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Johannes Fibiger - Wikipedia

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subsection</span> </button> <ul id="toc-Nobel_Prize_controversy-sublist" class="vector-toc-list"> <li id="toc-Nomination_and_selection" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Nomination_and_selection"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.1</span> <span>Nomination and selection</span> </div> </a> <ul id="toc-Nomination_and_selection-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Posthumous_criticism_and_disproof" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Posthumous_criticism_and_disproof"> <div class="vector-toc-text"> <span class="vector-toc-numb">4.2</span> <span>Posthumous criticism and disproof</span> </div> </a> <ul id="toc-Posthumous_criticism_and_disproof-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-See_also" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#See_also"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>See also</span> </div> </a> <ul id="toc-See_also-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Further_reading" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Further_reading"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>Further reading</span> </div> </a> <ul id="toc-Further_reading-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">8</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Toggle the table of contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Johannes Fibiger</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 55 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-55" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">55 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D9%8A%D9%88%D9%87%D8%A7%D9%86%D8%B3_%D9%81%D9%8A%D8%A8%D9%8A%D8%BA%D8%B1" title="يوهانس فيبيغر – Arabic" lang="ar" hreflang="ar" data-title="يوهانس فيبيغر" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%DA%98%D9%88%D9%87%D8%A7%D9%86%D8%B3_%D9%81%DB%8C%D8%A8%DB%8C%DA%AF%D8%B1" title="ژوهانس فیبیگر – South Azerbaijani" lang="azb" hreflang="azb" data-title="ژوهانس فیبیگر" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%87%E0%A6%AF%E0%A6%BC%E0%A7%8B%E0%A6%B9%E0%A7%8D%E0%A6%AF%E0%A6%BE%E0%A6%A8%E0%A7%87%E0%A6%B8_%E0%A6%AB%E0%A6%BF%E0%A6%AC%E0%A6%BF%E0%A6%97%E0%A6%BE%E0%A6%B0" title="ইয়োহ্যানেস ফিবিগার – Bangla" lang="bn" hreflang="bn" data-title="ইয়োহ্যানেস ফিবিগার" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%86%D0%B0%D1%85%D0%B0%D0%BD%D0%B5%D1%81_%D0%90%D0%BD%D0%B4%D1%80%D1%8D%D0%B0%D1%81_%D0%93%D1%80%D1%8B%D0%B1_%D0%A4%D1%96%D0%B1%D1%96%D0%B3%D0%B5%D1%80" title="Іаханес Андрэас Грыб Фібігер – Belarusian" lang="be" hreflang="be" data-title="Іаханес Андрэас Грыб Фібігер" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%99%D0%BE%D1%85%D0%B0%D0%BD%D0%B5%D1%81_%D0%A4%D0%B8%D0%B1%D0%B8%D0%B3%D0%B5%D1%80" title="Йоханес Фибигер – Bulgarian" lang="bg" hreflang="bg" data-title="Йоханес Фибигер" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger – Catalan" lang="ca" hreflang="ca" data-title="Johannes Andreas Grib Fibiger" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Czech" lang="cs" hreflang="cs" data-title="Johannes Fibiger" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-cy mw-list-item"><a href="https://cy.wikipedia.org/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger – Welsh" lang="cy" hreflang="cy" data-title="Johannes Andreas Grib Fibiger" data-language-autonym="Cymraeg" data-language-local-name="Welsh" class="interlanguage-link-target"><span>Cymraeg</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Danish" lang="da" hreflang="da" data-title="Johannes Fibiger" data-language-autonym="Dansk" data-language-local-name="Danish" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – German" lang="de" hreflang="de" data-title="Johannes Fibiger" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Estonian" lang="et" hreflang="et" data-title="Johannes Fibiger" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Spanish" lang="es" hreflang="es" data-title="Johannes Fibiger" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Basque" lang="eu" hreflang="eu" data-title="Johannes Fibiger" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%DA%98%D9%88%D9%87%D8%A7%D9%86%D8%B3_%D9%81%DB%8C%D8%A8%DB%8C%DA%AF%D8%B1" title="ژوهانس فیبیگر – Persian" lang="fa" hreflang="fa" data-title="ژوهانس فیبیگر" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – French" lang="fr" hreflang="fr" data-title="Johannes Fibiger" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Galician" lang="gl" hreflang="gl" data-title="Johannes Fibiger" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%9A%94%ED%95%98%EB%84%A4%EC%8A%A4_%ED%94%BC%EB%B9%84%EA%B2%8C%EB%A5%B4" title="요하네스 피비게르 – Korean" lang="ko" hreflang="ko" data-title="요하네스 피비게르" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger – Croatian" lang="hr" hreflang="hr" data-title="Johannes Andreas Grib Fibiger" data-language-autonym="Hrvatski" data-language-local-name="Croatian" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger – Ido" lang="io" hreflang="io" data-title="Johannes Andreas Grib Fibiger" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger – Indonesian" lang="id" hreflang="id" data-title="Johannes Andreas Grib Fibiger" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-is mw-list-item"><a href="https://is.wikipedia.org/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger – Icelandic" lang="is" hreflang="is" data-title="Johannes Andreas Grib Fibiger" data-language-autonym="Íslenska" data-language-local-name="Icelandic" class="interlanguage-link-target"><span>Íslenska</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger – Italian" lang="it" hreflang="it" data-title="Johannes Andreas Grib Fibiger" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%99%D7%95%D7%94%D7%A0%D7%A1_%D7%90%D7%A0%D7%93%D7%A8%D7%90%D7%A1_%D7%92%D7%A8%D7%99%D7%91_%D7%A4%D7%99%D7%91%D7%99%D7%92%D7%A8" title="יוהנס אנדראס גריב פיביגר – Hebrew" lang="he" hreflang="he" data-title="יוהנס אנדראס גריב פיביגר" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-ka mw-list-item"><a href="https://ka.wikipedia.org/wiki/%E1%83%98%E1%83%9D%E1%83%B0%E1%83%90%E1%83%9C%E1%83%94%E1%83%A1_%E1%83%A4%E1%83%98%E1%83%91%E1%83%98%E1%83%92%E1%83%94%E1%83%A0%E1%83%98" title="იოჰანეს ფიბიგერი – Georgian" lang="ka" hreflang="ka" data-title="იოჰანეს ფიბიგერი" data-language-autonym="ქართული" data-language-local-name="Georgian" class="interlanguage-link-target"><span>ქართული</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%99%D0%BE%D1%85%D0%B0%D0%BD%D0%BD%D0%B5%D1%81_%D0%90%D0%BD%D0%B4%D1%80%D0%B5%D0%B0%D1%81_%D0%93%D1%80%D0%B8%D0%B1_%D0%A4%D0%B8%D0%B1%D0%B8%D0%B3%D0%B5%D1%80" title="Йоханнес Андреас Гриб Фибигер – Kazakh" lang="kk" hreflang="kk" data-title="Йоханнес Андреас Гриб Фибигер" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Swahili" lang="sw" hreflang="sw" data-title="Johannes Fibiger" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Iohannes_Fibiger" title="Iohannes Fibiger – Latin" lang="la" hreflang="la" data-title="Iohannes Fibiger" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Hungarian" lang="hu" hreflang="hu" data-title="Johannes Fibiger" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-xmf mw-list-item"><a href="https://xmf.wikipedia.org/wiki/%E1%83%98%E1%83%9D%E1%83%B0%E1%83%90%E1%83%9C%E1%83%94%E1%83%A1_%E1%83%A4%E1%83%98%E1%83%91%E1%83%98%E1%83%92%E1%83%94%E1%83%A0%E1%83%98" title="იოჰანეს ფიბიგერი – Mingrelian" lang="xmf" hreflang="xmf" data-title="იოჰანეს ფიბიგერი" data-language-autonym="მარგალური" data-language-local-name="Mingrelian" class="interlanguage-link-target"><span>მარგალური</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%AC%D9%88%D9%87%D8%A7%D9%86%D8%B3_%D9%81%D9%8A%D8%A8%D9%8A%D8%AC%D8%B1" title="جوهانس فيبيجر – Egyptian Arabic" lang="arz" hreflang="arz" data-title="جوهانس فيبيجر" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%DA%98%D9%88%D9%87%D8%A7%D9%86%D8%B3_%D9%81%DB%8C%D8%A8%DB%8C%DA%AF%D8%B1" title="ژوهانس فیبیگر – Mazanderani" lang="mzn" hreflang="mzn" data-title="ژوهانس فیبیگر" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Malay" lang="ms" hreflang="ms" data-title="Johannes Fibiger" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger – Dutch" lang="nl" hreflang="nl" data-title="Johannes Andreas Grib Fibiger" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%83%A8%E3%83%8F%E3%83%8D%E3%82%B9%E3%83%BB%E3%83%95%E3%82%A3%E3%83%93%E3%82%B2%E3%83%AB" title="ヨハネス・フィビゲル – Japanese" lang="ja" hreflang="ja" data-title="ヨハネス・フィビゲル" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Johannes Fibiger" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Occitan" lang="oc" hreflang="oc" data-title="Johannes Fibiger" data-language-autonym="Occitan" data-language-local-name="Occitan" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Uzbek" lang="uz" hreflang="uz" data-title="Johannes Fibiger" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Uzbek" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D8%AC%D9%88%DB%81%D8%A7%D9%86%D8%B2_%D8%A7%DB%8C%D9%86%DA%88%D8%B1%DB%8C%D8%A7%D8%B3_%DA%AF%D8%B1%D8%A8_%D9%81%D8%A8%DA%AF%D8%B1" title="جوہانز اینڈریاس گرب فبگر – Western Punjabi" lang="pnb" hreflang="pnb" data-title="جوہانز اینڈریاس گرب فبگر" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Polish" lang="pl" hreflang="pl" data-title="Johannes Fibiger" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Portuguese" lang="pt" hreflang="pt" data-title="Johannes Fibiger" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Romanian" lang="ro" hreflang="ro" data-title="Johannes Fibiger" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-qu mw-list-item"><a href="https://qu.