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폴 L. 모드리치 - 위키백과, 우리 모두의 백과사전

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id="siteNotice"> <!-- CentralNotice --> </div> </div> <div class="pre-content heading-holder"> <div class="page-heading"> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">폴 L. 모드리치</span></h1> <div class="tagline"></div> </div> <nav class="page-actions-menu"> <ul id="p-views" class="page-actions-menu__list"> <li id="language-selector" class="page-actions-menu__list-item"><a role="button" href="https://ko-m-wikipedia-org.translate.goog/wiki/%ED%8F%B4_L._%EB%AA%A8%EB%93%9C%EB%A6%AC%EC%B9%98?_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB#p-lang" data-mw="interface" data-event-name="menu.languages" title="언어" class="cdx-button cdx-button--size-large cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--icon-only cdx-button--weight-quiet language-selector"> <span class="minerva-icon minerva-icon--language"></span> <span>언어</span> </a></li> <li id="page-actions-watch" class="page-actions-menu__list-item"><a role="button" 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cdx-button--weight-quiet edit-page menu__item--page-actions-edit"> <span class="minerva-icon minerva-icon--edit"></span> <span>편집</span> </a></li> </ul> </nav><!-- version 1.0.2 (change every time you update a partial) --> <div id="mw-content-subtitle"></div> </div> <div id="bodyContent" class="content"> <div id="mw-content-text" class="mw-body-content"> <script>function mfTempOpenSection(id){var block=document.getElementById("mf-section-"+id);block.className+=" open-block";block.previousSibling.className+=" open-block";}</script> <div class="mw-content-ltr mw-parser-output" lang="ko" dir="ltr"> <section class="mf-section-0" id="mf-section-0"> <p><span class="nowrap"></span></p> <style data-mw-deduplicate="TemplateStyles:r34311307">.mw-parser-output .infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}body.skin-minerva .mw-parser-output .infobox-header,body.skin-minerva .mw-parser-output .infobox-subheader,body.skin-minerva .mw-parser-output .infobox-above,body.skin-minerva .mw-parser-output .infobox-title,body.skin-minerva .mw-parser-output .infobox-image,body.skin-minerva .mw-parser-output .infobox-full-data,body.skin-minerva .mw-parser-output .infobox-below{text-align:center}</style> <p><b>폴 로런스 모드리치</b>(<span style="font-size: smaller;"><a href="https://ko-m-wikipedia-org.translate.goog/wiki/%EC%98%81%EC%96%B4?_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB" title="영어">영어</a>: </span><span lang="en">Paul Lawrence Modrich</span>, 1946년 6월 13일 ~ )는 미국의 생화학자이다. 2015년 <a href="https://ko-m-wikipedia-org.translate.goog/wiki/%EB%85%B8%EB%B2%A8_%ED%99%94%ED%95%99%EC%83%81?_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB" title="노벨 화학상">노벨 화학상</a>을 수상하였다.</p> <table class="infobox biography vcard vevent"> <tbody> <tr> <th colspan="2" class="infobox-above fn summary" style="text-align:center; font-size:125%;">폴 로런스 모드리치<br> <div style="color:gray;font-size:80%"> Paul Lawrence Modrich </div></th> </tr> <tr> <th scope="row" class="infobox-label">로마자 표기</th> <td class="infobox-data">Paul Lawrence Modrich</td> </tr> <tr> <th scope="row" class="infobox-label">출생</th> <td class="infobox-data">1946년 6월 13일</td> </tr> <tr> <th scope="row" class="infobox-label">성별</th> <td class="infobox-data">남성</td> </tr> <tr> <th scope="row" class="infobox-label">국적</th> <td class="infobox-data category">미국</td> </tr> <tr> <th scope="row" class="infobox-label">직업</th> <td class="infobox-data role">생화학자</td> </tr> <tr> <th scope="row" class="infobox-label">상훈</th> <td class="infobox-data">노벨 화학상 (2015)</td> </tr> </tbody> </table> <div id="toc" class="toc" role="navigation" aria-labelledby="mw-toc-heading"> <input type="checkbox" role="button" id="toctogglecheckbox" class="toctogglecheckbox" style="display:none"> <div class="toctitle" lang="ko" dir="ltr"> <h2 id="mw-toc-heading">목차</h2><span class="toctogglespan"><label class="toctogglelabel" for="toctogglecheckbox"></label></span> </div> <ul> <li class="toclevel-1 tocsection-1"><a href="https://ko-m-wikipedia-org.translate.goog/wiki/%ED%8F%B4_L._%EB%AA%A8%EB%93%9C%EB%A6%AC%EC%B9%98?_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB#%EC%83%9D%EC%95%A0"><span class="tocnumber">1</span> <span class="toctext">생애</span></a></li> <li class="toclevel-1 tocsection-2"><a href="https://ko-m-wikipedia-org.translate.goog/wiki/%ED%8F%B4_L._%EB%AA%A8%EB%93%9C%EB%A6%AC%EC%B9%98?_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB#%EC%97%B0%EA%B5%AC%EC%97%85%EC%A0%81"><span class="tocnumber">2</span> <span class="toctext">연구업적</span></a> <ul> <li class="toclevel-2 tocsection-3"><a href="https://ko-m-wikipedia-org.translate.goog/wiki/%ED%8F%B4_L._%EB%AA%A8%EB%93%9C%EB%A6%AC%EC%B9%98?