CINXE.COM

Paul Nurse - Simple English Wikipedia, the free encyclopedia

<!DOCTYPE html> <html class="client-nojs vector-feature-language-in-header-enabled vector-feature-language-in-main-page-header-disabled vector-feature-sticky-header-disabled vector-feature-page-tools-pinned-disabled vector-feature-toc-pinned-clientpref-1 vector-feature-main-menu-pinned-disabled vector-feature-limited-width-clientpref-1 vector-feature-limited-width-content-enabled vector-feature-custom-font-size-clientpref-1 vector-feature-appearance-pinned-clientpref-1 vector-feature-night-mode-disabled skin-theme-clientpref-day vector-toc-available" lang="en" dir="ltr"> <head> <meta charset="UTF-8"> <title>Paul Nurse - Simple English Wikipedia, the free encyclopedia</title> <script>(function(){var className="client-js vector-feature-language-in-header-enabled vector-feature-language-in-main-page-header-disabled vector-feature-sticky-header-disabled vector-feature-page-tools-pinned-disabled vector-feature-toc-pinned-clientpref-1 vector-feature-main-menu-pinned-disabled vector-feature-limited-width-clientpref-1 vector-feature-limited-width-content-enabled vector-feature-custom-font-size-clientpref-1 vector-feature-appearance-pinned-clientpref-1 vector-feature-night-mode-disabled skin-theme-clientpref-day vector-toc-available";var cookie=document.cookie.match(/(?:^|; )simplewikimwclientpreferences=([^;]+)/);if(cookie){cookie[1].split('%2C').forEach(function(pref){className=className.replace(new RegExp('(^| )'+pref.replace(/-clientpref-\w+$|[^\w-]+/g,'')+'-clientpref-\\w+( |$)'),'$1'+pref+'$2');});}document.documentElement.className=className;}());RLCONF={"wgBreakFrames":false,"wgSeparatorTransformTable":["",""],"wgDigitTransformTable":["",""],"wgDefaultDateFormat": "dmy","wgMonthNames":["","January","February","March","April","May","June","July","August","September","October","November","December"],"wgRequestId":"fe6b5749-c041-4ace-ac75-e69512daf091","wgCanonicalNamespace":"","wgCanonicalSpecialPageName":false,"wgNamespaceNumber":0,"wgPageName":"Paul_Nurse","wgTitle":"Paul Nurse","wgCurRevisionId":9574995,"wgRevisionId":9574995,"wgArticleId":291777,"wgIsArticle":true,"wgIsRedirect":false,"wgAction":"view","wgUserName":null,"wgUserGroups":["*"],"wgCategories":["Pages using the JsonConfig extension","Articles with hCards","Nobel Prize in Physiology or Medicine winners","1949 births","Living people","English biologists","English geneticists","Knights Bachelor","People from Norwich","Presidents of the Royal Society","British Nobel Prize winners","Alumni of the University of Birmingham","Alumni of the University of East Anglia","Rockefeller University faculty"],"wgPageViewLanguage":"en","wgPageContentLanguage":"en","wgPageContentModel":"wikitext", "wgRelevantPageName":"Paul_Nurse","wgRelevantArticleId":291777,"wgIsProbablyEditable":true,"wgRelevantPageIsProbablyEditable":true,"wgRestrictionEdit":[],"wgRestrictionMove":[],"wgNoticeProject":"wikipedia","wgCiteReferencePreviewsActive":false,"wgMediaViewerOnClick":true,"wgMediaViewerEnabledByDefault":true,"wgPopupsFlags":0,"wgVisualEditor":{"pageLanguageCode":"en","pageLanguageDir":"ltr","pageVariantFallbacks":"en"},"wgMFDisplayWikibaseDescriptions":{"search":true,"watchlist":true,"tagline":true,"nearby":true},"wgWMESchemaEditAttemptStepOversample":false,"wgWMEPageLength":4000,"wgRelatedArticlesCompat":[],"wgCentralAuthMobileDomain":false,"wgEditSubmitButtonLabelPublish":true,"wgULSPosition":"interlanguage","wgULSisCompactLinksEnabled":false,"wgVector2022LanguageInHeader":true,"wgULSisLanguageSelectorEmpty":false,"wgWikibaseItemId":"Q219716","wgCheckUserClientHintsHeadersJsApi":["brands","architecture","bitness","fullVersionList","mobile","model","platform","platformVersion"], "GEHomepageSuggestedEditsEnableTopics":true,"wgGETopicsMatchModeEnabled":false,"wgGEStructuredTaskRejectionReasonTextInputEnabled":false,"wgGELevelingUpEnabledForUser":false};RLSTATE={"ext.globalCssJs.user.styles":"ready","site.styles":"ready","user.styles":"ready","ext.globalCssJs.user":"ready","user":"ready","user.options":"loading","ext.cite.styles":"ready","skins.vector.search.codex.styles":"ready","skins.vector.styles":"ready","skins.vector.icons":"ready","jquery.makeCollapsible.styles":"ready","ext.wikimediamessages.styles":"ready","ext.visualEditor.desktopArticleTarget.noscript":"ready","ext.uls.interlanguage":"ready","wikibase.client.init":"ready","ext.wikimediaBadges":"ready"};RLPAGEMODULES=["ext.cite.ux-enhancements","site","mediawiki.page.ready","jquery.makeCollapsible","skins.vector.js","ext.centralNotice.geoIP","ext.centralNotice.startUp","ext.gadget.ReferenceTooltips","ext.gadget.refToolbar","ext.gadget.switcher","ext.urlShortener.toolbar", "ext.centralauth.centralautologin","mmv.bootstrap","ext.popups","ext.visualEditor.desktopArticleTarget.init","ext.visualEditor.targetLoader","ext.echo.centralauth","ext.eventLogging","ext.wikimediaEvents","ext.navigationTiming","ext.uls.interface","ext.cx.eventlogging.campaigns","ext.cx.uls.quick.actions","wikibase.client.vector-2022","ext.checkUser.clientHints","ext.growthExperiments.SuggestedEditSession","wikibase.sidebar.tracking"];</script> <script>(RLQ=window.RLQ||[]).push(function(){mw.loader.impl(function(){return["user.options@12s5i",function($,jQuery,require,module){mw.user.tokens.set({"patrolToken":"+\\","watchToken":"+\\","csrfToken":"+\\"}); }];});});</script> <link rel="stylesheet" href="/w/load.php?lang=en&amp;modules=ext.cite.styles%7Cext.uls.interlanguage%7Cext.visualEditor.desktopArticleTarget.noscript%7Cext.wikimediaBadges%7Cext.wikimediamessages.styles%7Cjquery.makeCollapsible.styles%7Cskins.vector.icons%2Cstyles%7Cskins.vector.search.codex.styles%7Cwikibase.client.init&amp;only=styles&amp;skin=vector-2022"> <script async="" src="/w/load.php?lang=en&amp;modules=startup&amp;only=scripts&amp;raw=1&amp;skin=vector-2022"></script> <meta name="ResourceLoaderDynamicStyles" content=""> <link rel="stylesheet" href="/w/load.php?lang=en&amp;modules=site.styles&amp;only=styles&amp;skin=vector-2022"> <meta name="generator" content="MediaWiki 1.44.0-wmf.4"> <meta name="referrer" content="origin"> <meta name="referrer" content="origin-when-cross-origin"> <meta name="robots" content="max-image-preview:standard"> <meta name="format-detection" content="telephone=no"> <meta property="og:image" content="https://upload.wikimedia.org/wikipedia/commons/thumb/a/ac/Paul_Nurse_portrait.jpg/1200px-Paul_Nurse_portrait.jpg"> <meta property="og:image:width" content="1200"> <meta property="og:image:height" content="1582"> <meta property="og:image" content="https://upload.wikimedia.org/wikipedia/commons/thumb/a/ac/Paul_Nurse_portrait.jpg/800px-Paul_Nurse_portrait.jpg"> <meta property="og:image:width" content="800"> <meta property="og:image:height" content="1054"> <meta property="og:image" content="https://upload.wikimedia.org/wikipedia/commons/thumb/a/ac/Paul_Nurse_portrait.jpg/640px-Paul_Nurse_portrait.jpg"> <meta property="og:image:width" content="640"> <meta property="og:image:height" content="843"> <meta name="viewport" content="width=1120"> <meta property="og:title" content="Paul Nurse - Simple English Wikipedia, the free encyclopedia"> <meta property="og:type" content="website"> <link rel="preconnect" href="//upload.wikimedia.org"> <link rel="alternate" media="only screen and (max-width: 640px)" href="//simple.m.wikipedia.org/wiki/Paul_Nurse"> <link rel="alternate" type="application/x-wiki" title="change this page" href="/w/index.php?title=Paul_Nurse&amp;action=edit"> <link rel="apple-touch-icon" href="/static/apple-touch/wikipedia.png"> <link rel="icon" href="/static/favicon/wikipedia.ico"> <link rel="search" type="application/opensearchdescription+xml" href="/w/rest.php/v1/search" title="Wikipedia (en)"> <link rel="EditURI" type="application/rsd+xml" href="//simple.wikipedia.org/w/api.php?action=rsd"> <link rel="canonical" href="https://simple.wikipedia.org/wiki/Paul_Nurse"> <link rel="license" href="https://creativecommons.org/licenses/by-sa/4.0/deed.simple"> <link rel="alternate" type="application/atom+xml" title="Wikipedia Atom feed" href="/w/index.php?title=Special:RecentChanges&amp;feed=atom"> <link rel="dns-prefetch" href="//meta.wikimedia.org" /> <link rel="dns-prefetch" href="//login.wikimedia.org"> </head> <body class="skin--responsive skin-vector skin-vector-search-vue mediawiki ltr sitedir-ltr mw-hide-empty-elt ns-0 ns-subject mw-editable page-Paul_Nurse rootpage-Paul_Nurse skin-vector-2022 action-view"><a class="mw-jump-link" href="#bodyContent">Jump to content</a> <div class="vector-header-container"> <header class="vector-header mw-header"> <div class="vector-header-start"> <nav class="vector-main-menu-landmark" aria-label="Site"> <div id="vector-main-menu-dropdown" class="vector-dropdown vector-main-menu-dropdown vector-button-flush-left vector-button-flush-right" > <input type="checkbox" id="vector-main-menu-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-main-menu-dropdown" class="vector-dropdown-checkbox " aria-label="Main menu" > <label id="vector-main-menu-dropdown-label" for="vector-main-menu-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-menu mw-ui-icon-wikimedia-menu"></span> <span class="vector-dropdown-label-text">Main menu</span> </label> <div class="vector-dropdown-content"> <div id="vector-main-menu-unpinned-container" class="vector-unpinned-container"> <div id="vector-main-menu" class="vector-main-menu vector-pinnable-element"> <div class="vector-pinnable-header vector-main-menu-pinnable-header vector-pinnable-header-unpinned" data-feature-name="main-menu-pinned" data-pinnable-element-id="vector-main-menu" data-pinned-container-id="vector-main-menu-pinned-container" data-unpinned-container-id="vector-main-menu-unpinned-container" > <div class="vector-pinnable-header-label">Main menu</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-main-menu.