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data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Gulingkan daftar isi" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Gulingkan daftar isi</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Mario Capecchi</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Pergi ke artikel dalam bahasa lain. Terdapat 54 bahasa" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-54" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">54 bahasa</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D9%85%D8%A7%D8%B1%D9%8A%D9%88_%D9%83%D8%A7%D8%A8%D9%8A%D9%83%D9%8A" title="ماريو كابيكي – Arab" lang="ar" hreflang="ar" data-title="ماريو كابيكي" data-language-autonym="العربية" data-language-local-name="Arab" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D9%85%D8%A7%D8%B1%D9%8A%D9%88_%D9%83%D8%A7%D8%A8%D9%8A%D9%83%D9%89" title="ماريو كابيكى – Arab Mesir" lang="arz" hreflang="arz" data-title="ماريو كابيكى" data-language-autonym="مصرى" data-language-local-name="Arab Mesir" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D9%85%D8%A7%D8%B1%DB%8C%D9%88_%DA%A9%D8%A7%D9%BE%DA%A9%DB%8C" title="ماریو کاپکی – South Azerbaijani" lang="azb" hreflang="azb" data-title="ماریو کاپکی" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%9C%D0%B0%D1%80%D1%8B%D0%B0_%D0%9A%D0%B0%D0%BF%D0%B5%D0%BA%D1%96" title="Марыа Капекі – Belarusia" lang="be" hreflang="be" data-title="Марыа Капекі" data-language-autonym="Беларуская" data-language-local-name="Belarusia" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%AE%E0%A6%BE%E0%A6%B0%E0%A6%BF%E0%A6%93_%E0%A6%95%E0%A6%BE%E0%A6%AA%E0%A7%87%E0%A6%95%E0%A7%8D%E0%A6%95%E0%A6%BF" title="মারিও কাপেক্কি – Bengali" lang="bn" hreflang="bn" data-title="মারিও কাপেক্কি" data-language-autonym="বাংলা" data-language-local-name="Bengali" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Katalan" lang="ca" hreflang="ca" data-title="Mario Capecchi" data-language-autonym="Català" data-language-local-name="Katalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Cheska" lang="cs" hreflang="cs" data-title="Mario Capecchi" data-language-autonym="Čeština" data-language-local-name="Cheska" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Dansk" lang="da" hreflang="da" data-title="Mario Capecchi" data-language-autonym="Dansk" data-language-local-name="Dansk" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Jerman" lang="de" hreflang="de" data-title="Mario Capecchi" data-language-autonym="Deutsch" data-language-local-name="Jerman" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Inggris" lang="en" hreflang="en" data-title="Mario Capecchi" data-language-autonym="English" data-language-local-name="Inggris" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Spanyol" lang="es" hreflang="es" data-title="Mario Capecchi" data-language-autonym="Español" data-language-local-name="Spanyol" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Esti" lang="et" hreflang="et" data-title="Mario Capecchi" data-language-autonym="Eesti" data-language-local-name="Esti" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Basque" lang="eu" hreflang="eu" data-title="Mario Capecchi" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D9%85%D8%A7%D8%B1%DB%8C%D9%88_%DA%A9%D8%A7%D9%BE%DA%A9%DB%8C" title="ماریو کاپکی – Persia" lang="fa" hreflang="fa" data-title="ماریو کاپکی" data-language-autonym="فارسی" data-language-local-name="Persia" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Suomi" lang="fi" hreflang="fi" data-title="Mario Capecchi" data-language-autonym="Suomi" data-language-local-name="Suomi" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Prancis" lang="fr" hreflang="fr" data-title="Mario Capecchi" data-language-autonym="Français" data-language-local-name="Prancis" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%9E%D7%A8%D7%99%D7%95_%D7%A7%D7%A4%D7%A7%D7%99" title="מריו קפקי – Ibrani" lang="he" hreflang="he" data-title="מריו קפקי" data-language-autonym="עברית" data-language-local-name="Ibrani" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%AE%E0%A4%BE%E0%A4%B0%E0%A4%BF%E0%A4%AF%E0%A5%8B_%E0%A4%B0%E0%A5%87%E0%A4%A8%E0%A4%BE%E0%A4%9F%E0%A5%8B_%E0%A4%95%E0%A5%88%E0%A4%AA%E0%A5%80%E0%A4%9A%E0%A5%80" title="मारियो रेनाटो कैपीची – Hindi" lang="hi" hreflang="hi" data-title="मारियो रेनाटो कैपीची" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Kroasia" lang="hr" hreflang="hr" data-title="Mario Capecchi" data-language-autonym="Hrvatski" data-language-local-name="Kroasia" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Hungaria" lang="hu" hreflang="hu" data-title="Mario Capecchi" data-language-autonym="Magyar" data-language-local-name="Hungaria" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-hy mw-list-item"><a href="https://hy.wikipedia.org/wiki/%D5%84%D5%A1%D6%80%D5%AB%D5%B8_%D4%BF%D5%A1%D5%BA%D5%A5%D5%AF%D5%AB" title="Մարիո Կապեկի – Armenia" lang="hy" hreflang="hy" data-title="Մարիո Կապեկի" data-language-autonym="Հայերեն" data-language-local-name="Armenia" class="interlanguage-link-target"><span>Հայերեն</span></a></li><li class="interlanguage-link interwiki-ia mw-list-item"><a href="https://ia.wikipedia.org/wiki/Mario_R_Capecchi" title="Mario R Capecchi – Interlingua" lang="ia" hreflang="ia" data-title="Mario R Capecchi" data-language-autonym="Interlingua" data-language-local-name="Interlingua" class="interlanguage-link-target"><span>Interlingua</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Ido" lang="io" hreflang="io" data-title="Mario Capecchi" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Italia" lang="it" hreflang="it" data-title="Mario Capecchi" data-language-autonym="Italiano" data-language-local-name="Italia" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%83%9E%E3%83%AA%E3%82%AA%E3%83%BB%E3%82%AB%E3%83%9A%E3%83%83%E3%82%AD" title="マリオ・カペッキ – Jepang" lang="ja" hreflang="ja" data-title="マリオ・カペッキ" data-language-autonym="日本語" data-language-local-name="Jepang" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-jv mw-list-item"><a href="https://jv.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Jawa" lang="jv" hreflang="jv" data-title="Mario Capecchi" data-language-autonym="Jawa" data-language-local-name="Jawa" class="interlanguage-link-target"><span>Jawa</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%9C%D0%B0%D1%80%D0%B8%D0%BE_%D0%9A%D0%B0%D0%BF%D0%B5%D0%BA%D0%BA%D0%B8" title="Марио Капекки – Kazakh" lang="kk" hreflang="kk" data-title="Марио Капекки" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EB%A7%88%EB%A6%AC%EC%98%A4_%EC%B9%B4%ED%8E%98%ED%82%A4" title="마리오 카페키 – Korea" lang="ko" hreflang="ko" data-title="마리오 카페키" data-language-autonym="한국어" data-language-local-name="Korea" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Marius_Capecchi" title="Marius Capecchi – Latin" lang="la" hreflang="la" data-title="Marius Capecchi" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-mg mw-list-item"><a href="https://mg.