/wiki/Gondwana" title="Gondwana">Gondwana</a> in <a href="/wiki/Pangaea" title="Pangaea">Pangaea</a>, and was located directly adjacent to East Africa. This demonstrates that early avemetatarsalians like <i>Yarasuchus</i> were geographically widespread in the Middle Triassic, as with other archosauriforms, in contrast to previous suggestions that pseudosuchians were more diverse<i>.</i> The close similarity between <i>Yarasuchus</i> and the Russian <i>Dongusuchus</i> further supports this, indicating <a href="/wiki/Biogeographical" class="mw-redirect" title="Biogeographical">biogeographical</a> affinities between India and Russia despite their widely separated <a href="/wiki/Palaeolatitude" class="mw-redirect" title="Palaeolatitude">palaeolatitudes</a>.<sup id="cite_ref-Teleo_3-13" class="reference"><a href="#cite_note-Teleo-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Teleo2_6-5" class="reference"><a href="#cite_note-Teleo2-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Ezcurra2016_15-4" class="reference"><a href="#cite_note-Ezcurra2016-15"><span class="cite-bracket">&#91;</span>15<span class="cite-bracket">&#93;</span></a></sup> </p><p>The sediments of the Yerrapalli Formation are interpreted as fluvial deposits, indicative of a broad, interchannel floodplain environment with seasonal ephemeral stream channels. The climate is thought to have been hot and dry with seasonal rainfall. This is consistent with the preservation state of the fossils, as the remains of <i>Yarasuchus</i> were found dismembered and disarticulated, suggesting the material was left exposed at the surface for a period before being buried by suspended fluvial sediments.<sup id="cite_ref-Taphonomy_21-0" class="reference"><a href="#cite_note-Taphonomy-21"><span class="cite-bracket">&#91;</span>21<span class="cite-bracket">&#93;</span></a></sup> There are few plant fossils known from the Yerrapalli Formation, however this is not believed to be due to it being an arid environment, but rather due to the hot and dry conditions being unsuitable to the fossilisation of plant material.<sup id="cite_ref-:0_4-10" class="reference"><a href="#cite_note-:0-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yarasuchus&amp;action=edit&amp;section=9" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-BS99-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-BS99_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-BS99_1-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-BS99_1-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text">Bandyopadhyay, S. and Sengupta, D. P. (1999). Middle Triassic vertebrate faunas from India. <i><a href="/wiki/Journal_of_African_Earth_Sciences" title="Journal of African Earth Sciences">Journal of African Earth Sciences</a></i> <b>29</b>: 233–241.</span> </li> <li id="cite_note-Sen,_2005-2"><span class="mw-cite-backlink">^ <a href="#cite_ref-Sen,_2005_2-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Sen,_2005_2-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Sen,_2005_2-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-Sen,_2005_2-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-Sen,_2005_2-4"><sup><i><b>e</b></i></sup></a> <a href="#cite_ref-Sen,_2005_2-5"><sup><i><b>f</b></i></sup></a> <a href="#cite_ref-Sen,_2005_2-6"><sup><i><b>g</b></i></sup></a> <a href="#cite_ref-Sen,_2005_2-7"><sup><i><b>h</b></i></sup></a> <a href="#cite_ref-Sen,_2005_2-8"><sup><i><b>i</b></i></sup></a> <a href="#cite_ref-Sen,_2005_2-9"><sup><i><b>j</b></i></sup></a> <a href="#cite_ref-Sen,_2005_2-10"><sup><i><b>k</b></i></sup></a> <a href="#cite_ref-Sen,_2005_2-11"><sup><i><b>l</b></i></sup></a> <a href="#cite_ref-Sen,_2005_2-12"><sup><i><b>m</b></i></sup></a> <a href="#cite_ref-Sen,_2005_2-13"><sup><i><b>n</b></i></sup></a> <a href="#cite_ref-Sen,_2005_2-14"><sup><i><b>o</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output 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(2013). <a rel="nofollow" class="external text" href="http://sp.lyellcollection.org/content/379/1/241">"Rauisuchia"</a>. <i><a href="/wiki/Geological_Society,_London,_Special_Publications" class="mw-redirect" title="Geological Society, London, Special Publications">Geological Society, London, Special Publications</a></i>. <b>379</b> (1): 241–274. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2013GSLSP.379..241N">2013GSLSP.379..241N</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1144%2FSP379.1">10.1144/SP379.1</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a>&#160;<a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:219193351">219193351</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Geological+Society%2C+London%2C+Special+Publications&amp;rft.atitle=Rauisuchia&amp;rft.volume=379&amp;rft.issue=1&amp;rft.pages=241-274&amp;rft.date=2013&amp;rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A219193351%23id-name%3DS2CID&amp;rft_id=info%3Adoi%2F10.1144%2FSP379.1&amp;rft_id=info%3Abibcode%2F2013GSLSP.379..241N&amp;rft.aulast=Nesbitt&amp;rft.aufirst=S.J.&amp;rft.au=Brusatte%2C+S.L.&amp;rft.au=Desojo%2C+J.B.&amp;rft.au=Liparini%2C+A.&amp;rft.au=De+Fran%C3%A7a%2C+M.A.G.&amp;rft.au=Weinbaum%2C+J.C.&amp;rft.au=Gower%2C+D.J.&amp;rft_id=http%3A%2F%2Fsp.lyellcollection.org%2Fcontent%2F379%2F1%2F241&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AYarasuchus" class="Z3988"></span></span> </li> <li id="cite_note-NSJ11-18"><span class="mw-cite-backlink"><b><a href="#cite_ref-NSJ11_18-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFNesbitt2011" class="citation journal cs1">Nesbitt, S.J. 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title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Sedimentary+Geology&amp;rft.atitle=Taphonomy+of+some+Gondwana+vertebrate+assemblages+of+India.&amp;rft.volume=147&amp;rft.issue=1%E2%80%932&amp;rft.pages=219-245&amp;rft.date=2002&amp;rft_id=info%3Adoi%2F10.1016%2FS0037-0738%2801%2900198-1&amp;rft_id=info%3Abibcode%2F2002SedG..147..219B&amp;rft.aulast=Brandyopadhyay&amp;rft.aufirst=S.&amp;rft.au=Roy+Chowdhury%2C+T.K.&amp;rft.au=Sengupta%2C+D.P.&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AYarasuchus" class="Z3988"></span></span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Yarasuchus&amp;action=edit&amp;section=10" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=148342"><i>Yarasuchus</i></a> in the <a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a></li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol ol,.mw-parser-output .hlist ol ul,.mw-parser-output .hlist ul dl,.mw-parser-output .hlist ul ol,.mw-parser-output .hlist ul ul{display:inline}.mw-parser-output .hlist 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.navbox-title{background-color:#ddf}.mw-parser-output .navbox-subgroup .navbox-group,.mw-parser-output .navbox-subgroup .navbox-abovebelow{background-color:#e6e6ff}.mw-parser-output .navbox-even{background-color:#f7f7f7}.mw-parser-output .navbox-odd{background-color:transparent}.mw-parser-output .navbox .hlist td dl,.mw-parser-output .navbox .hlist td ol,.mw-parser-output .navbox .hlist td ul,.mw-parser-output .navbox td.hlist dl,.mw-parser-output .navbox td.hlist ol,.mw-parser-output .navbox td.hlist ul{padding:0.125em 0}.mw-parser-output .navbox .navbar{display:block;font-size:100%}.mw-parser-output .navbox-title .navbar{float:left;text-align:left;margin-right:0.5em}body.skin--responsive .mw-parser-output .navbox-image img{max-width:none!important}@media print{body.ns-0 .mw-parser-output .navbox{display:none!important}}</style></div><div role="navigation" class="navbox" aria-labelledby="Avemetatarsalia_/_Pan-Aves" style="padding:3px"><table class="nowraplinks hlist mw-collapsible mw-collapsed navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Avemetatarsalia" title="Template:Avemetatarsalia"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Avemetatarsalia" title="Template talk:Avemetatarsalia"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Avemetatarsalia" title="Special:EditPage/Template:Avemetatarsalia"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Avemetatarsalia_/_Pan-Aves" style="font-size:114%;margin:0 4em"><a href="/wiki/Avemetatarsalia" title="Avemetatarsalia">Avemetatarsalia / Pan-Aves</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="*_Kingdom:_Animalia_*_Phylum:_Chordata_*_Class:_Sauropsida_*_Clade:_Archosauria" style="font-size:114%;margin:0 4em"><span style="font-weight:normal; font-style:normal"><span