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-main-menu.unpin">hide</button> </div> <div id="p-navigation" class="vector-menu mw-portlet mw-portlet-navigation" > <div class="vector-menu-heading"> Getting around </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="n-mainpage-description" class="mw-list-item"><a href="/wiki/Main_Page" title="Visit the main page [z]" accesskey="z"><span>Main page</span></a></li><li id="n-portal" class="mw-list-item"><a href="/wiki/Wikipedia:Simple_start" title="About the project, what you can do, where to find things"><span>Simple start</span></a></li><li id="n-Simple-talk" class="mw-list-item"><a href="/wiki/Wikipedia:Simple_talk"><span>Simple talk</span></a></li><li id="n-recentchanges" class="mw-list-item"><a href="/wiki/Special:RecentChanges" title="The list of new changes in the wiki. [r]" accesskey="r"><span>New changes</span></a></li><li id="n-randompage" class="mw-list-item"><a href="/wiki/Special:Random" title="Show any page [x]" accesskey="x"><span>Show any page</span></a></li><li id="n-help" class="mw-list-item"><a href="/wiki/Help:Contents" title="The place to get help"><span>Help</span></a></li><li id="n-contactpage" class="mw-list-item"><a href="//simple.wikipedia.org/wiki/Wikipedia:Contact_us"><span>Contact us</span></a></li><li id="n-aboutsite" class="mw-list-item"><a href="/wiki/Wikipedia:About"><span>About Wikipedia</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> <a href="/wiki/Main_Page" class="mw-logo"> <img class="mw-logo-icon" src="/static/images/icons/wikipedia.png" alt="" aria-hidden="true" height="50" width="50"> <span class="mw-logo-container skin-invert"> <img class="mw-logo-wordmark" alt="Wikipedia" src="/static/images/mobile/copyright/wikipedia-wordmark-en.svg" style="width: 7.5em; height: 1.125em;"> <img class="mw-logo-tagline" alt="The Free Encyclopedia" src="/static/images/mobile/copyright/wikipedia-tagline-simple.svg" width="101" height="18" style="width: 6.3125em; height: 1.125em;"> </span> </a> </div> <div class="vector-header-end"> <div id="p-search" role="search" class="vector-search-box-vue vector-search-box-collapses vector-search-box-show-thumbnail vector-search-box-auto-expand-width vector-search-box"> <a href="/wiki/Special:Search" class="cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only search-toggle" title="Search Wikipedia [f]" accesskey="f"><span class="vector-icon mw-ui-icon-search mw-ui-icon-wikimedia-search"></span> <span>Search</span> </a> <div class="vector-typeahead-search-container"> <div class="cdx-typeahead-search cdx-typeahead-search--show-thumbnail cdx-typeahead-search--auto-expand-width"> <form action="/w/index.php" id="searchform" class="cdx-search-input cdx-search-input--has-end-button"> <div id="simpleSearch" class="cdx-search-input__input-wrapper" data-search-loc="header-moved"> <div class="cdx-text-input cdx-text-input--has-start-icon"> <input class="cdx-text-input__input" type="search" name="search" placeholder="Search Wikipedia" aria-label="Search Wikipedia" autocapitalize="sentences" title="Search Wikipedia [f]" accesskey="f" id="searchInput" > <span class="cdx-text-input__icon cdx-text-input__start-icon"></span> </div> <input type="hidden" name="title" value="Special:Search"> </div> <button class="cdx-button cdx-search-input__end-button">Search</button> </form> </div> </div> </div> <nav class="vector-user-links vector-user-links-wide" aria-label="Personal tools"> <div class="vector-user-links-main"> <div id="p-vector-user-menu-preferences" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <div id="p-vector-user-menu-userpage" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <nav class="vector-appearance-landmark" aria-label="Appearance"> <div id="vector-appearance-dropdown" class="vector-dropdown " title="Change the appearance of the page&#039;s font size, width, and color" > <input type="checkbox" id="vector-appearance-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-appearance-dropdown" class="vector-dropdown-checkbox " aria-label="Appearance" > <label id="vector-appearance-dropdown-label" for="vector-appearance-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-appearance mw-ui-icon-wikimedia-appearance"></span> <span class="vector-dropdown-label-text">Appearance</span> </label> <div class="vector-dropdown-content"> <div id="vector-appearance-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <div id="p-vector-user-menu-notifications" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <div id="p-vector-user-menu-overflow" class="vector-menu mw-portlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-sitesupport-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="//donate.wikimedia.org/wiki/Special:FundraiserRedirector?utm_source=donate&amp;utm_medium=sidebar&amp;utm_campaign=C13_simple.wikipedia.org&amp;uselang=en" class=""><span>Give to Wikipedia</span></a> </li> <li id="pt-createaccount-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="/w/index.php?title=Special:CreateAccount&amp;returnto=Paul+Nurse" title="You are encouraged to create an account and log in; however, it is not mandatory" class=""><span>Create account</span></a> </li> <li id="pt-login-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="/w/index.php?title=Special:UserLogin&amp;returnto=Paul+Nurse" title="You are encouraged to log in; however, it is not mandatory [o]" accesskey="o" class=""><span>Log in</span></a> </li> </ul> </div> </div> </div> <div id="vector-user-links-dropdown" class="vector-dropdown vector-user-menu vector-button-flush-right vector-user-menu-logged-out" title="More options" > <input type="checkbox" id="vector-user-links-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-user-links-dropdown" class="vector-dropdown-checkbox " aria-label="Personal tools" > <label id="vector-user-links-dropdown-label" for="vector-user-links-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-ellipsis mw-ui-icon-wikimedia-ellipsis"></span> <span class="vector-dropdown-label-text">Personal tools</span> </label> <div class="vector-dropdown-content"> <div id="p-personal" class="vector-menu mw-portlet mw-portlet-personal user-links-collapsible-item" title="User menu" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-sitesupport" class="user-links-collapsible-item mw-list-item"><a href="//donate.wikimedia.org/wiki/Special:FundraiserRedirector?utm_source=donate&amp;utm_medium=sidebar&amp;utm_campaign=C13_simple.wikipedia.org&amp;uselang=en"><span>Give to Wikipedia</span></a></li><li id="pt-createaccount" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=Special:CreateAccount&amp;returnto=Paul+Nurse" title="You are encouraged to create an account and log in; however, it is not mandatory"><span class="vector-icon mw-ui-icon-userAdd mw-ui-icon-wikimedia-userAdd"></span> <span>Create account</span></a></li><li id="pt-login" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=Special:UserLogin&amp;returnto=Paul+Nurse" title="You are encouraged to log in; however, it is not mandatory [o]" accesskey="o"><span class="vector-icon mw-ui-icon-logIn mw-ui-icon-wikimedia-logIn"></span> <span>Log in</span></a></li> </ul> </div> </div> <div id="p-user-menu-anon-editor" class="vector-menu mw-portlet mw-portlet-user-menu-anon-editor" > <div class="vector-menu-heading"> Pages for logged out editors <a href="/wiki/Help:Introduction" aria-label="Learn more about editing"><span>learn more</span></a> </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-anoncontribs" class="mw-list-item"><a href="/wiki/Special:MyContributions" title="A list of edits made from this IP address [y]" accesskey="y"><span>Contributions</span></a></li><li id="pt-anontalk" class="mw-list-item"><a href="/wiki/Special:MyTalk" title="Discussion about changes from this IP address [n]" accesskey="n"><span>Talk</span></a></li> </ul> </div> </div> </div> </div> </nav> </div> </header> </div> <div class="mw-page-container"> <div class="mw-page-container-inner"> <div class="vector-sitenotice-container"> <div id="siteNotice"><!-- CentralNotice --></div> </div> <div class="vector-column-start"> <div class="vector-main-menu-container"> <div id="mw-navigation"> <nav id="mw-panel" class="vector-main-menu-landmark" aria-label="Site"> <div id="vector-main-menu-pinned-container" class="vector-pinned-container"> </div> </nav> </div> </div> <div class="vector-sticky-pinned-container"> <nav id="mw-panel-toc" aria-label="Contents" data-event-name="ui.sidebar-toc" class="mw-table-of-contents-container vector-toc-landmark"> <div id="vector-toc-pinned-container" class="vector-pinned-container"> <div id="vector-toc" class="vector-toc vector-pinnable-element"> <div class="vector-pinnable-header vector-toc-pinnable-header vector-pinnable-header-pinned" data-feature-name="toc-pinned" data-pinnable-element-id="vector-toc" > <h2 class="vector-pinnable-header-label">Contents</h2> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-toc.