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</div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Avram Hershko</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 54 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-54" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">54 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%A3%D9%81%D8%B1%D8%A7%D9%85_%D9%87%D9%8A%D8%B1%D8%B4%D9%83%D9%88" title="أفرام هيرشكو – Arabic" lang="ar" hreflang="ar" data-title="أفرام هيرشكو" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Avram_Her%C5%9Fko" title="Avram Herşko – Azerbaijani" lang="az" hreflang="az" data-title="Avram Herşko" data-language-autonym="Azərbaycanca" data-language-local-name="Azerbaijani" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D8%A7%D9%81%D8%B1%D8%A7%D9%85_%D9%87%D8%B1%D8%B4%DA%A9%D9%88" title="افرام هرشکو – South Azerbaijani" lang="azb" hreflang="azb" data-title="افرام هرشکو" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%86%E0%A6%AD%E0%A7%8D%E0%A6%B0%E0%A6%BE%E0%A6%AE_%E0%A6%B9%E0%A7%87%E0%A6%B0%E0%A7%8D%E0%A6%B6%E0%A6%95%E0%A7%8B" title="আভ্রাম হের্শকো – Bangla" lang="bn" hreflang="bn" data-title="আভ্রাম হের্শকো" data-language-autonym="বাংলা" data-language-local-name="Bangla" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%90%D1%9E%D1%80%D0%B0%D0%BC_%D0%93%D0%B5%D1%80%D1%88%D0%BA%D0%BE" title="Аўрам Гершко – Belarusian" lang="be" hreflang="be" data-title="Аўрам Гершко" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%90%D0%B2%D1%80%D0%B0%D0%BC_%D0%A5%D0%B5%D1%80%D1%88%D0%BA%D0%BE" title="Аврам Хершко – Bulgarian" lang="bg" hreflang="bg" data-title="Аврам Хершко" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Catalan" lang="ca" hreflang="ca" data-title="Avram Hershko" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Avram_Her%C5%A1ko" title="Avram Herško – Czech" lang="cs" hreflang="cs" data-title="Avram Herško" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Danish" lang="da" hreflang="da" data-title="Avram Hershko" data-language-autonym="Dansk" data-language-local-name="Danish" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – German" lang="de" hreflang="de" data-title="Avram Hershko" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Spanish" lang="es" hreflang="es" data-title="Avram Hershko" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Esperanto" lang="eo" hreflang="eo" data-title="Avram Hershko" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Basque" lang="eu" hreflang="eu" data-title="Avram Hershko" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%A7%D9%81%D8%B1%D8%A7%D9%85_%D9%87%D8%B1%D8%B4%DA%A9%D9%88" title="افرام هرشکو – Persian" lang="fa" hreflang="fa" data-title="افرام هرشکو" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – French" lang="fr" hreflang="fr" data-title="Avram Hershko" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Irish" lang="ga" hreflang="ga" data-title="Avram Hershko" data-language-autonym="Gaeilge" data-language-local-name="Irish" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-gd mw-list-item"><a href="https://gd.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Scottish Gaelic" lang="gd" hreflang="gd" data-title="Avram Hershko" data-language-autonym="Gàidhlig" data-language-local-name="Scottish Gaelic" class="interlanguage-link-target"><span>Gàidhlig</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Galician" lang="gl" hreflang="gl" data-title="Avram Hershko" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-hak mw-list-item"><a href="https://hak.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Hakka Chinese" lang="hak" hreflang="hak" data-title="Avram Hershko" data-language-autonym="客家語 / Hak-kâ-ngî" data-language-local-name="Hakka Chinese" class="interlanguage-link-target"><span>客家語 / Hak-kâ-ngî</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%95%84%EB%B8%8C%EB%9E%8C_%ED%97%A4%EB%A5%B4%EC%8A%88%EC%BD%94" title="아브람 헤르슈코 – Korean" lang="ko" hreflang="ko" data-title="아브람 헤르슈코" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%8F%E0%A4%B5%E0%A4%B0%E0%A4%AE_%E0%A4%B9%E0%A5%87%E0%A4%B0%E0%A5%8D%E0%A4%B6%E0%A4%95%E0%A5%8B" title="एवरम हेर्शको – Hindi" lang="hi" hreflang="hi" data-title="एवरम हेर्शको" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Ido" lang="io" hreflang="io" data-title="Avram Hershko" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Indonesian" lang="id" hreflang="id" data-title="Avram Hershko" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Italian" lang="it" hreflang="it" data-title="Avram Hershko" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%90%D7%91%D7%A8%D7%94%D7%9D_%D7%94%D7%A8%D7%A9%D7%A7%D7%95" title="אברהם הרשקו – Hebrew" lang="he" hreflang="he" data-title="אברהם הרשקו" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Swahili" lang="sw" hreflang="sw" data-title="Avram Hershko" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Latin" lang="la" hreflang="la" data-title="Avram Hershko" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Avram_Hersko" title="Avram Hersko – Hungarian" lang="hu" hreflang="hu" data-title="Avram Hersko" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%A3%D9%81%D8%B1%D8%A7%D9%85_%D9%87%D9%8A%D8%B1%D8%B4%D9%83%D9%88" title="أفرام هيرشكو – Egyptian Arabic" lang="arz" hreflang="arz" data-title="أفرام هيرشكو" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D8%A7%D9%81%D8%B1%D8%A7%D9%85_%D9%87%D8%B1%D8%B4%DA%A9%D9%88" title="افرام هرشکو – Mazanderani" lang="mzn" hreflang="mzn" data-title="افرام هرشکو" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Malay" lang="ms" hreflang="ms" data-title="Avram Hershko" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Dutch" lang="nl" hreflang="nl" data-title="Avram Hershko" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%A2%E3%83%96%E3%83%A9%E3%83%A0%E3%83%BB%E3%83%8F%E3%83%BC%E3%82%B7%E3%83%A5%E3%82%B3" title="アブラム・ハーシュコ – Japanese" lang="ja" hreflang="ja" data-title="アブラム・ハーシュコ" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Avram Hershko" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Occitan" lang="oc" hreflang="oc" data-title="Avram Hershko" data-language-autonym="Occitan" data-language-local-name="Occitan" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D8%A7%D9%88%D8%B1%D8%A7%D9%85_%DB%81%D8%B1%D8%B4%DA%A9%D9%88" title="اورام ہرشکو – Western Punjabi" lang="pnb" hreflang="pnb" data-title="اورام ہرشکو" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Low German" lang="nds" hreflang="nds" data-title="Avram Hershko" data-language-autonym="Plattdüütsch" data-language-local-name="Low German" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Awram_Herszko" title="Awram Herszko – Polish" lang="pl" hreflang="pl" data-title="Awram Herszko" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Portuguese" lang="pt" hreflang="pt" data-title="Avram Hershko" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Romanian" lang="ro" hreflang="ro" data-title="Avram Hershko" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%93%D0%B5%D1%80%D1%88%D0%BA%D0%BE,_%D0%90%D0%B2%D1%80%D0%B0%D0%BC" title="Гершко, Аврам – Russian" lang="ru" hreflang="ru" data-title="Гершко, Аврам" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Simple English" lang="en-simple" hreflang="en-simple" data-title="Avram Hershko" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Avram_Her%C5%A1ko" title="Avram Herško – Slovak" lang="sk" hreflang="sk" data-title="Avram Herško" data-language-autonym="Slovenčina" data-language-local-name="Slovak" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/Avram_Her%C5%A1ko" title="Avram Herško – Serbian" lang="sr" hreflang="sr" data-title="Avram Herško" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Finnish" lang="fi" hreflang="fi" data-title="Avram Hershko" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Avram_Hershko" title="Avram Hershko – Swedish" lang="sv" hreflang="sv" data-title="Avram Hershko" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a 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dir="ltr"><p class="mw-empty-elt"> </p> <div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Hungarian-Israeli biochemist</div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">The native form of this <a href="/wiki/Personal_name" title="Personal name">personal name</a> is <i>Herskó Ferenc Ábrahám</i>. This article uses <a href="/wiki/Personal_name#Western_name_order" title="Personal name">Western name order</a> when mentioning individuals.</div> <style data-mw-deduplicate="TemplateStyles:r1257001546">.mw-parser-output .infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox-full-data:not(.notheme)>div:not(.notheme)[style]{background:#1f1f23!important;color:#f8f9fa}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox-full-data:not(.notheme) div:not(.notheme){background:#1f1f23!important;color:#f8f9fa}}@media(min-width:640px){body.skin--responsive .mw-parser-output .infobox-table{display:table!important}body.skin--responsive .mw-parser-output .infobox-table>caption{display:table-caption!important}body.skin--responsive .mw-parser-output .infobox-table>tbody{display:table-row-group}body.skin--responsive .mw-parser-output .infobox-table tr{display:table-row!important}body.skin--responsive .mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox biography vcard"><tbody><tr><th colspan="2" class="infobox-above" style="font-size:125%;"><div class="fn">Avram Hershko</div></th></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Flickr_-_Government_Press_Office_(GPO)_-_Nobel_Laureate_Avram_Hershko.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Flickr_-_Government_Press_Office_%28GPO%29_-_Nobel_Laureate_Avram_Hershko.jpg/220px-Flickr_-_Government_Press_Office_%28GPO%29_-_Nobel_Laureate_Avram_Hershko.jpg" decoding="async" width="220" height="286" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Flickr_-_Government_Press_Office_%28GPO%29_-_Nobel_Laureate_Avram_Hershko.jpg/330px-Flickr_-_Government_Press_Office_%28GPO%29_-_Nobel_Laureate_Avram_Hershko.