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href="#Physiology"> <div class="vector-toc-text"> 3 Physiology </div> </a> <button aria-controls="toc-Physiology-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> Toggle Physiology subsection </button> <ul id="toc-Physiology-sublist" class="vector-toc-list"> <li id="toc-Symbiotic_relation" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Symbiotic_relation"> <div class="vector-toc-text"> 3.1 Symbiotic relation </div> </a> <ul id="toc-Symbiotic_relation-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Ecology" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Ecology"> <div class="vector-toc-text"> 3.2 Ecology </div> </a> <ul id="toc-Ecology-sublist" class="vector-toc-list"> <li id="toc-Miniature_ecosystem_and_holobiont_theory" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Miniature_ecosystem_and_holobiont_theory"> <div class="vector-toc-text"> 3.2.1 Miniature ecosystem and holobiont theory </div> </a> <ul id="toc-Miniature_ecosystem_and_holobiont_theory-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Lichenicolous_fungi" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Lichenicolous_fungi"> <div class="vector-toc-text"> 3.2.2 Lichenicolous fungi </div> </a> <ul id="toc-Lichenicolous_fungi-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Reaction_to_water" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Reaction_to_water"> <div class="vector-toc-text"> 3.3 Reaction to water </div> </a> <ul id="toc-Reaction_to_water-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Metabolites,_metabolite_structures_and_bioactivity" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Metabolites,_metabolite_structures_and_bioactivity"> <div class="vector-toc-text"> 3.4 Metabolites, metabolite structures and bioactivity </div> </a> <ul id="toc-Metabolites,_metabolite_structures_and_bioactivity-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Growth_rate" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Growth_rate"> <div class="vector-toc-text"> 3.5 Growth rate </div> </a> <ul id="toc-Growth_rate-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Life_span" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Life_span"> <div class="vector-toc-text"> 3.6 Life span </div> </a> <ul id="toc-Life_span-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Response_to_environmental_stress" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Response_to_environmental_stress"> <div class="vector-toc-text"> 3.7 Response to environmental stress </div> </a> <ul id="toc-Response_to_environmental_stress-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Reproduction_and_dispersal" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#Reproduction_and_dispersal"> <div class="vector-toc-text"> 4 Reproduction and dispersal </div> </a> <button aria-controls="toc-Reproduction_and_dispersal-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> Toggle Reproduction and dispersal subsection </button> <ul id="toc-Reproduction_and_dispersal-sublist" class="vector-toc-list"> <li id="toc-Vegetative_reproduction" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Vegetative_reproduction"> <div class="vector-toc-text"> 4.1 Vegetative reproduction </div> </a> <ul id="toc-Vegetative_reproduction-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Sexual_reproduction" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Sexual_reproduction"> <div class="vector-toc-text"> 4.2 Sexual reproduction </div> </a> <ul id="toc-Sexual_reproduction-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Taxonomy_and_classification" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#Taxonomy_and_classification"> <div class="vector-toc-text"> 5 Taxonomy and classification </div> </a> <button aria-controls="toc-Taxonomy_and_classification-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> Toggle Taxonomy and classification subsection </button> <ul id="toc-Taxonomy_and_classification-sublist" class="vector-toc-list"> <li id="toc-Fungi" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Fungi"> <div class="vector-toc-text"> 5.1 Fungi </div> </a> <ul id="toc-Fungi-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Photobionts" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Photobionts"> <div class="vector-toc-text"> 5.2 Photobionts </div> </a> <ul id="toc-Photobionts-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Controversy_over_classification_method_and_species_names" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Controversy_over_classification_method_and_species_names"> <div class="vector-toc-text"> 5.3 Controversy over classification method and species names </div> </a> <ul id="toc-Controversy_over_classification_method_and_species_names-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Diversity" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Diversity"> <div class="vector-toc-text"> 5.4 Diversity </div> </a> <ul id="toc-Diversity-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Identification_methods" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Identification_methods"> <div class="vector-toc-text"> 5.5 Identification methods </div> </a> <ul id="toc-Identification_methods-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Evolution_and_paleontology" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Evolution_and_paleontology"> <div class="vector-toc-text"> 5.6 Evolution and paleontology </div> </a> <ul id="toc-Evolution_and_paleontology-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Ecology_and_interactions_with_environment" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#Ecology_and_interactions_with_environment"> <div class="vector-toc-text"> 6 Ecology and interactions with environment </div> </a> <button aria-controls="toc-Ecology_and_interactions_with_environment-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> Toggle Ecology and interactions with environment subsection </button> <ul id="toc-Ecology_and_interactions_with_environment-sublist" class="vector-toc-list"> <li id="toc-Substrates_and_habitats" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Substrates_and_habitats"> <div class="vector-toc-text"> 6.1 Substrates and habitats </div> </a> <ul id="toc-Substrates_and_habitats-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Lichens_and_soils" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Lichens_and_soils"> <div class="vector-toc-text"> 6.2 Lichens and soils </div> </a> <ul id="toc-Lichens_and_soils-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Ecological_interactions" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Ecological_interactions"> <div class="vector-toc-text"> 6.3 Ecological interactions </div> </a> <ul id="toc-Ecological_interactions-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Effects_of_air_pollution" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Effects_of_air_pollution"> <div class="vector-toc-text"> 6.4 Effects of air pollution </div> </a> <ul id="toc-Effects_of_air_pollution-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Human_use" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#Human_use"> <div class="vector-toc-text"> 7 Human use </div> </a> <button aria-controls="toc-Human_use-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> Toggle Human use subsection </button> <ul id="toc-Human_use-sublist" class="vector-toc-list"> <li id="toc-Food" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Food"> <div class="vector-toc-text"> 7.1 Food </div> </a> <ul id="toc-Food-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Lichenometry" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Lichenometry"> <div class="vector-toc-text"> 7.2 Lichenometry </div> </a> <ul id="toc-Lichenometry-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Biodegradation" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Biodegradation"> <div class="vector-toc-text"> 7.3 Biodegradation </div> </a> <ul id="toc-Biodegradation-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Dyes" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Dyes"> <div class="vector-toc-text"> 7.4 Dyes </div> </a> <ul id="toc-Dyes-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Traditional_medicine_and_research" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Traditional_medicine_and_research"> <div class="vector-toc-text"> 7.5 Traditional medicine and research </div> </a> <ul id="toc-Traditional_medicine_and_research-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Aesthetic_appeal" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Aesthetic_appeal"> <div class="vector-toc-text"> 7.6 Aesthetic appeal </div> </a> <ul id="toc-Aesthetic_appeal-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-In_literature" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#In_literature"> <div class="vector-toc-text"> 7.7 In literature </div> </a> <ul id="toc-In_literature-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-History" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#History"> <div class="vector-toc-text"> 8 History </div> </a> <ul id="toc-History-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-See_also" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#See_also"> <div class="vector-toc-text"> 9 See also </div> </a> <ul id="toc-See_also-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Notes" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#Notes"> <div class="vector-toc-text"> 10 Notes </div> </a> <ul id="toc-Notes-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> 11 References </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" > Toggle the table of contents </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading">Lichen</h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 95 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-95" aria-hidden="true" > 95 languages </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-af mw-list-item"><a href="https://af.wikipedia.org/wiki/Korsmos" title="Korsmos – Afrikaans" lang="af" hreflang="af" data-title="Korsmos" data-language-autonym="Afrikaans" data-language-local-name="Afrikaans" class="interlanguage-link-target">Afrikaans</a></li><li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%A3%D8%B4%D9%86%D9%8A%D8%A7%D8%AA" title="أشنيات – Arabic" lang="ar" hreflang="ar" data-title="أشنيات" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target">العربية</a></li><li class="interlanguage-link interwiki-ast mw-list-item"><a href="https://ast.wikipedia.org/wiki/Lique" title="Lique – Asturian" lang="ast" hreflang="ast" data-title="Lique" data-language-autonym="Asturianu" data-language-local-name="Asturian" class="interlanguage-link-target">Asturianu</a></li><li class="interlanguage-link interwiki-atj mw-list-item"><a href="https://atj.wikipedia.org/wiki/Mirapakon" title="Mirapakon – Atikamekw" lang="atj" hreflang="atj" data-title="Mirapakon" data-language-autonym="Atikamekw" data-language-local-name="Atikamekw" class="interlanguage-link-target">Atikamekw</a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/%C5%9Eiby%C9%99l%C9%99r" title="Şibyələr – Azerbaijani" lang="az" hreflang="az" data-title="Şibyələr" data-language-autonym="Azərbaycanca" data-language-local-name="Azerbaijani" class="interlanguage-link-target">Azərbaycanca</a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D8%AF%D8%A7%D8%B4%E2%80%8C%D8%AC%D8%A7%D8%A8%D8%A7%D8%BA%DB%8C" title="داش‌جاباغی – South Azerbaijani" lang="azb" hreflang="azb" data-title="داش‌جاباغی" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target">تۆرکجه</a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%B2%E0%A6%BE%E0%A6%87%E0%A6%95%E0%A7%87%E0%A6%A8" title="লাইকেন – Bangla" lang="bn" hreflang="bn" data-title="লাইকেন" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target">বাংলা</a></li><li class="interlanguage-link interwiki-bjn mw-list-item"><a href="https://bjn.wikipedia.org/wiki/Lumut_karak" title="Lumut karak – Banjar" lang="bjn" hreflang="bjn" data-title="Lumut karak" data-language-autonym="Banjar" data-language-local-name="Banjar" class="interlanguage-link-target">Banjar</a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%9B%D1%96%D1%88%D0%B0%D0%B9%D0%BD%D1%96%D0%BA%D1%96" title="Лішайнікі – Belarusian" lang="be" hreflang="be" data-title="Лішайнікі" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target">Беларуская</a></li><li class="interlanguage-link interwiki-be-x-old mw-list-item"><a href="https://be-tarask.wikipedia.org/wiki/%D0%9B%D1%96%D1%88%D0%B0%D0%B9%D0%BD%D1%96%D0%BA%D1%96" title="Лішайнікі – Belarusian (Taraškievica orthography)" lang="be-tarask" hreflang="be-tarask" data-title="Лішайнікі" data-language-autonym="Беларуская (тарашкевіца)" data-language-local-name="Belarusian (Taraškievica orthography)" class="interlanguage-link-target">Беларуская (тарашкевіца)</a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%9B%D0%B8%D1%88%D0%B5%D0%B8" title="Лишеи – Bulgarian" lang="bg" hreflang="bg" data-title="Лишеи" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target">Български</a></li><li class="interlanguage-link interwiki-bs mw-list-item"><a href="https://bs.wikipedia.org/wiki/Li%C5%A1aj" title="Lišaj – Bosnian" lang="bs" hreflang="bs" data-title="Lišaj" data-language-autonym="Bosanski" data-language-local-name="Bosnian" class="interlanguage-link-target">Bosanski</a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Liquen" title="Liquen – Catalan" lang="ca" hreflang="ca" data-title="Liquen" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target">Català</a></li><li class="interlanguage-link interwiki-cs badge-Q17437796 badge-featuredarticle mw-list-item" title="featured article badge"><a href="https://cs.wikipedia.org/wiki/Li%C5%A1ejn%C3%ADk" title="Lišejník – Czech" lang="cs" hreflang="cs" data-title="Lišejník" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target">Čeština</a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Lav_(symbiotisk_organisme)" title="Lav (symbiotisk organisme) – Danish" lang="da" hreflang="da" data-title="Lav (symbiotisk organisme)" data-language-autonym="Dansk" data-language-local-name="Danish" class="interlanguage-link-target">Dansk</a></li><li class="interlanguage-link interwiki-ary mw-list-item"><a href="https://ary.wikipedia.org/wiki/%D8%AE%D8%B2" title="خز – Moroccan Arabic" lang="ary" hreflang="ary" data-title="خز" data-language-autonym="الدارجة" data-language-local-name="Moroccan Arabic" class="interlanguage-link-target">الدارجة</a></li><li class="interlanguage-link interwiki-de badge-Q17437796 badge-featuredarticle mw-list-item" title="featured article badge"><a href="https://de.wikipedia.org/wiki/Flechte" title="Flechte – German" lang="de" hreflang="de" data-title="Flechte" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target">Deutsch</a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Samblikud" title="Samblikud – Estonian" lang="et" hreflang="et" data-title="Samblikud" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target">Eesti</a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%9B%CE%B5%CE%B9%CF%87%CE%AE%CE%BD%CE%B1" title="Λειχήνα – Greek" lang="el" hreflang="el" data-title="Λειχήνα" data-language-autonym="Ελληνικά" data-language-local-name="Greek" class="interlanguage-link-target">Ελληνικά</a></li><li class="interlanguage-link interwiki-es badge-Q17437798 badge-goodarticle mw-list-item" title="good article badge"><a href="https://es.wikipedia.org/wiki/Liquen" title="Liquen – Spanish" lang="es" hreflang="es" data-title="Liquen" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target">Español</a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Likeno" title="Likeno – Esperanto" lang="eo" hreflang="eo" data-title="Likeno" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target">Esperanto</a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Liken" title="Liken – Basque" lang="eu" hreflang="eu" data-title="Liken" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target">Euskara</a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%DA%AF%D9%84%D8%B3%D9%86%DA%AF" title="گلسنگ – Persian" lang="fa" hreflang="fa" data-title="گلسنگ" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target">فارسی</a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Lichen" title="Lichen – French" lang="fr" hreflang="fr" data-title="Lichen" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target">Français</a></li><li class="interlanguage-link interwiki-fy mw-list-item"><a href="https://fy.wikipedia.org/wiki/Koarstmoas" title="Koarstmoas – Western Frisian" lang="fy" hreflang="fy" data-title="Koarstmoas" data-language-autonym="Frysk" data-language-local-name="Western Frisian" class="interlanguage-link-target">Frysk</a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/L%C3%A9icean" title="Léicean – Irish" lang="ga" hreflang="ga" data-title="Léicean" data-language-autonym="Gaeilge" data-language-local-name="Irish" class="interlanguage-link-target">Gaeilge</a></li><li class="interlanguage-link interwiki-gd mw-list-item"><a href="https://gd.wikipedia.org/wiki/Crotal" title="Crotal – Scottish Gaelic" lang="gd" hreflang="gd" data-title="Crotal" data-language-autonym="Gàidhlig" data-language-local-name="Scottish Gaelic" class="interlanguage-link-target">Gàidhlig</a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Lique" title="Lique – Galician" lang="gl" hreflang="gl" data-title="Lique" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target">Galego</a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%A7%80%EC%9D%98%EB%A5%98" title="지의류 – Korean" lang="ko" hreflang="ko" data-title="지의류" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target">한국어</a></li><li class="interlanguage-link interwiki-hy mw-list-item"><a href="https://hy.wikipedia.org/wiki/%D5%94%D5%A1%D6%80%D5%A1%D6%84%D5%B8%D5%BD%D5%A5%D6%80" title="Քարաքոսեր – Armenian" lang="hy" hreflang="hy" data-title="Քարաքոսեր" data-language-autonym="Հայերեն" data-language-local-name="Armenian" class="interlanguage-link-target">Հայերեն</a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%B2%E0%A4%BE%E0%A4%87%E0%A4%95%E0%A5%87%E0%A4%A8" title="लाइकेन – Hindi" lang="hi" hreflang="hi" data-title="लाइकेन" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target">हिन्दी</a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Li%C5%A1ajevi" title="Lišajevi – Croatian" lang="hr" hreflang="hr" data-title="Lišajevi" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target">Hrvatski</a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Likeno" title="Likeno – Ido" lang="io" hreflang="io" data-title="Likeno" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target">Ido</a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Lumut_kerak" title="Lumut kerak – Indonesian" lang="id" hreflang="id" data-title="Lumut kerak" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target">Bahasa Indonesia</a></li><li class="interlanguage-link interwiki-os mw-list-item"><a href="https://os.wikipedia.org/wiki/%D0%9B%D1%8B%D0%B2%D0%B7" title="Лывз – Ossetic" lang="os" hreflang="os" data-title="Лывз" data-language-autonym="Ирон" data-language-local-name="Ossetic" class="interlanguage-link-target">Ирон</a></li><li class="interlanguage-link interwiki-is mw-list-item"><a href="https://is.wikipedia.org/wiki/Fl%C3%A9ttur" title="Fléttur – Icelandic" lang="is" hreflang="is" data-title="Fléttur" data-language-autonym="Íslenska" data-language-local-name="Icelandic" class="interlanguage-link-target">Íslenska</a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Lichene" title="Lichene – Italian" lang="it" hreflang="it" data-title="Lichene" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target">Italiano</a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%97%D7%96%D7%96%D7%99%D7%AA" title="חזזית – Hebrew" lang="he" hreflang="he" data-title="חזזית" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target">עברית</a></li><li class="interlanguage-link interwiki-jv mw-list-item"><a href="https://jv.wikipedia.org/wiki/Lumut_kerak" title="Lumut kerak – Javanese" lang="jv" hreflang="jv" data-title="Lumut kerak" data-language-autonym="Jawa" data-language-local-name="Javanese" class="interlanguage-link-target">Jawa</a></li><li class="interlanguage-link interwiki-kn mw-list-item"><a href="https://kn.wikipedia.org/wiki/%E0%B2%95%E0%B2%B2%E0%B3%8D%E0%B2%B2%E0%B3%81%E0%B2%B9%E0%B3%82%E0%B2%B5%E0%B3%81_(%E0%B2%B2%E0%B3%88%E0%B2%95%E0%B2%A8%E0%B3%8D%E2%80%8C%E0%B2%97%E0%B2%B3%E0%B3%81)" title="ಕಲ್ಲುಹೂವು (ಲೈಕನ್‌ಗಳು) – Kannada" lang="kn" hreflang="kn" data-title="ಕಲ್ಲುಹೂವು (ಲೈಕನ್‌ಗಳು)" data-language-autonym="ಕನ್ನಡ" data-language-local-name="Kannada" class="interlanguage-link-target">ಕನ್ನಡ</a></li><li class="interlanguage-link interwiki-ka mw-list-item"><a href="https://ka.wikipedia.org/wiki/%E1%83%9A%E1%83%98%E1%83%A5%E1%83%94%E1%83%9C%E1%83%94%E1%83%91%E1%83%98" title="ლიქენები – Georgian" lang="ka" hreflang="ka" data-title="ლიქენები" data-language-autonym="ქართული" data-language-local-name="Georgian" class="interlanguage-link-target">ქართული</a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D2%9A%D1%8B%D0%BD%D0%B0%D0%BB%D0%B0%D1%80" title="Қыналар – Kazakh" lang="kk" hreflang="kk" data-title="Қыналар" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target">Қазақша</a></li><li class="interlanguage-link interwiki-kw mw-list-item"><a href="https://kw.wikipedia.org/wiki/Kenn_an_kerrek" title="Kenn an kerrek – Cornish" lang="kw" hreflang="kw" data-title="Kenn an kerrek" data-language-autonym="Kernowek" data-language-local-name="Cornish" class="interlanguage-link-target">Kernowek</a></li><li class="interlanguage-link interwiki-ht mw-list-item"><a href="https://ht.wikipedia.org/wiki/Liken" title="Liken – Haitian Creole" lang="ht" hreflang="ht" data-title="Liken" data-language-autonym="Kreyòl ayisyen" data-language-local-name="Haitian Creole" class="interlanguage-link-target">Kreyòl ayisyen</a></li><li class="interlanguage-link interwiki-ky mw-list-item"><a href="https://ky.wikipedia.org/wiki/%D0%AD%D2%A3%D0%B8%D0%BB%D1%87%D0%B5%D0%BA%D1%82%D0%B5%D1%80" title="Эңилчектер – Kyrgyz" lang="ky" hreflang="ky" data-title="Эңилчектер" data-language-autonym="Кыргызча" data-language-local-name="Kyrgyz" class="interlanguage-link-target">Кыргызча</a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Lichen" title="Lichen – Latin" lang="la" hreflang="la" data-title="Lichen" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target">Latina</a></li><li class="interlanguage-link interwiki-lv mw-list-item"><a href="https://lv.wikipedia.org/wiki/%C4%B6%C4%93rpji" title="Ķērpji – Latvian" lang="lv" hreflang="lv" data-title="Ķērpji" data-language-autonym="Latviešu" data-language-local-name="Latvian" class="interlanguage-link-target">Latviešu</a></li><li class="interlanguage-link interwiki-lb mw-list-item"><a href="https://lb.wikipedia.org/wiki/Flecht" title="Flecht – Luxembourgish" lang="lb" hreflang="lb" data-title="Flecht" data-language-autonym="Lëtzebuergesch" data-language-local-name="Luxembourgish" class="interlanguage-link-target">Lëtzebuergesch</a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Kerp%C4%97s" title="Kerpės – Lithuanian" lang="lt" hreflang="lt" data-title="Kerpės" data-language-autonym="Lietuvių" data-language-local-name="Lithuanian" class="interlanguage-link-target">Lietuvių</a></li><li class="interlanguage-link interwiki-li mw-list-item"><a href="https://li.wikipedia.org/wiki/Koorsmos" title="Koorsmos – Limburgish" lang="li" hreflang="li" data-title="Koorsmos" data-language-autonym="Limburgs" data-language-local-name="Limburgish" class="interlanguage-link-target">Limburgs</a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Zuzm%C3%B3k" title="Zuzmók – Hungarian" lang="hu" hreflang="hu" data-title="Zuzmók" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target">Magyar</a></li><li class="interlanguage-link interwiki-mk mw-list-item"><a href="https://mk.wikipedia.org/wiki/%D0%9B%D0%B8%D1%88%D0%B0%D0%B8" title="Лишаи – Macedonian" lang="mk" hreflang="mk" data-title="Лишаи" data-language-autonym="Македонски" data-language-local-name="Macedonian" class="interlanguage-link-target">Македонски</a></li><li class="interlanguage-link interwiki-mg mw-list-item"><a href="https://mg.wikipedia.org/wiki/Volombato" title="Volombato – Malagasy" lang="mg" hreflang="mg" data-title="Volombato" data-language-autonym="Malagasy" data-language-local-name="Malagasy" class="interlanguage-link-target">Malagasy</a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%95%E0%B5%BD%E0%B4%AA%E0%B5%8D%E0%B4%AA%E0%B4%BE%E0%B4%AF%E0%B5%BD" title="കൽപ്പായൽ – Malayalam" lang="ml" hreflang="ml" data-title="കൽപ്പായൽ" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target">മലയാളം</a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Kulampair" title="Kulampair – Malay" lang="ms" hreflang="ms" data-title="Kulampair" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target">Bahasa Melayu</a></li><li class="interlanguage-link interwiki-my mw-list-item"><a href="https://my.wikipedia.org/wiki/%E1%80%9E%E1%80%85%E1%80%BA%E1%80%80%E1%80%95%E1%80%BA%E1%80%99%E1%80%BE%E1%80%B1%E1%80%AC%E1%80%BA" title="သစ်ကပ်မှော် – Burmese" lang="my" hreflang="my" data-title="သစ်ကပ်မှော်" data-language-autonym="မြန်မာဘာသာ" data-language-local-name="Burmese" class="interlanguage-link-target">မြန်မာဘာသာ</a></li><li class="interlanguage-link interwiki-pcm mw-list-item"><a href="https://pcm.wikipedia.org/wiki/Laiken" title="Laiken – Nigerian Pidgin" lang="pcm" hreflang="pcm" data-title="Laiken" data-language-autonym="Naijá" data-language-local-name="Nigerian Pidgin" class="interlanguage-link-target">Naijá</a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Korstmos" title="Korstmos – Dutch" lang="nl" hreflang="nl" data-title="Korstmos" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target">Nederlands</a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E5%9C%B0%E8%A1%A3%E9%A1%9E" title="地衣類 – Japanese" lang="ja" hreflang="ja" data-title="地衣類" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target">日本語</a></li><li class="interlanguage-link interwiki-frr mw-list-item"><a href="https://frr.wikipedia.org/wiki/Hiasm%C3%B6%C3%B6sk" title="Hiasmöösk – Northern Frisian" lang="frr" hreflang="frr" data-title="Hiasmöösk" data-language-autonym="Nordfriisk" data-language-local-name="Northern Frisian" class="interlanguage-link-target">Nordfriisk</a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Lav" title="Lav – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Lav" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target">Norsk bokmål</a></li><li class="interlanguage-link interwiki-nn mw-list-item"><a href="https://nn.wikipedia.org/wiki/Lav" title="Lav – Norwegian Nynorsk" lang="nn" hreflang="nn" data-title="Lav" data-language-autonym="Norsk nynorsk" data-language-local-name="Norwegian Nynorsk" class="interlanguage-link-target">Norsk nynorsk</a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Liqu%C3%A8n" title="Liquèn – Occitan" lang="oc" hreflang="oc" data-title="Liquèn" data-language-autonym="Occitan" data-language-local-name="Occitan" class="interlanguage-link-target">Occitan</a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/Lishayniklar" title="Lishayniklar – Uzbek" lang="uz" hreflang="uz" data-title="Lishayniklar" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Uzbek" class="interlanguage-link-target">Oʻzbekcha / ўзбекча</a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D9%84%DB%8C%DA%86%D9%86" title="لیچن – Western Punjabi" lang="pnb" hreflang="pnb" data-title="لیچن" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target">پنجابی</a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Porosty" title="Porosty – Polish" lang="pl" hreflang="pl" data-title="Porosty" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target">Polski</a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/L%C3%ADquen" title="Líquen – Portuguese" lang="pt" hreflang="pt" data-title="Líquen" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target">Português</a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Lichen" title="Lichen – Romanian" lang="ro" hreflang="ro" data-title="Lichen" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target">Română</a></li><li class="interlanguage-link interwiki-qu mw-list-item"><a href="https://qu.wikipedia.org/wiki/Kusma_qara" title="Kusma qara – Quechua" lang="qu" hreflang="qu" data-title="Kusma qara" data-language-autonym="Runa Simi" data-language-local-name="Quechua" class="interlanguage-link-target">Runa Simi</a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9B%D0%B8%D1%88%D0%B0%D0%B9%D0%BD%D0%B8%D0%BA%D0%B8" title="Лишайники – Russian" lang="ru" hreflang="ru" data-title="Лишайники" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target">Русский</a></li><li class="interlanguage-link interwiki-sah mw-list-item"><a href="https://sah.wikipedia.org/wiki/%D0%9B%D0%B0%D0%B1%D1%8B%D0%BA%D1%82%D0%B0%D0%BB%D0%B0%D1%80" title="Лабыкталар – Yakut" lang="sah" hreflang="sah" data-title="Лабыкталар" data-language-autonym="Саха тыла" data-language-local-name="Yakut" class="interlanguage-link-target">Саха тыла</a></li><li class="interlanguage-link interwiki-sco mw-list-item"><a href="https://sco.wikipedia.org/wiki/Crottle" title="Crottle – Scots" lang="sco" hreflang="sco" data-title="Crottle" data-language-autonym="Scots" data-language-local-name="Scots" class="interlanguage-link-target">Scots</a></li><li class="interlanguage-link interwiki-sq mw-list-item"><a href="https://sq.wikipedia.org/wiki/Likenet" title="Likenet – Albanian" lang="sq" hreflang="sq" data-title="Likenet" data-language-autonym="Shqip" data-language-local-name="Albanian" class="interlanguage-link-target">Shqip</a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Lichen" title="Lichen – Simple English" lang="en-simple" hreflang="en-simple" data-title="Lichen" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target">Simple English</a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Li%C5%A1ajn%C3%ADk" title="Lišajník – Slovak" lang="sk" hreflang="sk" data-title="Lišajník" data-language-autonym="Slovenčina" data-language-local-name="Slovak" class="interlanguage-link-target">Slovenčina</a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Li%C5%A1aj" title="Lišaj – Slovenian" lang="sl" hreflang="sl" data-title="Lišaj" data-language-autonym="Slovenščina" data-language-local-name="Slovenian" class="interlanguage-link-target">Slovenščina</a></li><li class="interlanguage-link interwiki-ckb mw-list-item"><a href="https://ckb.wikipedia.org/wiki/%D9%82%DB%95%D9%88%D8%B2%DB%95%D8%A8%DB%95%D8%B1%D8%AF" title="قەوزەبەرد – Central Kurdish" lang="ckb" hreflang="ckb" data-title="قەوزەبەرد" data-language-autonym="کوردی" data-language-local-name="Central Kurdish" class="interlanguage-link-target">کوردی</a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%9B%D0%B8%D1%88%D0%B0%D1%98%D0%B8" title="Лишаји – Serbian" lang="sr" hreflang="sr" data-title="Лишаји" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target">Српски / srpski</a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/Li%C5%A1aj" title="Lišaj – Serbo-Croatian" lang="sh" hreflang="sh" data-title="Lišaj" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="Serbo-Croatian" class="interlanguage-link-target">Srpskohrvatski / српскохрватски</a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/J%C3%A4k%C3%A4l%C3%A4t" title="Jäkälät – Finnish" lang="fi" hreflang="fi" data-title="Jäkälät" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target">Suomi</a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Lavar" title="Lavar – Swedish" lang="sv" hreflang="sv" data-title="Lavar" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target">Svenska</a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%87%E0%AE%B2%E0%AF%88%E0%AE%95%E0%AF%8D%E0%AE%95%E0%AE%A9%E0%AF%8D" title="இலைக்கன் – Tamil" lang="ta" hreflang="ta" data-title="இலைக்கன்" data-language-autonym="தமிழ்" 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div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">For other uses, see <a href="/wiki/Lichen_(disambiguation)" class="mw-disambig" title="Lichen (disambiguation)">Lichen (disambiguation)</a>.