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class="vector-toc-numb">2</span> <span>Anatomy and morphology</span> </div> </a> <ul id="toc-Anatomy_and_morphology-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Habitat" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Habitat"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Habitat</span> </div> </a> <ul id="toc-Habitat-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Behavior" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Behavior"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Behavior</span> </div> </a> <ul id="toc-Behavior-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Reproduction" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Reproduction"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>Reproduction</span> </div> </a> <ul id="toc-Reproduction-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Diet" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Diet"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>Diet</span> </div> </a> <ul id="toc-Diet-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-In_captivity" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#In_captivity"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>In captivity</span> </div> </a> <ul id="toc-In_captivity-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">8</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet 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Available in 41 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-41" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">41 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D9%85%D8%A7%D8%AA%D8%A7%D9%85%D8%A7%D8%AA%D8%A7_(%D8%B3%D9%84%D8%AD%D9%81%D8%A7%D8%A9)" title="ماتاماتا (سلحفاة) – Arabic" lang="ar" hreflang="ar" data-title="ماتاماتا (سلحفاة)" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-gn mw-list-item"><a href="https://gn.wikipedia.org/wiki/Matamata" title="Matamata – Guarani" lang="gn" hreflang="gn" data-title="Matamata" data-language-autonym="Avañe'ẽ" data-language-local-name="Guarani" class="interlanguage-link-target"><span>Avañe'ẽ</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Sa%C3%A7aql%C4%B1_t%C4%B1sba%C4%9Fa" title="Saçaqlı tısbağa – Azerbaijani" lang="az" hreflang="az" data-title="Saçaqlı tısbağa" data-language-autonym="Azərbaycanca" data-language-local-name="Azerbaijani" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%AE%E0%A6%BE%E0%A6%9F%E0%A6%BE_%E0%A6%AE%E0%A6%BE%E0%A6%9F%E0%A6%BE" title="মাটা মাটা – Bangla" lang="bn" hreflang="bn" data-title="মাটা মাটা" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%9C%D0%B0%D1%82%D0%B0%D0%BC%D0%B0%D1%82%D0%B0" title="Матамата – Bulgarian" lang="bg" hreflang="bg" data-title="Матамата" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Matamata" title="Matamata – Catalan" lang="ca" hreflang="ca" data-title="Matamata" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-ceb mw-list-item"><a href="https://ceb.wikipedia.org/wiki/Chelus_fimbriatus" title="Chelus fimbriatus – Cebuano" lang="ceb" hreflang="ceb" data-title="Chelus fimbriatus" data-language-autonym="Cebuano" data-language-local-name="Cebuano" class="interlanguage-link-target"><span>Cebuano</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Matamata_t%C5%99%C3%A1snit%C3%A1" title="Matamata třásnitá – Czech" lang="cs" hreflang="cs" data-title="Matamata třásnitá" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Fransenschildkr%C3%B6te" title="Fransenschildkröte – German" lang="de" hreflang="de" data-title="Fransenschildkröte" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-nv mw-list-item"><a href="https://nv.wikipedia.org/wiki/Tsistee%C5%82_diyog%C3%AD" title="Tsisteeł diyogí – Navajo" lang="nv" hreflang="nv" data-title="Tsisteeł diyogí" data-language-autonym="Diné bizaad" data-language-local-name="Navajo" class="interlanguage-link-target"><span>Diné bizaad</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Chelus_fimbriata" title="Chelus fimbriata – Spanish" lang="es" hreflang="es" data-title="Chelus fimbriata" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Matamata" title="Matamata – Basque" lang="eu" hreflang="eu" data-title="Matamata" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D9%85%D8%A7%D8%AA%D8%A7%D9%85%D8%A7%D8%AA%D8%A7" title="ماتاماتا – Persian" lang="fa" hreflang="fa" data-title="ماتاماتا" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr badge-Q17437796 badge-featuredarticle mw-list-item" title="featured article badge"><a href="https://fr.wikipedia.org/wiki/Matamata" title="Matamata – French" lang="fr" hreflang="fr" data-title="Matamata" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Chelus_fimbriatus" title="Chelus fimbriatus – Galician" lang="gl" hreflang="gl" data-title="Chelus fimbriatus" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EB%A7%88%ED%83%80%EB%A7%88%ED%83%80%EA%B1%B0%EB%B6%81" title="마타마타거북 – Korean" lang="ko" hreflang="ko" data-title="마타마타거북" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Mata_mata" title="Mata mata – Croatian" lang="hr" hreflang="hr" data-title="Mata mata" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Chelus_fimbriata" title="Chelus fimbriata – Italian" lang="it" hreflang="it" data-title="Chelus fimbriata" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%9E%D7%98%D7%90%D7%9E%D7%98%D7%94" title="מטאמטה – Hebrew" lang="he" hreflang="he" data-title="מטאמטה" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-ka mw-list-item"><a href="https://ka.wikipedia.org/wiki/%E1%83%9B%E1%83%90%E1%83%A2%E1%83%90%E1%83%9B%E1%83%90%E1%83%A2%E1%83%90" title="მატამატა – Georgian" lang="ka" hreflang="ka" data-title="მატამატა" data-language-autonym="ქართული" data-language-local-name="Georgian" class="interlanguage-link-target"><span>ქართული</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Skarmalotasis_v%C4%97%C5%BElys" title="Skarmalotasis vėžlys – Lithuanian" lang="lt" hreflang="lt" data-title="Skarmalotasis vėžlys" data-language-autonym="Lietuvių" data-language-local-name="Lithuanian" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Cafrangos_tekn%C5%91s" title="Cafrangos teknős – Hungarian" lang="hu" hreflang="hu" data-title="Cafrangos teknős" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D9%85%D8%A7%D8%AA%D8%A7_%D9%85%D8%A7%D8%AA%D8%A7" title="ماتا ماتا – Egyptian Arabic" lang="arz" hreflang="arz" data-title="ماتا ماتا" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Mata_mata" title="Mata mata – Malay" lang="ms" hreflang="ms" data-title="Mata mata" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Matamata_(schildpad)" title="Matamata (schildpad) – Dutch" lang="nl" hreflang="nl" data-title="Matamata (schildpad)" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%83%9E%E3%82%BF%E3%83%9E%E3%82%BF" title="マタマタ – Japanese" lang="ja" hreflang="ja" data-title="マタマタ" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Matamata_(skilpadde)" title="Matamata (skilpadde) – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Matamata (skilpadde)" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D9%85%D8%A7%D9%B9%D8%A7_%D9%85%D8%A7%D9%B9%D8%A7" title="ماٹا ماٹا – Western Punjabi" lang="pnb" hreflang="pnb" data-title="ماٹا ماٹا" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Matamata" title="Matamata – Polish" lang="pl" hreflang="pl" data-title="Matamata" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Mata-mata" title="Mata-mata – Portuguese" lang="pt" hreflang="pt" data-title="Mata-mata" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-qu mw-list-item"><a href="https://qu.wikipedia.org/wiki/Mata-mata" title="Mata-mata – Quechua" lang="qu" hreflang="qu" data-title="Mata-mata" data-language-autonym="Runa Simi" data-language-local-name="Quechua" class="interlanguage-link-target"><span>Runa Simi</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%91%D0%B0%D1%85%D1%80%D0%BE%D0%BC%D1%87%D0%B0%D1%82%D0%B0%D1%8F_%D1%87%D0%B5%D1%80%D0%B5%D0%BF%D0%B0%D1%85%D0%B0" title="Бахромчатая черепаха – Russian" lang="ru" hreflang="ru" data-title="Бахромчатая черепаха" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Mata_mata" title="Mata mata – Simple English" lang="en-simple" hreflang="en-simple" data-title="Mata mata" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%9C%D0%B0%D1%82%D0%B0_%D0%BC%D0%B0%D1%82%D0%B0" title="Мата мата – Serbian" lang="sr" hreflang="sr" data-title="Мата мата" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Matamata_(kilpikonna)" title="Matamata (kilpikonna) – Finnish" lang="fi" hreflang="fi" data-title="Matamata (kilpikonna)" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Matamata_(sk%C3%B6ldpadda)" title="Matamata (sköldpadda) – Swedish" lang="sv" hreflang="sv" data-title="Matamata (sköldpadda)" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-th mw-list-item"><a href="https://th.wikipedia.org/wiki/%E0%B9%80%E0%B8%95%E0%B9%88%E0%B8%B2%E0%B8%A1%E0%B8%B2%E0%B8%95%E0%B8%B2%E0%B8%A1%E0%B8%B2%E0%B8%95%E0%B8%B2" title="เต่ามาตามาตา – Thai" lang="th" hreflang="th" data-title="เต่ามาตามาตา" data-language-autonym="ไทย" data-language-local-name="Thai" class="interlanguage-link-target"><span>ไทย</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%9C%D0%B0%D1%82%D0%B0%D0%BC%D0%B0%D1%82%D0%B0_%D0%B7%D0%B2%D0%B8%D1%87%D0%B0%D0%B9%D0%BD%D0%B0" title="Матамата звичайна – Ukrainian" lang="uk" hreflang="uk" data-title="Матамата звичайна" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Chelus_fimbriatus" title="Chelus fimbriatus – Vietnamese" lang="vi" hreflang="vi" data-title="Chelus fimbriatus" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-war mw-list-item"><a href="https://war.wikipedia.org/wiki/Chelus_fimbriatus" title="Chelus fimbriatus – Waray" lang="war" hreflang="war" data-title="Chelus fimbriatus" data-language-autonym="Winaray" data-language-local-name="Waray" class="interlanguage-link-target"><span>Winaray</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E6%A5%93%E8%91%89%E9%BE%9C" title="楓葉龜 – Chinese" lang="zh" hreflang="zh" data-title="楓葉龜" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q317386#sitelinks-wikipedia" title="Edit interlanguage links" class="wbc-editpage">Edit links</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Namespaces"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Mata_mata" title="View the content page [c]" accesskey="c"><span>Article</span></a></li><li id="ca-talk" class="vector-tab-noicon mw-list-item"><a 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<div class="mw-indicators"> </div> <div id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Species of freshwater turtle</div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">For other uses, see <a href="/wiki/Matamata_(disambiguation)" class="mw-disambig" title="Matamata (disambiguation)">Matamata (disambiguation)</a>.