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/df/Flickr_-_Government_Press_Office_%28GPO%29_-_Nobel_Laureate_Avram_Hershko.jpg/440px-Flickr_-_Government_Press_Office_%28GPO%29_-_Nobel_Laureate_Avram_Hershko.jpg 2x" data-file-width="1880" data-file-height="2444" /></a></span><div class="infobox-caption">Hershko in 1994</div></td></tr><tr><th scope="row" class="infobox-label">Born</th><td class="infobox-data"><div style="display:inline" class="nickname">Herskó Ferenc Ábrahám</div><br /><span style="display:none"> (<span class="bday">1937-12-31</span>) </span>December 31, 1937<span class="noprint ForceAgeToShow"> (age 86)</span><br /><div style="display:inline" class="birthplace"><a href="/wiki/Karcag" title="Karcag">Karcag</a>, <a href="/wiki/Kingdom_of_Hungary_(1920%E2%80%931946)" title="Kingdom of Hungary (1920–1946)">Hungary</a></div></td></tr><tr><th scope="row" class="infobox-label">Nationality</th><td class="infobox-data category"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol ol,.mw-parser-output .hlist ol ul,.mw-parser-output .hlist ul dl,.mw-parser-output .hlist ul ol,.mw-parser-output .hlist ul ul{display:inline}.mw-parser-output .hlist .mw-empty-li{display:none}.mw-parser-output .hlist dt::after{content:": "}.mw-parser-output .hlist dd::after,.mw-parser-output .hlist li::after{content:" · ";font-weight:bold}.mw-parser-output .hlist dd:last-child::after,.mw-parser-output .hlist dt:last-child::after,.mw-parser-output .hlist li:last-child::after{content:none}.mw-parser-output .hlist dd dd:first-child::before,.mw-parser-output .hlist dd dt:first-child::before,.mw-parser-output .hlist dd li:first-child::before,.mw-parser-output .hlist dt dd:first-child::before,.mw-parser-output .hlist dt dt:first-child::before,.mw-parser-output .hlist dt li:first-child::before,.mw-parser-output .hlist li dd:first-child::before,.mw-parser-output .hlist li dt:first-child::before,.mw-parser-output .hlist li li:first-child::before{content:" (";font-weight:normal}.mw-parser-output .hlist dd dd:last-child::after,.mw-parser-output .hlist dd dt:last-child::after,.mw-parser-output .hlist dd li:last-child::after,.mw-parser-output .hlist dt dd:last-child::after,.mw-parser-output .hlist dt dt:last-child::after,.mw-parser-output .hlist dt li:last-child::after,.mw-parser-output .hlist li dd:last-child::after,.mw-parser-output .hlist li dt:last-child::after,.mw-parser-output .hlist li li:last-child::after{content:")";font-weight:normal}.mw-parser-output .hlist ol{counter-reset:listitem}.mw-parser-output .hlist ol>li{counter-increment:listitem}.mw-parser-output .hlist ol>li::before{content:" "counter(listitem)"\a0 "}.mw-parser-output .hlist dd ol>li:first-child::before,.mw-parser-output .hlist dt ol>li:first-child::before,.mw-parser-output .hlist li ol>li:first-child::before{content:" ("counter(listitem)"\a0 "}</style><div class="hlist"><ul><li>Hungary</li><li>Israel</li></ul></div></td></tr><tr><th scope="row" class="infobox-label">Alma mater</th><td class="infobox-data"><a href="/wiki/Hebrew_University_of_Jerusalem" title="Hebrew University of Jerusalem">Hebrew University of Jerusalem</a></td></tr><tr><th scope="row" class="infobox-label">Known for</th><td class="infobox-data"><a href="/wiki/Ubiquitin" title="Ubiquitin">Ubiquitin</a>-mediated <a href="/wiki/Proteolysis#Protein_degradation" title="Proteolysis">protein degradation</a></td></tr><tr><th scope="row" class="infobox-label">Spouse</th><td class="infobox-data"><style data-mw-deduplicate="TemplateStyles:r1151524712">.mw-parser-output .marriage-line-margin2px{line-height:0;margin-bottom:-2px}.mw-parser-output .marriage-line-margin3px{line-height:0;margin-bottom:-3px}.mw-parser-output .marriage-display-ws{display:inline;white-space:nowrap}</style> <div class="marriage-display-ws"><div style="display:inline-block;line-height:normal;">Judith Leibowitz</div> <div style="display:inline-block;"></div>(<abbr title="married">m.</abbr> 1963)<wbr /></div></td></tr><tr><th scope="row" class="infobox-label">Children</th><td class="infobox-data">3</td></tr><tr><th scope="row" class="infobox-label">Awards</th><td class="infobox-data"><a href="/wiki/Weizmann_Prize" class="mw-redirect" title="Weizmann Prize">Weizmann Prize</a> (1987)<br /><a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Nobel Prize in Chemistry</a> (2004)</td></tr><tr><td colspan="2" class="infobox-full-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1257001546"><b>Scientific career</b></td></tr><tr><th scope="row" class="infobox-label">Fields</th><td class="infobox-data category"><a href="/wiki/Chemistry" title="Chemistry">Chemistry</a></td></tr><tr><th scope="row" class="infobox-label">Institutions</th><td class="infobox-data"><a href="/wiki/Technion_%E2%80%93_Israel_Institute_of_Technology" title="Technion – Israel Institute of Technology">Technion</a>, Haifa</td></tr><tr style="display:none"><td colspan="2"> </td></tr></tbody></table> <p><b>Avram Hershko</b> (<a href="/wiki/Hebrew_language" title="Hebrew language">Hebrew</a>: <span lang="he" dir="rtl">אברהם הרשקו</span>, <small><a href="/wiki/Romanization_of_Hebrew" title="Romanization of Hebrew">romanized</a>: </small><span title="Hebrew-language romanization"><i lang="he-Latn">Avraham Hershko</i></span>, <a href="/wiki/Hungarian_language" title="Hungarian language">Hungarian</a>: <i lang="hu">Herskó Ferenc Ábrahám</i>;<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> born December 31, 1937) is a Hungarian-Israeli <a href="/wiki/Biology" title="Biology">biochemist</a> who received the <a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Nobel Prize in Chemistry</a> in 2004. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Biography">Biography</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Avram_Hershko&action=edit&section=1" title="Edit section: Biography"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>He was born <b>Herskó Ferenc</b> in <a href="/wiki/Karcag" title="Karcag">Karcag</a>, <a href="/wiki/Kingdom_of_Hungary_(1920%E2%80%931946)" title="Kingdom of Hungary (1920–1946)">Hungary</a>, into a <a href="/wiki/Jews" title="Jews">Jewish</a> family,<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> the son of Shoshana/Margit 'Manci' (née Wulc) and Moshe Hershko, both teachers.<sup id="cite_ref-Nobel_3-0" class="reference"><a href="#cite_note-Nobel-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> During the <a href="/wiki/Second_World_War" class="mw-redirect" title="Second World War">Second World War</a>, his father was forced into <a href="/wiki/Labour_service_in_Hungary_during_World_War_II" title="Labour service in Hungary during World War II">labor service in the Hungarian army</a> and then taken as a prisoner by the <a href="/wiki/Soviet_Army" title="Soviet Army">Soviet Army</a>. For years, Avram's family didn't known anything about what had happened to his father. Avram, his mother and older brother Chaim/Laszlo 'Laci' were put in a <a href="/wiki/Ghetto" title="Ghetto">ghetto</a> in <a href="/wiki/Szolnok" title="Szolnok">Szolnok</a>. During the final days of the ghetto, most Jews were sent to be murdered in <a href="/wiki/Auschwitz_concentration_camp" title="Auschwitz concentration camp">Auschwitz</a>, but Avram and his family managed to board trains that took them to a <a href="/wiki/Nazi_concentration_camps" title="Nazi concentration camps">concentration camp</a> in <a href="/wiki/Austria_under_National_Socialism" class="mw-redirect" title="Austria under National Socialism">Austria</a>, where they were forced into labor until the end of the war. Avram and his mother and brother survived the war and returned to their home. His father returned as well, 4 years after they had last seen him.<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p><p>Hershko and his family emigrated to Israel in 1950 and settled in <a href="/wiki/Jerusalem" title="Jerusalem">Jerusalem</a>. He received his MD in 1965 and his PhD in 1969 from the <a href="/wiki/Hebrew_University_of_Jerusalem" title="Hebrew University of Jerusalem">Hebrew University of Jerusalem</a>-<a href="/wiki/Hadassah_Medical_Center" title="Hadassah Medical Center">Hadassah Medical Center</a>. He was a <a href="/wiki/Postdoctoral_researcher" title="Postdoctoral researcher">postdoctoral scholar</a> at the <a href="/wiki/University_of_California,_San_Francisco" title="University of California, San Francisco">University of California, San Francisco</a>. He is currently a Distinguished Professor at the Rappaport Faculty of Medicine at the <a href="/wiki/Technion_%E2%80%93_Israel_Institute_of_Technology" title="Technion – Israel Institute of Technology">Technion</a> in <a href="/wiki/Haifa" title="Haifa">Haifa</a> and a Distinguished Adjunct Professor at the <a href="/wiki/New_York_University_Grossman_School_of_Medicine" title="New York University Grossman School of Medicine">New York University Grossman School of Medicine</a>. </p><p>Along with <a href="/wiki/Aaron_Ciechanover" title="Aaron Ciechanover">Aaron Ciechanover</a> and <a href="/wiki/Irwin_Rose" title="Irwin Rose">Irwin Rose</a>, he was awarded the 2004 <a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Nobel Prize in Chemistry</a> for the discovery of <a href="/wiki/Ubiquitin" title="Ubiquitin">ubiquitin</a>-mediated <a href="/wiki/Proteolysis#Protein_degradation" title="Proteolysis">protein degradation</a>. The ubiquitin-<a href="/wiki/Proteasome" title="Proteasome">proteasome</a> system has a critical role in maintaining the homeostasis of cells and is believed to be involved in the development and progression of diseases such as cancer, muscular and neurological diseases, and immune and inflammatory responses. </p><p>His contributions to science directly helped cure one of his long-time friends of cancer.<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Honours_and_awards">Honours and awards</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Avram_Hershko&action=edit&section=2" title="Edit section: Honours and awards"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li>1987 – <a href="/wiki/Weizmann_Prize_for_Research_in_the_Exact_Sciences" title="Weizmann Prize for Research in the Exact Sciences">Weizmann Prize for Sciences</a><sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (August 2021)">citation needed</span></a></i>]</sup></li> <li>1994 – <a href="/wiki/Israel_Prize" title="Israel Prize">Israel Prize</a> in Biochemistry<sup id="cite_ref-prize_6-0" class="reference"><a href="#cite_note-prize-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup></li> <li>1999 – <a href="/wiki/Canada_Gairdner_International_Award" title="Canada Gairdner International Award">Canada Gairdner International Award</a> (with <a href="/wiki/Alexander_Varshavsky" title="Alexander Varshavsky">Alexander Varshavsky</a>)</li> <li>2000 – <a href="/wiki/Albert_Lasker_Award_for_Basic_Medical_Research" title="Albert Lasker Award for Basic Medical Research">Albert Lasker Award for Basic Medical Research</a> (with <a href="/wiki/Aaron_Ciechanover" title="Aaron Ciechanover">Aaron Ciechanover</a> and Varshavsky)</li> <li>2000 – <a href="/wiki/Alfred_P._Sloan_Jr._Prize" title="Alfred P. Sloan Jr. Prize">Alfred P. Sloan Jr. Prize</a> (with Varshavsky)</li> <li>2000 – Member, <a href="/wiki/Israel_Academy_of_Sciences_and_Humanities" title="Israel Academy of Sciences and Humanities">Israel Academy of Sciences and Humanities</a></li> <li>2001 – <a href="/wiki/Louisa_Gross_Horwitz_Prize" title="Louisa Gross Horwitz Prize">Louisa Gross Horwitz Prize</a> from <a href="/wiki/Columbia_University" title="Columbia University">Columbia University</a> (with Varshavsky)</li> <li>2001 – <a href="/wiki/Massry_Prize" title="Massry Prize">Massry Prize</a> from the <a href="/wiki/Keck_School_of_Medicine_of_USC" title="Keck School of Medicine of USC">Keck School of Medicine of USC</a>, <a href="/wiki/University_of_Southern_California" title="University of Southern California">University of Southern California</a> (with Varshavsky)</li> <li>2001 – <a href="/wiki/Wolf_Prize_in_Medicine" title="Wolf Prize in Medicine">Wolf Prize in Medicine</a> (with Varshavsky) for "the discovery of the <a href="/wiki/Ubiquitin" title="Ubiquitin">ubiquitin</a> system of intracellular <a href="/wiki/Proteolysis#Protein_degradation" title="Proteolysis">protein degradation</a> and the crucial functions of this system in cellular regulation."