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Malagasi" lang="mg" hreflang="mg" data-title="Mario Capecchi" data-language-autonym="Malagasy" data-language-local-name="Malagasi" class="interlanguage-link-target"><span>Malagasy</span></a></li><li class="interlanguage-link interwiki-mk mw-list-item"><a href="https://mk.wikipedia.org/wiki/%D0%9C%D0%B0%D1%80%D0%B8%D0%BE_%D0%9A%D0%B0%D0%BF%D0%B5%D0%BA%D0%B8" title="Марио Капеки – Makedonia" lang="mk" hreflang="mk" data-title="Марио Капеки" data-language-autonym="Македонски" data-language-local-name="Makedonia" class="interlanguage-link-target"><span>Македонски</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Melayu" lang="ms" hreflang="ms" data-title="Mario Capecchi" data-language-autonym="Bahasa Melayu" data-language-local-name="Melayu" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D9%85%D8%A7%D8%B1%DB%8C%D9%88_%DA%A9%D8%A7%D9%BE%DA%A9%DB%8C" title="ماریو کاپکی – Mazanderani" lang="mzn" hreflang="mzn" data-title="ماریو کاپکی" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Mario_Renato_Capecchi" title="Mario Renato Capecchi – Jerman Rendah" lang="nds" hreflang="nds" data-title="Mario Renato Capecchi" data-language-autonym="Plattdüütsch" data-language-local-name="Jerman Rendah" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Belanda" lang="nl" hreflang="nl" data-title="Mario Capecchi" data-language-autonym="Nederlands" data-language-local-name="Belanda" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Mario_R._Capecchi" title="Mario R. Capecchi – Bokmål Norwegia" lang="nb" hreflang="nb" data-title="Mario R. Capecchi" data-language-autonym="Norsk bokmål" data-language-local-name="Bokmål Norwegia" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Ositania" lang="oc" hreflang="oc" data-title="Mario Capecchi" data-language-autonym="Occitan" data-language-local-name="Ositania" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Polski" lang="pl" hreflang="pl" data-title="Mario Capecchi" data-language-autonym="Polski" data-language-local-name="Polski" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D9%85%D8%A7%D8%B1%DB%8C%D9%88_%DA%A9%D8%A7%D9%BE%DB%8C%DA%86%DB%8C" title="ماریو کاپیچی – Western Punjabi" lang="pnb" hreflang="pnb" data-title="ماریو کاپیچی" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Portugis" lang="pt" hreflang="pt" data-title="Mario Capecchi" data-language-autonym="Português" data-language-local-name="Portugis" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9A%D0%B0%D0%BF%D0%B5%D0%BA%D0%BA%D0%B8,_%D0%9C%D0%B0%D1%80%D0%B8%D0%BE" title="Капекки, Марио – Rusia" lang="ru" hreflang="ru" data-title="Капекки, Марио" data-language-autonym="Русский" data-language-local-name="Rusia" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Serbo-Kroasia" lang="sh" hreflang="sh" data-title="Mario Capecchi" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="Serbo-Kroasia" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Simple English" lang="en-simple" hreflang="en-simple" data-title="Mario Capecchi" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Sloven" lang="sl" hreflang="sl" data-title="Mario Capecchi" data-language-autonym="Slovenščina" data-language-local-name="Sloven" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Swedia" lang="sv" hreflang="sv" data-title="Mario Capecchi" data-language-autonym="Svenska" data-language-local-name="Swedia" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Swahili" lang="sw" hreflang="sw" data-title="Mario Capecchi" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Turki" lang="tr" hreflang="tr" data-title="Mario Capecchi" data-language-autonym="Türkçe" data-language-local-name="Turki" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://tt.wikipedia.org/wiki/%D0%9C%D0%B0%D1%80%D0%B8%D0%BE_%D0%9A%D0%B0%D0%BF%D0%B5%D0%BA%D0%BA%D0%B8" title="Марио Капекки – Tatar" lang="tt" hreflang="tt" data-title="Марио Капекки" data-language-autonym="Татарча / tatarça" data-language-local-name="Tatar" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%9C%D0%B0%D1%80%D1%96%D0%BE_%D0%9A%D0%B0%D0%BF%D0%B5%D0%BA%D0%BA%D1%96" title="Маріо Капеккі – Ukraina" lang="uk" hreflang="uk" data-title="Маріо Капеккі" data-language-autonym="Українська" data-language-local-name="Ukraina" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D9%85%D8%A7%D8%B1%DB%8C%D9%88_%DA%A9%DB%8C%D9%BE%DB%8C%DA%A9%DB%8C" title="ماریو کیپیکی – Urdu" lang="ur" hreflang="ur" data-title="ماریو کیپیکی" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vec mw-list-item"><a href="https://vec.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Venesia" lang="vec" hreflang="vec" data-title="Mario Capecchi" data-language-autonym="Vèneto" data-language-local-name="Venesia" class="interlanguage-link-target"><span>Vèneto</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Vietnam" lang="vi" hreflang="vi" data-title="Mario Capecchi" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnam" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/Mario_Capecchi" title="Mario Capecchi – Yoruba" lang="yo" hreflang="yo" data-title="Mario Capecchi" data-language-autonym="Yorùbá" data-language-local-name="Yoruba" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E9%A9%AC%E9%87%8C%E5%A5%A5%C2%B7%E5%8D%A1%E4%BD%A9%E5%A5%87" title="马里奥·卡佩奇 – Tionghoa" lang="zh" hreflang="zh" data-title="马里奥·卡佩奇" data-language-autonym="中文" data-language-local-name="Tionghoa" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q215574#sitelinks-wikipedia" title="Sunting pranala interwiki" class="wbc-editpage">Sunting pranala</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div 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.infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}body.skin-minerva .mw-parser-output .infobox-header,body.skin-minerva .mw-parser-output .infobox-subheader,body.skin-minerva .mw-parser-output .infobox-above,body.skin-minerva .mw-parser-output .infobox-title,body.skin-minerva .mw-parser-output .infobox-image,body.skin-minerva .mw-parser-output .infobox-full-data,body.skin-minerva .mw-parser-output .infobox-below{text-align:center}</style><table class="infobox biography vcard"><tbody><tr><th colspan="2" class="infobox-above" style="font-size:125%;"><span class="fn">Mario Capecchi</span></th></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/Berkas:Mario_Capecchi_UTHSCSA.JPG" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/0/07/Mario_Capecchi_UTHSCSA.JPG/220px-Mario_Capecchi_UTHSCSA.JPG" decoding="async" width="220" height="189" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/0/07/Mario_Capecchi_UTHSCSA.JPG 1.5x" data-file-width="298" data-file-height="256" /></a></span></td></tr><tr><th scope="row" class="infobox-label">Lahir</th><td class="infobox-data" style="padding-right:0em; padding-right:0em">6 Oktober 1937<span class="noprint"> (umur 87)</span><br /><span class="birthplace"><span class="flagicon"><span class="mw-image-border" typeof="mw:File"><a href="/wiki/Italia" title="Italia"><img alt="Italia" src="//upload.wikimedia.org/wikipedia/commons/thumb/0/03/Flag_of_Italy.svg/23px-Flag_of_Italy.svg.png" decoding="async" width="23" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/03/Flag_of_Italy.svg/35px-Flag_of_Italy.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/03/Flag_of_Italy.svg/45px-Flag_of_Italy.svg.png 2x" data-file-width="1500" data-file-height="1000" /></a></span></span> <a href="/wiki/Verona" title="Verona">Verona</a>, <a href="/wiki/Italia" title="Italia">Italia</a></span></td></tr><tr><th scope="row" class="infobox-label">Almamater</th><td class="infobox-data" style="padding-right:0em; padding-right:0em"><a href="/w/index.php?title=Antioch_College&action=edit&redlink=1" class="new" title="Antioch College (halaman belum tersedia)">Antioch College</a>, <a href="/wiki/Ohio" title="Ohio">Ohio</a><br /><a href="/wiki/Universitas_Harvard" title="Universitas Harvard">Universitas Harvard</a></td></tr><tr><th scope="row" class="infobox-label">Dikenal atas</th><td class="infobox-data" style="padding-right:0em; padding-right:0em"><i><a href="/w/index.php?title=Knockout_mouse&action=edit&redlink=1" class="new" title="Knockout mouse (halaman belum tersedia)">Knockout mouse</a></i></td></tr><tr><th scope="row" class="infobox-label">Penghargaan</th><td class="infobox-data" style="padding-right:0em; padding-right:0em"><a href="/wiki/Albert_Lasker_Award_for_Basic_Medical_Research" class="mw-redirect" title="Albert Lasker Award for Basic Medical Research">Albert Lasker Award for Basic Medical Research</a> (2001)<br /><a href="/wiki/Penghargaan_Wolf" title="Penghargaan Wolf">Penghargaan Wolf</a> dalam Kedokteran (2002)<br /><span typeof="mw:File"><a href="/wiki/Berkas:Nobel_prize_medal.