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Quechua" lang="qu" hreflang="qu" data-title="Johannes Fibiger" data-language-autonym="Runa Simi" data-language-local-name="Quechua" class="interlanguage-link-target"><span>Runa Simi</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%A4%D0%B8%D0%B1%D0%B8%D0%B3%D0%B5%D1%80,_%D0%99%D0%BE%D1%85%D0%B0%D0%BD%D0%BD%D0%B5%D1%81_%D0%90%D0%BD%D0%B4%D1%80%D0%B5%D0%B0%D1%81_%D0%93%D1%80%D0%B8%D0%B1" title="Фибигер, Йоханнес Андреас Гриб – Russian" lang="ru" hreflang="ru" data-title="Фибигер, Йоханнес Андреас Гриб" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Simple English" lang="en-simple" hreflang="en-simple" data-title="Johannes Fibiger" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger – Slovenian" lang="sl" hreflang="sl" data-title="Johannes Andreas Grib Fibiger" data-language-autonym="Slovenščina" data-language-local-name="Slovenian" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-ckb mw-list-item"><a href="https://ckb.wikipedia.org/wiki/%D8%AC%DB%86%DA%BE%D8%A7%D9%86%D8%B3_%D9%81%DB%8C%D8%A8%DA%AF%DB%95%D8%B1" title="جۆھانس فیبگەر – Central Kurdish" lang="ckb" hreflang="ckb" data-title="جۆھانس فیبگەر" data-language-autonym="کوردی" data-language-local-name="Central Kurdish" class="interlanguage-link-target"><span>کوردی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger – Finnish" lang="fi" hreflang="fi" data-title="Johannes Andreas Grib Fibiger" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Swedish" lang="sv" hreflang="sv" data-title="Johannes Fibiger" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://tt.wikipedia.org/wiki/%D0%99%D0%BE%D2%BB%D0%B0%D0%BD%D0%BD%D0%B5%D1%81_%D0%A4%D1%80%D0%B8%D0%B1%D0%B8%D0%B3%D0%B5%D1%80" title="Йоһаннес Фрибигер – Tatar" lang="tt" hreflang="tt" data-title="Йоһаннес Фрибигер" data-language-autonym="Татарча / tatarça" data-language-local-name="Tatar" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger – Turkish" lang="tr" hreflang="tr" data-title="Johannes Andreas Grib Fibiger" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%99%D0%BE%D0%B3%D0%B0%D0%BD%D0%B5%D1%81_%D0%A4%D1%96%D0%B1%D1%96%D0%B3%D0%B5%D1%80" title="Йоганес Фібігер – Ukrainian" lang="uk" hreflang="uk" data-title="Йоганес Фібігер" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D8%AC%D9%88%DB%81%D8%A7%D9%86%D8%B3_%D9%81%D8%A7%D8%A6%D8%A8%DA%AF%D8%B1" title="جوہانس فائبگر – Urdu" lang="ur" hreflang="ur" data-title="جوہانس فائبگر" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Vietnamese" lang="vi" hreflang="vi" data-title="Johannes Fibiger" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/Johannes_Fibiger" title="Johannes Fibiger – Yoruba" lang="yo" hreflang="yo" data-title="Johannes Fibiger" data-language-autonym="Yorùbá" data-language-local-name="Yoruba" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E7%B4%84%E7%BF%B0%E5%B0%BC%E6%96%AF%C2%B7%E8%8F%B2%E6%AF%94%E6%A0%BC" title="約翰尼斯·菲比格 – Chinese" lang="zh" hreflang="zh" data-title="約翰尼斯·菲比格" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div 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</div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Danish physician (1867–1928)</div> <p class="mw-empty-elt"> </p> <style data-mw-deduplicate="TemplateStyles:r1257001546">.mw-parser-output .infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox-full-data:not(.notheme)>div:not(.notheme)[style]{background:#1f1f23!important;color:#f8f9fa}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox-full-data:not(.notheme) div:not(.notheme){background:#1f1f23!important;color:#f8f9fa}}@media(min-width:640px){body.skin--responsive .mw-parser-output .infobox-table{display:table!important}body.skin--responsive .mw-parser-output .infobox-table>caption{display:table-caption!important}body.skin--responsive .mw-parser-output .infobox-table>tbody{display:table-row-group}body.skin--responsive .mw-parser-output .infobox-table tr{display:table-row!important}body.skin--responsive .mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox biography vcard"><tbody><tr><th colspan="2" class="infobox-above" style="font-size:125%;"><div class="fn">Johannes Fibiger</div></th></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:J_Fibiger.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/43/J_Fibiger.jpg/220px-J_Fibiger.jpg" decoding="async" width="220" height="325" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/43/J_Fibiger.jpg/330px-J_Fibiger.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/43/J_Fibiger.jpg/440px-J_Fibiger.jpg 2x" data-file-width="1040" data-file-height="1536" /></a></span></td></tr><tr><th scope="row" class="infobox-label">Born</th><td class="infobox-data"><div style="display:inline" class="nickname">Johannes Andreas Grib Fibiger</div><br /><span style="display:none">(<span class="bday">1867-04-23</span>)</span>23 April 1867<br /><div style="display:inline" class="birthplace"><a href="/wiki/Silkeborg" title="Silkeborg">Silkeborg</a>, Denmark</div></td></tr><tr><th scope="row" class="infobox-label">Died</th><td class="infobox-data">30 January 1928<span style="display:none">(1928-01-30)</span> (aged&#160;60)<br /><div style="display:inline" class="deathplace"><a href="/wiki/Copenhagen" title="Copenhagen">Copenhagen</a>, Denmark</div></td></tr><tr><th scope="row" class="infobox-label">Alma&#160;mater</th><td class="infobox-data">University of Copenhagen</td></tr><tr><th scope="row" class="infobox-label">Known&#160;for</th><td class="infobox-data">Induction of cancer using <i><a href="/wiki/Spiroptera_carcinoma" class="mw-redirect" title="Spiroptera carcinoma">Spiroptera carcinoma</a></i></td></tr><tr><th scope="row" class="infobox-label">Spouse</th><td class="infobox-data"><style data-mw-deduplicate="TemplateStyles:r1151524712">.mw-parser-output .marriage-line-margin2px{line-height:0;margin-bottom:-2px}.mw-parser-output .marriage-line-margin3px{line-height:0;margin-bottom:-3px}.mw-parser-output .marriage-display-ws{display:inline;white-space:nowrap}</style> <div class="marriage-display-ws"><div style="display:inline-block;line-height:normal;">Mathilde Fibiger</div>&#32;<div style="display:inline-block;">&#8203;</div>&#40;<abbr title="married">m.</abbr>&#160;1894&#41;<wbr />&#8203;</div></td></tr><tr><th scope="row" class="infobox-label">Children</th><td class="infobox-data">2</td></tr><tr><th scope="row" class="infobox-label">Awards</th><td class="infobox-data">1926 <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a></td></tr><tr><td colspan="2" class="infobox-full-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1257001546"><b>Scientific career</b></td></tr><tr><th scope="row" class="infobox-label">Fields</th><td class="infobox-data category"><a href="/wiki/Medicine" title="Medicine">Medicine</a><br /><a href="/wiki/Parasitology" title="Parasitology">Parasitology</a></td></tr><tr><th scope="row" class="infobox-label">Institutions</th><td class="infobox-data"><a href="/wiki/University_of_Copenhagen" title="University of Copenhagen">University of Copenhagen</a><br /><a href="/wiki/Royal_Danish_Army" title="Royal Danish Army">Royal Danish Army Medical Corps</a></td></tr><tr><th scope="row" class="infobox-label"><a href="/wiki/Author_citation_(zoology)" title="Author citation (zoology)">Author&#160;abbrev. (zoology)</a></th><td class="infobox-data">Fibiger</td></tr><tr style="display:none"><td colspan="2"> </td></tr></tbody></table> <p><b>Johannes Andreas Grib Fibiger</b> (23 April 1867 – 30 January 1928) was a Danish physician and professor of <a href="/wiki/Anatomical_pathology" title="Anatomical pathology">anatomical pathology</a> at the <a href="/wiki/University_of_Copenhagen" title="University of Copenhagen">University of Copenhagen</a>. He was the recipient of the 1926 <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a> "for his discovery of the <i>Spiroptera carcinoma</i>". He claimed to have shown that the <a href="/wiki/Nematode" title="Nematode">roundworm</a> which he called <i>Spiroptera carcinoma</i> (but is correctly named <i><a href="/wiki/Gongylonema_neoplasticum" title="Gongylonema neoplasticum">Gongylonema neoplasticum</a></i>) could cause stomach cancer (<a href="/wiki/Squamous_cell_carcinoma" class="mw-redirect" title="Squamous cell carcinoma">squamous cell carcinoma</a>) in rats and mice. His experimental results were later proven to be a case of mistaken conclusion. </p><p>While working at the Institute of Pathological Anatomy of University of Copenhagen, Fibiger discovered new roundworms in 1907 from wild rats. He suspected that the roundworms were responsible for stomach cancer in those rats. In 1913, he reported that he could experimentally induce cancer in healthy rats using the roundworms. His discovery was considered "the greatest contribution to experimental medicine" at the time.<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> In 1926, he was nominated for the Nobel Prize in Physiology or Medicine along with <a href="/wiki/Katsusaburo_Yamagiwa" class="mw-redirect" title="Katsusaburo Yamagiwa">Katsusaburo Yamagiwa</a>, who had experimentally induced carcinoma by painting crude coal tar on the inner surface of rabbits' ears in 1915. However, they were considered undeserving, and the 1926 prize was not given. In the next year Fibiger alone was retrospectively chosen for the 1926 Nobel Prize. </p><p>After his death, independent researches proved that <i>G. neoplasticum</i> cannot cause cancer. Tumours and cancer produced by Fibiger were due to <a href="/wiki/Vitamin_A_deficiency" title="Vitamin A deficiency">vitamin A deficiency</a>. Historical reassessment of Fibiger's data revealed that he had mistaken non-cancerous tumours for cancerous tumours. Erling Norrby, who had served as the Permanent Secretary of the <a href="/wiki/Royal_Swedish_Academy_of_Sciences" title="Royal Swedish Academy of Sciences">Royal Swedish Academy of Sciences</a> and Professor and Chairman of <a href="/wiki/Virology" title="Virology">Virology</a> at the <a href="/wiki/Karolinska_Institute" title="Karolinska Institute">Karolinska Institute</a>, in 2010 declared Fibiger's Nobel Prize as "one of the biggest blunders made by the Karolinska Institute."<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> </p><p>Fibiger's research method on diphtheria is regarded as the origin of an important research methodology in medicine known as controlled <a href="/wiki/Clinical_trial" title="Clinical trial">clinical trial</a>.<sup id="cite_ref-gluud_3-0" class="reference"><a href="#cite_note-gluud-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Biography">Biography</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Johannes_Fibiger&amp;action=edit&amp;section=1" title="Edit section: Biography"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Fibiger was born in Silkeborg, Midtjylland, Denmark. He was the second son of Christian Ludvig Wilhelm Fibiger and <a href="/wiki/Elfride_Fibiger" title="Elfride Fibiger">Elfride Fibiger</a> (<i>née</i> Müller). His father was a local physician and his mother, an author. He was named after his uncle, who was a clergyman and a poet.