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-toc.unpin">hide</button> </div> <ul class="vector-toc-contents" id="mw-panel-toc-list"> <li id="toc-mw-content-text" class="vector-toc-list-item vector-toc-level-1"> <a href="#" class="vector-toc-link"> <div class="vector-toc-text">Beginning</div> </a> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">1</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Toggle the table of contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Paul Nurse</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 49 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-49" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">49 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%A8%D9%88%D9%84_%D9%86%D8%B1%D8%B3" title="بول نرس – Arabic" lang="ar" hreflang="ar" data-title="بول نرس" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D9%BE%D8%A7%D8%A6%D9%88%D9%84_%D9%86%D8%B1%D8%B3" title="پائول نرس – South Azerbaijani" lang="azb" hreflang="azb" data-title="پائول نرس" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%AA%E0%A6%B2_%E0%A6%A8%E0%A6%BE%E0%A6%B0%E0%A7%8D%E0%A6%B8" title="পল নার্স – Bangla" lang="bn" hreflang="bn" data-title="পল নার্স" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%9F%D0%BE%D0%BB_%D0%9D%D0%B5%D1%80%D1%81" title="Пол Нерс – Belarusian" lang="be" hreflang="be" data-title="Пол Нерс" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%9F%D0%BE%D0%BB_%D0%9D%D1%8A%D1%80%D1%81" title="Пол Нърс – Bulgarian" lang="bg" hreflang="bg" data-title="Пол Нърс" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Catalan" lang="ca" hreflang="ca" data-title="Paul Nurse" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Czech" lang="cs" hreflang="cs" data-title="Paul Nurse" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Paul_M._Nurse" title="Paul M. Nurse – Danish" lang="da" hreflang="da" data-title="Paul M. Nurse" data-language-autonym="Dansk" data-language-local-name="Danish" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – German" lang="de" hreflang="de" data-title="Paul Nurse" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Paul_Maxime_Nurse" title="Paul Maxime Nurse – Estonian" lang="et" hreflang="et" data-title="Paul Maxime Nurse" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – English" lang="en" hreflang="en" data-title="Paul Nurse" data-language-autonym="English" data-language-local-name="English" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Spanish" lang="es" hreflang="es" data-title="Paul Nurse" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Basque" lang="eu" hreflang="eu" data-title="Paul Nurse" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D9%BE%D8%A7%D8%A6%D9%88%D9%84_%D9%86%D8%B1%D8%B3" title="پائول نرس – Persian" lang="fa" hreflang="fa" data-title="پائول نرس" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – French" lang="fr" hreflang="fr" data-title="Paul Nurse" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Paul_Maxime_Nurse" title="Paul Maxime Nurse – Irish" lang="ga" hreflang="ga" data-title="Paul Maxime Nurse" data-language-autonym="Gaeilge" data-language-local-name="Irish" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%ED%8F%B4_%EB%84%88%EC%8A%A4" title="폴 너스 – Korean" lang="ko" hreflang="ko" data-title="폴 너스" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Paul_Maxime_Nurse" title="Paul Maxime Nurse – Croatian" lang="hr" hreflang="hr" data-title="Paul Maxime Nurse" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Ido" lang="io" hreflang="io" data-title="Paul Nurse" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Paul_Maxime_Nurse" title="Paul Maxime Nurse – Indonesian" lang="id" hreflang="id" data-title="Paul Maxime Nurse" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-ia mw-list-item"><a href="https://ia.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Interlingua" lang="ia" hreflang="ia" data-title="Paul Nurse" data-language-autonym="Interlingua" data-language-local-name="Interlingua" class="interlanguage-link-target"><span>Interlingua</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Italian" lang="it" hreflang="it" data-title="Paul Nurse" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%A4%D7%95%D7%9C_%D7%A0%D7%A8%D7%A1" title="פול נרס – Hebrew" lang="he" hreflang="he" data-title="פול נרס" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%9F%D0%BE%D0%BB_%D0%9D%D0%B5%D1%80%D1%81" title="Пол Нерс – Kazakh" lang="kk" hreflang="kk" data-title="Пол Нерс" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Swahili" lang="sw" hreflang="sw" data-title="Paul Nurse" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Paulus_Nurse" title="Paulus Nurse – Latin" lang="la" hreflang="la" data-title="Paulus Nurse" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Hungarian" lang="hu" hreflang="hu" data-title="Paul Nurse" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%A8%D9%88%D9%84_%D9%86%D8%B1%D8%B3" title="بول نرس – Egyptian Arabic" lang="arz" hreflang="arz" data-title="بول نرس" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D9%BE%D8%A7%D8%A6%D9%88%D9%84_%D9%86%D8%B1%D8%B3" title="پائول نرس – Mazanderani" lang="mzn" hreflang="mzn" data-title="پائول نرس" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Malay" lang="ms" hreflang="ms" data-title="Paul Nurse" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Dutch" lang="nl" hreflang="nl" data-title="Paul Nurse" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%83%9D%E3%83%BC%E3%83%AB%E3%83%BB%E3%83%8A%E3%83%BC%E3%82%B9" title="ポール・ナース – Japanese" lang="ja" hreflang="ja" data-title="ポール・ナース" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Paul Nurse" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Occitan" lang="oc" hreflang="oc" data-title="Paul Nurse" data-language-autonym="Occitan" data-language-local-name="Occitan" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D9%BE%D8%A7%D9%84_%D9%86%D8%B1%D8%B3" title="پال نرس – Western Punjabi" lang="pnb" hreflang="pnb" data-title="پال نرس" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Paul_Maxime_Nurse" title="Paul Maxime Nurse – Low German" lang="nds" hreflang="nds" data-title="Paul Maxime Nurse" data-language-autonym="Plattdüütsch" data-language-local-name="Low German" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Polish" lang="pl" hreflang="pl" data-title="Paul Nurse" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Portuguese" lang="pt" hreflang="pt" data-title="Paul Nurse" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9D%D0%B5%D1%80%D1%81,_%D0%9F%D0%BE%D0%BB" title="Нерс, Пол – Russian" lang="ru" hreflang="ru" data-title="Нерс, Пол" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Slovak" lang="sk" hreflang="sk" data-title="Paul Nurse" data-language-autonym="Slovenčina" data-language-local-name="Slovak" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Slovenian" lang="sl" hreflang="sl" data-title="Paul Nurse" data-language-autonym="Slovenščina" data-language-local-name="Slovenian" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Finnish" lang="fi" hreflang="fi" data-title="Paul Nurse" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Swedish" lang="sv" hreflang="sv" data-title="Paul Nurse" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Turkish" lang="tr" hreflang="tr" data-title="Paul Nurse" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%9F%D0%BE%D0%BB_%D0%9D%D0%B5%D1%80%D1%81" title="Пол Нерс – Ukrainian" lang="uk" hreflang="uk" data-title="Пол Нерс" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D9%BE%D8%A7%D9%88%D9%84_%D9%86%D8%B1%D8%B3" title="پاول نرس – Urdu" lang="ur" hreflang="ur" data-title="پاول نرس" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Vietnamese" lang="vi" hreflang="vi" data-title="Paul Nurse" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/Paul_Nurse" title="Paul Nurse – Yoruba" lang="yo" hreflang="yo" data-title="Paul Nurse" data-language-autonym="Yorùbá" data-language-local-name="Yoruba" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E4%BF%9D%E7%BD%97%C2%B7%E7%BA%B3%E6%96%AF" title="保罗·纳斯 – Chinese" lang="zh" hreflang="zh" data-title="保罗·纳斯" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q219716#sitelinks-wikipedia" title="Edit interlanguage links" class="wbc-editpage">Change links</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Namespaces"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Paul_Nurse" title="View the content page [c]" accesskey="c"><span>Page</span></a></li><li id="ca-talk" class="vector-tab-noicon mw-list-item"><a href="/wiki/Talk:Paul_Nurse" rel="discussion" title="Discussion about the content page [t]" accesskey="t"><span>Talk</span></a></li> </ul> </div> </div> <div id="vector-variants-dropdown" class="vector-dropdown emptyPortlet" > <input type="checkbox" id="vector-variants-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-variants-dropdown" class="vector-dropdown-checkbox " aria-label="Change language variant" > <label id="vector-variants-dropdown-label" for="vector-variants-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">English</span> </label> <div class="vector-dropdown-content"> <div id="p-variants" class="vector-menu mw-portlet mw-portlet-variants emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> </div> </div> </nav> </div> <div id="right-navigation" class="vector-collapsible"> <nav aria-label="Views"> <div id="p-views" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-views" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-view" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Paul_Nurse"><span>Read</span></a></li><li id="ca-ve-edit" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Paul_Nurse&amp;veaction=edit" title="Edit this page [v]" accesskey="v"><span>Change</span></a></li><li id="ca-edit" class="collapsible vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Paul_Nurse&amp;action=edit" title="Edit the source code of this page [e]" accesskey="e"><span>Change source</span></a></li><li id="ca-history" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Paul_Nurse&amp;action=history" title="Past revisions of this page [h]" accesskey="h"><span>View history</span></a></li> </ul> </div> </div> </nav> <nav class="vector-page-tools-landmark" aria-label="Page tools"> <div id="vector-page-tools-dropdown" class="vector-dropdown vector-page-tools-dropdown" > <input type="checkbox" id="vector-page-tools-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-tools-dropdown" class="vector-dropdown-checkbox " aria-label="Tools" > <label id="vector-page-tools-dropdown-label" for="vector-page-tools-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">Tools</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-tools-unpinned-container" class="vector-unpinned-container"> <div id="vector-page-tools" class="vector-page-tools vector-pinnable-element"> <div class="vector-pinnable-header vector-page-tools-pinnable-header vector-pinnable-header-unpinned" data-feature-name="page-tools-pinned" data-pinnable-element-id="vector-page-tools" data-pinned-container-id="vector-page-tools-pinned-container" data-unpinned-container-id="vector-page-tools-unpinned-container" > <div class="vector-pinnable-header-label">Tools</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-page-tools.