<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup></li> <li>2002 – <a href="/wiki/The_EMET_Prize_for_Art,_Science_and_Culture" class="mw-redirect" title="The EMET Prize for Art, Science and Culture">The EMET Prize for Art, Science and Culture</a> in the category of Life Sciences (with Ciechanover and <a href="/wiki/Leo_Sachs" title="Leo Sachs">Leo Sachs</a>)</li> <li>2002 – <a href="/wiki/E.B._Wilson_Medal" title="E.B. Wilson Medal">E.B. Wilson Medal</a> (with Varshavsky)</li> <li>2003 – Foreign Associate, <a href="/wiki/National_Academy_of_Sciences" title="National Academy of Sciences">National Academy of Sciences</a>, USA</li> <li>2004 – Nobel Prize in Chemistry for his discovery with Ciechanover and <a href="/wiki/Irwin_Rose" title="Irwin Rose">Irwin Rose</a>, of ubiquitin-mediated protein degradation<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup></li> <li>2005 – Elected to the <a href="/wiki/American_Philosophical_Society" title="American Philosophical Society">American Philosophical Society</a><sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup></li></ul> <div class="mw-heading mw-heading2"><h2 id="Publications">Publications</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Avram_Hershko&action=edit&section=3" title="Edit section: Publications"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFHershkoCiechanoverHellerHaas1980" class="citation journal cs1">Hershko, A.; Ciechanover, A.; Heller, H.; Haas, A. L.; Rose, I. A. (April 1, 1980). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC348591">"Proposed role of ATP in protein breakdown: conjugation of protein with multiple chains of the polypeptide of ATP-dependent proteolysis"</a>. <i>Proceedings of the National Academy of Sciences</i>. <b>77</b> (4): 1783–1786. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1980PNAS...77.1783H">1980PNAS...77.1783H</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1073%2Fpnas.77.4.1783">10.1073/pnas.77.4.1783</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0027-8424">0027-8424</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC348591">348591</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/6990414">6990414</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Proceedings+of+the+National+Academy+of+Sciences&rft.atitle=Proposed+role+of+ATP+in+protein+breakdown%3A+conjugation+of+protein+with+multiple+chains+of+the+polypeptide+of+ATP-dependent+proteolysis.&rft.volume=77&rft.issue=4&rft.pages=1783-1786&rft.date=1980-04-01&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC348591%23id-name%3DPMC&rft_id=info%3Abibcode%2F1980PNAS...77.1783H&rft_id=info%3Apmid%2F6990414&rft_id=info%3Adoi%2F10.1073%2Fpnas.77.4.1783&rft.issn=0027-8424&rft.aulast=Hershko&rft.aufirst=A.&rft.au=Ciechanover%2C+A.&rft.au=Heller%2C+H.&rft.au=Haas%2C+A.+L.&rft.au=Rose%2C+I.+A.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC348591&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAvram+Hershko" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHershkoHellerEliasCiechanover1983" class="citation journal cs1">Hershko, A; Heller, H; Elias, S; Ciechanover, A (July 10, 1983). <a rel="nofollow" class="external text" href="https://doi.org/10.1016%2FS0021-9258%2820%2982050-X">"Components of ubiquitin-protein ligase system. Resolution, affinity purification, and role in protein breakdown"</a>. <i>The Journal of Biological Chemistry</i>. <b>258</b> (13): 8206–14. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1016%2FS0021-9258%2820%2982050-X">10.1016/S0021-9258(20)82050-X</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0021-9258">0021-9258</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/6305978">6305978</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+Journal+of+Biological+Chemistry&rft.atitle=Components+of+ubiquitin-protein+ligase+system.+Resolution%2C+affinity+purification%2C+and+role+in+protein+breakdown.&rft.volume=258&rft.issue=13&rft.pages=8206-14&rft.date=1983-07-10&rft.issn=0021-9258&rft_id=info%3Apmid%2F6305978&rft_id=info%3Adoi%2F10.1016%2FS0021-9258%2820%2982050-X&rft.aulast=Hershko&rft.aufirst=A&rft.au=Heller%2C+H&rft.au=Elias%2C+S&rft.au=Ciechanover%2C+A&rft_id=https%3A%2F%2Fdoi.org%2F10.1016%252FS0021-9258%252820%252982050-X&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAvram+Hershko" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHershkoLeshinskyGanothHeller1984" class="citation journal cs1">Hershko, A.; Leshinsky, E.; Ganoth, D.; Heller, H. (March 1, 1984). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC344969">"ATP-dependent degradation of ubiquitin-protein conjugates"</a>. <i>Proceedings of the National Academy of Sciences</i>. <b>81</b> (6): 1619–1623. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1984PNAS...81.1619H">1984PNAS...81.1619H</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1073%2Fpnas.81.6.1619">10.1073/pnas.81.6.1619</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0027-8424">0027-8424</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC344969">344969</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/6324208">6324208</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Proceedings+of+the+National+Academy+of+Sciences&rft.atitle=ATP-dependent+degradation+of+ubiquitin-protein+conjugates.&rft.volume=81&rft.issue=6&rft.pages=1619-1623&rft.date=1984-03-01&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC344969%23id-name%3DPMC&rft_id=info%3Abibcode%2F1984PNAS...81.1619H&rft_id=info%3Apmid%2F6324208&rft_id=info%3Adoi%2F10.1073%2Fpnas.81.6.1619&rft.issn=0027-8424&rft.aulast=Hershko&rft.aufirst=A.&rft.au=Leshinsky%2C+E.&rft.au=Ganoth%2C+D.&rft.au=Heller%2C+H.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC344969&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAvram+Hershko" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHershkoHellerEytanReiss1986" class="citation journal cs1">Hershko, A; Heller, H; Eytan, E; Reiss, Y (1986). <a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fs0021-9258%2818%2967192-3">"The protein substrate binding site of the ubiquitin-protein ligase system"</a>. <i>Journal of Biological Chemistry</i>. <b>261</b> (26). Elsevier BV: 11992–11999. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fs0021-9258%2818%2967192-3">10.1016/s0021-9258(18)67192-3</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0021-9258">0021-9258</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/3017957">3017957</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Journal+of+Biological+Chemistry&rft.atitle=The+protein+substrate+binding+site+of+the+ubiquitin-protein+ligase+system.&rft.volume=261&rft.issue=26&rft.pages=11992-11999&rft.date=1986&rft.issn=0021-9258&rft_id=info%3Apmid%2F3017957&rft_id=info%3Adoi%2F10.1016%2Fs0021-9258%2818%2967192-3&rft.aulast=Hershko&rft.aufirst=A&rft.au=Heller%2C+H&rft.au=Eytan%2C+E&rft.au=Reiss%2C+Y&rft_id=https%3A%2F%2Fdoi.org%2F10.1016%252Fs0021-9258%252818%252967192-3&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAvram+Hershko" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFGanothLeshinskyEytanHershko1988" class="citation journal cs1">Ganoth, D; Leshinsky, E; Eytan, E; Hershko, A (September 5, 1988). <a rel="nofollow" class="external text" href="https://doi.org/10.1016%2FS0021-9258%2818%2937771-8">"A multicomponent system that degrades proteins conjugated to ubiquitin. Resolution of factors and evidence for ATP-dependent complex formation"</a>. <i>The Journal of Biological Chemistry</i>. <b>263</b> (25): 12412–9. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1016%2FS0021-9258%2818%2937771-8">10.1016/S0021-9258(18)37771-8</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0021-9258">0021-9258</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/2842333">2842333</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+Journal+of+Biological+Chemistry&rft.atitle=A+multicomponent+system+that+degrades+proteins+conjugated+to+ubiquitin.+Resolution+of+factors+and+evidence+for+ATP-dependent+complex+formation.&rft.volume=263&rft.issue=25&rft.pages=12412-9&rft.date=1988-09-05&rft.issn=0021-9258&rft_id=info%3Apmid%2F2842333&rft_id=info%3Adoi%2F10.1016%2FS0021-9258%2818%2937771-8&rft.aulast=Ganoth&rft.aufirst=D&rft.au=Leshinsky%2C+E&rft.au=Eytan%2C+E&rft.au=Hershko%2C+A&rft_id=https%3A%2F%2Fdoi.org%2F10.1016%252FS0021-9258%252818%252937771-8&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAvram+Hershko" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFSudakinGanothDahanHeller1995" class="citation journal cs1">Sudakin, V; Ganoth, D; Dahan, A; Heller, H; Hershko, J; Luca, F C; Ruderman, J V; Hershko, A (1995). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC275828">"The cyclosome, a large complex containing cyclin-selective ubiquitin ligase activity, targets cyclins for destruction at the end of mitosis"</a>. <i>Molecular Biology of the Cell</i>. <b>6</b> (2). American Society for Cell Biology (ASCB): 185–197. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1091%2Fmbc.6.2.185">10.1091/mbc.6.2.185</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/1059-1524">1059-1524</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC275828">275828</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/7787245">7787245</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Molecular+Biology+of+the+Cell&rft.atitle=The+cyclosome%2C+a+large+complex+containing+cyclin-selective+ubiquitin+ligase+activity%2C+targets+cyclins+for+destruction+at+the+end+of+mitosis.&rft.volume=6&rft.issue=2&rft.pages=185-197&rft.date=1995&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC275828%23id-name%3DPMC&rft.issn=1059-1524&rft_id=info%3Apmid%2F7787245&rft_id=info%3Adoi%2F10.1091%2Fmbc.6.2.185&rft.aulast=Sudakin&rft.aufirst=V&rft.au=Ganoth%2C+D&rft.au=Dahan%2C+A&rft.au=Heller%2C+H&rft.au=Hershko%2C+J&rft.au=Luca%2C+F+C&rft.au=Ruderman%2C+J+V&rft.au=Hershko%2C+A&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC275828&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAvram+Hershko" class="Z3988"></span></li></ul> <div class="mw-heading mw-heading2"><h2 id="Involvement_with_biotechnology">Involvement with biotechnology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Avram_Hershko&action=edit&section=4" title="Edit section: Involvement with biotechnology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Hershko serves on the Scientific Advisory Board of <a href="/wiki/Oramed" title="Oramed">Oramed Pharmaceuticals</a>.<sup id="cite_ref-Oramed_Pharmaceuticals_2020_b459_10-0" class="reference"><a href="#cite_note-Oramed_Pharmaceuticals_2020_b459-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Avram_Hershko&action=edit&section=5" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239009302">.mw-parser-output .portalbox{padding:0;margin:0.5em 0;display:table;box-sizing:border-box;max-width:175px;list-style:none}.mw-parser-output .portalborder{border:1px solid var(--border-color-base,#a2a9b1);padding:0.1em;background:var(--background-color-neutral-subtle,#f8f9fa)}.mw-parser-output .portalbox-entry{display:table-row;font-size:85%;line-height:110%;height:1.9em;font-style:italic;font-weight:bold}.mw-parser-output .portalbox-image{display:table-cell;padding:0.2em;vertical-align:middle;text-align:center}.mw-parser-output .portalbox-link{display:table-cell;padding:0.2em 0.2em 0.2em 0.3em;vertical-align:middle}@media(min-width:720px){.mw-parser-output .portalleft{clear:left;float:left;margin:0.5em 1em 0.5em 0}.mw-parser-output .portalright{clear:right;float:right;margin:0.5em 0 0.5em 1em}}</style><ul role="navigation" aria-label="Portals" class="noprint portalbox portalborder portalright"> <li class="portalbox-entry"><span class="portalbox-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/6/69/P_vip.svg/28px-P_vip.svg.png" decoding="async" width="28" height="28" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/6/69/P_vip.svg/41px-P_vip.