</div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Lichen-covered_tree,_Tresco.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Lichen-covered_tree%2C_Tresco.jpg/220px-Lichen-covered_tree%2C_Tresco.jpg" decoding="async" width="220" height="337" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Lichen-covered_tree%2C_Tresco.jpg/330px-Lichen-covered_tree%2C_Tresco.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Lichen-covered_tree%2C_Tresco.jpg/440px-Lichen-covered_tree%2C_Tresco.jpg 2x" data-file-width="2128" data-file-height="3264" /></a><figcaption>A tree covered with leafy foliose lichens and shrubby fruticose lichens</figcaption></figure> <style data-mw-deduplicate="TemplateStyles:r1237032888/mw-parser-output/.tmulti">.mw-parser-output .tmulti .multiimageinner{display:flex;flex-direction:column}.mw-parser-output .tmulti .trow{display:flex;flex-direction:row;clear:left;flex-wrap:wrap;width:100%;box-sizing:border-box}.mw-parser-output .tmulti .tsingle{margin:1px;float:left}.mw-parser-output .tmulti .theader{clear:both;font-weight:bold;text-align:center;align-self:center;background-color:transparent;width:100%}.mw-parser-output .tmulti .thumbcaption{background-color:transparent}.mw-parser-output .tmulti .text-align-left{text-align:left}.mw-parser-output .tmulti .text-align-right{text-align:right}.mw-parser-output .tmulti .text-align-center{text-align:center}@media all and (max-width:720px){.mw-parser-output .tmulti .thumbinner{width:100%!important;box-sizing:border-box;max-width:none!important;align-items:center}.mw-parser-output .tmulti .trow{justify-content:center}.mw-parser-output .tmulti .tsingle{float:none!important;max-width:100%!important;box-sizing:border-box;text-align:center}.mw-parser-output .tmulti .tsingle .thumbcaption{text-align:left}.mw-parser-output .tmulti .trow>.thumbcaption{text-align:center}}@media screen{html.skin-theme-clientpref-night .mw-parser-output .tmulti .multiimageinner img{background-color:white}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .tmulti .multiimageinner img{background-color:white}}</style><div class="thumb tmulti tright"><div class="thumbinner multiimageinner" style="width:224px;max-width:224px"><div class="trow"><div class="theader" style="text-align:center">Common lichen growth forms</div></div><div class="trow"><div class="tsingle" style="width:222px;max-width:222px"><div class="thumbimage"><a href="/wiki/File:Letharia_vulpina_JHollinger_crop.jpg" class="mw-file-description"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/6e/Letharia_vulpina_JHollinger_crop.jpg/220px-Letharia_vulpina_JHollinger_crop.jpg" decoding="async" width="220" height="188" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/6e/Letharia_vulpina_JHollinger_crop.jpg/330px-Letharia_vulpina_JHollinger_crop.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/6e/Letharia_vulpina_JHollinger_crop.jpg/440px-Letharia_vulpina_JHollinger_crop.jpg 2x" data-file-width="1904" data-file-height="1628" /></a></div><div class="thumbcaption"><a href="/wiki/Letharia_vulpina" title="Letharia vulpina">Letharia vulpina</a>, wolf lichen, grows like a multiple-branched tuft or leafless mini-shrub, so it has a <a href="/wiki/Fruticose_lichen" title="Fruticose lichen">fruticose</a> growth form.</div></div></div><div class="trow"><div class="tsingle" style="width:222px;max-width:222px"><div class="thumbimage"><a href="/wiki/File:Flavoparmelia_caperata_-_Flickr_-_pellaea_(7).jpg" class="mw-file-description"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/5/54/Flavoparmelia_caperata_-_Flickr_-_pellaea_%287%29.jpg/220px-Flavoparmelia_caperata_-_Flickr_-_pellaea_%287%29.jpg" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/54/Flavoparmelia_caperata_-_Flickr_-_pellaea_%287%29.jpg/330px-Flavoparmelia_caperata_-_Flickr_-_pellaea_%287%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/54/Flavoparmelia_caperata_-_Flickr_-_pellaea_%287%29.jpg/440px-Flavoparmelia_caperata_-_Flickr_-_pellaea_%287%29.jpg 2x" data-file-width="3210" data-file-height="2407" /></a></div><div class="thumbcaption"><a href="/wiki/Flavoparmelia_caperata" title="Flavoparmelia caperata">Flavoparmelia caperata</a> has leaf-like structures, so it is <a href="/wiki/Foliose_lichen" title="Foliose lichen">foliose</a>.</div></div></div><div class="trow"><div class="tsingle" style="width:222px;max-width:222px"><div class="thumbimage"><a href="/wiki/File:Caloplaca_marina.JPG" class="mw-file-description"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/a/af/Caloplaca_marina.JPG/220px-Caloplaca_marina.JPG" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/af/Caloplaca_marina.JPG/330px-Caloplaca_marina.JPG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/af/Caloplaca_marina.JPG/440px-Caloplaca_marina.JPG 2x" data-file-width="2288" data-file-height="1712" /></a></div><div class="thumbcaption"><a href="/wiki/Caloplaca_marina" title="Caloplaca marina">Caloplaca marina</a> grows like an orange crust coating the rock, so it is <a href="/wiki/Crustose_lichen" title="Crustose lichen">crustose</a>.</div></div></div><div class="trow"><div class="tsingle" style="width:222px;max-width:222px"><div class="thumbimage"><a href="/wiki/File:Caloplaca_thallincola.jpg" class="mw-file-description"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/b/b9/Caloplaca_thallincola.jpg/220px-Caloplaca_thallincola.jpg" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/b/b9/Caloplaca_thallincola.jpg/330px-Caloplaca_thallincola.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/b/b9/Caloplaca_thallincola.jpg/440px-Caloplaca_thallincola.jpg 2x" data-file-width="800" data-file-height="600" /></a></div><div class="thumbcaption"><a href="/wiki/Variospora_thallincola" title="Variospora thallincola">Variospora thallincola</a> grows like a crust, and in a pattern that radiates outward from the center, so it has a <a href="/wiki/Crustose_lichen" title="Crustose lichen">crustose placodioid</a> growth form.</div></div></div><div class="trow"><div class="tsingle" style="width:222px;max-width:222px"><div class="thumbimage"><a href="/wiki/File:Chrysothrix_chlorina.jpg" class="mw-file-description"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/3c/Chrysothrix_chlorina.jpg/220px-Chrysothrix_chlorina.jpg" decoding="async" width="220" height="147" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/3c/Chrysothrix_chlorina.jpg/330px-Chrysothrix_chlorina.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/3c/Chrysothrix_chlorina.jpg/440px-Chrysothrix_chlorina.jpg 2x" data-file-width="3872" data-file-height="2592" /></a></div><div class="thumbcaption"><a href="/wiki/Chrysothrix_chlorina" title="Chrysothrix chlorina">Chrysothrix chlorina</a> grows like powder dusted on the rock, so it is <a href="/wiki/Leprose_lichen" class="mw-redirect" title="Leprose lichen">leprose</a>.</div></div></div><div class="trow"><div class="tsingle" style="width:222px;max-width:222px"><div class="thumbimage"><a href="/wiki/File:Blistered_Jelly_Lichen_(983550966).jpg" class="mw-file-description"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Blistered_Jelly_Lichen_%28983550966%29.jpg/220px-Blistered_Jelly_Lichen_%28983550966%29.jpg" decoding="async" width="220" height="176" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Blistered_Jelly_Lichen_%28983550966%29.jpg/330px-Blistered_Jelly_Lichen_%28983550966%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Blistered_Jelly_Lichen_%28983550966%29.jpg/440px-Blistered_Jelly_Lichen_%28983550966%29.jpg 2x" data-file-width="1000" data-file-height="800" /></a></div><div class="thumbcaption"><a href="/wiki/Collema_nigrescens" title="Collema nigrescens">Collema nigrescens</a> is <a href="/wiki/Gelatinous_lichen" class="mw-redirect" title="Gelatinous lichen">gelatinous</a>, without internal structure for its parts.</div></div></div></div></div> A lichen (<a href="/wiki/Help:IPA/English" title="Help:IPA/English">/ˈlaɪkən/</a> <a href="/wiki/Help:Pronunciation_respelling_key" title="Help:Pronunciation respelling key">LY-kən</a>, <style data-mw-deduplicate="TemplateStyles:r1177148991">.mw-parser-output .IPA-label-small{font-size:85%}.mw-parser-output .references .IPA-label-small,.mw-parser-output .infobox .IPA-label-small,.mw-parser-output .navbox .IPA-label-small{font-size:100%}</style><a href="/wiki/British_English" title="British English">UK</a> also <a href="/wiki/Help:IPA/English" title="Help:IPA/English">/ˈlɪtʃən/</a> <a href="/wiki/Help:Pronunciation_respelling_key" title="Help:Pronunciation respelling key">LITCH-ən</a>) is a hybrid <a href="/wiki/Colony_(biology)" title="Colony (biology)">colony</a> of <a href="/wiki/Algae" title="Algae">algae</a> or <a href="/wiki/Cyanobacteria" title="Cyanobacteria">cyanobacteria</a> living <a href="/wiki/Symbiotic" class="mw-redirect" title="Symbiotic">symbiotically</a> among <a href="/wiki/Hypha" title="Hypha">filaments</a> of multiple <a href="/wiki/Fungus" title="Fungus">fungi</a> species, along with <a href="/wiki/Yeast" title="Yeast">yeasts</a> and bacteria<a href="#cite_note-1">[1]</a><a href="#cite_note-2">[2]</a> embedded in the cortex or "skin", in a <a href="/wiki/Mutualism_(biology)" title="Mutualism (biology)">mutualistic</a> relationship.<a href="#cite_note-pmid_27445309-3">[3]</a><a href="#cite_note-WIL-4">[4]</a><a href="#cite_note-UCILABK-5">[5]</a><a href="#cite_note-LNA-6">[6]</a><a href="#cite_note-7">[7]</a> Lichens are the lifeform that first brought the term symbiosis (as symbiotismus<a href="#cite_note-8">[8]</a>) under biological context. Lichens have since been recognized as important actors in <a href="/wiki/Nutrient_cycling" class="mw-redirect" title="Nutrient cycling">nutrient cycling</a> and producers which many higher trophic feeders feed on, such as reindeer, gastropods, nematodes, mites, and springtails.<a href="#cite_note-9">[9]</a><a href="#cite_note-10">[10]</a><a href="#cite_note-11">[11]</a><a href="#cite_note-12">[12]</a> Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not <a href="/wiki/Plant" title="Plant">plants</a>. They may have tiny, leafless branches (<a href="/wiki/Fruticose" class="mw-redirect" title="Fruticose">fruticose</a>); flat leaf-like structures (<a href="/wiki/Foliose" class="mw-redirect" title="Foliose">foliose</a>); grow crust-like, adhering tightly to a surface (<a href="/wiki/Substrate_(biology)" title="Substrate (biology)">substrate</a>) like a thick coat of paint (<a href="/wiki/Crustose" title="Crustose">crustose</a>);<a href="#cite_note-ANBGLG-13">[13]</a> have a powder-like appearance (<a href="/wiki/Leprose_lichen" class="mw-redirect" title="Leprose lichen">leprose</a>); or other <a href="/wiki/Lichen_growth_forms" title="Lichen growth forms">growth forms</a>.<a href="#cite_note-LAL-14">[14]</a> A macrolichen is a lichen that is either bush-like or leafy; all other lichens are termed microlichens.<a href="#cite_note-WIL-4">[4]</a> Here, "macro" and "micro" do not refer to size, but to the growth form.<a href="#cite_note-WIL-4">[4]</a> <a href="/wiki/List_of_common_names_of_lichen_genera" title="List of common names of lichen genera">Common names for lichens</a> may contain the word <a href="/wiki/Moss" title="Moss">moss</a> (e.g., "<a href="/wiki/Reindeer_moss" class="mw-redirect" title="Reindeer moss">reindeer moss</a>", "<a href="/wiki/Iceland_moss" class="mw-redirect" title="Iceland moss">Iceland moss</a>"), and lichens may superficially look like and grow with <a href="/wiki/Moss" title="Moss">mosses</a>, but they are not closely related to mosses or any plant.<a href="#cite_note-LNA-6">[6]</a>: 3  Lichens do not have roots that absorb water and nutrients as plants do,<a href="#cite_note-SSFGCL-15">[15]</a>: 2  but like plants, they produce their own nutrition by <a href="/wiki/Photosynthesis" title="Photosynthesis">photosynthesis</a>.<a href="#cite_note-LLHE-16">[16]</a> When they grow on plants, they do not live as <a href="/wiki/Parasite" class="mw-redirect" title="Parasite">parasites</a>, but instead use the plant's surface as a substrate. Lichens occur from <a href="/wiki/Sea_level" title="Sea level">sea level</a> to high <a href="/wiki/Alpine_zone" class="mw-redirect" title="Alpine zone">alpine</a> elevations, in many environmental conditions, and can grow on almost any surface.<a href="#cite_note-LLHE-16">[16]</a><a href="#cite_note-17">[17]</a> They are abundant growing on bark, <a href="/wiki/Leaf" title="Leaf">leaves</a>, mosses, or other lichens<a href="#cite_note-SSFGCL-15">[15]</a> and hanging from branches "living on thin air" (<a href="/wiki/Epiphyte" title="Epiphyte">epiphytes</a>) in <a href="/wiki/Rain_forest" class="mw-redirect" title="Rain forest">rainforests</a> and in <a href="/wiki/Temperate_woodland" class="mw-redirect" title="Temperate woodland">temperate woodland</a>. They grow on rock, walls, <a href="/wiki/Gravestones" class="mw-redirect" title="Gravestones">gravestones</a>, <a href="/wiki/Roof" title="Roof">roofs</a>, exposed soil surfaces, rubber, bones, and in the soil as part of <a href="/wiki/Biological_soil_crust" title="Biological soil crust">biological soil crusts</a>. Various lichens have adapted to survive in some of the most extreme environments on Earth: <a href="/wiki/Arctic_tundra" class="mw-redirect" title="Arctic tundra">arctic tundra</a>, hot dry <a href="/wiki/Desert" title="Desert">deserts</a>, <a href="/wiki/Rocky_coast" class="mw-redirect" title="Rocky coast">rocky coasts</a>, and toxic <a href="/wiki/Slag" title="Slag">slag</a> heaps. They can even live inside solid rock, growing between the grains (<a href="/wiki/Endolithic_lichen" title="Endolithic lichen">endolithic</a>). There are about 20,000 known species.<a href="#cite_note-UCLS-18">[18]</a> <a href="/wiki/Lepraria" title="Lepraria">Some lichens</a> have lost the ability to reproduce sexually, yet continue to <a href="/wiki/Speciate" class="mw-redirect" title="Speciate">speciate</a>.<a href="#cite_note-SSFGCL-15">[15]</a><a href="#cite_note-LTRNASL-19">[19]</a> They can be seen as being relatively self-contained miniature <a href="/wiki/Ecosystem" title="Ecosystem">ecosystems</a>, where the fungi, algae, or cyanobacteria have the potential to engage with other <a href="/wiki/Microorganism" title="Microorganism">microorganisms</a> in a functioning system that may evolve as an even more complex <a href="/wiki/Holobiont" title="Holobiont">composite organism</a>.<a href="#cite_note-TT-20">[20]</a><a href="#cite_note-FESM-21">[21]</a><a href="#cite_note-SSSFD-22">[22]</a><a href="#cite_note-NPB-23">[23]</a> Lichens may be <a href="/wiki/Longevity" title="Longevity">long-lived</a>, with some considered to be among the oldest living things.<a href="#cite_note-LNA-6">[6]</a><a href="#cite_note-Morris2007-24">[24]</a> They are among the first living things to grow on fresh rock exposed after an event such as a landslide. The long life-span and slow and regular growth rate of some species can be used to date events (<a href="/wiki/Lichenometry" title="Lichenometry">lichenometry</a>). Lichens are a <a href="/wiki/Keystone_species" title="Keystone species">keystone species</a> in many ecosystems and benefit <a href="/wiki/Trees" class="mw-redirect" title="Trees">trees</a> and <a href="/wiki/Birds" class="mw-redirect" title="Birds">birds</a>.<a href="#cite_note-25">[25]</a> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Etymology_and_pronunciation">Etymology and pronunciation</h2>[<a href="/w/index.php?title=Lichen&action=edit&section=1" title="Edit section: Etymology and pronunciation">edit</a>]</div> The English word lichen derives from the <a href="/wiki/Greek_language" title="Greek language">Greek</a> λειχήν leichēn ("tree moss, lichen, lichen-like eruption on skin") via <a href="/wiki/Latin" title="Latin">Latin</a> lichen.<a href="#cite_note-26">[26]</a><a href="#cite_note-27">[27]</a><a href="#cite_note-28">[28]</a> The Greek noun, which literally means "licker", derives from the verb λείχειν leichein, "to lick".<a href="#cite_note-29">[29]</a><a href="#cite_note-Beekes-30">[30]</a> In American English, "lichen" is pronounced the same as the verb "liken" (<a href="/wiki/Help:IPA/English" title="Help:IPA/English">/ˈlaɪkən/</a>). In British English, both this pronunciation and one rhyming with "kitchen" (<a href="/wiki/Help:IPA/English" title="Help:IPA/English">/ˈlɪtʃən/</a>) are used.<a href="#cite_note-Cambridge-31">[31]</a><a href="#cite_note-Wordsworth-32">[32]</a><a href="#cite_note-33">[33]</a> <div class="mw-heading mw-heading2"><h2 id="Anatomy_and_morphology">Anatomy and morphology</h2>[<a href="/w/index.php?title=Lichen&action=edit&section=2" title="Edit section: Anatomy and morphology">edit</a>]</div> <div class="mw-heading mw-heading3"><h3 id="Growth_forms">Growth forms</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=3" title="Edit section: Growth forms">edit</a>]</div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Lichen_growth_forms" title="Lichen growth forms">Lichen growth forms</a></div> Lichens grow in a wide range of shapes and forms; this external appearance is known as their <a href="/wiki/Morphology_(biology)" title="Morphology (biology)">morphology</a>. The shape of a lichen is usually determined by the organization of the fungal filaments.<a href="#cite_note-MSULB-34">[34]</a> The nonreproductive tissues, or vegetative body parts, are called the <a href="/wiki/Thallus" title="Thallus">thallus</a>. Lichens are grouped by thallus type, since the thallus is usually the most visually prominent part of the lichen. Thallus growth forms typically correspond to a few basic internal structure types. <a href="/wiki/List_of_common_names_of_lichen_genera" title="List of common names of lichen genera">Common names for lichens</a> often come from a growth form or color that is typical of a lichen <a href="/wiki/Genus" title="Genus">genus</a>. Common groupings of lichen thallus growth forms are: <ol><li><a href="/wiki/Fruticose_lichen" title="Fruticose lichen">fruticose</a><a href="#cite_note-LNAILV-35">[35]</a><a href="#cite_note-ASLGPZ-36">[36]</a><a href="#cite_note-Dobson-37">[37]</a> – growing like a tuft or multiple-branched leafless mini-shrub, upright or hanging down, 3-dimensional branches with nearly round cross section (<a href="/wiki/Terete" title="Terete">terete</a>) or flattened</li> <li><a href="/wiki/Foliose_lichen" title="Foliose lichen">foliose</a><a href="#cite_note-LNAILV-35">[35]</a><a href="#cite_note-ASLGPZ-36">[36]</a> – growing in 2-dimensional, flat, leaf-like lobes</li> <li><a href="/wiki/Crustose_lichen" title="Crustose lichen">crustose</a><a href="#cite_note-ANBGLG-13">[13]</a><a href="#cite_note-LNAILV-35">[35]</a><a href="#cite_note-ASLGPZ-36">[36]</a> – crust-like, adhering tightly to a surface (<a href="/wiki/Substrate_(biology)" title="Substrate (biology)">substrate</a>) like a thick coat of paint</li> <li><a href="/wiki/Squamulose_lichen" title="Squamulose lichen">squamulose</a><a href="#cite_note-Dobson-37">[37]</a> – formed of small leaf-like scales crustose below but free at the tips</li> <li><a href="/wiki/Leprose_lichen" class="mw-redirect" title="Leprose lichen">leprose</a><a href="#cite_note-ASLTT-38">[38]</a> – powdery</li> <li><a href="/wiki/Gelatinous_lichen" class="mw-redirect" title="Gelatinous lichen">gelatinous</a> – jelly-like</li> <li><a href="/wiki/Filamentous_lichen" class="mw-redirect" title="Filamentous lichen">filamentous</a> – stringy or like matted hair</li> <li><a href="/wiki/Byssoid_lichen" class="mw-redirect" title="Byssoid lichen">byssoid</a> – wispy, like <a href="/wiki/Teased_wool" class="mw-redirect" title="Teased wool">teased wool</a></li> <li>structureless</li></ol> There are variations in growth types in a single lichen species, grey areas between the growth type descriptions, and overlapping between growth types, so some authors might describe lichens using different growth type descriptions. When a crustose lichen gets old, the center may start to crack up like old-dried paint, old-broken asphalt paving, or like the polygonal "islands" of cracked-up mud in a dried lakebed. This is called being <a href="/wiki/Rimose" title="Rimose">rimose</a> or <a href="/wiki/Areolate" class="mw-redirect" title="Areolate">areolate</a>, and the "island" pieces separated by the cracks are called areolas.<a href="#cite_note-LNAILV-35">[35]</a> The areolas appear separated, but are (or were)[<a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed">citation needed</a>] connected by an underlying <a href="/wiki/Prothallus#In_lichens" title="Prothallus">prothallus</a> or <a href="/wiki/Hypothallus" title="Hypothallus">hypothallus</a>.<a href="#cite_note-ASLTT-38">[38]</a> When a crustose lichen grows from a center and appears to radiate out, it is called crustose placodioid. When the edges of the areolas lift up from the substrate, it is called <a href="/wiki/Squamulose" class="mw-redirect" title="Squamulose">squamulose</a>.<a href="#cite_note-VMBMLF-39">[39]</a>: 159 <a href="#cite_note-Dobson-37">[37]</a> These growth form groups are not precisely defined. Foliose lichens may sometimes branch and appear to be fruticose. Fruticose lichens may have flattened branching parts and appear leafy. Squamulose lichens may appear where the edges lift up. Gelatinous lichens may appear leafy when dry.<a href="#cite_note-VMBMLF-39">[39]</a>: 159  The thallus is not always the part of the lichen that is most visually noticeable. Some lichens can grow inside solid rock between the grains (<a href="/wiki/Endolithic_lichen" title="Endolithic lichen">endolithic lichens</a>), with only the sexual fruiting part visible growing outside the rock.<a href="#cite_note-LNAILV-35">[35]</a> These may be dramatic in color or appearance.<a href="#cite_note-LNAILV-35">[35]</a> Forms of these sexual parts are not in the above growth form categories.<a href="#cite_note-LNAILV-35">[35]</a> The most visually noticeable reproductive parts are often circular, raised, plate-like or disc-like outgrowths, with crinkly edges, and are described in sections below. <div class="mw-heading mw-heading3"><h3 id="Color">Color</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=4" title="Edit section: Color">edit</a>]</div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Bipartite_and_tripartite_cyanolichens_(10.3897-mycokeys.6.3869)_Figure_1.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/9/9d/Bipartite_and_tripartite_cyanolichens_%2810.3897-mycokeys.6.3869%29_Figure_1.jpg/220px-Bipartite_and_tripartite_cyanolichens_%2810.3897-mycokeys.6.3869%29_Figure_1.jpg" decoding="async" width="220" height="152" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/9d/Bipartite_and_tripartite_cyanolichens_%2810.3897-mycokeys.6.3869%29_Figure_1.jpg/330px-Bipartite_and_tripartite_cyanolichens_%2810.3897-mycokeys.6.3869%29_Figure_1.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/9d/Bipartite_and_tripartite_cyanolichens_%2810.3897-mycokeys.6.3869%29_Figure_1.jpg/440px-Bipartite_and_tripartite_cyanolichens_%2810.3897-mycokeys.6.3869%29_Figure_1.jpg 2x" data-file-width="1512" data-file-height="1047" /></a><figcaption>Lichens</figcaption></figure> Lichens come in many colors.<a href="#cite_note-SSFGCL-15">[15]</a>: 4  Coloration is usually determined by the photosynthetic component.<a href="#cite_note-MSULB-34">[34]</a> Special pigments, such as yellow <a href="/wiki/Usnic_acid" title="Usnic acid">usnic acid</a>, give lichens a variety of colors, including reds, oranges, yellows, and browns, especially in exposed, dry habitats.<a href="#cite_note-SJCL-40">[40]</a> In the absence of special pigments, lichens are usually bright green to olive gray when wet, gray or grayish-green to brown when dry.<a href="#cite_note-SJCL-40">[40]</a> This is because moisture causes the surface skin (<a href="/wiki/Cortex_(lichen)" class="mw-redirect" title="Cortex (lichen)">cortex</a>) to become more transparent, exposing the green photobiont layer.<a href="#cite_note-SJCL-40">[40]</a> Different colored lichens covering large areas of exposed rock surfaces, or lichens covering or hanging from bark can be a spectacular display when the patches of diverse colors "come to life" or "glow" in brilliant displays following rain. Different colored lichens may inhabit different adjacent sections of a rock face, depending on the angle of exposure to light.<a href="#cite_note-SJCL-40">[40]</a> Colonies of lichens may be spectacular in appearance, dominating much of the surface of the visual landscape in forests and natural places, such as the <a href="/wiki/Lichens_of_the_Sierra_Nevada_(U.S.)#On_rock" title="Lichens of the Sierra Nevada (U.S.)">vertical "paint"</a> covering the vast rock faces of <a href="/wiki/Yosemite_National_Park" title="Yosemite National Park">Yosemite National Park</a>.<a href="#cite_note-LSN-41">[41]</a> Color is used in identification.<a href="#cite_note-JJMML-42">[42]</a>: 4  The color of a lichen changes depending on whether the lichen is wet or dry.<a href="#cite_note-JJMML-42">[42]</a> Color descriptions used for identification are based on the color that shows when the lichen is dry.<a href="#cite_note-JJMML-42">[42]</a> Dry lichens with a cyanobacterium as the photosynthetic partner tend to be dark grey, brown, or black.<a href="#cite_note-JJMML-42">[42]</a> The underside of the leaf-like lobes of foliose lichens is a different color from the top side (<a href="/wiki/Dorsiventral" title="Dorsiventral">dorsiventral</a>), often brown or black, sometimes white. A fruticose lichen may have flattened "branches", appearing similar to a foliose lichen, but the underside of a leaf-like structure on a fruticose lichen is the same color as the top side. The leaf-like lobes of a foliose lichen may branch, giving the appearance of a fruticose lichen, but the underside will be a different color from the top side.<a href="#cite_note-ASLTT-38">[38]</a> The sheen on some jelly-like gelatinous lichens is created by <a href="/wiki/Mucilaginous" class="mw-redirect" title="Mucilaginous">mucilaginous</a> secretions.<a href="#cite_note-MSULB-34">[34]</a> <div class="mw-heading mw-heading3"><h3 id="Internal_structure">Internal structure</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=5" title="Edit section: Internal structure">edit</a>]</div> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Lichen_cross_section_%E2%80%93_heteromeric_thallus.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/77/Lichen_cross_section_%E2%80%93_heteromeric_thallus.svg/220px-Lichen_cross_section_%E2%80%93_heteromeric_thallus.svg.png" decoding="async" width="220" height="227" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/77/Lichen_cross_section_%E2%80%93_heteromeric_thallus.svg/330px-Lichen_cross_section_%E2%80%93_heteromeric_thallus.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/77/Lichen_cross_section_%E2%80%93_heteromeric_thallus.svg/440px-Lichen_cross_section_%E2%80%93_heteromeric_thallus.svg.png 2x" data-file-width="516" data-file-height="532" /></a><figcaption>Schematic cross section of <a href="/wiki/Foliose_lichen" title="Foliose lichen">foliose lichen</a>: (a) The <a href="/wiki/Cortex_(lichen)" class="mw-redirect" title="Cortex (lichen)">cortex</a> is the outer layer of tightly woven fungus filaments (<a href="/wiki/Hyphae" class="mw-redirect" title="Hyphae">hyphae</a>) (b) This photobiont layer has photosynthesizing <a href="/wiki/Green_algae" title="Green algae">green algae</a> (c) Loosely packed hyphae in the medulla (d) A tightly woven lower cortex (e) Anchoring hyphae called <a href="/wiki/Rhizines" class="mw-redirect" title="Rhizines">rhizines</a> where the fungus attaches to the substrate</figcaption></figure> A lichen consists of a simple photosynthesizing organism, usually a <a href="/wiki/Green_alga" class="mw-redirect" title="Green alga">green alga</a> or <a href="/wiki/Cyanobacterium" class="mw-redirect" title="Cyanobacterium">cyanobacterium</a>, surrounded by filaments of a fungus. Generally, most of a lichen's bulk is made of interwoven fungal filaments,<a href="#cite_note-LMM-43">[43]</a> but this is reversed in filamentous and gelatinous lichens.<a href="#cite_note-MSULB-34">[34]</a> The fungus is called a mycobiont. The photosynthesizing organism is called a photobiont. Algal photobionts are called phycobionts.<a href="#cite_note-UNOLP-44">[44]</a> Cyanobacteria photobionts are called cyanobionts.