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-page-tools.unpin">hide</button> </div> <div id="p-cactions" class="vector-menu mw-portlet mw-portlet-cactions emptyPortlet vector-has-collapsible-items" title="More options" > <div class="vector-menu-heading"> Actions </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-more-view" class="selected vector-more-collapsible-item mw-list-item"><a href="/wiki/Paul_Nurse"><span>Read</span></a></li><li id="ca-more-ve-edit" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Paul_Nurse&amp;veaction=edit" title="Edit this page [v]" accesskey="v"><span>Change</span></a></li><li id="ca-more-edit" class="collapsible vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Paul_Nurse&amp;action=edit" title="Edit the source code of this page [e]" accesskey="e"><span>Change source</span></a></li><li id="ca-more-history" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Paul_Nurse&amp;action=history"><span>View history</span></a></li> </ul> </div> </div> <div id="p-tb" class="vector-menu mw-portlet mw-portlet-tb" > <div class="vector-menu-heading"> General </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="t-whatlinkshere" class="mw-list-item"><a href="/wiki/Special:WhatLinksHere/Paul_Nurse" title="A list of all wiki pages that link here [j]" accesskey="j"><span>What links here</span></a></li><li id="t-recentchangeslinked" class="mw-list-item"><a href="/wiki/Special:RecentChangesLinked/Paul_Nurse" rel="nofollow" title="New changes in pages linked from this page [k]" accesskey="k"><span>Related changes</span></a></li><li id="t-upload" class="mw-list-item"><a href="//commons.wikimedia.org/wiki/Special:UploadWizard" title="Upload files [u]" accesskey="u"><span>Upload file</span></a></li><li id="t-specialpages" class="mw-list-item"><a href="/wiki/Special:SpecialPages" title="A list of all special pages [q]" accesskey="q"><span>Special pages</span></a></li><li id="t-permalink" class="mw-list-item"><a href="/w/index.php?title=Paul_Nurse&amp;oldid=9574995" title="Permanent link to this revision of this page"><span>Permanent link</span></a></li><li id="t-info" class="mw-list-item"><a href="/w/index.php?title=Paul_Nurse&amp;action=info" title="More information about this page"><span>Page information</span></a></li><li id="t-cite" class="mw-list-item"><a href="/w/index.php?title=Special:CiteThisPage&amp;page=Paul_Nurse&amp;id=9574995&amp;wpFormIdentifier=titleform" title="Information on how to cite this page"><span>Cite this page</span></a></li><li id="t-urlshortener" class="mw-list-item"><a href="/w/index.php?title=Special:UrlShortener&amp;url=https%3A%2F%2Fsimple.wikipedia.org%2Fwiki%2FPaul_Nurse"><span>Get shortened URL</span></a></li><li id="t-urlshortener-qrcode" class="mw-list-item"><a href="/w/index.php?title=Special:QrCode&amp;url=https%3A%2F%2Fsimple.wikipedia.org%2Fwiki%2FPaul_Nurse"><span>Download QR code</span></a></li> </ul> </div> </div> <div id="p-coll-print_export" class="vector-menu mw-portlet mw-portlet-coll-print_export" > <div class="vector-menu-heading"> Print/export </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="coll-create_a_book" class="mw-list-item"><a href="/w/index.php?title=Special:Book&amp;bookcmd=book_creator&amp;referer=Paul+Nurse"><span>Make a book</span></a></li><li id="coll-download-as-rl" class="mw-list-item"><a href="/w/index.php?title=Special:DownloadAsPdf&amp;page=Paul_Nurse&amp;action=show-download-screen"><span>Download as PDF</span></a></li><li id="t-print" class="mw-list-item"><a href="/w/index.php?title=Paul_Nurse&amp;printable=yes" title="Printable version of this page [p]" accesskey="p"><span>Page for printing</span></a></li> </ul> </div> </div> <div id="p-wikibase-otherprojects" class="vector-menu mw-portlet mw-portlet-wikibase-otherprojects" > <div class="vector-menu-heading"> In other projects </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="wb-otherproject-link wb-otherproject-commons mw-list-item"><a href="https://commons.wikimedia.org/wiki/Category:Paul_Nurse" hreflang="en"><span>Wikimedia Commons</span></a></li><li id="t-wikibase" class="wb-otherproject-link wb-otherproject-wikibase-dataitem mw-list-item"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q219716" title="Link to connected data repository item [g]" accesskey="g"><span>Wikidata item</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> </div> </div> </div> <div class="vector-column-end"> <div class="vector-sticky-pinned-container"> <nav class="vector-page-tools-landmark" aria-label="Page tools"> <div id="vector-page-tools-pinned-container" class="vector-pinned-container"> </div> </nav> <nav class="vector-appearance-landmark" aria-label="Appearance"> <div id="vector-appearance-pinned-container" class="vector-pinned-container"> <div id="vector-appearance" class="vector-appearance vector-pinnable-element"> <div class="vector-pinnable-header vector-appearance-pinnable-header vector-pinnable-header-pinned" data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">Appearance</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">hide</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> <div id="mw-indicator-Nobel_prize_medal.svg" class="mw-indicator"><div class="mw-parser-output"><span typeof="mw:File"><a href="/wiki/Nobel_Prize" title="Nobel Prize"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/20px-Nobel_prize_medal.svg.png" decoding="async" width="20" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/30px-Nobel_prize_medal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/40px-Nobel_prize_medal.svg.png 2x" data-file-width="150" data-file-height="150" /></a></span></div></div> </div> <div id="siteSub" class="noprint">From Simple English Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><style data-mw-deduplicate="TemplateStyles:r9689576">.mw-parser-output .infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}body.skin-minerva .mw-parser-output .infobox-header,body.skin-minerva .mw-parser-output .infobox-subheader,body.skin-minerva .mw-parser-output .infobox-above,body.skin-minerva .mw-parser-output .infobox-title,body.skin-minerva .mw-parser-output .infobox-image,body.skin-minerva .mw-parser-output .infobox-full-data,body.skin-minerva .mw-parser-output .infobox-below{text-align:center}@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox-full-data div{background:#1f1f23!important;color:#f8f9fa}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox-full-data div{background:#1f1f23!important;color:#f8f9fa}}</style><table class="infobox biography vcard"><tbody><tr><th colspan="2" class="infobox-above" style="font-size:125%;"><div class="fn" style="display:inline-block">Sir Paul Nurse</div></th></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Paul_Nurse_portrait.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/ac/Paul_Nurse_portrait.jpg/220px-Paul_Nurse_portrait.jpg" decoding="async" width="220" height="290" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/ac/Paul_Nurse_portrait.jpg/330px-Paul_Nurse_portrait.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/ac/Paul_Nurse_portrait.jpg/440px-Paul_Nurse_portrait.jpg 2x" data-file-width="1648" data-file-height="2172" /></a></span><div class="infobox-caption">Paul Nurse</div></td></tr><tr><th scope="row" class="infobox-label">Born</th><td class="infobox-data"><span style="display:none"> (<span class="bday">1949-01-25</span>) </span>25 January 1949<span class="noprint ForceAgeToShow"> (age&#160;75)</span><br /><div style="display:inline" class="birthplace"><a href="/wiki/Norwich" title="Norwich">Norwich</a>, <a href="/wiki/Norfolk" title="Norfolk">Norfolk</a>, <a href="/wiki/England" title="England">England</a></div></td></tr><tr><th scope="row" class="infobox-label">Nationality</th><td class="infobox-data category"><a href="/wiki/British_people" title="British people">British</a></td></tr><tr><th scope="row" class="infobox-label">Alma&#160;mater</th><td class="infobox-data"><a href="/wiki/University_of_Birmingham" title="University of Birmingham">University of Birmingham</a><br /><a href="/w/index.php?title=University_of_East_Anglia&amp;action=edit&amp;redlink=1" class="new" title="University of East Anglia (not yet started)">University of East Anglia</a></td></tr><tr><th scope="row" class="infobox-label">Known&#160;for</th><td class="infobox-data"><a href="/wiki/Cell_cycle" class="mw-redirect" title="Cell cycle">Cell cycle</a> regulation</td></tr><tr><th scope="row" class="infobox-label">Awards</th><td class="infobox-data"><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" class="mw-redirect" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a> 2001<br />Copley Medal in 2005</td></tr><tr><td colspan="2" class="infobox-full-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r9689576"><b>Scientific career</b></td></tr><tr><th scope="row" class="infobox-label">Fields</th><td class="infobox-data category"><a href="/wiki/Genetics" title="Genetics">Genetics</a>/<a href="/wiki/Cell_biology" title="Cell biology">cell biology</a></td></tr><tr style="display:none"><td colspan="2"> </td></tr></tbody></table> <p><a href="/wiki/Knight_Bachelor" title="Knight Bachelor">Sir</a> <b>Paul Maxime Nurse</b> <small><a href="/wiki/Order_of_Merit" title="Order of Merit">OM</a> <a href="/wiki/Order_of_the_Companions_of_Honour" title="Order of the Companions of Honour">CH</a></small> <a href="/wiki/Royal_Society" class="mw-redirect" title="Royal Society"><small>PRS</small></a> (Paul Maxime Nurse, born <a href="/wiki/Norwich" title="Norwich">Norwich</a>, 25 January 1949) is a <a href="/wiki/British_people" title="British people">British</a> <a href="/wiki/Genetics" title="Genetics">geneticist</a> and <a href="/wiki/Cell_biology" title="Cell biology">cell biologist</a>.