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/6/69/P_vip.svg/55px-P_vip.svg.png 2x" data-file-width="1911" data-file-height="1944" /></span></span></span><span class="portalbox-link"><a href="/wiki/Portal:Biography" title="Portal:Biography">Biography portal</a></span></li><li class="portalbox-entry"><span class="portalbox-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/e/ed/Papapishu-Lab-icon-6.svg/28px-Papapishu-Lab-icon-6.svg.png" decoding="async" width="28" height="28" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/ed/Papapishu-Lab-icon-6.svg/42px-Papapishu-Lab-icon-6.svg.png 1.5x, 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portal</a></span></li></ul> <ul><li><a href="/wiki/List_of_Israel_Prize_recipients" title="List of Israel Prize recipients">List of Israel Prize recipients</a></li> <li><a href="/wiki/List_of_Israeli_Nobel_laureates" title="List of Israeli Nobel laureates">List of Israeli Nobel laureates</a></li> <li><a href="/wiki/List_of_Jewish_Nobel_laureates" title="List of Jewish Nobel laureates">List of Jewish Nobel laureates</a></li> <li><a href="/wiki/Science_and_technology_in_Israel" title="Science and technology in Israel">Science and technology in Israel</a></li> <li><a href="/wiki/Raz_Hershko" title="Raz Hershko">Raz Hershko</a> (born 1998), Israeli European champion and Olympic judoka</li></ul> <div style="clear:both;" class=""></div> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Avram_Hershko&action=edit&section=6" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20151006113721/http://mta.hu/articles/hungarys_nobel_prize_winners__25577">"Hungary's Nobel Prize Winners"</a>. <i>www.mta.hu</i>. Archived from <a rel="nofollow" class="external text" href="http://mta.hu/articles/hungarys_nobel_prize_winners__25577">the original</a> on October 6, 2015<span class="reference-accessdate">. Retrieved <span class="nowrap">September 22,</span> 2021</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=www.mta.hu&rft.atitle=Hungary%27s+Nobel+Prize+Winners&rft_id=http%3A%2F%2Fmta.hu%2Farticles%2Fhungarys_nobel_prize_winners__25577&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAvram+Hershko" class="Z3988"></span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-2">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFJINFO" class="citation web cs1">JINFO. <a rel="nofollow" class="external text" href="https://www.jinfo.org/Nobels_Chemistry.html">"Jewish Nobel Prize Winners in Chemistry"</a>. <i>www.jinfo.org</i><span class="reference-accessdate">. Retrieved <span class="nowrap">March 30,</span> 2023</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=www.jinfo.org&rft.atitle=Jewish+Nobel+Prize+Winners+in+Chemistry&rft.au=JINFO&rft_id=https%3A%2F%2Fwww.jinfo.org%2FNobels_Chemistry.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAvram+Hershko" class="Z3988"></span></span> </li> <li id="cite_note-Nobel-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-Nobel_3-0">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://www.nobelprize.org/laureate/780">Avram Hershko</a> on Nobelprize.org <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q232302#P8024" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span> including the Nobel Lecture <i>The Ubiquitin System for Protein Degradation and some of its Roles in the Control of the Cell Division Cycle</i></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.yadvashem.org/he/remembrance/survivors/hershko.html">"אברהם הרשקו"</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=%D7%90%D7%91%D7%A8%D7%94%D7%9D+%D7%94%D7%A8%D7%A9%D7%A7%D7%95&rft_id=https%3A%2F%2Fwww.yadvashem.org%2Fhe%2Fremembrance%2Fsurvivors%2Fhershko.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAvram+Hershko" class="Z3988"></span></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-5">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFFriedman2011" class="citation web cs1">Friedman, Sally (September 13, 2011). <a rel="nofollow" class="external text" href="https://archive.today/20120913041804/http://www.phillyburbs.com/news/local/burlington_county_times_news/nobel-prize-winner-s-discovery-helps-save-longtime-physician-friend/article_9ad53d9b-22f5-5fec-8956-92a7aae6a009.html">"Nobel Prize winner's discovery helps save longtime physician friend"</a>. <i><a href="/wiki/Burlington_County_Times" title="Burlington County Times">Burlington County Times</a></i>. phillyBurbs.com. Archived from <a rel="nofollow" class="external text" href="http://www.phillyburbs.com/news/local/burlington_county_times_news/nobel-prize-winner-s-discovery-helps-save-longtime-physician-friend/article_9ad53d9b-22f5-5fec-8956-92a7aae6a009.html">the original</a> on September 13, 2012<span class="reference-accessdate">. Retrieved <span class="nowrap">May 15,</span> 2012</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Burlington+County+Times&rft.atitle=Nobel+Prize+winner%27s+discovery+helps+save+longtime+physician+friend&rft.date=2011-09-13&rft.aulast=Friedman&rft.aufirst=Sally&rft_id=http%3A%2F%2Fwww.phillyburbs.com%2Fnews%2Flocal%2Fburlington_county_times_news%2Fnobel-prize-winner-s-discovery-helps-save-longtime-physician-friend%2Farticle_9ad53d9b-22f5-5fec-8956-92a7aae6a009.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAvram+Hershko" class="Z3988"></span></span> </li> <li id="cite_note-prize-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-prize_6-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1 cs1-prop-foreign-lang-source"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20081227153428/http://cms.education.gov.il/EducationCMS/Units/PrasIsrael/TashnagTashsab/TASNAG_TASNAT_Rikuz.htm?DictionaryKey=Tashnad">"Israel Prize Official Site – Recipients in 1994"</a> (in Hebrew). Archived from <a rel="nofollow" class="external text" href="http://cms.education.gov.il/EducationCMS/Units/PrasIsrael/TashnagTashsab/TASNAG_TASNAT_Rikuz.htm?DictionaryKey=Tashnad">the original</a> on December 27, 2008.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Israel+Prize+Official+Site+%E2%80%93+Recipients+in+1994&rft_id=http%3A%2F%2Fcms.education.gov.il%2FEducationCMS%2FUnits%2FPrasIsrael%2FTashnagTashsab%2FTASNAG_TASNAT_Rikuz.htm%3FDictionaryKey%3DTashnad&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAvram+Hershko" class="Z3988"></span></span> </li> <li id="cite_note-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-7">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20090226015735/http://www.wolffund.org.il/cat.asp?id=24&cat_title=MEDICINE">"Wolf Prize Recipients in Medicine"</a>. Archived from <a rel="nofollow" class="external text" href="http://www.wolffund.org.il/cat.asp?id=24&cat_title=MEDICINE">the original</a> on February 26, 2009.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Wolf+Prize+Recipients+in+Medicine&rft_id=http%3A%2F%2Fwww.wolffund.org.il%2Fcat.asp%3Fid%3D24%26cat_title%3DMEDICINE&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAvram+Hershko" class="Z3988"></span></span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-8">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFIddo_Genuth2005" class="citation web cs1">Iddo Genuth (July 29, 2005). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20051219160839/http://www.isracast.com/tech_news/101004_tech.htm">"Two Israeli Scientists Have Won The Nobel Prize In Chemistry"</a>. <i>isracast.com</i>. Archived from <a rel="nofollow" class="external text" href="http://www.isracast.com/tech_news/101004_tech.htm">the original</a> on December 19, 2005.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=isracast.com&rft.atitle=Two+Israeli+Scientists+Have+Won+The+Nobel+Prize+In+Chemistry&rft.date=2005-07-29&rft.au=Iddo+Genuth&rft_id=http%3A%2F%2Fwww.isracast.com%2Ftech_news%2F101004_tech.htm&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAvram+Hershko" class="Z3988"></span></span> </li> <li id="cite_note-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-9">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://search.amphilsoc.org/memhist/search?creator=Avram+Hershko&title=&subject=&subdiv=&mem=&year=&year-max=&dead=&keyword=&smode=advanced">"APS Member History"</a>. <i>search.amphilsoc.org</i><span class="reference-accessdate">. Retrieved <span class="nowrap">June 8,</span> 2021</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=search.amphilsoc.org&rft.atitle=APS+Member+History&rft_id=https%3A%2F%2Fsearch.amphilsoc.org%2Fmemhist%2Fsearch%3Fcreator%3DAvram%2BHershko%26title%3D%26subject%3D%26subdiv%3D%26mem%3D%26year%3D%26year-max%3D%26dead%3D%26keyword%3D%26smode%3Dadvanced&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAvram+Hershko" class="Z3988"></span></span> </li> <li id="cite_note-Oramed_Pharmaceuticals_2020_b459-10"><span class="mw-cite-backlink"><b><a href="#cite_ref-Oramed_Pharmaceuticals_2020_b459_10-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://oramed.com/nobel-laureate-oramed-sab-member-prof-avram-hershko-and-oramed-cso-dr-miriam-kidron-to-be-featured-on-biotalknology-webinar-oral-delivery-of-therapeutic-proteins-oramed-story-on-november-18-20/">"Nobel Laureate, Oramed SAB member Prof. Avram Hershko and Oramed CSO Dr. Miriam Kidron to be Featured on Biotalknology Webinar "Oral Delivery of Therapeutic Proteins – Oramed Story" on November 18, 2020"</a>. <i>Oramed Pharmaceuticals</i>. November 18, 2020<span class="reference-accessdate">. Retrieved <span class="nowrap">July 15,</span> 2023</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Oramed+Pharmaceuticals&rft.atitle=Nobel+Laureate%2C+Oramed+SAB+member+Prof.+Avram+Hershko+and+Oramed+CSO+Dr.+Miriam+Kidron+to+be+Featured+on+Biotalknology+Webinar+%22Oral+Delivery+of+Therapeutic+Proteins+%E2%80%93+Oramed+Story%22+on+November+18%2C+2020&rft.date=2020-11-18&rft_id=https%3A%2F%2Foramed.com%2Fnobel-laureate-oramed-sab-member-prof-avram-hershko-and-oramed-cso-dr-miriam-kidron-to-be-featured-on-biotalknology-webinar-oral-delivery-of-therapeutic-proteins-oramed-story-on-november-18-20%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3AAvram+Hershko" class="Z3988"></span></span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Avram_Hershko&action=edit&section=7" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1235681985">.mw-parser-output .side-box{margin:4px 0;box-sizing:border-box;border:1px solid #aaa;font-size:88%;line-height:1.25em;background-color:var(--background-color-interactive-subtle,#f8f9fa);display:flow-root}.mw-parser-output .side-box-abovebelow,.mw-parser-output .side-box-text{padding:0.25em 0.9em}.mw-parser-output .side-box-image{padding:2px 0 2px 0.9em;text-align:center}.mw-parser-output .side-box-imageright{padding:2px 0.9em 2px 0;text-align:center}@media(min-width:500px){.mw-parser-output .side-box-flex{display:flex;align-items:center}.mw-parser-output .side-box-text{flex:1;min-width:0}}@media(min-width:720px){.mw-parser-output .side-box{width:238px}.mw-parser-output .side-box-right{clear:right;float:right;margin-left:1em}.mw-parser-output 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src="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/30px-Commons-logo.svg.png" decoding="async" width="30" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/45px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/59px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></span></span></div> <div class="side-box-text plainlist">Wikimedia Commons has media related to <span style="font-weight: bold; font-style: italic;"><a href="https://commons.wikimedia.org/wiki/Category:Avram_Hershko" class="extiw" title="commons:Category:Avram Hershko">Avram Hershko</a></span>.