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dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Species of virus</div> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(250,250,190)"><i>Betacoronavirus gravedinis</i> </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:TEM_of_coronavirus_OC43.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/3d/TEM_of_coronavirus_OC43.jpg/220px-TEM_of_coronavirus_OC43.jpg" decoding="async" width="220" height="145" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/3d/TEM_of_coronavirus_OC43.jpg/330px-TEM_of_coronavirus_OC43.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/3d/TEM_of_coronavirus_OC43.jpg/440px-TEM_of_coronavirus_OC43.jpg 2x" data-file-width="700" data-file-height="460" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%"><a href="/wiki/Transmission_electron_micrograph" class="mw-redirect" title="Transmission electron micrograph">Transmission electron micrograph</a> of <a href="/wiki/Human_coronavirus_OC43" title="Human coronavirus OC43">human coronavirus OC43</a> </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(250,250,190)"><a href="/wiki/Virus_classification" title="Virus classification">Virus classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Embecovirus" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>(unranked): </td> <td><a href="/wiki/Virus" title="Virus">Virus</a> </td></tr> <tr> <td><i>Realm</i>: </td> <td><a href="/wiki/Riboviria" title="Riboviria"><i>Riboviria</i></a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Orthornavirae" title="Orthornavirae"><i>Orthornavirae</i></a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Pisuviricota" title="Pisuviricota"><i>Pisuviricota</i></a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Pisoniviricetes" title="Pisoniviricetes"><i>Pisoniviricetes</i></a> </td></tr> <tr> <td>Order: </td> <td><a href="/wiki/Nidovirales" title="Nidovirales"><i>Nidovirales</i></a> </td></tr> <tr> <td>Family: </td> <td><a href="/wiki/Coronaviridae" title="Coronaviridae"><i>Coronaviridae</i></a> </td></tr> <tr> <td>Genus: </td> <td><a href="/wiki/Betacoronavirus" title="Betacoronavirus"><i>Betacoronavirus</i></a> </td></tr> <tr> <td>Subgenus: </td> <td><a href="/wiki/Embecovirus" title="Embecovirus"><i>Embecovirus</i></a> </td></tr> <tr> <td>Species: </td> <td><div style="display:inline" class="species"><i><b>Betacoronavirus gravedinis</b></i></div> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(250,250,190)">Member viruses<sup id="cite_ref-ICTV_9_1-0" class="reference"><a href="#cite_note-ICTV_9-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </th></tr> <tr> <td colspan="2" style="text-align: left"> <ul><li><a href="/wiki/Bovine_coronavirus" title="Bovine coronavirus">Bovine coronavirus</a> <ul><li><a href="/w/index.php?title=Bovine_coronavirus_Mebus&action=edit&redlink=1" class="new" title="Bovine coronavirus Mebus (page does not exist)">Bovine coronavirus Mebus</a></li></ul></li> <li><a href="/w/index.php?title=Equine_coronavirus&action=edit&redlink=1" class="new" title="Equine coronavirus (page does not exist)">Equine coronavirus</a> <ul><li><a href="/w/index.php?title=Equine_coronavirus_NC99&action=edit&redlink=1" class="new" title="Equine coronavirus NC99 (page does not exist)">Equine coronavirus NC99</a></li></ul></li> <li><a href="/wiki/Human_coronavirus_OC43" title="Human coronavirus OC43">Human coronavirus OC43</a> <ul><li><a href="/w/index.php?title=Human_coronavirus_OC43_ATCC_VR-759&action=edit&redlink=1" class="new" title="Human coronavirus OC43 ATCC VR-759 (page does not exist)">Human coronavirus OC43 ATCC VR-759</a></li></ul></li> <li><a href="/w/index.php?title=Porcine_hemagglutinating_encephalomyelitis_virus&action=edit&redlink=1" class="new" title="Porcine hemagglutinating encephalomyelitis virus (page does not exist)">Porcine hemagglutinating encephalomyelitis virus</a> <ul><li><a href="/w/index.php?title=Porcine_hemagglutinating_encephalomyelitis_virus_VW572&action=edit&redlink=1" class="new" title="Porcine hemagglutinating encephalomyelitis virus VW572 (page does not exist)">Porcine hemagglutinating encephalomyelitis virus VW572</a></li></ul></li></ul> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(250,250,190)"><a href="/wiki/Synonym_(taxonomy)" title="Synonym (taxonomy)">Synonyms</a> </th></tr> <tr> <td colspan="2" style="text-align: left"> <ul><li>Betacoronavirus 