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vector-toc-level-2"> <a class="vector-toc-link" href="#Medico-legal_entomology"> <div class="vector-toc-text"> <span class="vector-toc-numb">3.2</span> <span>Medico-legal entomology</span> </div> </a> <ul id="toc-Medico-legal_entomology-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Ongoing_research" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Ongoing_research"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Ongoing research</span> </div> </a> <ul id="toc-Ongoing_research-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Systematics" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Systematics"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>Systematics</span> </div> </a> <ul id="toc-Systematics-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" 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Available in 31 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-31" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">31 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%A8%D9%88%D9%82%D9%8A%D8%A7%D8%AA_%D8%B6%D8%A7%D8%B1%D9%8A%D8%A9" title="بوقيات ضارية – Arabic" lang="ar" hreflang="ar" data-title="بوقيات ضارية" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%A1%D1%82%D1%80%D0%B0%D0%BA%D0%B0%D1%87%D1%8B" title="Стракачы – Belarusian" lang="be" hreflang="be" data-title="Стракачы" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Cl%C3%A8rids" title="Clèrids – Catalan" lang="ca" hreflang="ca" data-title="Clèrids" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-ceb mw-list-item"><a href="https://ceb.wikipedia.org/wiki/Cleridae" title="Cleridae – Cebuano" lang="ceb" hreflang="ceb" data-title="Cleridae" data-language-autonym="Cebuano" data-language-local-name="Cebuano" class="interlanguage-link-target"><span>Cebuano</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Pestrokrove%C4%8Dn%C3%ADkovit%C3%AD" title="Pestrokrovečníkovití – Czech" lang="cs" hreflang="cs" data-title="Pestrokrovečníkovití" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Buntk%C3%A4fer" title="Buntkäfer – German" lang="de" hreflang="de" data-title="Buntkäfer" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-nv mw-list-item"><a href="https://nv.wikipedia.org/wiki/Nd%C3%ADy%C3%ADliitsoh_bich%CA%BCosh" title="Ndíyíliitsoh bichʼosh – Navajo" lang="nv" hreflang="nv" data-title="Ndíyíliitsoh bichʼosh" data-language-autonym="Diné bizaad" data-language-local-name="Navajo" class="interlanguage-link-target"><span>Diné bizaad</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Sipelgmardiklased" title="Sipelgmardiklased – Estonian" lang="et" hreflang="et" data-title="Sipelgmardiklased" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Cleridae" title="Cleridae – Spanish" lang="es" hreflang="es" data-title="Cleridae" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Cleridae" title="Cleridae – Basque" lang="eu" hreflang="eu" data-title="Cleridae" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%B3%D9%88%D8%B3%DA%A9%E2%80%8C%D9%87%D8%A7%DB%8C_%D8%B1%D9%86%DA%AF%D8%A7%D8%B1%D9%86%DA%AF" title="سوسک‌های رنگارنگ – Persian" lang="fa" hreflang="fa" data-title="سوسک‌های رنگارنگ" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Cleridae" title="Cleridae – French" lang="fr" hreflang="fr" data-title="Cleridae" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%A7%D7%9C%D7%A8%D7%99%D7%AA%D7%99%D7%99%D7%9D" title="קלריתיים – Hebrew" lang="he" hreflang="he" data-title="קלריתיים" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-ka mw-list-item"><a href="https://ka.wikipedia.org/wiki/%E1%83%AD%E1%83%A0%E1%83%94%E1%83%9A%E1%83%90%E1%83%A1%E1%83%94%E1%83%91%E1%83%A0%E1%83%9C%E1%83%98" title="ჭრელასებრნი – Georgian" lang="ka" hreflang="ka" data-title="ჭრელასებრნი" data-language-autonym="ქართული" data-language-local-name="Georgian" class="interlanguage-link-target"><span>ქართული</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Ker%C5%A1vabaliai" title="Keršvabaliai – Lithuanian" lang="lt" hreflang="lt" data-title="Keršvabaliai" data-language-autonym="Lietuvių" data-language-local-name="Lithuanian" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Sz%C3%BAfarkasf%C3%A9l%C3%A9k" title="Szúfarkasfélék – Hungarian" lang="hu" hreflang="hu" data-title="Szúfarkasfélék" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%95%E0%B5%8D%E0%B4%B2%E0%B5%86%E0%B4%B1%E0%B4%BF%E0%B4%A1%E0%B5%87" title="ക്ലെറിഡേ – Malayalam" lang="ml" hreflang="ml" data-title="ക്ലെറിഡേ" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%A8%D9%88%D9%82%D9%8A%D8%A7%D8%AA_%D8%B6%D8%A7%D8%B1%D9%8A%D9%87" title="بوقيات ضاريه – Egyptian Arabic" lang="arz" hreflang="arz" data-title="بوقيات ضاريه" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Mierkevers" title="Mierkevers – Dutch" lang="nl" hreflang="nl" data-title="Mierkevers" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%AB%E3%83%83%E3%82%B3%E3%82%A6%E3%83%A0%E3%82%B7" title="カッコウムシ – Japanese" lang="ja" hreflang="ja" data-title="カッコウムシ" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Maurbiller" title="Maurbiller – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Maurbiller" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Przekraskowate" title="Przekraskowate – Polish" lang="pl" hreflang="pl" data-title="Przekraskowate" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Cleridae" title="Cleridae – Portuguese" lang="pt" hreflang="pt" data-title="Cleridae" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9F%D0%B5%D1%81%D1%82%D1%80%D1%8F%D0%BA%D0%B8" title="Пестряки – Russian" lang="ru" hreflang="ru" data-title="Пестряки" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Kirjokuoriaiset" title="Kirjokuoriaiset – Finnish" lang="fi" hreflang="fi" data-title="Kirjokuoriaiset" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Brokbaggar" title="Brokbaggar – Swedish" lang="sv" hreflang="sv" data-title="Brokbaggar" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Cleridae" title="Cleridae – Turkish" lang="tr" hreflang="tr" data-title="Cleridae" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%A1%D1%82%D1%80%D0%BE%D0%BA%D0%B0%D1%82%D0%BA%D0%B8_(%D0%B6%D1%83%D0%BA%D0%B8)" title="Строкатки (жуки) – Ukrainian" lang="uk" hreflang="uk" data-title="Строкатки (жуки)" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Cleridae" title="Cleridae – Vietnamese" lang="vi" hreflang="vi" data-title="Cleridae" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-war mw-list-item"><a href="https://war.wikipedia.org/wiki/Cleridae" title="Cleridae – Waray" lang="war" hreflang="war" data-title="Cleridae" data-language-autonym="Winaray" data-language-local-name="Waray" class="interlanguage-link-target"><span>Winaray</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E9%83%AD%E5%85%AC%E8%9F%B2%E7%A7%91" title="郭公蟲科 – Chinese" lang="zh" hreflang="zh" data-title="郭公蟲科" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q377988#sitelinks-wikipedia" title="Edit interlanguage links" class="wbc-editpage">Edit links</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Namespaces"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected vector-tab-noicon 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style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">Cleridae<br /><div style="font-size: 85%;">Temporal range: <span class="noprint"><span style="display:inline-block;"></span><span style="display:inline-block;">Middle Jurassic–Recent</span> <span style="display:inline-block;"></span><div id="Timeline-row" style="margin: 4px auto 0; clear:both; width:220px; padding:0px; height:18px; overflow:visible; white-space:nowrap; border:1px #666; border-style:solid none; position:relative; z-index:0; font-size:97%;"> <div style="position:absolute; height:100%; left:0px; width:207.23076923077px; padding-left:5px; text-align:left; background-color:rgb(254,217,106); background-image: linear-gradient(to right, rgba(255,255,255,1), rgba(254,217,106,1) 15%, rgba(254,217,106,1));"><a href="/wiki/Precambrian" title="Precambrian">PreꞒ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,160,86); left:37.636923076923px; width:18.073846153846px;"><a href="/wiki/Cambrian" title="Cambrian">Ꞓ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(0,146,112); left:55.710769230769px; width:14.08px;"><a href="/wiki/Ordovician" title="Ordovician">O</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(179,225,182); left:69.790769230769px; width:8.3261538461539px;"><a href="/wiki/Silurian" title="Silurian">S</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(203,140,55); left:78.116923076923px; width:20.409230769231px;"><a href="/wiki/Devonian" title="Devonian">D</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(103,165,153); left:98.526153846154px; width:20.307692307692px;"><a href="/wiki/Carboniferous" title="Carboniferous">C</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(240,64,40); left:118.83384615385px; width:15.907015384615px;"><a