<sup id="cite_ref-encyclo_4-0" class="reference"><a href="#cite_note-encyclo-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> His father died of internal bleeding when he was three years of age, after which the family moved to Copenhagen where his mother earned their living by writing. His mother established there the first cooking school, the Copenhagen Cooking School. </p><p>His namesake uncle supported Fibiger for his education. In 1883, at age 16, he passed his matriculation and was enrolled at the University of Copenhagen to study zoology and botany. He self-supported by teaching and working at the laboratories. He graduated in 1883. He continued for a medicine course and earned his medical degree in 1890. For a few months he worked as a physician at different hospitals, and also continued to study under <a href="/wiki/Robert_Koch" title="Robert Koch">Robert Koch</a> and <a href="/wiki/Emil_Adolf_von_Behring" class="mw-redirect" title="Emil Adolf von Behring">Emil Adolf von Behring</a> in Berlin. Between 1891 and 1894, he was assistant to C. J. Salomonsen at the Department of Bacteriology of Copenhagen University. From 1894 he joined the <a href="/wiki/Royal_Danish_Army" title="Royal Danish Army">Royal Danish Army Medical Corps</a>, where he served till 1897. It was during his army service that he completed his doctoral thesis <i>Research into the bacteriology of diphtheria</i>. The University of Copenhagen awarded him a doctorate degree in 1895. He continued the diphtheria research at Blegdamshospitalet in Copenhagen, working as a Junior Physician.<sup id="cite_ref-encyclo_4-1" class="reference"><a href="#cite_note-encyclo-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> In 1897, he was appointed <a href="/wiki/Prosector" title="Prosector">prosector</a> at the Institute of Pathological Anatomy of University of Copenhagen. He was promoted to full Professor and eventually its Director in 1900. He also served as Principal of the Laboratory of Clinical Bacteriology of the Army from 1890 to 1905), and Director of the Central Laboratory of the Army and Consultant Physician to the Army Medical Service in 1905.<sup id="cite_ref-nobel_5-0" class="reference"><a href="#cite_note-nobel-5"><span class="cite-bracket">&#91;</span>5<span class="cite-bracket">&#93;</span></a></sup> </p><p>Fibiger died of <a href="/wiki/Cardiac_arrest" title="Cardiac arrest">cardiac arrest</a> in Copenhagen on 30 January 1928.<sup id="cite_ref-nobel_5-1" class="reference"><a href="#cite_note-nobel-5"><span class="cite-bracket">&#91;</span>5<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Contributions">Contributions</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Johannes_Fibiger&amp;action=edit&amp;section=2" title="Edit section: Contributions"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Diphtheria_research">Diphtheria research</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Johannes_Fibiger&amp;action=edit&amp;section=3" title="Edit section: Diphtheria research"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Fibiger's doctoral research was on diphtheria. He developed more efficient method of growing bacteria in a laboratory setting. He discovered that there were two different forms (strains) of the diphtheria bacillus (<i><a href="/wiki/Corynebacterium_diphtheriae" title="Corynebacterium diphtheriae">Corynebacterium diphtheriae</a></i>) that produce two different symptoms, now called nasopharyngeal and cutaneous diphtheria. He also produced a blood serum against the disease.<sup id="cite_ref-encyclo_4-2" class="reference"><a href="#cite_note-encyclo-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> He was known for his methodological system of research. One of his experiments from 1898 in which he tested the blood serum for diphtheria is regarded by some as the first controlled <a href="/wiki/Clinical_trial" title="Clinical trial">clinical trial</a>.<sup id="cite_ref-gluud_3-1" class="reference"><a href="#cite_note-gluud-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> While working as a Junior Physician in Blegdamshospitalet, he tested his diphtheria serum among hundreds (484) patients. As would be in modern clinical trial, he separated serum-treated and untreated patients, and found that more of untreated patients died than the treated ones. According to an article in <i><a href="/wiki/The_British_Medical_Journal" class="mw-redirect" title="The British Medical Journal">The British Medical Journal</a></i> in 1998: </p> <style data-mw-deduplicate="TemplateStyles:r1244412712">.mw-parser-output .templatequote{overflow:hidden;margin:1em 0;padding:0 32px}.mw-parser-output .templatequotecite{line-height:1.5em;text-align:left;margin-top:0}@media(min-width:500px){.mw-parser-output .templatequotecite{padding-left:1.6em}}</style><blockquote class="templatequote"><p>[Fibiger's experiment in 1898] was the first clinical trial in which random allocation was used and emphasised as a pivotal methodological principle. This pioneering improvement in methodology, combined with a large number of patients and rigorous planning, conduct, and reporting, makes the trial a milestone in the history of clinical trials.<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup></p></blockquote> <div class="mw-heading mw-heading3"><h3 id="Cancer_and_parasitology_research">Cancer and parasitology research</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Johannes_Fibiger&amp;action=edit&amp;section=4" title="Edit section: Cancer and parasitology research"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>While studying <a href="/wiki/Tuberculosis" title="Tuberculosis">tuberculosis</a> in lab rats, Fibiger found <a href="/wiki/Tumor" class="mw-redirect" title="Tumor">tumors</a> in some wild rats collected from Dorpat (officially <a href="/wiki/Tartu" title="Tartu">Tartu</a>, now in Estonia) in 1907. Rats having stomach tumour (<a href="/wiki/Papilloma" title="Papilloma">papilloma</a>) also had nematodes and their eggs. He found that some tumours were <a href="/wiki/Metastatic" class="mw-redirect" title="Metastatic">metastatic</a> (cancerous). He hypothesised that the nematodes caused the stomach cancer. After years of investigation, he experimentally demonstrated in 1913 that the nematode could induce stomach cancer. He published his discovery in a series of three papers, and also presented them at the Académie Royale des Sciences et des Lettres de Danemark (<a href="/wiki/Royal_Danish_Academy_of_Sciences_and_Letters" title="Royal Danish Academy of Sciences and Letters">Royal Danish Academy of Sciences and Letters</a>), and Troisième Conférence Internationale pour l’Étude du Cancer (Third International Conference for Researches in Cancer) at Brussels the same year.<sup id="cite_ref-fibiger1919_7-0" class="reference"><a href="#cite_note-fibiger1919-7"><span class="cite-bracket">&#91;</span>7<span class="cite-bracket">&#93;</span></a></sup> He knew that the nematode was a new species, and provisionally named it <i>Spiroptera carcinom </i> in 1914. With Hjalmar Ditlevsen, of the Zoological Museum of the University of Copenhagen, he described it as <i>Spiroptera</i> (<i>Gongylonema</i>) <i>neoplastica</i> in 1914.<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">&#91;</span>8<span class="cite-bracket">&#93;</span></a></sup> Ditlevsen revised the description in 1918, and gave the final valid name <i>Gongylonema neoplasticum</i>.<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">&#91;</span>9<span class="cite-bracket">&#93;</span></a></sup> But Fibiger never used the formal scientific name, and persistently used <i>Spiroptera carcinoma</i>.<sup id="cite_ref-Lalchhandama_10-0" class="reference"><a href="#cite_note-Lalchhandama-10"><span class="cite-bracket">&#91;</span>10<span class="cite-bracket">&#93;</span></a></sup> </p><p>Fibiger's experiment was the first to show that helminth parasites can cause cancer, and that cancer (tumour) can be experimentally induced. His discovery was supported by the experiment of two Japanese scientists <a href="/wiki/Katsusaburo_Yamagiwa" class="mw-redirect" title="Katsusaburo Yamagiwa">Katsusaburo Yamagiwa</a> and Koichi Ichikawa in 1918. Yamagiwa and Ichikawa demonstrated the induction of cancer (<a href="/wiki/Carcinoma" title="Carcinoma">carcinoma</a>) in rabbits. They showed that it was relatively easy to produce cancer was by painting coal-tar on the inner surface of the ear.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">&#91;</span>11<span class="cite-bracket">&#93;</span></a></sup> A number independent experiments subsequently confirmed the cancer-inducing effect of coal-tar in mice.<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">&#91;</span>12<span class="cite-bracket">&#93;</span></a></sup> With such supporting evidence, Fibiger's work was regarded a milestone in cancer study.<sup id="cite_ref-Lalchhandama_10-1" class="reference"><a href="#cite_note-Lalchhandama-10"><span class="cite-bracket">&#91;</span>10<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">&#91;</span>13<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Personal_life">Personal life</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Johannes_Fibiger&amp;action=edit&amp;section=5" title="Edit section: Personal life"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Fibiger was married to Mathilde Fibiger (1863–1954). Mathilde was his cousin who came to help her mother's business while he was studying medicine. They got married on 4 August 1894. Fibiger had been suffering from colon cancer, and a month after he received his Nobel Prize, he died of heart attack on 30 January 1928 due to his worsening cancer. He was survived by his wife and two children.<sup id="cite_ref-nobel_5-2" class="reference"><a href="#cite_note-nobel-5"><span class="cite-bracket">&#91;</span>5<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">&#91;</span>14<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Nobel_Prize_controversy">Nobel Prize controversy</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Johannes_Fibiger&amp;action=edit&amp;section=6" title="Edit section: Nobel Prize controversy"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">See also: <a href="/wiki/Nobel_Prize_controversies" title="Nobel Prize controversies">Nobel Prize controversies</a></div> <div class="mw-heading mw-heading3"><h3 id="Nomination_and_selection">Nomination and selection</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Johannes_Fibiger&amp;action=edit&amp;section=7" title="Edit section: Nomination and selection"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Fibiger was nominated for the Nobel Prize in Physiology or Medicine 18 times from 1920. In 1926, he received two nominations along with Katsusaburo Yamagiwa.