1</li> <li><i>Bovine coronavirus</i><sup id="cite_ref-2008.100V_2-0" class="reference"><a href="#cite_note-2008.100V-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup></li> <li>canine respiratory coronavirus<sup id="cite_ref-2008.100V_2-1" class="reference"><a href="#cite_note-2008.100V-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup></li> <li><i>Equine coronavirus</i><sup id="cite_ref-2008.100V_2-2" class="reference"><a href="#cite_note-2008.100V-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup></li> <li><i>Human coronavirus OC43</i><sup id="cite_ref-2008.100V_2-3" class="reference"><a href="#cite_note-2008.100V-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup></li> <li><i>Human enteric coronavirus</i><sup id="cite_ref-2008.100V_2-4" class="reference"><a href="#cite_note-2008.100V-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup></li> <li><i>Porcine hemagglutinating encephalitis virus</i><sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup></li> <li><i>Porcine hemagglutinating encephalomyelitis virus</i><sup id="cite_ref-2008.100V_2-5" class="reference"><a href="#cite_note-2008.100V-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup></li></ul> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><i><b>Betacoronavirus gravedinis</b></i> <sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> (also called <i><b>Betacoronavirus 1</b></i>) is a species of <a href="/wiki/Coronavirus" title="Coronavirus">coronavirus</a> which infects humans and cattle.<sup id="cite_ref-ICTV_5-0" class="reference"><a href="#cite_note-ICTV-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> The infecting virus is an <a href="/wiki/Viral_envelope" title="Viral envelope">enveloped</a>, <a href="/wiki/Sense_(molecular_biology)" title="Sense (molecular biology)">positive-sense</a>, <a href="/wiki/RNA" title="RNA">single-stranded</a> <a href="/wiki/RNA_virus" title="RNA virus">RNA virus</a> and is a member of the genus <i><a href="/wiki/Betacoronavirus" title="Betacoronavirus">Betacoronavirus</a></i> and subgenus <i><a href="/wiki/Embecovirus" title="Embecovirus">Embecovirus</a>.<sup id="cite_ref-:0_7-0" class="reference"><a href="#cite_note-:0-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup></i> Like other embecoviruses, it has an additional shorter <a href="/wiki/Peplomer" class="mw-redirect" title="Peplomer">spike</a>-like surface protein called <a href="/wiki/Hemagglutinin_esterase" title="Hemagglutinin esterase">hemagglutinin esterase</a> (HE) as well as the larger <a href="/wiki/Coronavirus_spike_protein" title="Coronavirus spike protein">coronavirus spike protein</a>.<sup id="cite_ref-:03_8-0" class="reference"><a href="#cite_note-:03-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:022_9-0" class="reference"><a href="#cite_note-:022-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Betacoronavirus_1&action=edit&section=1" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-ICTV_9-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-ICTV_9_1-0">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFde_Groot2009" class="citation web cs1">de Groot, R.J; et al. (2009). <a rel="nofollow" class="external text" href="https://ictv.global/report_9th/RNApos/Nidovirales/Coronaviridae">"ICTV 9th Report (2011) New <i>Coronaviridae</i>"</a>. <i>International Committee on Taxonomy of Viruses (ICTV)</i>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20200123162412/https://talk.ictvonline.org/ictv-reports/ictv_9th_report/positive-sense-rna-viruses-2011/w/posrna_viruses/222/coronaviridae">Archived</a> from the original on 23 January 2020<span class="reference-accessdate">. Retrieved <span class="nowrap">23 January</span> 2020</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=International+Committee+on+Taxonomy+of+Viruses+%28ICTV%29&rft.atitle=ICTV+9th+Report+%282011%29+New+Coronaviridae&rft.date=2009&rft.aulast=de+Groot&rft.aufirst=R.J&rft_id=https%3A%2F%2Fictv.global%2Freport_9th%2FRNApos%2FNidovirales%2FCoronaviridae&rfr_id=info%3Asid%2Fen.wikipedia.org%3ABetacoronavirus+1" class="Z3988"></span></span> </li> <li id="cite_note-2008.100V-2"><span class="mw-cite-backlink">^ <a href="#cite_ref-2008.100V_2-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-2008.100V_2-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-2008.100V_2-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-2008.100V_2-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-2008.100V_2-4"><sup><i><b>e</b></i></sup></a> <a href="#cite_ref-2008.100V_2-5"><sup><i><b>f</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFde_Groot2009" class="citation web cs1">de Groot, Raoul J.; et al. (2009). <a rel="nofollow" class="external text" href="https://ictv.global/ictv/proposals/2008.085-122V.v4.Coronaviridae.pdf">"Revision of the family <i>Coronaviridae</i>"</a> <span class="cs1-format">(PDF)</span>. <i>International Committee on Taxonomy of Viruses (ICTV)</i>. p. 18. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20190207015126/https://talk.ictvonline.org/ICTV/proposals/2008.085-122V.v4.Coronaviridae.pdf">Archived</a> <span class="cs1-format">(PDF)</span> from the original on 7 February 2019<span class="reference-accessdate">. Retrieved <span class="nowrap">23 January</span> 2020</span>. <q>A new species, <i>Betacoronavirus 1</i>, is proposed to accommodate previously established <i>Betacoronavirus</i> species of humans, cattle, swine and horses (<i>Human enteric coronavirus</i>; <i>Human coronavirus OC43</i>; <i>Bovine coronavirus</i>; <i>Porcine hemagglutinating encephalomyelitis virus</i>; <i>Equine coronavirus</i>) and a newly recognized virus of dogs (canine respiratory coronavirus; Erles et al., 2007)</q></cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=International+Committee+on+Taxonomy+of+Viruses+%28ICTV%29&rft.atitle=Revision+of+the+family+Coronaviridae&rft.pages=18&rft.date=2009&rft.aulast=de+Groot&rft.aufirst=Raoul+J.&rft_id=https%3A%2F%2Fictv.global%2Fictv%2Fproposals%2F2008.085-122V.v4.Coronaviridae.pdf&rfr_id=info%3Asid%2Fen.wikipedia.org%3ABetacoronavirus+1" class="Z3988"></span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-3">^</a></b></span> <span class="reference-text">ICTV 5th Report Francki, R. I. B., Fauquet, C. M., Knudson, D. L. & Brown, F. (eds)(1991). Classification and nomenclature of viruses. Fifthreport of the International Committee on Taxonomy of Viruses. 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Retrieved <span class="nowrap">23 January</span> 2020</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=International+Committee+on+Taxonomy+of+Viruses+%28ICTV%29&rft.atitle=Virus+Taxonomy%3A+2018b+Release&rft.date=2019-03&rft_id=https%3A%2F%2Fictv.global%2Ftaxonomy&rfr_id=info%3Asid%2Fen.wikipedia.org%3ABetacoronavirus+1" class="Z3988"></span></span> </li> <li id="cite_note-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-6">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFLimNgTamLiu2016" class="citation journal cs1">Lim, Yvonne Xinyi; Ng, Yan Ling; Tam, James P.; Liu, Ding Xiang (2016-07-25). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5456285">"Human Coronaviruses: A Review of Virus–Host Interactions"</a>. <i>Diseases</i>. <b>4</b> (3): 26. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.3390%2Fdiseases4030026">10.3390/diseases4030026</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/2079-9721">2079-9721</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5456285">5456285</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/28933406">28933406</a>. <q>See Table 1.</q></cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Diseases&rft.atitle=Human+Coronaviruses%3A+A+Review+of+Virus%E2%80%93Host+Interactions&rft.volume=4&rft.issue=3&rft.pages=26&rft.date=2016-07-25&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5456285%23id-name%3DPMC&rft.issn=2079-9721&rft_id=info%3Apmid%2F28933406&rft_id=info%3Adoi%2F10.3390%2Fdiseases4030026&rft.aulast=Lim&rft.aufirst=Yvonne+Xinyi&rft.au=Ng%2C+Yan+Ling&rft.au=Tam%2C+James+P.&rft.au=Liu%2C+Ding+Xiang&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5456285&rfr_id=info%3Asid%2Fen.wikipedia.org%3ABetacoronavirus+1" class="Z3988"></span></span> </li> <li id="cite_note-:0-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-:0_7-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Undef&id=694003&lvl=3&keep=1&srchmode=1&unlock">"Taxonomy browser (Betacoronavirus 1)"</a>. <i>www.ncbi.nlm.nih.gov</i><span class="reference-accessdate">. Retrieved <span class="nowrap">2020-02-29</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=www.ncbi.nlm.nih.gov&rft.atitle=Taxonomy+browser+%28Betacoronavirus+1%29&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2FTaxonomy%2FBrowser%2Fwwwtax.cgi%3Fmode%3DUndef%26id%3D694003%26lvl%3D3%26keep%3D1%26srchmode%3D1%26unlock&rfr_id=info%3Asid%2Fen.wikipedia.org%3ABetacoronavirus+1" class="Z3988"></span></span> </li> <li id="cite_note-:03-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-:03_8-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWooHuangLauYuen2010" class="citation journal cs1">Woo, Patrick C. Y.; Huang, Yi; Lau, Susanna K. P.; Yuen, Kwok-Yung (2010-08-24). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3185738">"Coronavirus Genomics and Bioinformatics Analysis"</a>. <i>Viruses</i>. <b>2</b> (8): 1804–1820. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.3390%2Fv2081803">10.3390/v2081803</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/1999-4915">1999-4915</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3185738">3185738</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/21994708">21994708</a>. <q>In all members of Betacoronavirus subgroup A, a haemagglutinin esterase (HE) gene, which encodes a glycoprotein with neuraminate O-acetyl-esterase activity and the active site FGDS, is present downstream to ORF1ab and upstream to S gene (Figure 1).</q></cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Viruses&rft.atitle=Coronavirus+Genomics+and+Bioinformatics+Analysis&rft.volume=2&rft.issue=8&rft.pages=1804-1820&rft.date=2010-08-24&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3185738%23id-name%3DPMC&rft.issn=1999-4915&rft_id=info%3Apmid%2F21994708&rft_id=info%3Adoi%2F10.3390%2Fv2081803&rft.aulast=Woo&rft.aufirst=Patrick+C.+Y.&rft.au=Huang%2C+Yi&rft.au=Lau%2C+Susanna+K.+P.&rft.au=Yuen%2C+Kwok-Yung&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3185738&rfr_id=info%3Asid%2Fen.wikipedia.org%3ABetacoronavirus+1" class="Z3988"></span></span> </li> <li id="cite_note-:022-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-:022_9-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWooHuangLauYuen2010" class="citation journal cs1">Woo, Patrick C. Y.; Huang, Yi; Lau, Susanna K. P.; Yuen, Kwok-Yung (2010-08-24). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3185738">"Coronavirus Genomics and Bioinformatics Analysis"</a>. <i>Viruses</i>. <b>2</b> (8): 1804–1820. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.3390%2Fv2081803">10.3390/v2081803</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/1999-4915">1999-4915</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3185738">3185738</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/21994708">21994708</a>. <q>The presence of HE genes exclusively in members of Betacoronavirus subgroup A, but not members of Betacoronavirus subgroup B, C and D suggested that the recombination had probably occurred in the ancestor of members of Betacoronavirus subgroup A, after diverging from the ancestor of other subgroups of Betacoronavirus.</q></cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Viruses&rft.atitle=Coronavirus+Genomics+and+Bioinformatics+Analysis&rft.volume=2&rft.issue=8&rft.pages=1804-1820&rft.date=2010-08-24&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3185738%23id-name%3DPMC&rft.issn=1999-4915&rft_id=info%3Apmid%2F21994708&rft_id=info%3Adoi%2F10.3390%2Fv2081803&rft.aulast=Woo&rft.aufirst=Patrick+C.+Y.&rft.au=Huang%2C+Yi&rft.au=Lau%2C+Susanna+K.+P.&rft.au=Yuen%2C+Kwok-Yung&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3185738&rfr_id=info%3Asid%2Fen.wikipedia.org%3ABetacoronavirus+1" class="Z3988"></span></span> </li> </ol></div></div> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist 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href="/wiki/Template:Coronaviridae" title="Template:Coronaviridae"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Coronaviridae" title="Template talk:Coronaviridae"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Coronaviridae" title="Special:EditPage/Template:Coronaviridae"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Taxonomy_of_the_Coronaviridae" style="font-size:114%;margin:0 4em">Taxonomy of the <i><a href="/wiki/Coronaviridae" title="Coronaviridae">Coronaviridae</a></i></div></th></tr><tr><td class="navbox-abovebelow" colspan="2"><div>Higher taxonomy: <i><a href="/wiki/Riboviria" title="Riboviria">Riboviria</a></i> > <i><a href="/wiki/Orthornavirae" title="Orthornavirae">Orthornavirae</a></i> > <i><a href="/wiki/Pisuviricota" title="Pisuviricota">Pisuviricota</a></i> > <i><a