href="/wiki/Permian" title="Permian">P</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(129,43,146); left:134.74086153846px; width:17.092984615385px;"><a href="/wiki/Triassic" title="Triassic">T</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(52,178,201); left:151.83384615385px; width:19.089230769231px;"><a href="/wiki/Jurassic" title="Jurassic">J</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,198,78); left:170.92307692308px; width:26.738461538462px;"><a href="/wiki/Cretaceous" title="Cretaceous">K</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(253,154,82); left:197.66153846154px; width:14.543692307692px;"><a href="/wiki/Paleogene" title="Paleogene">Pg</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(255,230,25); left:212.20523076923px; width:6.9215384615385px;"><a href="/wiki/Neogene" title="Neogene">N</a></div> <div id="end-border" style="position:absolute; height:100%; background-color:#666; width:1px; left:219px"></div><div style="margin:0 auto; line-height:0; clear:both; width:220px; padding:0px; height:8px; overflow:visible; background-color:transparent; position:relative; top:-4px; z-index:100;"><div style="position:absolute; height:8px; left:160.87076923077px; width:59.129230769231px; background-color:#360; opacity:0.42;"></div> <div style="position:absolute; height:8px; left:160.87076923077px; width:59.129230769231px; background-color:#360; opacity:1;"></div> <div style="position:absolute; height:6px; top:1px; left:161.87076923077px; width:57.129230769231px; background-color:#6c3;"></div> </div> </div></span></div> </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:PZSL1852PlateAnnulosa27.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8e/PZSL1852PlateAnnulosa27.png/220px-PZSL1852PlateAnnulosa27.png" decoding="async" width="220" height="363" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8e/PZSL1852PlateAnnulosa27.png/330px-PZSL1852PlateAnnulosa27.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8e/PZSL1852PlateAnnulosa27.png/440px-PZSL1852PlateAnnulosa27.png 2x" data-file-width="1819" data-file-height="2999" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%">Some checkered beetles<br />described in the mid-19th century </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Cleridae" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Arthropod" title="Arthropod">Arthropoda</a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Insect" title="Insect">Insecta</a> </td></tr> <tr> <td>Order: </td> <td><a href="/wiki/Beetle" title="Beetle">Coleoptera</a> </td></tr> <tr> <td>Suborder: </td> <td><a href="/wiki/Polyphaga" title="Polyphaga">Polyphaga</a> </td></tr> <tr> <td>Infraorder: </td> <td><a href="/wiki/Cucujiformia" title="Cucujiformia">Cucujiformia</a> </td></tr> <tr> <td>Superfamily: </td> <td><a href="/wiki/Cleroidea" title="Cleroidea">Cleroidea</a> </td></tr> <tr> <td>Family: </td> <td><a class="mw-selflink selflink">Cleridae</a><br /><small><a href="/wiki/Pierre_Andr%C3%A9_Latreille" title="Pierre André Latreille">Latreille</a>, 1802</small> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">Subfamilies </th></tr> <tr> <td colspan="2" style="text-align: left"> <p><a href="/wiki/Clerinae" title="Clerinae">Clerinae</a> <small>Latreille, 1802</small><br /> <a href="/wiki/Enopliinae" title="Enopliinae">Enopliinae</a> <small>Gistel, 1856 (disputed)</small><br /> <a href="/wiki/Epiphloeinae" title="Epiphloeinae">Epiphloeinae</a> <small>Gistel, 1856 (disputed)</small><br /> <a href="/wiki/Hydnocerinae" title="Hydnocerinae">Hydnocerinae</a> <small>Spinola, 1844</small><br /> <a href="/w/index.php?title=Korynetinae&action=edit&redlink=1" class="new" title="Korynetinae (page does not exist)">Korynetinae</a> <small>Laporte, 1836</small><br /> <a href="/wiki/Tarsosteninae" title="Tarsosteninae">Tarsosteninae</a> <small>Jacquelin du Val, 1861 (disputed)</small><br /> <a href="/wiki/Thaneroclerinae" class="mw-redirect" title="Thaneroclerinae">Thaneroclerinae</a> <small>Chapin, 1921<sup class="noprint Inline-Template" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Verifiability" title="Wikipedia:Verifiability"><span title="The material near this tag needs to be fact-checked with the cited source(s). (April 2011)">verification needed</span></a></i>]</sup> (but see <a href="#Systematics">text</a>)</small><br /> <a href="/wiki/Tillinae" title="Tillinae">Tillinae</a> <small>Leach, 1815</small><br /> and see <a