<a href="#cite_note-UNOLP-44">[44]</a> The part of a lichen that is not involved in reproduction, the "body" or "vegetative tissue" of a lichen, is called the thallus. The thallus form is very different from any form where the fungus or alga are growing separately. The thallus is made up of filaments of the fungus called <a href="/wiki/Hyphae" class="mw-redirect" title="Hyphae">hyphae</a>. The filaments grow by branching then rejoining to create a mesh, which is called being "<a href="/wiki/Anastomosis" title="Anastomosis">anastomosed</a>". The mesh of fungal filaments may be dense or loose. Generally, the fungal mesh surrounds the algal or <a href="/wiki/Cyanobacterial" class="mw-redirect" title="Cyanobacterial">cyanobacterial</a> cells, often enclosing them within complex fungal tissues that are unique to lichen associations. The thallus may or may not have a protective "skin" of densely packed fungal filaments, often containing a second fungal species,<a href="#cite_note-pmid_27445309-3">[3]</a> which is called a cortex. Fruticose lichens have one cortex layer wrapping around the "branches". Foliose lichens have an upper cortex on the top side of the "leaf", and a separate lower cortex on the bottom side. Crustose and squamulose lichens have only an upper cortex, with the "inside" of the lichen in direct contact with the surface they grow on (the substrate). Even if the edges peel up from the substrate and appear flat and leaf-like, they lack a lower cortex, unlike foliose lichens. Filamentous, byssoid, leprose,<a href="#cite_note-ASLTT-38">[38]</a> gelatinous, and other lichens do not have a cortex; in other words, they are <a href="/wiki/Ecorticate" class="mw-redirect" title="Ecorticate">ecorticate</a>.<a href="#cite_note-ASLGGO-45">[45]</a> Fruticose, foliose, crustose, and squamulose lichens generally have up to three different types of tissue, <a href="/wiki/Cellular_differentiation" title="Cellular differentiation">differentiated</a> by having different densities of fungal filaments.<a href="#cite_note-LMM-43">[43]</a> The top layer, where the lichen contacts the environment, is called a cortex.<a href="#cite_note-LMM-43">[43]</a> The cortex is made of densely tightly woven, packed, and glued together (<a href="/wiki/Agglutination_(biology)" title="Agglutination (biology)">agglutinated</a>) fungal filaments.<a href="#cite_note-LMM-43">[43]</a> The dense packing makes the cortex act like a protective "skin", keeping other organisms out, and reducing the intensity of sunlight on the layers below.<a href="#cite_note-LMM-43">[43]</a> The cortex layer can be up to several hundred micrometers (μm) in thickness (less than a millimeter).<a href="#cite_note-Budel1996-46">[46]</a> The cortex may be further topped by an epicortex of secretions, not cells, 0.6–1 μm thick in <a href="/wiki/Parmeliaceae" title="Parmeliaceae">some lichens</a>.<a href="#cite_note-Budel1996-46">[46]</a> This secretion layer may or may not have pores.<a href="#cite_note-Budel1996-46">[46]</a> Below the cortex layer is a layer called the photobiontic layer or symbiont layer.<a href="#cite_note-ASLGPZ-36">[36]</a><a href="#cite_note-LMM-43">[43]</a> The symbiont layer has less densely packed fungal filaments, with the photosynthetic partner embedded in them.<a href="#cite_note-LMM-43">[43]</a> The less dense packing allows air circulation during photosynthesis, similar to the anatomy of a leaf.<a href="#cite_note-LMM-43">[43]</a> Each cell or group of cells of the photobiont is usually individually wrapped by hyphae, and in some cases penetrated by a <a href="/wiki/Haustorium" title="Haustorium">haustorium</a>.<a href="#cite_note-MSULB-34">[34]</a> In crustose and foliose lichens, algae in the photobiontic layer are diffuse among the fungal filaments, decreasing in gradation into the layer below. In fruticose lichens, the photobiontic layer is sharply distinct from the layer below.<a href="#cite_note-MSULB-34">[34]</a> The layer beneath the symbiont layer is called the <a href="/wiki/Medulla_(lichenology)" title="Medulla (lichenology)">medulla</a>. The medulla is less densely packed with fungal filaments than the layers above. In foliose lichens, as in <a href="/wiki/Peltigera" title="Peltigera">Peltigera</a>,<a href="#cite_note-VMBMLF-39">[39]</a>: 159  there is usually another densely packed layer of fungal filaments called the lower cortex.<a href="#cite_note-ASLTT-38">[38]</a><a href="#cite_note-LMM-43">[43]</a> Root-like fungal structures called rhizines (<a href="/wiki/Hypogymnia" title="Hypogymnia">usually</a>)<a href="#cite_note-VMBMLF-39">[39]</a>: 159  grow from the lower cortex to attach or anchor the lichen to the substrate.<a href="#cite_note-WIL-4">[4]</a><a href="#cite_note-ASLTT-38">[38]</a> Fruticose lichens have a single cortex wrapping all the way around the "stems" and "branches".<a href="#cite_note-VMBMLF-39">[39]</a> The medulla is the lowest layer, and may form a cottony white inner core for the branchlike thallus, or it may be hollow.<a href="#cite_note-VMBMLF-39">[39]</a>: 159  Crustose and squamulose lichens lack a lower cortex, and the medulla is in direct contact with the <a href="/wiki/Substrate_(biology)" title="Substrate (biology)">substrate</a> that the lichen grows on. In crustose areolate lichens, the edges of the areolas peel up from the substrate and appear leafy. In squamulose lichens the part of the lichen thallus that is not attached to the substrate may also appear leafy. But these leafy parts lack a lower cortex, which distinguishes crustose and squamulose lichens from foliose lichens.<a href="#cite_note-LMM-43">[43]</a> Conversely, foliose lichens may appear flattened against the substrate like a crustose lichen, but most of the leaf-like lobes can be lifted up from the substrate because it is separated from it by a tightly packed lower cortex.<a href="#cite_note-ASLTT-38">[38]</a> Gelatinous,<a href="#cite_note-VMBMLF-39">[39]</a>: 159  byssoid, and leprose lichens lack a cortex (are <a href="/wiki/Ecorticate" class="mw-redirect" title="Ecorticate">ecorticate</a>), and generally have only undifferentiated tissue, similar to only having a symbiont layer.[<a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed">citation needed</a>] In lichens that include both green algal and cyanobacterial symbionts, the cyanobacteria may be held on the upper or lower surface in small pustules called <a href="/wiki/Cephalodium" title="Cephalodium">cephalodia</a>. <a href="/wiki/Pruinia" class="mw-redirect" title="Pruinia">Pruinia</a> is a whitish coating on top of an upper surface.<a href="#cite_note-HPTCLGD-47">[47]</a> An <a href="/wiki/Epinecral_layer" class="mw-redirect" title="Epinecral layer">epinecral layer</a> is "a layer of horny dead fungal hyphae with indistinct <a href="/wiki/Lumen_(anatomy)" title="Lumen (anatomy)">lumina</a> in or near the cortex above the algal layer".<a href="#cite_note-HPTCLGD-47">[47]</a> In August 2016, it was reported that some macrolichens have more than one species of fungus in their tissues.<a href="#cite_note-pmid_27445309-3">[3]</a> <div class="mw-heading mw-heading2"><h2 id="Physiology">Physiology</h2>[<a href="/w/index.php?title=Lichen&action=edit&section=6" title="Edit section: Physiology">edit</a>]</div> <div class="mw-heading mw-heading3"><h3 id="Symbiotic_relation">Symbiotic relation</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=7" title="Edit section: Symbiotic relation">edit</a>]</div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Symbiosis_in_lichens" title="Symbiosis in lichens">Symbiosis in lichens</a></div> <style data-mw-deduplicate="TemplateStyles:r1244412712">.mw-parser-output .templatequote{overflow:hidden;margin:1em 0;padding:0 32px}.mw-parser-output .templatequotecite{line-height:1.5em;text-align:left;margin-top:0}@media(min-width:500px){.mw-parser-output .templatequotecite{padding-left:1.6em}}</style><blockquote class="templatequote">Lichens are fungi that have discovered agriculture<div class="templatequotecite">— <cite>Trevor Goward<a href="#cite_note-LNAILBE-48">[48]</a></cite></div></blockquote> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Abb4.1_Lichenes_lichens_interaction_fungus_alga_Mykobiont_Photobiont_2021_(M._Piepenbring).png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/9/91/Abb4.1_Lichenes_lichens_interaction_fungus_alga_Mykobiont_Photobiont_2021_%28M._Piepenbring%29.png/220px-Abb4.1_Lichenes_lichens_interaction_fungus_alga_Mykobiont_Photobiont_2021_%28M._Piepenbring%29.png" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/91/Abb4.1_Lichenes_lichens_interaction_fungus_alga_Mykobiont_Photobiont_2021_%28M._Piepenbring%29.png/330px-Abb4.1_Lichenes_lichens_interaction_fungus_alga_Mykobiont_Photobiont_2021_%28M._Piepenbring%29.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/91/Abb4.1_Lichenes_lichens_interaction_fungus_alga_Mykobiont_Photobiont_2021_%28M._Piepenbring%29.png/440px-Abb4.1_Lichenes_lichens_interaction_fungus_alga_Mykobiont_Photobiont_2021_%28M._Piepenbring%29.png 2x" data-file-width="3000" data-file-height="2250" /></a><figcaption>Lichens interaction</figcaption></figure> A lichen is a composite organism that emerges from <a href="/wiki/Algae" title="Algae">algae</a> or <a href="/wiki/Cyanobacteria" title="Cyanobacteria">cyanobacteria</a> living among the filaments (<a href="/wiki/Hyphae" class="mw-redirect" title="Hyphae">hyphae</a>) of the <a href="/wiki/Fungus" title="Fungus">fungi</a> in a mutually beneficial <a href="/wiki/Symbiotic" class="mw-redirect" title="Symbiotic">symbiotic</a> relationship. The fungi benefit from the carbohydrates produced by the algae or cyanobacteria via <a href="/wiki/Photosynthesis" title="Photosynthesis">photosynthesis</a>. The algae or cyanobacteria benefit by being protected from the environment by the filaments of the fungi, which also gather moisture and nutrients from the environment, and (usually) provide an anchor to it. Although some photosynthetic partners in a lichen can survive outside the lichen, the lichen symbiotic association extends the ecological range of both partners, whereby most descriptions of lichen associations describe them as symbiotic. Both partners gain water and mineral nutrients mainly from the atmosphere, through rain and dust. The fungal partner protects the alga by retaining water, serving as a larger capture area for mineral nutrients and, in some cases, provides minerals obtained from the <a href="/wiki/Substrate_(biology)" title="Substrate (biology)">substrate</a>. If a <a href="/wiki/Cyanobacteria" title="Cyanobacteria">cyanobacterium</a> is present, as a primary partner or another symbiont in addition to a green alga as in certain <a href="/wiki/Tripartite_symbiosis" title="Tripartite symbiosis">tripartite</a> lichens, they can <a href="/wiki/Nitrogen_fixation" title="Nitrogen fixation">fix atmospheric nitrogen</a>, complementing the activities of the green alga. In three different lineages the fungal partner has independently lost the mitochondrial gene atp9, which has key functions in mitochondrial energy production. The loss makes the fungi completely dependent on their symbionts.<a href="#cite_note-49">[49]</a> The algal or cyanobacterial cells are <a href="/wiki/Photosynthesis" title="Photosynthesis">photosynthetic</a> and, as in plants, they <a href="/wiki/Redox" title="Redox">reduce</a> atmospheric <a href="/wiki/Carbon_dioxide" title="Carbon dioxide">carbon dioxide</a> into organic carbon sugars to feed both symbionts. Phycobionts (algae) produce <a href="/wiki/Sugar_alcohols" class="mw-redirect" title="Sugar alcohols">sugar alcohols</a> (<a href="/wiki/Ribitol" title="Ribitol">ribitol</a>, <a href="/wiki/Sorbitol" title="Sorbitol">sorbitol</a>, and <a href="/wiki/Erythritol" title="Erythritol">erythritol</a>), which are absorbed by the mycobiont (fungus).<a href="#cite_note-UNOLP-44">[44]</a> Cyanobionts produce <a href="/wiki/Glucose" title="Glucose">glucose</a>.<a href="#cite_note-UNOLP-44">[44]</a> Lichenized fungal cells can make the photobiont "leak" out the products of photosynthesis, where they can then be absorbed by the fungus.<a href="#cite_note-SSFGCL-15">[15]</a>: 5  It appears many, probably the majority, of lichen also live in a symbiotic relationship with an order of <a href="/wiki/Basidiomycete" class="mw-redirect" title="Basidiomycete">basidiomycete</a> yeasts called <a href="/wiki/Cystobasidiomycetes" title="Cystobasidiomycetes">Cyphobasidiales</a>. The absence of this third partner could explain why growing lichen in the laboratory is difficult. The yeast cells are responsible for the formation of the characteristic cortex of the lichen thallus, and could also be important for its shape.<a href="#cite_note-50">[50]</a> The lichen combination of alga or cyanobacterium with a fungus has a very different form (morphology), physiology, and biochemistry than the component fungus, alga, or cyanobacterium growing by itself, naturally or in culture. The body (<a href="/wiki/Thallus" title="Thallus">thallus</a>) of most lichens is different from those of either the fungus or alga growing separately. When grown in the laboratory in the absence of its photobiont, a lichen fungus develops as a structureless, undifferentiated mass of fungal filaments (<a href="/wiki/Hyphae" class="mw-redirect" title="Hyphae">hyphae</a>). If combined with its photobiont under appropriate conditions, its characteristic form associated with the photobiont emerges, in the process called <a href="/wiki/Morphogenesis" title="Morphogenesis">morphogenesis</a>.<a href="#cite_note-LNA-6">[6]</a> In a few remarkable cases, a single lichen fungus can develop into two very different lichen forms when associating with either a green algal or a cyanobacterial symbiont. Quite naturally, these alternative forms were at first considered to be different species, until they were found growing in a conjoined manner.[<a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed">citation needed</a>] Evidence that lichens are examples of successful <a href="/wiki/Symbiosis" title="Symbiosis">symbiosis</a> is the fact that lichens can be found in almost every habitat and geographic area on the planet.<a href="#cite_note-TT-20">[20]</a> Two species in two genera of green algae are found in over 35% of all lichens, but can only rarely be found living on their own outside of a lichen.<a href="#cite_note-51">[51]</a> In a case where one fungal partner simultaneously had two green algae partners that outperform each other in different climates, this might indicate having more than one photosynthetic partner at the same time might enable the lichen to exist in a wider range of habitats and geographic locations.<a href="#cite_note-TT-20">[20]</a> At least one form of lichen, the North American beard-like lichens, are constituted of not two but three symbiotic partners: an ascomycetous fungus, a photosynthetic alga, and, unexpectedly, a basidiomycetous yeast.<a href="#cite_note-52">[52]</a> Phycobionts can have a net output of sugars with only water vapor.<a href="#cite_note-UNOLP-44">[44]</a> The thallus must be saturated with liquid water for cyanobionts to photosynthesize.<a href="#cite_note-UNOLP-44">[44]</a> Algae produce sugars that are absorbed by the fungus by diffusion into special fungal hyphae called <a href="/wiki/Appressoria" class="mw-redirect" title="Appressoria">appressoria</a> or <a href="/wiki/Haustoria" class="mw-redirect" title="Haustoria">haustoria</a> in contact with the wall of the algal cells.<a href="#cite_note-ELWWL-53">[53]</a> The appressoria or haustoria may produce a substance that increases permeability of the algal cell walls, and may penetrate the walls.<a href="#cite_note-ELWWL-53">[53]</a> The algae may contribute up to 80% of their sugar production to the fungus.<a href="#cite_note-ELWWL-53">[53]</a> <div class="mw-heading mw-heading3"><h3 id="Ecology">Ecology</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=8" title="Edit section: Ecology">edit</a>]</div> Lichen associations may be examples of <a href="/wiki/Mutualism_(biology)" title="Mutualism (biology)">mutualism</a> or <a href="/wiki/Commensalism" title="Commensalism">commensalism</a>, but the lichen relationship can be considered <a href="/wiki/Parasitism" title="Parasitism">parasitic</a><a href="#cite_note-Ahmadjian-54">[54]</a> under circumstances where the photosynthetic partner can exist in nature independently of the fungal partner, but not vice versa. Photobiont cells are routinely destroyed in the course of <a href="/wiki/Nutrient" title="Nutrient">nutrient</a> exchange. The association continues because reproduction of the photobiont cells matches the rate at which they are destroyed.<a href="#cite_note-Ahmadjian-54">[54]</a> The <a href="/wiki/Fungus" title="Fungus">fungus</a> surrounds the algal cells,<a href="#cite_note-LLHE-16">[16]</a> often enclosing them within complex fungal tissues unique to lichen associations. In many species the fungus penetrates the algal cell wall,<a href="#cite_note-LLHE-16">[16]</a> forming penetration pegs (<a href="/wiki/Haustoria" class="mw-redirect" title="Haustoria">haustoria</a>) similar to those produced by <a href="/wiki/Pathogenic_fungus" title="Pathogenic fungus">pathogenic fungi</a> that feed on a host.<a href="#cite_note-Dobson-37">[37]</a><a href="#cite_note-Honegger1988-55">[55]</a> <a href="/wiki/Cyanobacteria" title="Cyanobacteria">Cyanobacteria</a> in laboratory settings can grow faster when they are alone rather than when they are part of a lichen. <div class="mw-heading mw-heading4"><h4 id="Miniature_ecosystem_and_holobiont_theory">Miniature ecosystem and holobiont theory</h4>[<a href="/w/index.php?title=Lichen&action=edit&section=9" title="Edit section: Miniature ecosystem and holobiont theory">edit</a>]</div> Symbiosis in lichens is so well-balanced that lichens have been considered to be relatively self-contained miniature ecosystems in and of themselves.<a href="#cite_note-TT-20">[20]</a><a href="#cite_note-FESM-21">[21]</a> It is thought that lichens may be even more complex symbiotic systems that include non-photosynthetic bacterial communities performing other functions as partners in a <a href="/wiki/Holobiont" title="Holobiont">holobiont</a>.<a href="#cite_note-SSSFD-22">[22]</a><a href="#cite_note-NPB-23">[23]</a> Many lichens are very sensitive to environmental disturbances and can be used to cheaply<a href="#cite_note-LLHE-16">[16]</a> assess <a href="/wiki/Air_pollution" title="Air pollution">air pollution</a>,<a href="#cite_note-ferry-56">[56]</a><a href="#cite_note-rosehawksworth-57">[57]</a><a href="#cite_note-Hawksworthrose1976-58">[58]</a> <a href="/wiki/Ozone" title="Ozone">ozone</a> depletion, and metal contamination. Lichens have been used in making <a href="/wiki/Dye" title="Dye">dyes</a>, perfumes (<a href="/wiki/Oakmoss" title="Oakmoss">oakmoss</a>),<a href="#cite_note-OMAO-59">[59]</a> and in <a href="/wiki/Traditional_medicine" title="Traditional medicine">traditional medicines</a>. A few lichen species are eaten by insects<a href="#cite_note-LLHE-16">[16]</a> or larger animals, such as reindeer.<a href="#cite_note-60">[60]</a> Lichens are widely used as environmental indicators or bio-indicators. When air is very badly polluted with sulphur dioxide, there may be no lichens present; only some green algae can tolerate those conditions. If the air is clean, then shrubby, hairy and leafy lichens become abundant. A few lichen species can tolerate fairly high levels of pollution, and are commonly found in urban areas, on pavements, walls and tree bark. The most sensitive lichens are shrubby and leafy, while the most tolerant lichens are all crusty in appearance. Since industrialisation, many of the shrubby and leafy lichens such as <a href="/wiki/Ramalina" title="Ramalina">Ramalina</a>, <a href="/wiki/Usnea" title="Usnea">Usnea</a> and <a href="/wiki/Lobaria" title="Lobaria">Lobaria</a> species have very limited ranges, often being confined to the areas which have the cleanest air. <div class="mw-heading mw-heading4"><h4 id="Lichenicolous_fungi">Lichenicolous fungi</h4>[<a href="/w/index.php?title=Lichen&action=edit&section=10" title="Edit section: Lichenicolous fungi">edit</a>]</div> Some fungi can only be found living on lichens as <a href="/wiki/Obligate_parasite" title="Obligate parasite">obligate parasites</a>. These are referred to as <a href="/wiki/Lichenicolous_fungi" class="mw-redirect" title="Lichenicolous fungi">lichenicolous fungi</a>, and are a different species from the fungus living inside the lichen; thus they are not considered to be part of the lichen.<a href="#cite_note-LFIEB-61">[61]</a> <div class="mw-heading mw-heading3"><h3 id="Reaction_to_water">Reaction to water</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=11" title="Edit section: Reaction to water">edit</a>]</div> Moisture makes the cortex become more transparent.<a href="#cite_note-SSFGCL-15">[15]</a>: 4  This way, the algae can conduct photosynthesis when moisture is available, and is protected at other times. When the cortex is more transparent, the algae show more clearly and the lichen looks greener. <div class="mw-heading mw-heading3"><h3 id="Metabolites,_metabolite_structures_and_bioactivity">Metabolites, metabolite structures and bioactivity</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=12" title="Edit section: Metabolites, metabolite structures and bioactivity">edit</a>]</div> Lichens can show intense antioxidant activity.<a href="#cite_note-62">[62]</a><a href="#cite_note-63">[63]</a> <a href="/wiki/Secondary_metabolites" class="mw-redirect" title="Secondary metabolites">Secondary metabolites</a> are often deposited as crystals in the <a href="/wiki/Apoplast" title="Apoplast">apoplast</a>.<a href="#cite_note-HSLNA-64">[64]</a> Secondary metabolites are thought to play a role in preference for some substrates over others.<a href="#cite_note-HSLNA-64">[64]</a> <div class="mw-heading mw-heading3"><h3 id="Growth_rate">Growth rate</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=13" title="Edit section: Growth rate">edit</a>]</div> Lichens often have a regular but very slow growth rate of less than a millimeter per year. In crustose lichens, the area along the margin is where the most active growth is taking place.<a href="#cite_note-VMBMLF-39">[39]</a>: 159  Most crustose lichens grow only 1–2 mm in diameter per year. <div class="mw-heading mw-heading3"><h3 id="Life_span">Life span</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=14" title="Edit section: Life span">edit</a>]</div> Lichens may be <a href="/wiki/Longevity" title="Longevity">long-lived</a>, with some considered to be among the oldest living organisms.<a href="#cite_note-LNA-6">[6]</a><a href="#cite_note-Morris2007-24">[24]</a> Lifespan is difficult to measure because what defines the "same" individual lichen is not precise.<a href="#cite_note-ELNGDL-65">[65]</a> Lichens grow by vegetatively breaking off a piece, which may or may not be defined as the "same" lichen, and two lichens can merge, then becoming the "same" lichen.<a href="#cite_note-ELNGDL-65">[65]</a> One specimen of <a href="/wiki/Rhizocarpon_geographicum" title="Rhizocarpon geographicum">Rhizocarpon geographicum</a> on East <a href="/wiki/Baffin_Island" title="Baffin Island">Baffin Island</a> has an estimated age of 9500 years.<a href="#cite_note-66">[66]</a><a href="#cite_note-67">[67]</a> <a href="/wiki/Thallus" title="Thallus">Thalli</a> of Rhizocarpon geographicum and <a href="/wiki/Rhizocarpon" title="Rhizocarpon">Rhizocarpon</a> eupetraeoides/inarense in the central <a href="/wiki/Brooks_Range" title="Brooks Range">Brooks Range</a> of northern Alaska have been given a maximum possible age of 10,000–11,500 years.<a href="#cite_note-68">[68]</a><a href="#cite_note-69">[69]</a> <div class="mw-heading mw-heading3"><h3 id="Response_to_environmental_stress">Response to environmental stress</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=15" title="Edit section: Response to environmental stress">edit</a>]</div> Unlike simple dehydration in plants and animals, lichens may experience a complete loss of body water in dry periods.<a href="#cite_note-LLHE-16">[16]</a> Lichens are capable of surviving extremely low levels of <a href="/wiki/Water" title="Water">water</a> content (<a href="/wiki/Poikilohydric" class="mw-redirect" title="Poikilohydric">poikilohydric</a>).<a href="#cite_note-Nash2008-intro-70">[70]</a>: 5–6  They quickly absorb water when it becomes available again, becoming soft and fleshy.<a href="#cite_note-LLHE-16">[16]</a> In tests, lichen survived and showed remarkable results on the <a href="/wiki/Adaptive_capacity" title="Adaptive capacity">adaptation capacity</a> of <a href="/wiki/Photosynthesis" title="Photosynthesis">photosynthetic activity</a> within the <a href="/wiki/Simulation" title="Simulation">simulation time</a> of 34 days under <a href="/wiki/Life_on_Mars_(planet)#Life_on_Earth_under_Martian_conditions" class="mw-redirect" title="Life on Mars (planet)">Martian conditions</a> in the Mars Simulation Laboratory (MSL) maintained by the <a href="/wiki/German_Aerospace_Center" title="German Aerospace Center">German Aerospace Center</a> (DLR).<a href="#cite_note-Skymania-20120426-71">[71]</a><a href="#cite_note-72">[72]</a> The <a href="/wiki/European_Space_Agency" title="European Space Agency">European Space Agency</a> has discovered that lichens can survive unprotected in space. In an experiment led by Leopoldo Sancho from the Complutense University of Madrid, two species of lichen—<a href="/wiki/Rhizocarpon_geographicum" title="Rhizocarpon geographicum">Rhizocarpon geographicum</a> and <a href="/wiki/Rusavskia_elegans" title="Rusavskia elegans">Rusavskia elegans</a>—were sealed in a capsule and launched on a Russian Soyuz rocket 31 May 2005. Once in orbit, the capsules were opened and the lichens were directly exposed to the vacuum of space with its widely fluctuating temperatures and cosmic radiation. After 15 days, the lichens were brought back to earth and were found to be unchanged in their ability to photosynthesize.<a href="#cite_note-urlESA_-_Human_Spaceflight_and_Exploration_-_Lichen_survives_in_space-73">[73]</a><a href="#cite_note-Sancho2007-74">[74]</a> <div class="mw-heading mw-heading2"><h2 id="Reproduction_and_dispersal">Reproduction and dispersal</h2>[<a href="/w/index.php?title=Lichen&action=edit&section=16" title="Edit section: Reproduction and dispersal">edit</a>]</div> <div class="mw-heading mw-heading3"><h3 id="Vegetative_reproduction">Vegetative reproduction</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=17" title="Edit section: Vegetative reproduction">edit</a>]</div> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Lichen_reproduction1.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/da/Lichen_reproduction1.jpg/220px-Lichen_reproduction1.jpg" decoding="async" width="220" height="247" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/da/Lichen_reproduction1.jpg/330px-Lichen_reproduction1.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/da/Lichen_reproduction1.jpg/440px-Lichen_reproduction1.jpg 2x" data-file-width="1800" data-file-height="2022" /></a><figcaption>Xanthoparmelia sp. with dark-colored reproductive structures (disc-like <a href="/wiki/Apothecia" class="mw-redirect" title="Apothecia">apothecia</a>) at center, surrounded by a pale coloured vegetative <a href="/wiki/Thallus" title="Thallus">thallus</a>.</figcaption></figure> Many lichens reproduce asexually, either by a piece breaking off and growing on its own (<a href="/wiki/Vegetative_reproduction" title="Vegetative reproduction">vegetative reproduction</a>) or through the dispersal of <a href="/wiki/Diaspore_(botany)" title="Diaspore (botany)">diaspores</a> containing a few algal cells surrounded by fungal cells.<a href="#cite_note-WIL-4">[4]</a> Because of the relative lack of differentiation in the thallus, the line between diaspore formation and vegetative reproduction is often blurred. Fruticose lichens can fragment, and new lichens can grow from the fragment (<a href="/wiki/Vegetative_reproduction" title="Vegetative reproduction">vegetative reproduction</a>). Many lichens break up into fragments when they dry, dispersing themselves by wind action, to resume growth when moisture returns.<a href="#cite_note-Eichorn2005-75">[75]</a><a href="#cite_note-Cook1995-76">[76]</a> Soredia (singular: "soredium") are small groups of algal cells surrounded by fungal filaments that form in structures called soralia, from which the soredia can be dispersed by wind.<a href="#cite_note-WIL-4">[4]</a> Isidia (singular: "isidium") are branched, spiny, elongated, outgrowths from the thallus that break off for mechanical dispersal.<a href="#cite_note-WIL-4">[4]</a> Lichen propagules (<a href="/wiki/Diaspore_(botany)" title="Diaspore (botany)">diaspores</a>) typically contain cells from both partners, although the fungal components of so-called "fringe species" rely instead on algal cells dispersed by the "core species".<a href="#cite_note-RaiBergman2002-77">[77]</a> <div class="mw-heading mw-heading3"><h3 id="Sexual_reproduction">Sexual reproduction</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=18" title="Edit section: Sexual reproduction">edit</a>]</div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Lichen_reproduction.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/d7/Lichen_reproduction.jpg/220px-Lichen_reproduction.jpg" decoding="async" width="220" height="173" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/d7/Lichen_reproduction.jpg/330px-Lichen_reproduction.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/d7/Lichen_reproduction.jpg/440px-Lichen_reproduction.jpg 2x" data-file-width="579" data-file-height="456" /></a><figcaption>Disc-like <a href="/wiki/Apothecia" class="mw-redirect" title="Apothecia">apothecia</a> (left) and <a href="/wiki/Thallus" title="Thallus">thallus</a> (right) on a <a href="/wiki/Foliose_lichen" title="Foliose lichen">foliose lichen</a></figcaption></figure> Structures involved in reproduction often appear as discs, bumps, or squiggly lines on the surface of the thallus.<a href="#cite_note-SSFGCL-15">[15]</a>: 4  Though it has been argued that sexual reproduction in photobionts is selected against, there is strong evidence that suggests meiotic activities (sexual reproduction) in Trebouxia.<a href="#cite_note-78">[78]</a><a href="#cite_note-79">[79]</a> Many lichen fungi reproduce sexually like other fungi, producing spores formed by <a href="/wiki/Meiosis" title="Meiosis">meiosis</a> and fusion of gametes. Following dispersal, such fungal spores must meet with a compatible algal partner before a functional lichen can form. Some lichen fungi belong to the phylum <a href="/wiki/Basidiomycota" title="Basidiomycota">Basidiomycota</a> (basidiolichens) and produce <a href="/wiki/Mushroom" title="Mushroom">mushroom</a>-like reproductive structures resembling those of their nonlichenized relatives. Most lichen fungi belong to <a href="/wiki/Ascomycetes" class="mw-redirect" title="Ascomycetes">Ascomycetes</a> (ascolichens). Among the ascolichens, <a href="/wiki/Spore" title="Spore">spores</a> are produced in spore-producing structures called <a href="/wiki/Ascomata" class="mw-redirect" title="Ascomata">ascomata</a>.