style="font-size:85%;"> <ul><li>Kingdom: <a href="/wiki/Animal" title="Animal">Animalia</a></li> <li>Phylum: <a href="/wiki/Chordate" title="Chordate">Chordata</a></li> <li>Class: <a href="/wiki/Sauropsida" title="Sauropsida">Sauropsida</a></li> <li><i>Clade</i>: <a href="/wiki/Archosaur" title="Archosaur">Archosauria</a></li></ul> </span></span></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Sauropsida" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Sauropsida" title="Sauropsida">Sauropsida</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Sauropsida" title="Template:Sauropsida">Sauropsida</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Archosauromorpha" title="Archosauromorpha">Archosauromorpha</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Archosauromorpha" title="Template:Archosauromorpha">Archosauromorpha</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Avemetatarsalia" title="Avemetatarsalia">Avemetatarsalia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2" style="background:#ddddff;font-size:115%"><div><b><a href="/wiki/Avemetatarsalia" title="Avemetatarsalia">Avemetatarsalia</a></b></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Avemetatarsalia" scope="row" class="navbox-group" style="width:1%"><div style="position: absolute; transform: translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Avemetatarsalia" title="Avemetatarsalia">Avemetatarsalia</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Incertovenator" title="Incertovenator">Incertovenator</a></i>?</li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Mambachiton" title="Mambachiton">Mambachiton</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Aphanosauria" title="Aphanosauria">Aphanosauria</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Incertovenator" title="Incertovenator">Incertovenator</a></i>?</li> <li><i><a href="/wiki/Dongusuchus" title="Dongusuchus">Dongusuchus</a></i></li> <li><i><a href="/wiki/Spondylosoma" title="Spondylosoma">Spondylosoma</a></i></li> <li><i><a href="/wiki/Teleocrater" title="Teleocrater">Teleocrater</a></i></li> <li><i><a class="mw-selflink selflink">Yarasuchus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position: absolute; transform: translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Ornithodira" class="mw-redirect" title="Ornithodira">Ornithodira</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="†Pterosauromorpha" scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Pterosauromorpha" title="Pterosauromorpha">Pterosauromorpha</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Lagerpetidae" title="Lagerpetidae">Lagerpetidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Dromomeron" title="Dromomeron">Dromomeron</a></i></li> <li><i><a href="/wiki/Faxinalipterus" title="Faxinalipterus">Faxinalipterus</a></i>?</li> <li><i><a href="/wiki/Ixalerpeton" title="Ixalerpeton">Ixalerpeton</a></i></li> <li><i><a href="/wiki/Kongonaphon" title="Kongonaphon">Kongonaphon</a></i></li> <li><i><a href="/wiki/Lagerpeton" title="Lagerpeton">Lagerpeton</a></i></li> <li><i><a href="/wiki/Scleromochlus" title="Scleromochlus">Scleromochlus</a></i>?</li> <li><i><a href="/wiki/Venetoraptor" title="Venetoraptor">Venetoraptor</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Pterosaur" title="Pterosaur">Pterosauria</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Pterosauria" title="Template:Pterosauria">Pterosauria</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position: absolute; transform: translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Dinosauromorpha" title="Dinosauromorpha">Dinosauromorpha</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Lagosuchus" title="Lagosuchus">Lagosuchus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Marasuchus" title="Marasuchus">Marasuchus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Nyasasaurus" title="Nyasasaurus">Nyasasaurus</a></i>?</li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Saltopus" title="Saltopus">Saltopus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Silesauridae" title="Silesauridae">Silesauridae</a>?