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/20px-Nobel_prize_medal.svg.png" decoding="async" width="20" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/30px-Nobel_prize_medal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/40px-Nobel_prize_medal.svg.png 2x" data-file-width="150" data-file-height="150" /></a></span> <a href="/wiki/Penghargaan_Nobel_dalam_Fisiologi_atau_Kedokteran" class="mw-redirect" title="Penghargaan Nobel dalam Fisiologi atau Kedokteran">Penghargaan Nobel dalam Fisiologi atau Kedokteran</a> (2007)</td></tr><tr><td colspan="2" class="infobox-full-data" style="padding-right:0em; padding-right:0em"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r22657712"><b>Karier ilmiah</b></td></tr><tr><th scope="row" class="infobox-label">Bidang</th><td class="infobox-data category"><a href="/wiki/Genetika" title="Genetika">Genetika</a></td></tr><tr><th scope="row" class="infobox-label">Institusi</th><td class="infobox-data"><a href="/w/index.php?title=Harvard_School_of_Medicine&action=edit&redlink=1" class="new" title="Harvard School of Medicine (halaman belum tersedia)">Harvard School of Medicine</a><br /><a href="/wiki/Universitas_Utah" title="Universitas Utah">Universitas Utah</a></td></tr><tr style="display:none"><td colspan="2"> </td></tr></tbody></table> <p><b>Mario Renato Capecchi</b> (lahir 6 Oktober 1937) adalah seorang <a href="/w/index.php?title=Genetikawan&action=edit&redlink=1" class="new" title="Genetikawan (halaman belum tersedia)">genetikawan</a> molekuler berkebangsaan <a href="/wiki/Amerika_Serikat" title="Amerika Serikat">Amerika Serikat</a> yang dilahirkan di <a href="/wiki/Italia" title="Italia">Italia</a> dan pemenang bersama <a href="/wiki/Penghargaan_Nobel_dalam_Fisiologi" class="mw-redirect" title="Penghargaan Nobel dalam Fisiologi">Penghargaan Nobel dalam Fisiologi</a> tahun 2007. Saat ini ia adalah Profesor Luar Biasa <a href="/w/index.php?title=Genetika_manusia&action=edit&redlink=1" class="new" title="Genetika manusia (halaman belum tersedia)">genetika manusia</a> dan <a href="/wiki/Biologi" title="Biologi">biologi</a> di Sekolah Kedokteran <a href="/wiki/Universitas_Utah" title="Universitas Utah">Universitas Utah</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Tahun_awal">Tahun awal</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Mario_Capecchi&veaction=edit&section=1" title="Sunting bagian: Tahun awal" class="mw-editsection-visualeditor"><span>sunting</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Mario_Capecchi&action=edit&section=1" title="Sunting kode sumber bagian: Tahun awal"><span>sunting sumber</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Mario Capecchi dilahirkan di <a href="/wiki/Verona" title="Verona">Verona</a>, <a href="/wiki/Italia" title="Italia">Italia</a> pada 1937 dari Luciano Capecchi, seorang penerbang Italia yang kemudian dilaporkan hilang dalam tugas selagi mengoperasikan <a href="/w/index.php?title=Senapan_anti-pesawat&action=edit&redlink=1" class="new" title="Senapan anti-pesawat (halaman belum tersedia)">senapan anti-pesawat</a> di <a href="/wiki/Libya" title="Libya">Libya</a>, dan Lucy Ramberg, dilahirkan di Amerika dari seorang pelukis <a href="/wiki/Impresionis" class="mw-redirect" title="Impresionis">Impresionis</a> <a href="/w/index.php?title=Lucy_Dodd_Ramberg&action=edit&redlink=1" class="new" title="Lucy Dodd Ramberg (halaman belum tersedia)">Lucy Dodd Ramberg</a> dan arkeologis Jerman Walter Ramberg. </p><p>Pada <a href="/wiki/Perang_Dunia_II" title="Perang Dunia II">Perang Dunia II</a>, ibu Mario Capecchi dikirim ke kamp konsentrasi <a href="/wiki/Dachau" title="Dachau">Dachau</a> sebagai hukuman karena merupakan anggota dari kelompok anti <a href="/wiki/Fasisme" title="Fasisme">Fasis</a>, meninggalkan anaknya yang baru berumur 4 tahun ke asuhan keluarga petani Italia. Meskipun ibunya telah menjual harta-benda dan diberikan kepada keluarga petani tersebut untuk mengasuh anaknya namun uang tersebut habis dalam setahun dan keluarga tersebut tak mampu lagi merawat Capecchi. Capecchi bertahan hidup di jalanan di utara Italia selama empat tahun ke depan. </p><p>Pada 1945, ibunya dibebaskan dari kamp oleh pasukan AS yang kemudian mencari anaknya dan menemukannya setahun kemudian dalam keadaan sekarat karena kekurangan gizi di sebuah rumah sakit. </p><p>Ibu dan anak tersebut kemudian pindah ke <a href="/wiki/Pennsylvania" title="Pennsylvania">Pennsylvania</a>, Amerika Serikat. Di sana mereka tinggal di tempat paman Cappechi dari pihak ibu, <a href="/w/index.php?title=Edward_Ramberg&action=edit&redlink=1" class="new" title="Edward Ramberg (halaman belum tersedia)">Edward Ramberg</a>, seorang fisikawan di <a href="/wiki/RCA" title="RCA">RCA</a>. </p><p>Capecchi menerima gelar <a href="/w/index.php?title=B.S.&action=edit&redlink=1" class="new" title="B.S. (halaman belum tersedia)">B.S.nya</a> dalam bidang <a href="/wiki/Kimia" title="Kimia">kimia</a> dan <a href="/wiki/Fisika" title="Fisika">fisika</a> pada 1961 dari <a href="/w/index.php?title=Antioch_College&action=edit&redlink=1" class="new" title="Antioch College (halaman belum tersedia)">Antioch College</a> di <a href="/wiki/Ohio" title="Ohio">Ohio</a>. Dia menerima gelar <a href="/wiki/Ph.D." class="mw-redirect" title="Ph.D.">Ph.D.nya</a> dalam bidang <a href="/wiki/Biofisika" title="Biofisika">biofisika</a> pada 1967 dari <a href="/wiki/Universitas_Harvard" title="Universitas Harvard">Universitas Harvard</a> dengan <a href="/wiki/Tesis" title="Tesis">tesis</a> doktor di bawah bimbingan <a href="/wiki/James_D._Watson" class="mw-redirect" title="James D. Watson">James D. Watson</a>. Capecchi pernah menjadi anggota Junior Fellow of the Society of Fellows di Universitas Harvard dari 1967 sampai 1969. Pada 1969 dia menjadi Asisten Profesor di Departemen Biokimia <a href="/w/index.php?title=Sekolah_Medis_Harvard&action=edit&redlink=1" class="new" title="Sekolah Medis Harvard (halaman belum tersedia)">Sekolah Medis Harvard</a>. Dia dipromosi menjadi Profesor <i>Associate</i> pada 1971. Pada 1973 dia bergabung ke <a href="/wiki/Universitas_Utah" title="Universitas Utah">Universitas Utah</a>. Sejak 1988 Capecchi juga pernah menjadi seorang investigator di <a href="/w/index.php?title=Howard_Hughes_Medical_Institute&action=edit&redlink=1" class="new" title="Howard Hughes Medical Institute (halaman belum tersedia)">Howard Hughes Medical Institute</a>. Dia adalah seorang anggota <a href="/w/index.php?title=National_Academy_of_Sciences&action=edit&redlink=1" class="new" title="National Academy of Sciences (halaman belum tersedia)">National Academy of Sciences</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Knockout_mouse"><i>Knockout mouse</i></h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Mario_Capecchi&veaction=edit&section=2" title="Sunting bagian: Knockout mouse" class="mw-editsection-visualeditor"><span>sunting</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Mario_Capecchi&action=edit&section=2" title="Sunting kode sumber bagian: Knockout mouse"><span>sunting sumber</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Mario Capecchi terkenal karena kerjanya dalam penargetan gen sel punca dari embrio tikus yang memungkinkan teknologi <a href="/w/index.php?title=Transgenik&action=edit&redlink=1" class="new" title="Transgenik (halaman belum tersedia)">transgenik</a> termasuk <a href="/wiki/Kloning" title="Kloning">kloning</a> dan modifikasi genetika. Hasil kerja ini dicapai melalui usaha <a href="/wiki/Martin_J._Evans" class="mw-redirect" title="Martin J. Evans">Martin J. Evans</a> dan <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a> yang meneliti<i> <a href="/w/index.php?title=Knockout_mouse&action=edit&redlink=1" class="new" title="Knockout mouse (halaman belum tersedia)">knockout mouse</a></i>. Hasil kerja mereka membuat mereka dianugerahi Penghargaan Nobel 2007 untuk kedokteran atau fisiologi. </p> <div class="mw-heading mw-heading2"><h2 id="Pranala_luar">Pranala luar</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Mario_Capecchi&veaction=edit&section=3" title="Sunting bagian: Pranala luar" class="mw-editsection-visualeditor"><span>sunting</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Mario_Capecchi&action=edit&section=3" title="Sunting kode sumber bagian: Pranala luar"><span>sunting sumber</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><span style="font-size:0.95em; font-weight:bold; color:#777; cursor:help;" title="Bahasa Inggris" lang="Inggris">(Inggris)</span> <a rel="nofollow" class="external text" href="http://www.hhmi.org/research/investigators/capecchi_bio.html">Howard Hughes Medical Institute biography</a></li> <li><span style="font-size:0.95em; font-weight:bold; color:#777; cursor:help;" title="Bahasa Inggris" lang="Inggris">(Inggris)</span> <a rel="nofollow" class="external text" href="http://www.genetics.utah.edu/faculty/mcapecchi.html">Eccles Institute of Human Genetics biography</a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20071011011144/http://www.genetics.utah.edu/faculty/mcapecchi.html">Diarsipkan</a> 2007-10-11 di <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback Machine</a>.