CINXE.COM

<!DOCTYPE html><html lang="en-US"><head><title>Just a moment...</title><meta http-equiv="Content-Type" content="text/html; charset=UTF-8"><meta http-equiv="X-UA-Compatible" content="IE=Edge"><meta name="robots" content="noindex,nofollow"><meta name="viewport" content="width=device-width,initial-scale=1"><style>*{box-sizing:border-box;margin:0;padding:0}html{line-height:1.15;-webkit-text-size-adjust:100%;color:#313131;font-family:system-ui,-apple-system,BlinkMacSystemFont,Segoe UI,Roboto,Helvetica Neue,Arial,Noto Sans,sans-serif,Apple Color Emoji,Segoe UI Emoji,Segoe UI Symbol,Noto Color Emoji}body{display:flex;flex-direction:column;height:100vh;min-height:100vh}.main-content{margin:8rem auto;max-width:60rem;padding-left:1.5rem}@media (width <= 720px){.main-content{margin-top:4rem}}.h2{font-size:1.5rem;font-weight:500;line-height:2.25rem}@media (width <= 720px){.h2{font-size:1.25rem;line-height:1.5rem}}#challenge-error-text{background-image:url(data:image/svg+xml;base64,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);background-repeat:no-repeat;background-size:contain;padding-left:34px}@media (prefers-color-scheme:dark){body{background-color:#222;color:#d9d9d9}}</style><meta http-equiv="refresh" content="390"></head><body class="no-js"><div class="main-wrapper" role="main"><div class="main-content"><noscript><div class="h2"><span id="challenge-error-text">Enable JavaScript and cookies to continue</span></div></noscript></div></div><script>(function(){window._cf_chl_opt={cvId: '3',cZone: "onlinelibrary.wiley.com",cType: 'managed',cRay: '8e6f84d20a8781f8',cH: 'bJSPERd2So6KoxWfnm3PsXcM4DOfiQKMjVHAljpVz_s-1732347707-1.2.1.1-3tTgbfiZTCPDKbgnU3fwL0YksbI7Yw1bkiNxoOqhhwufme_R9IqvSZ5rk9aP_cLR',cUPMDTk: "\/doi\/full\/10.1002\/jmv.29029?__cf_chl_tk=AjBn.00ByCYrO57l8PJKQgjisLHdMZx51gO3NuROkbE-1732347707-1.0.1.1-YZ8TFBr1XMrnu_AiAZR91RjbQRn77Yn0frXyCcOLzEc",cFPWv: 'g',cITimeS: '1732347707',cTTimeMs: '1000',cMTimeMs: '390000',cTplC: 0,cTplV: 5,cTplB: 'cf',cK: "",fa: "\/doi\/full\/10.1002\/jmv.29029?