<sup id="cite_ref-nobelnom_15-0" class="reference"><a href="#cite_note-nobelnom-15"><span class="cite-bracket">&#91;</span>15<span class="cite-bracket">&#93;</span></a></sup> Folke Henschen and Hilding Bergstrand were appointed by the Nobel Committee as assessors. They disagreed on each other's conclusion. Henschen was in favour of the nomination and concluded that "the experimental carcinoma is worthy of the Nobel Prize. It should therefore be just if the prize would be divided between Johannes Fibiger, the discoverer of the experimental spiroptera carcinoma, and Katsusaburo Yamagiwa, the discoverer of the experimental tar carcinoma." But Bergstrand opposed to it, concluding that "an experimental confirmation of a previously known fact... [referring to the prevalence of cancer among chimney sweeps and factory workers, an already established medical fact at the time<sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">&#91;</span>16<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">&#91;</span>17<span class="cite-bracket">&#93;</span></a></sup>] can, in this case, not be considered... that one cannot, at this point, find much support for the possibility that the work of Fibiger and Yamagiwa will have great importance in the solving of the riddle of cancer. Under such circumstances I do not consider these discoveries worthy of the Nobel Prize." The Nobel Committee could not make the final decision and decided not to give the prize for 1926. In the next year, Fibiger received seven nominations, and Yamagiwa was then excluded. Instead there were two other nominees, <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Otto Heinrich Warburg</a> and <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Julius Wagner-Jauregg</a>. In addition to Henschen and Bergstrand, Einar Hammersten was appointed as the third assessor. On Fibiger, Bergstrand was of the same opinion, Hammersten was in favour. Hence the three nominees were recommended for the award. The Nobel Committee decided to give the 1926 prize jointly to Fibiger and Warburg, and the 1927 prize to Wagner-Jauregg. However, the authority at the Karolinska Institute disagreed with the recommendation for Warburg for undisclosed reasons, and Fibiger became the sole winner.<sup id="cite_ref-stolt04_18-0" class="reference"><a href="#cite_note-stolt04-18"><span class="cite-bracket">&#91;</span>18<span class="cite-bracket">&#93;</span></a></sup> (Warburg was eventually awarded the prize in 1931.<sup id="cite_ref-Lalchhandama_10-2" class="reference"><a href="#cite_note-Lalchhandama-10"><span class="cite-bracket">&#91;</span>10<span class="cite-bracket">&#93;</span></a></sup>) </p> <div class="mw-heading mw-heading3"><h3 id="Posthumous_criticism_and_disproof">Posthumous criticism and disproof</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Johannes_Fibiger&amp;action=edit&amp;section=8" title="Edit section: Posthumous criticism and disproof"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Fibiger's discovery was unchallenged during his lifetime. The only serious criticism was by F. D. Bullock and G. L. Rohdenburg. In 1918, they argued that the cancerous tumours produced in Fibiger's experiments were similar to non-cancerous tumours. Such tumours easily developed in epithelial cells.<sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">&#91;</span>19<span class="cite-bracket">&#93;</span></a></sup> Fibiger defended "That these tumors are true carcinomata cannot, thus, be doubted, and the fact that they may occur in younger animals does not diminish our right to range them among the true malignant neoplasms."<sup id="cite_ref-fibiger1919_7-1" class="reference"><a href="#cite_note-fibiger1919-7"><span class="cite-bracket">&#91;</span>7<span class="cite-bracket">&#93;</span></a></sup> </p><p>It was only after Fibiger's death that the importance of vitamin A in cancer development was fully appreciated. Fibiger had used rats which were fed vitamin A-less diet. It was found that Vitamin A deficiency alone can cause tumours and cancers. The parasites had merely caused the tissue irritation (chronic inflammation) that drove the damaged cells into cancer; any tissue irritation could have induced the tumours.<sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">&#91;</span>20<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">&#91;</span>21<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">&#91;</span>22<span class="cite-bracket">&#93;</span></a></sup> The major challenge came from <a href="/w/index.php?title=Richard_Douglas_Passey&amp;action=edit&amp;redlink=1" class="new" title="Richard Douglas Passey (page does not exist)">Richard Douglas Passey</a>, with his colleagues A. Léese, and J.C. Knox. They reported a new finding in 1935 that <i>S. carcinoma</i> do not cause cancer in rats. They concluded that Fibiger probably had mistaken metaplasia (a non-cancerous tumour) with malignant neoplasia (true cancer).<sup id="cite_ref-23" class="reference"><a href="#cite_note-23"><span class="cite-bracket">&#91;</span>23<span class="cite-bracket">&#93;</span></a></sup> In 1937, W. Cramer experimentally showed that Fibiger's tumour were not cancerous.<sup id="cite_ref-24" class="reference"><a href="#cite_note-24"><span class="cite-bracket">&#91;</span>24<span class="cite-bracket">&#93;</span></a></sup> The final disproof was shown by Claude R. Hitchcock and E. T. Bell. In 1952, they repeated Fibiger's experiments using advanced microscopy and histology, and conclusively demonstrated that the tumours due to <i>G. neoplasticum</i> in rats were non-cancerous, and that the tumours were primarily due to vitamin A deficiency.<sup id="cite_ref-25" class="reference"><a href="#cite_note-25"><span class="cite-bracket">&#91;</span>25<span class="cite-bracket">&#93;</span></a></sup> </p><p>It is also claimed that exclusion of Yamagiwa was another mistake. Yamagiwa's work has become the primary basis for induction of tumours in cancer research.<sup id="cite_ref-26" class="reference"><a href="#cite_note-26"><span class="cite-bracket">&#91;</span>26<span class="cite-bracket">&#93;</span></a></sup> Because of this, some consider Fibiger's Nobel Prize to be undeserved particularly because Yamagiwa did not receive the prize.<sup id="cite_ref-27" class="reference"><a href="#cite_note-27"><span class="cite-bracket">&#91;</span>27<span class="cite-bracket">&#93;</span></a></sup> <i><a href="/wiki/Encyclop%C3%A6dia_Britannica" title="Encyclopædia Britannica">Encyclopædia Britannica</a></i><span class="nowrap" style="padding-left:0.1em;">&#39;s</span> guide to Nobel Prizes in cancer research mentions Yamagiwa's work as a milestone without mentioning Fibiger.<sup id="cite_ref-28" class="reference"><a href="#cite_note-28"><span class="cite-bracket">&#91;</span>28<span class="cite-bracket">&#93;</span></a></sup> </p><p>Although <i>G. neoplasticum</i> is non-carcinogenic, <a href="/wiki/Carcinogenic_parasite" title="Carcinogenic parasite">other helminth parasites</a> such as <i><a href="/wiki/Schistosoma_haematobium" title="Schistosoma haematobium">Schistosoma haematobium</a></i>, <i><a href="/wiki/Opisthorchis_viverrini" title="Opisthorchis viverrini">Opisthorchis viverrini</a></i> and <i><a href="/wiki/Clonorchis_sinensis" title="Clonorchis sinensis">Clonorchis sinensis</a></i> are now established to cause cancer in humans.<sup id="cite_ref-29" class="reference"><a href="#cite_note-29"><span class="cite-bracket">&#91;</span>29<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-30" class="reference"><a href="#cite_note-30"><span class="cite-bracket">&#91;</span>30<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Johannes_Fibiger&amp;action=edit&amp;section=9" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="/wiki/Carcinogenic_parasite" title="Carcinogenic parasite">Carcinogenic parasite</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Johannes_Fibiger&amp;action=edit&amp;section=10" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width" style="column-width: 30em;"> <ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFWernstedt1927" class="citation web cs1">Wernstedt, W. (1927). <a rel="nofollow" class="external text" href="https://www.nobelprize.org/nobel_prizes/medicine/laureates/1926/press.html">"Award Ceremony Speech"</a>. <i>www.nobelprize.org</i>. Nobel Media AB<span class="reference-accessdate">. Retrieved <span class="nowrap">1 May</span> 2017</span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=unknown&amp;rft.jtitle=www.nobelprize.org&amp;rft.atitle=Award+Ceremony+Speech&amp;rft.date=1927&amp;rft.aulast=Wernstedt&amp;rft.aufirst=W.&amp;rft_id=https%3A%2F%2Fwww.nobelprize.org%2Fnobel_prizes%2Fmedicine%2Flaureates%2F1926%2Fpress.html&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AJohannes+Fibiger" class="Z3988"></span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-2">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFNorrby2010" class="citation book cs1">Norrby, Erling (2010). <i>Nobel Prizes and Life Sciences</i>. 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"Of Fibiger and fables: a cautionary tale of cockroaches and <i>Helicobacter pylori</i>". <i>J Clin Gastroenterol</i>. <b>33</b> (3): 177–179. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1097%2F00004836-200109000-00001">10.1097/00004836-200109000-00001</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/11500602">11500602</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=J+Clin+Gastroenterol&amp;rft.atitle=Of+Fibiger+and+fables%3A+a+cautionary+tale+of+cockroaches+and+Helicobacter+pylori&amp;rft.volume=33&amp;rft.issue=3&amp;rft.pages=177-179&amp;rft.date=2001&amp;rft_id=info%3Adoi%2F10.1097%2F00004836-200109000-00001&amp;rft_id=info%3Apmid%2F11500602&amp;rft.aulast=Modlin&amp;rft.aufirst=IM&amp;rft.au=Kidd%2C+M&amp;rft.au=Hinoue%2C+T&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AJohannes+Fibiger" class="Z3988"></span></span> </li> <li id="cite_note-23"><span class="mw-cite-backlink"><b><a href="#cite_ref-23">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPasseyLeeseKnox1935" class="citation journal cs1">Passey, R. D.; Leese, A.; Knox, J. C. (1935). "Spiroptera cancer and diet deficiency". <i>Journal of Pathology and Bacteriology</i>. <b>40</b> (1): 198–199.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Journal+of+Pathology+and+Bacteriology&amp;rft.atitle=Spiroptera+cancer+and+diet+deficiency&amp;rft.volume=40&amp;rft.issue=1&amp;rft.pages=198-199&amp;rft.date=1935&amp;rft.aulast=Passey&amp;rft.aufirst=R.+D.&amp;rft.au=Leese%2C+A.&amp;rft.au=Knox%2C+J.+C.&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AJohannes+Fibiger" class="Z3988"></span></span> </li> <li id="cite_note-24"><span class="mw-cite-backlink"><b><a href="#cite_ref-24">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCramer1937" class="citation journal cs1">Cramer, W. (1937). <a rel="nofollow" class="external text" href="https://doi.org/10.1158%2Fajc.1937.537">"Papillomatosis in the forestomach of the rat and its bearing on the work of Fibiger"</a>. <i>The American Journal of Cancer</i>. <b>31</b> (4): 537–555. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1158%2Fajc.1937.537">10.1158/ajc.1937.537</a></span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=The+American+Journal+of+Cancer&amp;rft.atitle=Papillomatosis+in+the+forestomach+of+the+rat+and+its+bearing+on+the+work+of+Fibiger&amp;rft.volume=31&amp;rft.issue=4&amp;rft.pages=537-555&amp;rft.date=1937&amp;rft_id=info%3Adoi%2F10.1158%2Fajc.1937.537&amp;rft.aulast=Cramer&amp;rft.aufirst=W.&amp;rft_id=https%3A%2F%2Fdoi.org%2F10.1158%252Fajc.1937.537&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AJohannes+Fibiger" class="Z3988"></span></span> </li> <li id="cite_note-25"><span class="mw-cite-backlink"><b><a href="#cite_ref-25">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHitchcockBell1952" class="citation journal cs1">Hitchcock, Claude R.; Bell, E. T. (1952). "Studies on the nematode parasite, <i>Gongylonema Neoplasticum</i> (<i>Spiroptera Neoplasticum</i>), and avitaminosis a in the forestomach of rats: Comparison with Fibiger's results". <i>Journal of the National Cancer Institute</i>. <b>12</b> (6): 1345–1387. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fjnci%2F12.6.1345">10.1093/jnci/12.6.1345</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/14939031">14939031</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Journal+of+the+National+Cancer+Institute&amp;rft.atitle=Studies+on+the+nematode+parasite%2C+Gongylonema+Neoplasticum+%28Spiroptera+Neoplasticum%29%2C+and+avitaminosis+a+in+the+forestomach+of+rats%3A+Comparison+with+Fibiger%27s+results&amp;rft.volume=12&amp;rft.issue=6&amp;rft.pages=1345-1387&amp;rft.date=1952&amp;rft_id=info%3Adoi%2F10.1093%2Fjnci%2F12.6.1345&amp;rft_id=info%3Apmid%2F14939031&amp;rft.aulast=Hitchcock&amp;rft.aufirst=Claude+R.&amp;rft.au=Bell%2C+E.+T.&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AJohannes+Fibiger" class="Z3988"></span></span> </li> <li id="cite_note-26"><span class="mw-cite-backlink"><b><a href="#cite_ref-26">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation journal cs1"><a rel="nofollow" class="external text" href="https://doi.org/10.3322%2Fcanjclin.27.3.172">"Katsusaburo Yamagiwa (1863–1930)"</a>. <i>CA: A Cancer Journal for Clinicians</i>. <b>27</b> (3): 172–173. 1977. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.3322%2Fcanjclin.27.3.172">10.3322/canjclin.27.3.172</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/406017">406017</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a>&#160;<a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:36283426">36283426</a>. <q>Yamagiwa, then Director of the Department of Pathology at Tokyo Imperial University Medical School, had theorized that repetition or continuation of chronic irritation caused precancerous alterations in previously normal epithelium. If the irritant continued its action, carcinoma could result. These data, publicly presented at a special meeting of the Tokyo Medical Society and reprinted below, focused attention on chemical carcinogenesis. Further more, his experimental method provided researchers with a means of producing cancer in the laboratory and anticipated investigation of specific carcinogenic agents and the precise way in which they acted. Within a decade, Keller and associates extracted a highly potent carcinogenic hydrocarbon from coal tar. Dr. Yamagiwa had begun a new era in cancer research.</q></cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=CA%3A+A+Cancer+Journal+for+Clinicians&amp;rft.atitle=Katsusaburo+Yamagiwa+%281863%E2%80%931930%29&amp;rft.volume=27&amp;rft.issue=3&amp;rft.pages=172-173&amp;rft.date=1977&amp;rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A36283426%23id-name%3DS2CID&amp;rft_id=info%3Apmid%2F406017&amp;rft_id=info%3Adoi%2F10.3322%2Fcanjclin.27.3.172&amp;rft_id=https%3A%2F%2Fdoi.org%2F10.3322%252Fcanjclin.27.3.172&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AJohannes+Fibiger" class="Z3988"></span></span> </li> <li id="cite_note-27"><span class="mw-cite-backlink"><b><a href="#cite_ref-27">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFJames_R._Bartholomew" class="citation web cs1">James R. Bartholomew. <a rel="nofollow" class="external text" href="http://chemeducator.org/bibs/0010006/1060478mr.htm">"Katsusaburo Yamagiwa's Nobel candidacy: Physiology or medicine in the 1920s"</a>. <q>explores the candidacy of Yamagiwa, who had developed the world's first efficient method for producing cancer artificially in the laboratory by swabbing coal tar on rabbits' ears, which had stimulated activity among cancer researchers worldwide. Johannes Fibiger of Denmark, who discovered how to use parasites to cause cancer in rats two years before Yamagiwa's achievement, received the prize, probably because nominations were often greatly influenced by acquaintanceship, geography, and the marginalization that distance from other centers imposed on the Japanese.</q></cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rft.genre=unknown&amp;rft.btitle=Katsusaburo+Yamagiwa%27s+Nobel+candidacy%3A+Physiology+or+medicine+in+the+1920s&amp;rft.au=James+R.+Bartholomew&amp;rft_id=http%3A%2F%2Fchemeducator.org%2Fbibs%2F0010006%2F1060478mr.htm&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AJohannes+Fibiger" class="Z3988"></span></span> </li> <li id="cite_note-28"><span class="mw-cite-backlink"><b><a href="#cite_ref-28">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20120323104136/http://www.britannica.com/nobelprize/article-224787">Guide to Nobel Prize</a>. Britannica.com. Retrieved on 25 September 2010.</span> </li> <li id="cite_note-29"><span class="mw-cite-backlink"><b><a href="#cite_ref-29">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFFriedReddyMayer2010" class="citation journal cs1">Fried B, Reddy A, Mayer D (2010). "Helminths in human carcinogenesis". <i>Cancer Lett</i>. <b>305</b> (2): 239–249. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.canlet.2010.07.008">10.1016/j.canlet.2010.07.008</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/20667649">20667649</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Cancer+Lett&amp;rft.atitle=Helminths+in+human+carcinogenesis&amp;rft.volume=305&amp;rft.issue=2&amp;rft.pages=239-249&amp;rft.date=2010&amp;rft_id=info%3Adoi%2F10.1016%2Fj.canlet.2010.07.008&amp;rft_id=info%3Apmid%2F20667649&amp;rft.aulast=Fried&amp;rft.aufirst=B&amp;rft.au=Reddy%2C+A&amp;rft.au=Mayer%2C+D&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AJohannes+Fibiger" class="Z3988"></span></span> </li> <li id="cite_note-30"><span class="mw-cite-backlink"><b><a href="#cite_ref-30">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFYoungCampbellHallJex2010" class="citation journal cs1">Young ND, Campbell BE, Hall RS, Jex AR, Cantacessi C, Laha T, Sohn WM, Sripa B, Loukas A, Brindley PJ, Gasser RB (2010). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2889816">"Unlocking the transcriptomes of two carcinogenic parasites, <i>Clonorchis sinensis</i> and <i>Opisthorchis viverrini</i>"</a>. <i>PLOS Negl Trop Dis</i>. <b>4</b> (6): e719. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1371%2Fjournal.pntd.0000719">10.1371/journal.pntd.0000719</a></span>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a>&#160;<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2889816">2889816</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/20582164">20582164</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=PLOS+Negl+Trop+Dis&amp;rft.atitle=Unlocking+the+transcriptomes+of+two+carcinogenic+parasites%2C+Clonorchis+sinensis+and+Opisthorchis+viverrini&amp;rft.volume=4&amp;rft.issue=6&amp;rft.pages=e719&amp;rft.date=2010&amp;rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC2889816%23id-name%3DPMC&amp;rft_id=info%3Apmid%2F20582164&amp;rft_id=info%3Adoi%2F10.1371%2Fjournal.pntd.0000719&amp;rft.aulast=Young&amp;rft.aufirst=ND&amp;rft.au=Campbell%2C+BE&amp;rft.au=Hall%2C+RS&amp;rft.au=Jex%2C+AR&amp;rft.au=Cantacessi%2C+C&amp;rft.au=Laha%2C+T&amp;rft.au=Sohn%2C+WM&amp;rft.au=Sripa%2C+B&amp;rft.au=Loukas%2C+A&amp;rft.au=Brindley%2C+PJ&amp;rft.au=Gasser%2C+RB&amp;rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC2889816&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AJohannes+Fibiger" class="Z3988"></span></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="Further_reading">Further reading</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Johannes_Fibiger&amp;action=edit&amp;section=11" title="Edit section: Further reading"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><i>Nobel Lectures, Physiology or Medicine 1922–1941</i>, Elsevier Publishing Company, Amsterdam, 1965.</li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFModlinKiddHinoue2001" class="citation journal cs1">Modlin, I M; Kidd, M; Hinoue, T (2001). "Of Fibiger and fables: a cautionary tale of cockroaches and Helicobacter pylori". <i>J. Clin. Gastroenterol</i>. <b>33</b> (3) (published September 2001): 177–9. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1097%2F00004836-200109000-00001">10.1097/00004836-200109000-00001</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/11500602">11500602</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=J.+Clin.+Gastroenterol.&amp;rft.atitle=Of+Fibiger+and+fables%3A+a+cautionary+tale+of+cockroaches+and+Helicobacter+pylori&amp;rft.volume=33&amp;rft.issue=3&amp;rft.pages=177-9&amp;rft.date=2001&amp;rft_id=info%3Adoi%2F10.1097%2F00004836-200109000-00001&amp;rft_id=info%3Apmid%2F11500602&amp;rft.aulast=Modlin&amp;rft.aufirst=I+M&amp;rft.au=Kidd%2C+M&amp;rft.au=Hinoue%2C+T&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AJohannes+Fibiger" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFRaju1998" class="citation journal cs1">Raju, T N (1998). "The Nobel chronicles. 1926: Johannes Andreas Grib Fibiger (1867–1928)". <i><a href="/wiki/The_Lancet" title="The Lancet">The Lancet</a></i>. <b>352</b> (9140) (published 14 November 1998): 1635. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2FS0140-6736%2898%2900067-1">10.1016/S0140-6736(98)00067-1</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/9843145">9843145</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a>&#160;<a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:54400144">54400144</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=The+Lancet&amp;rft.atitle=The+Nobel+chronicles.+1926%3A+Johannes+Andreas+Grib+Fibiger+%281867%E2%80%931928%29&amp;rft.volume=352&amp;rft.issue=9140&amp;rft.pages=1635&amp;rft.date=1998&amp;rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A54400144%23id-name%3DS2CID&amp;rft_id=info%3Apmid%2F9843145&amp;rft_id=info%3Adoi%2F10.1016%2FS0140-6736%2898%2900067-1&amp;rft.aulast=Raju&amp;rft.aufirst=T+N&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AJohannes+Fibiger" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFStolleyLasky1992" class="citation journal cs1">Stolley, P D; Lasky, T (1992). "Johannes Fibiger and his Nobel Prize for the hypothesis that a worm causes stomach cancer". <i>Ann. Intern. Med</i>. <b>116</b> (9) (published 1 May 1992): 765–9. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.7326%2F0003-4819-116-9-765">10.7326/0003-4819-116-9-765</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/1558350">1558350</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Ann.+Intern.+Med.&amp;rft.atitle=Johannes+Fibiger+and+his+Nobel+Prize+for+the+hypothesis+that+a+worm+causes+stomach+cancer&amp;rft.volume=116&amp;rft.issue=9&amp;rft.pages=765-9&amp;rft.date=1992&amp;rft_id=info%3Adoi%2F10.7326%2F0003-4819-116-9-765&amp;rft_id=info%3Apmid%2F1558350&amp;rft.aulast=Stolley&amp;rft.aufirst=P+D&amp;rft.au=Lasky%2C+T&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AJohannes+Fibiger" class="Z3988"></span></li></ul> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Johannes_Fibiger&amp;action=edit&amp;section=12" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a rel="nofollow" class="external text" href="https://www.nobelprize.org/laureate/316">Johannes Fibiger</a> on Nobelprize.org <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q216324#P8024" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span> including the Nobel Lecture on 12 December 1927 <i>Investigations on Spiroptera Carcinoma and the Experimental Induction of Cancer</i></li> <li><a rel="nofollow" class="external text" href="http://www.whonamedit.com/doctor.cfm/1443.html">Biography from <i>WhoNamedIt</i></a></li> <li><a rel="nofollow" class="external text" href="http://www.yourdictionary.com/johannes-fibiger">Biography from <i>YourDictionary</i></a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20140201201612/http://www.yourdictionary.com/johannes-fibiger">Archived</a> 1 February 2014 at the <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback 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aria-labelledby="Laureates_of_the_Nobel_Prize_in_Physiology_or_Medicine" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar 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template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Nobel_Prize_in_Physiology_or_Medicine" title="Special:EditPage/Template:Nobel Prize in Physiology or Medicine"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Laureates_of_the_Nobel_Prize_in_Physiology_or_Medicine" style="font-size:114%;margin:0 4em"><a href="/wiki/List_of_Nobel_laureates_in_Physiology_or_Medicine" title="List of Nobel laureates in Physiology or Medicine">Laureates</a> of the <a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Nobel Prize in Physiology or Medicine</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">1901–1925</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1901: <a href="/wiki/Emil_von_Behring" title="Emil von Behring">Emil Behring</a></span></li> <li><span class="nowrap">1902: <a href="/wiki/Ronald_Ross" title="Ronald Ross">Ronald Ross</a></span></li> <li><span class="nowrap">1903: <a href="/wiki/Niels_Ryberg_Finsen" title="Niels Ryberg Finsen">Niels Finsen</a></span></li> <li><span class="nowrap">1904: <a href="/wiki/Ivan_Pavlov" title="Ivan Pavlov">Ivan Pavlov</a></span></li> <li><span class="nowrap">1905: <a href="/wiki/Robert_Koch" title="Robert Koch">Robert Koch</a></span></li> <li><span class="nowrap">1906: <a href="/wiki/Camillo_Golgi" title="Camillo Golgi">Camillo Golgi</a> / <a href="/wiki/Santiago_Ram%C3%B3n_y_Cajal" title="Santiago Ramón y Cajal">Santiago Ramón y Cajal</a></span></li> <li><span class="nowrap">1907: <a href="/wiki/Charles_Louis_Alphonse_Laveran" title="Charles Louis Alphonse Laveran">Alphonse Laveran</a></span></li> <li><span class="nowrap">1908: <a href="/wiki/%C3%89lie_Metchnikoff" title="Élie Metchnikoff">Élie Metchnikoff</a> / <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Paul Ehrlich</a></span></li> <li><span class="nowrap">1909: <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Emil Kocher</a></span></li> <li><span class="nowrap">1910: <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Albrecht Kossel</a></span></li> <li><span class="nowrap">1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Allvar Gullstrand</a></span></li> <li><span class="nowrap">1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Alexis Carrel</a></span></li> <li><span class="nowrap">1913: <a href="/wiki/Charles_Richet" title="Charles Richet">Charles Richet</a></span></li> <li><span class="nowrap">1914: <a href="/wiki/R%C3%B3bert_B%C3%A1r%C3%A1ny" class="mw-redirect" title="Róbert Bárány">Róbert Bárány</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1915</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1916</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1917</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1918</span></span></li> <li><span class="nowrap">1919: <a href="/wiki/Jules_Bordet" title="Jules Bordet">Jules Bordet</a></span></li> <li><span class="nowrap">1920: <a href="/wiki/August_Krogh" title="August Krogh">August Krogh</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1921</span></span></li> <li><span class="nowrap">1922: <a href="/wiki/Archibald_Hill" title="Archibald Hill">Archibald Hill</a> / <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Otto Meyerhof</a></span></li> <li><span class="nowrap">1923: <a href="/wiki/Frederick_Banting" title="Frederick Banting">Frederick Banting</a> / <a href="/wiki/John_Macleod_(physiologist)" title="John Macleod (physiologist)">John Macleod</a></span></li> <li><span class="nowrap">1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Willem Einthoven</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1925</span></span></li></ul> </div></td><td class="noviewer navbox-image" rowspan="5" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/80px-Nobel_Prize.png" decoding="async" width="80" height="79" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/120px-Nobel_Prize.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/160px-Nobel_Prize.png 2x" data-file-width="500" data-file-height="492" /></span></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1926–1950</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1926: <a class="mw-selflink selflink">Johannes Fibiger</a></span></li> <li><span class="nowrap">1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Julius Wagner-Jauregg</a></span></li> <li><span class="nowrap">1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Charles Nicolle</a></span></li> <li><span class="nowrap">1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Christiaan Eijkman</a> / <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Frederick Gowland Hopkins</a></span></li> <li><span class="nowrap">1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Karl Landsteiner</a></span></li> <li><span class="nowrap">1931: <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Otto Warburg</a></span></li> <li><span class="nowrap">1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Charles Scott Sherrington</a> / <a href="/wiki/Edgar_Adrian" title="Edgar Adrian">Edgar Adrian</a></span></li> <li><span class="nowrap">1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Thomas Morgan</a></span></li> <li><span class="nowrap">1934: <a href="/wiki/George_Whipple" title="George Whipple">George Whipple</a> / <a href="/wiki/George_Minot" title="George Minot">George Minot</a> / <a href="/wiki/William_P._Murphy" title="William P. Murphy">William Murphy</a></span></li> <li><span class="nowrap">1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Hans Spemann</a></span></li> <li><span class="nowrap">1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Henry Dale</a> / <a href="/wiki/Otto_Loewi" title="Otto Loewi">Otto Loewi</a></span></li> <li><span class="nowrap">1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Albert Szent-Györgyi</a></span></li> <li><span class="nowrap">1938: <a href="/wiki/Corneille_Heymans" title="Corneille Heymans">Corneille Heymans</a></span></li> <li><span class="nowrap">1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Gerhard Domagk</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1940</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1941</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1942</span></span></li> <li><span class="nowrap">1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Henrik Dam</a> / <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Edward Doisy</a></span></li> <li><span class="nowrap">1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Joseph Erlanger</a> / <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Herbert Gasser</a></span></li> <li><span class="nowrap">1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Alexander Fleming</a> / <a href="/wiki/Ernst_Chain" title="Ernst Chain">Ernst Chain</a> / <a href="/wiki/Howard_Florey" title="Howard Florey">Howard Florey</a></span></li> <li><span class="nowrap">1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Hermann Muller</a></span></li> <li><span class="nowrap">1947: <a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">Carl Cori</a> / <a href="/wiki/Gerty_Cori" title="Gerty Cori">Gerty Cori</a> / <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Bernardo Houssay</a></span></li> <li><span class="nowrap">1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Paul Müller</a></span></li> <li><span class="nowrap">1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Walter Hess</a> / <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">António Egas Moniz</a></span></li> <li><span class="nowrap">1950: <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Edward Kendall</a> / <a href="/wiki/Tadeusz_Reichstein" title="Tadeusz Reichstein">Tadeusz Reichstein</a> / <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Philip Hench</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1951–1975</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Max Theiler</a></span></li> <li><span class="nowrap">1952: <a href="/wiki/Selman_Waksman" title="Selman Waksman">Selman Waksman</a></span></li> <li><span class="nowrap">1953: <a href="/wiki/Hans_Krebs_(biochemist)" title="Hans Krebs (biochemist)">Hans Krebs</a> / <a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Fritz Lipmann</a></span></li> <li><span class="nowrap">1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">John Enders</a> / <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Thomas Weller</a> / <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Frederick Robbins</a></span></li> <li><span class="nowrap">1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Hugo Theorell</a></span></li> <li><span