</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Agnosphitys" title="Agnosphitys">Agnosphitys</a></i></li> <li><i><a href="/wiki/Asilisaurus" title="Asilisaurus">Asilisaurus</a></i></li> <li><i><a href="/wiki/Lewisuchus" title="Lewisuchus">Lewisuchus</a></i></li> <li><i><a href="/wiki/Pisanosaurus" title="Pisanosaurus">Pisanosaurus</a></i>?</li> <li><i><a href="/wiki/Soumyasaurus" title="Soumyasaurus">Soumyasaurus</a></i></li> <li><i><a href="/wiki/Technosaurus" title="Technosaurus">Technosaurus</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Sulcimentisauria" class="mw-redirect" title="Sulcimentisauria">Sulcimentisauria</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Amanasaurus" title="Amanasaurus">Amanasaurus</a></i></li> <li><i><a href="/wiki/Diodorus_scytobrachion" title="Diodorus scytobrachion">Diodorus</a></i></li> <li><i><a href="/wiki/Eucoelophysis" title="Eucoelophysis">Eucoelophysis</a></i></li> <li><i><a href="/wiki/Gamatavus" title="Gamatavus">Gamatavus</a></i></li> <li><i><a href="/wiki/Gondwanax" title="Gondwanax">Gondwanax</a></i></li> <li><i><a href="/wiki/Ignotosaurus" title="Ignotosaurus">Ignotosaurus</a></i></li> <li><i><a href="/wiki/Kwanasaurus" title="Kwanasaurus">Kwanasaurus</a></i></li> <li><i><a href="/wiki/Lutungutali" title="Lutungutali">Lutungutali</a></i></li> <li><i><a href="/wiki/Sacisaurus" title="Sacisaurus">Sacisaurus</a></i></li> <li><i><a href="/wiki/Silesaurus" title="Silesaurus">Silesaurus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position: absolute; transform: translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Dinosaur" title="Dinosaur">Dinosauria</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="†Ornithischia" scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Ornithischia" title="Ornithischia">Ornithischia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Ornithischia" title="Template:Ornithischia">Ornithischia</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position: absolute; transform: translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Saurischia" title="Saurischia">Saurischia</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Alwalkeria" title="Alwalkeria">Alwalkeria</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Chindesaurus" title="Chindesaurus">Chindesaurus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Daemonosaurus" title="Daemonosaurus">Daemonosaurus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Eodromaeus" title="Eodromaeus">Eodromaeus</a></i></li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Saltopus" title="Saltopus">Saltopus</a></i>?</li> <li><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><a href="/wiki/Tawa_hallae" title="Tawa hallae">Tawa</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Herrerasauria" class="mw-redirect" title="Herrerasauria">Herrerasauria</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Chindesaurus" title="Chindesaurus">Chindesaurus</a></i>?</li> <li><i><a href="/wiki/Daemonosaurus" title="Daemonosaurus">Daemonosaurus</a></i>?</li> <li><i><a href="/wiki/Tawa_hallae" title="Tawa hallae">Tawa</a></i>?</li> <li><i><a href="/wiki/Caseosaurus" title="Caseosaurus">Caseosaurus</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Herrerasauridae" title="Herrerasauridae">Herrerasauridae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Gnathovorax" title="Gnathovorax">Gnathovorax</a></i></li> <li><i><a href="/wiki/Herrerasaurus" title="Herrerasaurus">Herrerasaurus</a></i></li> <li><i><a href="/wiki/Sanjuansaurus" title="Sanjuansaurus">Sanjuansaurus</a></i></li> <li><i><a href="/wiki/Staurikosaurus" title="Staurikosaurus">Staurikosaurus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Eusaurischia" class="mw-redirect" title="Eusaurischia">Eusaurischia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><a href="/wiki/Sauropodomorpha" title="Sauropodomorpha">Sauropodomorpha</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Sauropodomorpha" title="Template:Sauropodomorpha">Sauropodomorpha</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Theropoda" title="Theropoda">Theropoda</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Theropoda" title="Template:Theropoda">Theropoda</a></li> <li>includes <a href="/wiki/Bird" title="Bird">birds</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Teleocrater_v1.png" class="mw-file-description" title="Teleocrater rhadinus"><img alt="Teleocrater rhadinus" src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a6/Teleocrater_v1.png/140px-Teleocrater_v1.png" decoding="async" width="140" height="37" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/a6/Teleocrater_v1.png/210px-Teleocrater_v1.