href="#Systematics">below</a> </p> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><b>Cleridae</b> are a family of <a href="/wiki/Beetle" title="Beetle">beetles</a> of the superfamily <a href="/wiki/Cleroidea" title="Cleroidea">Cleroidea</a>. They are commonly known as <b>checkered beetles</b>. The family Cleridae has a worldwide distribution, and a variety of <a href="/wiki/Habitat" title="Habitat">habitats</a> and feeding preferences. </p><p>Cleridae have many <a href="/wiki/Ecological_niche" title="Ecological niche">niches</a> and feeding habits. Most genera are <a href="/wiki/Predaceous" class="mw-redirect" title="Predaceous">predaceous</a> and feed on other beetles and larvae; however other genera are <a href="/wiki/Scavengers" class="mw-redirect" title="Scavengers">scavengers</a> or <a href="/wiki/Pollen" title="Pollen">pollen</a> feeders. Clerids have elongated bodies with bristly hairs, are usually bright colored, and have variable <a href="/wiki/Antenna_(biology)" title="Antenna (biology)">antennae</a>. Checkered beetles range in length between 3 and 24 millimetres (0.12 and 0.94 in). Cleridae can be identified based on their 5–5–5 <a href="/wiki/Tarsal_formula" title="Tarsal formula">tarsal formula</a>, division of <a href="/wiki/Sternite" class="mw-redirect" title="Sternite">sternites</a>, and the absence of a special type of vesicle. Female Cleridae lay between <span style="white-space:nowrap;">28–42</span> <a href="/wiki/Egg_(biology)" class="mw-redirect" title="Egg (biology)">eggs</a> at a time predominately under the <a href="/wiki/Bark_(botany)" title="Bark (botany)">bark</a> of trees. Larvae are predaceous and feed vigorously before pupation and subsequently emergence as adults. </p><p>Clerids have a minor significance in <a href="/wiki/Forensic_entomology" title="Forensic entomology">forensic entomology</a>. Some species are occasionally found on carrion in the later dry stages of <a href="/wiki/Decomposition" title="Decomposition">decay</a>. Also, some species are pests (<a href="/wiki/Stored_product_entomology" class="mw-redirect" title="Stored product entomology">stored product entomology</a>) and are found infesting various food products. Research efforts related to Cleridae have focused primarily on using certain species as biological controls. This is a very effective technique for controlling <a href="/wiki/Bark_beetle" title="Bark beetle">bark beetles</a> due to the voracious appetite of many clerid species. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Description">Description</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cleridae&action=edit&section=1" title="Edit section: Description"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Enoclerus_ichneumoneus.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/89/Enoclerus_ichneumoneus.jpg/170px-Enoclerus_ichneumoneus.jpg" decoding="async" width="170" height="225" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/89/Enoclerus_ichneumoneus.jpg/255px-Enoclerus_ichneumoneus.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/89/Enoclerus_ichneumoneus.jpg/340px-Enoclerus_ichneumoneus.jpg 2x" data-file-width="579" data-file-height="768" /></a><figcaption>Narrow pronotum in <i><a href="/wiki/Enoclerus_ichneumoneus" title="Enoclerus ichneumoneus">Enoclerus ichneumoneus</a></i> (<a href="/wiki/Clerinae" title="Clerinae">Clerinae</a>)</figcaption></figure> <div class="mw-heading mw-heading3"><h3 id="Appearance">Appearance</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cleridae&action=edit&section=2" title="Edit section: Appearance"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Generally, checkered beetles are elongated and <a href="/wiki/Oval" title="Oval">oval</a> in shape and range from 3–24 millimetres (0.12–0.94 in) in length.<sup id="cite_ref-Johnson_1-0" class="reference"><a href="#cite_note-Johnson-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> Their entire bodies are covered with bristly hairs and many display an ornate body color pattern.<sup id="cite_ref-Johnson_1-1" class="reference"><a href="#cite_note-Johnson-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> These often brightly color patterns can be red, yellow, orange, or blue.<sup id="cite_ref-White_2-0" class="reference"><a href="#cite_note-White-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> The antennae are clubbed at the tip for most species, but others can be "clubbed, saw-tooth, or thread-like."