__cf_chl_f_tk=AjBn.00ByCYrO57l8PJKQgjisLHdMZx51gO3NuROkbE-1732347707-1.0.1.1-YZ8TFBr1XMrnu_AiAZR91RjbQRn77Yn0frXyCcOLzEc",md: "raykiI_6QET9CF6tsJ01wofC7iut42wn.bzxtI.v5yE-1732347707-1.2.1.1-qXodHx_rtj7FPatV3IWH6sWmQpx8YA9svdbf.rM.3DGlM08j9aCRQ.WPERt1DTM46OabPUCejUSIrs7SysNhnxIfFf9zNDWFkZBpXKSTvj46fBCx4U9DVHJEW0hIznqPfgMSUFZw7CnjA3KRmd7ysLtyB261vZt5tSykQErDSMWBAAtcCp8Cofv12m.aqWv1zNUvdlKAPLT6OsUBGO4GjN5c2.ZZBl5XZSDCYnV_JPzxK_SEvJevefgCOyEbuJ1vkZQxovWNbzz_PKAleQ5ZKpxE9MXS11rhPd1i_ePFzlJocFpJZKuG6dA4jue4_vOzSgk3g66YyiwrNwSpb6Emsvw4ek92gsiG_SXgTSHSB78rpuzyr0KOLKelndHhGoDFtdk19X_7jIN2FwcIW0MAmA2iJDmDuS6qIKoJeG0ZA5bmNRGQV9taMkWEldTCZJGS3h9M.qjFyo_wUws2XdBUvmqbiGPwiQJ9G8nwsZB5nBRNIrNsyI.1H2kDpMCq7hzXlf.zixygQ.IdEpildzPaN_DBC8yt1swUPNEZsV.C5RuV6a1p9MYoYoboXgnwlhGzSlfWZHm3Wlx2m0qwRR7gu_4wO_804J6MtnBc1zC4o5KSdjABMV1mvTgIMsnBX4k2EP4OniCsp1pdjVxHSRZWFwWZRGKGRoQctWCnvg1xsUu0UyGAhhp1Vvj893Suvte3YHzzK8NdwPLbeRPRIM1ObpkQmNteRbLWxxmaxDA32BJ.eTNT.L.QZz90bgriwMZUSPm7cdEkjVndM9yZgw5Yi1LXBx5tT1cCTYtLJq9HmJ8upKQDEDCVGz.KzXrjbGbQOIOlRLSOdHAGbftqnsJ1KIZiVU3Zbc7TzaOyt.5XGjDSL.NlGoHYMzXSj1ljtgcuDfu8GkSiLR06M6XmNAh6EdqBa22j3MOmX7KRoJFobnCyectdfNSGyLW5iU5XmzjQNhcjQDDNMxvC_0dLAWLzETlIgtU6LXsrzBe33u4DAYX7jEhWqUC9Jg6fJ3jLlumLZ.X9606yaScg_mGJABM3eClEFz9..BpHAiJOloV2iSRHASOKaoHr7k4VlYpaMy9lyQ.TNeKJA5jaWWgwIqwrzHsR3E8APexppEdRYJuJdF5ZDnT_ud5eQQ93a.XL9kUfWVzkUOTeKxiGcK9DUtDnH8rwOspqh7IAm.cBfELalU.axrf4vQ7sfg389cqmeplH.KAm2eIGHSu8mkCnKmKkkbHUQpH0wzM1bbmSjTjQQC4hLcJqu68cwjMFV9oc9tWwAbgX75YItVSFzSt50w6lL.TZfdHVeaGH.UPOSlj6svuSrCFMx7kRxZ12heOf6UBbLN4pj0UizadDdutMq2.Ulm8Xs8ssS7neFDq1acMSu.PkZsl2XJTlV7On.PXRRhoQ4r9P5OCbXLbERNS.UCnwe_8aiGLu6SS_FQ.ykKazwpR.fIgu0Pl5kNssYcsbMv3_._tSLGmrl7PnfiqusYK2jyi2FTcD.UZiL6B5GpC3x6rXHB.PzQFtv_jXaEw0wr7XX0TteKqJ.TYswaaT37WDdPdBap5ykdnuZOrb_WJT_.Bj6FiSfOAIzimOmJE_SR_bavkcN9MFzXieWctpd_f60Z1NG3COw.W5C7_GXiA45_3W6MIhBy4JHLOcl8sEl_W2JFlCxvhIddfDEE2UP9s8NXtSbJpRSL00fkaNR7BKIy_UdnGBVmbuEzIzccr02lcjpyItX5P_gkcZJY_zWlciBM_6FLHzncOZfiDe6yp1Hi2Lsv1g3xNDnyGy32zR_hyGqWcY4GZjcfoqx2QO7UMlzs2yZa1ZigWOYJUNC92ra4weTKYuc.SNwdwY28Owr7AQQCFDQP_i2iu8rMniuxIFUWqTurm3r1CCN6