</div></div> </div> <ul><li><a rel="nofollow" class="external text" href="https://www.nobelprize.org/laureate/780">Avram Hershko</a> on Nobelprize.org <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q232302#P8024" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/20px-OOjs_UI_icon_edit-ltr-progressive.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span> including the Nobel Lecture <i>The Ubiquitin System for Protein Degradation and some of its Roles in the Control of the Cell Division Cycle</i></li> <li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20130327222727/http://md.technion.ac.il/lecturers/lecturer_desc.asp?lecturerID=33&departmentID=2&contentCatID=13">Website at the Technion</a></li> <li><a rel="nofollow" class="external text" href="https://www.ibiology.org/profiles/lessons-life-science/">Avram Hershko's Short Talk: "Lessons from My Life in Science"</a></li> <li><a rel="nofollow" class="external text" href="http://www.crookedtimber.org/archives/002639.html">"Hungarian" Nobel Prize winners</a> Crooked Timber</li> <li><a rel="nofollow" class="external text" href="https://www.jewishvirtuallibrary.org/jsource/biography/Hershko.html">Avram Hershko</a> Jewish Virtual Library</li> <li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20050417005712/http://www.facs.org/education/gs2004/gs38hershko.pdf">Ubiquitin-Mediated Protein Degradation: From the lab to the bedside</a> Dan Hersko</li> <li><a rel="nofollow" class="external text" href="http://www.cumc.columbia.edu/horwitz/">The Official Site of Louisa Gross Horwitz Prize</a></li></ul> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style 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/></span></span></span> <a href="/wiki/List_of_Israeli_Nobel_laureates" title="List of Israeli Nobel laureates">Israeli Nobel laureates</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1966: <a href="/wiki/Shmuel_Yosef_Agnon" title="Shmuel Yosef Agnon">Shmuel Yosef Agnon</a> (<a href="/wiki/Nobel_Prize_in_Literature" title="Nobel Prize in Literature">Literature</a>)</span></li> <li><span class="nowrap">1978: <a href="/wiki/Menachem_Begin" title="Menachem Begin">Menachem Begin</a> (<a href="/wiki/Nobel_Prize_in_Peace" class="mw-redirect" title="Nobel Prize in Peace">Peace</a>)</span></li> <li><span class="nowrap">1994: <a href="/wiki/Shimon_Peres" title="Shimon Peres">Shimon Peres</a> / <a href="/wiki/Yitzhak_Rabin" title="Yitzhak Rabin">Yitzhak Rabin</a> (<a href="/wiki/Nobel_Prize_in_Peace" class="mw-redirect" title="Nobel Prize in Peace">Peace</a>)</span></li> <li><span class="nowrap">2002: <i><a href="/wiki/Daniel_Kahneman" title="Daniel Kahneman">Daniel Kahneman</a> (<a href="/wiki/Nobel_Memorial_Prize_in_Economic_Sciences" title="Nobel Memorial Prize in Economic Sciences">Economics</a>)</i></span></li> <li><span class="nowrap">2004: <a href="/wiki/Aaron_Ciechanover" title="Aaron Ciechanover">Aaron Ciechanover</a> / <a class="mw-selflink selflink">Avram Hershko</a> (<a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Chemistry</a>)</span></li> <li><span class="nowrap">2005: <i><a href="/wiki/Robert_Aumann" title="Robert Aumann">Robert Aumann</a> (<a href="/wiki/Nobel_Memorial_Prize_in_Economic_Sciences" title="Nobel Memorial Prize in Economic Sciences">Economics</a>)</i></span></li> <li><span class="nowrap">2009: <a href="/wiki/Ada_Yonath" title="Ada Yonath">Ada Yonath</a> (<a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Chemistry</a>)</span></li> <li><span class="nowrap">2011: <a href="/wiki/Dan_Shechtman" title="Dan Shechtman">Dan Shechtman</a> (<a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Chemistry</a>)</span></li> <li><span class="nowrap">2013: <a href="/wiki/Michael_Levitt_(biophysicist)" title="Michael Levitt (biophysicist)">Michael Levitt</a> / <a href="/wiki/Arieh_Warshel" title="Arieh Warshel">Arieh Warshel</a> (<a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Chemistry</a>)</span></li> <li><span class="nowrap">2021: <i><a href="/wiki/Joshua_Angrist" title="Joshua Angrist">Joshua Angrist</a> (<a href="/wiki/Nobel_Memorial_Prize_in_Economic_Sciences" title="Nobel Memorial Prize in Economic Sciences">Economics</a>)</i></span></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div><i>Italics</i> indicate a Nobel Memorial Prize, i.e. not one of the original Prizes bequested by <a href="/wiki/Alfred_Nobel" title="Alfred Nobel">Alfred Nobel</a>.</div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><style data-mw-deduplicate="TemplateStyles:r1239334494">@media screen{html.skin-theme-clientpref-night .mw-parser-output div:not(.notheme)>.tmp-color,html.skin-theme-clientpref-night .mw-parser-output p>.tmp-color,html.skin-theme-clientpref-night .mw-parser-output table:not(.notheme) .tmp-color{color:inherit!important}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output div:not(.notheme)>.tmp-color,html.skin-theme-clientpref-os .mw-parser-output p>.tmp-color,html.skin-theme-clientpref-os .mw-parser-output table:not(.notheme) .tmp-color{color:inherit!important}}</style><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239334494"></div><div role="navigation" class="navbox" aria-labelledby="Laureates_of_the_Nobel_Prize_in_Chemistry" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Nobel_Prize_in_Chemistry" title="Template:Nobel Prize in Chemistry"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Nobel_Prize_in_Chemistry" title="Template talk:Nobel Prize in Chemistry"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Nobel_Prize_in_Chemistry" title="Special:EditPage/Template:Nobel Prize in Chemistry"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Laureates_of_the_Nobel_Prize_in_Chemistry" style="font-size:114%;margin:0 4em"><a href="/wiki/List_of_Nobel_laureates_in_Chemistry" title="List of Nobel laureates in Chemistry">Laureates</a> of the <a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Nobel Prize in Chemistry</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">1901–1925</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1901: <a href="/wiki/Jacobus_Henricus_van_%27t_Hoff" title="Jacobus Henricus van 't Hoff">Jacobus van 't Hoff</a></span></li> <li><span class="nowrap">1902: <a href="/wiki/Emil_Fischer" title="Emil Fischer">Emil Fischer</a></span></li> <li><span class="nowrap">1903: <a href="/wiki/Svante_Arrhenius" title="Svante Arrhenius">Svante Arrhenius</a></span></li> <li><span class="nowrap">1904: <a href="/wiki/William_Ramsay" title="William Ramsay">William Ramsay</a></span></li> <li><span class="nowrap">1905: <a href="/wiki/Adolf_von_Baeyer" title="Adolf von Baeyer">Adolf von Baeyer</a></span></li> <li><span class="nowrap">1906: <a href="/wiki/Henri_Moissan" title="Henri Moissan">Henri Moissan</a></span></li> <li><span class="nowrap">1907: <a href="/wiki/Eduard_Buchner" title="Eduard Buchner">Eduard Buchner</a></span></li> <li><span class="nowrap">1908: <a href="/wiki/Ernest_Rutherford" title="Ernest Rutherford">Ernest Rutherford</a></span></li> <li><span class="nowrap">1909: <a href="/wiki/Wilhelm_Ostwald" title="Wilhelm Ostwald">Wilhelm Ostwald</a></span></li> <li><span class="nowrap">1910: <a href="/wiki/Otto_Wallach" title="Otto Wallach">Otto Wallach</a></span></li> <li><span class="nowrap">1911: <a href="/wiki/Marie_Curie" title="Marie Curie">Marie Curie</a></span></li> <li><span class="nowrap">1912: <a href="/wiki/Victor_Grignard" title="Victor Grignard">Victor Grignard</a> / <a href="/wiki/Paul_Sabatier_(chemist)" title="Paul Sabatier (chemist)">Paul Sabatier</a></span></li> <li><span class="nowrap">1913: <a href="/wiki/Alfred_Werner" title="Alfred Werner">Alfred Werner</a></span></li> <li><span class="nowrap">1914: <a href="/wiki/Theodore_William_Richards" title="Theodore William Richards">Theodore Richards</a></span></li> <li><span class="nowrap">1915: <a href="/wiki/Richard_Willst%C3%A4tter" title="Richard Willstätter">Richard Willstätter</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1916</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1917</span></span></li> <li><span class="nowrap">1918: <a href="/wiki/Fritz_Haber" title="Fritz Haber">Fritz Haber</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1919</span></span></li> <li><span class="nowrap">1920: <a href="/wiki/Walther_Nernst" title="Walther Nernst">Walther Nernst</a></span></li> <li><span class="nowrap">1921: <a href="/wiki/Frederick_Soddy" title="Frederick Soddy">Frederick Soddy</a></span></li> <li><span class="nowrap">1922: <a href="/wiki/Francis_William_Aston" title="Francis William Aston">Francis Aston</a></span></li> <li><span class="nowrap">1923: <a href="/wiki/Fritz_Pregl" title="Fritz Pregl">Fritz Pregl</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1924</span></span></li> <li><span class="nowrap">1925: <a href="/wiki/Richard_Adolf_Zsigmondy" title="Richard Adolf Zsigmondy">Richard Zsigmondy</a></span></li></ul> </div></td><td class="noviewer navbox-image" rowspan="5" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/80px-Nobel_Prize.png" decoding="async" width="80" height="79" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/120px-Nobel_Prize.