<a href="#cite_note-SSFGCL-15">[15]</a> The most common types of ascomata are the <a href="/wiki/Apothecium" class="mw-redirect" title="Apothecium">apothecium</a> (plural: apothecia) and <a href="/wiki/Perithecium" class="mw-redirect" title="Perithecium">perithecium</a> (plural: perithecia).<a href="#cite_note-SSFGCL-15">[15]</a>: 14  Apothecia are usually cups or plate-like discs located on the top surface of the lichen thallus. When apothecia are shaped like squiggly line segments instead of like discs, they are called lirellae.<a href="#cite_note-SSFGCL-15">[15]</a>: 14  Perithecia are shaped like flasks that are immersed in the lichen thallus tissue, which has a small hole for the spores to escape the flask, and appear like black dots on the lichen surface.<a href="#cite_note-SSFGCL-15">[15]</a>: 14  The three most common spore body types are raised discs called <a href="/wiki/Ascocarp" title="Ascocarp">apothecia</a> (singular: apothecium), bottle-like cups with a small hole at the top called perithecia (singular: perithecium), and pycnidia (singular: pycnidium), shaped like perithecia but without asci (an <a href="/wiki/Ascus" title="Ascus">ascus</a> is the structure that contains and releases the sexual spores in fungi of the <a href="/wiki/Ascomycota" title="Ascomycota">Ascomycota</a>).<a href="#cite_note-80">[80]</a> The apothecium has a layer of exposed spore-producing cells called <a href="/wiki/Ascus" title="Ascus">asci</a> (singular: ascus), and is usually a different color from the thallus tissue.<a href="#cite_note-SSFGCL-15">[15]</a>: 14  When the apothecium has an outer margin, the margin is called the exciple.<a href="#cite_note-SSFGCL-15">[15]</a>: 14  When the exciple has a color similar to colored thallus tissue the apothecium or lichen is called lecanorine, meaning similar to members of the genus <a href="/wiki/Lecanora" title="Lecanora">Lecanora</a>.<a href="#cite_note-SSFGCL-15">[15]</a>: 14  When the exciple is blackened like carbon it is called lecideine meaning similar to members of the genus <a href="/wiki/Lecidea" title="Lecidea">Lecidea</a>.<a href="#cite_note-SSFGCL-15">[15]</a>: 14  When the margin is pale or colorless it is called biatorine.<a href="#cite_note-SSFGCL-15">[15]</a>: 14  <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Pilophorus_acicularis_109164.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/72/Pilophorus_acicularis_109164.jpg/220px-Pilophorus_acicularis_109164.jpg" decoding="async" width="220" height="199" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/72/Pilophorus_acicularis_109164.jpg/330px-Pilophorus_acicularis_109164.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/72/Pilophorus_acicularis_109164.jpg/440px-Pilophorus_acicularis_109164.jpg 2x" data-file-width="1193" data-file-height="1078" /></a><figcaption>Crust-like thallus with <a href="/wiki/Pseudopodetia" class="mw-redirect" title="Pseudopodetia">pseudopodetia</a></figcaption></figure> A "<a href="/wiki/Podetium" title="Podetium">podetium</a>" (plural: <a href="/wiki/Podetia" class="mw-redirect" title="Podetia">podetia</a>) is a lichenized stalk-like structure of the fruiting body rising from the thallus, associated with some fungi that produce a fungal <a href="/wiki/Apothecium" class="mw-redirect" title="Apothecium">apothecium</a>.<a href="#cite_note-ASLGPZ-36">[36]</a> Since it is part of the reproductive tissue, podetia are not considered part of the main body (thallus), but may be visually prominent.<a href="#cite_note-ASLGPZ-36">[36]</a> The podetium may be branched, and sometimes cup-like. They usually bear the fungal <a href="/wiki/Pycnidia" class="mw-redirect" title="Pycnidia">pycnidia</a> or <a href="/wiki/Apothecia" class="mw-redirect" title="Apothecia">apothecia</a> or both.<a href="#cite_note-ASLGPZ-36">[36]</a> Many lichens have <a href="/wiki/Apothecia" class="mw-redirect" title="Apothecia">apothecia</a> that are visible to the naked eye.<a href="#cite_note-WIL-4">[4]</a> Most lichens produce abundant sexual structures.<a href="#cite_note-Murtagh-81">[81]</a> Many species appear to disperse only by sexual spores.<a href="#cite_note-Murtagh-81">[81]</a> For example, the crustose lichens Graphis scripta and Ochrolechia parella produce no symbiotic vegetative propagules. Instead, the lichen-forming fungi of these species reproduce sexually by self-fertilization (i.e. they are <a href="/wiki/Homothallic" class="mw-redirect" title="Homothallic">homothallic</a>). This breeding system may enable successful reproduction in harsh environments.<a href="#cite_note-Murtagh-81">[81]</a> Mazaedia (singular: mazaedium) are apothecia shaped like a <a href="/wiki/Pin#Straight_pins" title="Pin">dressmaker's pin</a> in <a href="/wiki/Pin_lichen" class="mw-redirect" title="Pin lichen">pin lichens</a>, where the fruiting body is a brown or black mass of loose ascospores enclosed by a cup-shaped exciple, which sits on top of a tiny stalk.<a href="#cite_note-SSFGCL-15">[15]</a>: 15  <div class="mw-heading mw-heading2"><h2 id="Taxonomy_and_classification">Taxonomy and classification</h2>[<a href="/w/index.php?title=Lichen&action=edit&section=19" title="Edit section: Taxonomy and classification">edit</a>]</div> Lichens are classified by the fungal component. Lichen species are given the same scientific name (<a href="/wiki/Binomial_name" class="mw-redirect" title="Binomial name">binomial name</a>) as the fungus species in the lichen. Lichens are being integrated into the classification schemes for fungi. The alga bears its own scientific name, which bears no relationship to that of the lichen or fungus.<a href="#cite_note-Kirk_pp.378-81-82">[82]</a> There are about 20,000 identified lichen species,<a href="#cite_note-Lücking_et_al._2017-83">[83]</a><a href="#cite_note-Lücking_Hodkinson_Leavitt_2017-84">[84]</a> and taxonomists have estimated that the total number of lichen species (including those yet undiscovered) might be as high as 28,000.<a href="#cite_note-Lücking_et_al._2009-85">[85]</a> Nearly 20% of known fungal species are associated with lichens.<a href="#cite_note-ELWWL-53">[53]</a> "Lichenized fungus" may refer to the entire lichen, or to just the fungus. This may cause confusion without context. A particular fungus species may form lichens with different algae species, giving rise to what appear to be different lichen species, but which are still classified (as of 2014) as the same lichen species.<a href="#cite_note-FSSD-86">[86]</a> Formerly, some lichen taxonomists placed lichens in their own division, the Mycophycophyta, but this practice is no longer accepted because the components belong to separate <a href="/wiki/Biological_classification" class="mw-redirect" title="Biological classification">lineages</a>. Neither the ascolichens nor the basidiolichens form <a href="/wiki/Monophyletic" class="mw-redirect" title="Monophyletic">monophyletic</a> lineages in their respective fungal phyla, but they do form several major solely or primarily lichen-forming groups within each phylum.<a href="#cite_note-Lutzoni_2004-87">[87]</a> Even more unusual than basidiolichens is the fungus <a href="/wiki/Geosiphon" title="Geosiphon">Geosiphon pyriforme</a>, a member of the <a href="/wiki/Glomeromycota" title="Glomeromycota">Glomeromycota</a> that is unique in that it encloses a cyanobacterial symbiont inside its cells. <a href="/wiki/Geosiphon" title="Geosiphon">Geosiphon</a> is not usually considered to be a lichen, and its peculiar symbiosis was not recognized for many years. The genus is more closely allied to <a href="/wiki/Mycorrhiza" title="Mycorrhiza">endomycorrhizal</a> genera. Fungi from <a href="/wiki/Verrucariales" title="Verrucariales">Verrucariales</a> also form marine lichens with the <a href="/wiki/Brown_algae" title="Brown algae">brown algae</a> Petroderma maculiforme,<a href="#cite_note-88">[88]</a> and have a symbiotic relationship with <a href="/wiki/Seaweed" title="Seaweed">seaweed</a> (such as <a href="/wiki/Ascophyllum" title="Ascophyllum">rockweed</a>) and <a href="/wiki/Blidingia_minima" title="Blidingia minima">Blidingia minima</a>, where the algae are the dominant components. The fungi is thought to help the rockweeds to resist desiccation when exposed to air.<a href="#cite_note-89">[89]</a><a href="#cite_note-90">[90]</a> In addition, lichens can also use <a href="/wiki/Yellow-green_algae" title="Yellow-green algae">yellow-green algae</a> (<a href="/wiki/Heterococcus" title="Heterococcus">Heterococcus</a>) as their symbiotic partner.<a href="#cite_note-91">[91]</a> Lichens independently emerged from fungi associating with algae and cyanobacteria multiple times throughout history.<a href="#cite_note-92">[92]</a> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Lichen_foliac%C3%A9.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/40/Lichen_foliac%C3%A9.jpg/220px-Lichen_foliac%C3%A9.jpg" decoding="async" width="220" height="219" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/40/Lichen_foliac%C3%A9.jpg/330px-Lichen_foliac%C3%A9.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/40/Lichen_foliac%C3%A9.jpg/440px-Lichen_foliac%C3%A9.jpg 2x" data-file-width="759" data-file-height="754" /></a><figcaption>Mycobiont</figcaption></figure> <div class="mw-heading mw-heading3"><h3 id="Fungi">Fungi</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=20" title="Edit section: Fungi">edit</a>]</div> The fungal component of a lichen is called the mycobiont. The mycobiont may be an <a href="/wiki/Ascomycete" class="mw-redirect" title="Ascomycete">Ascomycete</a> or <a href="/wiki/Basidiomycete" class="mw-redirect" title="Basidiomycete">Basidiomycete</a>.<a href="#cite_note-UCLS-18">[18]</a> The associated lichens are called either <a href="/wiki/Ascomycota" title="Ascomycota">ascolichens</a> or <a href="/wiki/Basidiolichen" title="Basidiolichen">basidiolichens</a>, respectively. Living as a <a href="/wiki/Symbiosis" title="Symbiosis">symbiont</a> in a lichen appears to be a successful way for a fungus to derive essential nutrients, since about 20% of all fungal species have acquired this mode of life.<a href="#cite_note-Hawksworth1988-93">[93]</a> Thalli produced by a given fungal symbiont with its differing partners may be similar,[<a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed">citation needed</a>] and the secondary metabolites identical,[<a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed">citation needed</a>] indicating[<a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed">citation needed</a>] that the fungus has the dominant role in determining the morphology of the lichen. But the same mycobiont with different photobionts may also produce very different growth forms.<a href="#cite_note-FSSD-86">[86]</a> Lichens are known in which there is one fungus associated with two or even three algal species. Although each lichen thallus generally appears homogeneous, some evidence seems to suggest that the fungal component may consist of more than one genetic individual of that species.[<a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed">citation needed</a>] Two or more fungal species can interact to form the same lichen.<a href="#cite_note-Rikkinen1995-94">[94]</a> The following table lists the <a href="/wiki/Order_(biology)" title="Order (biology)">orders</a> and <a href="/wiki/Family_(biology)" title="Family (biology)">families</a> of fungi that include lichen-forming species. <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol ol,.mw-parser-output .hlist ol ul,.mw-parser-output .hlist ul dl,.mw-parser-output .hlist ul ol,.mw-parser-output .hlist ul ul{display:inline}.mw-parser-output .hlist .mw-empty-li{display:none}.mw-parser-output .hlist dt::after{content:": "}.mw-parser-output .hlist dd::after,.mw-parser-output .hlist li::after{content:" · ";font-weight:bold}.mw-parser-output .hlist dd:last-child::after,.mw-parser-output .hlist dt:last-child::after,.mw-parser-output .hlist li:last-child::after{content:none}.mw-parser-output .hlist dd dd:first-child::before,.mw-parser-output .hlist dd dt:first-child::before,.mw-parser-output .hlist dd li:first-child::before,.mw-parser-output .hlist dt dd:first-child::before,.mw-parser-output .hlist dt dt:first-child::before,.mw-parser-output .hlist dt li:first-child::before,.mw-parser-output .hlist li dd:first-child::before,.mw-parser-output .hlist li dt:first-child::before,.mw-parser-output .hlist li li:first-child::before{content:" (";font-weight:normal}.mw-parser-output .hlist dd dd:last-child::after,.mw-parser-output .hlist dd dt:last-child::after,.mw-parser-output .hlist dd li:last-child::after,.mw-parser-output .hlist dt dd:last-child::after,.mw-parser-output .hlist dt dt:last-child::after,.mw-parser-output .hlist dt li:last-child::after,.mw-parser-output .hlist li dd:last-child::after,.mw-parser-output .hlist li dt:last-child::after,.mw-parser-output .hlist li li:last-child::after{content:")";font-weight:normal}.mw-parser-output .hlist ol{counter-reset:listitem}.mw-parser-output .hlist ol>li{counter-increment:listitem}.mw-parser-output .hlist ol>li::before{content:" "counter(listitem)"\a0 "}.mw-parser-output .hlist dd ol>li:first-child::before,.mw-parser-output .hlist dt ol>li:first-child::before,.mw-parser-output .hlist li ol>li:first-child::before{content:" ("counter(listitem)"\a0 "}</style><style data-mw-deduplicate="TemplateStyles:r1236075235">.mw-parser-output .navbox{box-sizing:border-box;border:1px solid #a2a9b1;width:100%;clear:both;font-size:88%;text-align:center;padding:1px;margin:1em auto 0}.mw-parser-output .navbox .navbox{margin-top:0}.mw-parser-output .navbox+.navbox,.mw-parser-output .navbox+.navbox-styles+.navbox{margin-top:-1px}.mw-parser-output .navbox-inner,.mw-parser-output .navbox-subgroup{width:100%}.mw-parser-output .navbox-group,.mw-parser-output .navbox-title,.mw-parser-output .navbox-abovebelow{padding:0.25em 1em;line-height:1.5em;text-align:center}.mw-parser-output .navbox-group{white-space:nowrap;text-align:right}.mw-parser-output .navbox,.mw-parser-output .navbox-subgroup{background-color:#fdfdfd}.mw-parser-output .navbox-list{line-height:1.5em;border-color:#fdfdfd}.mw-parser-output .navbox-list-with-group{text-align:left;border-left-width:2px;border-left-style:solid}.mw-parser-output tr+tr>.navbox-abovebelow,.mw-parser-output tr+tr>.navbox-group,.mw-parser-output tr+tr>.navbox-image,.mw-parser-output tr+tr>.navbox-list{border-top:2px solid #fdfdfd}.mw-parser-output .navbox-title{background-color:#ccf}.mw-parser-output .navbox-abovebelow,.mw-parser-output .navbox-group,.mw-parser-output .navbox-subgroup .navbox-title{background-color:#ddf}.mw-parser-output .navbox-subgroup .navbox-group,.mw-parser-output .navbox-subgroup .navbox-abovebelow{background-color:#e6e6ff}.mw-parser-output .navbox-even{background-color:#f7f7f7}.mw-parser-output .navbox-odd{background-color:transparent}.mw-parser-output .navbox .hlist td dl,.mw-parser-output .navbox .hlist td ol,.mw-parser-output .navbox .hlist td ul,.mw-parser-output .navbox td.hlist dl,.mw-parser-output .navbox td.hlist ol,.mw-parser-output .navbox td.hlist ul{padding:0.125em 0}.mw-parser-output .navbox .navbar{display:block;font-size:100%}.mw-parser-output .navbox-title .navbar{float:left;text-align:left;margin-right:0.5em}body.skin--responsive .mw-parser-output .navbox-image img{max-width:none!important}@media print{body.ns-0 .mw-parser-output .navbox{display:none!important}}</style></div><div role="navigation" class="navbox" aria-labelledby="Taxonomy_of_the_Lichen_families" style="padding:3px"><table class="nowraplinks mw-collapsible mw-collapsed navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2" style="background:#ADD8E6 ;"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Lichen_family_taxonomy" title="Template:Lichen family taxonomy"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Lichen_family_taxonomy" title="Template talk:Lichen family taxonomy"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Lichen_family_taxonomy" title="Special:EditPage/Template:Lichen family taxonomy"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Taxonomy_of_the_Lichen_families" style="font-size:114%;margin:0 4em">Taxonomy of the <a class="mw-selflink selflink">Lichen</a> families</div></th></tr><tr><th scope="row" class="navbox-group" style="width:7em;text-align:left;background:#e6e6ff"><a href="/wiki/Ascomycota" title="Ascomycota">Ascomycota</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Arthoniomycetes" title="Arthoniomycetes">Arthoniomycetes</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:13em;text-align:left;"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Arthoniales" scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Arthoniales" title="Arthoniales">Arthoniales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Arthoniaceae" title="Arthoniaceae">Arthoniaceae</a></li> <li><a href="/wiki/Chrysothricaceae" class="mw-redirect" title="Chrysothricaceae">Chrysothricaceae</a></li> <li><a href="/wiki/Melaspileaceae" title="Melaspileaceae">Melaspileaceae</a></li> <li><a href="/wiki/Roccellaceae" title="Roccellaceae">Roccellaceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Dothideomycetes" title="Dothideomycetes">Dothideomycetes</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:13em;text-align:left;"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Capnodiales" title="Capnodiales">Capnodiales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Capnodiaceae" title="Capnodiaceae">Capnodiaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Dothideales" title="Dothideales">Dothideales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Dacampiaceae" title="Dacampiaceae">Dacampiaceae</a></li> <li><a href="/wiki/Xanthopyreniaceae" title="Xanthopyreniaceae">Xanthopyreniaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Pleosporales" title="Pleosporales">Pleosporales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Melanommataceae" title="Melanommataceae">Melanommataceae</a></li> <li><a href="/wiki/Mytilinidiaceae" title="Mytilinidiaceae">Mytilinidiaceae</a></li> <li><a href="/wiki/Pleomassariaceae" title="Pleomassariaceae">Pleomassariaceae</a></li> <li><a href="/wiki/Pleosporaceae" title="Pleosporaceae">Pleosporaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;;background:#E5E4E2"><a href="/wiki/Incertae_sedis" title="Incertae sedis">Incertae sedis</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Epigloeaceae" class="mw-redirect" title="Epigloeaceae">Epigloeaceae</a></li> <li><a href="/wiki/Arthopyreniaceae" class="mw-redirect" title="Arthopyreniaceae">Arthopyreniaceae</a></li> <li><a href="/wiki/Didymosphaeriaceae" title="Didymosphaeriaceae">Didymosphaeriaceae</a></li> <li><a href="/wiki/Lichenotheliaceae" title="Lichenotheliaceae">Lichenotheliaceae</a></li> <li><a href="/wiki/Microthyriaceae" title="Microthyriaceae">Microthyriaceae</a></li> <li><a href="/wiki/Mycosphaerellaceae" title="Mycosphaerellaceae">Mycosphaerellaceae</a></li> <li><a href="/wiki/Naetrocymbaceae" title="Naetrocymbaceae">Naetrocymbaceae</a></li> <li><a href="/wiki/Parmulariaceae" title="Parmulariaceae">Parmulariaceae</a></li> <li><a href="/wiki/Pseudoperisporiaceae" title="Pseudoperisporiaceae">Pseudoperisporiaceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Eurotiomycetes" title="Eurotiomycetes">Eurotiomycetes</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:13em;text-align:left;"><a href="/wiki/Chaetothyriomycetidae" title="Chaetothyriomycetidae">Chaetothyriomycetidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Chaetothyriales" title="Chaetothyriales">Chaetothyriales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Chaetothyriaceae" title="Chaetothyriaceae">Chaetothyriaceae</a></li> <li><a href="/wiki/Coccodiniaceae" title="Coccodiniaceae">Coccodiniaceae</a></li> <li><a href="/w/index.php?title=Cyphellophoraceae&action=edit&redlink=1" class="new" title="Cyphellophoraceae (page does not exist)">Cyphellophoraceae</a></li> <li><a href="/wiki/Epibryaceae" class="mw-redirect" title="Epibryaceae">Epibryaceae</a></li> <li><a href="/wiki/Herpotrichiellaceae" title="Herpotrichiellaceae">Herpotrichiellaceae</a></li> <li><a href="/wiki/Lyrommataceae" class="mw-redirect" title="Lyrommataceae">Lyrommataceae</a></li> <li><a href="/wiki/Microtheliopsidaceae" title="Microtheliopsidaceae">Microtheliopsidaceae</a></li> <li><a href="/wiki/Pyrenotrichaceae" title="Pyrenotrichaceae">Pyrenotrichaceae</a></li> <li><a href="/w/index.php?title=Trichomeriaceae&action=edit&redlink=1" class="new" title="Trichomeriaceae (page does not exist)">Trichomeriaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Pyrenulales" title="Pyrenulales">Pyrenulales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Monoblastiaceae" title="Monoblastiaceae">Monoblastiaceae</a></li> <li><a href="/wiki/Pyrenulaceae" title="Pyrenulaceae">Pyrenulaceae</a></li> <li><a href="/wiki/Requienellaceae" title="Requienellaceae">Requienellaceae</a></li> <li><a href="/wiki/Trypetheliaceae" title="Trypetheliaceae">Trypetheliaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Verrucariales" title="Verrucariales">Verrucariales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Adelococcaceae" title="Adelococcaceae">Adelococcaceae</a></li> <li><a href="/wiki/Verrucariaceae" title="Verrucariaceae">Verrucariaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;;background:#E5E4E2"><a href="/wiki/Incertae_sedis" title="Incertae sedis">Incertae sedis</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Strigulaceae" title="Strigulaceae">Strigulaceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:13em;text-align:left;;background:#E5E4E2"><a href="/wiki/Incertae_sedis" title="Incertae sedis">Incertae sedis</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Mycocaliciales" title="Mycocaliciales">Mycocaliciales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Mycocaliciaceae" title="Mycocaliciaceae">Mycocaliciaceae</a></li> <li><a href="/wiki/Sphinctrinaceae" title="Sphinctrinaceae">Sphinctrinaceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Lecanoromycetes" title="Lecanoromycetes">Lecanoromycetes</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:13em;text-align:left;"><a href="/wiki/Acarosporomycetidae" class="mw-redirect" title="Acarosporomycetidae">Acarosporomycetidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Acarosporales" scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Acarosporales" title="Acarosporales">Acarosporales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Acarosporaceae" title="Acarosporaceae">Acarosporaceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:13em;text-align:left;"><a href="/wiki/Lecanoromycetidae" title="Lecanoromycetidae">Lecanoromycetidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Caliciales" title="Caliciales">Caliciales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Caliciaceae" title="Caliciaceae">Caliciaceae</a></li> <li><a href="/wiki/Physciaceae" title="Physciaceae">Physciaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Lecanorales" title="Lecanorales">Lecanorales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Biatorellaceae" title="Biatorellaceae">Biatorellaceae</a></li> <li><a href="/wiki/Bruceomycetaceae" title="Bruceomycetaceae">Bruceomycetaceae</a></li> <li><a href="/wiki/Catillariaceae" title="Catillariaceae">Catillariaceae</a></li> <li><a href="/wiki/Cladoniaceae" title="Cladoniaceae">Cladoniaceae</a></li> <li><a href="/wiki/Gypsoplacaceae" class="mw-redirect" title="Gypsoplacaceae">Gypsoplacaceae</a></li> <li><a href="/wiki/Haematommataceae" class="mw-redirect" title="Haematommataceae">Haematommataceae</a></li> <li><a href="/wiki/Lecanoraceae" title="Lecanoraceae">Lecanoraceae</a></li> <li><a href="/wiki/Malmideaceae" title="Malmideaceae">Malmideaceae</a></li> <li><a href="/wiki/Megalariaceae" class="mw-redirect" title="Megalariaceae">Megalariaceae</a></li> <li><a href="/wiki/Parmeliaceae" title="Parmeliaceae">Parmeliaceae</a></li> <li><a href="/wiki/Pilocarpaceae" title="Pilocarpaceae">Pilocarpaceae</a></li> <li><a href="/wiki/Psilolechiaceae" class="mw-redirect" title="Psilolechiaceae">Psilolechiaceae</a></li> <li><a href="/wiki/Psoraceae" title="Psoraceae">Psoraceae</a></li> <li><a href="/wiki/Ramalinaceae" title="Ramalinaceae">Ramalinaceae</a></li> <li><a href="/wiki/Ramboldiaceae" class="mw-redirect" title="Ramboldiaceae">Ramboldiaceae</a></li> <li><a href="/wiki/Scoliciosporaceae" title="Scoliciosporaceae">Scoliciosporaceae</a></li> <li><a href="/wiki/Sphaerophoraceae" title="Sphaerophoraceae">Sphaerophoraceae</a></li> <li><a href="/wiki/Tephromelataceae" title="Tephromelataceae">Tephromelataceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Leprocaulales" class="mw-redirect" title="Leprocaulales">Leprocaulales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Leprocaulaceae" title="Leprocaulaceae">Leprocaulaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Peltigerales" title="Peltigerales">Peltigerales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Coccocarpiaceae" title="Coccocarpiaceae">Coccocarpiaceae</a></li> <li><a href="/wiki/Collemataceae" title="Collemataceae">Collemataceae</a></li> <li><a href="/wiki/Koerberiaceae" title="Koerberiaceae">Koerberiaceae</a></li> <li><a href="/wiki/Massalongiaceae" title="Massalongiaceae">Massalongiaceae</a></li> <li><a href="/wiki/Pannariaceae" title="Pannariaceae">Pannariaceae</a></li> <li><a href="/wiki/Peltigeraceae" title="Peltigeraceae">Peltigeraceae</a></li> <li><a href="/wiki/Placynthiaceae" title="Placynthiaceae">Placynthiaceae</a></li> <li><a href="/wiki/Vahliellaceae" class="mw-redirect" title="Vahliellaceae">Vahliellaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Rhizocarpales" title="Rhizocarpales">Rhizocarpales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Rhizocarpaceae" title="Rhizocarpaceae">Rhizocarpaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Sporastatiales" class="mw-redirect" title="Sporastatiales">Sporastatiales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Sporastatiaceae" title="Sporastatiaceae">Sporastatiaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Teloschistales" title="Teloschistales">Teloschistales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Brigantiaeaceae" title="Brigantiaeaceae">Brigantiaeaceae</a></li> <li><a href="/wiki/Megalosporaceae" title="Megalosporaceae">Megalosporaceae</a></li> <li><a href="/wiki/Teloschistaceae" title="Teloschistaceae">Teloschistaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;;background:#E5E4E2"><a href="/wiki/Incertae_sedis" title="Incertae sedis">Incertae sedis</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Coniocybaceae" title="Coniocybaceae">Coniocybaceae</a></li> <li><a href="/wiki/Fuscideaceae" title="Fuscideaceae">Fuscideaceae</a></li> <li><a href="/wiki/Phlyctidaceae" title="Phlyctidaceae">Phlyctidaceae</a></li> <li><a href="/wiki/Umbilicariaceae" title="Umbilicariaceae">Umbilicariaceae</a></li> <li><a href="/wiki/Vezdaeaceae" class="mw-redirect" title="Vezdaeaceae">Vezdaeaceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:13em;text-align:left;"><a href="/wiki/Ostropomycetidae" title="Ostropomycetidae">Ostropomycetidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Agyriales" class="mw-redirect" title="Agyriales">Agyriales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agyriaceae" title="Agyriaceae">Agyriaceae</a></li> <li><a href="/wiki/Anamylopsoraceae" class="mw-redirect" title="Anamylopsoraceae">Anamylopsoraceae</a></li> <li><a href="/wiki/Schaereriaceae" class="mw-redirect" title="Schaereriaceae">Schaereriaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Baeomycetales" title="Baeomycetales">Baeomycetales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Arctomiaceae" title="Arctomiaceae">Arctomiaceae</a></li> <li><a href="/wiki/Arthrorhaphidaceae" class="mw-redirect" title="Arthrorhaphidaceae">Arthrorhaphidaceae</a></li> <li><a href="/wiki/Baeomycetaceae" class="mw-redirect" title="Baeomycetaceae">Baeomycetaceae</a></li> <li><a href="/wiki/Cameroniaceae" class="mw-redirect" title="Cameroniaceae">Cameroniaceae</a></li> <li><a href="/wiki/Hymeneliaceae" title="Hymeneliaceae">Hymeneliaceae</a></li> <li><a href="/w/index.php?title=Protothelenellaceae&action=edit&redlink=1" class="new" title="Protothelenellaceae (page does not exist)">Protothelenellaceae</a></li> <li><a href="/wiki/Trapeliaceae" title="Trapeliaceae">Trapeliaceae</a></li> <li><a href="/wiki/Xylographaceae" title="Xylographaceae">Xylographaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Graphidales" title="Graphidales">Graphidales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Diploschistaceae" class="mw-redirect" title="Diploschistaceae">Diploschistaceae</a></li> <li><a href="/wiki/Fissurinaceae" class="mw-redirect" title="Fissurinaceae">Fissurinaceae</a></li> <li><a href="/wiki/Gomphillaceae" title="Gomphillaceae">Gomphillaceae</a></li> <li><a href="/wiki/Graphidaceae" title="Graphidaceae">Graphidaceae</a></li> <li><a href="/wiki/Redonographaceae" class="mw-redirect" title="Redonographaceae">Redonographaceae</a></li> <li><a href="/wiki/Thelotremataceae" class="mw-redirect" title="Thelotremataceae">Thelotremataceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Gyalectales" title="Gyalectales">Gyalectales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Coenogoniaceae" class="mw-redirect" title="Coenogoniaceae">Coenogoniaceae</a></li> <li><a href="/wiki/Gyalectaceae" title="Gyalectaceae">Gyalectaceae</a></li> <li><a href="/wiki/Phlyctidaceae" title="Phlyctidaceae">Phlyctidaceae</a></li> <li><a href="/wiki/Sagiolechiaceae" title="Sagiolechiaceae">Sagiolechiaceae</a></li> <li><a href="/wiki/Trichotheliaceae" class="mw-redirect" title="Trichotheliaceae">Trichotheliaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Odontotrematales" class="mw-redirect" title="Odontotrematales">Odontotrematales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Odontotremataceae" title="Odontotremataceae">Odontotremataceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Ostropales" title="Ostropales">Ostropales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Phaneromycetaceae" class="mw-redirect" title="Phaneromycetaceae">Phaneromycetaceae</a></li> <li><a