<sup id="cite_ref-illpres_1-0" class="reference"><a href="#cite_note-illpres-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> </p><p>He was awarded the 2001 <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" class="mw-redirect" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a>, with <a href="/wiki/Leland_Hartwell" class="mw-redirect" title="Leland Hartwell">Leland Hartwell</a> and <a href="/wiki/Timothy_Hunt" class="mw-redirect" title="Timothy Hunt">Timothy Hunt</a>, for their discoveries about <a href="/wiki/Cell_division" title="Cell division">cell division</a>. </p><p>They discovered cycle of cell division is regulated by <a href="/w/index.php?title=Cyclin&amp;action=edit&amp;redlink=1" class="new" title="Cyclin (not yet started)">cyclin</a> and cyclin dependent <a href="/w/index.php?title=Kinase&amp;action=edit&amp;redlink=1" class="new" title="Kinase (not yet started)">kinases</a>.<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> </p><p>Sir Paul is the current <a href="/wiki/President_of_the_Royal_Society" class="mw-redirect" title="President of the Royal Society">President of the Royal Society</a> and Chief Executive and Director of the <a href="/w/index.php?title=UK_Centre_for_Medical_Research_and_Innovation&amp;action=edit&amp;redlink=1" class="new" title="UK Centre for Medical Research and Innovation (not yet started)">UK Centre for Medical Research and Innovation</a>. </p><p>He is also <a href="/wiki/Emeritus" title="Emeritus">Emeritus</a> Professor and <a href="/w/index.php?title=Head_of_the_Laboratory&amp;action=edit&amp;redlink=1" class="new" title="Head of the Laboratory (not yet started)">Head of the Laboratory</a> for <a href="/wiki/Yeast" title="Yeast">Yeast</a> genetics and <a href="/wiki/Cell_biology" title="Cell biology">Cell Biology</a> at the <a href="/wiki/Rockefeller_University" title="Rockefeller University">Rockefeller University</a>, <a href="/wiki/New_York_City" title="New York City">New York City</a>.<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> </p><p>Nurse was awarded the <a href="/wiki/Royal_Society" class="mw-redirect" title="Royal Society">Royal Society</a>'s <a href="/wiki/Copley_Medal" title="Copley Medal">Copley Medal</a> in 2005. </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Paul_Nurse&amp;veaction=edit&amp;section=1" title="Change section: References" class="mw-editsection-visualeditor"><span>change</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Paul_Nurse&amp;action=edit&amp;section=1" title="Edit section&#039;s source code: References"><span>change source</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r9724317">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-illpres-1"><span class="mw-cite-backlink"><a href="#cite_ref-illpres_1-0">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.nobelprize.org/nobel_prizes/medicine/laureates/2001/illpres/index.html">The Nobel Prize in Physiology or Medicine 2001</a> Illustrated Lecture</span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><a href="#cite_ref-2">↑</a></span> <span class="reference-text">Nurse, Paul 2004. <i>The great ideas of biology</i>. The Romanes Lecture for 2003, Oxford University Press. <style data-mw-deduplicate="TemplateStyles:r9724332">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/0-19-951897-1" title="Special:BookSources/0-19-951897-1">0-19-951897-1</a></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><a href="#cite_ref-3">↑</a></span> <span class="reference-text">Cyclins are a family of proteins that control the way cells go through the cell cycle by activating cyclin-dependent kinase enzymes. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r9724332"><cite id="CITEREFGalderisi_UJori_FPGiordano_A2003" class="citation journal cs1">Galderisi U; Jori FP; Giordano A (2003). "Cell cycle regulation and neural differentiation". <i>Oncogene</i>. <b>22</b> (33): 5208–19. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2Fsj.onc.1206558">10.1038/sj.onc.1206558</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/12910258">12910258</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a>&#160;<a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:19528945">19528945</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Oncogene&amp;rft.atitle=Cell+cycle+regulation+and+neural+differentiation&amp;rft.volume=22&amp;rft.issue=33&amp;rft.pages=5208-19&amp;rft.date=2003&amp;rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A19528945%23id-name%3DS2CID&amp;rft_id=info%3Apmid%2F12910258&amp;rft_id=info%3Adoi%2F10.1038%2Fsj.onc.1206558&amp;rft.au=Galderisi+U&amp;rft.au=Jori+FP&amp;rft.au=Giordano+A&amp;rfr_id=info%3Asid%2Fsimple.wikipedia.org%3APaul+Nurse" class="Z3988"></span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><a href="#cite_ref-4">↑</a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r9724332"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20110419182754/http://www.rockefeller.edu/research/faculty/abstract.php?id=316">"Rockefeller University: Sir Paul Nurse"</a>. Archived from <a rel="nofollow" class="external text" href="http://www.rockefeller.edu/research/faculty/abstract.php?id=316">the original</a> on 2011-04-19<span class="reference-accessdate">. Retrieved <span class="nowrap">2011-04-18</span></span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=unknown&amp;rft.btitle=Rockefeller+University%3A+Sir+Paul+Nurse&amp;rft_id=http%3A%2F%2Fwww.rockefeller.edu%2Fresearch%2Ffaculty%2Fabstract.php%3Fid%3D316&amp;rfr_id=info%3Asid%2Fsimple.wikipedia.org%3APaul+Nurse" class="Z3988"></span></span> </li> </ol></div></div> <div class="navbox-styles nomobile"><style data-mw-deduplicate="TemplateStyles:r9122492">.mw-parser-output .navbox{box-sizing:border-box;border:1px solid #a2a9b1;width:100%;clear:both;font-size:88%;text-align:center;padding:1px;margin:1em auto 0}.mw-parser-output .navbox .navbox{margin-top:0}.mw-parser-output .navbox+.navbox,.mw-parser-output .navbox+.navbox-styles+.navbox{margin-top:-1px}.mw-parser-output .navbox-inner,.mw-parser-output .navbox-subgroup{width:100%}.mw-parser-output .navbox-group,.mw-parser-output .navbox-title,.mw-parser-output .navbox-abovebelow{padding:0.25em 1em;line-height:1.5em;text-align:center}.mw-parser-output .navbox-group{white-space:nowrap;text-align:right}.mw-parser-output .navbox,.mw-parser-output .navbox-subgroup{background-color:#fdfdfd}.mw-parser-output .navbox-list{line-height:1.5em;border-color:#fdfdfd}.mw-parser-output .navbox-list-with-group{text-align:left;border-left-width:2px;border-left-style:solid}.mw-parser-output tr+tr>.navbox-abovebelow,.mw-parser-output tr+tr>.navbox-group,.mw-parser-output tr+tr>.navbox-image,.mw-parser-output tr+tr>.navbox-list{border-top:2px solid #fdfdfd}.mw-parser-output .navbox-title{background-color:#ccf}.mw-parser-output .navbox-abovebelow,.mw-parser-output .navbox-group,.mw-parser-output .navbox-subgroup .navbox-title{background-color:#ddf}.mw-parser-output .navbox-subgroup .navbox-group,.mw-parser-output .navbox-subgroup .navbox-abovebelow{background-color:#e6e6ff}.mw-parser-output .navbox-even{background-color:#f7f7f7}.mw-parser-output .navbox-odd{background-color:transparent}.mw-parser-output .navbox .hlist td dl,.mw-parser-output .navbox .hlist td ol,.mw-parser-output .navbox .hlist td ul,.mw-parser-output .navbox td.hlist dl,.mw-parser-output .navbox td.hlist ol,.mw-parser-output .navbox td.hlist ul{padding:0.125em 0}.mw-parser-output .navbox .navbar{display:block;font-size:100%}.mw-parser-output .navbox-title .navbar{float:left;text-align:left;margin-right:0.5em}</style></div><div role="navigation" class="navbox" aria-labelledby="Winners_of_the_Nobel_Prize_in_Physiology_or_Medicine" style="padding:3px"><table class="nowraplinks mw-collapsible mw-collapsed navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2" style="background:#d3d3d3;"><style data-mw-deduplicate="TemplateStyles:r9122447">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Nobel_Prize_in_Physiology_or_Medicine" title="Template:Nobel Prize in Physiology or Medicine"><abbr title="View this template" style=";background:#d3d3d3;;background:none transparent;border:none;box-shadow:none;padding:0;">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Nobel_Prize_in_Physiology_or_Medicine" title="Template talk:Nobel Prize in Physiology or Medicine"><abbr title="Discuss this template" style=";background:#d3d3d3;;background:none transparent;border:none;box-shadow:none;padding:0;">t</abbr></a></li><li class="nv-edit"><a class="external