</div> <p class="mw-empty-elt"> </p> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">Mata mata<br /><div style="font-size: 85%;">Temporal range: <span class="noprint"><span style="display:inline-block;"></span><span style="display:inline-block;">Pliocene–Recent</span> <span style="display:inline-block;"></span><div id="Timeline-row" style="margin: 4px auto 0; clear:both; width:220px; padding:0px; height:18px; overflow:visible; white-space:nowrap; border:1px #666; border-style:solid none; position:relative; z-index:0; font-size:97%;"> <div style="position:absolute; height:100%; left:0px; width:207.23076923077px; padding-left:5px; text-align:left; background-color:rgb(254,217,106); background-image: linear-gradient(to right, rgba(255,255,255,1), rgba(254,217,106,1) 15%, rgba(254,217,106,1));"><a href="/wiki/Precambrian" title="Precambrian">PreꞒ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,160,86); left:37.636923076923px; width:18.073846153846px;"><a href="/wiki/Cambrian" title="Cambrian">Ꞓ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(0,146,112); left:55.710769230769px; width:14.08px;"><a href="/wiki/Ordovician" title="Ordovician">O</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(179,225,182); left:69.790769230769px; width:8.3261538461539px;"><a href="/wiki/Silurian" title="Silurian">S</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(203,140,55); left:78.116923076923px; width:20.409230769231px;"><a href="/wiki/Devonian" title="Devonian">D</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(103,165,153); left:98.526153846154px; width:20.307692307692px;"><a href="/wiki/Carboniferous" title="Carboniferous">C</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(240,64,40); left:118.83384615385px; width:15.907015384615px;"><a href="/wiki/Permian" title="Permian">P</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(129,43,146); left:134.74086153846px; width:17.092984615385px;"><a href="/wiki/Triassic" title="Triassic">T</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(52,178,201); left:151.83384615385px; width:19.089230769231px;"><a href="/wiki/Jurassic" title="Jurassic">J</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,198,78); left:170.92307692308px; width:26.738461538462px;"><a href="/wiki/Cretaceous" title="Cretaceous">K</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(253,154,82); left:197.66153846154px; width:14.543692307692px;"><a href="/wiki/Paleogene" title="Paleogene">Pg</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(255,230,25); left:212.20523076923px; width:6.9215384615385px;"><a href="/wiki/Neogene" title="Neogene">N</a></div> <div id="end-border" style="position:absolute; height:100%; background-color:#666; width:1px; left:219px"></div><div style="margin:0 auto; line-height:0; clear:both; width:220px; padding:0px; height:8px; overflow:visible; background-color:transparent; position:relative; top:-4px; z-index:100;"><div style="position:absolute; height:8px; left:218.19498461538px; width:1.8050153846154px; background-color:#360; opacity:0.42;"></div> <div style="position:absolute; height:8px; left:218.19498461538px; width:1.8050153846154px; background-color:#360; opacity:1;"></div> <div style="position:absolute; height:6px; top:1px; left:219.19498461538px; width:-0.19498461538462px; background-color:#6c3;"></div> </div> </div></span></div> </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:2009_Chelus_fimbriatus.JPG" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/7f/2009_Chelus_fimbriatus.JPG/220px-2009_Chelus_fimbriatus.JPG" decoding="async" width="220" height="147" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/7f/2009_Chelus_fimbriatus.JPG/330px-2009_Chelus_fimbriatus.JPG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/7f/2009_Chelus_fimbriatus.JPG/440px-2009_Chelus_fimbriatus.JPG 2x" data-file-width="4272" data-file-height="2848" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%">In Shanghai Aquarium, China </td></tr> <tr style="color:inherit; text-align: center; background-color: rgb(235,235,210)"> <th colspan="2"><div style="text-align: center"><a href="/wiki/Conservation_status" title="Conservation status">Conservation status</a></div> </th></tr> <tr> <td colspan="2"><div style="text-align: center"><span class="mw-default-size skin-invert-image" typeof="mw:File/Frameless"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/5/5a/Status_iucn3.1_LC.svg/220px-Status_iucn3.1_LC.svg.png" decoding="async" width="220" height="59" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/5a/Status_iucn3.1_LC.svg/330px-Status_iucn3.1_LC.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/5a/Status_iucn3.1_LC.svg/440px-Status_iucn3.1_LC.svg.png 2x" data-file-width="512" data-file-height="137" /></span></span><br /><a href="/wiki/Least_Concern" class="mw-redirect" title="Least Concern">Least Concern</a> <small> (<a href="/wiki/IUCN_Red_List" title="IUCN Red List">IUCN 3.1</a>)<sup id="cite_ref-ttwg_1-0" class="reference"><a href="#cite_note-ttwg-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup></small></div> </td></tr> <tr> <td colspan="2"><div style="text-align: center"><a href="/wiki/CITES" title="CITES">CITES</a> Appendix II<small> (<a href="/wiki/CITES" title="CITES">CITES</a>)<sup id="cite_ref-CITES_2-0" class="reference"><a href="#cite_note-CITES-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup></small></div> </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Chelus" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Chordate" title="Chordate">Chordata</a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Reptile" title="Reptile">Reptilia</a> </td></tr> <tr> <td>Order: </td> <td><a href="/wiki/Turtle" title="Turtle">Testudines</a> </td></tr> <tr> <td>Suborder: </td> <td><a href="/wiki/Pleurodira" title="Pleurodira">Pleurodira</a> </td></tr> <tr> <td>Family: </td> <td><a href="/wiki/Chelidae" title="Chelidae">Chelidae</a> </td></tr> <tr> <td>Genus: </td> <td><a href="/wiki/Chelus" title="Chelus"><i>Chelus</i></a> </td></tr> <tr> <td>Species: </td> <td><div style="display:inline" class="species"><i><b>C. fimbriata</b></i></div> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Binomial_nomenclature" title="Binomial nomenclature">Binomial name</a> </th></tr> <tr> <td colspan="2" style="text-align: center"><b><span class="binomial"><span style="font-weight:normal;"></span><i>Chelus fimbriata</i></span></b><br /><div style="font-size: 85%;">(<a href="/wiki/Johann_Gottlob_Schneider" class="mw-redirect" title="Johann Gottlob Schneider">Schneider</a>, 1783)<sup id="cite_ref-sch83_3-0" class="reference"><a href="#cite_note-sch83-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup></div> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Synonym_(taxonomy)" title="Synonym (taxonomy)">Synonyms</a><sup id="cite_ref-ttwg_1-1" class="reference"><a href="#cite_note-ttwg-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Fritz_2007_6-0" class="reference"><a href="#cite_note-Fritz_2007-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </th></tr> <tr> <td colspan="2" style="text-align: left"> <style data-mw-deduplicate="TemplateStyles:r1214851843">.mw-parser-output .hidden-begin{box-sizing:border-box;width:100%;padding:5px;border:none;font-size:95%}.mw-parser-output .hidden-title{font-weight:bold;line-height:1.6;text-align:left}.mw-parser-output .hidden-content{text-align:left}@media all and (max-width:500px){.mw-parser-output .hidden-begin{width:auto!important;clear:none!important;float:none!important}}</style><div class="hidden-begin mw-collapsible mw-collapsible-leftside-toggle mw-collapsed" style=""><div class="hidden-title skin-nightmode-reset-color" style=""><span style="font-size:85%;">Species synonymy</span></div><div class="hidden-content mw-collapsible-content" style=""> <ul><li><i>Testudo terrestris</i> <small>Fermin, 1765</small> <i>Nomen rejectum</i><sup id="cite_ref-iczn1963_4-0" class="reference"><a href="#cite_note-iczn1963-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup></li> <li><i>Testudo fimbriata</i> <small>Schneider, 1783</small><sup id="cite_ref-sch83_3-1" class="reference"><a href="#cite_note-sch83-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup></li> <li><i>Testudo fimbria</i> <small>Gmelin, 1789</small> <small><i>nomen novum</i></small></li> <li><i>Testudo matamata</i> <small>Bruguière, 1792</small> <i>nomen novum</i></li> <li><i>Testudo bispinosa</i> <small>Daudin, 1801</small> <i>nomen novum</i></li> <li><i>Emydes matamata</i> <small>Brongniart, 1805</small></li> <li><i>Chelus fimbriata</i> <small>(Schneider, 1783)</small> <small>recombination</small><sup id="cite_ref-dum1806_5-0" class="reference"><a href="#cite_note-dum1806-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup></li> <li><i>Testudo rapara</i> <small>Gray, 1831</small> <small><i>nomen novum</i></small></li> <li><i>Testudo raparara</i> <small>Gray, 1844</small> <small><i>nomen novum</i></small></li> <li><i>Testudo raxarara</i> <small>Gray, 1856</small> <small><i>nomen novum</i></small></li> <li><i>Chelys boulengerii</i> <small>Baur, 1890</small> <small><i>nomen novum</i></small></li></ul> </div></div> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p>The <b>mata mata</b>, <b>mata-mata</b>, or <b>matamata</b> (<i><b>Chelus fimbriata</b></i>)<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> is a South American <a href="/wiki/Species" title="Species">species</a> of freshwater <a href="/wiki/Turtle" title="Turtle">turtle</a> found in the <a href="/wiki/Amazon_basin" title="Amazon basin">Amazon basin</a> and river system of the eastern <a href="/wiki/Guianas" class="mw-redirect" title="Guianas">Guianas</a>. It was formerly believed to also occur in the <a href="/wiki/Orinoco" title="Orinoco">Orinoco</a> basin, western Guianas and upper <a href="/wiki/Rio_Negro_(Amazon)" title="Rio Negro (Amazon)">Rio Negro</a>–<a href="/wiki/Branco_River" title="Branco River">Branco</a> system, but in 2020 these populations were found to belong to a separate species, <i><a href="/wiki/Chelus_orinocensis" title="Chelus orinocensis">Chelus orinocensis</a></i> (Orinoco mata mata).<sup id="cite_ref-ttwg_1-2" class="reference"><a href="#cite_note-ttwg-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Vargas-Ramírez2020_8-0" class="reference"><a href="#cite_note-Vargas-Ramírez2020-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> Subsequently, some authorities have modified the common name of <i>Chelus fimbriata</i> to <b>Amazon mata mata</b>.<sup id="cite_ref-ttwg_1-3" class="reference"><a href="#cite_note-ttwg-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> These two are the only <a href="/wiki/Extant_taxon" class="mw-redirect" title="Extant taxon">extant</a> species in the <a href="/wiki/Genus" title="Genus">genus</a> <i><a href="/wiki/Chelus" title="Chelus">Chelus</a></i>.<sup id="cite_ref-ttwg_1-4" class="reference"><a href="#cite_note-ttwg-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Vargas-Ramírez2020_8-1" class="reference"><a href="#cite_note-Vargas-Ramírez2020-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Taxonomy">Taxonomy</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Mata_mata&action=edit&section=1" title="Edit section: Taxonomy"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The mata mata first became known to western scientists when it was described by French <a href="/wiki/Naturalist" class="mw-redirect" title="Naturalist">naturalist</a> <a href="/wiki/Pierre_Barr%C3%A8re" title="Pierre Barrère">Pierre Barrère</a> in 1741 as a "large land turtle with spiky and ridged scales" (translation).