href="/wiki/Pisoniviricetes" title="Pisoniviricetes">Pisoniviricetes</a></i> > <i><a href="/wiki/Nidovirales" title="Nidovirales">Nidovirales</a></i> > <i><a href="/wiki/Cornidovirineae" class="mw-redirect" title="Cornidovirineae">Cornidovirineae</a></i> > <i>Coronaviridae</i></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible expanded navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Orthocoronavirinae" style="font-size:114%;margin:0 4em"><i><a href="/wiki/Orthocoronavirinae" class="mw-redirect" title="Orthocoronavirinae">Orthocoronavirinae</a></i></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Alphacoronavirus" title="Alphacoronavirus">Alphacoronavirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Colacovirus" title="Colacovirus">Colacovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>Bat coronavirus CDPHE15</i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Decacovirus" title="Decacovirus">Decacovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Bat_coronavirus_HKU10" title="Bat coronavirus HKU10">Bat coronavirus HKU10</a></i></li> <li><i><a href="/wiki/Rhinolophus_ferrumequinum_alphacoronavirus_HuB-2013" title="Rhinolophus ferrumequinum alphacoronavirus HuB-2013">Rhinolophus ferrumequinum alphacoronavirus HuB-2013</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Duvinacovirus" class="mw-redirect" title="Duvinacovirus">Duvinacovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>Human coronavirus 229E</i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Luchacovirus" title="Luchacovirus">Luchacovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>Lucheng Rn rat coronavirus</i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Minacovirus" title="Minacovirus">Minacovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>Mink coronavirus 1</i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Minunacovirus" title="Minunacovirus">Minunacovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Miniopterus_bat_coronavirus_1" title="Miniopterus bat coronavirus 1">Miniopterus bat coronavirus 1</a></i></li> <li><i><a href="/wiki/Miniopterus_bat_coronavirus_HKU8" title="Miniopterus bat coronavirus HKU8">Miniopterus bat coronavirus HKU8</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Myotacovirus" title="Myotacovirus">Myotacovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>Myotis ricketti alphacoronavirus Sax-2011</i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Nyctacovirus" title="Nyctacovirus">Nyctacovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Nyctalus_velutinus_alphacoronavirus_SC-2013" title="Nyctalus velutinus alphacoronavirus SC-2013">Nyctalus velutinus alphacoronavirus SC-2013</a></i></li> <li><i><a href="/wiki/Pipistrellus_kuhlii_coronavirus_3398" title="Pipistrellus kuhlii coronavirus 3398">Pipistrellus kuhlii coronavirus 3398</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Pedacovirus" title="Pedacovirus">Pedacovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Porcine_epidemic_diarrhea_virus" title="Porcine epidemic diarrhea virus">Porcine epidemic diarrhea virus</a></i></li> <li><i><a href="/wiki/Scotophilus_bat_coronavirus_512" title="Scotophilus bat coronavirus 512">Scotophilus bat coronavirus 512</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Rhinacovirus" class="mw-redirect" title="Rhinacovirus">Rhinacovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>Rhinolophus bat coronavirus HKU2</i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Setracovirus" title="Setracovirus">Setracovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Human_coronavirus_NL63" title="Human coronavirus NL63">Human coronavirus NL63</a></i></li> <li><i><a href="/wiki/NL63-related_bat_coronavirus_strain_BtKYNL63-9b" title="NL63-related bat coronavirus strain BtKYNL63-9b">NL63-related bat coronavirus strain BtKYNL63-9b</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Soracovirus" title="Soracovirus">Soracovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>Sorex araneus coronavirus T14</i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Sunacovirus" title="Sunacovirus">Sunacovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>Suncus murinus coronavirus X74</i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Tegacovirus" class="mw-redirect" title="Tegacovirus">Tegacovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>Alphacoronavirus 