_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB#%EB%8C%80%EC%9E%A5%EA%B7%A0(E.coli)%EC%97%90%EC%84%9C%EC%9D%98_DNA_mismatch_repair"><span class="tocnumber">2.1</span> <span class="toctext">대장균(<i>E.coli</i>)에서의 DNA mismatch repair</span></a></li> <li class="toclevel-2 tocsection-4"><a href="https://ko-m-wikipedia-org.translate.goog/wiki/%ED%8F%B4_L._%EB%AA%A8%EB%93%9C%EB%A6%AC%EC%B9%98?_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB#%EC%A7%84%ED%95%B5%EC%84%B8%ED%8F%AC%EC%97%90%EC%84%9C%EC%9D%98_mismatch_repair"><span class="tocnumber">2.2</span> <span class="toctext">진핵세포에서의 mismatch repair</span></a></li> </ul></li> <li class="toclevel-1 tocsection-5"><a href="https://ko-m-wikipedia-org.translate.goog/wiki/%ED%8F%B4_L._%EB%AA%A8%EB%93%9C%EB%A6%AC%EC%B9%98?_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB#%EC%B0%B8%EA%B3%A0_%EC%9E%90%EB%A3%8C"><span class="tocnumber">3</span> <span class="toctext">참고 자료</span></a></li> <li class="toclevel-1 tocsection-6"><a href="https://ko-m-wikipedia-org.translate.goog/wiki/%ED%8F%B4_L._%EB%AA%A8%EB%93%9C%EB%A6%AC%EC%B9%98?_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB#%EC%99%B8%EB%B6%80_%EB%A7%81%ED%81%AC"><span class="tocnumber">4</span> <span class="toctext">외부 링크</span></a></li> </ul> </div> </section> <div class="mw-heading mw-heading2 section-heading" onclick="mfTempOpenSection(1)"> <span class="indicator mf-icon mf-icon-expand mf-icon--small"></span> <h2 id="생애"><span id=".EC.83.9D.EC.95.A0"></span>생애</h2><span class="mw-editsection"> <a role="button" href="https://ko-m-wikipedia-org.translate.goog/w/index.php?title=%ED%8F%B4_L._%EB%AA%A8%EB%93%9C%EB%A6%AC%EC%B9%98&amp;action=edit&amp;section=1&amp;_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB" title="부분 편집: 생애" class="cdx-button cdx-button--size-large cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--icon-only cdx-button--weight-quiet "> <span class="minerva-icon minerva-icon--edit"></span> <span>편집</span> </a> </span> </div> <section class="mf-section-1 collapsible-block" id="mf-section-1"> <p>미국 뉴 멕시코 주 북부의 작은 마을에서 자란 폴 모드리치는 어렸을 적 생물학 선생님인 아버지로부터 DNA에 대해 듣게 된다. 이후, 그 DNA라는 것은 그의 삶의 중심이 되게 된다. 스탠포드 대학에서의 박사과정, 하버드 대학에서의 박사 후 연구과정, 그리고 듀크대 에서의 교수과정 내내 그는 DNA에 영향을 주는 효소들을 연구했다. 그리고 1970년대 말 DNA repair에 연관된 효소를 연구하기 시작하게 된다.</p> </section> <div class="mw-heading mw-heading2 section-heading" onclick="mfTempOpenSection(2)"> <span class="indicator mf-icon mf-icon-expand mf-icon--small"></span> <h2 id="연구업적"><span id=".EC.97.B0.EA.B5.AC.EC.97.85.EC.A0.81"></span>연구업적</h2><span class="mw-editsection"> <a role="button" href="https://ko-m-wikipedia-org.translate.goog/w/index.php?title=%ED%8F%B4_L._%EB%AA%A8%EB%93%9C%EB%A6%AC%EC%B9%98&amp;action=edit&amp;section=2&amp;_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB" title="부분 편집: 연구업적" class="cdx-button cdx-button--size-large cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--icon-only cdx-button--weight-quiet "> <span class="minerva-icon minerva-icon--edit"></span> <span>편집</span> </a> </span> </div> <section class="mf-section-2 collapsible-block" id="mf-section-2"> <figure class="mw-default-size" typeof="mw:File/Thumb"> <a href="https://ko-m-wikipedia-org.translate.goog/wiki/%ED%8C%8C%EC%9D%BC:DNA_mismatch_repair_Ecoli.png?_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB" class="mw-file-description"> <noscript> <img src="//upload.wikimedia.org/wikipedia/commons/2/26/DNA_mismatch_repair_Ecoli.png" decoding="async" width="194" height="889" class="mw-file-element" data-file-width="194" data-file-height="889"> </noscript><span class="lazy-image-placeholder" style="width: 194px;height: 889px;" data-src="//upload.wikimedia.org/wikipedia/commons/2/26/DNA_mismatch_repair_Ecoli.png" data-width="194" data-height="889" data-class="mw-file-element">&nbsp;</span></a> <figcaption> <i>E.coli</i>의 DNA mismatch repair mechanism 모식도 </figcaption> </figure> <p>인간은 엄마의 난자로부터 23개, 아빠의 정자로부터 23개의 염색체(chromosome)을 받은 후 많고 많고 셀 수 없을 정도로 수많은 세포분열(cell division)을 하게 된다. 하나의 수정란에서 시작된 원본의 genome set은 어떻게 이 수많은 복제에도 불구하고 놀라울 정도로 온전하게 유지될 수 있는 걸까? 바로 DNA repair mechanism이다. 게놈(Genome)을 감시(monitoring)하고 수선(repair) 해준다는 이 단백질들을 밝혀낸 과학자들이 2015년 노벨 화학상을 받게 되었다. 그들 중 폴 모드리치는 mismatch repair를 밝혀내었다.