oeNNzOX0fKlELDwRHND666PzBqhTAISpHdhVqMkBltnhvkU9_b4c3fIrpGQLsqFXsQjhO_Qiu37PFMLWfU2nZ.4kPmvGsTixHrcaVcRU2jp.1Ojzil4otKUtXcpTKdMka.lf2iR.GXYWOwH_s9uTICGJcKYPBRgmn1H4AeApN4V4nNyXbeq8qjHYazK81j4IKwP9Pj1ZaGd3ziJqY50NVP.mvpa27Fxk7X_Ykh5h6hDmPs7rZc.hv60CV0VACxQ8lBpOz__ZjnzlqMS24XYt64akhHlyJJL_WafdcgHOoxCLtlvnCivG_VxL6Kvk8pKzhoQPSYZsmD4CSVDGBD2Y1VvpV4d2CDqPJ.vZa7pnIIDGotDVf6WRli2Jn7QUtufCTpDtQLuz1EjXWNswMLgd4KzhFqNMwIJNCAbntcVvWEMj1fBd4lJ9xR5r4JhBq0w9SdQnT0d9PDjKMqX1tJDJu1yNZF8kcdapjxjpjjf3A0s0fuqO.bYR7BRZQsWHx_EtqVurbZJ9POxlWVAV03KuW.6m4iNAhNwX3WtH3jHfmk7Yv3XPRDyvrhZ704NCSi7PW1a4IH6DmAv.uD4QKjRzvau5Sqwc9EpJPb4nT44z8PmxGzTJ0i1g.pxzLvgUdzuC7QJATo_5EQap1d3Y3MzqVgoB1ie_rCqOO_qSy4Cc43I2Q6.ktPXwrS4.oJO2eOc0jis6LhVSXluUZIiOFdGr4pABfeuMtS4ornPF6qCdxJ8HjDzcBGmrNDCPwzFivR.k4KpIAN95a4OD2FFbMC9jocN1VEO1dWPi8mvgsXgc9wIDyBjggdLty7KrVp7lNn6hln5_UiuLv3ISGIYKFvseUfY0T.QiJ0UE7cpGT5n3WWtLf.8T3beufyQ7ZPrBdrUei_OjaGtZJGNh6dulpsGzbVQBMY7PJR_BoENOKoZ_Lhf52TZ4svWh3pZnEW1vHHjyhRP71E_pRsEwIgw16mh8ijV5MNDOKVt6xbvHQpxWu18Td6O6OGFs7YnogTTA3JUf3VOITcwEUTgoATZSxbEZd6InAfkUhFOrMRqg4C.4r0a98VFmm3AA25aJtoAJLcwh2aZZOhrR4",mdrd: "82NwHKk3T8r41K1yabPsnV6UNUZCadE.rf8KFkva2.M-1732347707-1.2.1.1-3g76Mpf0svy5GTmLkQSj5IDOS4miJOVkQ0CI6KLGDZWlaCqmGR049mWTL2syJ9wui.GaAmbLZfUTn4Kri6ujY9iBKn5DUM3I3oVZ9ItWmFcOKbIxLf_OiWuY4MUMAzJor5eonh9CQcXN9Sw3Lqn8tcBebeV7raUtTZWz03R2vYUaiIdE9cUrI7Yp38CZkiX3evRp9qIobNyKMBJA6itlTbMZDL_KN7xOjH4z6LTCaHtG7BnQdGKGfQTFSsl_Y2d6O3S0YfkcUHGx.r1TlHnS2iGxyeKbbs7JKPadqOh4kLaa2Qad_nJYIH1.CZCn5q2rVai8soj5r1nAQfp_QHoxZLvDHHQHptqMk9c1yGtOOFB8.abWaLslzUFTgsJUJBi2zOjPwHJF66Izavnh0GlDrq_rwAYG3k2bk6vWd_98vPdGtxDsUmtr9ZPDEdpkVHf2T1IO2ZA3hSBOChOdI4SXqVJM.040lOG8T2BKabYqEvImrJNW1WEzRM1qCD56C_C9ShjxpmF831AxXQqpObfUlu_rPSO94KjERMYw11nBcUU_0NJ8gO8Bhiqn09xyjmmNSBHfiOfi1MC4FIbGUwVC252uZ.mcP2Aejmw2vmwVNmNS20F9lCcqYNUB7n_akCHdnGOqUvpoiG1Nsu1vwF0v9JJPUo6MJsKLbieB8FWoPixr6QuqP5E6NNZfucKgaQKyjHC.STAnNazSOhdsh_lNhR.v7VdnO9.x88dB1tEJMzenOq12BxV5Y5e2j6avvdz0Ag_pvQeapHFMOfmMEG2DqydN7kr.OUqwwohot6r20My7DYxUB2fvLmymILEwUN0wSCKqUMM8U__neJ5SoVBfVv1f.JosVMiZc.NV7WAF2KHSMRPJc9uZox6U