</li> <li><span style="font-size:0.95em; font-weight:bold; color:#777; cursor:help;" title="Bahasa Inggris" lang="Inggris">(Inggris)</span> <a rel="nofollow" class="external text" href="http://www.neuroscience.med.utah.edu/Faculty/Capecchi.html">University of Utah biography</a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20071011005858/http://www.neuroscience.med.utah.edu/Faculty/Capecchi.html">Diarsipkan</a> 2007-10-11 di <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback Machine</a>.</li> <li><span style="font-size:0.95em; font-weight:bold; color:#777; cursor:help;" title="Bahasa Inggris" lang="Inggris">(Inggris)</span> <a rel="nofollow" class="external text" href="http://www.dnai.org/d/index.html?m=3,5">Interviews with Mario Capecchi from Dolan DNA Learning Center's <i>DNA Interactive</i></a></li> <li><span style="font-size:0.95em; font-weight:bold; color:#777; cursor:help;" title="Bahasa Inggris" lang="Inggris">(Inggris)</span> <a rel="nofollow" class="external text" href="http://www.dnaftb.org/dnaftb/41/concept/index.html">Capecchi animation from Dolan DNA Learning Center's <i>DNA from the Beginning</i></a></li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r23782733">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist 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title="Pembicaraan Templat:Penghargaan Nobel Fisiologi atau Kedokteran"><abbr title="Diskusikan templat ini">b</abbr></a></li><li class="nv-sunting"><a class="external text" href="https://id.wikipedia.org/w/index.php?title=Templat:Penghargaan_Nobel_Fisiologi_atau_Kedokteran&action=edit"><abbr title="Sunting templat ini">s</abbr></a></li></ul></div><div id="Penerima_Penghargaan_Nobel_Fisiologi_atau_Kedokteran" style="font-size:114%;margin:0 4em"><a href="/wiki/Daftar_penerima_Nobel_Fisiologi_atau_Kedokteran" title="Daftar penerima Nobel Fisiologi atau Kedokteran">Penerima</a> <a href="/wiki/Nobel_Fisiologi_atau_Kedokteran" class="mw-redirect" title="Nobel Fisiologi atau Kedokteran">Penghargaan Nobel Fisiologi atau Kedokteran</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">1901–1925</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1901: <a href="/wiki/Emil_von_Behring" class="mw-redirect" title="Emil von Behring">Emil Behring</a></span></li> <li><span class="nowrap">1902: <a href="/wiki/Ronald_Ross" title="Ronald Ross">Ronald Ross</a></span></li> <li><span class="nowrap">1903: <a href="/wiki/Niels_Ryberg_Finsen" title="Niels Ryberg Finsen">Niels Finsen</a></span></li> <li><span class="nowrap">1904: <a href="/wiki/Ivan_Pavlov" title="Ivan Pavlov">Ivan Pavlov</a></span></li> <li><span class="nowrap">1905: <a href="/wiki/Robert_Koch" title="Robert Koch">Robert Koch</a></span></li> <li><span class="nowrap">1906: <a href="/wiki/Camillo_Golgi" title="Camillo Golgi">Camillo Golgi</a> / <a href="/wiki/Santiago_Ram%C3%B3n_y_Cajal" title="Santiago Ramón y Cajal">Santiago Ramón y Cajal</a></span></li> <li><span class="nowrap">1907: <a href="/wiki/Charles_Louis_Alphonse_Laveran" class="mw-redirect" title="Charles Louis Alphonse Laveran">Alphonse Laveran</a></span></li> <li><span class="nowrap">1908: <a href="/wiki/%C3%89lie_Metchnikoff" class="mw-redirect" title="Élie Metchnikoff">Élie Metchnikoff</a> / <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Paul Ehrlich</a></span></li> <li><span class="nowrap">1909: <a href="/wiki/Emil_Theodor_Kocher" class="mw-redirect" title="Emil Theodor Kocher">Emil Kocher</a></span></li> <li><span class="nowrap">1910: <a href="/wiki/Albrecht_Kossel" class="mw-redirect" title="Albrecht Kossel">Albrecht Kossel</a></span></li> <li><span class="nowrap">1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Allvar Gullstrand</a></span></li> <li><span class="nowrap">1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Alexis Carrel</a></span></li> <li><span class="nowrap">1913: <a href="/wiki/Charles_Richet" class="mw-redirect" title="Charles Richet">Charles Richet</a></span></li> <li><span class="nowrap">1914: <a href="/wiki/R%C3%B3bert_B%C3%A1r%C3%A1ny" title="Róbert Bárány">Róbert Bárány</a></span></li> <li><span class="nowrap"><span style="color:darkgrey">1915</span></span></li> <li><span class="nowrap"><span style="color:darkgrey">1916</span></span></li> <li><span class="nowrap"><span style="color:darkgrey">1917</span></span></li> <li><span class="nowrap"><span style="color:darkgrey">1918</span></span></li> <li><span class="nowrap">1919: <a href="/wiki/Jules_Bordet" class="mw-redirect" title="Jules Bordet">Jules Bordet</a></span></li> <li><span class="nowrap">1920: <a href="/wiki/August_Krogh" class="mw-redirect" title="August Krogh">August Krogh</a></span></li> <li><span class="nowrap"><span style="color:darkgrey">1921</span></span></li> <li><span class="nowrap">1922: <a href="/wiki/Archibald_Hill" title="Archibald Hill">Archibald Hill</a> / <a href="/wiki/Otto_Fritz_Meyerhof" class="mw-redirect" title="Otto Fritz Meyerhof">Otto Meyerhof</a></span></li> <li><span class="nowrap">1923: <a href="/wiki/Frederick_Banting" class="mw-redirect" title="Frederick Banting">Frederick Banting</a> / <a href="/wiki/John_James_Rickard_Macleod" title="John James Rickard Macleod">John Macleod</a></span></li> <li><span class="nowrap">1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Willem Einthoven</a></span></li> <li><span class="nowrap"><span style="color:darkgrey">1925</span></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1926–1950</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1926: <a href="/wiki/Johannes_Fibiger" class="mw-redirect" title="Johannes Fibiger">Johannes Fibiger</a></span></li> <li><span class="nowrap">1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Julius Wagner-Jauregg</a></span></li> <li><span class="nowrap">1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Charles Nicolle</a></span></li> <li><span class="nowrap">1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Christiaan Eijkman</a> / <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Frederick Gowland Hopkins</a></span></li> <li><span class="nowrap">1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Karl Landsteiner</a></span></li> <li><span class="nowrap">1931: <a href="/wiki/Otto_Heinrich_Warburg" class="mw-redirect" title="Otto Heinrich Warburg">Otto Warburg</a></span></li> <li><span class="nowrap">1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Charles Scott Sherrington</a> / <a href="/wiki/Edgar_Adrian" class="mw-redirect" title="Edgar Adrian">Edgar Adrian</a></span></li> <li><span class="nowrap">1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Thomas Morgan</a></span></li> <li><span class="nowrap">1934: <a href="/wiki/George_Whipple" title="George Whipple">George Whipple</a> / <a href="/wiki/George_Minot" title="George Minot">George Minot</a> / <a href="/wiki/William_P._Murphy" class="mw-redirect" title="William P. Murphy">William Murphy</a></span></li> <li><span class="nowrap">1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Hans Spemann</a></span></li> <li><span class="nowrap">1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Henry Dale</a> / <a href="/wiki/Otto_Loewi" title="Otto Loewi">Otto Loewi</a></span></li> <li><span class="nowrap">1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Albert Szent-Györgyi</a></span></li> <li><span class="nowrap">1938: <a href="/wiki/Corneille_Heymans" title="Corneille Heymans">Corneille Heymans</a></span></li> <li><span class="nowrap">1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Gerhard