href="/wiki/Spirographaceae" class="mw-redirect" title="Spirographaceae">Spirographaceae</a></li> <li><a href="/wiki/Stictidaceae" title="Stictidaceae">Stictidaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Pertusariales" title="Pertusariales">Pertusariales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agyriaceae" title="Agyriaceae">Agyriaceae</a></li> <li><a href="/wiki/Coccotremataceae" title="Coccotremataceae">Coccotremataceae</a></li> <li><a href="/wiki/Icmadophilaceae" title="Icmadophilaceae">Icmadophilaceae</a></li> <li><a href="/wiki/Megasporaceae" title="Megasporaceae">Megasporaceae</a></li> <li><a href="/wiki/Microcaliciaceae" class="mw-redirect" title="Microcaliciaceae">Microcaliciaceae</a></li> <li><a href="/wiki/Ochrolechiaceae" title="Ochrolechiaceae">Ochrolechiaceae</a></li> <li><a href="/wiki/Pertusariaceae" title="Pertusariaceae">Pertusariaceae</a></li> <li><a href="/wiki/Varicellariaceae" class="mw-redirect" title="Varicellariaceae">Varicellariaceae</a></li> <li><a href="/w/index.php?title=Variolariaceae&action=edit&redlink=1" class="new" title="Variolariaceae (page does not exist)">Variolariaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Sarrameanales" class="mw-redirect" title="Sarrameanales">Sarrameanales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Sarrameanaceae" title="Sarrameanaceae">Sarrameanaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Schaereriales" class="mw-redirect" title="Schaereriales">Schaereriales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Schaereriaceae" class="mw-redirect" title="Schaereriaceae">Schaereriaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Thelenellales" class="mw-redirect" title="Thelenellales">Thelenellales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Thelenellaceae" title="Thelenellaceae">Thelenellaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;;background:#E5E4E2"><a href="/wiki/Incertae_sedis" title="Incertae sedis">Incertae sedis</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Arctomiaceae" title="Arctomiaceae">Arctomiaceae</a></li> <li><a href="/wiki/Hymeneliaceae" title="Hymeneliaceae">Hymeneliaceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Leotiomycetes" title="Leotiomycetes">Leotiomycetes</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Leotiomycetidae" scope="row" class="navbox-group" style="width:13em;text-align:left;"><a href="/wiki/Leotiomycetidae" title="Leotiomycetidae">Leotiomycetidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Helotiales" title="Helotiales">Helotiales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/w/index.php?title=Amicodiscaceae&action=edit&redlink=1" class="new" title="Amicodiscaceae (page does not exist)">Amicodiscaceae</a></li> <li><a href="/wiki/Arachnopezizaceae" title="Arachnopezizaceae">Arachnopezizaceae</a></li> <li><a href="/w/index.php?title=Bryoglossaceae&action=edit&redlink=1" class="new" title="Bryoglossaceae (page does not exist)">Bryoglossaceae</a></li> <li><a href="/wiki/Cenangiaceae" title="Cenangiaceae">Cenangiaceae</a></li> <li><a href="/wiki/Chlorociboriaceae" title="Chlorociboriaceae">Chlorociboriaceae</a></li> <li><a href="/w/index.php?title=Chlorospleniaceae&action=edit&redlink=1" class="new" title="Chlorospleniaceae (page does not exist)">Chlorospleniaceae</a></li> <li><a href="/w/index.php?title=Chrysodiscaceae&action=edit&redlink=1" class="new" title="Chrysodiscaceae (page does not exist)">Chrysodiscaceae</a></li> <li><a href="/wiki/Dermateaceae" title="Dermateaceae">Dermateaceae</a></li> <li><a href="/w/index.php?title=Discinellaceae&action=edit&redlink=1" class="new" title="Discinellaceae (page does not exist)">Discinellaceae</a></li> <li><a href="/w/index.php?title=Drepanopezizaceae&action=edit&redlink=1" class="new" title="Drepanopezizaceae (page does not exist)">Drepanopezizaceae</a></li> <li><a href="/w/index.php?title=Gelatinodiscaceae&action=edit&redlink=1" class="new" title="Gelatinodiscaceae (page does not exist)">Gelatinodiscaceae</a></li> <li><a href="/w/index.php?title=Godroniaceae&action=edit&redlink=1" class="new" title="Godroniaceae (page does not exist)">Godroniaceae</a></li> <li><a href="/w/index.php?title=Hamatocanthoscyphaceae&action=edit&redlink=1" class="new" title="Hamatocanthoscyphaceae (page does not exist)">Hamatocanthoscyphaceae</a></li> <li><a href="/wiki/Helotiaceae" title="Helotiaceae">Helotiaceae</a></li> <li><a href="/wiki/Hemiphacidiaceae" class="mw-redirect" title="Hemiphacidiaceae">Hemiphacidiaceae</a></li> <li><a href="/w/index.php?title=Heterosphaeriaceae&action=edit&redlink=1" class="new" title="Heterosphaeriaceae (page does not exist)">Heterosphaeriaceae</a></li> <li><a href="/wiki/Hyaloscyphaceae" title="Hyaloscyphaceae">Hyaloscyphaceae</a></li> <li><a href="/w/index.php?title=Hydrocinaceae&action=edit&redlink=1" class="new" title="Hydrocinaceae (page does not exist)">Hydrocinaceae</a></li> <li><a href="/w/index.php?title=Lachnaceae&action=edit&redlink=1" class="new" title="Lachnaceae (page does not exist)">Lachnaceae</a></li> <li><a href="/wiki/Loramycetaceae" title="Loramycetaceae">Loramycetaceae</a></li> <li><a href="/w/index.php?title=Mitrulaceae&action=edit&redlink=1" class="new" title="Mitrulaceae (page does not exist)">Mitrulaceae</a></li> <li><a href="/wiki/Mollisiaceae" title="Mollisiaceae">Mollisiaceae</a></li> <li><a href="/w/index.php?title=Neolauriomycetaceae&action=edit&redlink=1" class="new" title="Neolauriomycetaceae (page does not exist)">Neolauriomycetaceae</a></li> <li><a href="/w/index.php?title=Pleuroascaceae&action=edit&redlink=1" class="new" title="Pleuroascaceae (page does not exist)">Pleuroascaceae</a></li> <li><a href="/w/index.php?title=Ploettnerulaceae&action=edit&redlink=1" class="new" title="Ploettnerulaceae (page does not exist)">Ploettnerulaceae</a></li> <li><a href="/wiki/Rutstroemiaceae" title="Rutstroemiaceae">Rutstroemiaceae</a></li> <li><a href="/wiki/Sclerotiniaceae" title="Sclerotiniaceae">Sclerotiniaceae</a></li> <li><a href="/w/index.php?title=Solenopeziaceae&action=edit&redlink=1" class="new" title="Solenopeziaceae (page does not exist)">Solenopeziaceae</a></li> <li><a href="/w/index.php?title=Tympanidaceae&action=edit&redlink=1" class="new" title="Tympanidaceae (page does not exist)">Tympanidaceae</a></li> <li><a href="/wiki/Vibrisseaceae" title="Vibrisseaceae">Vibrisseaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Lahmiales" class="mw-redirect" title="Lahmiales">Lahmiales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Lahmiaceae" class="mw-redirect" title="Lahmiaceae">Lahmiaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/w/index.php?title=Lauriomycetales&action=edit&redlink=1" class="new" title="Lauriomycetales (page does not exist)">Lauriomycetales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/w/index.php?title=Lauriomycetaceae&action=edit&redlink=1" class="new" title="Lauriomycetaceae (page does not exist)">Lauriomycetaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Leotiales" title="Leotiales">Leotiales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/w/index.php?title=Cochlearomycetaceae&action=edit&redlink=1" class="new" title="Cochlearomycetaceae (page does not exist)">Cochlearomycetaceae</a></li> <li><a href="/wiki/Leotiaceae" title="Leotiaceae">Leotiaceae</a></li> <li><a href="/w/index.php?title=Mniaeciaceae&action=edit&redlink=1" class="new" title="Mniaeciaceae (page does not exist)">Mniaeciaceae</a></li> <li><a href="/w/index.php?title=Tympanidaceae&action=edit&redlink=1" class="new" title="Tympanidaceae (page does not exist)">Tympanidaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/w/index.php?title=Marthamycetales&action=edit&redlink=1" class="new" title="Marthamycetales (page does not exist)">Marthamycetales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Marthamycetaceae" title="Marthamycetaceae">Marthamycetaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Medeolariales" title="Medeolariales">Medeolariales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Medeolariaceae" class="mw-redirect" title="Medeolariaceae">Medeolariaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/w/index.php?title=Micraspidales&action=edit&redlink=1" class="new" title="Micraspidales (page does not exist)">Micraspidales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/w/index.php?title=Micraspidaceae&action=edit&redlink=1" class="new" title="Micraspidaceae (page does not exist)">Micraspidaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/w/index.php?title=Phacidiales&action=edit&redlink=1" class="new" title="Phacidiales (page does not exist)">Phacidiales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/w/index.php?title=Helicogoniaceae&action=edit&redlink=1" class="new" title="Helicogoniaceae (page does not exist)">Helicogoniaceae</a></li> <li><a href="/wiki/Phacidiaceae" title="Phacidiaceae">Phacidiaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Rhytismatales" title="Rhytismatales">Rhytismatales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cudoniaceae" title="Cudoniaceae">Cudoniaceae</a></li> <li><a href="/wiki/Rhytismataceae" title="Rhytismataceae">Rhytismataceae</a></li> <li><a href="/w/index.php?title=Triblidiaceae&action=edit&redlink=1" class="new" title="Triblidiaceae (page does not exist)">Triblidiaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Thelebolales" title="Thelebolales">Thelebolales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pseudeurotiaceae" title="Pseudeurotiaceae">Pseudeurotiaceae</a></li> <li><a href="/wiki/Thelebolaceae" class="mw-redirect" title="Thelebolaceae">Thelebolaceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Lichinomycetes" class="mw-redirect" title="Lichinomycetes">Lichinomycetes</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:13em;text-align:left;"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Lichinales" scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Lichinales" title="Lichinales">Lichinales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Gloeoheppiaceae" title="Gloeoheppiaceae">Gloeoheppiaceae</a></li> <li><a href="/wiki/Lichinaceae" title="Lichinaceae">Lichinaceae</a></li> <li><a href="/wiki/Peltulaceae" class="mw-redirect" title="Peltulaceae">Peltulaceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Sordariomycetes" title="Sordariomycetes">Sordariomycetes</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:13em;text-align:left;"><a href="/wiki/Diaporthomycetidae" title="Diaporthomycetidae">Diaporthomycetidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:13em;text-align:left;"><a href="/wiki/Hypocreomycetidae" title="Hypocreomycetidae">Hypocreomycetidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Coronophorales" title="Coronophorales">Coronophorales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Nitschkiaceae" title="Nitschkiaceae">Nitschkiaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Hypocreales" title="Hypocreales">Hypocreales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bionectriaceae" title="Bionectriaceae">Bionectriaceae</a></li> <li><a href="/wiki/Nectriaceae" title="Nectriaceae">Nectriaceae</a></li> <li><a href="/wiki/Niessliaceae" title="Niessliaceae">Niessliaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Microascales" title="Microascales">Microascales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Microascaceae" title="Microascaceae">Microascaceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:13em;text-align:left;"><a href="/wiki/Xylariales" title="Xylariales">Xylariomycetidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Xylariales" title="Xylariales">Xylariales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Hyponectriaceae" title="Hyponectriaceae">Hyponectriaceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:13em;text-align:left;;background:#E5E4E2"><a href="/wiki/Incertae_sedis" title="Incertae sedis">Incertae sedis</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Phyllachorales" title="Phyllachorales">Phyllachorales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Phyllachoraceae" title="Phyllachoraceae">Phyllachoraceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;;background:#E5E4E2"><a href="/wiki/Incertae_sedis" title="Incertae sedis">Incertae sedis</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Obryzaceae" title="Obryzaceae">Obryzaceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;;background:#E5E4E2"><a href="/wiki/Incertae_sedis" title="Incertae sedis">Incertae sedis</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:13em;text-align:left;background:#E5E4E2"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Lahmiales" class="mw-redirect" title="Lahmiales">Lahmiales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Lahmiales" class="mw-redirect" title="Lahmiales">Lahmiaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;;background:#E5E4E2"><a href="/wiki/Incertae_sedis" title="Incertae sedis">Incertae sedis</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Aspidotheliaceae" class="mw-redirect" title="Aspidotheliaceae">Aspidotheliaceae</a></li> <li><a href="/wiki/Mastodiaceae" class="mw-redirect" title="Mastodiaceae">Mastodiaceae</a></li> <li><a href="/wiki/Thelenellaceae" title="Thelenellaceae">Thelenellaceae</a></li> <li><a href="/wiki/Baeomycetales" title="Baeomycetales">Baeomycetaceae</a></li> <li><a href="/wiki/Coccotremataceae" title="Coccotremataceae">Coccotremataceae</a></li> <li><a href="/wiki/Thelocarpaceae" title="Thelocarpaceae">Thelocarpaceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:7em;text-align:left;background:#e6e6ff"><a href="/wiki/Basidiomycota" title="Basidiomycota">Basidiomycota</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Basidiomycota" title="Basidiomycota">Basidiomycetes</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:13em;text-align:left;"><a href="/wiki/Agaricomycetidae" title="Agaricomycetidae">Agaricomycetidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Agaricales" title="Agaricales">Agaricales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Hygrophoraceae" title="Hygrophoraceae">Hygrophoraceae</a></li> <li><a href="/wiki/Tricholomataceae" title="Tricholomataceae">Tricholomataceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Atheliales" title="Atheliales">Atheliales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Atheliales" title="Atheliales">Atheliaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Boletales" title="Boletales">Boletales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Coniophoraceae" title="Coniophoraceae">Coniophoraceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Cantharellales" title="Cantharellales">Cantharellales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Clavulinaceae" title="Clavulinaceae">Clavulinaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Jaapiales" class="mw-redirect" title="Jaapiales">Jaapiales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Jaapiaceae" class="mw-redirect" title="Jaapiaceae">Jaapiaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Lepidostromatales" title="Lepidostromatales">Lepidostromatales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Lepidostromataceae" class="mw-redirect" title="Lepidostromataceae">Lepidostromataceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:13em;text-align:left;"><a href="/wiki/Tremellomycetidae" class="mw-redirect" title="Tremellomycetidae">Tremellomycetidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Tremellales" title="Tremellales">Tremellales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Syzygosporaceae" class="mw-redirect" title="Syzygosporaceae">Syzygosporaceae</a></li> <li><a href="/wiki/Tremellaceae" title="Tremellaceae">Tremellaceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Urediniomycetes" class="mw-redirect" title="Urediniomycetes">Urediniomycetes</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:13em;text-align:left;"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Atractiellomycetes" title="Atractiellomycetes">Atractiellales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Chionosphaeraceae" title="Chionosphaeraceae">Chionosphaeraceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;"><a href="/wiki/Rust_(fungus)" title="Rust (fungus)">Uredinales</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Pucciniaceae" title="Pucciniaceae">Pucciniaceae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em;text-align:left;;background:#E5E4E2"><a href="/wiki/Platygloeales" title="Platygloeales">Platygloeales</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Platygloeaceae" title="Platygloeaceae">Platygloeaceae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd hlist" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2" style="background:#ADD8E6"><div id="References" style="font-size:114%;margin:0 4em">References</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0;background:transparent;color:inherit;"><div style="padding:0px"><table class="navbox-columns-table" style="border-spacing: 0px; text-align:left;width:auto; margin-left:auto; margin-right:auto;"><tbody><tr style="vertical-align:top"><td style="width:5em;">   </td><td class="navbox-list" style="padding:0px;text-align: left;width:50%;"><div> <ul> <li><cite id="CITEREFAndersonEkman2005" class="citation journal cs1">Anderson, Heidi L.; Ekman, Stefan (2005). "Disintegration of the Micareaceae (lichenized Ascomycota): a molecular phylogeny based on mitochondrial rDNA sequences". Mycological Research. 109 (1): 21–30. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1017%2FS0953756204001625">10.1017/S0953756204001625</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/15736860">15736860</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Mycological+Research&rft.atitle=Disintegration+of+the+Micareaceae+%28lichenized+Ascomycota%29%3A+a+molecular+phylogeny+based+on+mitochondrial+rDNA+sequences&rft.volume=109&rft.issue=1&rft.pages=21-30&rft.date=2005&rft_id=info%3Adoi%2F10.1017%2FS0953756204001625&rft_id=info%3Apmid%2F15736860&rft.aulast=Anderson&rft.aufirst=Heidi+L.&rft.au=Ekman%2C+Stefan&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"></li> <li>CABI Bioscience Databases. Available online at <a rel="nofollow" class="external free" href="http://www.indexfungorum.org/">http://www.indexfungorum.org/</a>.</li> <li><cite id="CITEREFErtzLawreySikaroodiGillevet2008" class="citation journal cs1">Ertz, Damien; Lawrey, James D.; Sikaroodi, Masoumeh; Gillevet, Patrick M.; Fischer, Eberhard; Killmann, Dorothee; Sérusiaux, Emmanuël (2008). <a rel="nofollow" class="external text" href="http://orbi.ulg.ac.be/handle/2268/22808">"A new lineage of lichenized basidiomycetes inferred from a two-gene phylogeny: The Lepidostromataceae with three species from the tropics"</a>. American Journal of Botany. 95 (12): 1548–1556. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.3732%2Fajb.0800232">10.3732/ajb.0800232</a>. <a href="/wiki/Hdl_(identifier)" class="mw-redirect" title="Hdl (identifier)">hdl</a>:<a rel="nofollow" class="external text" href="https://hdl.handle.net/2268%2F22808">2268/22808</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/21628162">21628162</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=American+Journal+of+Botany&rft.atitle=A+new+lineage+of+lichenized+basidiomycetes+inferred+from+a+two-gene+phylogeny%3A+The+Lepidostromataceae+with+three+species+from+the+tropics&rft.volume=95&rft.issue=12&rft.pages=1548-1556&rft.date=2008&rft_id=info%3Ahdl%2F2268%2F22808&rft_id=info%3Apmid%2F21628162&rft_id=info%3Adoi%2F10.3732%2Fajb.0800232&rft.aulast=Ertz&rft.aufirst=Damien&rft.au=Lawrey%2C+James+D.&rft.au=Sikaroodi%2C+Masoumeh&rft.au=Gillevet%2C+Patrick+M.&rft.au=Fischer%2C+Eberhard&rft.au=Killmann%2C+Dorothee&rft.au=S%C3%A9rusiaux%2C+Emmanu%C3%ABl&rft_id=http%3A%2F%2Forbi.ulg.ac.be%2Fhandle%2F2268%2F22808&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"></li> <li><cite id="CITEREFEkmanAndersenWedin2008" class="citation journal cs1">Ekman, Stefan; Andersen, Heidi L.; Wedin, Mats (2008). "The limitations of ancestral state reconstruction and the evolution of the ascus in the Lecanorales (lichenized Ascomycota)". Systematic Biology. 57 (1): 141–156. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1080%2F10635150801910451">10.1080/10635150801910451</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/18300027">18300027</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Systematic+Biology&rft.atitle=The+limitations+of+ancestral+state+reconstruction+and+the+evolution+of+the+ascus+in+the+Lecanorales+%28lichenized+Ascomycota%29&rft.volume=57&rft.issue=1&rft.pages=141-156&rft.date=2008&rft_id=info%3Adoi%2F10.1080%2F10635150801910451&rft_id=info%3Apmid%2F18300027&rft.aulast=Ekman&rft.aufirst=Stefan&rft.au=Andersen%2C+Heidi+L.&rft.au=Wedin%2C+Mats&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"></li> </ul> </div></td><td class="navbox-list" style="border-left:2px solid #fdfdfd;padding:0px;text-align: left;width:50%;"><div> <ul> <li><cite id="CITEREFEkman2001" class="citation journal cs1">Ekman, Stefan (2001). "Molecular phylogeny of the Bacidiaceae (Lecanorales, lichenized Ascomycota)". Mycological Research. 105 (7): 783–797. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1017%2FS0953756201004269">10.1017/S0953756201004269</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Mycological+Research&rft.atitle=Molecular+phylogeny+of+the+Bacidiaceae+%28Lecanorales%2C+lichenized+Ascomycota%29&rft.volume=105&rft.issue=7&rft.pages=783-797&rft.date=2001&rft_id=info%3Adoi%2F10.1017%2FS0953756201004269&rft.aulast=Ekman&rft.aufirst=Stefan&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"></li> <li><cite id="CITEREFGrubeWinka2002" class="citation journal cs1">Grube, Martin; Winka, Katarina (2002). "Progress in understanding the evolution and classification of lichenized ascomycetes". Mycologist. 16 (2): 67–76. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1017%2FS0269915X02002069">10.1017/S0269915X02002069</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Mycologist&rft.atitle=Progress+in+understanding+the+evolution+and+classification+of+lichenized+ascomycetes&rft.volume=16&rft.issue=2&rft.pages=67-76&rft.date=2002&rft_id=info%3Adoi%2F10.1017%2FS0269915X02002069&rft.aulast=Grube&rft.aufirst=Martin&rft.au=Winka%2C+Katarina&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"></li> <li><cite id="CITEREFLiuHall2004" class="citation journal cs1">Liu, Yajuan J.; Hall, Benjamin D. (2004). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC384777">"Body plan evolution of ascomycetes, as inferred from an RNA polymerase II phylogeny"</a>. <a href="/wiki/Proceedings_of_the_National_Academy_of_Sciences" class="mw-redirect" title="Proceedings of the National Academy of Sciences">PNAS</a>. 101 (13): 4507–4512. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2004PNAS..101.4507L">2004PNAS..101.4507L</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1073%2Fpnas.0400938101">10.1073/pnas.0400938101</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC384777">384777</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/15070748">15070748</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=PNAS&rft.atitle=Body+plan+evolution+of+ascomycetes%2C+as+inferred+from+an+RNA+polymerase+II+phylogeny&rft.volume=101&rft.issue=13&rft.pages=4507-4512&rft.date=2004&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC384777%23id-name%3DPMC&rft_id=info%3Apmid%2F15070748&rft_id=info%3Adoi%2F10.1073%2Fpnas.0400938101&rft_id=info%3Abibcode%2F2004PNAS..101.4507L&rft.aulast=Liu&rft.aufirst=Yajuan+J.&rft.au=Hall%2C+Benjamin+D.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC384777&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"></li> <li><cite id="CITEREFSchmittYamamotoLumbsch2006" class="citation journal cs1">Schmitt, I.; Yamamoto, Y.; Lumbsch, H. T. (2006). "Phylogeny of Pertusariales (Ascomycotina): Resurrection of Ochrolechiaceae and new circumscription of Megasporaceae". Journal of the Hattori Botanical Laboratory. 100: 753–764.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Journal+of+the+Hattori+Botanical+Laboratory&rft.atitle=Phylogeny+of+Pertusariales+%28Ascomycotina%29%3A+Resurrection+of+Ochrolechiaceae+and+new+circumscription+of+Megasporaceae&rft.volume=100&rft.pages=753-764&rft.date=2006&rft.aulast=Schmitt&rft.aufirst=I.&rft.au=Yamamoto%2C+Y.&rft.au=Lumbsch%2C+H.+T.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"></li> <li><cite id="CITEREFStaigerKalbGrube2006" class="citation journal cs1">Staiger, Bettina; Kalb, Klaus; Grube, Martin (2006). "Phylogeny and phenotypic variation in the lichen family Graphidaceae (Ostropomycetidae, Ascomycota)". Mycological Research. 110 (7): 765–772. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.mycres.2006.05.003">10.1016/j.mycres.2006.05.003</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/16876697">16876697</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Mycological+Research&rft.atitle=Phylogeny+and+phenotypic+variation+in+the+lichen+family+Graphidaceae+%28Ostropomycetidae%2C+Ascomycota%29&rft.volume=110&rft.issue=7&rft.pages=765-772&rft.date=2006&rft_id=info%3Adoi%2F10.1016%2Fj.mycres.2006.05.003&rft_id=info%3Apmid%2F16876697&rft.aulast=Staiger&rft.aufirst=Bettina&rft.au=Kalb%2C+Klaus&rft.au=Grube%2C+Martin&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"></li> </ul> </div></td></tr></tbody></table></div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="mw-heading mw-heading3"><h3 id="Photobionts">Photobionts</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=21" title="Edit section: Photobionts">edit</a>]</div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Bipartite_and_tripartite_cyanolichens_(10.3897-mycokeys.6.3869)_Figure_1_lowerleft.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/9/90/Bipartite_and_tripartite_cyanolichens_%2810.3897-mycokeys.6.3869%29_Figure_1_lowerleft.jpg/220px-Bipartite_and_tripartite_cyanolichens_%2810.3897-mycokeys.6.3869%29_Figure_1_lowerleft.jpg" decoding="async" width="220" height="152" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/90/Bipartite_and_tripartite_cyanolichens_%2810.3897-mycokeys.6.3869%29_Figure_1_lowerleft.jpg/330px-Bipartite_and_tripartite_cyanolichens_%2810.3897-mycokeys.6.3869%29_Figure_1_lowerleft.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/90/Bipartite_and_tripartite_cyanolichens_%2810.3897-mycokeys.6.3869%29_Figure_1_lowerleft.jpg/440px-Bipartite_and_tripartite_cyanolichens_%2810.3897-mycokeys.6.3869%29_Figure_1_lowerleft.jpg 2x" data-file-width="756" data-file-height="524" /></a><figcaption>Photobionts</figcaption></figure> The <a href="/wiki/Photosynthetic" class="mw-redirect" title="Photosynthetic">photosynthetic</a> partner in a lichen is called a photobiont. The photobionts in lichens come from a variety of simple <a href="/wiki/Prokaryotic" class="mw-redirect" title="Prokaryotic">prokaryotic</a> and <a href="/wiki/Eukaryotic" class="mw-redirect" title="Eukaryotic">eukaryotic</a> organisms. In the majority of lichens the photobiont is a green alga (<a href="/wiki/Chlorophyta" title="Chlorophyta">Chlorophyta</a>) or a <a href="/wiki/Cyanobacteria" title="Cyanobacteria">cyanobacterium</a>. In some lichens both types are present; in such cases, the alga is typically the primary partner, with the cyanobacteria being located in cryptic pockets.<a href="#cite_note-95">[95]</a> Algal photobionts are called phycobionts, while cyanobacterial photobionts are called cyanobionts.<a href="#cite_note-UNOLP-44">[44]</a> About 90% of all known lichens have phycobionts, and about 10% have cyanobionts.<a href="#cite_note-UNOLP-44">[44]</a> Approximately 100 species of photosynthetic partners from 40<a href="#cite_note-UNOLP-44">[44]</a> genera and five distinct classes (prokaryotic: <a href="/wiki/Cyanophyceae" class="mw-redirect" title="Cyanophyceae">Cyanophyceae</a>; eukaryotic: <a href="/wiki/Trebouxiophyceae" title="Trebouxiophyceae">Trebouxiophyceae</a>, <a href="/wiki/Phaeophyceae" class="mw-redirect" title="Phaeophyceae">Phaeophyceae</a>, <a href="/wiki/Chlorophyceae" title="Chlorophyceae">Chlorophyceae</a>) have been found to associate with the lichen-forming fungi.<a href="#cite_note-Friedl1996-96">[96]</a> Common <a href="/wiki/Algal" class="mw-redirect" title="Algal">algal</a> photobionts are from the genera <a href="/wiki/Trebouxia" title="Trebouxia">Trebouxia</a>, <a href="/wiki/Trentepohlia_(alga)" title="Trentepohlia (alga)">Trentepohlia</a>, <a href="/w/index.php?title=Pseudotrebouxia&action=edit&redlink=1" class="new" title="Pseudotrebouxia (page does not exist)">Pseudotrebouxia</a>, or <a href="/wiki/Myrmecia_(alga)" title="Myrmecia (alga)">Myrmecia</a>. Trebouxia is the most common genus of green algae in lichens, occurring in about 40% of all lichens. "Trebouxioid" means either a photobiont that is in the genus Trebouxia, or resembles a member of that genus, and is therefore presumably a member of the class <a href="/wiki/Trebouxiophyceae" title="Trebouxiophyceae">Trebouxiophyceae</a>.<a href="#cite_note-ASLGPZ-36">[36]</a> The second most commonly represented green alga genus is <a href="/wiki/Trentepohlia_(alga)" title="Trentepohlia (alga)">Trentepohlia</a>.<a href="#cite_note-Dobson-37">[37]</a> Overall, about 100 species of eukaryotes are known to occur as photobionts in lichens. All the algae are probably able to exist independently in nature as well as in the lichen.<a href="#cite_note-Rikkinen1995-94">[94]</a> A "<a href="/wiki/Cyanolichen" title="Cyanolichen">cyanolichen</a>" is a lichen with a <a href="/wiki/Cyanobacteria" title="Cyanobacteria">cyanobacterium</a> as its main photosynthetic component (photobiont).<a href="#cite_note-ASLGAF-97">[97]</a> Most cyanolichen are also ascolichens, but a few basidiolichen like <a href="/wiki/Dictyonema" title="Dictyonema">Dictyonema</a> and <a href="/wiki/Acantholichen" title="Acantholichen">Acantholichen</a> have cyanobacteria as their partner.<a href="#cite_note-98">[98]</a> The most commonly occurring cyanobacterium <a href="/wiki/Genus" title="Genus">genus</a> is <a href="/wiki/Nostoc" title="Nostoc">Nostoc</a>.<a href="#cite_note-Rikkinen1995-94">[94]</a> Other<a href="#cite_note-Dobson-37">[37]</a> common <a href="/wiki/Cyanobacterium" class="mw-redirect" title="Cyanobacterium">cyanobacterium</a> photobionts are from <a href="/wiki/Scytonema" title="Scytonema">Scytonema</a>.<a href="#cite_note-UCLS-18">[18]</a> Many cyanolichens are small and black, and have <a href="/wiki/Limestone" title="Limestone">limestone</a> as the substrate.