class="nowrap">1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">André Cournand</a> / <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Werner Forssmann</a> / <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Dickinson W. Richards</a></span></li> <li><span class="nowrap">1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Daniel Bovet</a></span></li> <li><span class="nowrap">1958: <a href="/wiki/George_Beadle" title="George Beadle">George Beadle</a> / <a href="/wiki/Edward_Tatum" title="Edward Tatum">Edward Tatum</a> / <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Joshua Lederberg</a></span></li> <li><span class="nowrap">1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Severo Ochoa</a> / <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Arthur Kornberg</a></span></li> <li><span class="nowrap">1960: <a href="/wiki/Macfarlane_Burnet" title="Macfarlane Burnet">Frank Burnet</a> / <a href="/wiki/Peter_Medawar" title="Peter Medawar">Peter Medawar</a></span></li> <li><span class="nowrap">1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Georg von Békésy</a></span></li> <li><span class="nowrap">1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Francis Crick</a> / <a href="/wiki/James_Watson" title="James Watson">James Watson</a> / <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Maurice Wilkins</a></span></li> <li><span class="nowrap">1963: <a href="/wiki/John_Eccles_(neurophysiologist)" title="John Eccles (neurophysiologist)">John Eccles</a> / <a href="/wiki/Alan_Hodgkin" title="Alan Hodgkin">Alan Hodgkin</a> / <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Andrew Huxley</a></span></li> <li><span class="nowrap">1964: <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Konrad Bloch</a> / <a href="/wiki/Feodor_Lynen" title="Feodor Lynen">Feodor Lynen</a></span></li> <li><span class="nowrap">1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">François Jacob</a> / <a href="/wiki/Andr%C3%A9_Michel_Lwoff" title="André Michel Lwoff">André Lwoff</a> / <a href="/wiki/Jacques_Monod" title="Jacques Monod">Jacques Monod</a></span></li> <li><span class="nowrap">1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Francis Rous</a> / <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Charles B. Huggins</a></span></li> <li><span class="nowrap">1967: <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Ragnar Granit</a> / <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Haldan Hartline</a> / <a href="/wiki/George_Wald" title="George Wald">George Wald</a></span></li> <li><span class="nowrap">1968: <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Robert W. Holley</a> / <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Har Khorana</a> / <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Marshall Nirenberg</a></span></li> <li><span class="nowrap">1969: <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Max Delbrück</a> / <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Alfred Hershey</a> / <a href="/wiki/Salvador_Luria" title="Salvador Luria">Salvador Luria</a></span></li> <li><span class="nowrap">1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Bernard Katz</a> / <a href="/wiki/Ulf_von_Euler" title="Ulf von Euler">Ulf von Euler</a> / <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Julius Axelrod</a></span></li> <li><span class="nowrap">1971: <a href="/wiki/Earl_Wilbur_Sutherland_Jr." title="Earl Wilbur Sutherland Jr.">Earl Sutherland Jr.</a></span></li> <li><span class="nowrap">1972: <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Gerald Edelman</a> / <a href="/wiki/Rodney_Robert_Porter" class="mw-redirect" title="Rodney Robert Porter">Rodney Porter</a></span></li> <li><span class="nowrap">1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Karl von Frisch</a> / <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Konrad Lorenz</a> / <a href="/wiki/Nikolaas_Tinbergen" title="Nikolaas Tinbergen">Nikolaas Tinbergen</a></span></li> <li><span class="nowrap">1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Albert Claude</a> / <a href="/wiki/Christian_de_Duve" title="Christian de Duve">Christian de Duve</a> / <a href="/wiki/George_Emil_Palade" title="George Emil Palade">George Palade</a></span></li> <li><span class="nowrap">1975: <a href="/wiki/David_Baltimore" title="David Baltimore">David Baltimore</a> / <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Renato Dulbecco</a> / <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Howard Temin</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1976–2000</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1976: <a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Baruch Blumberg</a> / <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Daniel Gajdusek</a></span></li> <li><span class="nowrap">1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Roger Guillemin</a> / <a href="/wiki/Andrew_Schally" title="Andrew Schally">Andrew Schally</a> / <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Rosalyn Yalow</a></span></li> <li><span class="nowrap">1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Werner Arber</a> / <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Daniel Nathans</a> / <a href="/wiki/Hamilton_O._Smith" title="Hamilton O. Smith">Hamilton O. Smith</a></span></li> <li><span class="nowrap">1979: <a href="/wiki/Allan_MacLeod_Cormack" title="Allan MacLeod Cormack">Allan Cormack</a> / <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Godfrey Hounsfield</a></span></li> <li><span class="nowrap">1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Baruj Benacerraf</a> / <a href="/wiki/Jean_Dausset" title="Jean Dausset">Jean Dausset</a> / <a href="/wiki/George_Davis_Snell" title="George Davis Snell">George Snell</a></span></li> <li><span class="nowrap">1981: <a href="/wiki/Roger_Wolcott_Sperry" title="Roger Wolcott Sperry">Roger Sperry</a> / <a href="/wiki/David_H._Hubel" title="David H. Hubel">David H. Hubel</a> / <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Torsten Wiesel</a></span></li> <li><span class="nowrap">1982: <a href="/wiki/Sune_Bergstr%C3%B6m" title="Sune Bergström">Sune Bergström</a> / <a href="/wiki/Bengt_I._Samuelsson" title="Bengt I. Samuelsson">Bengt I. Samuelsson</a> / <a href="/wiki/John_Vane" title="John Vane">John Vane</a></span></li> <li><span class="nowrap">1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a></span></li> <li><span class="nowrap">1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Niels Jerne</a> / <a href="/wiki/Georges_J._F._K%C3%B6hler" title="Georges J. F. Köhler">Georges Köhler</a> / <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a></span></li> <li><span class="nowrap">1985: <a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Michael Brown</a> / <a href="/wiki/Joseph_L._Goldstein" title="Joseph L. Goldstein">Joseph L. Goldstein</a></span></li> <li><span class="nowrap">1986: <a href="/wiki/Stanley_Cohen_(biochemist)" title="Stanley Cohen (biochemist)">Stanley Cohen</a> / <a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Rita Levi-Montalcini</a></span></li> <li><span class="nowrap">1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Susumu Tonegawa</a></span></li> <li><span class="nowrap">1988: <a href="/wiki/James_Black_(pharmacologist)" title="James Black (pharmacologist)">James W. Black</a> / <a href="/wiki/Gertrude_B._Elion" title="Gertrude B. Elion">Gertrude B. Elion</a> / <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">George H. Hitchings</a></span></li> <li><span class="nowrap">1989: <a href="/wiki/J._Michael_Bishop" title="J. Michael Bishop">J. Michael Bishop</a> / <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Harold E. Varmus</a></span></li> <li><span class="nowrap">1990: <a href="/wiki/Joseph_Murray" class="mw-redirect" title="Joseph Murray">Joseph Murray</a> / <a href="/wiki/E._Donnall_Thomas" title="E. Donnall Thomas">E. Donnall Thomas</a></span></li> <li><span class="nowrap">1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Erwin Neher</a> / <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Bert Sakmann</a></span></li> <li><span class="nowrap">1992: <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Edmond Fischer</a> / <a href="/wiki/Edwin_G._Krebs" title="Edwin G. Krebs">Edwin G. Krebs</a></span></li> <li><span class="nowrap">1993: <a href="/wiki/Richard_J._Roberts" title="Richard J. Roberts">Richard J. Roberts</a> / <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Phillip Sharp</a></span></li> <li><span class="nowrap">1994: <a href="/wiki/Alfred_G._Gilman" title="Alfred G. Gilman">Alfred G. Gilman</a> / <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Martin Rodbell</a></span></li> <li><span class="nowrap">1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Edward B. Lewis</a> / <a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Christiane Nüsslein-Volhard</a> / <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Eric F. Wieschaus</a></span></li> <li><span class="nowrap">1996: <a href="/wiki/Peter_C._Doherty" class="mw-redirect" title="Peter C. Doherty">Peter C. Doherty</a> / <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Rolf M. Zinkernagel</a></span></li> <li><span class="nowrap">1997: <a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Stanley B. Prusiner</a></span></li> <li><span class="nowrap">1998: <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Robert F. Furchgott</a> / <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Louis Ignarro</a> / <a href="/wiki/Ferid_Murad" title="Ferid Murad">Ferid Murad</a></span></li> <li><span class="nowrap">1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Günter Blobel</a></span></li> <li><span class="nowrap">2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Arvid Carlsson</a> / <a href="/wiki/Paul_Greengard" title="Paul Greengard">Paul Greengard</a> / <a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric Kandel</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2001–present</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">2001: <a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Leland H. Hartwell</a> / <a href="/wiki/Tim_Hunt" title="Tim Hunt">Tim Hunt</a> / <a href="/wiki/Paul_Nurse" title="Paul Nurse">Paul Nurse</a></span></li> <li><span class="nowrap">2002: <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Sydney Brenner</a> / <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">H. Robert Horvitz</a> / <a href="/wiki/John_Sulston" title="John Sulston">John E. Sulston</a></span></li> <li><span class="nowrap">2003: <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Paul Lauterbur</a> / <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Peter Mansfield</a></span></li> <li><span