<sup id="cite_ref-Johnson_1-2" class="reference"><a href="#cite_note-Johnson-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-White_2-1" class="reference"><a href="#cite_note-White-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> The pronotum region is nearly cylindrical and characteristically narrower than the <a href="/wiki/Elytra" class="mw-redirect" title="Elytra">elytra</a> (special hardened front wings), while the head is as wide or wider than the <a href="/wiki/Prothorax" title="Prothorax">pronotum</a>.<sup id="cite_ref-White_2-2" class="reference"><a href="#cite_note-White-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> Their elytra have tiny pits or depressions, and never expose more than two <a href="/wiki/Tergites" class="mw-redirect" title="Tergites">tergites</a> (<a href="/wiki/Dorsum_(biology)" class="mw-redirect" title="Dorsum (biology)">dorsal</a> plates).<sup id="cite_ref-Johnson_1-3" class="reference"><a href="#cite_note-Johnson-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Identification">Identification</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cleridae&action=edit&section=3" title="Edit section: Identification"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Clerid beetles fall under the suborder <a href="/wiki/Polyphaga" title="Polyphaga">Polyphaga</a>. Key characteristics of Polyphaga are that the hind <a href="/wiki/Arthropod_coxa" class="mw-redirect" title="Arthropod coxa">coxa</a> (base of the leg), do not divide the first and second abdominal/ventral plates which are known as <a href="/wiki/Sternites" class="mw-redirect" title="Sternites">sternites</a>. Also, the <a href="/wiki/Notopleuron" title="Notopleuron">notopleural</a> suture (found under the pronotal shield) is not present.<sup id="cite_ref-Johnson_1-4" class="reference"><a href="#cite_note-Johnson-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> To further identify Clerid beetles, a few additional characteristics need to be examined. </p> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Tillus_elongatus_tarsus.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/f0/Tillus_elongatus_tarsus.jpg/220px-Tillus_elongatus_tarsus.jpg" decoding="async" width="220" height="98" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/f0/Tillus_elongatus_tarsus.jpg/330px-Tillus_elongatus_tarsus.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/f0/Tillus_elongatus_tarsus.jpg/440px-Tillus_elongatus_tarsus.jpg 2x" data-file-width="681" data-file-height="303" /></a><figcaption>5 rear leg tarsomeres of <i><a href="/wiki/Tillus_elongatus" title="Tillus elongatus">Tillus elongatus</a></i> (<a href="/wiki/Tillinae" title="Tillinae">Tillinae</a>)</figcaption></figure> <p>Clerid beetles have unique legs that help to distinguish them from other families. Their <a href="/wiki/Arthropod_leg" title="Arthropod leg">tarsal</a> formula is 5–5–5, meaning that on each of the front, middle and hind legs there are 5 tarsomeres (individual subsegments of the feet/tarsi).<sup id="cite_ref-Johnson_1-5" class="reference"><a href="#cite_note-Johnson-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> One or more of these subsegments on each leg is typically lobed, and the 4th tarsomere is normally difficult to distinguish. Furthermore, an important feature that eliminates many other families of beetles is that clerids' front coxae (base of the leg) expose the second segment of the legs known as the trochanter.<sup id="cite_ref-Johnson_1-6" class="reference"><a href="#cite_note-Johnson-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p><p>The second defining characteristic of the family Cleridae is that clerids never have eversible vesicles (small usually hidden balloon-like structures thought to be scent glands) on their <a href="/wiki/Abdomen" title="Abdomen">abdomen</a> and <a href="/wiki/Pronotum" class="mw-redirect" title="Pronotum">pronotum</a>. This characteristic distinguishes them from a similar family <a href="/wiki/Melyridae" title="Melyridae">Melyridae</a> which sometimes has these glands.<sup id="cite_ref-Johnson_1-7" class="reference"><a href="#cite_note-Johnson-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> This trait is very important in correctly differentiating checkered beetles from Melyridae. </p> <div class="mw-heading mw-heading2"><h2 id="Distribution_and_ecology">Distribution and ecology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cleridae&action=edit&section=4" title="Edit section: Distribution and ecology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Trichodes_ornatus_on_a_flower.