</p> <div class="mw-heading mw-heading3"> <h3 id="대장균(E.coli)에서의_DNA_mismatch_repair"><span id=".EB.8C.80.EC.9E.A5.EA.B7.A0.28E.coli.29.EC.97.90.EC.84.9C.EC.9D.98_DNA_mismatch_repair"></span>대장균(<i>E.coli</i>)에서의 DNA mismatch repair</h3><span class="mw-editsection"> <a role="button" href="https://ko-m-wikipedia-org.translate.goog/w/index.php?title=%ED%8F%B4_L._%EB%AA%A8%EB%93%9C%EB%A6%AC%EC%B9%98&amp;action=edit&amp;section=3&amp;_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB" title="부분 편집: 대장균(E.coli)에서의 DNA mismatch repair" class="cdx-button cdx-button--size-large cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--icon-only cdx-button--weight-quiet "> <span class="minerva-icon minerva-icon--edit"></span> <span>편집</span> </a> </span> </div> <p>1983년부터 윤곽을 드러내기 시작한 그의 업적은 DNA 메틸화(DNA methylation)이 정말로 가닥 특이적(strand specific)한 mismatch repair를 제시한다는 발견으로 시작된다. 이어서 그는 <i>E.coli</i> 추출물에서 생체 밖(in vitro)상으로 DNA mismatch repair를 분석할 수 있는 정량법(assay)를 구축했다. 이 정량법(assay)를 기반으로, 그는 수선기작(repair mechanism)을 하나하나 밝히기 시작했다. Repair activity는 ATP, methylation state, MutH, MutL, MutS, UvrD에 영향을 받는다는 것을 밝혔으며, 이 수선 유전자(repair gene)들을 순수하게 분리해내 각각의 특징을 파헤치는 논문들을 연달아 내기 시작했다. 이렇게 쌓인 mismatch repair연구에 대한 경험들을 바탕으로 1989년에 그의 획기적인 논문이 발표된다. 이 연구로 생체 밖(in vitro)상에서 DNA mismatch repair를 완벽하게 재구성해 냈다. DNA mismatch repair를 완벽하게 재구성을 하기 위해서는 DNA polymerase III, exonuclease I, DNA ligase, 그리고 정제된 MutH, MutL, MutS, UvrD, SSB protein이 필요하다고 그는 설명했다. 또한 이 요소들은 함께 mismatch repair를 해내는데, mismatch가 일어난 장소에서부터 어느 정도 거리가 있는, GATC 서열 위에 한 가닥만 메틸화(methylation)된 hemimethylated 가닥이 방향을 지시한다는 것을 이 논문을 통해 설명했다. Mismatch repair system의 중심역할을 하는 MutS가 왓슨-크릭(Watson-Crick) 염기쌍에 위배되는 mismatch를 인식하고, 이후 MutL과 복합체(complex)를 이룬다. MutL은 중개자(mediator)역할을 하는데, MutH, MutS와 모두 상호작용한다. MutS, MutL 복합체가 MutH를 부르고(recruit), 이 MutH는 hemimethylated GATC site에 붙는다. 이 과정을 통해 가닥특이성(strand-specificity)가 생긴다. 메틸(methyl)기가 붙어있는 가닥은 parental strand이고 메틸(methyl)기가 없는 것은 새로 합성된 daughter strand가 되기 때문에 양 가닥을 구분 가능하다. 원래는 약햤던 MutH의 endonuclease activity를 활성화되고 그 결과 MutH는 methyl기가 없는 가닥(strand)의 GATC site 부근에 틈(nick)을 만들게 된다. 다음 helicase인 UvrD의 도움을 받아 이중가닥을 풀어가면서 Endonuclease I이 메틸(methyl)기가 없는 가닥을 분해해나간다. 틈(nick)부터 mismatch site 바로 넘어서까지 분해된다. 이렇게 생긴 틈새(gap)는 DNA pol III와 DNA ligase로 채워진다. 이렇게 mismatch가 원본의 주형으로 교정된다.</p> <div class="mw-heading mw-heading3"> <h3 id="진핵세포에서의_mismatch_repair"><span id=".EC.A7.84.ED.95.B5.EC.84.B8.ED.8F.AC.EC.97.90.EC.84.9C.EC.9D.98_mismatch_repair"></span>진핵세포에서의 mismatch repair</h3><span class="mw-editsection"> <a role="button" href="https://ko-m-wikipedia-org.translate.goog/w/index.php?title=%ED%8F%B4_L._%EB%AA%A8%EB%93%9C%EB%A6%AC%EC%B9%98&amp;action=edit&amp;section=4&amp;_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB" title="부분 편집: 진핵세포에서의 mismatch repair" class="cdx-button cdx-button--size-large cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--icon-only cdx-button--weight-quiet "> <span class="minerva-icon minerva-icon--edit"></span> <span>편집</span> </a> </span> </div> <p>모드리치는 대장균(<i>E.coli</i>)의 수선기작(repair mechanism)을 밝혀낸 후 진핵생물에서의 mismatch repair를 연구하기 위해 인간과 초파리의 세포에서의 mismatch repair system을 계속해서 연구해나갔다. 진핵세포(eukaryotic)에서의 system은 MutS, MutL 등의 효소와 상동적으로 대장균(<i>E.coli</i>)에서의 mismatch repair system 이 잘 보존되어있지만, 대장균(<i>E.coli</i>)와 한 가지 차이점이 있다. 메틸화(methylation)로 가닥특이적(strand specific)하게 수선(repair)되던 대장균(<i>E.coli</i>)과 달리 인간(human), 초파리(drosophila)의 경우 메틸화(methylation)가 가닥 특이적 수선(strand specific repair)에 관여하지 않는다는 것이다. 그대신 strand break로 가닥특이적(strand specific)으로 수선(repair)이 일어났다는 것이 실험을 통해 밝혀졌다.</p> </section> <div class="mw-heading mw-heading2 section-heading" onclick="mfTempOpenSection(3)"> <span class="indicator mf-icon mf-icon-expand mf-icon--small"></span> <h2 id="참고_자료"><span id=".EC.B0.B8.EA.B3.A0_.EC.9E.90.EB.A3.8C"></span>참고 자료</h2><span class="mw-editsection"> <a role="button" href="https://ko-m-wikipedia-org.translate.goog/w/index.php?title=%ED%8F%B4_L._%EB%AA%A8%EB%93%9C%EB%A6%AC%EC%B9%98&amp;action=edit&amp;section=5&amp;_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB" title="부분 편집: 참고 자료" class="cdx-button cdx-button--size-large cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--icon-only cdx-button--weight-quiet "> <span class="minerva-icon minerva-icon--edit"></span> <span>편집</span> </a> </span> </div> <section class="mf-section-3 collapsible-block" id="mf-section-3"> <ul> <li><a rel="nofollow" class="external free" href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://www.biochem.duke.edu/paul-l-modrich-primary">https://www.biochem.duke.edu/paul-l-modrich-primary</a> <a rel="nofollow" class="external text" href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://web.archive.org/web/20151127135513/https://www.biochem.duke.edu/paul-l-modrich-primary">Archived</a> 2015년 11월 27일 - <a href="https://ko-m-wikipedia-org.translate.goog/wiki/%EC%9B%A8%EC%9D%B4%EB%B0%B1_%EB%A8%B8%EC%8B%A0?