text" href="https://simple.wikipedia.org/w/index.php?title=Template:Nobel_Prize_in_Physiology_or_Medicine&amp;action=edit"><abbr title="Edit this template" style=";background:#d3d3d3;;background:none transparent;border:none;box-shadow:none;padding:0;">e</abbr></a></li></ul></div><div id="Winners_of_the_Nobel_Prize_in_Physiology_or_Medicine" style="font-size:114%;margin:0 4em">Winners of the <a href="/wiki/Nobel_Prize" title="Nobel Prize">Nobel Prize</a> in <a href="/wiki/List_of_Nobel_Prize_winners_in_Physiology_or_Medicine" title="List of Nobel Prize winners in Physiology or Medicine">Physiology or Medicine</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1901 – 1925</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><div> <p><a href="/wiki/Emil_Adolf_von_Behring" title="Emil Adolf von Behring">Behring</a> (1901)&#160;<b>&#183;</b>&#32; <a href="/wiki/Ronald_Ross" title="Ronald Ross">Ross</a> (1902)&#160;<b>&#183;</b>&#32; <a href="/wiki/Niels_Ryberg_Finsen" title="Niels Ryberg Finsen">Finsen</a> (1903)&#160;<b>&#183;</b>&#32; <a href="/wiki/Ivan_Pavlov" title="Ivan Pavlov">Pavlov</a> (1904)&#160;<b>&#183;</b>&#32; <a href="/wiki/Robert_Koch" title="Robert Koch">Koch</a> (1905)&#160;<b>&#183;</b>&#32; <a href="/wiki/Camillo_Golgi" title="Camillo Golgi">Golgi</a> / <a href="/wiki/Santiago_Ram%C3%B3n_y_Cajal" title="Santiago Ramón y Cajal">Ramón y Cajal</a> (1906)&#160;<b>&#183;</b>&#32; <a href="/wiki/Alphonse_Laveran" title="Alphonse Laveran">Laveran</a> (1907)&#160;<b>&#183;</b>&#32; <a href="/wiki/Ilya_Mechnikov" title="Ilya Mechnikov">Mechnikov</a> / <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Ehrlich</a> (1908)&#160;<b>&#183;</b>&#32; <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Kocher</a> (1909)&#160;<b>&#183;</b>&#32; <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Kossel</a> (1910)&#160;<b>&#183;</b>&#32; <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Gullstrand</a> (1911)&#160;<b>&#183;</b>&#32; <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Carrel</a> (1912)&#160;<b>&#183;</b>&#32; <a href="/wiki/Charles_Robert_Richet" title="Charles Robert Richet">Richet</a> (1913)&#160;<b>&#183;</b>&#32; <a href="/wiki/Robert_B%C3%A1r%C3%A1ny" title="Robert Bárány">Bárány</a> (1914)&#160;<b>&#183;</b>&#32; <a href="/wiki/Jules_Bordet" title="Jules Bordet">Bordet</a> (1919)&#160;<b>&#183;</b>&#32; <a href="/wiki/August_Krogh" title="August Krogh">Krogh</a> (1920)&#160;<b>&#183;</b>&#32; <a href="/wiki/Archibald_Hill" title="Archibald Hill">Hill</a> / <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Meyerhof</a> (1922)&#160;<b>&#183;</b>&#32; <a href="/wiki/Frederick_Banting" title="Frederick Banting">Banting</a> / <a href="/wiki/John_Macleod" title="John Macleod">Macleod</a> (1923)&#160;<b>&#183;</b>&#32; <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Einthoven</a> (1924) </p> </div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1926 – 1950</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><div> <p><a href="/wiki/Johannes_Fibiger" title="Johannes Fibiger">Fibiger</a> (1926)&#160;<b>&#183;</b>&#32; <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Wagner-Jauregg</a> (1927)&#160;<b>&#183;</b>&#32; <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Nicolle</a> (1928)&#160;<b>&#183;</b>&#32; <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Eijkman</a> / <a href="/wiki/Frederick_Hopkins" title="Frederick Hopkins">Hopkins</a> (1929)&#160;<b>&#183;</b>&#32; <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Landsteiner</a> (1930)&#160;<b>&#183;</b>&#32; <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Warburg</a> (1931)&#160;<b>&#183;</b>&#32; <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Sherrington</a> / <a href="/wiki/Edgar_Adrian,_1st_Baron_Adrian" title="Edgar Adrian, 1st Baron Adrian">Adrian</a> (1932)&#160;<b>&#183;</b>&#32; <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Morgan</a> (1933)&#160;<b>&#183;</b>&#32; <a href="/wiki/George_Whipple" title="George Whipple">Whipple</a> / <a href="/wiki/George_Minot" title="George Minot">Minot</a> / <a href="/wiki/William_P._Murphy" title="William P. Murphy">Murphy</a> (1934)&#160;<b>&#183;</b>&#32; <a href="/wiki/Hans_Spemann" title="Hans Spemann">Spemann</a> (1935)&#160;<b>&#183;</b>&#32; <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Dale</a> / <a href="/wiki/Otto_Loewi" title="Otto Loewi">Loewi</a> (1936)&#160;<b>&#183;</b>&#32; <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Szent-Györgyi</a> (1937)&#160;<b>&#183;</b>&#32; <a href="/wiki/Corneille_Heymans" title="Corneille Heymans">Heymans</a> (1938)&#160;<b>&#183;</b>&#32; <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Domagk</a> (1939)&#160;<b>&#183;</b>&#32; <a href="/wiki/Henrik_Dam" title="Henrik Dam">Dam</a> / <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Doisy</a> (1943)&#160;<b>&#183;</b>&#32; <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Erlanger</a> / <a href="/wiki/Herbert_Gasser" title="Herbert Gasser">Gasser</a> (1944)&#160;<b>&#183;</b>&#32; <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Fleming</a> / <a href="/wiki/Ernst_Chain" title="Ernst Chain">Chain</a> / <a href="/wiki/Howard_Florey" title="Howard Florey">Florey</a> (1945)&#160;<b>&#183;</b>&#32; <a href="/wiki/H._J._Muller" title="H. J. Muller">Muller</a> (1946)&#160;<b>&#183;</b>&#32; <a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">C.Cori</a> / <a href="/wiki/Gerty_Cori" title="Gerty Cori">G.Cori</a> / <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Houssay</a> (1947)&#160;<b>&#183;</b>&#32; <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Müller</a> (1948)&#160;<b>&#183;</b>&#32; <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Hess</a> / <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">Moniz</a> (1949)&#160;<b>&#183;</b>&#32; <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Kendall</a> / <a href="/wiki/Tadeus_Reichstein" title="Tadeus Reichstein">Reichstein</a> / <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Hench</a> (1950) </p> </div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1951 – 1975</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><div> <p><a href="/wiki/Max_Theiler" title="Max Theiler">Theiler</a> (1951)&#160;<b>&#183;</b>&#32; <a href="/wiki/Selman_Waksman" title="Selman Waksman">Waksman</a> (1952)&#160;<b>&#183;</b>&#32; <a href="/wiki/Hans_Krebs" title="Hans Krebs">Krebs</a> / <a href="/wiki/Fritz_Albert_Lipmann" class="mw-redirect" title="Fritz Albert Lipmann">Lipmann</a> (1953)&#160;<b>&#183;</b>&#32; <a href="/wiki/John_Franklin_Enders" class="mw-redirect" title="John Franklin Enders">Enders</a> / <a href="/wiki/Thomas_Huckle_Weller" class="mw-redirect" title="Thomas Huckle Weller">Weller</a> / <a href="/wiki/Frederick_Chapman_Robbins" class="mw-redirect" title="Frederick Chapman Robbins">Robbins</a> (1954)&#160;<b>&#183;</b>&#32; <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Theorell</a> (1955)&#160;<b>&#183;</b>&#32; <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" class="mw-redirect" title="André Frédéric Cournand">Cournand</a> / <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Forssmann</a> / <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Richards</a> (1956)&#160;<b>&#183;</b>&#32; <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Bovet</a> (1957)&#160;<b>&#183;</b>&#32; <a href="/wiki/George_Beadle" title="George Beadle">Beadle</a> / <a href="/wiki/Edward_Tatum" title="Edward Tatum">Tatum</a> / <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Lederberg</a> (1958)&#160;<b>&#183;</b>&#32; <a href="/wiki/Severo_Ochoa" class="mw-redirect" title="Severo Ochoa">Ochoa</a> / <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Kornberg</a> (1959)&#160;<b>&#183;</b>&#32; <a href="/wiki/Frank_Macfarlane_Burnet" title="Frank Macfarlane Burnet">Burnet</a> / <a href="/wiki/Peter_Medawar" title="Peter Medawar">Medawar</a> (1960)&#160;<b>&#183;</b>&#32; <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Békésy</a> (1961)&#160;<b>&#183;</b>&#32; <a href="/wiki/Francis_Crick" title="Francis Crick">Crick</a> / <a href="/wiki/James_D._Watson" title="James D. Watson">Watson</a> / <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Wilkins</a> (1962)&#160;<b>&#183;</b>&#32; <a href="/wiki/John_Eccles_(neurophysiologist)" title="John Eccles (neurophysiologist)">Eccles</a> / <a href="/wiki/Alan_Hodgkin" title="Alan Hodgkin">Hodgkin</a> / <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Huxley</a> (1963)&#160;<b>&#183;</b>&#32; <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Bloch</a> / <a href="/wiki/Feodor_Felix_Konrad_Lynen" class="mw-redirect" title="Feodor Felix Konrad Lynen">Lynen</a> (1964)&#160;<b>&#183;</b>&#32; <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">Jacob</a> / <a href="/wiki/Andr%C3%A9_Michel_Lwoff" class="mw-redirect" title="André Michel Lwoff">Lwoff</a> / <a href="/wiki/Jacques_Monod" title="Jacques Monod">Monod</a> (1965)&#160;<b>&#183;</b>&#32; <a href="/wiki/Francis_Peyton_Rous" class="mw-redirect" title="Francis Peyton Rous">Rous</a> / <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Huggins</a> (1966)&#160;<b>&#183;</b>&#32; <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Granit</a> / <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Hartline</a> / <a href="/wiki/George_Wald" title="George Wald">Wald</a> (1967)&#160;<b>&#183;</b>&#32; <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Holley</a> / <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Khorana</a> / <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Nirenberg</a> (1968)&#160;<b>&#183;</b>&#32; <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Delbrück</a> / <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Hershey</a> / <a