<sup id="cite_ref-Espenshade_9-0" class="reference"><a href="#cite_note-Espenshade-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> It was first scientifically described as <i>Testudo terrestris</i> in 1765 by Spanish zoologist <a href="/wiki/Ferm%C3%ADn_Zan%C3%B3n_Cervera" title="Fermín Zanón Cervera">Fermín Zanón Cervera</a>, but the same scientific name was used for describing the <a href="/wiki/Greek_tortoise" title="Greek tortoise">Greek tortoise</a> by <a href="/wiki/Peter_Forssk%C3%A5l" title="Peter Forsskål">Peter Forsskål</a> posthumously in 1775. In 1783, the mata mata was scientifically described again, this time as <i>Testudo fimbriata</i> by German naturalist <a href="/wiki/Johann_Gottlob_Schneider" class="mw-redirect" title="Johann Gottlob Schneider">Johann Gottlob Schneider</a>. In 1805, it was moved to its own genus as <i>Chelys fimbriata</i> by <a href="/wiki/Andr%C3%A9_Marie_Constant_Dum%C3%A9ril" title="André Marie Constant Duméril">André Marie Constant Duméril</a>.<sup id="cite_ref-ttwg_1-5" class="reference"><a href="#cite_note-ttwg-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-ReptileDatabase_10-0" class="reference"><a href="#cite_note-ReptileDatabase-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> From the mid 1700s to the late 1800s, altogether 15 different scientific names were used for the species.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> Since the late 1800s, most authorities have used the name <i>Chelys fimbriata</i>. In 1963, the <a href="/wiki/International_Commission_on_Zoological_Nomenclature" title="International Commission on Zoological Nomenclature">International Commission on Zoological Nomenclature</a> resolved the issue of <i>Testudo terrestris</i> (Fermín 1765) by declaring it a <a href="/wiki/Nomen_oblitum" title="Nomen oblitum">nomen oblitum</a> (meaning that only <i>Testudo terrestris</i> Forsskål 1775 is valid and a synonym for the Greek tortoise), while also declaring that <i>Chelys fimbriata</i> is a <a href="/wiki/Conserved_name" title="Conserved name">conserved name</a> for the mata mata.<sup id="cite_ref-ttwg_1-6" class="reference"><a href="#cite_note-ttwg-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-ReptileDatabase_10-1" class="reference"><a href="#cite_note-ReptileDatabase-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p><p>In 1995, distinct morphological differences were found between the populations of the Amazon and Orinoco basins.<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> In 2020, this was supported by a <a href="/wiki/Genetics" title="Genetics">genetic</a> analysis, which showed a deep split between two of its populations. The authors proposed that the mostly Orinoco population be assigned to a new species, <i><a href="/wiki/Chelus_orinocensis" title="Chelus orinocensis">Chelus orinocensis</a></i>, with the Amazon population retaining the <i>Chelus fimbriatus</i>. However, <i>Chelus fimbriatus</i> is absent from the upper Rio Negro–Branco system (the northernmost part of the Amazon basin), which instead is inhabited by <i>Chelus orinocensis</i>.<sup id="cite_ref-Vargas-Ramírez2020_8-2" class="reference"><a href="#cite_note-Vargas-Ramírez2020-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Anatomy_and_morphology">Anatomy and morphology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Mata_mata&action=edit&section=2" title="Edit section: Anatomy and morphology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Mata_Mata_1911.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/b/b5/Mata_Mata_1911.jpg/220px-Mata_Mata_1911.jpg" decoding="async" width="220" height="220" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/b/b5/Mata_Mata_1911.jpg/330px-Mata_Mata_1911.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/b/b5/Mata_Mata_1911.jpg/440px-Mata_Mata_1911.jpg 2x" data-file-width="489" data-file-height="490" /></a><figcaption>Top view of the mata mata turtle</figcaption></figure> <p>The mata mata is a large, sedentary turtle with a large, triangular, flattened head with many <a href="/wiki/Tubercle_(anatomy)" class="mw-redirect" title="Tubercle (anatomy)">tubercles</a> and flaps of skin, and a "spike" on its long and tubular snout.<sup id="cite_ref-Espenshade_9-1" class="reference"><a href="#cite_note-Espenshade-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> Three <a href="/wiki/Barbel_(anatomy)" class="mw-redirect" title="Barbel (anatomy)">barbels</a> occur on the chin and four additional filamentous barbels at the upper jaw, which is neither hooked nor notched.<sup id="cite_ref-Bartlett_13-0" class="reference"><a href="#cite_note-Bartlett-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p><p>The mata mata's brown or black, oblong <a href="/wiki/Carapace" title="Carapace">carapace</a> can measure up to 95 cm (37 in) at adult age.<sup id="cite_ref-description_14-0" class="reference"><a href="#cite_note-description-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> The full adult weight is 21 kg (46 lb).<sup id="cite_ref-description_14-1" class="reference"><a href="#cite_note-description-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> The mata mata's <a href="/wiki/Plastron" class="mw-redirect" title="Plastron">plastron</a> is reduced, narrowed, hingeless, shortened towards the front, and deeply notched at the rear with narrow bridges.<sup id="cite_ref-Bartlett_13-1" class="reference"><a href="#cite_note-Bartlett-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> These may be meant to allow the turtle to resemble a piece of bark, camouflaging it from possible predators.<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> The plastron and bridges are cream to yellow or brown.<sup id="cite_ref-Bartlett_13-2" class="reference"><a href="#cite_note-Bartlett-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> The head, neck, tail, and limbs are grayish brown on adults.<sup id="cite_ref-Bartlett_13-3" class="reference"><a href="#cite_note-Bartlett-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> The neck is longer than the vertebra under its carapace and is fringed with small skin flaps along both sides.<sup id="cite_ref-Bartlett_13-4" class="reference"><a href="#cite_note-Bartlett-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> Hatchlings show a pink to reddish tinge in the underside edge of their carapaces and plastrons that gradually disappear as they grow. </p><p>Each forefoot has five webbed claws. Males have concave plastrons and longer, thicker tails than females.<sup id="cite_ref-Bartlett_13-5" class="reference"><a href="#cite_note-Bartlett-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Habitat">Habitat</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Mata_mata&action=edit&section=3" title="Edit section: Habitat"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The mata mata inhabits slow moving streams, <a href="/wiki/Stagnant_(water)" class="mw-redirect" title="Stagnant (water)">stagnant</a> pools, marshes, and swamps ranging into northern <a href="/wiki/Bolivia" title="Bolivia">Bolivia</a>, eastern <a href="/wiki/Peru" title="Peru">Peru</a>, eastern <a href="/wiki/Ecuador" title="Ecuador">Ecuador</a>, southeastern <a href="/wiki/Colombia" title="Colombia">Colombia</a>, the eastern <a href="/wiki/Guianas" class="mw-redirect" title="Guianas">Guianas</a>, and northern and central <a href="/wiki/Brazil" title="Brazil">Brazil</a>.<sup id="cite_ref-ttwg_1-7" class="reference"><a href="#cite_note-ttwg-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> The mata mata is strictly an aquatic species but it prefers standing in shallow water where its snout can reach the surface to breathe.<sup id="cite_ref-Rosenfeld_16-0" class="reference"><a href="#cite_note-Rosenfeld-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Behavior">Behavior</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Mata_mata&action=edit&section=4" title="Edit section: Behavior"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Chelus_fimbriatus_close.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/6/6e/Chelus_fimbriatus_close.jpg/220px-Chelus_fimbriatus_close.jpg" decoding="async" width="220" height="259" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/6e/Chelus_fimbriatus_close.jpg/330px-Chelus_fimbriatus_close.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/6e/Chelus_fimbriatus_close.jpg/440px-Chelus_fimbriatus_close.jpg 2x" data-file-width="800" data-file-height="941" /></a><figcaption>Head, in profile</figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Chelus_fimbriatus.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/1/11/Chelus_fimbriatus.jpg/220px-Chelus_fimbriatus.jpg" decoding="async" width="220" height="127" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/11/Chelus_fimbriatus.jpg/330px-Chelus_fimbriatus.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/11/Chelus_fimbriatus.jpg/440px-Chelus_fimbriatus.jpg 2x" data-file-width="2000" data-file-height="1156" /></a><figcaption></figcaption></figure> <p>The appearance of the mata mata's shell resembles a piece of bark, and its head resembles fallen leaves.<sup id="cite_ref-Cogger_17-0" class="reference"><a href="#cite_note-Cogger-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> As it remains motionless in the water, its <a href="/wiki/Skin" title="Skin">skin</a> flaps enable it to blend into the surrounding vegetation until a fish comes close.<sup id="cite_ref-Cogger_17-1" class="reference"><a href="#cite_note-Cogger-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> The mata mata thrusts out its head and opens its large mouth as wide as possible, creating a low-pressure vacuum that sucks the prey into its mouth, known as <a href="/wiki/Suction_feeding" class="mw-redirect" title="Suction feeding">suction feeding</a>.<sup id="cite_ref-Cogger_17-2" class="reference"><a href="#cite_note-Cogger-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> The mata mata snaps its mouth shut, the water is slowly expelled, and the fish is swallowed whole; the mata mata cannot chew due to the way its mouth is constructed.