_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB" title="웨이백 머신">웨이백 머신</a></li> <li><a rel="nofollow" class="external free" href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=http://www.nobelprize.org/nobel_prizes/chemistry/laureates/2015/">http://www.nobelprize.org/nobel_prizes/chemistry/laureates/2015/</a></li> <li><a rel="nofollow" class="external free" href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=http://www.hhmi.org/research/dna-mismatch-repair-and-genetic-stability">http://www.hhmi.org/research/dna-mismatch-repair-and-genetic-stability</a> <a rel="nofollow" class="external text" href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://web.archive.org/web/20151114085659/http://www.hhmi.org/research/dna-mismatch-repair-and-genetic-stability">Archived</a> 2015년 11월 14일 - <a href="https://ko-m-wikipedia-org.translate.goog/wiki/%EC%9B%A8%EC%9D%B4%EB%B0%B1_%EB%A8%B8%EC%8B%A0?_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB" title="웨이백 머신">웨이백 머신</a></li> <li>Su, S.S. and P. Modrich, Escherichia coli mutS-encoded protein binds to mismatched DNA base pairs. Proc Natl Acad Sci U S A, 1983. 83〔14〕:p.4639-43.</li> <li>Holmes, J., Jr., S. Clark, and P. Modrich, Strand-specific mismatch correction in nuclear extracts of human and Drosophila melanogaster cell lines. Proc Natl Acad Sci U S A, 1990. 87〔15〕:p.5837-41.</li> <li>Scientific Background on the Nobel Prize in Chemistry 2015</li> <li>Pukkila, P.J., et al., Effects of high levels of DNA adenine methylation on methyl-directed mismatch repair in Escherichia coli. Genetics, 1983. 104(4): p. 571-82.</li> <li>Lu, A.L., S. Clark, and P. Modrich, Methyl-directed repair of DNA base-pair mismatches in vitro. Proc Natl Acad Sci U S A, 1983. 80(15): p. 4639-43.</li> <li>Su, S.S. and P. Modrich, Escherichia coli mutS-encoded protein binds to mismatched DNA base pairs. Proc Natl Acad Sci U S A, 1986. 83(14): p. 5057-61.</li> <li>Grilley, M., et al., Isolation and characterization of the Escherichia coli mutL gene product. J Biol Chem, 1989. 264(2): p. 1000-4.</li> <li>Welsh, K.M., et al., Isolation and characterization of the Escherichia coli mutH gene product. J Biol Chem, 1987. 262(32): p. 15624-9.</li> <li>Lahue, R.S., K.G. Au, and P. Modrich, DNA mismatch correction in a defined system. Science, 1989. 245(4914): p. 160-4.</li> </ul> </section> <div class="mw-heading mw-heading2 section-heading" onclick="mfTempOpenSection(4)"> <span class="indicator mf-icon mf-icon-expand mf-icon--small"></span> <h2 id="외부_링크"><span id=".EC.99.B8.EB.B6.80_.EB.A7.81.ED.81.AC"></span>외부 링크</h2><span class="mw-editsection"> <a role="button" href="https://ko-m-wikipedia-org.translate.goog/w/index.php?title=%ED%8F%B4_L._%EB%AA%A8%EB%93%9C%EB%A6%AC%EC%B9%98&amp;action=edit&amp;section=6&amp;_x_tr_sl=auto&amp;_x_tr_tl=en&amp;_x_tr_hl=en-GB" title="부분 편집: 외부 링크" class="cdx-button cdx-button--size-large cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--icon-only cdx-button--weight-quiet "> <span class="minerva-icon minerva-icon--edit"></span> <span>편집</span> </a> </span> </div> <section class="mf-section-4 collapsible-block" id="mf-section-4"> <ul> <li><span typeof="mw:File"><span> <noscript> <img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/12px-Commons-logo.svg.png" decoding="async" width="12" height="16" class="mw-file-element" data-file-width="1024" data-file-height="1376"> </noscript><span class="lazy-image-placeholder" style="width: 12px;height: 16px;" data-src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/12px-Commons-logo.svg.png" data-alt="" data-width="12" data-height="16" data-srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/18px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/24px-Commons-logo.svg.png 2x" data-class="mw-file-element">&nbsp;</span></span></span> 위키미디어 공용에 <span class="plainlinks"><a class="external text" href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://commons.wikimedia.org/wiki/Category:Paul_L._Modrich?uselang%3Dko">폴 L. 모드리치</a></span> 관련 미디어 분류가 있습니다.</li> </ul> <div class="navbox-styles"> <style data-mw-deduplicate="TemplateStyles:r36480591">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol ol,.mw-parser-output .hlist ol ul,.mw-parser-output .hlist ul dl,.mw-parser-output .hlist ul ol,.mw-parser-output .hlist ul ul{display:inline}.mw-parser-output .hlist .mw-empty-li{display:none}.mw-parser-output .hlist dt::after{content:": "}.mw-parser-output .hlist dd::after,.mw-parser-output .hlist li::after{content:" · ";font-weight:bold}.mw-parser-output .hlist dd:last-child::after,.mw-parser-output .hlist dt:last-child::after,.mw-parser-output .hlist li:last-child::after{content:none}.mw-parser-output .hlist dd dd:first-child::before,.mw-parser-output .hlist