[<a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed">citation needed</a>] Another cyanolichen group, the <a href="/wiki/Jelly_lichen" class="mw-redirect" title="Jelly lichen">jelly lichens</a> of the genera <a href="/wiki/Collema" title="Collema">Collema</a> or <a href="/wiki/Leptogium" title="Leptogium">Leptogium</a> are gelatinous and live on moist soils. Another group of large and <a href="/wiki/Foliose" class="mw-redirect" title="Foliose">foliose</a> species including <a href="/wiki/Peltigera" title="Peltigera">Peltigera</a>, <a href="/wiki/Lobaria" title="Lobaria">Lobaria</a>, and <a href="/wiki/Degelia" title="Degelia">Degelia</a> are grey-blue, especially when dampened or wet. Many of these characterize the <a href="/w/index.php?title=Lobarion&action=edit&redlink=1" class="new" title="Lobarion (page does not exist)">Lobarion</a> communities of higher rainfall areas in western Britain, e.g., in the <a href="/wiki/Celtic_rain_forest" class="mw-redirect" title="Celtic rain forest">Celtic rain forest</a>. Strains of cyanobacteria found in various cyanolichens are often closely related to one another.<a href="#cite_note-SM-99">[99]</a> They differ from the most closely related free-living strains.<a href="#cite_note-SM-99">[99]</a> The lichen association is a close symbiosis. It extends the ecological range of both partners but is not always obligatory for their growth and reproduction in natural environments, since many of the algal symbionts can live independently. A prominent example is the alga <a href="/wiki/Trentepohlia_(alga)" title="Trentepohlia (alga)">Trentepohlia</a>, which forms orange-coloured populations on tree trunks and suitable rock faces. Lichen propagules (<a href="/wiki/Diaspore_(botany)" title="Diaspore (botany)">diaspores</a>) typically contain cells from both partners, although the fungal components of so-called "fringe species" rely instead on algal cells dispersed by the "core species".<a href="#cite_note-RaiBergman2002-77">[77]</a> The same cyanobiont species can occur in association with different fungal species as lichen partners.<a href="#cite_note-100">[100]</a> The same phycobiont species can occur in association with different fungal species as lichen partners.<a href="#cite_note-UNOLP-44">[44]</a> More than one phycobiont may be present in a single thallus.<a href="#cite_note-UNOLP-44">[44]</a> A single lichen may contain several algal <a href="/wiki/Genotype" title="Genotype">genotypes</a>.<a href="#cite_note-101">[101]</a><a href="#cite_note-102">[102]</a> These multiple genotypes may better enable response to adaptation to environmental changes, and enable the lichen to inhabit a wider range of environments.<a href="#cite_note-103">[103]</a> <div class="mw-heading mw-heading3"><h3 id="Controversy_over_classification_method_and_species_names">Controversy over classification method and species names</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=22" title="Edit section: Controversy over classification method and species names">edit</a>]</div> There are about 20,000 known lichen <a href="/wiki/Species_(biology)" class="mw-redirect" title="Species (biology)">species</a>.<a href="#cite_note-UCLS-18">[18]</a> But what is meant by "species" is different from what is meant by biological species in plants, animals, or fungi, where being the same species implies that there is a common <a href="/wiki/Ancestral_lineage" class="mw-redirect" title="Ancestral lineage">ancestral lineage</a>.<a href="#cite_note-UCLS-18">[18]</a> Because lichens are combinations of members of two or even three different biological <a href="/wiki/Kingdom_(biology)" title="Kingdom (biology)">kingdoms</a>, these components must have a different ancestral lineage from each other. By convention, lichens are still called "species" anyway, and are classified according to the species of their fungus, not the species of the algae or cyanobacteria. Lichens are given the same scientific name (<a href="/wiki/Binomial_name" class="mw-redirect" title="Binomial name">binomial name</a>) as the fungus in them, which may cause some confusion. The alga bears its own scientific name, which has no relationship to the name of the lichen or fungus.<a href="#cite_note-Kirk_pp.378-81-82">[82]</a> Depending on context, "lichenized fungus" may refer to the entire lichen, or to the fungus when it is in the lichen, which can be grown in culture in isolation from the algae or cyanobacteria. Some algae and cyanobacteria are found naturally living outside of the lichen. The fungal, algal, or cyanobacterial component of a lichen can be grown by itself in culture. When growing by themselves, the fungus, algae, or cyanobacteria have very different properties than those of the lichen. Lichen properties such as growth form, physiology, and biochemistry, are very different from the combination of the properties of the fungus and the algae or cyanobacteria. The same fungus growing in combination with different algae or cyanobacteria, can produce lichens that are very different in most properties, meeting non-DNA criteria for being different "species". Historically, these different combinations were classified as different species. When the fungus is identified as being the same using modern DNA methods, these apparently different species get reclassified as the same species under the current (2014) convention for classification by fungal component. This has led to debate about this classification convention. These apparently different "species" have their own independent evolutionary history.<a href="#cite_note-WIL-4">[4]</a><a href="#cite_note-FSSD-86">[86]</a> There is also debate as to the appropriateness of giving the same binomial name to the fungus, and to the lichen that combines that fungus with an alga or cyanobacterium (<a href="/wiki/Synecdoche" title="Synecdoche">synecdoche</a>). This is especially the case when combining the same fungus with different algae or cyanobacteria produces dramatically different lichen organisms, which would be considered different species by any measure other than the DNA of the fungal component. If the whole lichen produced by the same fungus growing in association with different algae or cyanobacteria, were to be classified as different "species", the number of "lichen species" would be greater. <div class="mw-heading mw-heading3"><h3 id="Diversity">Diversity</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=23" title="Edit section: Diversity">edit</a>]</div> The largest number of lichenized fungi occur in the <a href="/wiki/Ascomycota" title="Ascomycota">Ascomycota</a>, with about 40% of species forming such an association.<a href="#cite_note-Kirk_pp.378-81-82">[82]</a> Some of these lichenized fungi occur in orders with nonlichenized fungi that live as <a href="/wiki/Saprotroph" class="mw-redirect" title="Saprotroph">saprotrophs</a> or <a href="/wiki/Parasite" class="mw-redirect" title="Parasite">plant parasites</a> (for example, the <a href="/wiki/Leotiales" title="Leotiales">Leotiales</a>, <a href="/wiki/Dothideales" title="Dothideales">Dothideales</a>, and <a href="/wiki/Pezizales" title="Pezizales">Pezizales</a>). Other lichen fungi occur in only five <a href="/wiki/Order_(biology)" title="Order (biology)">orders</a> in which all members are engaged in this habit (Orders <a href="/wiki/Graphidales" title="Graphidales">Graphidales</a>, <a href="/wiki/Gyalectales" title="Gyalectales">Gyalectales</a>, <a href="/wiki/Peltigerales" title="Peltigerales">Peltigerales</a>, <a href="/wiki/Pertusariales" title="Pertusariales">Pertusariales</a>, and <a href="/wiki/Teloschistales" title="Teloschistales">Teloschistales</a>). Overall, about 98% of lichens have an ascomycetous mycobiont.<a href="#cite_note-Lutzoni_Pagel_Reeb_pp._937–940-104">[104]</a> Next to the Ascomycota, the largest number of lichenized fungi occur in the unassigned <a href="/wiki/Fungi_imperfecti" title="Fungi imperfecti">fungi imperfecti</a>, a catch-all category for fungi whose sexual form of reproduction has never been observed.[<a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed">citation needed</a>] Comparatively few <a href="/wiki/Basidiomycetes" class="mw-redirect" title="Basidiomycetes">basidiomycetes</a> are lichenized, but these include <a href="/wiki/Agarics" class="mw-redirect" title="Agarics">agarics</a>, such as species of <a href="/wiki/Lichenomphalia" title="Lichenomphalia">Lichenomphalia</a>, <a href="/wiki/Clavarioid_fungi" title="Clavarioid fungi">clavarioid fungi</a>, such as species of <a href="/wiki/Multiclavula" title="Multiclavula">Multiclavula</a>, and <a href="/wiki/Corticioid_fungi" title="Corticioid fungi">corticioid fungi</a>, such as species of <a href="/wiki/Dictyonema" title="Dictyonema">Dictyonema</a>. <div class="mw-heading mw-heading3"><h3 id="Identification_methods">Identification methods</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=24" title="Edit section: Identification methods">edit</a>]</div> Lichen identification uses growth form, microscopy and reactions to chemical tests. The outcome of the "Pd test" is called "Pd", which is also used as an abbreviation for the chemical used in the test, <a href="/wiki/Para-phenylenediamine" class="mw-redirect" title="Para-phenylenediamine">para-phenylenediamine</a>.<a href="#cite_note-ASLGPZ-36">[36]</a> If putting a drop on a lichen turns an area bright yellow to orange, this helps identify it as belonging to either the genus <a href="/wiki/Cladonia" title="Cladonia">Cladonia</a> or <a href="/wiki/Lecanora" title="Lecanora">Lecanora</a>.<a href="#cite_note-ASLGPZ-36">[36]</a> <div class="mw-heading mw-heading3"><h3 id="Evolution_and_paleontology">Evolution and paleontology</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=25" title="Edit section: Evolution and paleontology">edit</a>]</div> The fossil record for lichens is poor.<a href="#cite_note-UCMPLFR-105">[105]</a> The extreme habitats that lichens dominate, such as tundra, mountains, and deserts, are not ordinarily conducive to producing fossils.<a href="#cite_note-UCMPLFR-105">[105]</a><a href="#cite_note-urlFossil_Record_of_Lichens-106">[106]</a> There are fossilized lichens embedded in amber. The fossilized Anzia is found in pieces of amber in northern Europe and dates back approximately 40 million years.<a href="#cite_note-Poinar1992-107">[107]</a> Lichen fragments are also found in fossil leaf beds, such as <a href="/wiki/Lobaria" title="Lobaria">Lobaria</a> from Trinity County in northern California, US, dating back to the early to middle <a href="/wiki/Miocene" title="Miocene">Miocene</a>.<a href="#cite_note-Peterson2000-108">[108]</a> The oldest fossil lichen in which both symbiotic partners have been recovered is <a href="/wiki/Winfrenatia" title="Winfrenatia">Winfrenatia</a>, an early zygomycetous (<a href="/wiki/Glomeromycota" title="Glomeromycota">Glomeromycotan</a>) lichen symbiosis that may have involved controlled parasitism,[<a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed">citation needed</a>] is permineralized in the <a href="/wiki/Rhynie_Chert" class="mw-redirect" title="Rhynie Chert">Rhynie Chert</a> of Scotland, dating from early <a href="/wiki/Devonian" title="Devonian">Early Devonian</a>, about 400 million years ago.<a href="#cite_note-Taylor1995-109">[109]</a> The slightly older fossil <a href="/wiki/Spongiophyton" title="Spongiophyton">Spongiophyton</a> has also been interpreted as a lichen on morphological<a href="#cite_note-Taylor2004-110">[110]</a> and isotopic<a href="#cite_note-Jahren2003-111">[111]</a> grounds, although the isotopic basis is decidedly shaky.<a href="#cite_note-Fletcher2004-112">[112]</a> It has been demonstrated that <a href="/wiki/Silurian" title="Silurian">Silurian</a>-<a href="/wiki/Devonian" title="Devonian">Devonian</a> fossils <a href="/wiki/Nematothallus" title="Nematothallus">Nematothallus</a><a href="#cite_note-113">[113]</a> and <a href="/wiki/Prototaxites" title="Prototaxites">Prototaxites</a><a href="#cite_note-114">[114]</a> were lichenized. Thus lichenized <a href="/wiki/Ascomycota" title="Ascomycota">Ascomycota</a> and <a href="/wiki/Basidiomycota" title="Basidiomycota">Basidiomycota</a> were a component of <a href="/wiki/Llandovery_epoch" class="mw-redirect" title="Llandovery epoch">Early Silurian</a>-<a href="/wiki/Devonian" title="Devonian">Devonian</a> terrestrial ecosystems.<a href="#cite_note-Karatygin2009-115">[115]</a><a href="#cite_note-Karatygin2007-116">[116]</a> Newer research suggests that lichen evolved after the evolution of land plants.<a href="#cite_note-117">[117]</a> The ancestral ecological state of both <a href="/wiki/Ascomycota" title="Ascomycota">Ascomycota</a> and <a href="/wiki/Basidiomycota" title="Basidiomycota">Basidiomycota</a> was probably <a href="/wiki/Saprobe" class="mw-redirect" title="Saprobe">saprobism</a>, and independent lichenization events may have occurred multiple times.<a href="#cite_note-118">[118]</a><a href="#cite_note-Schoch2009-119">[119]</a> In 1995, Gargas and colleagues proposed that there were at least five independent origins of lichenization; three in the basidiomycetes and at least two in the Ascomycetes.<a href="#cite_note-Gargas1995-120">[120]</a> Lutzoni et al. (2001) suggest lichenization probably evolved earlier and was followed by multiple independent losses. Some non-lichen-forming fungi may have secondarily lost the ability to form a lichen association. As a result, lichenization has been viewed as a highly successful nutritional strategy.<a href="#cite_note-Honegger1998-121">[121]</a><a href="#cite_note-Wedin2004-122">[122]</a> Lichenized <a href="/wiki/Glomeromycota" title="Glomeromycota">Glomeromycota</a> may extend well back into the Precambrian. Lichen-like fossils consisting of coccoid cells (<a href="/wiki/Cyanobacteria" title="Cyanobacteria">cyanobacteria</a>?) and thin filaments (mucoromycotinan <a href="/wiki/Glomeromycota" title="Glomeromycota">Glomeromycota</a>?) are permineralized in marine <a href="/wiki/Phosphorite" title="Phosphorite">phosphorite</a> of the <a href="/wiki/Doushantuo_Formation" title="Doushantuo Formation">Doushantuo Formation</a> in southern China. These fossils are thought to be 551 to 635 million years old or <a href="/wiki/Ediacaran" title="Ediacaran">Ediacaran</a>.<a href="#cite_note-Yuan2005-123">[123]</a> Ediacaran <a href="/wiki/Acritarchs" class="mw-redirect" title="Acritarchs">acritarchs</a> also have many similarities with <a href="/wiki/Glomeromycota" title="Glomeromycota">Glomeromycotan</a> vesicles and spores.<a href="#cite_note-124">[124]</a> It has also been claimed that <a href="/wiki/Ediacaran_biota" title="Ediacaran biota">Ediacaran fossils</a> including <a href="/wiki/Dickinsonia" title="Dickinsonia">Dickinsonia</a>,<a href="#cite_note-Retallack2007-125">[125]</a> were lichens,<a href="#cite_note-Retallack1994-126">[126]</a> although this claim is controversial.<a href="#cite_note-Switek2012-127">[127]</a> Endosymbiotic <a href="/wiki/Glomeromycota" title="Glomeromycota">Glomeromycota</a> comparable with living <a href="/wiki/Geosiphon" title="Geosiphon">Geosiphon</a> may extend back into the <a href="/wiki/Proterozoic" title="Proterozoic">Proterozoic</a> in the form of 1500 million year old <a href="/wiki/Horodyskia" title="Horodyskia">Horodyskia</a><a href="#cite_note-128">[128]</a> and 2200 million year old <a href="/wiki/Diskagma" title="Diskagma">Diskagma</a>.<a href="#cite_note-Retallack-2013-129">[129]</a> Discovery of these fossils suggest that fungi developed symbiotic partnerships with photoautotrophs long before the evolution of vascular plants, though the Ediacaran lichen hypothesis is largely rejected due to an inappropriate definition of lichens based on taphonomy and substrate ecology.<a href="#cite_note-130">[130]</a> However, a 2019 study by the same scientist who rejected the Ediacaran lichen hypothesis, Nelsen, used new time-calibrated phylogenies to conclude that there is no evidence of lichen before the existence of vascular plants.<a href="#cite_note-131">[131]</a> Lecanoromycetes, one of the most common classes of lichen-forming fungi, diverged from its ancestor, which may have also been lichen forming, around 258 million years ago, during the late Paleozoic period. However, the closely related clade Euritiomycetes appears to have become lichen-forming only 52 million years ago, during the early Cenozoic period.<a href="#cite_note-132">[132]</a> <div class="mw-heading mw-heading2"><h2 id="Ecology_and_interactions_with_environment">Ecology and interactions with environment</h2>[<a href="/w/index.php?title=Lichen&action=edit&section=26" title="Edit section: Ecology and interactions with environment">edit</a>]</div> <div class="mw-heading mw-heading3"><h3 id="Substrates_and_habitats">Substrates and habitats</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=27" title="Edit section: Substrates and habitats">edit</a>]</div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Der_Heilige_Dominikus_am_Nordturm_des_Regensburger_Doms.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/d3/Der_Heilige_Dominikus_am_Nordturm_des_Regensburger_Doms.jpg/170px-Der_Heilige_Dominikus_am_Nordturm_des_Regensburger_Doms.jpg" decoding="async" width="170" height="227" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/d3/Der_Heilige_Dominikus_am_Nordturm_des_Regensburger_Doms.jpg/255px-Der_Heilige_Dominikus_am_Nordturm_des_Regensburger_Doms.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/d3/Der_Heilige_Dominikus_am_Nordturm_des_Regensburger_Doms.jpg/340px-Der_Heilige_Dominikus_am_Nordturm_des_Regensburger_Doms.jpg 2x" data-file-width="1500" data-file-height="2000" /></a><figcaption>Lichens on a limestone statue on a tower of <a href="/wiki/Regensburg_Cathedral" title="Regensburg Cathedral">Regensburg Cathedral</a></figcaption></figure> Lichens grow on and in a wide range of substrates and habitats, including some of the most extreme conditions on earth.<a href="#cite_note-133">[133]</a> They are abundant growing on bark, leaves, and hanging from <a href="/wiki/Epiphyte" title="Epiphyte">epiphyte</a> branches in <a href="/wiki/Rain_forest" class="mw-redirect" title="Rain forest">rain forests</a> and in <a href="/wiki/Temperate_woodland" class="mw-redirect" title="Temperate woodland">temperate woodland</a>. They grow on bare rock, walls, gravestones, roofs, and exposed soil surfaces. They can survive in some of the most extreme environments on Earth: <a href="/wiki/Arctic_tundra" class="mw-redirect" title="Arctic tundra">arctic tundra</a>, hot dry <a href="/wiki/Desert" title="Desert">deserts</a>, rocky coasts, and toxic <a href="/wiki/Slag_heap" class="mw-redirect" title="Slag heap">slag heaps</a>. They can live inside solid rock, growing between the grains, and in the soil as part of a <a href="/wiki/Biological_soil_crust" title="Biological soil crust">biological soil crust</a> in arid habitats such as deserts. Some lichens do not grow on anything, living out their lives blowing about the environment.<a href="#cite_note-WIL-4">[4]</a> When growing on mineral surfaces, some lichens slowly decompose their substrate by chemically degrading and physically disrupting the minerals, contributing to the process of <a href="/wiki/Weathering" title="Weathering">weathering</a> by which rocks are gradually turned into soil. While this contribution to weathering is usually benign, it can cause problems for artificial stone structures. For example, there is an ongoing lichen growth problem on <a href="/wiki/Mount_Rushmore_National_Memorial" class="mw-redirect" title="Mount Rushmore National Memorial">Mount Rushmore National Memorial</a> that requires the employment of mountain-climbing conservators to clean the monument.<a href="#cite_note-WaPo11Jul2005-134">[134]</a> Lichens are not <a href="/wiki/Parasites" class="mw-redirect" title="Parasites">parasites</a> on the plants they grow on, but only use them as a substrate. The fungi of some lichen species may "take over" the algae of other lichen species.<a href="#cite_note-LLHE-16">[16]</a><a href="#cite_note-ABGPPU-135">[135]</a> Lichens make their own food from their photosynthetic parts and by absorbing minerals from the environment.<a href="#cite_note-LLHE-16">[16]</a> Lichens growing on leaves may have the appearance of being parasites on the leaves, but they are not. Some lichens in Diploschistes parasitise other lichens. Diploschistes muscorum starts its development in the tissue of a host Cladonia species.<a href="#cite_note-Honegger1988-55">[55]</a>: 30 <a href="#cite_note-Dobson-37">[37]</a>: 171  <figure class="mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Lichen_A_01.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8e/Lichen_A_01.jpg/165px-Lichen_A_01.jpg" decoding="async" width="165" height="124" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8e/Lichen_A_01.jpg/248px-Lichen_A_01.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8e/Lichen_A_01.jpg/331px-Lichen_A_01.jpg 2x" data-file-width="5712" data-file-height="4284" /></a><figcaption> Lichen on a fallen branch</figcaption></figure> In the arctic tundra, lichens, together with <a href="/wiki/Mosses" class="mw-redirect" title="Mosses">mosses</a> and <a href="/wiki/Liverworts" class="mw-redirect" title="Liverworts">liverworts</a>, make up the majority of the <a href="/wiki/Ground_cover" class="mw-redirect" title="Ground cover">ground cover</a>, which helps insulate the ground and may provide forage for grazing animals. An example is "<a href="/wiki/Reindeer_moss" class="mw-redirect" title="Reindeer moss">reindeer moss</a>", which is a lichen, not a moss.<a href="#cite_note-LLHE-16">[16]</a> There are only two species of known permanently submerged lichens; <a href="/w/index.php?title=Hydrothyria_venosa&action=edit&redlink=1" class="new" title="Hydrothyria venosa (page does not exist)">Hydrothyria venosa</a> is found in fresh water environments, and <a href="/wiki/Verrucaria_serpuloides" title="Verrucaria serpuloides">Verrucaria serpuloides</a> is found in marine environments.<a href="#cite_note-136">[136]</a> A crustose lichen that grows on rock is called a <a href="/wiki/Saxicolous_lichen" title="Saxicolous lichen">saxicolous lichen</a>.<a href="#cite_note-ASLGPZ-36">[36]</a><a href="#cite_note-VMBMLF-39">[39]</a>: 159  Crustose lichens that grow on the rock are <a href="/wiki/Epilithic" class="mw-redirect" title="Epilithic">epilithic</a>, and those that grow immersed inside rock, growing between the crystals with only their fruiting bodies exposed to the air, are called <a href="/wiki/Endolithic_lichen" title="Endolithic lichen">endolithic lichens</a>.<a href="#cite_note-LNAILV-35">[35]</a><a href="#cite_note-VMBMLF-39">[39]</a>: 159 <a href="#cite_note-ASLGAF-97">[97]</a> A crustose lichen that grows on bark is called a <a href="/wiki/Corticolous_lichen" title="Corticolous lichen">corticolous lichen</a>.<a href="#cite_note-VMBMLF-39">[39]</a>: 159  A lichen that grows on wood from which the bark has been stripped is called a <a href="/wiki/Lignicolous_lichen" title="Lignicolous lichen">lignicolous lichen</a>.<a href="#cite_note-ASLGGO-45">[45]</a> Lichens that grow immersed inside plant tissues are called <a href="/w/index.php?title=Endophloidic_lichen&action=edit&redlink=1" class="new" title="Endophloidic lichen (page does not exist)">endophloidic lichens</a> or <a href="/w/index.php?title=Endophloidal_lichen&action=edit&redlink=1" class="new" title="Endophloidal lichen (page does not exist)">endophloidal lichens</a>.<a href="#cite_note-LNAILV-35">[35]</a><a href="#cite_note-VMBMLF-39">[39]</a>: 159  Lichens that use leaves as substrates, whether the leaf is still on the tree or on the ground, are called <a href="/wiki/Epiphyllous_lichen" class="mw-redirect" title="Epiphyllous lichen">epiphyllous</a> or <a href="/wiki/Foliicolous_lichen" title="Foliicolous lichen">foliicolous</a>.<a href="#cite_note-UNOLP-44">[44]</a> A <a href="/wiki/Terricolous_lichen" title="Terricolous lichen">terricolous lichen</a> grows on the soil as a substrate. Many squamulose lichens are terricolous.<a href="#cite_note-VMBMLF-39">[39]</a>: 159  <a href="/wiki/Umbilicate_lichen" title="Umbilicate lichen">Umbilicate lichens</a> are foliose lichens that are attached to the substrate at only one point.<a href="#cite_note-LNAILV-35">[35]</a> A <a href="/wiki/Vagrant_lichen" title="Vagrant lichen">vagrant lichen</a> is not attached to a substrate at all, and lives its life being blown around by the wind. <div class="mw-heading mw-heading3"><h3 id="Lichens_and_soils">Lichens and soils</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=28" title="Edit section: Lichens and soils">edit</a>]</div> In addition to distinct physical mechanisms by which lichens break down raw stone, studies indicate lichens attack stone chemically, entering newly chelated minerals into the ecology. The substances exuded by lichens, known for their strong ability to bind and sequester metals, along with the common formation of new minerals, especially metal <a href="/wiki/Oxalate" title="Oxalate">oxalates</a>, and the traits of the substrates they alter, all highlight the important role lichens play in the process of chemical <a href="/wiki/Weathering" title="Weathering">weathering</a>.<a href="#cite_note-catena-137">[137]</a> Over time, this activity creates new fertile soil from stone. Lichens may be <a href="/wiki/Lichens_and_nitrogen_cycling" title="Lichens and nitrogen cycling">important in contributing nitrogen</a> to soils in some deserts through being eaten, along with their rock substrate, by snails, which then defecate, putting the nitrogen into the soils.<a href="#cite_note-138">[138]</a> Lichens help bind and stabilize soil sand in dunes.<a href="#cite_note-WIL-4">[4]</a> In deserts and semi-arid areas, lichens are part of extensive, living <a href="/wiki/Biological_soil_crust" title="Biological soil crust">biological soil crusts</a>, essential for maintaining the soil structure.<a href="#cite_note-WIL-4">[4]</a> <div class="mw-heading mw-heading3"><h3 id="Ecological_interactions">Ecological interactions</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=29" title="Edit section: Ecological interactions">edit</a>]</div> Lichens are <a href="/wiki/Pioneer_species" title="Pioneer species">pioneer species</a>, among the first living things to grow on bare rock or areas denuded of life by a disaster.<a href="#cite_note-WIL-4">[4]</a> Lichens may have to compete with plants for access to sunlight, but because of their small size and slow growth, they thrive in places where higher plants have difficulty growing. Lichens are often the <a href="/wiki/Ecological_succession" title="Ecological succession">first to settle</a> in places lacking soil, constituting the sole vegetation in some extreme environments such as those found at high mountain elevations and at high latitudes.<a href="#cite_note-139">[139]</a> Some survive in the tough conditions of deserts, and others on frozen soil of the Arctic regions.<a href="#cite_note-Oksanen2006-140">[140]</a> A major ecophysiological advantage of lichens is that they are <a href="/wiki/Poikilohydry" title="Poikilohydry">poikilohydric</a> (poikilo- variable, hydric- relating to water), meaning that though they have little control over the status of their hydration, they can tolerate irregular and extended periods of severe <a href="/wiki/Desiccation" title="Desiccation">desiccation</a>. Like some <a href="/wiki/Moss" title="Moss">mosses</a>, <a href="/wiki/Marchantiophyta" title="Marchantiophyta">liverworts</a>, <a href="/wiki/Fern" title="Fern">ferns</a> and a few <a href="/wiki/Resurrection_plant" title="Resurrection plant">resurrection plants</a>, upon desiccation, lichens enter a metabolic suspension or stasis (known as <a href="/wiki/Cryptobiosis" title="Cryptobiosis">cryptobiosis</a>) in which the cells of the lichen symbionts are dehydrated to a degree that halts most biochemical activity. In this cryptobiotic state, lichens can survive wider extremes of temperature, radiation and drought in the harsh environments they often inhabit. <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Moss_lichen.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Moss_lichen.jpg/220px-Moss_lichen.jpg" decoding="async" width="220" height="125" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/63/Moss_lichen.jpg/330px-Moss_lichen.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/63/Moss_lichen.jpg/440px-Moss_lichen.jpg 2x" data-file-width="1448" data-file-height="822" /></a><figcaption>Lichens suppress the growth of mosses and higher plants around them</figcaption></figure> Lichens do not have roots and do not need to tap continuous reservoirs of water like most higher plants, thus they can grow in locations impossible for most plants, such as bare rock, sterile soil or sand, and various artificial structures such as walls, roofs, and monuments. Many lichens also grow as <a href="/wiki/Epiphyte" title="Epiphyte">epiphytes</a> (epi- on the surface, phyte- plant) on plants, particularly on the trunks and branches of trees. When growing on plants, lichens are not <a href="/wiki/Parasite" class="mw-redirect" title="Parasite">parasites</a>; they do not consume any part of the plant nor poison it. Lichens produce <a href="/wiki/Allelopathy" title="Allelopathy">allelopathic</a> chemicals that inhibit the growth of mosses. Some ground-dwelling lichens, such as members of the subgenus <a href="/wiki/Cladoniaceae" title="Cladoniaceae">Cladina</a> (reindeer lichens), produce allelopathic chemicals that leach into the soil and inhibit the germination of seeds, spruce and other plants.<a href="#cite_note-141">[141]</a> Stability (that is, longevity) of their <a href="/wiki/Substrate_(biology)" title="Substrate (biology)">substrate</a> is a major factor of lichen habitats. Most lichens grow on stable rock surfaces or the bark of old trees, but many others grow on soil and sand. In these latter cases, lichens are often an important part of soil stabilization; indeed, in some desert ecosystems, <a href="/wiki/Vascular_plant" title="Vascular plant">vascular (higher) plant</a> seeds cannot become established except in places where lichen crusts stabilize the sand and help retain water. Lichens may be eaten by some animals, such as <a href="/wiki/Reindeer" title="Reindeer">reindeer</a>, living in <a href="/wiki/Arctic" title="Arctic">arctic</a> regions. The <a href="/wiki/Larva" title="Larva">larvae</a> of a number of <a href="/wiki/Lepidoptera" title="Lepidoptera">Lepidoptera</a> species feed exclusively on lichens. These include <a href="/wiki/Common_footman" title="Common footman">common footman</a> and <a href="/wiki/Marbled_beauty" title="Marbled beauty">marbled beauty</a>. They are very low in protein and high in carbohydrates, making them unsuitable for some animals. The <a href="/wiki/Northern_flying_squirrel" title="Northern flying squirrel">Northern flying squirrel</a> uses it for nesting, food and winter water. <div class="mw-heading mw-heading3"><h3 id="Effects_of_air_pollution">Effects of air pollution</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=30" title="Edit section: Effects of air pollution">edit</a>]</div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Lobaria_pulmonaria_010108c.