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/e/ed/Nobel_Prize.png/160px-Nobel_Prize.png 2x" data-file-width="500" data-file-height="492" /></span></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1926–1950</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1926: <a href="/wiki/Theodor_Svedberg" title="Theodor Svedberg">Theodor Svedberg</a></span></li> <li><span class="nowrap">1927: <a href="/wiki/Heinrich_Otto_Wieland" title="Heinrich Otto Wieland">Heinrich Wieland</a></span></li> <li><span class="nowrap">1928: <a href="/wiki/Adolf_Windaus" title="Adolf Windaus">Adolf Windaus</a></span></li> <li><span class="nowrap">1929: <a href="/wiki/Arthur_Harden" title="Arthur Harden">Arthur Harden</a> / <a href="/wiki/Hans_von_Euler-Chelpin" title="Hans von Euler-Chelpin">Hans von Euler-Chelpin</a></span></li> <li><span class="nowrap">1930: <a href="/wiki/Hans_Fischer" title="Hans Fischer">Hans Fischer</a></span></li> <li><span class="nowrap">1931: <a href="/wiki/Carl_Bosch" title="Carl Bosch">Carl Bosch</a> / <a href="/wiki/Friedrich_Bergius" title="Friedrich Bergius">Friedrich Bergius</a></span></li> <li><span class="nowrap">1932: <a href="/wiki/Irving_Langmuir" title="Irving Langmuir">Irving Langmuir</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1933</span></span></li> <li><span class="nowrap">1934: <a href="/wiki/Harold_Urey" title="Harold Urey">Harold Urey</a></span></li> <li><span class="nowrap">1935: <a href="/wiki/Fr%C3%A9d%C3%A9ric_Joliot-Curie" title="Frédéric Joliot-Curie">Frédéric Joliot-Curie</a> / <a href="/wiki/Ir%C3%A8ne_Joliot-Curie" title="Irène Joliot-Curie">Irène Joliot-Curie</a></span></li> <li><span class="nowrap">1936: <a href="/wiki/Peter_Debye" title="Peter Debye">Peter Debye</a></span></li> <li><span class="nowrap">1937: <a href="/wiki/Norman_Haworth" title="Norman Haworth">Norman Haworth</a> / <a href="/wiki/Paul_Karrer" title="Paul Karrer">Paul Karrer</a></span></li> <li><span class="nowrap">1938: <a href="/wiki/Richard_Kuhn" title="Richard Kuhn">Richard Kuhn</a></span></li> <li><span class="nowrap">1939: <a href="/wiki/Adolf_Butenandt" title="Adolf Butenandt">Adolf Butenandt</a> / <a href="/wiki/Leopold_Ru%C5%BEi%C4%8Dka" title="Leopold Ružička">Leopold Ružička</a></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1940</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1941</span></span></li> <li><span class="nowrap"><span class="tmp-color" style="color:darkgrey">1942</span></span></li> <li><span class="nowrap">1943: <a href="/wiki/George_de_Hevesy" title="George de Hevesy">George de Hevesy</a></span></li> <li><span class="nowrap">1944: <a href="/wiki/Otto_Hahn" title="Otto Hahn">Otto Hahn</a></span></li> <li><span class="nowrap">1945: <a href="/wiki/Artturi_Ilmari_Virtanen" title="Artturi Ilmari Virtanen">Artturi Virtanen</a></span></li> <li><span class="nowrap">1946: <a href="/wiki/James_B._Sumner" title="James B. Sumner">James B. Sumner</a> / <a href="/wiki/John_Howard_Northrop" title="John Howard Northrop">John Northrop</a> / <a href="/wiki/Wendell_Meredith_Stanley" title="Wendell Meredith Stanley">Wendell Meredith Stanley</a></span></li> <li><span class="nowrap">1947: <a href="/wiki/Robert_Robinson_(chemist)" title="Robert Robinson (chemist)">Robert Robinson</a></span></li> <li><span class="nowrap">1948: <a href="/wiki/Arne_Tiselius" title="Arne Tiselius">Arne Tiselius</a></span></li> <li><span class="nowrap">1949: <a href="/wiki/William_Giauque" title="William Giauque">William Giauque</a></span></li> <li><span class="nowrap">1950: <a href="/wiki/Otto_Diels" title="Otto Diels">Otto Diels</a> / <a href="/wiki/Kurt_Alder" title="Kurt Alder">Kurt Alder</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1951–1975</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1951: <a href="/wiki/Edwin_McMillan" title="Edwin McMillan">Edwin McMillan</a> / <a href="/wiki/Glenn_T._Seaborg" title="Glenn T. Seaborg">Glenn T. Seaborg</a></span></li> <li><span class="nowrap">1952: <a href="/wiki/Archer_Martin" title="Archer Martin">Archer Martin</a> / <a href="/wiki/Richard_Laurence_Millington_Synge" title="Richard Laurence Millington Synge">Richard Synge</a></span></li> <li><span class="nowrap">1953: <a href="/wiki/Hermann_Staudinger" title="Hermann Staudinger">Hermann Staudinger</a></span></li> <li><span class="nowrap">1954: <a href="/wiki/Linus_Pauling" title="Linus Pauling">Linus Pauling</a></span></li> <li><span class="nowrap">1955: <a href="/wiki/Vincent_du_Vigneaud" title="Vincent du Vigneaud">Vincent du Vigneaud</a></span></li> <li><span class="nowrap">1956: <a href="/wiki/Cyril_Norman_Hinshelwood" class="mw-redirect" title="Cyril Norman Hinshelwood">Cyril Hinshelwood</a> / <a href="/wiki/Nikolay_Semyonov" title="Nikolay Semyonov">Nikolay Semyonov</a></span></li> <li><span class="nowrap">1957: <a href="/wiki/Alexander_R._Todd" class="mw-redirect" title="Alexander R. Todd">Alexander Todd</a></span></li> <li><span class="nowrap">1958: <a href="/wiki/Frederick_Sanger" title="Frederick Sanger">Frederick Sanger</a></span></li> <li><span class="nowrap">1959: <a href="/wiki/Jaroslav_Heyrovsk%C3%BD" title="Jaroslav Heyrovský">Jaroslav Heyrovský</a></span></li> <li><span class="nowrap">1960: <a href="/wiki/Willard_Libby" title="Willard Libby">Willard Libby</a></span></li> <li><span class="nowrap">1961: <a href="/wiki/Melvin_Calvin" title="Melvin Calvin">Melvin Calvin</a></span></li> <li><span class="nowrap">1962: <a href="/wiki/Max_Perutz" title="Max Perutz">Max Perutz</a> / <a href="/wiki/John_Kendrew" title="John Kendrew">John Kendrew</a></span></li> <li><span class="nowrap">1963: <a href="/wiki/Karl_Ziegler" title="Karl Ziegler">Karl Ziegler</a> / <a href="/wiki/Giulio_Natta" title="Giulio Natta">Giulio Natta</a></span></li> <li><span class="nowrap">1964: <a href="/wiki/Dorothy_Hodgkin" title="Dorothy Hodgkin">Dorothy Hodgkin</a></span></li> <li><span class="nowrap">1965: <a href="/wiki/Robert_Burns_Woodward" title="Robert Burns Woodward">Robert Woodward</a></span></li> <li><span class="nowrap">1966: <a href="/wiki/Robert_S._Mulliken" title="Robert S. Mulliken">Robert S. Mulliken</a></span></li> <li><span class="nowrap">1967: <a href="/wiki/Manfred_Eigen" title="Manfred Eigen">Manfred Eigen</a> / <a href="/wiki/Ronald_George_Wreyford_Norrish" title="Ronald George Wreyford Norrish">Ronald Norrish</a> / <a href="/wiki/George_Porter" title="George Porter">George Porter</a></span></li> <li><span class="nowrap">1968: <a href="/wiki/Lars_Onsager" title="Lars Onsager">Lars Onsager</a></span></li> <li><span class="nowrap">1969: <a href="/wiki/Derek_Barton" title="Derek Barton">Derek Barton</a> / <a href="/wiki/Odd_Hassel" title="Odd Hassel">Odd Hassel</a></span></li> <li><span class="nowrap">1970: <a href="/wiki/Luis_Federico_Leloir" title="Luis Federico Leloir">Luis Federico Leloir</a></span></li> <li><span class="nowrap">1971: <a href="/wiki/Gerhard_Herzberg" title="Gerhard Herzberg">Gerhard Herzberg</a></span></li> <li><span class="nowrap">1972: <a href="/wiki/Christian_B._Anfinsen" title="Christian B. Anfinsen">Christian B. Anfinsen</a> / <a href="/wiki/Stanford_Moore" title="Stanford Moore">Stanford Moore</a> / <a href="/wiki/William_Howard_Stein" title="William Howard Stein">William Stein</a></span></li> <li><span class="nowrap">1973: <a href="/wiki/Ernst_Otto_Fischer" title="Ernst Otto Fischer">Ernst Otto Fischer</a> / <a href="/wiki/Geoffrey_Wilkinson" title="Geoffrey Wilkinson">Geoffrey Wilkinson</a></span></li> <li><span class="nowrap">1974: <a href="/wiki/Paul_Flory" title="Paul Flory">Paul Flory</a></span></li> <li><span class="nowrap">1975: <a href="/wiki/John_Cornforth" title="John Cornforth">John Cornforth</a> / <a href="/wiki/Vladimir_Prelog" title="Vladimir Prelog">Vladimir Prelog</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1976–2000</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">1976: <a href="/wiki/William_Lipscomb" title="William Lipscomb">William Lipscomb</a></span></li> <li><span class="nowrap">1977: <a href="/wiki/Ilya_Prigogine" title="Ilya Prigogine">Ilya Prigogine</a></span></li> <li><span class="nowrap">1978: <a href="/wiki/Peter_D._Mitchell" title="Peter D. Mitchell">Peter D. Mitchell</a></span></li> <li><span class="nowrap">1979: <a href="/wiki/Herbert_C._Brown" title="Herbert C. Brown">Herbert C. Brown</a> / <a href="/wiki/Georg_Wittig" title="Georg Wittig">Georg Wittig</a></span></li> <li><span class="nowrap">1980: <a href="/wiki/Paul_Berg" title="Paul Berg">Paul Berg</a> / <a href="/wiki/Walter_Gilbert" title="Walter Gilbert">Walter Gilbert</a> / <a href="/wiki/Frederick_Sanger" title="Frederick Sanger">Frederick Sanger</a></span></li> <li><span class="nowrap">1981: <a href="/wiki/Kenichi_Fukui" title="Kenichi Fukui">Kenichi Fukui</a> / <a href="/wiki/Roald_Hoffmann" title="Roald Hoffmann">Roald Hoffmann</a></span></li> <li><span class="nowrap">1982: <a href="/wiki/Aaron_Klug" title="Aaron Klug">Aaron Klug</a></span></li> <li><span class="nowrap">1983: <a href="/wiki/Henry_Taube" title="Henry Taube">Henry Taube</a></span></li> <li><span class="nowrap">1984: <a href="/wiki/Robert_Bruce_Merrifield" title="Robert Bruce Merrifield">Robert Merrifield</a></span></li> <li><span class="nowrap">1985: <a href="/wiki/Herbert_A._Hauptman" title="Herbert A. Hauptman">Herbert A. Hauptman</a> / <a href="/wiki/Jerome_Karle" title="Jerome Karle">Jerome Karle</a></span></li> <li><span class="nowrap">1986: <a href="/wiki/Dudley_R._Herschbach" title="Dudley R. Herschbach">Dudley R. Herschbach</a> / <a href="/wiki/Yuan_T._Lee" title="Yuan T. Lee">Yuan T. Lee</a> / <a href="/wiki/John_Polanyi" title="John Polanyi">John Polanyi</a></span></li> <li><span class="nowrap">1987: <a href="/wiki/Donald_J._Cram" title="Donald J. Cram">Donald J. Cram</a> / <a href="/wiki/Jean-Marie_Lehn" title="Jean-Marie Lehn">Jean-Marie Lehn</a> / <a href="/wiki/Charles_J._Pedersen" title="Charles J. Pedersen">Charles J. Pedersen</a></span></li> <li><span class="nowrap">1988: <a href="/wiki/Johann_Deisenhofer" title="Johann Deisenhofer">Johann Deisenhofer</a> / <a href="/wiki/Robert_Huber" title="Robert Huber">Robert Huber</a> / <a href="/wiki/Hartmut_Michel" title="Hartmut Michel">Hartmut Michel</a></span></li> <li><span class="nowrap">1989: <a href="/wiki/Sidney_Altman" title="Sidney Altman">Sidney Altman</a> / <a href="/wiki/Thomas_Cech" title="Thomas Cech">Thomas Cech</a></span></li> <li><span class="nowrap">1990: <a href="/wiki/Elias_James_Corey" title="Elias James Corey">Elias Corey</a></span></li> <li><span class="nowrap">1991: <a href="/wiki/Richard_R._Ernst" title="Richard R. Ernst">Richard R. Ernst</a></span></li> <li><span class="nowrap">1992: <a href="/wiki/Rudolph_A._Marcus" title="Rudolph A. Marcus">Rudolph A. Marcus</a></span></li> <li><span class="nowrap">1993: <a href="/wiki/Kary_Mullis" title="Kary Mullis">Kary Mullis</a> / <a href="/wiki/Michael_Smith_(chemist)" title="Michael Smith (chemist)">Michael Smith</a></span></li> <li><span class="nowrap">1994: <a href="/wiki/George_Andrew_Olah" title="George Andrew Olah">George Olah</a></span></li> <li><span class="nowrap">1995: <a href="/wiki/Paul_J._Crutzen" title="Paul J. Crutzen">Paul J. Crutzen</a> / <a href="/wiki/Mario_Molina" title="Mario Molina">Mario Molina</a> / <a href="/wiki/F._Sherwood_Rowland" title="F. Sherwood Rowland">F. Sherwood Rowland</a></span></li> <li><span class="nowrap">1996: <a href="/wiki/Robert_Curl" title="Robert Curl">Robert Curl</a> / <a href="/wiki/Harry_Kroto" title="Harry Kroto">Harold Kroto</a> / <a href="/wiki/Richard_Smalley" title="Richard Smalley">Richard Smalley</a></span></li> <li><span class="nowrap">1997: <a href="/wiki/Paul_D._Boyer" title="Paul D. Boyer">Paul D. Boyer</a> / <a href="/wiki/John_E._Walker" title="John E. Walker">John E. Walker</a> / <a href="/wiki/Jens_Christian_Skou" title="Jens Christian Skou">Jens Christian Skou</a></span></li> <li><span class="nowrap">1998: <a href="/wiki/Walter_Kohn" title="Walter Kohn">Walter Kohn</a> / <a href="/wiki/John_Pople" title="John Pople">John Pople</a></span></li> <li><span class="nowrap">1999: <a href="/wiki/Ahmed_Zewail" title="Ahmed Zewail">Ahmed Zewail</a></span></li> <li><span class="nowrap">2000: <a href="/wiki/Alan_J._Heeger" title="Alan J. Heeger">Alan J. Heeger</a> / <a href="/wiki/Alan_MacDiarmid" title="Alan MacDiarmid">Alan MacDiarmid</a> / <a href="/wiki/Hideki_Shirakawa" title="Hideki