<sup id="cite_ref-Cogger_17-3" class="reference"><a href="#cite_note-Cogger-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Reproduction">Reproduction</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Mata_mata&action=edit&section=5" title="Edit section: Reproduction"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Males display for females by extending their limbs, lunging their heads toward the females with mouths agape, and moving the lateral flaps on their heads. Nesting occurs from October through December in the <a href="/wiki/Upper_Amazon" class="mw-redirect" title="Upper Amazon">Upper Amazon</a>. The 12 to 28 brittle, spherical, 35 mm-diameter <a href="/wiki/Egg_(biology)" class="mw-redirect" title="Egg (biology)">eggs</a> are deposited in a clutch. </p> <div class="mw-heading mw-heading2"><h2 id="Diet">Diet</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Mata_mata&action=edit&section=6" title="Edit section: Diet"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1251242444">.mw-parser-output .ambox{border:1px solid #a2a9b1;border-left:10px solid #36c;background-color:#fbfbfb;box-sizing:border-box}.mw-parser-output .ambox+link+.ambox,.mw-parser-output .ambox+link+style+.ambox,.mw-parser-output .ambox+link+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+style+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+link+.ambox{margin-top:-1px}html body.mediawiki .mw-parser-output .ambox.mbox-small-left{margin:4px 1em 4px 0;overflow:hidden;width:238px;border-collapse:collapse;font-size:88%;line-height:1.25em}.mw-parser-output .ambox-speedy{border-left:10px solid #b32424;background-color:#fee7e6}.mw-parser-output .ambox-delete{border-left:10px solid #b32424}.mw-parser-output .ambox-content{border-left:10px solid #f28500}.mw-parser-output .ambox-style{border-left:10px solid #fc3}.mw-parser-output .ambox-move{border-left:10px solid #9932cc}.mw-parser-output .ambox-protection{border-left:10px solid #a2a9b1}.mw-parser-output .ambox .mbox-text{border:none;padding:0.25em 0.5em;width:100%}.mw-parser-output .ambox .mbox-image{border:none;padding:2px 0 2px 0.5em;text-align:center}.mw-parser-output .ambox .mbox-imageright{border:none;padding:2px 0.5em 2px 0;text-align:center}.mw-parser-output .ambox .mbox-empty-cell{border:none;padding:0;width:1px}.mw-parser-output .ambox .mbox-image-div{width:52px}@media(min-width:720px){.mw-parser-output .ambox{margin:0 10%}}@media print{body.ns-0 .mw-parser-output .ambox{display:none!important}}</style><table class="box-Expand_section plainlinks metadata ambox mbox-small-left ambox-content" role="presentation"><tbody><tr><td class="mbox-image"><span typeof="mw:File"><a href="/wiki/File:Wiki_letter_w_cropped.svg" class="mw-file-description"><img alt="[icon]" src="//upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Wiki_letter_w_cropped.svg/20px-Wiki_letter_w_cropped.svg.png" decoding="async" width="20" height="14" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Wiki_letter_w_cropped.svg/30px-Wiki_letter_w_cropped.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Wiki_letter_w_cropped.svg/40px-Wiki_letter_w_cropped.svg.png 2x" data-file-width="44" data-file-height="31" /></a></span></td><td class="mbox-text"><div class="mbox-text-span">This section <b>needs expansion</b>. You can help by <a class="external text" href="https://en.wikipedia.org/w/index.php?title=Mata_mata&action=edit&section=">adding to it</a>. <span class="date-container"><i>(<span class="date">March 2018</span>)</i></span></div></td></tr></tbody></table> <p>The mata mata is carnivorous, feeding almost exclusively upon aquatic <a href="/wiki/Invertebrates" class="mw-redirect" title="Invertebrates">invertebrates</a> (such as <a href="/wiki/Worm" title="Worm">worms</a>, <a href="/wiki/Mussel" title="Mussel">mussels</a>, <a href="/wiki/Crustacean" title="Crustacean">crustaceans</a> and <a href="/wiki/Insect" title="Insect">insects</a>)<sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-sta.uwi.edu_20-0" class="reference"><a href="#cite_note-sta.uwi.edu-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> and <a href="/wiki/Fish" title="Fish">fish</a>.<sup id="cite_ref-Espenshade_9-2" class="reference"><a href="#cite_note-Espenshade-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Cogger_17-4" class="reference"><a href="#cite_note-Cogger-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> On rare occasions, it may feed on small <a href="/wiki/Bird" title="Bird">birds</a>, <a href="/wiki/Amphibian" title="Amphibian">amphibians</a> or small <a href="/wiki/Mammal" title="Mammal">mammals</a> that have entered the water.<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-sta.uwi.edu_20-1" class="reference"><a href="#cite_note-sta.uwi.edu-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> When the stomach content of 20 wild mata mata turtles was examined it consisted exclusively of small fish. The turtles predominantly feed at night in muddy water with limited visibility. However the turtle is well adapted to hunting in these conditions. The mata mata has very fine eyesight with eyes that reflect light, similar to other nocturnal reptiles. In addition, the skin flaps on the neck are also extremely sensitive and help the mata mata detect nearby movement.<sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup> </p><p>Mata mata turtles use a specific method of seizing their prey. They will move the prey into shallower areas of water, surround the prey, and wave their front legs to prevent them from escaping. Once surrounded, the mata mata turtles will open their mouths and contract their pharynx, causing a rush of water that pushes the prey into their mouth.<sup id="cite_ref-23" class="reference"><a href="#cite_note-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="In_captivity">In captivity</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Mata_mata&action=edit&section=7" title="Edit section: In captivity"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Mata_mata_NC.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4d/Mata_mata_NC.jpg/220px-Mata_mata_NC.jpg" decoding="async" width="220" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4d/Mata_mata_NC.jpg/330px-Mata_mata_NC.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4d/Mata_mata_NC.jpg/440px-Mata_mata_NC.jpg 2x" data-file-width="4320" data-file-height="3240" /></a><figcaption>Mata mata turtles on exhibit at the <a href="/wiki/North_Carolina_Museum_of_Natural_Sciences" title="North Carolina Museum of Natural Sciences">North Carolina Museum of Natural Sciences</a>, the United States</figcaption></figure> <p>Mata mata turtles are readily available in the exotic pet trade and are quite expensive to obtain. Due to their unique appearance, they make interesting display animals. They also grow quite large. However, mata matas are not active hunters, so, like the <a href="/wiki/Alligator_snapping_turtle" title="Alligator snapping turtle">alligator snapping turtle</a>, they need less space than a large, active species. </p><p>As with all aquatic turtles, water quality is one of the keys to keeping this species successfully in captivity. Warm, acidic water is the best type used with a high <a href="/wiki/Tannin-stained_waters" class="mw-redirect" title="Tannin-stained waters">tannin content</a> that should be maintained all year round. Moderate to heavy filtration is recommended.<sup id="cite_ref-24" class="reference"><a href="#cite_note-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup> Author David Fogel considers his captive mata mata turtles to be quite intelligent. For example, he has observed one turtle positioning itself near the spray bar of the aquarium at feeding time so that floating food is pushed beneath the water's surface where the turtle can catch it more easily.<sup id="cite_ref-25" class="reference"><a href="#cite_note-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Mata_mata&action=edit&section=8" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-ttwg-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-ttwg_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-ttwg_1-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-ttwg_1-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-ttwg_1-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-ttwg_1-4"><sup><i><b>e</b></i></sup></a> <a href="#cite_ref-ttwg_1-5"><sup><i><b>f</b></i></sup></a> <a href="#cite_ref-ttwg_1-6"><sup><i><b>g</b></i></sup></a> <a href="#cite_ref-ttwg_1-7"><sup><i><b>h</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFRhodinInversonRogerFritz2021" class="citation journal cs1">Rhodin, Anders G.J.; Inverson, John B.; Roger, Bour; Fritz, Uwe; Georges, Arthur; Shaffer, H. 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(January 1995). <a rel="nofollow" class="external text" href="https://chelonian.org/wp-content/uploads/file/CCB%20Vol%201%20No%204%20(1995)/Sanchez-Villagra_etal_1995.pdf">"Geographic variation in the matamata turtle, <i>Chelus fimbriatus</i>, with observations on its shell morphology and morphometry"</a> <span class="cs1-format">(PDF)</span>. <i>Chelonian Conservation and Biology</i>. <b>1</b>: 292–300.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Chelonian+Conservation+and+Biology&rft.atitle=Geographic+variation+in+the+matamata+turtle%2C+Chelus+fimbriatus%2C+with+observations+on+its+shell+morphology+and+morphometry&rft.volume=1&rft.pages=292-300&rft.date=1995-01&rft.aulast=Sanchez-Vilaga&rft.aufirst=Marcelo+R.&rft.au=Prichard%2C+Peter+C.+H.&rft.au=Paolillo%2C+Alfredo&rft.au=Linares%2C+Omar+J.&rft_id=https%3A%2F%2Fchelonian.org%2Fwp-content%2Fuploads%2Ffile%2FCCB%2520Vol%25201%2520No%25204%2520%281995%29%2FSanchez-Villagra_etal_1995.pdf&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMata+mata" class="Z3988"></span></span> </li> <li id="cite_note-Bartlett-13"><span class="mw-cite-backlink">^ <a href="#cite_ref-Bartlett_13-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Bartlett_13-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Bartlett_13-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-Bartlett_13-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-Bartlett_13-4"><sup><i><b>e</b></i></sup></a> <a href="#cite_ref-Bartlett_13-5"><sup><i><b>f</b></i></sup></a></span> <span class="reference-text"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBartlett2007" class="citation cs2">Bartlett, Dick (2007), "The Matamata", <i>Reptiles Magazine</i>, <b>15</b> (12): 18–20</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Reptiles+Magazine&rft.atitle=The+Matamata&rft.volume=15&rft.issue=12&rft.pages=18-20&rft.date=2007&rft.aulast=Bartlett&rft.aufirst=Dick&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMata+mata" class="Z3988"></span></span> </li> <li id="cite_note-description-14"><span class="mw-cite-backlink">^ <a href="#cite_ref-description_14-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-description_14-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><span class="languageicon">(in French)</span> <i>Toutes les tortues du monde</i> by Franck Bonin, Bernard Devaux and Alain Dupré, second edition (1998), editions Delachaux and Niestlé/WWF.</span> </li> <li id="cite_note-15"><span class="mw-cite-backlink"><b><a href="#cite_ref-15">^</a></b></span> <span class="reference-text">Encyclopedia of Animals: Mammals, Birds, Reptiles and Amphibians, Harold G. 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Turtle and Tortoise Preservation group's Turtles of the world series.