_XGu4qxEk5cwZD9ehZWeoDfDK8cqp6sC6sDWF0wm58fmy9NBxSawdmc0CsC5OqIEvGvWpgH1frMEQ2G2AlRQQFMRLlKA4fyhP0iwozRKGcKU9Dp8k317AhMbdeGvYO01g.cEX_ZrBIZBjYMbKHEXSCnmtIsLmAeiERfJNOxqjXKgEBp9KvSJeP7wAlpH3QEoTvXjeG8eNrK5tNEPUUs3mHPAn6ME6.LRM1Q0.kD0Ftxhj8xJEd.krSyp_.qCQ4Ptt4c2cxEHHNmWfJaVJI1XM8BFM.l4np_AjVjuyzyVP3t2j73ghJnYPdw_up7hifRt0CkBh427sJbmEFvC7yGvHRmet6locKUCMmcectbbXR_c6txM5vnQeQCMnLgI1NCAD9567L1hVkuzkhuD9MbDG20ECuMN2nTmtjDEik8XhLh.aGHNmdwZsLm36c4zNq8fJgSzWr3u90Qj09fNy4UkCG3UVBBG0Z8dJBDnxJtSmAyBhUJ85GjcMFbWVRaKZexP6lz7wDFmYCYtVOr4a0.SA1Ve_nMZvaMcfH0Q3Fvm59qN32twsbhuR4hoe7zHdx4JbitD4K.dyLfDYy2dU9G362gXL1CzyFEVR7I5qCtBy5KasylY2nEp9hxlQwAkGIIiLAbD3gB0VoQCLgyodu.GqxbUAQ_ia294CbHYo2UzFgR2J06_zh62Chyq1Dw2MIj0.hLADn0tgDmBKD5T2TJYI96U1amOFBhQDUAsykKuRU5sG_72hhNH0DCbQPWBqa8eC2vnamwsdXncH5XSMhu5nt2Rw6iHULr3q.YfEIdmOLJShYQFqbAJsUs0HhyRn5hZ0hDS9dypFXD.jM6YntnhFF98UXJJzmT1Xelgh_c7r93g9Z3rRNcrhJDMMaI_FZrAc232I9_tzZGa.ALSZ74cQSUL4J3r2NLdCL2sBHphK0oVpZnmL3r0XsaswliDojWw4opuUuBiBKfv1vvymWv6T8TTaKaECibY_fVQ5tNwpcqj0sCbUc3s_hsY9kfhDYn0BydqjBA81uV5uxI6dARahLDZjjwsJ7hKRcQAbs8Cm8yOgbzgs9UdXGOhMlyq2wjvxFNZuh_bjT8GBr1MMkv9nbyAw8KjRw"};var cpo = document.createElement('script');cpo.src = '/cdn-cgi/challenge-platform/h/g/orchestrate/chl_page/v1?ray=8e6f84d20a8781f8';window._cf_chl_opt.cOgUHash = location.hash === '' && location.href.indexOf('#') !== -1 ? '#' : location.hash;window._cf_chl_opt.cOgUQuery = location.search === '' && location.href.slice(0, location.href.length - window._cf_chl_opt.cOgUHash.length).indexOf('?') !== -1 ? '?' : location.search;if (window.history && window.history.replaceState) {var ogU = location.pathname + window._cf_chl_opt.cOgUQuery + window._cf_chl_opt.cOgUHash;history.replaceState(null, null, "\/doi\/full\/10.1002\/jmv.29029?__cf_chl_rt_tk=AjBn.00ByCYrO57l8PJKQgjisLHdMZx51gO3NuROkbE-1732347707-1.0.1.1-YZ8TFBr1XMrnu_AiAZR91RjbQRn77Yn0frXyCcOLzEc" + window._cf_chl_opt.cOgUHash);cpo.onload = function() {history.replaceState(null, null, ogU);}}document.getElementsByTagName('head')[0].appendChild(cpo);}());</script></body></html>