dd dt:first-child::before,.mw-parser-output .hlist dd li:first-child::before,.mw-parser-output .hlist dt dd:first-child::before,.mw-parser-output .hlist dt dt:first-child::before,.mw-parser-output .hlist dt li:first-child::before,.mw-parser-output .hlist li dd:first-child::before,.mw-parser-output .hlist li dt:first-child::before,.mw-parser-output .hlist li li:first-child::before{content:" (";font-weight:normal}.mw-parser-output .hlist dd dd:last-child::after,.mw-parser-output .hlist dd dt:last-child::after,.mw-parser-output .hlist dd li:last-child::after,.mw-parser-output .hlist dt dd:last-child::after,.mw-parser-output .hlist dt dt:last-child::after,.mw-parser-output .hlist dt li:last-child::after,.mw-parser-output .hlist li dd:last-child::after,.mw-parser-output .hlist li dt:last-child::after,.mw-parser-output .hlist li li:last-child::after{content:")";font-weight:normal}.mw-parser-output .hlist ol{counter-reset:listitem}.mw-parser-output .hlist ol>li{counter-increment:listitem}.mw-parser-output .hlist ol>li::before{content:" "counter(listitem)"\a0 "}.mw-parser-output .hlist dd ol>li:first-child::before,.mw-parser-output .hlist dt ol>li:first-child::before,.mw-parser-output .hlist li ol>li:first-child::before{content:" ("counter(listitem)"\a0 "}.mw-parser-output .hlist-pipe dd:after,.mw-parser-output .hlist-pipe li:after{content:" | ";font-weight:normal}.mw-parser-output .hlist-hyphen dd:after,.mw-parser-output .hlist-hyphen li:after{content:" - 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Modrich – 카탈로니아어" lang="ca" hreflang="ca" data-title="Paul L. Modrich" data-language-autonym="Català" data-language-local-name="카탈로니아어" class="interlanguage-link-target"><span>Català</span></a></li> <li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://cs.wikipedia.org/wiki/Paul_Modrich" title="Paul Modrich – 체코어" lang="cs" hreflang="cs" data-title="Paul Modrich" data-language-autonym="Čeština" data-language-local-name="체코어" class="interlanguage-link-target"><span>Čeština</span></a></li> <li class="interlanguage-link interwiki-da mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://da.wikipedia.org/wiki/Paul_L._Modrich" title="Paul L. Modrich – 덴마크어" lang="da" hreflang="da" data-title="Paul L. Modrich" data-language-autonym="Dansk" data-language-local-name="덴마크어" class="interlanguage-link-target"><span>Dansk</span></a></li> <li class="interlanguage-link interwiki-de mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://de.wikipedia.org/wiki/Paul_Modrich" title="Paul Modrich – 독일어" lang="de" hreflang="de" data-title="Paul Modrich" data-language-autonym="Deutsch" data-language-local-name="독일어" class="interlanguage-link-target"><span>Deutsch</span></a></li> <li class="interlanguage-link interwiki-en mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://en.wikipedia.org/wiki/Paul_L._Modrich" title="Paul L. Modrich – 영어" lang="en" hreflang="en" data-title="Paul L. Modrich" data-language-autonym="English" data-language-local-name="영어" class="interlanguage-link-target"><span>English</span></a></li> <li class="interlanguage-link interwiki-es mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://es.wikipedia.org/wiki/Paul_L._Modrich" title="Paul L. Modrich – 스페인어" lang="es" hreflang="es" data-title="Paul L. Modrich" data-language-autonym="Español" data-language-local-name="스페인어" class="interlanguage-link-target"><span>Español</span></a></li> <li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://eu.wikipedia.org/wiki/Paul_Modrich" title="Paul Modrich – 바스크어" lang="eu" hreflang="eu" data-title="Paul Modrich" data-language-autonym="Euskara" data-language-local-name="바스크어" class="interlanguage-link-target"><span>Euskara</span></a></li> <li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://fa.wikipedia.org/wiki/%25D9%25BE%25D8%25A7%25D9%2584_%25D9%2585%25D9%2588%25D8%25AF%25D8%25B1%25DB%258C%25DA%2586" title="پال مودریچ – 페르시아어" lang="fa" hreflang="fa" data-title="پال مودریچ" data-language-autonym="فارسی" data-language-local-name="페르시아어" class="interlanguage-link-target"><span>فارسی</span></a></li> <li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://fi.wikipedia.org/wiki/Paul_Modrich" title="Paul Modrich – 핀란드어" lang="fi" hreflang="fi" data-title="Paul Modrich" data-language-autonym="Suomi" data-language-local-name="핀란드어" class="interlanguage-link-target"><span>Suomi</span></a></li> <li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://fr.wikipedia.org/wiki/Paul_L._Modrich" title="Paul L. Modrich – 프랑스어" lang="fr" hreflang="fr" data-title="Paul L. Modrich" data-language-autonym="Français" data-language-local-name="프랑스어" class="interlanguage-link-target"><span>Français</span></a></li> <li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://ga.wikipedia.org/wiki/Paul_L._Modrich" title="Paul