href="/wiki/Salvador_Luria" title="Salvador Luria">Luria</a> (1969)&#160;<b>&#183;</b>&#32; <a href="/wiki/Bernard_Katz" title="Bernard Katz">Katz</a> / <a href="/wiki/Ulf_von_Euler" title="Ulf von Euler">Euler</a> / <a href="/wiki/Julius_Axelrod" class="mw-redirect" title="Julius Axelrod">Axelrod</a> (1970)&#160;<b>&#183;</b>&#32; <a href="/wiki/Earl_Wilbur_Sutherland_Jr." title="Earl Wilbur Sutherland Jr.">Sutherland</a> (1971)&#160;<b>&#183;</b>&#32; <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Edelman</a> / <a href="/wiki/Rodney_Robert_Porter" class="mw-redirect" title="Rodney Robert Porter">Porter</a> (1972)&#160;<b>&#183;</b>&#32; <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Frisch</a> / <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Lorenz</a> / <a href="/wiki/Niko_Tinbergen" title="Niko Tinbergen">Tinbergen</a> (1973)&#160;<b>&#183;</b>&#32; <a href="/wiki/Albert_Claude" title="Albert Claude">Claude</a> / <a href="/wiki/Christian_de_Duve" title="Christian de Duve">Duve</a> / <a href="/wiki/George_Emil_Palade" title="George Emil Palade">Palade</a> (1974)&#160;<b>&#183;</b>&#32; <a href="/wiki/David_Baltimore" title="David Baltimore">Baltimore</a> / <a href="/wiki/Howard_Temin" title="Howard Temin">Temin</a> / <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Dulbecco</a> (1975) </p> </div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1976 – 2000</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><div> <p><a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Blumberg</a> / <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Gajdusek</a> (1976)&#160;<b>&#183;</b>&#32; <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Guillemin</a> / <a href="/wiki/Andrew_Schally" title="Andrew Schally">Schally</a> / <a href="/wiki/Rosalyn_Yalow" title="Rosalyn Yalow">Yalow</a> (1977)&#160;<b>&#183;</b>&#32; <a href="/wiki/Werner_Arber" title="Werner Arber">Arber</a> / <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Nathans</a> / <a href="/wiki/Hamilton_Smith" title="Hamilton Smith">Smith</a> (1978)&#160;<b>&#183;</b>&#32; <a href="/wiki/Allan_McLeod_Cormack" title="Allan McLeod Cormack">Cormack</a> / <a href="/wiki/Godfrey_Hounsfield" class="mw-redirect" title="Godfrey Hounsfield">Hounsfield</a> (1979)&#160;<b>&#183;</b>&#32; <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Benacerraf</a> / <a href="/wiki/Jean_Dausset" title="Jean Dausset">Dausset</a> / <a href="/wiki/George_Davis_Snell" title="George Davis Snell">Snell</a> (1980)&#160;<b>&#183;</b>&#32; <a href="/wiki/Roger_Sperry" class="mw-redirect" title="Roger Sperry">Sperry</a> / <a href="/wiki/David_Hubel" title="David Hubel">Hubel</a> / <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Wiesel</a> (1981)&#160;<b>&#183;</b>&#32; <a href="/wiki/Sune_Bergstr%C3%B6m" title="Sune Bergström">Bergström</a> / <a href="/wiki/Bengt_I._Samuelsson" title="Bengt I. Samuelsson">Samuelsson</a> / <a href="/wiki/John_Robert_Vane" class="mw-redirect" title="John Robert Vane">Vane</a> (1982)&#160;<b>&#183;</b>&#32; <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">McClintock</a> (1983)&#160;<b>&#183;</b>&#32; <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Jerne</a> / <a href="/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler">Köhler</a> / <a href="/wiki/C%C3%A9sar_Milstein" class="mw-redirect" title="César Milstein">Milstein</a> (1984)&#160;<b>&#183;</b>&#32; <a href="/wiki/Michael_Stuart_Brown" class="mw-redirect" title="Michael Stuart Brown">Brown</a> / <a href="/wiki/Joseph_L._Goldstein" title="Joseph L. Goldstein">Goldstein</a> (1985)&#160;<b>&#183;</b>&#32; <a href="/wiki/Stanley_Cohen" title="Stanley Cohen">Stanley Cohen</a> / <a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Levi-Montalcini</a> (1986)&#160;<b>&#183;</b>&#32; <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Tonegawa</a> (1987)&#160;<b>&#183;</b>&#32; <a href="/wiki/James_Black" title="James Black">Black</a> / <a href="/wiki/Gertrude_B._Elion" title="Gertrude B. Elion">Elion</a> / <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">Hitchings</a> (1988)&#160;<b>&#183;</b>&#32; <a href="/wiki/J._Michael_Bishop" title="J. Michael Bishop">Bishop</a> / <a href="/wiki/Harold_Varmus" title="Harold Varmus">Varmus</a> (1989)&#160;<b>&#183;</b>&#32; <a href="/wiki/Joseph_Murray" title="Joseph Murray">Murray</a> / <a href="/wiki/E._Donnall_Thomas" title="E. Donnall Thomas">Thomas</a> (1990)&#160;<b>&#183;</b>&#32; <a href="/wiki/Erwin_Neher" title="Erwin Neher">Neher</a> / <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Sakmann</a> (1991)&#160;<b>&#183;</b>&#32; <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Fischer</a> / <a href="/wiki/Edwin_G._Krebs" title="Edwin G. Krebs">Krebs</a> (1992)&#160;<b>&#183;</b>&#32; <a href="/wiki/Richard_Roberts" title="Richard Roberts">Roberts</a> / <a href="/wiki/Phillip_Sharp" title="Phillip Sharp">Sharp</a> (1993)&#160;<b>&#183;</b>&#32; <a href="/wiki/Alfred_G._Gilman" title="Alfred G. Gilman">Gilman</a> / <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Rodbell</a> (1994)&#160;<b>&#183;</b>&#32; <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Lewis</a> / <a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Nüsslein-Volhard</a> / <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Wieschaus</a> (1995)&#160;<b>&#183;</b>&#32; <a href="/wiki/Peter_C._Doherty" title="Peter C. Doherty">Doherty</a> / <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Zinkernagel</a> (1996)&#160;<b>&#183;</b>&#32; <a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Prusiner</a> (1997)&#160;<b>&#183;</b>&#32; <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Furchgott</a> / <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Ignarro</a> / <a href="/wiki/Ferid_Murad" title="Ferid Murad">Murad</a> (1998)&#160;<b>&#183;</b>&#32; <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Blobel</a> (1999)&#160;<b>&#183;</b>&#32; <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Carlsson</a> / <a href="/wiki/Paul_Greengard" title="Paul Greengard">Greengard</a> / <a href="/wiki/Eric_Kandel" title="Eric Kandel">Kandel</a> (2000) </p> </div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">2001 – present</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><div> <p><a href="/wiki/Lee_Hartwell" title="Lee Hartwell">Hartwell</a> / <a href="/wiki/Tim_Hunt" title="Tim Hunt">Hunt</a> / <a class="mw-selflink selflink">Nurse</a> (2001)&#160;<b>&#183;</b>&#32; <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Brenner</a> / <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">Horvitz</a> / <a href="/wiki/John_Sulston" title="John Sulston">Sulston</a> (2002)&#160;<b>&#183;</b>&#32; <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Lauterbur</a> / <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Mansfield</a> (2003)&#160;<b>&#183;</b>&#32; <a href="/wiki/Richard_Axel" title="Richard Axel">Axel</a> / <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Buck</a> (2004)&#160;<b>&#183;</b>&#32; <a href="/wiki/Barry_Marshall" title="Barry Marshall">Marshall</a> / <a href="/wiki/Robin_Warren" title="Robin Warren">Warren</a> (2005)&#160;<b>&#183;</b>&#32; <a href="/wiki/Andrew_Fire" title="Andrew Fire">Fire</a> / <a href="/wiki/Craig_Mello" title="Craig Mello">Mello</a> (2006)&#160;<b>&#183;</b>&#32; <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Capecchi</a> / <a href="/wiki/Martin_Evans" title="Martin Evans">Evans</a> / <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Smithies</a> (2007)&#160;<b>&#183;</b>&#32; <a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Barré-Sinoussi</a> / <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Montagnier</a> / <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">zur Hausen</a> (2008)&#160;<b>&#183;</b>&#32; <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Blackburn</a> / <a href="/wiki/Carol_Greider" title="Carol Greider">Greider</a> / <a href="/wiki/Jack_Szostak" title="Jack Szostak">Szostak</a> (2009)&#160;<b>&#183;</b>&#32; <a href="/wiki/Robert_Edwards" title="Robert Edwards">Edwards</a> (2010)&#160;<b>&#183;</b>&#32; <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Beutler</a> / <a href="/wiki/Jules_Hoffmann" title="Jules Hoffmann">Hoffmann</a>; <a href="/wiki/Ralph_Steinman" title="Ralph Steinman">Steinman</a> (2011)&#160;<b>&#183;</b>&#32; <a href="/wiki/John_Gurdon" title="John Gurdon">Gurdon</a> / <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Yamanaka</a> (2012)&#160;<b>&#183;</b>&#32; <a href="/wiki/James_Rothman" title="James Rothman">Rothman</a> / <a href="/wiki/Randy_Schekman" title="Randy Schekman">Schekman</a> / <a href="/wiki/Thomas_C._S%C3%BCdhof" title="Thomas C. Südhof">Südhof</a> (2013)&#160;<b>&#183;</b>&#32; <a href="/wiki/John_O%27Keefe" title="John O&#39;Keefe">O'Keefe</a> / <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">M-B. Moser</a> / <a href="/wiki/Edvard_Moser" title="Edvard Moser">E. Moser</a> (2014)&#160;<b>&#183;</b>&#32; <a href="/wiki/William_C._Campbell_(scientist)" title="William C. Campbell (scientist)">Campbell</a> / <a href="/wiki/Satoshi_%C5%8Cmura" title="Satoshi Ōmura">Ōmura</a> / <a href="/wiki/Tu_Youyou" title="Tu Youyou">Youyou</a> (2015)&#160;<b>&#183;</b>&#32; <a href="/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi">Ohsumi</a> (2016)&#160;<b>&#183;</b>&#32; <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Hall</a> / <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Rosbash</a> / <a href="/wiki/Michael_W._Young" title="Michael W. Young">Young</a> (2017)&#160;<b>&#183;</b>&#32; <a href="/wiki/James_P._Allison" title="James P. Allison">Allison</a> / <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Honjo</a> (2018)&#160;<b>&#183;</b>&#32; <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Semenza</a> / <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Ratcliffe</a> / <a href="/wiki/William_Kaelin_Jr." title="William Kaelin Jr.">Kaelin Jr.