Domagk</a></span></li> <li><span class="nowrap"><span style="color:darkgrey">1940</span></span></li> <li><span class="nowrap"><span style="color:darkgrey">1941</span></span></li> <li><span class="nowrap"><span style="color:darkgrey">1942</span></span></li> <li><span class="nowrap">1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Henrik Dam</a> / <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Edward Doisy</a></span></li> <li><span class="nowrap">1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Joseph Erlanger</a> / <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Herbert Gasser</a></span></li> <li><span class="nowrap">1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Alexander Fleming</a> / <a href="/wiki/Ernst_Chain" title="Ernst Chain">Ernst Chain</a> / <a href="/wiki/Howard_Florey" class="mw-redirect" title="Howard Florey">Howard Florey</a></span></li> <li><span class="nowrap">1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Hermann Muller</a></span></li> <li><span class="nowrap">1947: <a href="/wiki/Carl_Ferdinand_Cori" class="mw-redirect" title="Carl Ferdinand Cori">Carl Cori</a> / <a href="/wiki/Gerty_Cori" title="Gerty Cori">Gerty Cori</a> / <a href="/wiki/Bernardo_Houssay" class="mw-redirect" title="Bernardo Houssay">Bernardo Houssay</a></span></li> <li><span class="nowrap">1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Paul Müller</a></span></li> <li><span class="nowrap">1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Walter Hess</a> / <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">António Egas Moniz</a></span></li> <li><span class="nowrap">1950: <a href="/wiki/Edward_Calvin_Kendall" class="mw-redirect" title="Edward Calvin Kendall">Edward Kendall</a> / <a href="/wiki/Tadeusz_Reichstein" class="mw-redirect" title="Tadeusz Reichstein">Tadeusz Reichstein</a> / <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Philip Hench</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1951–1975</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Max Theiler</a></span></li> <li><span class="nowrap">1952: <a href="/wiki/Selman_Waksman" class="mw-redirect" title="Selman Waksman">Selman Waksman</a></span></li> <li><span class="nowrap">1953: <a href="/wiki/Hans_Adolf_Krebs" title="Hans Adolf Krebs">Hans Krebs</a> / <a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Fritz Lipmann</a></span></li> <li><span class="nowrap">1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">John Enders</a> / <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Thomas Weller</a> / <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Frederick Robbins</a></span></li> <li><span class="nowrap">1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Hugo Theorell</a></span></li> <li><span class="nowrap">1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" class="mw-redirect" title="André Frédéric Cournand">André Cournand</a> / <a href="/wiki/Werner_Forssmann" class="mw-redirect" title="Werner Forssmann">Werner Forssmann</a> / <a href="/wiki/Dickinson_W._Richards" class="mw-redirect" title="Dickinson W. Richards">Dickinson W. Richards</a></span></li> <li><span class="nowrap">1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Daniel Bovet</a></span></li> <li><span class="nowrap">1958: <a href="/wiki/George_Beadle" title="George Beadle">George Beadle</a> / <a href="/wiki/Edward_Tatum" title="Edward Tatum">Edward Tatum</a> / <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Joshua Lederberg</a></span></li> <li><span class="nowrap">1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Severo Ochoa</a> / <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Arthur Kornberg</a></span></li> <li><span class="nowrap">1960: <a href="/wiki/Macfarlane_Burnet" class="mw-redirect" title="Macfarlane Burnet">Frank Burnet</a> / <a href="/wiki/Peter_Medawar" class="mw-redirect" title="Peter Medawar">Peter Medawar</a></span></li> <li><span class="nowrap">1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Georg von Békésy</a></span></li> <li><span class="nowrap">1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Francis Crick</a> / <a href="/wiki/James_Watson" class="mw-redirect" title="James Watson">James Watson</a> / <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Maurice Wilkins</a></span></li> <li><span class="nowrap">1963: <a href="/wiki/John_Carew_Eccles" title="John Carew Eccles">John Eccles</a> / <a href="/wiki/Alan_Hodgkin" title="Alan Hodgkin">Alan Hodgkin</a> / <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Andrew Huxley</a></span></li> <li><span class="nowrap">1964: <a href="/wiki/Konrad_Emil_Bloch" class="mw-redirect" title="Konrad Emil Bloch">Konrad Bloch</a> / <a href="/wiki/Feodor_Lynen" class="mw-redirect" title="Feodor Lynen">Feodor Lynen</a></span></li> <li><span class="nowrap">1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">François Jacob</a> / <a href="/wiki/Andr%C3%A9_Michel_Lwoff" class="mw-redirect" title="André Michel Lwoff">André Lwoff</a> / <a href="/wiki/Jacques_Monod" class="mw-redirect" title="Jacques Monod">Jacques Monod</a></span></li> <li><span class="nowrap">1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Francis Rous</a> / <a href="/wiki/Charles_Brenton_Huggins" class="mw-redirect" title="Charles Brenton Huggins">Charles B. Huggins</a></span></li> <li><span class="nowrap">1967: <a href="/wiki/Ragnar_Granit" class="mw-redirect" title="Ragnar Granit">Ragnar Granit</a> / <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Haldan Hartline</a> / <a href="/wiki/George_Wald" title="George Wald">George Wald</a></span></li> <li><span class="nowrap">1968: <a href="/wiki/Robert_W._Holley" class="mw-redirect" title="Robert W. Holley">Robert W. Holley</a> / <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Har Khorana</a> / <a href="/wiki/Marshall_Warren_Nirenberg" class="mw-redirect" title="Marshall Warren Nirenberg">Marshall Nirenberg</a></span></li> <li><span class="nowrap">1969: <a href="/wiki/Max_Delbr%C3%BCck" class="mw-redirect" title="Max Delbrück">Max Delbrück</a> / <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Alfred Hershey</a> / <a href="/wiki/Salvador_Luria" class="mw-redirect" title="Salvador Luria">Salvador Luria</a></span></li> <li><span class="nowrap">1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Bernard Katz</a> / <a href="/wiki/Ulf_von_Euler" title="Ulf von Euler">Ulf von Euler</a> / <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Julius Axelrod</a></span></li> <li><span class="nowrap">1971: <a href="/wiki/Earl_Wilbur_Sutherland_Jr." class="mw-redirect" title="Earl Wilbur Sutherland Jr.">Earl Sutherland Jr.</a></span></li> <li><span class="nowrap">1972: <a href="/wiki/Gerald_Edelman" class="mw-redirect" title="Gerald Edelman">Gerald Edelman</a> / <a href="/wiki/Rodney_Robert_Porter" title="Rodney Robert Porter">Rodney Porter</a></span></li> <li><span class="nowrap">1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Karl von Frisch</a> / <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Konrad Lorenz</a> / <a href="/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen">Nikolaas Tinbergen</a></span></li> <li><span class="nowrap">1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Albert Claude</a> / <a href="/wiki/Christian_de_Duve" title="Christian de Duve">Christian de Duve</a> / <a href="/wiki/George_Emil_Palade" class="mw-redirect" title="George Emil Palade">George Palade</a></span></li> <li><span class="nowrap">1975: <a href="/wiki/David_Baltimore" title="David Baltimore">David Baltimore</a> / <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Renato Dulbecco</a> / <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Howard Temin</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1976–2000</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1976: <a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Baruch Blumberg</a> / <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Daniel Gajdusek</a></span></li> <li><span class="nowrap">1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Roger Guillemin</a> / <a