L. Modrich – 아일랜드어" lang="ga" hreflang="ga" data-title="Paul L. Modrich" data-language-autonym="Gaeilge" data-language-local-name="아일랜드어" class="interlanguage-link-target"><span>Gaeilge</span></a></li> <li class="interlanguage-link interwiki-gd mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://gd.wikipedia.org/wiki/Paul_Modrich" title="Paul Modrich – 스코틀랜드 게일어" lang="gd" hreflang="gd" data-title="Paul Modrich" data-language-autonym="Gàidhlig" data-language-local-name="스코틀랜드 게일어" class="interlanguage-link-target"><span>Gàidhlig</span></a></li> <li class="interlanguage-link interwiki-hak mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://hak.wikipedia.org/wiki/Paul_L._Modrich" title="Paul L. Modrich – 하카어" lang="hak" hreflang="hak" data-title="Paul L. Modrich" data-language-autonym="客家語 / Hak-kâ-ngî" data-language-local-name="하카어" class="interlanguage-link-target"><span>客家語 / Hak-kâ-ngî</span></a></li> <li class="interlanguage-link interwiki-he mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://he.wikipedia.org/wiki/%25D7%25A4%25D7%2595%25D7%259C_%25D7%259E%25D7%2595%25D7%2593%25D7%25A8%25D7%2599%25D7%25A5%2527" title="פול מודריץ' – 히브리어" lang="he" hreflang="he" data-title="פול מודריץ'" data-language-autonym="עברית" data-language-local-name="히브리어" class="interlanguage-link-target"><span>עברית</span></a></li> <li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://hr.wikipedia.org/wiki/Paul_Modrich" title="Paul Modrich – 크로아티아어" lang="hr" hreflang="hr" data-title="Paul Modrich" data-language-autonym="Hrvatski" data-language-local-name="크로아티아어" class="interlanguage-link-target"><span>Hrvatski</span></a></li> <li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://hu.wikipedia.org/wiki/Paul_Modrich" title="Paul Modrich – 헝가리어" lang="hu" hreflang="hu" data-title="Paul Modrich" data-language-autonym="Magyar" data-language-local-name="헝가리어" class="interlanguage-link-target"><span>Magyar</span></a></li> <li class="interlanguage-link interwiki-hy mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://hy.wikipedia.org/wiki/%25D5%258A%25D5%25B8%25D5%25AC_%25D5%2584%25D5%25B8%25D5%25A4%25D6%2580%25D5%25AB%25D5%25B9" title="Պոլ Մոդրիչ – 아르메니아어" lang="hy" hreflang="hy" data-title="Պոլ Մոդրիչ" data-language-autonym="Հայերեն" data-language-local-name="아르메니아어" class="interlanguage-link-target"><span>Հայերեն</span></a></li> <li class="interlanguage-link interwiki-id mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://id.wikipedia.org/wiki/Paul_L._Modrich" title="Paul L. Modrich – 인도네시아어" lang="id" hreflang="id" data-title="Paul L. Modrich" data-language-autonym="Bahasa Indonesia" data-language-local-name="인도네시아어" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li> <li class="interlanguage-link interwiki-io mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://io.wikipedia.org/wiki/Paul_L._Modrich" title="Paul L. Modrich – 이도어" lang="io" hreflang="io" data-title="Paul L. Modrich" data-language-autonym="Ido" data-language-local-name="이도어" class="interlanguage-link-target"><span>Ido</span></a></li> <li class="interlanguage-link interwiki-it mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://it.wikipedia.org/wiki/Paul_Modrich" title="Paul Modrich – 이탈리아어" lang="it" hreflang="it" data-title="Paul Modrich" data-language-autonym="Italiano" data-language-local-name="이탈리아어" class="interlanguage-link-target"><span>Italiano</span></a></li> <li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://ja.wikipedia.org/wiki/%25E3%2583%259D%25E3%2583%25BC%25E3%2583%25AB%25E3%2583%25BB%25E3%2583%25A2%25E3%2583%2589%25E3%2583%25AA%25E3%2583%2583%25E3%2583%2581" title="ポール・モドリッチ – 일본어" lang="ja" hreflang="ja" data-title="ポール・モドリッチ" data-language-autonym="日本語" data-language-local-name="일본어" class="interlanguage-link-target"><span>日本語</span></a></li> <li class="interlanguage-link interwiki-ka mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://ka.wikipedia.org/wiki/%25E1%2583%259E%25E1%2583%259D%25E1%2583%259A_%25E1%2583%259B%25E1%2583%259D%25E1%2583%2593%25E1%2583%25A0%25E1%2583%2598%25E1%2583%25A9%25E1%2583%2598" title="პოლ მოდრიჩი – 조지아어" lang="ka" hreflang="ka" data-title="პოლ მოდრიჩი" data-language-autonym="ქართული" data-language-local-name="조지아어" class="interlanguage-link-target"><span>ქართული</span></a></li> <li class="interlanguage-link interwiki-la mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://la.wikipedia.org/wiki/Paulus_Modrich" title="Paulus Modrich – 라틴어" lang="la" hreflang="la" data-title="Paulus