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Matamatas%3A+The+Natural+History%2C+Captive+Care+and+Breeding+of+Chelus+fimbriatus&rft.pub=Turtle+and+Tortoise+Preservation+group%27s+Turtles+of+the+world+series&rft.date=2011&rft.aulast=Fogel&rft.aufirst=David&rfr_id=info%3Asid%2Fen.wikipedia.org%3AMata+mata" class="Z3988"></span></span> </li> </ol></div></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 0;box-sizing:border-box;border:1px solid #aaa;font-size:88%;line-height:1.25em;background-color:var(--background-color-interactive-subtle,#f8f9fa);display:flow-root}.mw-parser-output .side-box-abovebelow,.mw-parser-output .side-box-text{padding:0.25em 0.9em}.mw-parser-output .side-box-image{padding:2px 0 2px 0.9em;text-align:center}.mw-parser-output .side-box-imageright{padding:2px 0.9em 2px 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abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Pleurodira" title="Template:Pleurodira"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Pleurodira" title="Template talk:Pleurodira"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Pleurodira" title="Special:EditPage/Template:Pleurodira"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Side-necked_turtle_species" style="font-size:114%;margin:0 4em">Side-necked <a href="/wiki/Turtle" title="Turtle">turtle</a> species</div></th></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li>Kingdom: <a href="/wiki/Animal" title="Animal">Animalia</a></li> <li>Phylum: <a href="/wiki/Chordate" title="Chordate">Chordata</a></li> <li>Class: <a href="/wiki/Reptile" title="Reptile">Reptilia</a></li> <li>Subclass: <a href="/wiki/Anapsid" title="Anapsid">Anapsida</a></li> <li>Order: <a href="/wiki/Turtle" title="Turtle">Testudines</a></li> <li>Suborder: <a href="/wiki/Pleurodira" title="Pleurodira">Pleurodira</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:5em"><a href="/wiki/Family_(biology)" title="Family (biology)">Family</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Subfamily" scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Subfamily" title="Subfamily">Subfamily</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Genus" scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Genus" title="Genus">Genus</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Subgenus" scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Subgenus" title="Subgenus">Subgenus</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td class="navbox-abovebelow" colspan="2" style="text-align:left;font-weight:bold;"><div id="Species"><a href="/wiki/Species" title="Species">Species</a></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:5em"><a href="/wiki/Chelidae" title="Chelidae">Chelidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Chelidae" title="Chelidae">Chelinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Acanthochelys" title="Acanthochelys">Acanthochelys</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Big-headed_pantanal_swamp_turtle" title="Big-headed pantanal swamp turtle">Acanthochelys macrocephala</a></i></li> <li><i><a href="/wiki/Chaco_side-necked_turtle" title="Chaco side-necked turtle">Acanthochelys pallidipectoris</a></i></li> <li><i><a href="/wiki/Brazilian_radiolated_swamp_turtle" title="Brazilian radiolated swamp turtle">Acanthochelys radiolata</a></i></li> <li><i><a href="/wiki/Black_spine-neck_swamp_turtle" title="Black spine-neck swamp turtle">Acanthochelys spixii</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Chelus" title="Chelus">Chelus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a class="mw-selflink selflink">Chelus fimbriata</a></i></li> <li><i><a href="/wiki/Chelus_orinocensis" title="Chelus orinocensis">Chelus orinocensis</a></i></li> <li>†<i><a href="/w/index.php?title=Chelus_colombiana&action=edit&redlink=1" class="new" title="Chelus colombiana (page does not exist)">Chelus colombiana</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Mesoclemmys" title="Mesoclemmys">Mesoclemmys</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Dahl%27s_toad-headed_turtle" title="Dahl's toad-headed turtle">Mesoclemmys dahli</a></i></li> <li><i><a href="/wiki/Mesoclemmys_gibba" title="Mesoclemmys gibba">Mesoclemmys gibba</a></i></li> <li><i><a href="/w/index.php?title=Mesoclemmys_jurutiensis&action=edit&redlink=1" class="new" title="Mesoclemmys jurutiensis (page does not exist)">Mesoclemmys jurutiensis</a></i></li> <li><i><a href="/wiki/Mesoclemmys_nasuta" title="Mesoclemmys nasuta">Mesoclemmys nasuta</a></i></li> <li><i><a href="/wiki/Mesoclemmys_perplexa" title="Mesoclemmys perplexa">Mesoclemmys perplexa</a></i></li> <li><i><a href="/wiki/Mesoclemmys_raniceps" title="Mesoclemmys raniceps">Mesoclemmys raniceps</a></i></li> <li><i><a href="/w/index.php?title=Mesoclemmys_sabiniparaensis&action=edit&redlink=1" class="new" title="Mesoclemmys sabiniparaensis (page does not exist)">Mesoclemmys sabiniparaensis</a></i></li> <li><i><a href="/wiki/Mesoclemmys_tuberculata" title="Mesoclemmys tuberculata">Mesoclemmys tuberculata</a></i></li> <li><i><a href="/wiki/Vanderhaege%27s_toad-headed_turtle" title="Vanderhaege's toad-headed turtle">Mesoclemmys vanderhaegei</a></i></li> <li>†<i><a href="/w/index.php?title=Mesoclemmys_vanegasorum&action=edit&redlink=1" class="new" title="Mesoclemmys vanegasorum (page does not exist)">Mesoclemmys vanegasorum</a></i></li> <li><i><a href="/wiki/Zulia_toad-headed_sideneck" title="Zulia toad-headed sideneck">Mesoclemmys zuliae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Phrynops" title="Phrynops">Phrynops</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Phrynops_geoffroanus" title="Phrynops geoffroanus">Phrynops geoffroanus</a></i></li> <li><i><a href="/wiki/Phrynops_hilarii" title="Phrynops hilarii">Phrynops hilarii</a></i></li> <li>†<i><a href="/wiki/Phrynops_paranaensis" title="Phrynops paranaensis">Phrynops paranaensis</a></i></li> <li><i><a href="/wiki/Phrynops_tuberosus" title="Phrynops tuberosus">Phrynops tuberosus</a></i></li> <li><i><a href="/wiki/Phrynops_williamsi" title="Phrynops williamsi">Phrynops williamsi</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Twist-necked_turtle" title="Twist-necked turtle">Platemys</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Twist-necked_turtle" title="Twist-necked turtle">Platemys platycephala</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Hoge%27s_side-necked_turtle" title="Hoge's side-necked turtle">Ranacephala</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Hoge%27s_side-necked_turtle" title="Hoge's side-necked turtle">Ranacephala hogei</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Red_side-necked_turtle" title="Red side-necked turtle">Rhinemys</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Red_side-necked_turtle" title="Red side-necked turtle">Rhinemys rufipes</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Chelidae#Classification" title="Chelidae">Chelodininae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Chelodina" title="Chelodina">Chelodina</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Chelodina" title="Chelodina">Chelodina</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Chelodina_canni" title="Chelodina canni">Chelodina canni</a></i></li> <li><i><a href="/wiki/Chelodina_gunaleni" title="Chelodina gunaleni">Chelodina gunaleni</a></i></li> <li><i><a href="/wiki/Eastern_long-necked_turtle" title="Eastern long-necked turtle">Chelodina longicollis</a></i></li> <li><i><a href="/wiki/Roti_Island_snake-necked_turtle" title="Roti Island snake-necked turtle">Chelodina mccordi</a></i></li> <li>†<i><a href="/wiki/Chelodina_murrayi" title="Chelodina murrayi">Chelodina murrayi</a></i></li> <li><i><a href="/wiki/New_Guinea_snake-necked_turtle" title="New Guinea snake-necked turtle">Chelodina novaeguineae</a></i></li> <li><i><a href="/wiki/Pritchard%27s_snake-necked_turtle" title="Pritchard's snake-necked turtle">Chelodina pritchardi</a></i></li> <li><i><a href="/wiki/Reimann%27s_snake-necked_turtle" title="Reimann's snake-necked turtle">Chelodina reimanni</a></i></li> <li><i><a href="/wiki/Chelodina_steindachneri" title="Chelodina steindachneri">Chelodina steindachneri</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Chelodina" title="Chelodina">Chelydera</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li>†<i><a href="/wiki/Chelodina_alanrixi" title="Chelodina alanrixi">Chelodina alanrixi</a></i></li> <li><i><a href="/wiki/Chelodina_burrungandjii" title="Chelodina burrungandjii">Chelodina burrungandjii</a></i></li> <li><i><a href="/wiki/Chelodina_expansa" title="Chelodina expansa">Chelodina expansa</a></i></li> <li><i><a href="/wiki/Chelodina_insculpta" title="Chelodina insculpta">Chelodina insculpta</a></i></li> <li><i><a href="/wiki/Chelodina_kuchlingi" title="Chelodina kuchlingi">Chelodina kuchlingi</a></i></li> <li><i><a href="/wiki/Chelodina_kurrichalpongo" title="Chelodina kurrichalpongo">Chelodina kurrichalpongo</a></i></li> <li><i><a href="/wiki/Parker%27s_snake-necked_turtle" title="Parker's snake-necked turtle">Chelodina parkeri</a></i></li> <li><i><a href="/wiki/Northern_snake-necked_turtle" title="Northern snake-necked turtle">Chelodina rugosa</a></i></li> <li><i><a href="/wiki/Chelodina_walloyarrina" title="Chelodina walloyarrina">Chelodina walloyarrina</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Chelodina" title="Chelodina">Macrochelodina</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Southwestern_snake-necked_turtle" class="mw-redirect" title="Southwestern snake-necked turtle">Chelodina oblonga</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Elseya" title="Elseya">Elseya</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Elseya" title="Elseya">Elseya</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Elseya_dentata" title="Elseya dentata">Elseya dentata</a></i></li> <li><i><a href="/wiki/Elseya_branderhorsti" title="Elseya branderhorsti">Elseya branderhorsti</a></i></li> <li><i><a href="/wiki/Elseya_flaviventralis" title="Elseya flaviventralis">Elseya flaviventralis</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Elseya" title="Elseya">Pelocomastes</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Elseya_albagula" title="Elseya albagula">Elseya albagula</a></i></li> <li><i><a href="/wiki/Irwin%27s_turtle" title="Irwin's turtle">Elseya irwini</a></i></li> <li><i><a href="/wiki/Gulf_snapping_turtle" title="Gulf snapping turtle">Elseya lavarackorum</a></i></li> <li>†<i><a href="/wiki/Elseya_nadibajagu" title="Elseya nadibajagu">Elseya nadibajagu</a></i></li> <li>†<i><a href="/wiki/Elseya_uberrima" title="Elseya uberrima">Elseya uberrima</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Elseya" title="Elseya">Hanwarachelys</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Elseya_caelatus" title="Elseya caelatus">Elseya caelatus</a></i></li> <li><i><a href="/wiki/New_Guinea_snapping_turtle" title="New Guinea snapping turtle">Elseya novaeguineae</a></i></li> <li><i><a href="/wiki/Elseya_orestiad" class="mw-redirect" title="Elseya orestiad">Elseya orestiad</a></i></li> <li><i><a href="/wiki/Elseya_rhodini" title="Elseya rhodini">Elseya rhodini</a></i></li> <li><i><a href="/wiki/Elseya_schultzei" title="Elseya schultzei">Elseya schultzei</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Mary_River_turtle" title="Mary River turtle">Elusor</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Mary_River_turtle" title="Mary River turtle">Elusor macrurus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Emydura" title="Emydura">Emydura</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Emydura_australis" title="Emydura australis">Emydura australis</a></i></li> <li><i><a href="/wiki/Emydura_gunaleni" title="Emydura gunaleni">Emydura gunaleni</a></i></li> <li><i><a href="/wiki/Emydura_macquarii" title="Emydura macquarii">Emydura macquarii</a></i></li> <li><i><a href="/wiki/Red-bellied_short-necked_turtle" title="Red-bellied short-necked turtle">Emydura subglobosa</a></i></li> <li><i><a href="/wiki/Emydura_tanybaraga" class="mw-redirect" title="Emydura tanybaraga">Emydura tanybaraga</a></i></li> <li><i><a href="/wiki/Emydura_victoriae" title="Emydura victoriae">Emydura victoriae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Myuchelys" title="Myuchelys">Myuchelys</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Namoi_River_snapping_turtle" title="Namoi River snapping turtle">Myuchelys