</a> (2019)&#160;<b>&#183;</b>&#32; <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Atler</a> / <a href="/wiki/Michael_Houghton" title="Michael Houghton">Houghton</a> / <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Rice</a> (2020)&#160;<b>&#183;</b>&#32; <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Patapoutian</a> / <a href="/wiki/David_Julius" title="David Julius">Julius</a> (2021)&#160;<b>&#183;</b>&#32; <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Pääbo</a> (2022)&#160;<b>&#183;</b>&#32; <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Karikó</a> / <a href="/wiki/Drew_Weissman" title="Drew Weissman">Weissman</a> (2023)&#160;<b>&#183;</b>&#32; <a href="/wiki/Victor_Ambros" title="Victor Ambros">Ambros</a> / <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Ruvkun</a> (2024) </p> </div></div></td></tr></tbody></table></div> <p class="mw-empty-elt"> </p> <!-- NewPP limit report Parsed by mw‐web.eqiad.main‐6b6c9bdc8b‐4lcdd Cached time: 20241103183819 Cache expiry: 2592000 Reduced expiry: false Complications: [vary‐revision‐sha1] CPU time usage: 0.363 seconds Real time usage: 0.711 seconds Preprocessor visited node count: 2495/1000000 Post‐expand include size: 55853/2097152 bytes Template argument size: 2638/2097152 bytes Highest expansion depth: 16/100 Expensive parser function count: 1/500 Unstrip recursion depth: 1/20 Unstrip post‐expand size: 16490/5000000 bytes Lua time usage: 0.169/10.000 seconds Lua memory usage: 4642173/52428800 bytes Number of Wikibase entities loaded: 1/400 --> <!-- Transclusion expansion time report (%,ms,calls,template) 100.00% 449.375 1 -total 48.51% 218.009 1 Template:Infobox_scientist 46.65% 209.639 1 Template:Infobox_person 30.22% 135.823 1 Template:Reflist 18.61% 83.609 5 Template:Br_separated_entries 15.64% 70.298 1 Template:Birth_date_and_age 13.33% 59.880 1 Template:ISBN 12.58% 56.548 1 Template:Nobel_Prize_in_Physiology_or_Medicine 12.48% 56.090 1 Template:Cite_journal 11.83% 53.181 1 Template:Navbox --> <!-- Saved in parser cache with key simplewiki:pcache:idhash:291777-0!canonical and timestamp 20241103183819 and revision id 9574995. Rendering was triggered because: page-view --> </div><!--esi <esi:include src="/esitest-fa8a495983347898/content" /> --><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Retrieved from "<a dir="ltr" href="https://simple.wikipedia.org/w/index.php?title=Paul_Nurse&amp;oldid=9574995">https://simple.wikipedia.org/w/index.php?title=Paul_Nurse&amp;oldid=9574995</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Special:Categories" title="Special:Categories">Categories</a>: <ul><li><a href="/wiki/Category:Nobel_Prize_in_Physiology_or_Medicine_winners" title="Category:Nobel Prize in Physiology or Medicine winners">Nobel Prize in Physiology or Medicine winners</a></li><li><a href="/wiki/Category:1949_births" title="Category:1949 births">1949 births</a></li><li><a href="/wiki/Category:Living_people" title="Category:Living people">Living people</a></li><li><a href="/wiki/Category:English_biologists" title="Category:English biologists">English biologists</a></li><li><a href="/wiki/Category:English_geneticists" title="Category:English geneticists">English geneticists</a></li><li><a href="/wiki/Category:Knights_Bachelor" title="Category:Knights Bachelor">Knights Bachelor</a></li><li><a href="/wiki/Category:People_from_Norwich" title="Category:People from Norwich">People from Norwich</a></li><li><a href="/wiki/Category:Presidents_of_the_Royal_Society" title="Category:Presidents of the Royal Society">Presidents of the Royal Society</a></li><li><a href="/wiki/Category:British_Nobel_Prize_winners" title="Category:British Nobel Prize winners">British Nobel Prize winners</a></li><li><a href="/wiki/Category:Alumni_of_the_University_of_Birmingham" title="Category:Alumni of the University of Birmingham">Alumni of the University of Birmingham</a></li><li><a href="/wiki/Category:Alumni_of_the_University_of_East_Anglia" title="Category:Alumni of the University of East Anglia">Alumni of the University of East Anglia</a></li><li><a href="/wiki/Category:Rockefeller_University_faculty" title="Category:Rockefeller University faculty">Rockefeller University faculty</a></li></ul></div><div id="mw-hidden-catlinks" class="mw-hidden-catlinks mw-hidden-cats-hidden">Hidden categories: <ul><li><a href="/wiki/Category:Pages_using_the_JsonConfig_extension" title="Category:Pages using the JsonConfig extension">Pages using the JsonConfig extension</a></li><li><a href="/wiki/Category:Articles_with_hCards" title="Category:Articles with hCards">Articles with hCards</a></li></ul></div></div> </div> </main> </div> <div class="mw-footer-container"> <footer id="footer" class="mw-footer" > <ul id="footer-info"> <li id="footer-info-lastmod"> This page was last changed on 1 June 2024, at 04:40.</li> <li id="footer-info-copyright">Text is available under the <a rel="nofollow" class="external text" href="//creativecommons.org/licenses/by-sa/4.0/">Creative Commons Attribution-ShareAlike License</a> and the <a href="/wiki/Wikipedia:Text_of_the_GNU_Free_Documentation_License" title="Wikipedia:Text of the GNU Free Documentation License">GFDL</a>; additional terms may apply. See <a class="external text" href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Terms_of_Use">Terms of Use</a> for details.</li> </ul> <ul id="footer-places"> <li id="footer-places-privacy"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Privacy_policy">Privacy policy</a></li> <li id="footer-places-about"><a href="/wiki/Wikipedia:About">About Wikipedia</a></li> <li id="footer-places-disclaimers"><a href="/wiki/Wikipedia:General_disclaimer">Disclaimers</a></li> <li id="footer-places-wm-codeofconduct"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Universal_Code_of_Conduct">Code of Conduct</a></li> <li id="footer-places-developers"><a href="https://developer.wikimedia.org">Developers</a></li> <li id="footer-places-statslink"><a href="https://stats.wikimedia.org/#/simple.wikipedia.org">Statistics</a></li> <li id="footer-places-cookiestatement"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Cookie_statement">Cookie statement</a></li> <li id="footer-places-mobileview"><a href="//simple.m.wikipedia.org/w/index.php?title=Paul_Nurse&amp;mobileaction=toggle_view_mobile" class="noprint stopMobileRedirectToggle">Mobile view</a></li> </ul> <ul id="footer-icons" class="noprint"> <li id="footer-copyrightico"><a href="https://wikimediafoundation.org/" class="cdx-button cdx-button--fake-button cdx-button--size-large cdx-button--fake-button--enabled"><img src="/static/images/footer/wikimedia-button.svg" width="84" height="29" alt="Wikimedia Foundation" loading="lazy"></a></li> <li id="footer-poweredbyico"><a href="https://www.mediawiki.org/" class="cdx-button cdx-button--fake-button cdx-button--size-large cdx-button--fake-button--enabled"><img src="/w/resources/assets/poweredby_mediawiki.svg" alt="Powered by MediaWiki" width="88" height="31" loading="lazy"></a></li> </ul> </footer> </div> </div> </div> <div class="vector-settings" id="p-dock-bottom"> <ul></ul> </div><script>(RLQ=window.RLQ||[]).push(function(){mw.config.set({"wgHostname":"mw-web.codfw.main-6b7f745dd4-sgqbm","wgBackendResponseTime":147,"wgPageParseReport":{"limitreport":{"cputime":"0.363","walltime":"0.711","ppvisitednodes":{"value":2495,"limit":1000000},"postexpandincludesize":{"value":55853,"limit":2097152},"templateargumentsize":{"value":2638,"limit":2097152},"expansiondepth":{"value":16,"limit":100},"expensivefunctioncount":{"value":1,"limit":500},"unstrip-depth":{"value":1,"limit":20},"unstrip-size":{"value":16490,"limit":5000000},"entityaccesscount":{"value":1,"limit":400},"timingprofile":["100.00% 449.375 1 -total"," 48.51% 218.009 1 Template:Infobox_scientist"," 46.65% 209.639 1 Template:Infobox_person"," 30.22% 135.823 1 Template:Reflist"," 18.61% 83.609 5 Template:Br_separated_entries"," 15.64% 70.298 1 Template:Birth_date_and_age"," 13.33% 59.880 1 Template:ISBN"," 12.58% 56.548 1 Template:Nobel_Prize_in_Physiology_or_Medicine"," 12.48% 56.090 1 Template:Cite_journal"," 11.83% 53.181 1 Template:Navbox"]},"scribunto":{"limitreport-timeusage":{"value":"0.169","limit":"10.000"},"limitreport-memusage":{"value":4642173,"limit":52428800}},"cachereport":{"origin":"mw-web.eqiad.main-6b6c9bdc8b-4lcdd","timestamp":"20241103183819","ttl":2592000,"transientcontent":false}}});});</script> <script type="application/ld+json">{"@context":"https:\/\/schema.org","@type":"Article","name":"Paul Nurse","url":"https:\/\/simple.wikipedia.org\/wiki\/Paul_Nurse","sameAs":"http:\/\/www.wikidata.org\/entity\/Q219716","mainEntity":"http:\/\/www.wikidata.org\/entity\/Q219716","author":{"@type":"Organization","name":"Contributors to Wikimedia projects"},"publisher":{"@type":"Organization","name":"Wikimedia Foundation, Inc.","logo":{"@type":"ImageObject","url":"https:\/\/www.wikimedia.org\/static\/images\/wmf-hor-googpub.png"}},"datePublished":"2011-04-18T09:35:46Z","dateModified":"2024-06-01T04:40:18Z","image":"https:\/\/upload.wikimedia.org\/wikipedia\/commons\/a\/ac\/Paul_Nurse_portrait.jpg","headline":"Nobel prize winning British biochemist"}</script> </body> </html>

Pages: 1 2 3 4 5 6 7 8 9 10