href="/wiki/Andrew_Schally" class="mw-redirect" title="Andrew Schally">Andrew Schally</a> / <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Rosalyn Yalow</a></span></li> <li><span class="nowrap">1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Werner Arber</a> / <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Daniel Nathans</a> / <a href="/wiki/Hamilton_O._Smith" class="mw-redirect" title="Hamilton O. Smith">Hamilton O. Smith</a></span></li> <li><span class="nowrap">1979: <a href="/wiki/Allan_M._Cormack" title="Allan M. Cormack">Allan Cormack</a> / <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Godfrey Hounsfield</a></span></li> <li><span class="nowrap">1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Baruj Benacerraf</a> / <a href="/wiki/Jean_Dausset" class="mw-redirect" title="Jean Dausset">Jean Dausset</a> / <a href="/wiki/George_Davis_Snell" class="mw-redirect" title="George Davis Snell">George Snell</a></span></li> <li><span class="nowrap">1981: <a href="/wiki/Roger_Wolcott_Sperry" title="Roger Wolcott Sperry">Roger Sperry</a> / <a href="/wiki/David_H._Hubel" class="mw-redirect" title="David H. Hubel">David H. Hubel</a> / <a href="/wiki/Torsten_Wiesel" class="mw-redirect" title="Torsten Wiesel">Torsten Wiesel</a></span></li> <li><span class="nowrap">1982: <a href="/wiki/Sune_Bergstr%C3%B6m" title="Sune Bergström">Sune Bergström</a> / <a href="/wiki/Bengt_I._Samuelsson" class="mw-redirect" title="Bengt I. Samuelsson">Bengt I. Samuelsson</a> / <a href="/wiki/John_Vane" class="mw-redirect" title="John Vane">John Vane</a></span></li> <li><span class="nowrap">1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a></span></li> <li><span class="nowrap">1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Niels Jerne</a> / <a href="/wiki/Georges_J._F._K%C3%B6hler" class="mw-redirect" title="Georges J. F. Köhler">Georges Köhler</a> / <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a></span></li> <li><span class="nowrap">1985: <a href="/wiki/Michael_Stuart_Brown" class="mw-redirect" title="Michael Stuart Brown">Michael Brown</a> / <a href="/wiki/Joseph_L._Goldstein" class="mw-redirect" title="Joseph L. Goldstein">Joseph L. Goldstein</a></span></li> <li><span class="nowrap">1986: <a href="/wiki/Stanley_Cohen_(biokimiawan)" title="Stanley Cohen (biokimiawan)">Stanley Cohen</a> / <a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Rita Levi-Montalcini</a></span></li> <li><span class="nowrap">1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Susumu Tonegawa</a></span></li> <li><span class="nowrap">1988: <a href="/wiki/James_Whyte_Black" title="James Whyte Black">James W. Black</a> / <a href="/wiki/Gertrude_B._Elion" class="mw-redirect" title="Gertrude B. Elion">Gertrude B. Elion</a> / <a href="/wiki/George_H._Hitchings" class="mw-redirect" title="George H. Hitchings">George H. Hitchings</a></span></li> <li><span class="nowrap">1989: <a href="/wiki/J._Michael_Bishop" class="mw-redirect" title="J. Michael Bishop">J. Michael Bishop</a> / <a href="/wiki/Harold_E._Varmus" class="mw-redirect" title="Harold E. Varmus">Harold E. Varmus</a></span></li> <li><span class="nowrap">1990: <a href="/wiki/Joseph_Murray" class="mw-redirect" title="Joseph Murray">Joseph Murray</a> / <a href="/wiki/E._Donnall_Thomas" class="mw-redirect" title="E. Donnall Thomas">E. Donnall Thomas</a></span></li> <li><span class="nowrap">1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Erwin Neher</a> / <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Bert Sakmann</a></span></li> <li><span class="nowrap">1992: <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Edmond Fischer</a> / <a href="/wiki/Edwin_G._Krebs" class="mw-redirect" title="Edwin G. Krebs">Edwin G. Krebs</a></span></li> <li><span class="nowrap">1993: <a href="/wiki/Richard_J._Roberts" title="Richard J. Roberts">Richard J. Roberts</a> / <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Phillip Sharp</a></span></li> <li><span class="nowrap">1994: <a href="/wiki/Alfred_G._Gilman" title="Alfred G. Gilman">Alfred G. Gilman</a> / <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Martin Rodbell</a></span></li> <li><span class="nowrap">1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Edward B. Lewis</a> / <a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Christiane Nüsslein-Volhard</a> / <a href="/wiki/Eric_F._Wieschaus" class="mw-redirect" title="Eric F. Wieschaus">Eric F. Wieschaus</a></span></li> <li><span class="nowrap">1996: <a href="/wiki/Peter_C._Doherty" class="mw-redirect" title="Peter C. Doherty">Peter C. Doherty</a> / <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Rolf M. Zinkernagel</a></span></li> <li><span class="nowrap">1997: <a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Stanley B. Prusiner</a></span></li> <li><span class="nowrap">1998: <a href="/wiki/Robert_F._Furchgott" class="mw-redirect" title="Robert F. Furchgott">Robert F. Furchgott</a> / <a href="/wiki/Louis_Ignarro" class="mw-redirect" title="Louis Ignarro">Louis Ignarro</a> / <a href="/wiki/Ferid_Murad" title="Ferid Murad">Ferid Murad</a></span></li> <li><span class="nowrap">1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Günter Blobel</a></span></li> <li><span class="nowrap">2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Arvid Carlsson</a> / <a href="/wiki/Paul_Greengard" title="Paul Greengard">Paul Greengard</a> / <a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric Kandel</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2001–sekarang</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">2001: <a href="/wiki/Leland_H._Hartwell" class="mw-redirect" title="Leland H. Hartwell">Leland H. Hartwell</a> / <a href="/wiki/Tim_Hunt" class="mw-redirect" title="Tim Hunt">Tim Hunt</a> / <a href="/wiki/Paul_Nurse" class="mw-redirect" title="Paul Nurse">Paul Nurse</a></span></li> <li><span class="nowrap">2002: <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Sydney Brenner</a> / <a href="/wiki/H._Robert_Horvitz" class="mw-redirect" title="H. Robert Horvitz">H. Robert Horvitz</a> / <a href="/wiki/John_Sulston" class="mw-redirect" title="John Sulston">John E. Sulston</a></span></li> <li><span class="nowrap">2003: <a href="/wiki/Paul_Lauterbur" class="mw-redirect" title="Paul Lauterbur">Paul Lauterbur</a> / <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Peter Mansfield</a></span></li> <li><span class="nowrap">2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Richard Axel</a> / <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Linda B. Buck</a></span></li> <li><span class="nowrap">2005: <a href="/wiki/Barry_Marshall" class="mw-redirect" title="Barry Marshall">Barry Marshall</a> / <a href="/wiki/Robin_Warren" class="mw-redirect" title="Robin Warren">Robin Warren</a></span></li> <li><span class="nowrap">2006: <a href="/wiki/Andrew_Fire" class="mw-redirect" title="Andrew Fire">Andrew Fire</a> / <a href="/wiki/Craig_Mello" class="mw-redirect" title="Craig Mello">Craig Mello</a></span></li> <li><span class="nowrap">2007: <a class="mw-selflink selflink">Mario Capecchi</a> / <a href="/wiki/Martin_Evans" title="Martin Evans">Martin Evans</a> / <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a></span></li> <li><span class="nowrap">2008: <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Harald zur Hausen</a> / <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Luc Montagnier</a> / <a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Françoise Barré-Sinoussi</a></span></li> <li><span class="nowrap">2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Elizabeth Blackburn</a> / <a href="/wiki/Carol_W._Greider" title="Carol W. Greider">Carol W. Greider</a> / <a href="/wiki/Jack_W._Szostak" title="Jack W. Szostak">Jack W. Szostak</a></span></li> <li><span class="nowrap">2010: <a href="/wiki/Robert_Edwards" title="Robert Edwards">Robert G. Edwards</a></span></li> <li><span class="nowrap">2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Bruce Beutler</a> / <a href="/wiki/Jules_A._Hoffmann" title="Jules A. Hoffmann">Jules A. Hoffmann</a> / <a href="/wiki/Ralph_M._Steinman" title="Ralph