belli</a></i></li> <li><i><a href="/wiki/Bellinger_River_snapping_turtle" title="Bellinger River snapping turtle">Myuchelys georgesi</a></i></li> <li><i><a href="/wiki/Saw-shelled_turtle" title="Saw-shelled turtle">Myuchelys latisternum</a></i></li> <li><i><a href="/wiki/Manning_River_snapping_turtle" title="Manning River snapping turtle">Myuchelys purvisi</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Rheodytes" title="Rheodytes">Rheodytes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Rheodytes" title="Rheodytes">Rheodytes leukops</a></i></li> <li>†<i><a href="/wiki/Rheodytes_devisi" title="Rheodytes devisi">Rheodytes devisi</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Chelidae" title="Chelidae">Hydromedusinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Hydromedusa" scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Hydromedusa" title="Hydromedusa">Hydromedusa</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Brazilian_snake-necked_turtle" title="Brazilian snake-necked turtle">Hydromedusa maximiliani</a></i></li> <li><i><a href="/wiki/Argentine_snake-necked_turtle" title="Argentine snake-necked turtle">Hydromedusa tectifera</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Western_swamp_turtle" title="Western swamp turtle">Pseudemydurinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Pseudemydura" scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Western_swamp_turtle" title="Western swamp turtle">Pseudemydura</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Western_swamp_turtle" title="Western swamp turtle">Pseudemydura umbrina</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:5em"><a href="/wiki/Pelomedusidae" title="Pelomedusidae">Pelomedusidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/African_helmeted_turtle" title="African helmeted turtle">Pelomedusa</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/African_helmeted_turtle" title="African helmeted turtle">Pelomedusa subrufa</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Pelusios" title="Pelusios">Pelusios</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Adanson%27s_mud_turtle" title="Adanson's mud turtle">Pelusios adansonii</a></i></li> <li><i><a href="/wiki/Okavango_mud_turtle" title="Okavango mud turtle">Pelusios bechuanicus</a></i></li> <li><i><a href="/wiki/Turkana_mud_turtle" title="Turkana mud turtle">Pelusios broadleyi</a></i></li> <li><i><a href="/wiki/African_keeled_mud_turtle" title="African keeled mud turtle">Pelusios carinatus</a></i></li> <li><i><a href="/wiki/West_African_mud_turtle" title="West African mud turtle">Pelusios castaneus</a></i></li> <li><i><a href="/wiki/Yellow-bellied_mud_turtle" title="Yellow-bellied mud turtle">Pelusios castanoides</a></i></li> <li><i><a href="/wiki/Central_African_mud_turtle" title="Central African mud turtle">Pelusios chapini</a></i></li> <li><i><a href="/wiki/Ivory_Coast_mud_turtle" title="Ivory Coast mud turtle">Pelusios cupulatta</a></i></li> <li><i><a href="/wiki/African_forest_turtle" title="African forest turtle">Pelusios gabonensis</a></i></li> <li><i><a href="/wiki/Pelusios_marani" title="Pelusios marani">Pelusios marani</a></i></li> <li><i><a href="/wiki/African_dwarf_mud_turtle" title="African dwarf mud turtle">Pelusios nanus</a></i></li> <li><i><a href="/wiki/West_African_black_turtle" title="West African black turtle">Pelusios niger</a></i></li> <li><i><a href="/wiki/Variable_mud_turtle" title="Variable mud turtle">Pelusios rhodesianus</a></i></li> <li><i><a href="/wiki/Serrated_hinged_terrapin" title="Serrated hinged terrapin">Pelusios sinuatus</a></i></li> <li><i><a href="/wiki/East_African_black_mud_turtle" title="East African black mud turtle">Pelusios subniger</a></i></li> <li><i><a href="/wiki/Upemba_mud_turtle" title="Upemba mud turtle">Pelusios upembae</a></i></li> <li><i><a href="/wiki/Williams%27_mud_turtle" title="Williams' mud turtle">Pelusios williamsi</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:5em"><a href="/wiki/Podocnemididae" title="Podocnemididae">Podocnemididae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Madagascan_big-headed_turtle" title="Madagascan big-headed turtle">Erymnochelys</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Madagascan_big-headed_turtle" title="Madagascan big-headed turtle">Erymnochelys madagascariensis</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Big-headed_Amazon_River_turtle" title="Big-headed Amazon River turtle">Peltocephalus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Big-headed_Amazon_River_turtle" title="Big-headed Amazon River turtle">Peltocephalus dumerilianus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><i><a href="/wiki/Podocnemis" title="Podocnemis">Podocnemis</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Red-headed_Amazon_River_turtle" title="Red-headed Amazon River turtle">Podocnemis erythrocephala</a></i></li> <li><i><a href="/wiki/Arrau_turtle" title="Arrau turtle">Podocnemis expansa</a></i></li> <li><i><a href="/wiki/Magdalena_River_turtle" title="Magdalena River turtle">Podocnemis lewyana</a></i></li> <li><i><a href="/wiki/Six-tubercled_Amazon_River_turtle" title="Six-tubercled Amazon River turtle">Podocnemis sextuberculata</a></i></li> <li><i><a href="/wiki/Yellow-spotted_river_turtle" title="Yellow-spotted river turtle">Podocnemis unifilis</a></i></li> <li><i><a href="/wiki/Savanna_side-necked_turtle" title="Savanna side-necked turtle">Podocnemis vogli</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li>Phylogenetic arrangement of turtles based on <a rel="nofollow" class="external text" href="https://iucn-tftsg.org/wp-content/uploads/crm.8.checklist.atlas_.v9.2021.e3.pdf"><i>Turtles of the World 2021 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status</i></a>. Key: †=extinct.</li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Testudines" style="padding:3px"><table class="nowraplinks hlist mw-collapsible mw-collapsed navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Testudines" title="Template:Testudines"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Testudines" title="Template talk:Testudines"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Testudines" title="Special:EditPage/Template:Testudines"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Testudines" style="font-size:114%;margin:0 4em"><a href="/wiki/Turtle" title="Turtle">Testudines</a></div></th></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li>Kingdom: <a href="/wiki/Animal" title="Animal">Animalia</a></li> <li>Phylum: <a href="/wiki/Chordate" title="Chordate">Chordata</a></li> <li>Class: <a href="/wiki/Reptile" title="Reptile">Reptilia</a></li> <li><i>Clade</i>: <a href="/wiki/Diapsid" title="Diapsid">Diapsida</a></li> <li>Order: <a href="/wiki/Turtle" title="Turtle">Testudines</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:5em"><a href="/wiki/Order_(biology)#Hierarchy_of_ranks" title="Order (biology)">Suborder</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Superfamily" scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Taxonomic_rank" title="Taxonomic rank">Superfamily</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Family" scope="row" class="navbox-group" style="width:11em"><a href="/wiki/List_of_Testudines_families" title="List of Testudines families">Family</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td class="navbox-abovebelow" colspan="2" style="text-align:left;"><div id="Genus"><b><a href="/wiki/Genus" title="Genus">Genus</a></b></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:5em"><a href="/wiki/Cryptodira" title="Cryptodira">Cryptodira</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Sea_turtle" title="Sea turtle">Chelonioidea</a><br />(Sea turtles)</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Cheloniidae" scope="row" class="navbox-group" style="width:11em"><a href="/wiki/Cheloniidae" title="Cheloniidae">Cheloniidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li>†<i><a href="/wiki/Allopleuron" title="Allopleuron">Allopleuron</a></i></li> <li><i><a href="/wiki/Loggerhead_sea_turtle" title="Loggerhead sea turtle">Caretta</a></i></li> <li>†<i><a href="/wiki/Carolinochelys" title="Carolinochelys">Carolinochelys</a></i></li> <li><i><a href="/wiki/Green_sea_turtle" title="Green sea turtle">Chelonia</a></i></li> <li>†<i><a href="/wiki/Eochelone" title="Eochelone">Eochelone</a></i></li> <li><i><a href="/wiki/Hawksbill_sea_turtle" title="Hawksbill sea turtle">Eretmochelys</a></i></li> <li>†<i><a href="/wiki/Gigantatypus" title="Gigantatypus">Gigantatypus</a></i></li> <li>†<i><a href="/wiki/Glarichelys" title="Glarichelys">Glarichelys</a></i></li> <li>†<i><a href="/wiki/Itilochelys" title="Itilochelys">Itilochelys</a></i></li> <li><i><a href="/wiki/Ridley_sea_turtle" title="Ridley sea turtle">Lepidochelys</a></i></li> <li>†<i><a href="/wiki/Mexichelys" title="Mexichelys">Mexichelys</a></i></li> <li>†<i><a href="/wiki/Miocaretta" title="Miocaretta">Miocaretta</a></i></li> <li><i><a href="/wiki/Flatback_sea_turtle" title="Flatback sea turtle">Natator</a></i></li> <li>†<i><a href="/wiki/Pacifichelys" title="Pacifichelys">Pacifichelys</a></i></li> <li>†<i><a href="/wiki/Syllomus" title="Syllomus">Syllomus</a></i></li> <li>†<i><a href="/wiki/Tasbacka" title="Tasbacka">Tasbacka</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:11em"><a href="/wiki/Dermochelyidae" title="Dermochelyidae">Dermochelyidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li>†<i><a href="/wiki/Arabemys" title="Arabemys">Arabemys</a></i></li> <li>†<i><a href="/wiki/Corsochelys" title="Corsochelys">Corsochelys</a></i></li> <li>†<i><a href="/wiki/Cosmochelys" title="Cosmochelys">Cosmochelys</a></i></li> <li><i><a href="/wiki/Leatherback_sea_turtle" title="Leatherback sea turtle">Dermochelys</a></i></li> <li>†<i><a href="/wiki/Eosphargis" title="Eosphargis">Eosphargis</a></i></li> <li>†<i><a href="/wiki/Mesodermochelys" title="Mesodermochelys">Mesodermochelys</a></i></li> <li>†<i><a href="/wiki/Psephophorus" title="Psephophorus">Psephophorus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:11em"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li>†<i><a href="/wiki/Euclastes" title="Euclastes">Euclastes</a></i></li> <li>†<i><a href="/wiki/Peritresius" title="Peritresius">Peritresius</a></i></li> <li>†<i><a href="/wiki/Procolpochelys" title="Procolpochelys">Procolpochelys</a></i></li> <li>†<i><a href="/wiki/Protosphargis" title="Protosphargis">Protosphargis</a></i></li> <li>†<i><a href="/wiki/Puppigerus" title="Puppigerus">Puppigerus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Kinosternoidea" title="Kinosternoidea">Kinosternoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Dermatemydidae" scope="row" class="navbox-group" style="width:11em"><a href="/wiki/Dermatemydidae" title="Dermatemydidae">Dermatemydidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Hickatee" title="Hickatee">Dermatemys</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:11em"><a href="/wiki/Kinosternidae" title="Kinosternidae">Kinosternidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Narrow-bridged_musk_turtle" title="Narrow-bridged musk turtle">Claudius</a></i></li> <li>†<i><a href="/wiki/Hoplochelys" title="Hoplochelys">Hoplochelys</a></i></li> <li><i><a