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/a6/Teleocrater_v1.png/280px-Teleocrater_v1.png 2x" data-file-width="1800" data-file-height="481" /></a></span> <p><span typeof="mw:File"><a href="/wiki/File:Kongonaphon_life_restoration.png" class="mw-file-description" title="Kongonaphon kely"><img alt="Kongonaphon kely" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/7c/Kongonaphon_life_restoration.png/140px-Kongonaphon_life_restoration.png" decoding="async" width="140" height="61" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/7c/Kongonaphon_life_restoration.png/210px-Kongonaphon_life_restoration.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/7c/Kongonaphon_life_restoration.png/280px-Kongonaphon_life_restoration.png 2x" data-file-width="2118" data-file-height="921" /></a></span> <span typeof="mw:File"><a href="/wiki/File:Marasuchus.jpg" class="mw-file-description" title="Marasuchus lilloensis"><img alt="Marasuchus lilloensis" src="//upload.wikimedia.org/wikipedia/commons/thumb/2/21/Marasuchus.jpg/140px-Marasuchus.jpg" decoding="async" width="140" height="71" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/21/Marasuchus.jpg/210px-Marasuchus.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/21/Marasuchus.jpg/280px-Marasuchus.jpg 2x" data-file-width="3275" data-file-height="1668" /></a></span> <span typeof="mw:File"><a href="/wiki/File:Diodorus_restoration.jpg" class="mw-file-description" title="Diodorus scytobrachion"><img alt="Diodorus scytobrachion" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/9f/Diodorus_restoration.jpg/140px-Diodorus_restoration.jpg" decoding="async" width="140" height="70" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/9f/Diodorus_restoration.jpg/210px-Diodorus_restoration.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/9f/Diodorus_restoration.jpg/280px-Diodorus_restoration.jpg 2x" data-file-width="1572" data-file-height="782" /></a></span> </p> <span typeof="mw:File"><a href="/wiki/File:Herresaurus.png" class="mw-file-description" title="Herrerasaurus ischigualastensis"><img alt="Herrerasaurus ischigualastensis" src="//upload.wikimedia.org/wikipedia/commons/thumb/f/fd/Herresaurus.png/140px-Herresaurus.png" decoding="async" width="140" height="49" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/fd/Herresaurus.png/210px-Herresaurus.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/fd/Herresaurus.png/280px-Herresaurus.png 2x" data-file-width="7500" data-file-height="2625" /></a></span></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li>See also: <a href="/wiki/Ornithoscelida" title="Ornithoscelida">Ornithoscelida</a></li> <li><a href="/wiki/Phytodinosauria" title="Phytodinosauria">Phytodinosauria</a></li> <li><span class="noviewer" typeof="mw:File"><span title="Category"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/16px-Symbol_category_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/23px-Symbol_category_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/9/96/Symbol_category_class.svg/31px-Symbol_category_class.svg.png 2x" data-file-width="180" data-file-height="185" /></span></span> <a href="/wiki/Category:Avemetatarsalians" title="Category:Avemetatarsalians">Category</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Yarasuchus</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q28647753" class="extiw" title="wikidata:Q28647753">Q28647753</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/4822224">4822224</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&amp;id=1041543">1041543</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=3605931">3605931</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=148342">148342</a></span></span></li></ul> </div></td></tr></tbody></table></div> <!-- NewPP limit report Parsed by mw‐api‐ext.codfw.main‐7556f8b5dd‐d5jgw Cached time: 20241124180435 Cache 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