Modrich" data-language-autonym="Latina" data-language-local-name="라틴어" class="interlanguage-link-target"><span>Latina</span></a></li> <li class="interlanguage-link interwiki-mk mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://mk.wikipedia.org/wiki/%25D0%259F%25D0%25BE%25D0%25BB_%25D0%259C%25D0%25BE%25D0%25B4%25D1%2580%25D0%25B8%25D1%2587" title="Пол Модрич – 마케도니아어" lang="mk" hreflang="mk" data-title="Пол Модрич" data-language-autonym="Македонски" data-language-local-name="마케도니아어" class="interlanguage-link-target"><span>Македонски</span></a></li> <li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://ms.wikipedia.org/wiki/Paul_L._Modrich" title="Paul L. Modrich – 말레이어" lang="ms" hreflang="ms" data-title="Paul L. Modrich" data-language-autonym="Bahasa Melayu" data-language-local-name="말레이어" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li> <li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://mzn.wikipedia.org/wiki/%25D9%25BE%25D8%25A7%25D9%2584_%25D9%2585%25D9%2588%25D8%25AF%25D8%25B1%25DB%258C%25DA%2586" title="پال مودریچ – 마잔데라니어" lang="mzn" hreflang="mzn" data-title="پال مودریچ" data-language-autonym="مازِرونی" data-language-local-name="마잔데라니어" class="interlanguage-link-target"><span>مازِرونی</span></a></li> <li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://nds.wikipedia.org/wiki/Paul_Lawrence_Modrich" title="Paul Lawrence Modrich – 저지 독일어" lang="nds" hreflang="nds" data-title="Paul Lawrence Modrich" data-language-autonym="Plattdüütsch" data-language-local-name="저지 독일어" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li> <li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://nl.wikipedia.org/wiki/Paul_Modrich" title="Paul Modrich – 네덜란드어" lang="nl" hreflang="nl" data-title="Paul Modrich" data-language-autonym="Nederlands" data-language-local-name="네덜란드어" class="interlanguage-link-target"><span>Nederlands</span></a></li> <li class="interlanguage-link interwiki-no mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://no.wikipedia.org/wiki/Paul_L._Modrich" title="Paul L. Modrich – 노르웨이어(보크말)" lang="nb" hreflang="nb" data-title="Paul L. Modrich" data-language-autonym="Norsk bokmål" data-language-local-name="노르웨이어(보크말)" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li> <li class="interlanguage-link interwiki-pa mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://pa.wikipedia.org/wiki/%25E0%25A8%25AA%25E0%25A9%258C%25E0%25A8%25B2_%25E0%25A8%2590%25E0%25A8%25B2._%25E0%25A8%25AE%25E0%25A9%258B%25E0%25A8%25A1%25E0%25A8%25B0%25E0%25A8%25BF%25E0%25A8%2595" title="ਪੌਲ ਐਲ. ਮੋਡਰਿਕ – 펀잡어" lang="pa" hreflang="pa" data-title="ਪੌਲ ਐਲ. ਮੋਡਰਿਕ" data-language-autonym="ਪੰਜਾਬੀ" data-language-local-name="펀잡어" class="interlanguage-link-target"><span>ਪੰਜਾਬੀ</span></a></li> <li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://pl.wikipedia.org/wiki/Paul_Modrich" title="Paul Modrich – 폴란드어" lang="pl" hreflang="pl" data-title="Paul Modrich" data-language-autonym="Polski" data-language-local-name="폴란드어" class="interlanguage-link-target"><span>Polski</span></a></li> <li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://pnb.wikipedia.org/wiki/%25D9%25BE%25D8%25A7%25D9%2584_%25D9%2584%25D8%25A7%25D8%25B1%25DB%258C%25D9%2586%25D8%25B3_%25D9%2585%25D9%2588%25DA%2588%25D8%25B1%25DB%258C%25DA%2586" title="پال لارینس موڈریچ – Western Punjabi" lang="pnb" hreflang="pnb" data-title="پال لارینس موڈریچ" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li> <li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://pt.wikipedia.org/wiki/Paul_Modrich" title="Paul Modrich – 포르투갈어" lang="pt" hreflang="pt" data-title="Paul Modrich" data-language-autonym="Português" data-language-local-name="포르투갈어" class="interlanguage-link-target"><span>Português</span></a></li> <li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://ro.wikipedia.org/wiki/Paul_Modrich" title="Paul Modrich – 루마니아어" lang="ro" hreflang="ro" data-title="Paul Modrich" data-language-autonym="Română" data-language-local-name="루마니아어" class="interlanguage-link-target"><span>Română</span></a></li> <li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://ru.wikipedia.org/wiki/%25D0%259C%25D0%25BE%25D0%25B4%25D1%2580%25D0%25B8%25D1%2587,_%25D0%259F%25D0%25BE%25D0%25BB" title="Модрич, Пол – 러시아어" lang="ru" hreflang="ru" data-title="Модрич, Пол" data-language-autonym="Русский" data-language-local-name="러시아어" class="interlanguage-link-target"><span>Русский</span></a></li> <li class="interlanguage-link interwiki-scn mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://scn.wikipedia.org/wiki/Paul_L._Modrich" title="Paul L. Modrich – 시칠리아어" lang="scn" hreflang="scn" data-title="Paul