href="/wiki/Kinosternon" title="Kinosternon">Kinosternon</a></i></li> <li><i><a href="/wiki/Staurotypus" title="Staurotypus">Staurotypus</a></i></li> <li><i><a href="/wiki/Sternotherus" title="Sternotherus">Sternotherus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Testudinoidea" title="Testudinoidea">Testudinoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:11em"><a href="/wiki/Emydidae" title="Emydidae">Emydidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li>†<i><a href="/wiki/Acherontemys" title="Acherontemys">Acherontemys</a></i></li> <li><i><a href="/wiki/Chrysemys" title="Chrysemys">Chrysemys</a></i></li> <li><i><a href="/wiki/Clemmys" title="Clemmys">Clemmys</a></i></li> <li><i><a href="/wiki/Deirochelys" title="Deirochelys">Deirochelys</a></i></li> <li><i><a href="/wiki/Emys" title="Emys">Emys</a></i></li> <li><i><a href="/wiki/Actinemys" title="Actinemys">Actinemys</a></i></li> <li><i><a href="/wiki/Emydoidea" class="mw-redirect" title="Emydoidea">Emydoidea</a></i></li> <li><i><a href="/wiki/Glyptemys" title="Glyptemys">Glyptemys</a></i></li> <li><i><a href="/wiki/Graptemys" title="Graptemys">Graptemys</a></i></li> <li><i><a href="/wiki/Diamondback_terrapin" title="Diamondback terrapin">Malaclemys</a></i></li> <li><i><a href="/wiki/Pseudemys" title="Pseudemys">Pseudemys</a></i></li> <li><i><a href="/wiki/Box_turtle" title="Box turtle">Terrapene</a></i></li> <li><i><a href="/wiki/Trachemys" title="Trachemys">Trachemys</a></i></li> <li>†<i><a href="/w/index.php?title=Wilburemys&action=edit&redlink=1" class="new" title="Wilburemys (page does not exist)">Wilburemys</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:11em"><a href="/wiki/Geoemydidae" title="Geoemydidae">Geoemydidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Batagur" title="Batagur">Batagur</a></i></li> <li>†<i><a href="/wiki/Banhxeochelys" title="Banhxeochelys">Banhxeochelys</a></i></li> <li><i><a href="/wiki/Asian_box_turtle" title="Asian box turtle">Cuora</a></i></li> <li><i><a href="/wiki/Cyclemys" title="Cyclemys">Cyclemys</a></i></li> <li><i><a href="/wiki/Black_pond_turtle" title="Black pond turtle">Geoclemys</a></i></li> <li><i><a href="/wiki/Geoemyda" title="Geoemyda">Geoemyda</a></i></li> <li><i><a href="/wiki/Brahminy_river_turtle" title="Brahminy river turtle">Hardella</a></i></li> <li><i><a href="/wiki/Heosemys" title="Heosemys">Heosemys</a></i></li> <li><i><a href="/wiki/Sulawesi_forest_turtle" title="Sulawesi forest turtle">Leucocephalon</a></i></li> <li><i><a href="/wiki/Malayemys" title="Malayemys">Malayemys</a></i></li> <li><i><a href="/wiki/Mauremys" title="Mauremys">Mauremys</a></i></li> <li><i><a href="/wiki/Melanochelys" title="Melanochelys">Melanochelys</a></i></li> <li><i><a href="/wiki/Morenia" title="Morenia">Morenia</a></i></li> <li><i><a href="/wiki/Malayan_flat-shelled_turtle" title="Malayan flat-shelled turtle">Notochelys</a></i></li> <li><i><a href="/wiki/Malaysian_giant_turtle" title="Malaysian giant turtle">Orlitia</a></i></li> <li><i><a href="/wiki/Pangshura" title="Pangshura">Pangshura</a></i></li> <li><i><a href="/wiki/Rhinoclemmys" title="Rhinoclemmys">Rhinoclemmys</a></i></li> <li><i><a href="/wiki/Sacalia" title="Sacalia">Sacalia</a></i></li> <li><i><a href="/wiki/Siebenrockiella" title="Siebenrockiella">Siebenrockiella</a></i></li> <li><i><a href="/wiki/Cane_turtle" title="Cane turtle">Vijayachelys</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:11em"> <a href="/wiki/Big-headed_turtle" title="Big-headed turtle">Platysternidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Big-headed_turtle" title="Big-headed turtle">Platysternon</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:11em"><a href="/wiki/Tortoise" title="Tortoise">Testudinidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Aldabrachelys" title="Aldabrachelys">Aldabrachelys</a></i></li> <li><i><a href="/wiki/Astrochelys" title="Astrochelys">Astrochelys</a></i></li> <li><i><a href="/wiki/Centrochelys" title="Centrochelys">Centrochelys</a></i></li> <li><i><a href="/wiki/Chelonoidis" title="Chelonoidis">Chelonoidis</a></i></li> <li><i><a href="/wiki/Angulate_tortoise" title="Angulate tortoise">Chersina</a></i></li> <li><i><a href="/wiki/Cylindraspis" title="Cylindraspis">Cylindraspis</a></i></li> <li>†<i><a href="/w/index.php?title=Cymatholcus&action=edit&redlink=1" class="new" title="Cymatholcus (page does not exist)">Cymatholcus</a></i></li> <li>†<i><a href="/w/index.php?title=Floridemys&action=edit&redlink=1" class="new" title="Floridemys (page does not exist)">Floridemys</a></i></li> <li><i><a href="/wiki/Geochelone" title="Geochelone">Geochelone</a></i></li> <li><i><a href="/wiki/Gopherus" title="Gopherus">Gopherus</a></i></li> <li>†<i><a href="/wiki/Hadrianus_(turtle)" title="Hadrianus (turtle)">Hadrianus</a></i></li> <li>†<i><a href="/wiki/Hesperotestudo" title="Hesperotestudo">Hesperotestudo</a></i></li> <li><i><a href="/wiki/Homopus" title="Homopus">Homopus</a></i></li> <li><i><a href="/wiki/Indotestudo" title="Indotestudo">Indotestudo</a></i></li> <li><i><a href="/wiki/Kinixys" title="Kinixys">Kinixys</a></i></li> <li><i><a href="/wiki/Pancake_tortoise" title="Pancake tortoise">Malacochersus</a></i></li> <li><i><a href="/wiki/Manouria" title="Manouria">Manouria</a></i></li> <li>†<i><a href="/wiki/Megalochelys" title="Megalochelys">Megalochelys</a></i></li> <li>†<i><a href="/w/index.php?title=Oligopherus&action=edit&redlink=1" class="new" title="Oligopherus (page does not exist)">Oligopherus</a></i></li> <li><i><a href="/wiki/Psammobates" title="Psammobates">Psammobates</a></i></li> <li><i><a href="/wiki/Pyxis_(turtle)" title="Pyxis (turtle)">Pyxis</a></i></li> <li>†<i><a href="/wiki/Solitudo" title="Solitudo">Solitudo</a></i></li> <li><i><a href="/wiki/Leopard_tortoise" title="Leopard tortoise">Stigmochelys</a></i></li> <li>†<i><a href="/wiki/Stylemys" title="Stylemys">Stylemys</a></i></li> <li><i><a href="/wiki/Testudo_(genus)" title="Testudo (genus)">Testudo</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Trionychia" title="Trionychia">Trionychia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:11em"><a href="/wiki/Pig-nosed_turtle" title="Pig-nosed turtle">Carettochelyidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li>†<i><a href="/wiki/Allaeochelys" title="Allaeochelys">Allaeochelys</a></i></li> <li>†<i><a href="/wiki/Anosteira" title="Anosteira">Anosteira</a></i></li> <li><i><a href="/wiki/Pig-nosed_turtle" title="Pig-nosed turtle">Carettochelys</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:11em"><a href="/wiki/Trionychidae" title="Trionychidae">Trionychidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Amyda" title="Amyda">Amyda</a></i></li> <li><i><a href="/wiki/Apalone" title="Apalone">Apalone</a></i></li> <li>†<i><a href="/wiki/Axestemys" title="Axestemys">Axestemys</a></i></li> <li><i><a href="/wiki/Chitra_(turtle)" title="Chitra (turtle)">Chitra</a></i></li> <li><i><a href="/wiki/Cyclanorbis" title="Cyclanorbis">Cyclanorbis</a></i></li> <li><i><a href="/wiki/Cycloderma" title="Cycloderma">Cycloderma</a></i></li> <li>†<i><a href="/wiki/Drazinderetes" title="Drazinderetes">Drazinderetes</a></i></li> <li><i><a href="/wiki/Malayan_softshell_turtle" title="Malayan softshell turtle">Dogania</a></i></li> <li>†<i><a href="/wiki/Gilmoremys" title="Gilmoremys">Gilmoremys</a></i></li> <li>†<i><a href="/wiki/Hutchemys" title="Hutchemys">Hutchemys</a></i></li> <li>†<i><a href="/wiki/Khunnuchelys" title="Khunnuchelys">Khunnuchelys</a></i></li> <li><i><a href="/wiki/Lissemys" title="Lissemys">Lissemys</a></i></li> <li><i><a href="/wiki/Nilssonia_(turtle)" title="Nilssonia (turtle)">Nilssonia</a></i></li> <li><i><a href="/wiki/Wattle-necked_softshell_turtle" title="Wattle-necked softshell turtle">Palea</a></i></li> <li>†<i><a href="/wiki/Palaeoamyda" title="Palaeoamyda">Palaeoamyda</a></i></li> <li><i><a href="/wiki/Pelochelys" title="Pelochelys">Pelochelys</a></i></li> <li><i><a href="/wiki/Pelodiscus" title="Pelodiscus">Pelodiscus</a></i></li> <li><i><a href="/wiki/Rafetus" title="Rafetus">Rafetus</a></i></li> <li><i><a href="/wiki/Trionyx" title="Trionyx">Trionyx</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:11em"> </th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li>†<i><a href="/wiki/Basilochelys" title="Basilochelys">Basilochelys</a></i></li> <li>†<i><a href="/wiki/Sinaspideretes" title="Sinaspideretes">Sinaspideretes</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Chelydridae" scope="row" class="navbox-group" style="width:11em"><a href="/wiki/Chelydridae" title="Chelydridae">Chelydridae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Chelydra" title="Chelydra">Chelydra</a></i></li> <li>†<i><a href="/wiki/Chelydrops" title="Chelydrops">Chelydrops</a></i></li> <li>†<i><a href="/wiki/Chelydropsis" title="Chelydropsis">Chelydropsis</a></i></li> <li>†<i><a href="/wiki/Emarginachelys" title="Emarginachelys">Emarginachelys</a></i></li> <li>†<i><a href="/wiki/Macrocephalochelys" title="Macrocephalochelys">Macrocephalochelys</a></i></li> <li><i><a href="/wiki/Macrochelys" title="Macrochelys">Macrochelys</a></i></li> <li>†<i><a href="/wiki/Planiplastron" title="Planiplastron">Planiplastron</a></i></li> <li>†<i><a href="/wiki/Protochelydra" title="Protochelydra">Protochelydra</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:11em">†<a href="/wiki/Nanhsiungchelyidae" title="Nanhsiungchelyidae">Nanhsiungchelyidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li>†<i><a href="/wiki/Anomalochelys" title="Anomalochelys">Anomalochelys</a></i></li> <li>†<i><a href="/wiki/Basilemys" title="Basilemys">Basilemys</a></i></li> <li>†<i><a href="/wiki/Jiangxichelys" title="Jiangxichelys">Jiangxichelys</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:11em">†<a href="/wiki/Protostegidae" title="Protostegidae">Protostegidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li>†<i><a href="/wiki/Alienochelys" title="Alienochelys">Alienochelys</a></i></li> <li>†<i><a href="/wiki/Archelon" title="Archelon">Archelon</a></i></li> <li>†<i><a href="/wiki/Atlantochelys" title="Atlantochelys">Atlantochelys</a></i></li> <li>†<i><a href="/wiki/Bouliachelys" title="Bouliachelys">Bouliachelys</a></i></li> <li>†<i><a href="/wiki/Calcarichelys" title="Calcarichelys">Calcarichelys</a></i></li> <li>†<i><a href="/wiki/Cratochelone" title="Cratochelone">Cratochelone</a></i></li> <li>†<i><a href="/wiki/Desmatochelys" title="Desmatochelys">Desmatochelys</a></i></li> <li>†<i><a href="/wiki/Iserosaurus" title="Iserosaurus">Iserosaurus</a></i></li> <li>†<i><a href="/wiki/Notochelone" title="Notochelone">Notochelone</a></i></li> <li>†<i><a href="/wiki/Ocepechelon" title="Ocepechelon">Ocepechelon</a></i></li> <li>†<i><a href="/wiki/Pneumatoarthrus" title="Pneumatoarthrus">Pneumatoarthrus</a></i></li> <li>†<i><a href="/wiki/Protostega" title="Protostega">Protostega</a></i></li> <li>†<i><a href="/wiki/Rhinochelys" title="Rhinochelys">Rhinochelys</a></i></li> <li>†<i><a href="/wiki/Santanachelys" title="Santanachelys">Santanachelys</a></i></li> <li>†<i><a href="/wiki/Terlinguachelys" title="Terlinguachelys">Terlinguachelys</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:11em"> </th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li>†<i><a href="/wiki/Adocus" title="Adocus">Adocus</a></i></li> <li>†<i><a