1</i> <ul><li><a href="/wiki/Feline_coronavirus" title="Feline coronavirus">Feline coronavirus</a></li> <li><a href="/wiki/Canine_coronavirus" title="Canine coronavirus">Canine coronavirus</a></li> <li><a href="/wiki/Canine_coronavirus_HuPn-2018" title="Canine coronavirus HuPn-2018">Canine coronavirus HuPn-2018</a></li> <li><a href="/wiki/Transmissible_gastroenteritis_virus" title="Transmissible gastroenteritis virus">Transmissible gastroenteritis virus</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Betacoronavirus" title="Betacoronavirus">Betacoronavirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Embecovirus" title="Embecovirus">Embecovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a class="mw-selflink selflink">Betacoronavirus 1</a></i> <ul><li><a href="/wiki/Bovine_coronavirus" title="Bovine coronavirus">Bovine coronavirus</a></li> <li><a href="/wiki/Human_coronavirus_OC43" title="Human coronavirus OC43">Human coronavirus OC43</a></li></ul></li> <li><i><a href="/wiki/China_Rattus_coronavirus_HKU24" title="China Rattus coronavirus HKU24">China Rattus coronavirus HKU24</a></i></li> <li><i><a href="/wiki/Human_coronavirus_HKU1" title="Human coronavirus HKU1">Human coronavirus HKU1</a></i></li> <li><i><a href="/wiki/Murine_coronavirus" title="Murine coronavirus">Murine coronavirus</a></i> <ul><li><a href="/wiki/Rat_coronavirus" title="Rat coronavirus">Rat coronavirus</a></li></ul></li> <li><i><a href="/wiki/Myodes_coronavirus_2JL14" title="Myodes coronavirus 2JL14">Myodes coronavirus 2JL14</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Hibecovirus" title="Hibecovirus">Hibecovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>Bat Hp-betacoronavirus Zhejiang2013</i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Merbecovirus" title="Merbecovirus">Merbecovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Hedgehog_coronavirus_1" title="Hedgehog coronavirus 1">Hedgehog coronavirus 1</a></i></li> <li><i><a href="/wiki/Middle_East_respiratory_syndrome-related_coronavirus" class="mw-redirect" title="Middle East respiratory syndrome-related coronavirus">Middle East respiratory syndrome-related coronavirus</a></i> <ul><li><a href="/wiki/London1_novel_CoV/2012" title="London1 novel CoV/2012">London1_novel CoV/2012</a></li> <li><a href="/wiki/MERS_coronavirus_EMC/2012" title="MERS coronavirus EMC/2012">MERS coronavirus EMC/2012</a></li></ul></li> <li><i><a href="/wiki/Pipistrellus_bat_coronavirus_HKU5" title="Pipistrellus bat coronavirus HKU5">Pipistrellus bat coronavirus HKU5</a></i></li> <li><i><a href="/wiki/Tylonycteris_bat_coronavirus_HKU4" title="Tylonycteris bat coronavirus HKU4">Tylonycteris bat coronavirus HKU4</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Nobecovirus" title="Nobecovirus">Nobecovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Eidolon_bat_coronavirus_C704" title="Eidolon bat coronavirus C704">Eidolon bat coronavirus C704</a></i></li> <li><i><a href="/wiki/Rousettus_bat_coronavirus_GCCDC1" title="Rousettus bat coronavirus GCCDC1">Rousettus bat coronavirus GCCDC1</a></i></li> <li><i><a href="/wiki/Rousettus_bat_coronavirus_HKU9" title="Rousettus bat coronavirus HKU9">Rousettus bat coronavirus HKU9</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Sarbecovirus" class="mw-redirect" title="Sarbecovirus">Sarbecovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/SARS-related_coronavirus" title="SARS-related coronavirus">SARS-related coronavirus</a></i> <ul><li><a href="/wiki/Severe_acute_respiratory_syndrome_coronavirus_1" class="mw-redirect" title="Severe acute respiratory syndrome coronavirus 1">SARS-CoV-1</a></li> <li><a href="/wiki/SARS-CoV-2" title="SARS-CoV-2">SARS-CoV-2</a> <ul><li><a href="/wiki/Variants_of_SARS-CoV-2" title="Variants of SARS-CoV-2">Variants</a></li></ul></li> <li><a href="/w/index.php?title=BANAL-52&action=edit&redlink=1" class="new" title="BANAL-52 (page does not exist)">BANAL-52</a></li> <li><a href="/wiki/Bat_SARS-like_coronavirus_WIV1" title="Bat SARS-like coronavirus WIV1">Bat SARS-like coronavirus WIV1</a></li> <li><a href="/wiki/RacCS203" title="RacCS203">RacCS203</a></li> <li><a href="/wiki/RaTG13" title="RaTG13">RaTG13</a></li> <li><a href="/wiki/RmYN02" title="RmYN02">RmYN02</a></li> <li><a href="/wiki/RpYN06" title="RpYN06">RpYN06</a></li> <li><a