M. Steinman">Ralph M. Steinman</a> (<i>anumerta</i>)</span></li> <li><span class="nowrap">2012: <a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> / <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Shinya Yamanaka</a></span></li> <li><span class="nowrap">2013: <a href="/wiki/James_Rothman" class="mw-redirect" title="James Rothman">James Rothman</a> / <a href="/wiki/Randy_Schekman" title="Randy Schekman">Randy Schekman</a> / <a href="/wiki/Thomas_C._S%C3%BCdhof" title="Thomas C. Südhof">Thomas C. Südhof</a></span></li> <li><span class="nowrap">2014: <a href="/wiki/John_O%27Keefe" title="John O'Keefe">John O'Keefe</a> / <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">May-Britt Moser</a> / <a href="/wiki/Edvard_Moser" title="Edvard Moser">Edvard Moser</a></span></li> <li><span class="nowrap">2015: <a href="/wiki/William_C._Campbell_(ilmuwan)" class="mw-redirect" title="William C. Campbell (ilmuwan)">William C. Campbell</a> / <a href="/wiki/Satoshi_%C5%8Cmura" title="Satoshi Ōmura">Satoshi Ōmura</a> / <a href="/wiki/Tu_Youyou" title="Tu Youyou">Tu Youyou</a></span></li> <li><span class="nowrap">2016: <a href="/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi">Yoshinori Ohsumi</a></span></li> <li><span class="nowrap">2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Jeffrey C. Hall</a> / <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Michael Rosbash</a> / <a href="/wiki/Michael_W._Young" title="Michael W. Young">Michael W. Young</a></span></li> <li><span class="nowrap">2018: <a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a> / <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Tasuku Honjo</a></span></li> <li><span class="nowrap">2019: <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Gregg L. Semenza</a> / <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Peter J. Ratcliffe</a> / <a href="/wiki/William_Kaelin_Jr." title="William Kaelin Jr.">William Kaelin Jr.</a></span></li> <li><span class="nowrap">2020: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Harvey J. Alter</a> / <a href="/wiki/Michael_Houghton" title="Michael Houghton">Michael Houghton</a> / <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Charles M. Rice</a></span></li> <li><span class="nowrap">2021: <a href="/wiki/David_Julius" title="David Julius">David Julius</a> / <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Ardem Patapoutian</a></span></li> <li><span class="nowrap">2022: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Svante Pääbo</a></span></li> <li><span class="nowrap">2023: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Katalin Karikó</a> / <a href="/wiki/Drew_Weissman" title="Drew Weissman">Drew Weissman</a></span></li> <li><span class="nowrap">2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a> / <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r23782733"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r25847331"></div><div role="navigation" class="navbox" aria-labelledby="Daftar_penerima_penghargaan_Wolf_dalam_bidang_kedokteran" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r18590415"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-lihat"><a href="/wiki/Templat:Penghargaan_Wolf_bidang_Kedokteran" title="Templat:Penghargaan Wolf bidang Kedokteran"><abbr title="Lihat templat ini">l</abbr></a></li><li class="nv-bicara"><a href="/w/index.php?title=Pembicaraan_Templat:Penghargaan_Wolf_bidang_Kedokteran&action=edit&redlink=1" class="new" title="Pembicaraan Templat:Penghargaan Wolf bidang Kedokteran (halaman belum tersedia)"><abbr title="Diskusikan templat ini">b</abbr></a></li><li class="nv-sunting"><a class="external text" href="https://id.wikipedia.org/w/index.php?title=Templat:Penghargaan_Wolf_bidang_Kedokteran&action=edit"><abbr title="Sunting templat ini">s</abbr></a></li></ul></div><div id="Daftar_penerima_penghargaan_Wolf_dalam_bidang_kedokteran" style="font-size:114%;margin:0 4em">Daftar penerima <a href="/wiki/Penghargaan_Wolf" title="Penghargaan Wolf">penghargaan Wolf</a> dalam bidang <a href="/wiki/Daftar_penerima_penghargaan_Wolf_dalam_bidang_kedokteran" class="mw-redirect" title="Daftar penerima penghargaan Wolf dalam bidang kedokteran">kedokteran</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">1970-an</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/George_Davis_Snell" class="mw-redirect" title="George Davis Snell">George Snell</a> / <a href="/wiki/Jean_Dausset" class="mw-redirect" title="Jean Dausset">Jean Dausset</a> / <a href="/w/index.php?title=Jon_van_Rood&action=edit&redlink=1" class="new" title="Jon van Rood (halaman belum tersedia)">Jon J. van Rood</a> (1978)</li> <li><a href="/wiki/Roger_Wolcott_Sperry" title="Roger Wolcott Sperry">Roger Sperry</a> / <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Arvid Carlsson</a> / <a href="/w/index.php?title=Oleh_Hornykiewicz&action=edit&redlink=1" class="new" title="Oleh Hornykiewicz (halaman belum tersedia)">Oleh Hornykiewicz</a> (1979)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1980-an</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a> / <a href="/w/index.php?title=Leo_Sachs&action=edit&redlink=1" class="new" title="Leo Sachs (halaman belum tersedia)">Leo Sachs</a> / <a href="/w/index.php?title=James_Learmonth_Gowans&action=edit&redlink=1" class="new" title="James Learmonth Gowans (halaman belum tersedia)">James L. Gowans</a> (1980)</li> <li><a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a> / <a href="/wiki/Stanley_Norman_Cohen" title="Stanley Norman Cohen">Stanley Norman Cohen</a> (1981)</li> <li><a href="/w/index.php?title=Jean-Pierre_Changeux&action=edit&redlink=1" class="new" title="Jean-Pierre Changeux (halaman belum tersedia)">Jean-Pierre Changeux</a> / <a href="/w/index.php?title=Solomon_H._Snyder&action=edit&redlink=1" class="new" title="Solomon H. Snyder (halaman belum tersedia)">Solomon H. Snyder</a> / <a href="/wiki/James_Whyte_Black" title="James Whyte Black">James W. Black</a> (1982)</li> <li><a href="/w/index.php?title=Donald_F._Steiner&action=edit&redlink=1" class="new" title="Donald F. Steiner (halaman belum tersedia)">Donald F. Steiner</a> (1984/5)</li> <li><a href="/w/index.php?title=Osamu_Hayaishi&action=edit&redlink=1" class="new" title="Osamu Hayaishi (halaman belum tersedia)">Osamu Hayaishi</a> (1986)</li> <li><a href="/w/index.php?title=Pedro_Cuatrecasas&action=edit&redlink=1" class="new" title="Pedro Cuatrecasas (halaman belum tersedia)">Pedro Cuatrecasas</a> / <a href="/w/index.php?title=Meir_Wilchek&action=edit&redlink=1" class="new" title="Meir Wilchek (halaman belum tersedia)">Meir Wilchek</a> (1987)</li> <li><a href="/w/index.php?title=Henri_G._Hers&action=edit&redlink=1" class="new" title="Henri G. Hers (halaman belum tersedia)">Henri G. Hers</a> / <a href="/w/index.php?title=Elizabeth_F._Neufeld&action=edit&redlink=1" class="new" title="Elizabeth F. Neufeld (halaman belum tersedia)">Elizabeth F. Neufeld</a> (1988)</li> <li><a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> / <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Edward B. Lewis</a> (1989)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1990-an</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/w/index.php?title=Maclyn_McCarty&action=edit&redlink=1" class="new" title="Maclyn McCarty (halaman belum tersedia)">Maclyn McCarty</a> (1990)</li> <li><a href="/w/index.php?title=Seymour_Benzer&action=edit&redlink=1" class="new" title="Seymour Benzer (halaman belum tersedia)">Seymour Benzer</a> (1991)</li> <li><a href="/w/index.php?title=Judah_Folkman&action=edit&redlink=1" class="new" title="Judah Folkman (halaman belum tersedia)">Judah Folkman</a> (1992)</li> <li><a href="/wiki/Michael_Berridge" title="Michael Berridge">Michael Berridge</a> / <a href="/w/index.php?title=Yasutomi_Nishizuka&action=edit&redlink=1" class="new" title="Yasutomi Nishizuka (halaman belum tersedia)">Yasutomi Nishizuka</a> (1994/5)</li> <li><a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Stanley B. Prusiner</a> (1995/6)</li> <li><a href="/w/index.php?title=Mary_F._Lyon&action=edit&redlink=1" class="new" title="Mary