Shirakawa">Hideki Shirakawa</a></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2001–present</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span class="nowrap">2001: <a href="/wiki/William_Standish_Knowles" title="William Standish Knowles">William Knowles</a> / <a href="/wiki/Ry%C5%8Dji_Noyori" title="Ryōji Noyori">Ryoji Noyori</a> / <a href="/wiki/Karl_Barry_Sharpless" title="Karl Barry Sharpless">K. Barry Sharpless</a></span></li> <li><span class="nowrap">2002: <a href="/wiki/John_B._Fenn" title="John B. Fenn">John B. Fenn</a> / <a href="/wiki/Koichi_Tanaka" title="Koichi Tanaka">Koichi Tanaka</a> / <a href="/wiki/Kurt_W%C3%BCthrich" title="Kurt Wüthrich">Kurt Wüthrich</a></span></li> <li><span class="nowrap">2003: <a href="/wiki/Peter_Agre" title="Peter Agre">Peter Agre</a> / <a href="/wiki/Roderick_MacKinnon" title="Roderick MacKinnon">Roderick MacKinnon</a></span></li> <li><span class="nowrap">2004: <a href="/wiki/Aaron_Ciechanover" title="Aaron Ciechanover">Aaron Ciechanover</a> / <a class="mw-selflink selflink">Avram Hershko</a> / <a href="/wiki/Irwin_Rose" title="Irwin Rose">Irwin Rose</a></span></li> <li><span class="nowrap">2005: <a href="/wiki/Robert_H._Grubbs" title="Robert H. Grubbs">Robert H. Grubbs</a> / <a href="/wiki/Richard_R._Schrock" title="Richard R. Schrock">Richard R. Schrock</a> / <a href="/wiki/Yves_Chauvin" title="Yves Chauvin">Yves Chauvin</a></span></li> <li><span class="nowrap">2006: <a href="/wiki/Roger_D._Kornberg" title="Roger D. Kornberg">Roger D. Kornberg</a></span></li> <li><span class="nowrap">2007: <a href="/wiki/Gerhard_Ertl" title="Gerhard Ertl">Gerhard Ertl</a></span></li> <li><span class="nowrap">2008: <a href="/wiki/Osamu_Shimomura" title="Osamu Shimomura">Osamu Shimomura</a> / <a href="/wiki/Martin_Chalfie" title="Martin Chalfie">Martin Chalfie</a> / <a href="/wiki/Roger_Y._Tsien" title="Roger Y. Tsien">Roger Y. Tsien</a></span></li> <li><span class="nowrap">2009: <a href="/wiki/Venki_Ramakrishnan" title="Venki Ramakrishnan">Venkatraman Ramakrishnan</a> / <a href="/wiki/Thomas_A._Steitz" title="Thomas A. Steitz">Thomas A. Steitz</a> / <a href="/wiki/Ada_Yonath" title="Ada Yonath">Ada E. Yonath</a></span></li> <li><span class="nowrap">2010: <a href="/wiki/Richard_F._Heck" title="Richard F. Heck">Richard F. Heck</a> / <a href="/wiki/Akira_Suzuki" title="Akira Suzuki">Akira Suzuki</a> / <a href="/wiki/Ei-ichi_Negishi" title="Ei-ichi Negishi">Ei-ichi Negishi</a></span></li> <li><span class="nowrap">2011: <a href="/wiki/Dan_Shechtman" title="Dan Shechtman">Dan Shechtman</a></span></li> <li><span class="nowrap">2012: <a href="/wiki/Robert_Lefkowitz" title="Robert Lefkowitz">Robert Lefkowitz</a> / <a href="/wiki/Brian_Kobilka" title="Brian Kobilka">Brian Kobilka</a></span></li> <li><span class="nowrap">2013: <a href="/wiki/Martin_Karplus" title="Martin Karplus">Martin Karplus</a> / <a href="/wiki/Michael_Levitt_(biophysicist)" title="Michael Levitt (biophysicist)">Michael Levitt</a> / <a href="/wiki/Arieh_Warshel" title="Arieh Warshel">Arieh Warshel</a></span></li> <li><span class="nowrap">2014: <a href="/wiki/Eric_Betzig" title="Eric Betzig">Eric Betzig</a> / <a href="/wiki/Stefan_Hell" title="Stefan Hell">Stefan Hell</a> / <a href="/wiki/William_E._Moerner" title="William E. Moerner">William E. Moerner</a></span></li> <li><span class="nowrap">2015: <a href="/wiki/Tomas_Lindahl" title="Tomas Lindahl">Tomas Lindahl</a> / <a href="/wiki/Paul_L._Modrich" title="Paul L. Modrich">Paul L. Modrich</a> / <a href="/wiki/Aziz_Sancar" title="Aziz Sancar">Aziz Sancar</a></span></li> <li><span class="nowrap">2016: <a href="/wiki/Jean-Pierre_Sauvage" title="Jean-Pierre Sauvage">Jean-Pierre Sauvage</a> / <a href="/wiki/Fraser_Stoddart" title="Fraser Stoddart">Fraser Stoddart</a> / <a href="/wiki/Ben_Feringa" title="Ben Feringa">Ben Feringa</a></span></li> <li><span class="nowrap">2017: <a href="/wiki/Jacques_Dubochet" title="Jacques Dubochet">Jacques Dubochet</a> / <a href="/wiki/Joachim_Frank" title="Joachim Frank">Joachim Frank</a> / <a href="/wiki/Richard_Henderson_(biologist)" title="Richard Henderson (biologist)">Richard Henderson</a></span></li> <li><span class="nowrap">2018: <a href="/wiki/Frances_Arnold" title="Frances Arnold">Frances Arnold</a> / <a href="/wiki/Gregory_Winter" title="Gregory Winter">Gregory Winter</a> / <a href="/wiki/George_Smith_(chemist)" title="George Smith (chemist)">George Smith</a></span></li> <li><span class="nowrap">2019: <a href="/wiki/John_B._Goodenough" title="John B. Goodenough">John B. Goodenough</a> / <a href="/wiki/M._Stanley_Whittingham" title="M. Stanley Whittingham">M. Stanley Whittingham</a> / <a href="/wiki/Akira_Yoshino" title="Akira Yoshino">Akira Yoshino</a></span></li> <li><span class="nowrap">2020: <a href="/wiki/Emmanuelle_Charpentier" title="Emmanuelle Charpentier">Emmanuelle Charpentier</a> / <a href="/wiki/Jennifer_Doudna" title="Jennifer Doudna">Jennifer Doudna</a></span></li> <li><span class="nowrap">2021: <a href="/wiki/David_MacMillan" title="David MacMillan">David MacMillan</a> / <a href="/wiki/Benjamin_List" title="Benjamin List">Benjamin List</a></span></li> <li><span class="nowrap">2022: <a href="/wiki/Carolyn_Bertozzi" title="Carolyn Bertozzi">Carolyn R. Bertozzi</a> / <a href="/wiki/Morten_P._Meldal" title="Morten P. Meldal">Morten P. Meldal</a> / <a href="/wiki/Karl_Barry_Sharpless" title="Karl Barry Sharpless">Karl Barry Sharpless</a></span></li> <li><span class="nowrap">2023: <a href="/wiki/Moungi_Bawendi" title="Moungi Bawendi">Moungi G. Bawendi</a> / <a href="/wiki/Louis_E._Brus" title="Louis E. Brus">Louis E. Brus</a> / <a href="/wiki/Alexey_Ekimov" title="Alexey Ekimov">Alexei I. Ekimov</a></span></li> <li><span class="nowrap">2024: <a href="/wiki/David_Baker_(biochemist)" title="David Baker (biochemist)">David Baker</a> / <a href="/wiki/Demis_Hassabis" title="Demis Hassabis">Demis Hassabis</a> / <a href="/wiki/John_M._Jumper" title="John M. Jumper">John M. Jumper</a></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><style data-mw-deduplicate="TemplateStyles:r886047488">.mw-parser-output .nobold{font-weight:normal}</style></div><div role="navigation" class="navbox" aria-labelledby="2004_Nobel_Prize_laureates" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:2004_Nobel_Prize_winners" title="Template:2004 Nobel Prize winners"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:2004_Nobel_Prize_winners" title="Template talk:2004 Nobel Prize winners"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:2004_Nobel_Prize_winners" title="Special:EditPage/Template:2004 Nobel Prize winners"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="2004_Nobel_Prize_laureates" style="font-size:114%;margin:0 4em">2004 <a href="/wiki/Nobel_Prize" title="Nobel Prize">Nobel Prize</a> laureates</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Chemistry</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Aaron_Ciechanover" title="Aaron Ciechanover">Aaron Ciechanover</a> (Israel)</li> <li><a class="mw-selflink selflink">Avram Hershko</a> (Israel)</li> <li><a href="/wiki/Irwin_Rose" title="Irwin Rose">Irwin Rose</a> (United States)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Literature" title="Nobel Prize in Literature">Literature</a> (<a href="/wiki/2004_Nobel_Prize_in_Literature" title="2004 Nobel Prize in Literature">2004</a>)</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Elfriede_Jelinek" title="Elfriede Jelinek">Elfriede Jelinek</a> (Austria)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Peace_Prize" title="Nobel Peace Prize">Peace</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Wangari_Maathai" class="mw-redirect" title="Wangari Maathai">Wangari Maathai</a> (Kenya)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physics" title="Nobel Prize in Physics">Physics</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/David_Gross" title="David Gross">David Gross</a> (United States)</li> <li><a href="/wiki/Hugh_David_Politzer" title="Hugh David Politzer">Hugh David Politzer</a> (United States)</li> <li><a href="/wiki/Frank_Wilczek" title="Frank Wilczek">Frank Wilczek</a> (United States)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Physiology or Medicine</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Richard_Axel" title="Richard Axel">Richard Axel</a> (United States)</li> <li><a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Linda B. Buck</a> (United States)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Memorial_Prize_in_Economic_Sciences" title="Nobel Memorial Prize in Economic Sciences">Economic Sciences</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Finn_E._Kydland" title="Finn E. Kydland">Finn E. Kydland</a> (Norway)</li> <li><a href="/wiki/Edward_C._Prescott" title="Edward C. Prescott">Edward C. Prescott</a> (United States)</li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <dl><dt><span class="nobold"><a href="/wiki/List_of_Nobel_laureates" title="List of Nobel laureates">Nobel Prize recipients</a></span></dt> <dd><a href="/wiki/Template:1999_Nobel_Prize_winners" title="Template:1999 Nobel Prize winners">1999</a></dd> <dd><a href="/wiki/Template:2000_Nobel_Prize_winners" title="Template:2000 Nobel Prize winners">2000</a></dd> <dd><a href="/wiki/Template:2001_Nobel_Prize_winners" title="Template:2001 Nobel Prize winners">2001</a></dd> <dd><a href="/wiki/Template:2002_Nobel_Prize_winners" title="Template:2002 Nobel Prize winners">2002</a></dd> <dd><a href="/wiki/Template:2003_Nobel_Prize_winners" title="Template:2003 Nobel Prize winners">2003</a></dd> <dd><b>2004 </b></dd> <dd><a href="/wiki/Template:2005_Nobel_Prize_winners" title="Template:2005 Nobel Prize winners">2005</a></dd> <dd><a href="/wiki/Template:2006_Nobel_Prize_winners" title="Template:2006 Nobel Prize winners">2006</a></dd> <dd><a href="/wiki/Template:2007_Nobel_Prize_winners" title="Template:2007 Nobel Prize winners">2007</a></dd> <dd><a href="/wiki/Template:2008_Nobel_Prize_winners" title="Template:2008 Nobel Prize winners">2008</a></dd> <dd><a href="/wiki/Template:2009_Nobel_Prize_winners" title="Template:2009 Nobel Prize winners">2009</a></dd></dl> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Laureates_of_the_Wolf_Prize_in_Medicine" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2" style="background:#d3d3d3;"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Wolf_Prize_in_Medicine" title="Template:Wolf Prize in Medicine"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Wolf_Prize_in_Medicine" title="Template talk:Wolf Prize in Medicine"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Wolf_Prize_in_Medicine" title="Special:EditPage/Template:Wolf Prize in Medicine"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Laureates_of_the_Wolf_Prize_in_Medicine" style="font-size:114%;margin:0 4em">Laureates of the <a href="/wiki/Wolf_Prize_in_Medicine" title="Wolf Prize in Medicine">Wolf Prize in Medicine</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1970s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/George_Davis_Snell" title="George Davis Snell">George Snell</a> / <a href="/wiki/Jean_Dausset" title="Jean Dausset">Jean Dausset</a> / <a href="/wiki/Jon_van_Rood" title="Jon van Rood">Jon J. van