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/da/Lobaria_pulmonaria_010108c.jpg/220px-Lobaria_pulmonaria_010108c.jpg" decoding="async" width="220" height="164" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/da/Lobaria_pulmonaria_010108c.jpg/330px-Lobaria_pulmonaria_010108c.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/da/Lobaria_pulmonaria_010108c.jpg/440px-Lobaria_pulmonaria_010108c.jpg 2x" data-file-width="2800" data-file-height="2085" /></a><figcaption>Some lichens, like the foliose <a href="/wiki/Lobaria_pulmonaria" title="Lobaria pulmonaria">Lobaria pulmonaria</a>, are sensitive to air pollution.</figcaption></figure> If lichens are exposed to air pollutants at all times, without any <a href="/wiki/Deciduous" title="Deciduous">deciduous</a> parts, they are unable to avoid the accumulation of pollutants. Also lacking <a href="/wiki/Stomata" class="mw-redirect" title="Stomata">stomata</a> and a <a href="/wiki/Plant_cuticle" title="Plant cuticle">cuticle</a>, lichens may absorb <a href="/wiki/Aerosol" title="Aerosol">aerosols</a> and gases over the entire thallus surface from which they may readily <a href="/wiki/Diffusion" title="Diffusion">diffuse</a> to the photobiont layer.<a href="#cite_note-Nash2008-142">[142]</a> Because lichens do not possess roots, their primary source of most <a href="/wiki/Chemical_element" title="Chemical element">elements</a> is the air, and therefore elemental levels in lichens often reflect the accumulated composition of ambient air. The processes by which atmospheric deposition occurs include <a href="/wiki/Fog" title="Fog">fog</a> and <a href="/wiki/Dew" title="Dew">dew</a>, gaseous absorption, and dry deposition.<a href="#cite_note-Knops1991-143">[143]</a> Consequently, environmental studies with lichens emphasize their feasibility as effective <a href="/wiki/Bioindicator" title="Bioindicator">biomonitors</a> of atmospheric quality.<a href="#cite_note-Nash2008-142">[142]</a> Not all lichens are equally sensitive to <a href="/wiki/Air_pollution" title="Air pollution">air pollutants</a>, so different lichen species show different levels of sensitivity to specific atmospheric pollutants.<a href="#cite_note-hogan-144">[144]</a> The sensitivity of a lichen to air pollution is directly related to the energy needs of the mycobiont, so that the stronger the dependency of the mycobiont on the photobiont, the more sensitive the lichen is to air pollution.<a href="#cite_note-Beltman1980-145">[145]</a> Upon exposure to air pollution, the photobiont may use metabolic energy for repair of its cellular structures that would otherwise be used for maintenance of its photosynthetic activity, therefore leaving less metabolic energy available for the mycobiont. The alteration of the balance between the photobiont and mycobiont can lead to the breakdown of the symbiotic association. Therefore, lichen decline may result not only from the accumulation of toxic substances, but also from altered nutrient supplies that favor one symbiont over the other.<a href="#cite_note-Nash2008-142">[142]</a> This interaction between lichens and air pollution has been used as a means of monitoring air quality since 1859, with more systematic methods developed by <a href="/wiki/William_Nylander_(botanist)" title="William Nylander (botanist)">William Nylander</a> in 1866.<a href="#cite_note-WIL-4">[4]</a> <div class="mw-heading mw-heading2"><h2 id="Human_use">Human use</h2>[<a href="/w/index.php?title=Lichen&action=edit&section=31" title="Edit section: Human use">edit</a>]</div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Further information: <a href="/wiki/Ethnolichenology" title="Ethnolichenology">Ethnolichenology</a></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Hiroshige_II_-_Kishu_kumano_iwatake_tori_-_Shokoku_meisho_hyakkei.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/Hiroshige_II_-_Kishu_kumano_iwatake_tori_-_Shokoku_meisho_hyakkei.jpg/170px-Hiroshige_II_-_Kishu_kumano_iwatake_tori_-_Shokoku_meisho_hyakkei.jpg" decoding="async" width="170" height="257" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/Hiroshige_II_-_Kishu_kumano_iwatake_tori_-_Shokoku_meisho_hyakkei.jpg/255px-Hiroshige_II_-_Kishu_kumano_iwatake_tori_-_Shokoku_meisho_hyakkei.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/Hiroshige_II_-_Kishu_kumano_iwatake_tori_-_Shokoku_meisho_hyakkei.jpg/340px-Hiroshige_II_-_Kishu_kumano_iwatake_tori_-_Shokoku_meisho_hyakkei.jpg 2x" data-file-width="5084" data-file-height="7692" /></a><figcaption><a href="/wiki/Iwatake" class="mw-redirect" title="Iwatake">Iwatake</a> (Umbilicaria esculenta) gathering at Kumano in Kishū, by <a href="/wiki/Hiroshige_II" title="Hiroshige II">Hiroshige II</a></figcaption></figure> <div class="mw-heading mw-heading3"><h3 id="Food">Food</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=32" title="Edit section: Food">edit</a>]</div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Edible_lichen" title="Edible lichen">Edible lichen</a></div> Lichens are eaten by many different cultures across the world. Although some lichens are only eaten in times of <a href="/wiki/Famine" title="Famine">famine</a>, others are a <a href="/wiki/Staple_food" title="Staple food">staple food</a> or even a <a href="/wiki/Delicacy" title="Delicacy">delicacy</a>. Two obstacles are often encountered when eating lichens: lichen <a href="/wiki/Polysaccharide" title="Polysaccharide">polysaccharides</a> are generally indigestible to humans, and lichens usually contain mildly toxic <a href="/wiki/Secondary_compound" class="mw-redirect" title="Secondary compound">secondary compounds</a> that should be removed before eating. Very few lichens are poisonous, but those high in <a href="/wiki/Vulpinic_acid" title="Vulpinic acid">vulpinic acid</a> or <a href="/wiki/Usnic_acid" title="Usnic acid">usnic acid</a> are toxic.<a href="#cite_note-Emmerich1993-146">[146]</a> Most poisonous lichens are yellow.[<a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed">citation needed</a>] In the past, <a href="/wiki/Iceland_moss" class="mw-redirect" title="Iceland moss">Iceland moss</a> (Cetraria islandica) was an important source of food for humans in northern Europe, and was cooked as a bread, porridge, pudding, soup, or salad. <a href="/wiki/Bryoria_fremontii" title="Bryoria fremontii">Bryoria fremontii</a> (edible horsehair lichen) was an important food in parts of North America, where it was usually <a href="/wiki/Pit_barbecue" title="Pit barbecue">pitcooked</a>. Northern peoples in North America and Siberia traditionally eat the partially digested <a href="/wiki/Cladonia_rangiferina" title="Cladonia rangiferina">reindeer lichen</a> (Cladina spp.) after they remove it from the <a href="/wiki/Rumen" title="Rumen">rumen</a> of caribou or reindeer that have been killed. <a href="/wiki/Rock_tripe" title="Rock tripe">Rock tripe</a> (Umbilicaria spp. and Lasalia spp.) is a lichen that has frequently been used as an emergency food in North America, and one species, <a href="/wiki/Umbilicaria_esculenta" title="Umbilicaria esculenta">Umbilicaria esculenta</a>, (iwatake in Japanese) is used in a variety of traditional Korean and Japanese foods. <div class="mw-heading mw-heading3"><h3 id="Lichenometry">Lichenometry</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=33" title="Edit section: Lichenometry">edit</a>]</div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Lichenometry" title="Lichenometry">Lichenometry</a></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Lichen_forming_a_Hidden_Mickey.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/9/9e/Lichen_forming_a_Hidden_Mickey.jpg/220px-Lichen_forming_a_Hidden_Mickey.jpg" decoding="async" width="220" height="170" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/9e/Lichen_forming_a_Hidden_Mickey.jpg/330px-Lichen_forming_a_Hidden_Mickey.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/9e/Lichen_forming_a_Hidden_Mickey.jpg/440px-Lichen_forming_a_Hidden_Mickey.jpg 2x" data-file-width="800" data-file-height="619" /></a><figcaption>Lichen grown in a <a href="/wiki/Mickey_Mouse" title="Mickey Mouse">Mickey Mouse</a> shape</figcaption></figure> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Lichen_Mickey_13_years_later.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a5/Lichen_Mickey_13_years_later.jpg/220px-Lichen_Mickey_13_years_later.jpg" decoding="async" width="220" height="170" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/a5/Lichen_Mickey_13_years_later.jpg/330px-Lichen_Mickey_13_years_later.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/a5/Lichen_Mickey_13_years_later.jpg/440px-Lichen_Mickey_13_years_later.jpg 2x" data-file-width="801" data-file-height="619" /></a><figcaption>13 years later</figcaption></figure> Lichenometry is a technique used to determine the age of exposed rock surfaces based on the size of lichen thalli. Introduced by Beschel in the 1950s,<a href="#cite_note-Beschel1950-147">[147]</a> the technique has found many applications. it is used in <a href="/wiki/Archaeology" title="Archaeology">archaeology</a>, <a href="/wiki/Palaeontology" class="mw-redirect" title="Palaeontology">palaeontology</a>, and <a href="/wiki/Geomorphology" title="Geomorphology">geomorphology</a>. It uses the presumed regular but slow rate of lichen growth to determine the <a href="/wiki/Surface_exposure_dating" title="Surface exposure dating">age of exposed rock</a>.<a href="#cite_note-LSN-41">[41]</a>: 9 <a href="#cite_note-148">[148]</a> Measuring the diameter (or other size measurement) of the largest lichen of a species on a rock surface indicates the length of time since the rock surface was first exposed. Lichen can be preserved on old rock faces for up to[<a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed">citation needed</a>] 10,000 years, providing the maximum age limit of the technique, though it is most accurate (within 10% error) when applied to surfaces that have been exposed for less than 1,000 years.<a href="#cite_note-149">[149]</a> Lichenometry is especially useful for dating surfaces less than 500 years old, as <a href="/wiki/Radiocarbon_dating" title="Radiocarbon dating">radiocarbon dating</a> techniques are less accurate over this period.<a href="#cite_note-innes-150">[150]</a> The lichens most commonly used for lichenometry are those of the genera <a href="/wiki/Rhizocarpon" title="Rhizocarpon">Rhizocarpon</a> (e.g. the species <a href="/wiki/Rhizocarpon_geographicum" title="Rhizocarpon geographicum">Rhizocarpon geographicum</a>, map lichen) and <a href="/wiki/Xanthoria" title="Xanthoria">Xanthoria</a>. <div class="mw-heading mw-heading3"><h3 id="Biodegradation">Biodegradation</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=34" title="Edit section: Biodegradation">edit</a>]</div> Lichens have been shown to degrade <a href="/wiki/Polyester_resin" title="Polyester resin">polyester resins</a>, as can be seen in archaeological sites in the Roman city of <a href="/wiki/Baelo_Claudia" title="Baelo Claudia">Baelo Claudia</a> in Spain.<a href="#cite_note-151">[151]</a> Lichens can accumulate several environmental pollutants such as lead, copper, and <a href="/wiki/Radionuclide" title="Radionuclide">radionuclides</a>.<a href="#cite_note-gadd-152">[152]</a> Some species of lichen, such as <a href="/wiki/Parmelia_sulcata" title="Parmelia sulcata">Parmelia sulcata</a> (called a hammered shield lichen, among other names) and <a href="/wiki/Lobaria_pulmonaria" title="Lobaria pulmonaria">Lobaria pulmonaria</a> (lung lichen), and many in the <a href="/wiki/Cladonia" title="Cladonia">Cladonia genus</a>, have been shown to produce <a href="/wiki/Serine_protease" title="Serine protease">serine proteases</a> capable of the degradation of pathogenic forms of <a href="/wiki/Prion_protein" class="mw-redirect" title="Prion protein">prion protein</a> (PrP), which may be useful in treating contaminated environmental reservoirs.<a href="#cite_note-153">[153]</a><a href="#cite_note-154">[154]</a><a href="#cite_note-155">[155]</a> <div class="mw-heading mw-heading3"><h3 id="Dyes">Dyes</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=35" title="Edit section: Dyes">edit</a>]</div> Many lichens produce secondary compounds, including <a href="/wiki/Biological_pigment" title="Biological pigment">pigments</a> that reduce harmful amounts of sunlight and powerful toxins that deter <a href="/wiki/Herbivore" title="Herbivore">herbivores</a> or kill bacteria. These compounds are very useful for lichen identification, and have had economic importance as <a href="/wiki/Dye" title="Dye">dyes</a> such as <a href="/wiki/Cudbear" class="mw-redirect" title="Cudbear">cudbear</a> or primitive <a href="/wiki/Antibiotics" class="mw-redirect" title="Antibiotics">antibiotics</a>. A <a href="/wiki/PH_indicator" title="PH indicator">pH indicator</a> (which can indicate acidic or basic substances) called <a href="/wiki/Litmus" title="Litmus">litmus</a> is a dye extracted from the lichen <a href="/wiki/Roccella_tinctoria" title="Roccella tinctoria">Roccella tinctoria</a> ("dyer's weed")<a href="#cite_note-156">[156]</a> by boiling. It gives its name to the well-known <a href="/wiki/Litmus_test_(chemistry)" class="mw-redirect" title="Litmus test (chemistry)">litmus test</a>. <a href="/wiki/Traditional_dyes_of_the_Scottish_Highlands" title="Traditional dyes of the Scottish Highlands">Traditional dyes of the Scottish Highlands</a> for <a href="/wiki/Harris_tweed" title="Harris tweed">Harris tweed</a><a href="#cite_note-WIL-4">[4]</a> and other traditional cloths were made from lichens, including the orange <a href="/wiki/Xanthoria_parietina" title="Xanthoria parietina">Xanthoria parietina</a> ("common orange lichen") and the grey foliaceous <a href="/wiki/Parmelia_saxatilis" title="Parmelia saxatilis">Parmelia saxatilis</a> common on rocks and known colloquially as "crottle". There are reports dating almost 2,000 years old of lichens being used to make purple and red dyes.<a href="#cite_note-Casselman1999-157">[157]</a> Of great historical and commercial significance are lichens belonging to the family <a href="/wiki/Roccellaceae" title="Roccellaceae">Roccellaceae</a>, commonly called orchella weed or orchil. <a href="/wiki/Orcein" title="Orcein">Orcein</a> and other lichen dyes have largely been replaced by <a href="/wiki/Dye#Natural_and_synthetic" title="Dye">synthetic versions</a>. <div class="mw-heading mw-heading3"><h3 id="Traditional_medicine_and_research">Traditional medicine and research</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=36" title="Edit section: Traditional medicine and research">edit</a>]</div> Historically, in <a href="/wiki/Traditional_medicine" title="Traditional medicine">traditional medicine</a> of Europe, <a href="/wiki/Lobaria_pulmonaria" title="Lobaria pulmonaria">Lobaria pulmonaria</a> was collected in large quantities as "lungwort", due to its lung-like appearance (the "<a href="/wiki/Doctrine_of_signatures" title="Doctrine of signatures">doctrine of signatures</a>" suggesting that herbs can treat body parts that they physically resemble).Similarly, <a href="/wiki/Peltigera_leucophlebia" title="Peltigera leucophlebia">Peltigera leucophlebia</a> ("ruffled freckled pelt") was used as a supposed cure for <a href="/wiki/Candidiasis" title="Candidiasis">thrush</a>, due to the resemblance of its cephalodia to the appearance of the disease.<a href="#cite_note-Dobson-37">[37]</a> Lichens produce <a href="/wiki/Metabolite" title="Metabolite">metabolites</a> being researched for their potential therapeutic or diagnostic value.<a href="#cite_note-158">[158]</a> Some metabolites produced by lichens are structurally and functionally similar to <a href="/wiki/Broad-spectrum_antibiotics" class="mw-redirect" title="Broad-spectrum antibiotics">broad-spectrum antibiotics</a> while few are associated respectively to antiseptic similarities.<a href="#cite_note-morton-159">[159]</a> <a href="/wiki/Usnic_acid" title="Usnic acid">Usnic acid</a> is the most commonly studied metabolite produced by lichens.<a href="#cite_note-morton-159">[159]</a> It is also under research as a <a href="/wiki/Bactericide" title="Bactericide">bactericidal</a> agent against <a href="/wiki/Escherichia_coli" title="Escherichia coli">Escherichia coli</a> and <a href="/wiki/Staphylococcus_aureus" title="Staphylococcus aureus">Staphylococcus aureus</a>.<a href="#cite_note-160">[160]</a> <div class="mw-heading mw-heading3"><h3 id="Aesthetic_appeal">Aesthetic appeal</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=37" title="Edit section: Aesthetic appeal">edit</a>]</div> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:CladonioPinetum.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/41/CladonioPinetum.jpg/220px-CladonioPinetum.jpg" decoding="async" width="220" height="149" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/41/CladonioPinetum.jpg/330px-CladonioPinetum.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/41/CladonioPinetum.jpg/440px-CladonioPinetum.jpg 2x" data-file-width="1740" data-file-height="1175" /></a><figcaption>Pine forest with <a href="/wiki/Cladonia" title="Cladonia">Cladonia</a> lichen ground-cover</figcaption></figure> Colonies of lichens may be spectacular in appearance, dominating the surface of the visual landscape as part of the aesthetic appeal to visitors of <a href="/wiki/Yosemite_National_Park" title="Yosemite National Park">Yosemite National Park</a>, <a href="/wiki/Sequoia_National_Park" title="Sequoia National Park">Sequoia National Park</a>, and the <a href="/wiki/Bay_of_Fires" title="Bay of Fires">Bay of Fires</a>.<a href="#cite_note-LSN-41">[41]</a>: 2  <a href="/wiki/Caloplaca" title="Caloplaca">Orange</a> and <a href="/wiki/Xanthoria" title="Xanthoria">yellow</a> lichens add to the ambience of desert trees, tundras, and rocky seashores. Intricate webs of lichens <a href="/wiki/Usnea" title="Usnea">hanging from tree branches</a> add a mysterious aspect to forests. Fruticose lichens are used in <a href="/wiki/Model_railroading" class="mw-redirect" title="Model railroading">model railroading</a><a href="#cite_note-161">[161]</a> and other modeling hobbies as a material for making miniature trees and shrubs. <div class="mw-heading mw-heading3"><h3 id="In_literature">In literature</h3>[<a href="/w/index.php?title=Lichen&action=edit&section=38" title="Edit section: In literature">edit</a>]</div> In early <a href="/wiki/Midrashic" class="mw-redirect" title="Midrashic">Midrashic</a> literature, the Hebrew word "vayilafeth" in <a href="/wiki/Book_of_Ruth" title="Book of Ruth">Ruth</a> 3:8 is explained as referring to <a href="/wiki/Ruth_(biblical_figure)" title="Ruth (biblical figure)">Ruth</a> entwining herself around <a href="/wiki/Boaz" title="Boaz">Boaz</a> like lichen.<a href="#cite_note-162">[162]</a> The 10th century Arab physician <a href="/wiki/Al-Tamimi,_the_physician" class="mw-redirect" title="Al-Tamimi, the physician">Al-Tamimi</a> mentions lichens dissolved in <a href="/wiki/Vinegar" title="Vinegar">vinegar</a> and <a href="/wiki/Rose_water" title="Rose water">rose water</a> being used in his day for the treatment of skin diseases and rashes.<a href="#cite_note-163">[163]</a> The plot of <a href="/wiki/John_Wyndham" title="John Wyndham">John Wyndham</a>'s science fiction novel <a href="/wiki/Trouble_with_Lichen" title="Trouble with Lichen">Trouble with Lichen</a> revolves around an anti-aging chemical extracted from a lichen. <div class="mw-heading mw-heading2"><h2 id="History">History</h2>[<a href="/w/index.php?title=Lichen&action=edit&section=39" title="Edit section: History">edit</a>]</div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Haeckel_Lichenes.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/39/Haeckel_Lichenes.jpg/220px-Haeckel_Lichenes.jpg" decoding="async" width="220" height="307" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/39/Haeckel_Lichenes.jpg/330px-Haeckel_Lichenes.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/39/Haeckel_Lichenes.jpg/440px-Haeckel_Lichenes.jpg 2x" data-file-width="2364" data-file-height="3294" /></a><figcaption>"Lichenes" fancifully drawn by <a href="/wiki/Ernst_Haeckel" title="Ernst Haeckel">Ernst Haeckel</a> to emphasize his ideas of symmetry in his <a href="/wiki/Kunstformen_der_Natur" title="Kunstformen der Natur">Artforms of Nature</a>, 1904</figcaption></figure> Although lichens had been recognized as organisms for quite some time, it was not until 1867, when Swiss botanist <a href="/wiki/Simon_Schwendener" title="Simon Schwendener">Simon Schwendener</a> proposed his dual theory of lichens, that lichens are a combination of fungi with algae or cyanobacteria, whereby the true nature of the lichen association began to emerge.<a href="#cite_note-Honegger2000-164">[164]</a> Schwendener's hypothesis, which at the time lacked experimental evidence, arose from his extensive analysis of the anatomy and development in lichens, algae, and fungi using a <a href="/wiki/Light_microscopy" class="mw-redirect" title="Light microscopy">light microscope</a>. Many of the leading lichenologists at the time, such as <a href="/wiki/James_Mascall_Morrison_Crombie" title="James Mascall Morrison Crombie">James Crombie</a> and <a href="/wiki/William_Nylander_(botanist)" title="William Nylander (botanist)">Nylander</a>, rejected Schwendener's hypothesis because the consensus was that all living organisms were autonomous.<a href="#cite_note-Honegger2000-164">[164]</a> Other prominent biologists, such as <a href="/wiki/Heinrich_Anton_de_Bary" title="Heinrich Anton de Bary">Heinrich Anton de Bary</a>, <a href="/wiki/Albert_Bernhard_Frank" title="Albert Bernhard Frank">Albert Bernhard Frank</a>, <a href="/wiki/Beatrix_Potter" title="Beatrix Potter">Beatrix Potter</a>, <a href="/wiki/Melchior_Treub" title="Melchior Treub">Melchior Treub</a> and <a href="/wiki/Hermann_Hellriegel" title="Hermann Hellriegel">Hermann Hellriegel</a>, were not so quick to reject Schwendener's ideas and the concept soon spread into other areas of study, such as microbial, plant, animal and human pathogens.<a href="#cite_note-Honegger2000-164">[164]</a><a href="#cite_note-Treub1873-165">[165]</a><a href="#cite_note-166">[166]</a> When the complex relationships between pathogenic microorganisms and their hosts were finally identified, Schwendener's hypothesis began to gain popularity. Further experimental proof of the dual nature of lichens was obtained when <a href="/w/index.php?title=Eugen_Thomas&action=edit&redlink=1" class="new" title="Eugen Thomas (page does not exist)">Eugen Thomas</a> published his results in 1939 on the first successful re-synthesis experiment.<a href="#cite_note-Honegger2000-164">[164]</a> In the 2010s, a new facet of the fungi–algae partnership was discovered. <a href="/wiki/Toby_Spribille" title="Toby Spribille">Toby Spribille</a> and colleagues found that many types of lichen that were long thought to be <a href="/wiki/Ascomycota" title="Ascomycota">ascomycete</a>–algae pairs were actually ascomycete–<a href="/wiki/Basidiomycota" title="Basidiomycota">basidiomycete</a>–algae trios. The third symbiotic partner in many lichens is a basidiomycete yeast.<a href="#cite_note-pmid_27445309-3">[3]</a><a href="#cite_note-Yong_2016-07-21-167">[167]</a> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2>[<a href="/w/index.php?title=Lichen&action=edit&section=40" title="Edit section: See also">edit</a>]</div> <style data-mw-deduplicate="TemplateStyles:r1239009302">.mw-parser-output .portalbox{padding:0;margin:0.5em 0;display:table;box-sizing:border-box;max-width:175px;list-style:none}.mw-parser-output .portalborder{border:1px solid var(--border-color-base,#a2a9b1);padding:0.1em;background:var(--background-color-neutral-subtle,#f8f9fa)}.mw-parser-output .portalbox-entry{display:table-row;font-size:85%;line-height:110%;height:1.9em;font-style:italic;font-weight:bold}.mw-parser-output .portalbox-image{display:table-cell;padding:0.2em;vertical-align:middle;text-align:center}.mw-parser-output .portalbox-link{display:table-cell;padding:0.2em 0.2em 0.2em 0.3em;vertical-align:middle}@media(min-width:720px){.mw-parser-output .portalleft{clear:left;float:left;margin:0.5em 1em 0.5em 0}.mw-parser-output .portalright{clear:right;float:right;margin:0.5em 0 0.5em 1em}}</style><ul role="navigation" aria-label="Portals" class="noprint portalbox portalborder portalright"> <li class="portalbox-entry"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/42/Karl_Johanssvamp%2C_Iduns_kokbok.png/25px-Karl_Johanssvamp%2C_Iduns_kokbok.png" decoding="async" width="25" height="28" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/42/Karl_Johanssvamp%2C_Iduns_kokbok.png/38px-Karl_Johanssvamp%2C_Iduns_kokbok.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/42/Karl_Johanssvamp%2C_Iduns_kokbok.png/51px-Karl_Johanssvamp%2C_Iduns_kokbok.png 2x" data-file-width="1105" data-file-height="1215" /><a href="/wiki/Portal:Fungi" title="Portal:Fungi">Fungi portal</a></li><li class="portalbox-entry"><a href="/wiki/File:Issoria_lathonia.jpg" class="mw-file-description"><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Issoria_lathonia.jpg/32px-Issoria_lathonia.jpg" decoding="async" width="32" height="23" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Issoria_lathonia.jpg/48px-Issoria_lathonia.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Issoria_lathonia.jpg/64px-Issoria_lathonia.jpg 2x" data-file-width="629" data-file-height="445" /></a><a href="/wiki/Portal:Biology" title="Portal:Biology">Biology portal</a></li></ul> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Outline_of_lichens" title="Outline of lichens">Outline of lichens</a></div> <ul><li><a href="/wiki/Lichenology" title="Lichenology">Lichenology</a></li> <li><a href="/wiki/Lichens_and_nitrogen_cycling" title="Lichens and nitrogen cycling">Lichens and nitrogen cycling</a></li> <li><a href="/wiki/Mycophycobiosis" title="Mycophycobiosis">Mycophycobiosis</a> - a symbiosis where a fungus lives in the macroscopic thallus of fresh water and marine algae; technically not a lichen but a similar phenomenon where fungi and algae are in symbiosis</li></ul> <div class="mw-heading mw-heading2"><h2 id="Notes">Notes</h2>[<a href="/w/index.php?title=Lichen&action=edit&section=41" title="Edit section: Notes">edit</a>]</div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> </div> <div class="mw-heading mw-heading2"><h2 id="References">References</h2>[<a href="/w/index.php?title=Lichen&action=edit&section=42" title="Edit section: References">edit</a>]</div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239543626"><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-1"><a href="#cite_ref-1">^</a> <style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFGrubeCardinalede_CastroMüller2009" class="citation journal cs1">Grube, Martin; 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Environmental Microbiology (Submitted manuscript). 13 (3): 806–818. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2011EnvMi..13..806C">2011EnvMi..13..806C</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1111%2Fj.1462-2920.2010.02386.x">10.1111/j.1462-2920.2010.02386.x</a>. <a href="/wiki/Hdl_(identifier)" class="mw-redirect" title="Hdl (identifier)">hdl</a>:<a rel="nofollow" class="external text" href="https://hdl.handle.net/10251%2F60269">10251/60269</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/21134099">21134099</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Environmental+Microbiology&rft.atitle=Two+Trebouxia+algae+with+different+physiological+performances+are+ever-present+in+lichen+thalli+of+Ramalina+farinacea.+Coexistence+versus+competition%3F&rft.volume=13&rft.issue=3&rft.pages=806-818&rft.date=2011&rft_id=info%3Ahdl%2F10251%2F60269&rft_id=info%3Apmid%2F21134099&rft_id=info%3Adoi%2F10.1111%2Fj.1462-2920.2010.02386.x&rft_id=info%3Abibcode%2F2011EnvMi..13..806C&rft.aulast=Casano&rft.aufirst=L.+M.&rft.au=Del+Campo%2C+E.+M.&rft.au=Garc%C3%ADa-Breijo%2C+F.+J.&rft.au=Reig-Armi%C3%B1ana%2C+J&rft.au=Gasulla%2C+F&rft.au=Del+Hoyo%2C+A&rft.au=Gu%C3%A9ra%2C+A&rft.au=Barreno%2C+E&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-FESM-21">^ <a href="#cite_ref-FESM_21-0">a</a> <a href="#cite_ref-FESM_21-1">b</a> <a href="/wiki/Rosmarie_Honegger" title="Rosmarie Honegger">Honegger, R.</a> (1991) Fungal evolution: symbiosis and morphogenesis, Symbiosis as a Source of Evolutionary Innovation, Margulis, L., and Fester, R. (eds). Cambridge, MA, US: The MIT Press, pp. 319–340. </li> <li id="cite_note-SSSFD-22">^ <a href="#cite_ref-SSSFD_22-0">a</a> <a href="#cite_ref-SSSFD_22-1">b</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFGrubeCardinaleDe_CastroMüller2009" class="citation journal cs1">Grube, M; Cardinale, M; De Castro, J. V. Jr.; Müller, H; Berg, G (2009). <a rel="nofollow" class="external text" href="https://doi.org/10.1038%2Fismej.2009.63">"Species-specific structural and functional diversity of bacterial communities in lichen symbioses"</a>. The ISME Journal. 3 (9): 1105–1115. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2009ISMEJ...3.1105G">2009ISMEJ...3.1105G</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2Fismej.2009.63">10.1038/ismej.2009.63</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/19554038">19554038</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+ISME+Journal&rft.atitle=Species-specific+structural+and+functional+diversity+of+bacterial+communities+in+lichen+symbioses&rft.volume=3&rft.issue=9&rft.pages=1105-1115&rft.date=2009&rft_id=info%3Apmid%2F19554038&rft_id=info%3Adoi%2F10.1038%2Fismej.2009.63&rft_id=info%3Abibcode%2F2009ISMEJ...3.1105G&rft.aulast=Grube&rft.aufirst=M&rft.au=Cardinale%2C+M&rft.au=De+Castro%2C+J.+V.+Jr.