L. Modrich" data-language-autonym="Sicilianu" data-language-local-name="시칠리아어" class="interlanguage-link-target"><span>Sicilianu</span></a></li> <li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://sh.wikipedia.org/wiki/Paul_L._Modrich" title="Paul L. Modrich – 세르비아-크로아티아어" lang="sh" hreflang="sh" data-title="Paul L. Modrich" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="세르비아-크로아티아어" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li> <li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://simple.wikipedia.org/wiki/Paul_Modrich" title="Paul Modrich – Simple English" lang="en-simple" hreflang="en-simple" data-title="Paul Modrich" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li> <li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://sk.wikipedia.org/wiki/Paul_Modrich" title="Paul Modrich – 슬로바키아어" lang="sk" hreflang="sk" data-title="Paul Modrich" data-language-autonym="Slovenčina" data-language-local-name="슬로바키아어" class="interlanguage-link-target"><span>Slovenčina</span></a></li> <li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://sv.wikipedia.org/wiki/Paul_Modrich" title="Paul Modrich – 스웨덴어" lang="sv" hreflang="sv" data-title="Paul Modrich" data-language-autonym="Svenska" data-language-local-name="스웨덴어" class="interlanguage-link-target"><span>Svenska</span></a></li> <li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://sw.wikipedia.org/wiki/Paul_Modrich" title="Paul Modrich – 스와힐리어" lang="sw" hreflang="sw" data-title="Paul Modrich" data-language-autonym="Kiswahili" data-language-local-name="스와힐리어" class="interlanguage-link-target"><span>Kiswahili</span></a></li> <li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://ta.wikipedia.org/wiki/%25E0%25AE%25AA%25E0%25AE%25B5%25E0%25AF%2581%25E0%25AE%25B2%25E0%25AF%258D_%25E0%25AE%25AE%25E0%25AF%258B%25E0%25AE%259F%25E0%25AF%258D%25E0%25AE%25B0%25E0%25AE%25BF%25E0%25AE%259A%25E0%25AF%258D" title="பவுல் மோட்ரிச் – 타밀어" lang="ta" hreflang="ta" data-title="பவுல் மோட்ரிச்" data-language-autonym="தமிழ்" data-language-local-name="타밀어" class="interlanguage-link-target"><span>தமிழ்</span></a></li> <li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://tr.wikipedia.org/wiki/Paul_Modrich" title="Paul Modrich – 터키어" lang="tr" hreflang="tr" data-title="Paul Modrich" data-language-autonym="Türkçe" data-language-local-name="터키어" class="interlanguage-link-target"><span>Türkçe</span></a></li> <li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://tt.wikipedia.org/wiki/%25D0%259F%25D0%25BE%25D0%25BB_%25D0%259C%25D0%25BE%25D0%25B4%25D1%2580%25D0%25B8%25D1%2587" title="Пол Модрич – 타타르어" lang="tt" hreflang="tt" data-title="Пол Модрич" data-language-autonym="Татарча / tatarça" data-language-local-name="타타르어" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li> <li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://uk.wikipedia.org/wiki/%25D0%259F%25D0%25BE%25D0%25BB_%25D0%259C%25D0%25BE%25D0%25B4%25D1%2580%25D0%25B8%25D1%2587" title="Пол Модрич – 우크라이나어" lang="uk" hreflang="uk" data-title="Пол Модрич" data-language-autonym="Українська" data-language-local-name="우크라이나어" class="interlanguage-link-target"><span>Українська</span></a></li> <li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://ur.wikipedia.org/wiki/%25D9%25BE%25D8%25A7%25D9%2584_%25D8%25A7%25DB%258C%25D9%2584_%25D9%2585%25D9%2588%25DA%2588%25D8%25B1%25DA%2586" title="پال ایل موڈرچ – 우르두어" lang="ur" hreflang="ur" data-title="پال ایل موڈرچ" data-language-autonym="اردو" data-language-local-name="우르두어" class="interlanguage-link-target"><span>اردو</span></a></li> <li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://vi.wikipedia.org/wiki/Paul_L._Modrich" title="Paul L. Modrich – 베트남어" lang="vi" hreflang="vi" data-title="Paul L. Modrich" data-language-autonym="Tiếng Việt" data-language-local-name="베트남어" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li> <li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://translate.google.com/website?sl=auto&amp;tl=en&amp;hl=en-GB&amp;u=https://zh.wikipedia.org/wiki/%25E4%25BF%259D%25E7%25BD%2597%25C2%25B7%25E8%258E%25AB%25E5%25BE%25B7%25E9%2587%258C%25E5%25A5%2587" title="保罗·莫德里奇 – 중국어" lang="zh" hreflang="zh" data-title="保罗·莫德里奇" data-language-autonym="中文" data-language-local-name="중국어" class="interlanguage-link-target"><span>中文</span></a></li> </ul> </section> </div> <div class="minerva-footer-logo"> <img src="/static/images/mobile/copyright/wikipedia-wordmark-ko.svg" alt="위키백과" width="120" height="28" style="width: 7.5em; height: 1.75em;"> </div> <ul id="footer-info" class="footer-info hlist hlist-separated"> <li id="footer-info-lastmod">이 문서는 2024년 11월 22일 (금) 14:45에 마지막으로 편집되었습니다.</li> <li id="footer-info-copyright">별도로 명시하지 않은 경우, 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