F. Lyon (halaman belum tersedia)">Mary F. Lyon</a> (1996/7)</li> <li><a href="/wiki/Michael_Sela" title="Michael Sela">Michael Sela</a> / <a href="/w/index.php?title=Ruth_Arnon&action=edit&redlink=1" class="new" title="Ruth Arnon (halaman belum tersedia)">Ruth Arnon</a> (1998)</li> <li><a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric Kandel</a> (1999)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2000-an</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Avram_Hershko" title="Avram Hershko">Avram Hershko</a> / <a href="/w/index.php?title=Alexander_Varshavsky&action=edit&redlink=1" class="new" title="Alexander Varshavsky (halaman belum tersedia)">Alexander Varshavsky</a> (2001)</li> <li><a href="/w/index.php?title=Ralph_L._Brinster&action=edit&redlink=1" class="new" title="Ralph L. Brinster (halaman belum tersedia)">Ralph L. Brinster</a> / <a class="mw-selflink selflink">Mario Capecchi</a> / <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a> (2002/3)</li> <li><a href="/w/index.php?title=Robert_Weinberg&action=edit&redlink=1" class="new" title="Robert Weinberg (halaman belum tersedia)">Robert Weinberg</a> / <a href="/wiki/Roger_Y._Tsien" class="mw-redirect" title="Roger Y. Tsien">Roger Y. Tsien</a> (2004)</li> <li><a href="/w/index.php?title=Alexander_Levitzki&action=edit&redlink=1" class="new" title="Alexander Levitzki (halaman belum tersedia)">Alexander Levitzki</a> / <a href="/w/index.php?title=Anthony_R._Hunter&action=edit&redlink=1" class="new" title="Anthony R. Hunter (halaman belum tersedia)">Anthony R. Hunter</a> / <a href="/w/index.php?title=Anthony_Pawson&action=edit&redlink=1" class="new" title="Anthony Pawson (halaman belum tersedia)">Anthony Pawson</a> (2005)</li> <li><a href="/w/index.php?title=Howard_Cedar&action=edit&redlink=1" class="new" title="Howard Cedar (halaman belum tersedia)">Howard Cedar</a> / <a href="/w/index.php?title=Aharon_Razin&action=edit&redlink=1" class="new" title="Aharon Razin (halaman belum tersedia)">Aharon Razin</a> (2008)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2010-an</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/w/index.php?title=Axel_Ullrich&action=edit&redlink=1" class="new" title="Axel Ullrich (halaman belum tersedia)">Axel Ullrich</a> (2010)</li> <li><a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Shinya Yamanaka</a> / <a href="/w/index.php?title=Rudolf_Jaenisch&action=edit&redlink=1" class="new" title="Rudolf Jaenisch (halaman belum tersedia)">Rudolf Jaenisch</a> (2011)</li> <li><a href="/w/index.php?title=Ronald_M._Evans&action=edit&redlink=1" class="new" title="Ronald M. Evans (halaman belum tersedia)">Ronald M. Evans</a> (2012)</li> <li><a href="/w/index.php?title=Nahum_Sonenberg&action=edit&redlink=1" class="new" title="Nahum Sonenberg (halaman belum tersedia)">Nahum Sonenberg</a> / <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a> / <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a> (2014)</li> <li><a href="/w/index.php?title=John_Kappler&action=edit&redlink=1" class="new" title="John Kappler (halaman belum tersedia)">John Kappler</a> / <a href="/w/index.php?title=Philippa_Marrack&action=edit&redlink=1" class="new" title="Philippa Marrack (halaman belum tersedia)">Philippa Marrack</a> / <a href="/w/index.php?title=Jeffrey_V._Ravetch&action=edit&redlink=1" class="new" title="Jeffrey V. Ravetch (halaman belum tersedia)">Jeffrey V. Ravetch</a> (2015)</li> <li><a href="/w/index.php?title=Lewis_C._Cantley&action=edit&redlink=1" class="new" title="Lewis C. Cantley (halaman belum tersedia)">Lewis C. Cantley</a> / <a href="/w/index.php?title=C._Ronald_Kahn&action=edit&redlink=1" class="new" title="C. Ronald Kahn (halaman belum tersedia)">C. Ronald Kahn</a> (2016)</li> <li><a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a> (2017)</li> <li><a href="/wiki/Jeffrey_M._Friedman" title="Jeffrey M. Friedman">Jeffrey M. Friedman</a> (2019)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2020-an</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Emmanuelle_Charpentier" title="Emmanuelle Charpentier">Emmanuelle Charpentier</a> / <a href="/wiki/Jennifer_Doudna" title="Jennifer Doudna">Jennifer Doudna</a> (2020)</li></ul> </div> </td></tr><tr><td class="navbox-abovebelow" colspan="2"><div><div> <p>Templat penghargaan Wolf : <a href="/wiki/Templat:Penghargaan_Wolf_bidang_Agrikultur" class="mw-redirect" title="Templat:Penghargaan Wolf bidang Agrikultur">agrikultur</a><span style="font-weight:bold;"> ·</span>  <a href="/wiki/Templat:Penghargaan_Wolf_bidang_Kimia" title="Templat:Penghargaan Wolf bidang Kimia">kimia</a><span style="font-weight:bold;"> ·</span>  <a href="/wiki/Templat:Penghargaan_Wolf_bidang_Matematika" title="Templat:Penghargaan Wolf bidang Matematika">matematika</a><span style="font-weight:bold;"> ·</span>  <a href="/wiki/Templat:Penghargaan_Wolf_bidang_Kedokteran" title="Templat:Penghargaan Wolf bidang Kedokteran">kedokteran</a><span style="font-weight:bold;"> ·</span>  <a href="/wiki/Templat:Penghargaan_Wolf_bidang_Fisika" title="Templat:Penghargaan Wolf bidang Fisika">fisika</a><span style="font-weight:bold;"> ·</span>  <a href="/wiki/Templat:Penghargaan_Wolf_bidang_Seni" title="Templat:Penghargaan Wolf bidang Seni">seni</a> </p> </div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r23782733"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r25847331"></div><div role="navigation" class="navbox authority-control" aria-labelledby="Pengawasan_otoritas_frameless_&#124;text-top_&#124;10px_&#124;alt=Sunting_ini_di_Wikidata_&#124;link=https&#58;//www.wikidata.org/wiki/Q215574#identifiers&#124;Sunting_ini_di_Wikidata" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Pengawasan_otoritas_frameless_&#124;text-top_&#124;10px_&#124;alt=Sunting_ini_di_Wikidata_&#124;link=https&#58;//www.wikidata.org/wiki/Q215574#identifiers&#124;Sunting_ini_di_Wikidata" style="font-size:114%;margin:0 4em"><a href="/wiki/Bantuan:Pengawasan_otoritas" title="Bantuan:Pengawasan otoritas">Pengawasan otoritas</a> <span class="mw-valign-text-top" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q215574#identifiers" title="Sunting ini di Wikidata"><img alt="Sunting ini di Wikidata" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">Umum</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://d-nb.info/gnd/1217732470">Integrated Authority File (Jerman)</a></span></li> <li><a href="/wiki/ISNI_(identifier)" class="mw-redirect" title="ISNI (identifier)">ISNI</a> <ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://isni.org/isni/0000000029897908">1</a></span></li></ul></li> <li><a href="/wiki/ORCID_(identifier)" class="mw-redirect" title="ORCID (identifier)">ORCID</a> <ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://orcid.org/0000-0002-9591-4993">1</a></span></li></ul></li> <li><a href="/wiki/VIAF_(identifier)" class="mw-redirect" title="VIAF (identifier)">VIAF</a> <ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://viaf.org/viaf/201366364">1</a></span></li></ul></li> <li><span class="nowrap"><a rel="nofollow" class="external text" href="https://www.worldcat.org/identities/containsVIAFID/201366364">WorldCat (via VIAF)</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Perpustakaan nasional</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/names/n90604233">Amerika Serikat</a></span></li> <li><span class="uid"><a rel="nofollow" class="external text" href="http://mak.bn.org.pl/cgi-bin/KHW/makwww.exe?BM=1&NU=1&IM=4&WI=9810678350505606">Polandia</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Basis data ilmiah</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://ci.nii.ac.jp/author/DA04475098?l=en">CiNii (Jepang)</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Lain-lain</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/SUDOC_(identifier)" class="mw-redirect" title="SUDOC (identifier)">SUDOC (Prancis)</a> <ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.idref.fr/130721328">1</a></span></li></ul></li></ul> </div></td></tr></tbody></table></div>    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