href="/wiki/Argillochelys" title="Argillochelys">Argillochelys</a></i></li> <li>†<i><a href="/wiki/Bashuchelys" title="Bashuchelys">Bashuchelys</a></i></li> <li>†<i><a href="/wiki/Ctenochelys" title="Ctenochelys">Ctenochelys</a></i></li> <li>†<i><a href="/wiki/Prionochelys" title="Prionochelys">Prionochelys</a></i></li> <li>†<i><a href="/wiki/Toxochelys" title="Toxochelys">Toxochelys</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:5em"><a href="/wiki/Pleurodira" title="Pleurodira">Pleurodira</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:11em">†<a href="/wiki/Araripemydidae" title="Araripemydidae">Araripemydidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li>†<i><a href="/wiki/Araripemys" title="Araripemys">Araripemys</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:11em">†<a href="/wiki/Bothremydidae" title="Bothremydidae">Bothremydidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li>†<i><a href="/wiki/Araiochelys" title="Araiochelys">Araiochelys</a></i></li> <li>†<i><a href="/wiki/Arenila" title="Arenila">Arenila</a></i></li> <li>†<i><a href="/wiki/Azabbaremys" title="Azabbaremys">Azabbaremys</a></i></li> <li>†<i><a href="/wiki/Bothremys" title="Bothremys">Bothremys</a></i></li> <li>†<i><a href="/wiki/Cearachelys" title="Cearachelys">Cearachelys</a></i></li> <li>†<i><a href="/wiki/Chedighaii" title="Chedighaii">Chedighaii</a></i></li> <li>†<i><a href="/wiki/Chupacabrachelys" title="Chupacabrachelys">Chupacabrachelys</a></i></li> <li>†<i><a href="/wiki/Eotaphrosphys" title="Eotaphrosphys">Eotaphrosphys</a></i></li> <li>†<i><a href="/wiki/Foxemys" title="Foxemys">Foxemys</a></i></li> <li>†<i><a href="/wiki/Galianemys" title="Galianemys">Galianemys</a></i></li> <li>†<i><a href="/wiki/Ilatardia" title="Ilatardia">Ilatardia</a></i></li> <li>†<i><a href="/wiki/Inaechelys" title="Inaechelys">Inaechelys</a></i></li> <li>†<i><a href="/wiki/Itapecuruemys" title="Itapecuruemys">Itapecuruemys</a></i></li> <li>†<i><a href="/wiki/Jainemys" title="Jainemys">Jainemys</a></i></li> <li>†<i><a href="/wiki/Kinkonychelys" title="Kinkonychelys">Kinkonychelys</a></i></li> <li>†<i><a href="/wiki/Kurmademys" title="Kurmademys">Kurmademys</a></i></li> <li>†<i><a href="/wiki/Labrostochelys" title="Labrostochelys">Labrostochelys</a></i></li> <li>†<i><a href="/wiki/Nigeremys" title="Nigeremys">Nigeremys</a></i></li> <li>†<i><a href="/wiki/Phosphatochelys" title="Phosphatochelys">Phosphatochelys</a></i></li> <li>†<i><a href="/wiki/Polysternon" title="Polysternon">Polysternon</a></i></li> <li>†<i><a href="/wiki/Puentemys" title="Puentemys">Puentemys</a></i></li> <li>†<i><a href="/wiki/Rosasia" title="Rosasia">Rosasia</a></i></li> <li>†<i><a href="/wiki/Rhothonemys" title="Rhothonemys">Rhothonemys</a></i></li> <li>†<i><a href="/wiki/Sankuchemys" title="Sankuchemys">Sankuchemys</a></i></li> <li>†<i><a href="/wiki/Taphrosphys" title="Taphrosphys">Taphrosphys</a></i></li> <li>†<i><a href="/wiki/Ummulisani" title="Ummulisani">Ummulisani</a></i></li> <li>†<i><a href="/wiki/Zolhafah" title="Zolhafah">Zolhafah</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:11em"><a href="/wiki/Chelidae" title="Chelidae">Chelidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Acanthochelys" title="Acanthochelys">Acanthochelys</a></i></li> <li><i><a href="/wiki/Chelodina" title="Chelodina">Chelodina</a></i></li> <li><i><a class="mw-selflink selflink">Chelus</a></i></li> <li><i><a href="/wiki/Elseya" title="Elseya">Elseya</a></i></li> <li><i><a href="/wiki/Mary_River_turtle" title="Mary River turtle">Elusor</a></i></li> <li><i><a href="/wiki/Emydura" title="Emydura">Emydura</a></i></li> <li><i><a href="/wiki/Hydromedusa" title="Hydromedusa">Hydromedusa</a></i></li> <li>†<i><a href="/wiki/Lomalatachelys" title="Lomalatachelys">Lomalatachelys</a></i></li> <li><i><a href="/wiki/Mesoclemmys" title="Mesoclemmys">Mesoclemmys</a></i></li> <li><i><a href="/wiki/Myuchelys" title="Myuchelys">Myuchelys</a></i></li> <li><i><a href="/wiki/Phrynops" title="Phrynops">Phrynops</a></i></li> <li><i><a href="/wiki/Twist-necked_turtle" title="Twist-necked turtle">Platemys</a></i></li> <li>†<i><a href="/wiki/Prochelidella" title="Prochelidella">Prochelidella</a></i></li> <li><i><a href="/wiki/Western_swamp_turtle" title="Western swamp turtle">Pseudemydura</a></i></li> <li><i><a href="/wiki/Rheodytes" title="Rheodytes">Rheodytes</a></i></li> <li><i><a href="/wiki/Red_side-necked_turtle" title="Red side-necked turtle">Rhinemys</a></i></li> <li>†<i><a href="/wiki/Yaminuechelys" title="Yaminuechelys">Yaminuechelys</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:11em"><a href="/wiki/Pelomedusidae" title="Pelomedusidae">Pelomedusidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/African_helmeted_turtle" title="African helmeted turtle">Pelomedusa</a></i></li> <li><i><a href="/wiki/Pelusios" title="Pelusios">Pelusios</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:11em"><a href="/wiki/Podocnemididae" title="Podocnemididae">Podocnemididae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li>†<i><a href="/wiki/Albertwoodemys" title="Albertwoodemys">Albertwoodemys</a></i></li> <li>†<i><a href="/wiki/Bauruemys" title="Bauruemys">Bauruemys</a></i></li> <li>†<i><a href="/wiki/Brontochelys" title="Brontochelys">Brontochelys</a></i></li> <li>†<i><a href="/wiki/Caninemys" title="Caninemys">Caninemys</a></i></li> <li>†<i><a href="/wiki/Carbonemys" title="Carbonemys">Carbonemys</a></i></li> <li>†<i><a href="/wiki/Cerrejonemys" title="Cerrejonemys">Cerrejonemys</a></i></li> <li>†<i><a href="/wiki/Cordichelys" title="Cordichelys">Cordichelys</a></i></li> <li><i><a href="/wiki/Madagascan_big-headed_turtle" title="Madagascan big-headed turtle">Erymnochelys</a></i></li> <li>†<i><a href="/wiki/Lapparentemys" title="Lapparentemys">Lapparentemys</a></i></li> <li>†<i><a href="/wiki/Latentemys" title="Latentemys">Latentemys</a></i></li> <li><i><a href="/wiki/Big-headed_Amazon_River_turtle" title="Big-headed Amazon River turtle">Peltocephalus</a></i></li> <li><i><a href="/wiki/Podocnemis" title="Podocnemis">Podocnemis</a></i></li> <li>†<i><a href="/wiki/Stupendemys" title="Stupendemys">Stupendemys</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:11em">†<a href="/wiki/Sahonachelyidae" title="Sahonachelyidae">Sahonachelyidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li>†<i><a href="/wiki/Sahonachelys" title="Sahonachelys">Sahonachelys </a></i></li> <li>†<i><a href="/wiki/Sokatra" title="Sokatra">Sokatra</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:5em"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:11em"> </th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li>†<i><a href="/wiki/Caribemys" class="mw-redirect" title="Caribemys">Caribemys</a></i></li> <li>†<i><a href="/wiki/Caririemys" title="Caririemys">Caririemys</a></i></li> <li>†<i><a href="/wiki/Tacuarembemys" title="Tacuarembemys">Tacuarembemys</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li>Phylogenetic arrangement of turtles based on <a rel="nofollow" class="external text" href="http://images.turtleconservancy.org/documents/2017/crm-7-checklist-atlas-v8-2017.pdf"><i>Turtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status</i></a>. † = extinct.</li></ul> <ul><li>See also <a href="/wiki/List_of_Testudines_families" title="List of Testudines families">List of Testudines families</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Chelus</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q14566138" class="extiw" title="wikidata:Q14566138">Q14566138</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Chelus" class="extiw" title="wikispecies:Chelus">Chelus</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Barcode_of_Life_Data_System" title="Barcode of Life Data System">BOLD</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://www.boldsystems.org/index.php/TaxBrowser_TaxonPage?taxid=171553">171553</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Catalogue_of_Life" title="Catalogue of Life">CoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.catalogueoflife.org/data/taxon/87DY8">87DY8</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/83728">83728</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/2441926">2441926</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/INaturalist" title="INaturalist">iNaturalist</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://inaturalist.org/taxa/39602">39602</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&id=1380855">1380855</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Integrated_Taxonomic_Information_System" title="Integrated Taxonomic Information System">ITIS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=208720">208720</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=44494">44494</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=235772">235772</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=37611">37611</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/World_Register_of_Marine_Species" title="World Register of Marine Species">WoRMS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.marinespecies.org/aphia.php?p=taxdetails&id=1358274">1358274</a></span></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Chelus fimbriatus</i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q317386" class="extiw" title="wikidata:Q317386">Q317386</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Chelus_fimbriata" class="extiw" title="wikispecies:Chelus fimbriata">Chelus fimbriata</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/795410">795410</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/2441927">2441927</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/INaturalist" title="INaturalist">iNaturalist</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://inaturalist.org/taxa/39603">39603</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&id=11008285">11008285</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Invasive_Species_Compendium" title="Invasive Species Compendium">ISC</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.cabi.org/isc/datasheet/113209">113209</a></span></span></li> <li><span style="white-space:nowrap;">NAS: <span class="uid"><a rel="nofollow" class="external text" href="https://nas.er.usgs.gov/queries/FactSheet.aspx?speciesID=1223">1223</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Observation.org" title="Observation.org">Observation.org</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://observation.org/species/98042/">98042</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=235773">235773</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Reptile_Database" title="Reptile Database">RD</a>: <span class="uid"><a class="external text" href="https://wikidata-externalid-url.toolforge.org/?url_prefix=https%3A%2F%2Freptile-database.reptarium.cz%2Fspecies%3F&id=genus%3DChelus%26species%3Dfimbriata">fimbriata</a></span></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox authority-control" aria-label="Navbox" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Help:Authority_control" title="Help:Authority control">Authority control databases</a>: National <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q317386#identifiers" title="Edit this at Wikidata"><img alt="Edit this at 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