&rft.au=M%C3%BCller%2C+H&rft.au=Berg%2C+G&rft_id=https%3A%2F%2Fdoi.org%2F10.1038%252Fismej.2009.63&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-NPB-23">^ <a href="#cite_ref-NPB_23-0">a</a> <a href="#cite_ref-NPB_23-1">b</a> Barreno, E., Herrera-Campos, M., García-Breijo, F., Gasulla, F., and Reig-Armiñana, J. (2008) <a rel="nofollow" class="external text" href="https://www.researchgate.net/publication/258264055">"Non photosynthetic bacteria associated to cortical structures on Ramalina and Usnea thalli from Mexico"</a>. Asilomar, Pacific Grove, CA, USA: Abstracts IAL 6- ABLS Joint Meeting. </li> <li id="cite_note-Morris2007-24">^ <a href="#cite_ref-Morris2007_24-0">a</a> <a href="#cite_ref-Morris2007_24-1">b</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMorrisPurvis2007" class="citation book cs1">Morris J, Purvis W (2007). Lichens (Life). London: The Natural History Museum. p. 19. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-565-09153-8" title="Special:BookSources/978-0-565-09153-8"><bdi>978-0-565-09153-8</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Lichens+%28Life%29&rft.place=London&rft.pages=19&rft.pub=The+Natural+History+Museum&rft.date=2007&rft.isbn=978-0-565-09153-8&rft.aulast=Morris&rft.aufirst=J&rft.au=Purvis%2C+W&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-25"><a href="#cite_ref-25">^</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.nps.gov/articles/lichen-and-our-air.htm">"Lichen - The Little Things That Matter (U.S. National Park Service)"</a>. www.nps.gov. Retrieved 17 November 2023.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=www.nps.gov&rft.atitle=Lichen+-+The+Little+Things+That+Matter+%28U.S.+National+Park+Service%29&rft_id=https%3A%2F%2Fwww.nps.gov%2Farticles%2Flichen-and-our-air.htm&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-26"><a href="#cite_ref-26">^</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHarper" class="citation web cs1">Harper, Douglas. <a rel="nofollow" class="external text" href="https://www.etymonline.com/?term=lichen">"lichen"</a>. <a href="/wiki/Online_Etymology_Dictionary" title="Online Etymology Dictionary">Online Etymology Dictionary</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Online+Etymology+Dictionary&rft.atitle=lichen&rft.aulast=Harper&rft.aufirst=Douglas&rft_id=https%3A%2F%2Fwww.etymonline.com%2F%3Fterm%3Dlichen&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-27"><a href="#cite_ref-27">^</a> <a rel="nofollow" class="external text" href="http://www.perseus.tufts.edu/hopper/text?doc=Perseus:text:1999.04.0059:entry=lichen">lichen</a>. Charlton T. Lewis and Charles Short. <a href="/wiki/A_Latin_Dictionary" title="A Latin Dictionary">A Latin Dictionary</a> on <a href="/wiki/Perseus_Project" class="mw-redirect" title="Perseus Project">Perseus Project</a>. </li> <li id="cite_note-28"><a href="#cite_ref-28">^</a> <a rel="nofollow" class="external text" href="https://www.perseus.tufts.edu/hopper/text?doc=Perseus:text:1999.04.0057:entry=leixh/n">λειχήν</a>. <a href="/wiki/Henry_Liddell" title="Henry Liddell">Liddell, Henry George</a>; <a href="/wiki/Robert_Scott_(philologist)" title="Robert Scott (philologist)">Scott, Robert</a>; <a href="/wiki/A_Greek%E2%80%93English_Lexicon" title="A Greek–English Lexicon">A Greek–English Lexicon</a> at the <a href="/wiki/Perseus_Project" class="mw-redirect" title="Perseus Project">Perseus Project</a>. </li> <li id="cite_note-29"><a href="#cite_ref-29">^</a> <a rel="nofollow" class="external text" href="https://www.perseus.tufts.edu/hopper/text?doc=Perseus:text:1999.04.0057:entry=lei/xw">λείχειν</a> in <a href="/wiki/Henry_Liddell" title="Henry Liddell">Liddell</a> and <a href="/wiki/Robert_Scott_(philologist)" title="Robert Scott (philologist)">Scott</a>. </li> <li id="cite_note-Beekes-30"><a href="#cite_ref-Beekes_30-0">^</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBeekes2010" class="citation book cs1"><a href="/wiki/Robert_S._P._Beekes" title="Robert S. P. Beekes">Beekes, Robert S. P.</a> (2010). "s.v. λειχήν, λείχω". Etymological Dictionary of Greek. Leiden Indo-European Etymological Dictionary Series. Vol. 1. With the assistance of Lucien van Beek. Leiden, Boston: Brill. pp. 846–47. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/9789004174184" title="Special:BookSources/9789004174184"><bdi>9789004174184</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=s.v.+%CE%BB%CE%B5%CE%B9%CF%87%CE%AE%CE%BD%2C+%CE%BB%CE%B5%CE%AF%CF%87%CF%89&rft.btitle=Etymological+Dictionary+of+Greek&rft.place=Leiden%2C+Boston&rft.series=Leiden+Indo-European+Etymological+Dictionary+Series&rft.pages=846-47&rft.pub=Brill&rft.date=2010&rft.isbn=9789004174184&rft.aulast=Beekes&rft.aufirst=Robert+S.+P.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-Cambridge-31"><a href="#cite_ref-Cambridge_31-0">^</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://dictionary.cambridge.org/pronunciation/english/lichen">"Lichen"</a>. Cambridge Dictionary. Retrieved 8 September 2022.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Lichen&rft.pub=Cambridge+Dictionary&rft_id=https%3A%2F%2Fdictionary.cambridge.org%2Fpronunciation%2Fenglish%2Flichen&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-Wordsworth-32"><a href="#cite_ref-Wordsworth_32-0">^</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWordsworth2012" class="citation web cs1">Wordsworth, Dot (17 November 2012). <a rel="nofollow" class="external text" href="https://www.spectator.co.uk/article/lichen">"Lichen"</a>. spectator.co.uk. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20141223053140/http://www.spectator.co.uk/life/mind-your-language/8757911/lichen/">Archived</a> from the original on 23 December 2014. Retrieved 8 September 2022.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Lichen&rft.pub=spectator.co.uk&rft.date=2012-11-17&rft.aulast=Wordsworth&rft.aufirst=Dot&rft_id=https%3A%2F%2Fwww.spectator.co.uk%2Farticle%2Flichen&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-33"><a href="#cite_ref-33">^</a> The Oxford English Dictionary cites only the "liken" pronunciation: <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFReference-OED-lichen" class="citation encyclopaedia cs1"><a rel="nofollow" class="external text" href="https://www.oed.com/search/dictionary/?q=lichen">"lichen"</a>. <a href="/wiki/Oxford_English_Dictionary" title="Oxford English Dictionary">Oxford English Dictionary</a> (Online ed.). <a href="/wiki/Oxford_University_Press" title="Oxford University Press">Oxford University Press</a>. Retrieved 10 January 2018.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=lichen&rft.btitle=Oxford+English+Dictionary&rft.edition=Online&rft.pub=Oxford+University+Press&rft_id=https%3A%2F%2Fwww.oed.com%2Fsearch%2Fdictionary%2F%3Fq%3Dlichen&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> (Subscription or <a rel="nofollow" class="external text" href="https://www.oed.com/public/login/loggingin#withyourlibrary">participating institution membership</a> required.) </li> <li id="cite_note-MSULB-34">^ <a href="#cite_ref-MSULB_34-0">a</a> <a href="#cite_ref-MSULB_34-1">b</a> <a href="#cite_ref-MSULB_34-2">c</a> <a href="#cite_ref-MSULB_34-3">d</a> <a href="#cite_ref-MSULB_34-4">e</a> <a href="#cite_ref-MSULB_34-5">f</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20111005224410/https://www.msu.edu/course/bot/423/Plntlist8mosslichen.html">"Lichens and Bryophytes, Michigan State University, 10-25-99"</a>. Archived from <a rel="nofollow" class="external text" href="https://www.msu.edu/course/bot/423/Plntlist8mosslichen.html">the original</a> on 5 October 2011. Retrieved 10 October 2014.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Lichens+and+Bryophytes%2C+Michigan+State+University%2C+10-25-99&rft_id=https%3A%2F%2Fwww.msu.edu%2Fcourse%2Fbot%2F423%2FPlntlist8mosslichen.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-LNAILV-35">^ <a href="#cite_ref-LNAILV_35-0">a</a> <a href="#cite_ref-LNAILV_35-1">b</a> <a href="#cite_ref-LNAILV_35-2">c</a> <a href="#cite_ref-LNAILV_35-3">d</a> <a href="#cite_ref-LNAILV_35-4">e</a> <a href="#cite_ref-LNAILV_35-5">f</a> <a href="#cite_ref-LNAILV_35-6">g</a> <a href="#cite_ref-LNAILV_35-7">h</a> <a href="#cite_ref-LNAILV_35-8">i</a> <a href="#cite_ref-LNAILV_35-9">j</a> Lichen Vocabulary, Lichens of North America Information, Sylvia and Stephen Sharnoff, <a rel="nofollow" class="external autonumber" href="http://www.lichen.com/vocabulary.html">[1]</a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20150120063843/http://www.lichen.com/vocabulary.html">Archived</a> 20 January 2015 at the <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback Machine</a> </li> <li id="cite_note-ASLGPZ-36">^ <a href="#cite_ref-ASLGPZ_36-0">a</a> <a href="#cite_ref-ASLGPZ_36-1">b</a> <a href="#cite_ref-ASLGPZ_36-2">c</a> <a href="#cite_ref-ASLGPZ_36-3">d</a> <a href="#cite_ref-ASLGPZ_36-4">e</a> <a href="#cite_ref-ASLGPZ_36-5">f</a> <a href="#cite_ref-ASLGPZ_36-6">g</a> <a href="#cite_ref-ASLGPZ_36-7">h</a> <a href="#cite_ref-ASLGPZ_36-8">i</a> <a href="#cite_ref-ASLGPZ_36-9">j</a> <a href="#cite_ref-ASLGPZ_36-10">k</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.lichens.lastdragon.org/faq/glossary3.html">"Alan Silverside's Lichen Glossary (p-z), Alan Silverside"</a>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20141031095023/http://www.lichens.lastdragon.org/faq/glossary3.html">Archived</a> from the original on 31 October 2014. Retrieved 10 October 2014.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Alan+Silverside%27s+Lichen+Glossary+%28p-z%29%2C+Alan+Silverside&rft_id=http%3A%2F%2Fwww.lichens.lastdragon.org%2Ffaq%2Fglossary3.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-Dobson-37">^ <a href="#cite_ref-Dobson_37-0">a</a> <a href="#cite_ref-Dobson_37-1">b</a> <a href="#cite_ref-Dobson_37-2">c</a> <a href="#cite_ref-Dobson_37-3">d</a> <a href="#cite_ref-Dobson_37-4">e</a> <a href="#cite_ref-Dobson_37-5">f</a> <a href="#cite_ref-Dobson_37-6">g</a> <a href="#cite_ref-Dobson_37-7">h</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFDobson2011" class="citation book cs1">Dobson, F.S. (2011). Lichens, an illustrated guide to the British and Irish species. Slough, UK: Richmond Publishing Co. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/9780855463151" title="Special:BookSources/9780855463151"><bdi>9780855463151</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Lichens%2C+an+illustrated+guide+to+the+British+and+Irish+species&rft.place=Slough%2C+UK&rft.pub=Richmond+Publishing+Co.&rft.date=2011&rft.isbn=9780855463151&rft.aulast=Dobson&rft.aufirst=F.S.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-ASLTT-38">^ <a href="#cite_ref-ASLTT_38-0">a</a> <a href="#cite_ref-ASLTT_38-1">b</a> <a href="#cite_ref-ASLTT_38-2">c</a> <a href="#cite_ref-ASLTT_38-3">d</a> <a href="#cite_ref-ASLTT_38-4">e</a> <a href="#cite_ref-ASLTT_38-5">f</a> <a href="#cite_ref-ASLTT_38-6">g</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.lichens.lastdragon.org/faq/lichenthallustypes.html#foliose">"Foliose lichens, Lichen Thallus Types, Allan Silverside"</a>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20141019112429/http://www.lichens.lastdragon.org/faq/lichenthallustypes.html#foliose">Archived</a> from the original on 19 October 2014. Retrieved 10 October 2014.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Foliose+lichens%2C+Lichen+Thallus+Types%2C+Allan+Silverside&rft_id=http%3A%2F%2Fwww.lichens.lastdragon.org%2Ffaq%2Flichenthallustypes.html%23foliose&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-VMBMLF-39">^ <a href="#cite_ref-VMBMLF_39-0">a</a> <a href="#cite_ref-VMBMLF_39-1">b</a> <a href="#cite_ref-VMBMLF_39-2">c</a> <a href="#cite_ref-VMBMLF_39-3">d</a> <a href="#cite_ref-VMBMLF_39-4">e</a> <a href="#cite_ref-VMBMLF_39-5">f</a> <a href="#cite_ref-VMBMLF_39-6">g</a> <a href="#cite_ref-VMBMLF_39-7">h</a> <a href="#cite_ref-VMBMLF_39-8">i</a> <a href="#cite_ref-VMBMLF_39-9">j</a> <a href="#cite_ref-VMBMLF_39-10">k</a> <a href="#cite_ref-VMBMLF_39-11">l</a> <a href="#cite_ref-VMBMLF_39-12">m</a> Mosses Lichens & Ferns of Northwest North America, Dale H. Vitt, Janet E. Marsh, Robin B. Bovey, Lone Pine Publishing Company, <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/0-295-96666-1" title="Special:BookSources/0-295-96666-1">0-295-96666-1</a> </li> <li id="cite_note-SJCL-40">^ <a href="#cite_ref-SJCL_40-0">a</a> <a href="#cite_ref-SJCL_40-1">b</a> <a href="#cite_ref-SJCL_40-2">c</a> <a href="#cite_ref-SJCL_40-3">d</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20150510220114/http://www.saguaro-juniper.com/i_and_i/lichens/lichens.html">"Lichens, Saguaro-Juniper Corporation"</a>. Archived from <a rel="nofollow" class="external text" href="http://www.saguaro-juniper.com/i_and_i/lichens/lichens.html">the original</a> on 10 May 2015. Retrieved 10 October 2014.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Lichens%2C+Saguaro-Juniper+Corporation&rft_id=http%3A%2F%2Fwww.saguaro-juniper.com%2Fi_and_i%2Flichens%2Flichens.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-LSN-41">^ <a href="#cite_ref-LSN_41-0">a</a> <a href="#cite_ref-LSN_41-1">b</a> <a href="#cite_ref-LSN_41-2">c</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMcCuneGrenonMartinMutch2007" class="citation journal cs1">McCune, B.; Grenon, J.; Martin, E.; Mutch, L.S.; Martin, E.P. (March 2007). "Lichens in relation to management issues in the Sierra Nevada national parks". North American Fungi. 2: 1–39. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.2509%2Fpnwf.2007.002.003">10.2509/pnwf.2007.002.003</a> (inactive 1 November 2024).</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=North+American+Fungi&rft.atitle=Lichens+in+relation+to+management+issues+in+the+Sierra+Nevada+national+parks&rft.volume=2&rft.pages=1-39&rft.date=2007-03&rft_id=info%3Adoi%2F10.2509%2Fpnwf.2007.002.003&rft.aulast=McCune&rft.aufirst=B.&rft.au=Grenon%2C+J.&rft.au=Martin%2C+E.&rft.au=Mutch%2C+L.S.&rft.au=Martin%2C+E.P.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"><code class="cs1-code">{{<a href="/wiki/Template:Cite_journal" title="Template:Cite journal">cite journal</a>}}</code>: CS1 maint: DOI inactive as of November 2024 (<a href="/wiki/Category:CS1_maint:_DOI_inactive_as_of_November_2024" title="Category:CS1 maint: DOI inactive as of November 2024">link</a>) </li> <li id="cite_note-JJMML-42">^ <a href="#cite_ref-JJMML_42-0">a</a> <a href="#cite_ref-JJMML_42-1">b</a> <a href="#cite_ref-JJMML_42-2">c</a> <a href="#cite_ref-JJMML_42-3">d</a> Michigan Lichens, Julie Jones Medlin, B. Jain Publishers, 1996, <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/0877370397" title="Special:BookSources/0877370397">0877370397</a>, 9780877370390, <a rel="nofollow" class="external autonumber" href="https://books.google.com/books?id=-Jo0yP2-69UC&pg=PA8">[2]</a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20161124144607/https://books.google.com/books?id=-Jo0yP2-69UC&pg=PA8">Archived</a> 24 November 2016 at the <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback Machine</a> </li> <li id="cite_note-LMM-43">^ <a href="#cite_ref-LMM_43-0">a</a> <a href="#cite_ref-LMM_43-1">b</a> <a href="#cite_ref-LMM_43-2">c</a> <a href="#cite_ref-LMM_43-3">d</a> <a href="#cite_ref-LMM_43-4">e</a> <a href="#cite_ref-LMM_43-5">f</a> <a href="#cite_ref-LMM_43-6">g</a> <a href="#cite_ref-LMM_43-7">h</a> <a href="#cite_ref-LMM_43-8">i</a> <a href="#cite_ref-LMM_43-9">j</a> Lichens: More on Morphology, University of California Museum of Paleontology, <a rel="nofollow" class="external autonumber" href="http://www.ucmp.berkeley.edu/fungi/lichens/lichenmm.html">[3]</a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20150228172027/http://www.ucmp.berkeley.edu/fungi/lichens/lichenmm.html">Archived</a> 28 February 2015 at the <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback Machine</a> </li> <li id="cite_note-UNOLP-44">^ <a href="#cite_ref-UNOLP_44-0">a</a> <a href="#cite_ref-UNOLP_44-1">b</a> <a href="#cite_ref-UNOLP_44-2">c</a> <a href="#cite_ref-UNOLP_44-3">d</a> <a href="#cite_ref-UNOLP_44-4">e</a> <a href="#cite_ref-UNOLP_44-5">f</a> <a href="#cite_ref-UNOLP_44-6">g</a> <a href="#cite_ref-UNOLP_44-7">h</a> <a href="#cite_ref-UNOLP_44-8">i</a> <a href="#cite_ref-UNOLP_44-9">j</a> <a href="#cite_ref-UNOLP_44-10">k</a> <a href="#cite_ref-UNOLP_44-11">l</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20141006145803/http://www.unomaha.edu/lichens/Bio%204350%20PDF/Photobionts.pdf">"Lichen Photobionts, University of Nebraska Omaha"</a> (PDF). Archived from <a rel="nofollow" class="external text" href="http://www.unomaha.edu/lichens/Bio%204350%20PDF/Photobionts.pdf">the original</a> (PDF) on 6 October 2014.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Lichen+Photobionts%2C+University+of+Nebraska+Omaha&rft_id=http%3A%2F%2Fwww.unomaha.edu%2Flichens%2FBio%25204350%2520PDF%2FPhotobionts.pdf&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-ASLGGO-45">^ <a href="#cite_ref-ASLGGO_45-0">a</a> <a href="#cite_ref-ASLGGO_45-1">b</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.lichens.lastdragon.org/faq/glossary2.html">"Alan Silverside's Lichen Glossary (g-o), Alan Silverside"</a>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20141102105836/http://www.lichens.lastdragon.org/faq/glossary2.html">Archived</a> from the original on 2 November 2014. Retrieved 10 October 2014.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Alan+Silverside%27s+Lichen+Glossary+%28g-o%29%2C+Alan+Silverside&rft_id=http%3A%2F%2Fwww.lichens.lastdragon.org%2Ffaq%2Fglossary2.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-Budel1996-46">^ <a href="#cite_ref-Budel1996_46-0">a</a> <a href="#cite_ref-Budel1996_46-1">b</a> <a href="#cite_ref-Budel1996_46-2">c</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBüdel,_B.Scheidegger,_C.1996" class="citation book cs1">Büdel, B.; Scheidegger, C. (1996). "Thallus morphology and anatomy". Lichen Biology. pp. 37–64. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1017%2FCBO9780511790478.005">10.1017/CBO9780511790478.005</a>. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/9780511790478" title="Special:BookSources/9780511790478"><bdi>9780511790478</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=Thallus+morphology+and+anatomy&rft.btitle=Lichen+Biology&rft.pages=37-64&rft.date=1996&rft_id=info%3Adoi%2F10.1017%2FCBO9780511790478.005&rft.isbn=9780511790478&rft.au=B%C3%BCdel%2C+B.&rft.au=Scheidegger%2C+C.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-HPTCLGD-47">^ <a href="#cite_ref-HPTCLGD_47-0">a</a> <a href="#cite_ref-HPTCLGD_47-1">b</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHeiđmarssonHeidmarsson1996" class="citation journal cs1">Heiđmarsson, Starri; Heidmarsson, Starri (1996). 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Catherine; Schneider, Kevin; Stabentheiner, Edith; Toome-Heller, Merje; Thor, Göran; Mayrhofer, Helmut; Johannesson, Hanna; McCutcheon, John P. (29 July 2016). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5793994">"Basidiomycete yeasts in the cortex of ascomycete macrolichens"</a>. 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New York: Watson-Guptill Publications. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-8230-5727-6" title="Special:BookSources/978-0-8230-5727-6"><bdi>978-0-8230-5727-6</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Wild+color%3A+%5Bthe+complete+guide+to+making+and+using+natural+dyes%5D&rft.place=New+York&rft.pub=Watson-Guptill+Publications&rft.date=1999&rft.isbn=978-0-8230-5727-6&rft.au=Casselman%2C+Karen+Leigh&rft.au=Dean%2C+Jenny&rft_id=https%3A%2F%2Farchive.org%2Fdetails%2Fwildcolor00dean&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-158"><a href="#cite_ref-158">^</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMuller2001" class="citation journal cs1">Muller, K (2001). "Pharmaceutically Relevant Metabolites from Lichens". Applied Microbiology and Biotechnology. 56 (1–2): 9–10. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1007%2Fs002530100684">10.1007/s002530100684</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/11499952">11499952</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:11958438">11958438</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Applied+Microbiology+and+Biotechnology&rft.atitle=Pharmaceutically+Relevant+Metabolites+from+Lichens&rft.volume=56&rft.issue=1%E2%80%932&rft.pages=9-10&rft.date=2001&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A11958438%23id-name%3DS2CID&rft_id=info%3Apmid%2F11499952&rft_id=info%3Adoi%2F10.1007%2Fs002530100684&rft.aulast=Muller&rft.aufirst=K&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-morton-159">^ <a href="#cite_ref-morton_159-0">a</a> <a href="#cite_ref-morton_159-1">b</a> Morton, E.; Winters, J. and Smith, L. (2010). <a rel="nofollow" class="external text" href="https://deepblue.lib.umich.edu/bitstream/handle/2027.42/78342/Morton_Winters_Smith_2010.pdf?sequence=1">"An Analysis of Antiseptic and Antibiotic Properties of Variously Treated Mosses and Lichens"</a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20170820033126/https://deepblue.lib.umich.edu/bitstream/handle/2027.42/78342/Morton_Winters_Smith_2010.pdf?sequence=1">Archived</a> 20 August 2017 at the <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback Machine</a>. University of Michigan Biological Station </li> <li id="cite_note-160"><a href="#cite_ref-160">^</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBustinza1952" class="citation journal cs1">Bustinza, F. (1952). "Antibacterial Substances from Lichens". Economic Botany. 6 (4): 402–406. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1952EcBot...6..402B">1952EcBot...6..402B</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1007%2Fbf02984888">10.1007/bf02984888</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:39335883">39335883</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Economic+Botany&rft.atitle=Antibacterial+Substances+from+Lichens&rft.volume=6&rft.issue=4&rft.pages=402-406&rft.date=1952&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A39335883%23id-name%3DS2CID&rft_id=info%3Adoi%2F10.1007%2Fbf02984888&rft_id=info%3Abibcode%2F1952EcBot...6..402B&rft.aulast=Bustinza&rft.aufirst=F.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-161"><a href="#cite_ref-161">^</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20141015193251/http://www.themodelrailroader.com/landscaping/using-lichen-on-your-model-railway-layout/">"Themodelrailroader.com"</a>. Archived from <a rel="nofollow" class="external text" href="http://yourdiorama.com/landscaping/using-lichen-on-your-model-railway-layout/">the original</a> on 15 October 2014. Retrieved 10 October 2014.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Themodelrailroader.com&rft_id=http%3A%2F%2Fyourdiorama.com%2Flandscaping%2Fusing-lichen-on-your-model-railway-layout%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-162"><a href="#cite_ref-162">^</a> Thus explained by Rabbi <a href="/wiki/Enoch_Zundel_ben_Joseph" title="Enoch Zundel ben Joseph">Enoch Zundel ben Joseph</a>, in his commentary Etz Yosef ("Tree of Joseph"), on Sefer <a href="/wiki/Midrash_Rabbah" class="mw-redirect" title="Midrash Rabbah">Midrash Rabbah</a>, vol. 2, New York 1987, s.v. Ruth Rabba 6:3 </li> <li id="cite_note-163"><a href="#cite_ref-163">^</a> Zohar Amar and Yaron Serri, The Land of Israel and Syria as Described by Al-Tamimi, Ramat-Gan 2004, pp. 56, 108–109 <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/965-226-252-8" title="Special:BookSources/965-226-252-8">965-226-252-8</a> (Hebrew) </li> <li id="cite_note-Honegger2000-164">^ <a href="#cite_ref-Honegger2000_164-0">a</a> <a href="#cite_ref-Honegger2000_164-1">b</a> <a href="#cite_ref-Honegger2000_164-2">c</a> <a href="#cite_ref-Honegger2000_164-3">d</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHonegger_R.2000" class="citation journal cs1">Honegger R. (2000). "Simon Schwender (1829–1919) and the dual hypothesis in lichens". Bryologist. 103 (2): 307–313. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1639%2F0007-2745%282000%29103%5B0307%3ASSATDH%5D2.0.CO%3B2">10.1639/0007-2745(2000)103[0307:SSATDH]2.0.CO;2</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0007-2745">0007-2745</a>. <a href="/wiki/JSTOR_(identifier)" class="mw-redirect" title="JSTOR (identifier)">JSTOR</a> <a rel="nofollow" class="external text" href="https://www.jstor.org/stable/3244159">3244159</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:84580224">84580224</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Bryologist&rft.atitle=Simon+Schwender+%281829%E2%80%931919%29+and+the+dual+hypothesis+in+lichens&rft.volume=103&rft.issue=2&rft.pages=307-313&rft.date=2000&rft.issn=0007-2745&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A84580224%23id-name%3DS2CID&rft_id=https%3A%2F%2Fwww.jstor.org%2Fstable%2F3244159%23id-name%3DJSTOR&rft_id=info%3Adoi%2F10.1639%2F0007-2745%282000%29103%5B0307%3ASSATDH%5D2.0.CO%3B2&rft.au=Honegger+R.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-Treub1873-165"><a href="#cite_ref-Treub1873_165-0">^</a> Treub, Melchior (1873) Onderzoekingen over de natuur der lichenen. Dissertation Leiden University. </li> <li id="cite_note-166"><a href="#cite_ref-166">^</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWakeford2002" class="citation book cs1">Wakeford, Tom (2002). <a rel="nofollow" class="external text" href="http://worldcat.org/oclc/922973303">Liaisons of Life : From Hornworts to Hippos, How the Unassuming Microbe Has Driven Evolution</a>. John Wiley & Sons, Incorporated. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-470-24400-5" title="Special:BookSources/978-0-470-24400-5"><bdi>978-0-470-24400-5</bdi></a>. <a href="/wiki/OCLC_(identifier)" class="mw-redirect" title="OCLC (identifier)">OCLC</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/oclc/922973303">922973303</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Liaisons+of+Life+%3A+From+Hornworts+to+Hippos%2C+How+the+Unassuming+Microbe+Has+Driven+Evolution.&rft.pub=John+Wiley+%26+Sons%2C+Incorporated&rft.date=2002&rft_id=info%3Aoclcnum%2F922973303&rft.isbn=978-0-470-24400-5&rft.aulast=Wakeford&rft.aufirst=Tom&rft_id=http%3A%2F%2Fworldcat.org%2Foclc%2F922973303&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> <li id="cite_note-Yong_2016-07-21-167"><a href="#cite_ref-Yong_2016-07-21_167-0">^</a> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFYong2016" class="citation cs2"><a href="/wiki/Ed_Yong" title="Ed Yong">Yong, Ed</a> (21 July 2016), <a rel="nofollow" class="external text" href="https://www.theatlantic.com/science/archive/2016/07/how-a-guy-from-a-montana-trailer-park-upturned-150-years-of-biology/491702/">"How a guy from a Montana trailer park overturned 150 years of biology"</a>, The Atlantic, <a rel="nofollow" class="external text" href="https://web.archive.org/web/20170723161812/https://www.theatlantic.com/science/archive/2016/07/how-a-guy-from-a-montana-trailer-park-upturned-150-years-of-biology/491702/">archived</a> from the original on 23 July 2017, retrieved 23 July 2017.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+Atlantic&rft.atitle=How+a+guy+from+a+Montana+trailer+park+overturned+150+years+of+biology&rft.date=2016-07-21&rft.aulast=Yong&rft.aufirst=Ed&rft_id=https%3A%2F%2Fwww.theatlantic.com%2Fscience%2Farchive%2F2016%2F07%2Fhow-a-guy-from-a-montana-trailer-park-upturned-150-years-of-biology%2F491702%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3ALichen" class="Z3988"> </li> </ol></div></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 0;box-sizing:border-box;border:1px solid #aaa;font-size:88%;line-height:1.25em;background-color:var(--background-color-interactive-subtle,#f8f9fa);display:flow-root}.mw-parser-output .side-box-abovebelow,.mw-parser-output .side-box-text{padding:0.25em 0.9em}.mw-parser-output .side-box-image{padding:2px 0 2px 0.9em;text-align:center}.mw-parser-output .side-box-imageright{padding:2px 0.9em 2px 0;text-align:center}@media(min-width:500px){.mw-parser-output .side-box-flex{display:flex;align-items:center}.mw-parser-output .side-box-text{flex:1;min-width:0}}@media(min-width:720px){.mw-parser-output .side-box{width:238px}.mw-parser-output .side-box-right{clear:right;float:right;margin-left:1em}.mw-parser-output .side-box-left{margin-right:1em}}</style><style data-mw-deduplicate="TemplateStyles:r1237033735">@media print{body.ns-0 .mw-parser-output .sistersitebox{display:none!important}}@media screen{html.skin-theme-clientpref-night .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}</style><div class="side-box side-box-right plainlinks sistersitebox"><style data-mw-deduplicate="TemplateStyles:r1126788409">.mw-parser-output .plainlist ol,.mw-parser-output .plainlist ul{line-height:inherit;list-style:none;margin:0;padding:0}.mw-parser-output .plainlist ol li,.mw-parser-output .plainlist ul li{margin-bottom:0}</style> <div class="side-box-flex"> <div class="side-box-image"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/30px-Commons-logo.svg.png" decoding="async" width="30" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/45px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/59px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></div> <div class="side-box-text plainlist">Wikimedia Commons has media related to <a href="https://commons.wikimedia.org/wiki/Category:Lichens" class="extiw" title="commons:Category:Lichens">Lichens</a>.</div></div> </div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1235681985"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1237033735"><div class="side-box side-box-right plainlinks sistersitebox"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"> <div class="side-box-flex"> <div class="side-box-image"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/99/Wiktionary-logo-en-v2.svg/40px-Wiktionary-logo-en-v2.svg.png" decoding="async" width="40" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/99/Wiktionary-logo-en-v2.svg/60px-Wiktionary-logo-en-v2.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/99/Wiktionary-logo-en-v2.svg/80px-Wiktionary-logo-en-v2.svg.png 2x" data-file-width="512" data-file-height="512" /></div> <div class="side-box-text plainlist">Look up <a href="https://en.wiktionary.org/wiki/Special:Search/lichen" class="extiw" title="wiktionary:Special:Search/lichen">lichen</a> in Wiktionary, the free dictionary.</div></div> </div> <div class="navbox-styles"><link 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