class="nowrap">2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Richard Axel</a> / <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Linda B. Buck</a></span></li> <li><span class="nowrap">2005: <a href="/wiki/Barry_Marshall" title="Barry Marshall">Barry Marshall</a> / <a href="/wiki/Robin_Warren" title="Robin Warren">Robin Warren</a></span></li> <li><span class="nowrap">2006: <a href="/wiki/Andrew_Fire" title="Andrew Fire">Andrew Fire</a> / <a href="/wiki/Craig_Mello" title="Craig Mello">Craig Mello</a></span></li> <li><span class="nowrap">2007: <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Mario Capecchi</a> / <a href="/wiki/Martin_Evans" title="Martin Evans">Martin Evans</a> / <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a></span></li> <li><span class="nowrap">2008: <a href="/wiki/Harald_zur_Hausen" title="Harald zur Hausen">Harald zur Hausen</a> / <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Luc Montagnier</a> / <a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Françoise Barré-Sinoussi</a></span></li> <li><span class="nowrap">2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Elizabeth Blackburn</a> / <a href="/wiki/Carol_W._Greider" title="Carol W. Greider">Carol W. Greider</a> / <a href="/wiki/Jack_W._Szostak" title="Jack W. Szostak">Jack W. Szostak</a></span></li> <li><span class="nowrap">2010: <a href="/wiki/Robert_Edwards_(physiologist)" title="Robert Edwards (physiologist)">Robert G. Edwards</a></span></li> <li><span class="nowrap">2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Bruce Beutler</a> / <a href="/wiki/Jules_A._Hoffmann" title="Jules A. Hoffmann">Jules A. Hoffmann</a> / <a href="/wiki/Ralph_M._Steinman" title="Ralph M. Steinman">Ralph M. Steinman</a> (<i>posthumously</i>)</span></li> <li><span class="nowrap">2012: <a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> / <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Shinya Yamanaka</a></span></li> <li><span class="nowrap">2013: <a href="/wiki/James_Rothman" title="James Rothman">James Rothman</a> / <a href="/wiki/Randy_Schekman" title="Randy Schekman">Randy Schekman</a> / <a href="/wiki/Thomas_C._S%C3%BCdhof" title="Thomas C. Südhof">Thomas C. Südhof</a></span></li> <li><span class="nowrap">2014: <a href="/wiki/John_O%27Keefe_(neuroscientist)" title="John O&#39;Keefe (neuroscientist)">John O'Keefe</a> / <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">May-Britt Moser</a> / <a href="/wiki/Edvard_Moser" title="Edvard Moser">Edvard Moser</a></span></li> <li><span class="nowrap">2015: <a href="/wiki/William_C._Campbell_(scientist)" title="William C. Campbell (scientist)">William C. Campbell</a> / <a href="/wiki/Satoshi_%C5%8Cmura" title="Satoshi Ōmura">Satoshi Ōmura</a> / <a href="/wiki/Tu_Youyou" title="Tu Youyou">Tu Youyou</a></span></li> <li><span class="nowrap">2016: <a href="/wiki/Yoshinori_Ohsumi" title="Yoshinori Ohsumi">Yoshinori Ohsumi</a></span></li> <li><span class="nowrap">2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Jeffrey C. Hall</a> / <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Michael Rosbash</a> / <a href="/wiki/Michael_W._Young" title="Michael W. Young">Michael W. Young</a></span></li> <li><span class="nowrap">2018: <a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a> / <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Tasuku Honjo</a></span></li> <li><span class="nowrap">2019: <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Gregg L. Semenza</a> / <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Peter J. Ratcliffe</a> / <a href="/wiki/William_Kaelin_Jr." title="William Kaelin Jr.">William Kaelin Jr.</a></span></li> <li><span class="nowrap"><a href="/wiki/2020_Nobel_Prize_in_Physiology_or_Medicine" title="2020 Nobel Prize in Physiology or Medicine">2020</a>: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Harvey J. Alter</a> / <a href="/wiki/Michael_Houghton_(virologist)" class="mw-redirect" title="Michael Houghton (virologist)">Michael Houghton</a> / <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Charles M. Rice</a></span></li> <li><span class="nowrap"><a href="/wiki/2021_Nobel_Prize_in_Physiology_or_Medicine" title="2021 Nobel Prize in Physiology or Medicine">2021</a>: <a href="/wiki/David_Julius" title="David Julius">David Julius</a> / <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Ardem Patapoutian</a></span></li> <li><span class="nowrap"><a href="/wiki/2022_Nobel_Prize_in_Physiology_or_Medicine" title="2022 Nobel Prize in Physiology or Medicine">2022</a>: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Svante Pääbo</a></span></li> <li><span class="nowrap"><a href="/wiki/2023_Nobel_Prize_in_Physiology_or_Medicine" title="2023 Nobel Prize in Physiology or Medicine">2023</a>: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Katalin Karikó</a> / <a href="/wiki/Drew_Weissman" title="Drew Weissman">Drew Weissman</a></span></li> <li><span class="nowrap">2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a> / <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><style data-mw-deduplicate="TemplateStyles:r886047488">.mw-parser-output .nobold{font-weight:normal}</style></div><div role="navigation" class="navbox" aria-labelledby="1926_Nobel_Prize_laureates" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:1926_Nobel_Prize_winners" title="Template:1926 Nobel Prize winners"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:1926_Nobel_Prize_winners" title="Template talk:1926 Nobel Prize winners"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:1926_Nobel_Prize_winners" title="Special:EditPage/Template:1926 Nobel Prize winners"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="1926_Nobel_Prize_laureates" style="font-size:114%;margin:0 4em">1926 <a href="/wiki/Nobel_Prize" title="Nobel Prize">Nobel Prize</a> laureates</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Chemistry</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Theodor_Svedberg" title="Theodor Svedberg">Theodor Svedberg</a> (Sweden)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Literature" title="Nobel Prize in Literature">Literature</a> (<a href="/wiki/1926_Nobel_Prize_in_Literature" title="1926 Nobel Prize in Literature">1926</a>)</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Grazia_Deledda" title="Grazia Deledda">Grazia Deledda</a> (Italy)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Peace_Prize" title="Nobel Peace Prize">Peace</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Aristide_Briand" title="Aristide Briand">Aristide Briand</a> (France)</li> <li><a href="/wiki/Gustav_Stresemann" title="Gustav Stresemann">Gustav Stresemann</a> (Germany)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physics" title="Nobel Prize in Physics">Physics</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Jean_Baptiste_Perrin" title="Jean Baptiste Perrin">Jean Baptiste Perrin</a> (France)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Physiology or Medicine</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a class="mw-selflink selflink">Johannes Fibiger</a> (Denmark)</li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <dl><dt><span class="nobold"><a href="/wiki/List_of_Nobel_laureates" title="List of Nobel laureates">Nobel Prize recipients</a></span></dt> <dd><a href="/wiki/Template:1921_Nobel_Prize_winners" title="Template:1921 Nobel Prize winners">1921</a></dd> <dd><a href="/wiki/Template:1922_Nobel_Prize_winners" title="Template:1922 Nobel Prize winners">1922</a></dd> <dd><a href="/wiki/Template:1923_Nobel_Prize_winners" title="Template:1923 Nobel Prize winners">1923</a></dd> <dd><a href="/wiki/Template:1924_Nobel_Prize_winners" title="Template:1924 Nobel Prize winners">1924</a></dd> <dd><a href="/wiki/Template:1925_Nobel_Prize_winners" title="Template:1925 Nobel Prize winners">1925</a></dd> <dd><b>1926 </b></dd> <dd><a href="/wiki/Template:1927_Nobel_Prize_winners" title="Template:1927 Nobel Prize winners">1927</a></dd> <dd><a href="/wiki/Template:1928_Nobel_Prize_winners" title="Template:1928 Nobel Prize winners">1928</a></dd> <dd><a href="/wiki/Template:1929_Nobel_Prize_winners" title="Template:1929 Nobel Prize winners">1929</a></dd> <dd><a href="/wiki/Template:1930_Nobel_Prize_winners" title="Template:1930 Nobel Prize winners">1930</a></dd> <dd><a href="/wiki/Template:1931_Nobel_Prize_winners" title="Template:1931 Nobel Prize winners">1931</a></dd></dl> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox authority-control" aria-labelledby="Authority_control_databases_frameless&amp;#124;text-top&amp;#124;10px&amp;#124;alt=Edit_this_at_Wikidata&amp;#124;link=https&amp;#58;//www.wikidata.org/wiki/Q216324#identifiers&amp;#124;class=noprint&amp;#124;Edit_this_at_Wikidata" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Authority_control_databases_frameless&amp;#124;text-top&amp;#124;10px&amp;#124;alt=Edit_this_at_Wikidata&amp;#124;link=https&amp;#58;//www.wikidata.org/wiki/Q216324#identifiers&amp;#124;class=noprint&amp;#124;Edit_this_at_Wikidata" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Authority_control" title="Help:Authority control">Authority control databases</a> <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q216324#identifiers" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, 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style="width:1%">National</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://d-nb.info/gnd/117507245">Germany</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/no2006059391">United States</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://catalogue.bnf.fr/ark:/12148/cb112214321">France</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://data.bnf.fr/ark:/12148/cb112214321">BnF data</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://aleph.nkp.cz/F/?func=find-c&amp;local_base=aut&amp;ccl_term=ica=nlk20000079376&amp;CON_LNG=ENG">Czech Republic</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="http://data.bibliotheken.nl/id/thes/p115150390">Netherlands</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://libris.kb.se/gdsvvwl01bwcntc">Sweden</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="http://olduli.nli.org.il/F/?func=find-b&amp;local_base=NLX10&amp;find_code=UID&amp;request=987007594408105171">Israel</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">People</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.deutsche-biographie.de/pnd117507245.html?language=en">Deutsche Biographie</a></span></li></ul></div></td></tr></tbody></table></div> <!-- NewPP limit report Parsed by mw‐web.eqiad.main‐5dc468848‐kbrbs Cached time: 20241122164820 Cache expiry: 2592000 Reduced expiry: false Complications: 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