Rood</a> (1978)</li> <li><a href="/wiki/Roger_Wolcott_Sperry" title="Roger Wolcott Sperry">Roger Sperry</a> / <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Arvid Carlsson</a> / <a href="/wiki/Oleh_Hornykiewicz" title="Oleh Hornykiewicz">Oleh Hornykiewicz</a> (1979)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1980s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a> / <a href="/wiki/Leo_Sachs" title="Leo Sachs">Leo Sachs</a> / <a href="/wiki/James_Learmonth_Gowans" title="James Learmonth Gowans">James L. Gowans</a> (1980)</li> <li><a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a> / <a href="/wiki/Stanley_Norman_Cohen" title="Stanley Norman Cohen">Stanley Norman Cohen</a> (1981)</li> <li><a href="/wiki/Jean-Pierre_Changeux" title="Jean-Pierre Changeux">Jean-Pierre Changeux</a> / <a href="/wiki/Solomon_H._Snyder" title="Solomon H. Snyder">Solomon H. Snyder</a> / <a href="/wiki/James_Black_(pharmacologist)" title="James Black (pharmacologist)">James W. Black</a> (1982)</li> <li><a href="/wiki/Donald_F._Steiner" title="Donald F. Steiner">Donald F. Steiner</a> (1984/5)</li> <li><a href="/wiki/Osamu_Hayaishi" title="Osamu Hayaishi">Osamu Hayaishi</a> (1986)</li> <li><a href="/wiki/Pedro_Cuatrecasas" title="Pedro Cuatrecasas">Pedro Cuatrecasas</a> / <a href="/wiki/Meir_Wilchek" title="Meir Wilchek">Meir Wilchek</a> (1987)</li> <li><a href="/wiki/Henri_G._Hers" title="Henri G. Hers">Henri G. Hers</a> / <a href="/wiki/Elizabeth_F._Neufeld" title="Elizabeth F. Neufeld">Elizabeth F. Neufeld</a> (1988)</li> <li><a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> / <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Edward B. Lewis</a> (1989)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1990s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Maclyn_McCarty" title="Maclyn McCarty">Maclyn McCarty</a> (1990)</li> <li><a href="/wiki/Seymour_Benzer" title="Seymour Benzer">Seymour Benzer</a> (1991)</li> <li><a href="/wiki/Judah_Folkman" title="Judah Folkman">Judah Folkman</a> (1992)</li> <li><a href="/wiki/Michael_Berridge" title="Michael Berridge">Michael Berridge</a> / <a href="/wiki/Yasutomi_Nishizuka" title="Yasutomi Nishizuka">Yasutomi Nishizuka</a> (1994/5)</li> <li><a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Stanley B. Prusiner</a> (1995/6)</li> <li><a href="/wiki/Mary_F._Lyon" title="Mary F. Lyon">Mary F. Lyon</a> (1996/7)</li> <li><a href="/wiki/Michael_Sela" title="Michael Sela">Michael Sela</a> / <a href="/wiki/Ruth_Arnon" title="Ruth Arnon">Ruth Arnon</a> (1998)</li> <li><a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric Kandel</a> (1999)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">2000s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a class="mw-selflink selflink">Avram Hershko</a> / <a href="/wiki/Alexander_Varshavsky" title="Alexander Varshavsky">Alexander Varshavsky</a> (2001)</li> <li><a href="/wiki/Ralph_L._Brinster" title="Ralph L. Brinster">Ralph L. Brinster</a> / <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Mario Capecchi</a> / <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a> (2002/3)</li> <li><a href="/wiki/Robert_Weinberg_(biologist)" title="Robert Weinberg (biologist)">Robert Weinberg</a> / <a href="/wiki/Roger_Y._Tsien" title="Roger Y. Tsien">Roger Y. Tsien</a> (2004)</li> <li><a href="/wiki/Alexander_Levitzki" title="Alexander Levitzki">Alexander Levitzki</a> / <a href="/wiki/Anthony_R._Hunter" title="Anthony R. Hunter">Anthony R. Hunter</a> / <a href="/wiki/Tony_Pawson_(biochemist)" title="Tony Pawson (biochemist)">Tony Pawson</a> (2005)</li> <li><a href="/wiki/Howard_Cedar" title="Howard Cedar">Howard Cedar</a> / <a href="/wiki/Aharon_Razin" title="Aharon Razin">Aharon Razin</a> (2008)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">2010s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Axel_Ullrich" title="Axel Ullrich">Axel Ullrich</a> (2010)</li> <li><a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Shinya Yamanaka</a> / <a href="/wiki/Rudolf_Jaenisch" title="Rudolf Jaenisch">Rudolf Jaenisch</a> (2011)</li> <li><a href="/wiki/Ronald_M._Evans" title="Ronald M. Evans">Ronald M. Evans</a> (2012)</li> <li><a href="/wiki/Nahum_Sonenberg" title="Nahum Sonenberg">Nahum Sonenberg</a> / <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a> / <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a> (2014)</li> <li><a href="/wiki/John_Kappler" title="John Kappler">John Kappler</a> / <a href="/wiki/Philippa_Marrack" title="Philippa Marrack">Philippa Marrack</a> / <a href="/wiki/Jeffrey_V._Ravetch" title="Jeffrey V. Ravetch">Jeffrey V. Ravetch</a> (2015)</li> <li><a href="/wiki/Lewis_C._Cantley" title="Lewis C. Cantley">Lewis C. Cantley</a> / <a href="/wiki/C._Ronald_Kahn" title="C. Ronald Kahn">C. Ronald Kahn</a> (2016)</li> <li><a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a> (2017)</li> <li><a href="/wiki/Jeffrey_M._Friedman" title="Jeffrey M. Friedman">Jeffrey M. Friedman</a> (2019)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">2020s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Emmanuelle_Charpentier" title="Emmanuelle Charpentier">Emmanuelle Charpentier</a> / <a href="/wiki/Jennifer_Doudna" title="Jennifer Doudna">Jennifer Doudna</a> (2020)</li> <li><a href="/wiki/Joan_Steitz" class="mw-redirect" title="Joan Steitz">Joan Steitz</a> / <a href="/wiki/Lynne_Maquat" class="mw-redirect" title="Lynne Maquat">Lynne Elizabeth Maquat</a> / <a href="/wiki/Adrian_R._Krainer" class="mw-redirect" title="Adrian R. Krainer">Adrian Krainer</a> (2021)</li> <li><a href="/wiki/Daniel_J._Drucker" title="Daniel J. Drucker">Daniel J. Drucker</a> (2023)</li> <li><a href="/wiki/Botond_Roska" title="Botond Roska">Botond Roska</a> / <a href="/wiki/Jos%C3%A9-Alain_Sahel" title="José-Alain Sahel">José-Alain Sahel</a> (2024)</li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Hungarian_or_Hungarian-American_Nobel_Laureates_23x15px&#124;border_&#124;alt=&#124;link=" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Hungarian_Nobel_Laureates" title="Template:Hungarian Nobel Laureates"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Hungarian_Nobel_Laureates" title="Template talk:Hungarian Nobel Laureates"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Hungarian_Nobel_Laureates" title="Special:EditPage/Template:Hungarian Nobel Laureates"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Hungarian_or_Hungarian-American_Nobel_Laureates_23x15px&#124;border_&#124;alt=&#124;link=" style="font-size:114%;margin:0 4em"><a href="/wiki/Hungarian_people" class="mw-redirect" title="Hungarian people">Hungarian</a> or Hungarian-American <a href="/wiki/List_of_Nobel_laureates" title="List of Nobel laureates">Nobel Laureates</a> <span class="flagicon"><span class="mw-image-border" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c1/Flag_of_Hungary.svg/23px-Flag_of_Hungary.svg.png" decoding="async" width="23" height="12" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c1/Flag_of_Hungary.svg/35px-Flag_of_Hungary.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c1/Flag_of_Hungary.svg/46px-Flag_of_Hungary.svg.png 2x" data-file-width="1200" data-file-height="600" /></span></span></span></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Chemistry" title="Nobel Prize in Chemistry">Chemistry</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Richard_Adolf_Zsigmondy" title="Richard Adolf Zsigmondy">Zsigmondy Richárd</a> (1925)</li> <li><a href="/wiki/George_de_Hevesy" title="George de Hevesy">Hevesy György</a> (1943)</li> <li><a href="/wiki/John_Polanyi" title="John Polanyi">Polányi János</a> (1986)</li> <li><a href="/wiki/George_Andrew_Olah" title="George Andrew Olah">Oláh György</a> (1994)</li> <li><a class="mw-selflink selflink">Herskó Ferenc</a> (2004)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Literature" title="Nobel Prize in Literature">Literature</a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Imre_Kert%C3%A9sz" title="Imre Kertész">Kertész Imre</a> (2002)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physics" title="Nobel Prize in Physics">Physics</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Philipp_Lenard" title="Philipp Lenard">Lénárd Fülöp</a> (1905)</li> <li><a href="/wiki/Eugene_Wigner" title="Eugene Wigner">Wigner Jenő</a> (1963)</li> <li><a href="/wiki/Dennis_Gabor" title="Dennis Gabor">Gábor Dénes</a> (1971)</li> <li><a href="/wiki/Ferenc_Krausz" title="Ferenc Krausz">Krausz Ferenc</a> (2023)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Prize_in_Physiology_or_Medicine" title="Nobel Prize in Physiology or Medicine">Physiology or Medicine</a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/R%C3%B3bert_B%C3%A1r%C3%A1ny" class="mw-redirect" title="Róbert Bárány">Bárány Róbert</a> (1914)</li> <li><a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Szent-Györgyi Albert</a> (1937)</li> <li><a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Békésy György</a> (1961)</li> <li><a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Karikó Katalin</a> (2023)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Nobel_Memorial_Prize_in_Economic_Sciences" title="Nobel Memorial Prize in Economic Sciences">Economic Sciences</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/John_Harsanyi" title="John Harsanyi">Harsányi János</a> (1994)</li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox authority-control" aria-labelledby="Authority_control_databases_frameless&#124;text-top&#124;10px&#124;alt=Edit_this_at_Wikidata&#124;link=https&#58;//www.wikidata.org/wiki/Q232302#identifiers&#124;class=noprint&#124;Edit_this_at_Wikidata" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Authority_control_databases_frameless&#124;text-top&#124;10px&#124;alt=Edit_this_at_Wikidata&#124;link=https&#58;//www.wikidata.org/wiki/Q232302#identifiers&#124;class=noprint&#124;Edit_this_at_Wikidata" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Authority_control" title="Help:Authority control">Authority control databases</a> <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q232302#identifiers" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" 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class="navbox-group" style="width:1%">National</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://d-nb.info/gnd/1103034480">Germany</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/n88163753">United States</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://libris.kb.se/0xbdhggj3rv3tg4">Sweden</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://dbn.bn.org.pl/descriptor-details/9810693356505606">Poland</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="http://olduli.nli.org.il/F/?func=find-b&local_base=NLX10&find_code=UID&request=987007460173305171">Israel</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Academics</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.scopus.com/authid/detail.uri?authorId=7006965675">Scopus</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.idref.fr/080905234">IdRef</a></span></li></ul></div></td></tr></tbody></table></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Retrieved from "<a dir="ltr" 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