href="/wiki/SHC014-CoV" title="SHC014-CoV">SHC014-CoV</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Gammacoronavirus" title="Gammacoronavirus">Gammacoronavirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Brangacovirus" title="Brangacovirus">Brangacovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>Goose coronavirus CB17</i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Cegacovirus" class="mw-redirect" title="Cegacovirus">Cegacovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Beluga_whale_coronavirus_SW1" title="Beluga whale coronavirus SW1">Beluga whale coronavirus SW1</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Igacovirus" title="Igacovirus">Igacovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Avian_coronavirus" title="Avian coronavirus">Avian coronavirus</a></i></li> <li><i><a href="/wiki/Avian_coronavirus_9203" title="Avian coronavirus 9203">Avian coronavirus 9203</a></i></li> <li><i><a href="/wiki/Duck_coronavirus_2714" title="Duck coronavirus 2714">Duck coronavirus 2714</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Deltacoronavirus" title="Deltacoronavirus">Deltacoronavirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Andecovirus" title="Andecovirus">Andecovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>Wigeon coronavirus HKU20</i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Buldecovirus" title="Buldecovirus">Buldecovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Bulbul_coronavirus_HKU11" title="Bulbul coronavirus HKU11">Bulbul coronavirus HKU11</a></i></li> <li><i><a href="/wiki/Common_moorhen_coronavirus_HKU21" title="Common moorhen coronavirus HKU21">Common moorhen coronavirus HKU21</a></i></li> <li><i><a href="/wiki/Coronavirus_HKU15" title="Coronavirus HKU15">Coronavirus HKU15</a></i></li> <li><i><a href="/wiki/Munia_coronavirus_HKU13" title="Munia coronavirus HKU13">Munia coronavirus HKU13</a></i></li> <li><i><a href="/wiki/White-eye_coronavirus_HKU16" title="White-eye coronavirus HKU16">White-eye coronavirus HKU16</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:8em"><i><a href="/wiki/Herdecovirus" title="Herdecovirus">Herdecovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>Night heron coronavirus HKU19</i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible expanded navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Letovirinae" style="font-size:114%;margin:0 4em"><i><a href="/wiki/Letovirinae" title="Letovirinae">Letovirinae</a></i></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Alphaletovirus" scope="row" class="navbox-group" style="width:9em"><i>Alphaletovirus</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Milecovirus" scope="row" class="navbox-group" style="width:8em"><i>Milecovirus</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>Microhyla letovirus 1</i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div>Source: <a href="/wiki/International_Committee_on_Taxonomy_of_Viruses" title="International Committee on Taxonomy of Viruses">ICTV</a> –– <span class="noviewer" typeof="mw:File"><a href="/wiki/File:Wikispecies-logo.svg" class="mw-file-description" title="Wikispecies page"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/14px-Wikispecies-logo.svg.png" decoding="async" width="14" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/20px-Wikispecies-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/27px-Wikispecies-logo.svg.png 2x" data-file-width="941" data-file-height="1103" /></a></span> <a href="https://species.wikimedia.org/wiki/Special:Search/Coronaviridae" class="extiw" title="wikispecies:Special:Search/Coronaviridae">Wikispecies</a></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Betacoronavirus 1</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q18965332" class="extiw" title="wikidata:Q18965332">Q18965332</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Betacoronavirus_1" class="extiw" title="wikispecies:Betacoronavirus 1">Betacoronavirus 1</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Catalogue_of_Life" title="Catalogue of Life">CoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.catalogueoflife.org/data/taxon/LNPX">LNPX</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/11621326">11621326</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&id=11459240">11459240</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=694003">694003</a></span></span></li></ul> 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