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/d4/Trichodes_ornatus_on_a_flower.jpg/220px-Trichodes_ornatus_on_a_flower.jpg" decoding="async" width="220" height="174" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/d4/Trichodes_ornatus_on_a_flower.jpg/330px-Trichodes_ornatus_on_a_flower.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/d4/Trichodes_ornatus_on_a_flower.jpg/440px-Trichodes_ornatus_on_a_flower.jpg 2x" data-file-width="768" data-file-height="606" /></a><figcaption><i><a href="/wiki/Trichodes_ornatus" title="Trichodes ornatus">Trichodes ornatus</a></i> (<a href="/wiki/Clerinae" title="Clerinae">Clerinae</a>) on a flower</figcaption></figure> <p>Cleridae can be found in the Americas, Africa, Europe, the Middle East and even in Australia. There are approximately 3,500 species in the world and about 500 species in North America.<sup id="cite_ref-Byrd_3-0" class="reference"><a href="#cite_note-Byrd-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Due to this wide distribution there are many different <a href="/wiki/Habitats" class="mw-redirect" title="Habitats">habitats</a> in which the checkered beetles can be found. </p><p>Many of the species are known as "flower visitors", that prey on other <a href="/wiki/Flower" title="Flower">flower</a> visiting <a href="/wiki/Insects" class="mw-redirect" title="Insects">insects</a> and also feed on <a href="/wiki/Pollen" title="Pollen">pollen</a>. These species are found in moist, sunny environments where flowering plants are found in abundance.<sup id="cite_ref-Gerstmeier_4-0" class="reference"><a href="#cite_note-Gerstmeier-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p><p>Another habitat commonly inhabited by clerid beetles is <a href="/wiki/Trees" class="mw-redirect" title="Trees">trees</a>. These "tree living species" are found in forests across the world with various climates and an array of easily preyed upon insects. They seek protection under the bark and hunt for other insects above and below the <a href="/wiki/Bark_(botany)" title="Bark (botany)">bark</a>.<sup id="cite_ref-Gerstmeier_4-1" class="reference"><a href="#cite_note-Gerstmeier-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> The primary source of prey for these bark living hunters is bark beetles. </p><p>The third type of clerid beetles is the "nest robbing species" which live in shrubbery and in trees. Unlike the tree living species, these species do not actually <a href="/wiki/Burrow" title="Burrow">burrow</a> into the bark. Nest robbing species typically hunt termite, bee, and wasp larvae, and one particular species has been noted to prey primarily on grasshopper egg masses.<sup id="cite_ref-Gerstmeier_4-2" class="reference"><a href="#cite_note-Gerstmeier-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Not all nest robbing species actively hunt live prey, some species for example prefer to feed only on dead honey bee larvae and adults.<sup id="cite_ref-Gerstmeier_4-3" class="reference"><a href="#cite_note-Gerstmeier-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Feeding_habits">Feeding habits</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cleridae&action=edit&section=5" title="Edit section: Feeding habits"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The Cleridae contains many species of predaceous beetles that feed on other beetles and beetle larvae in their natural habitat.<sup id="cite_ref-Byrd_3-1" class="reference"><a href="#cite_note-Byrd-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> The most common prey item for checkered beetles are bark beetles and wood boring beetles.<sup id="cite_ref-Bellows_5-0" class="reference"><a href="#cite_note-Bellows-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p><p>In general, the bulk of adult Cleridae feed mainly on other adult beetles while the larvae stage feed on other beetle larvae. Some checkered beetles are known to have an extremely voracious appetite with some larvae able to consume "several times their own body weight" in a day.<sup id="cite_ref-U.S_6-0" class="reference"><a href="#cite_note-U.S-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p><p>Although most species of checkered beetles are predaceous in nature, some are scavengers and others have been found feeding on flower pollen.<sup id="cite_ref-McNamara_7-0" class="reference"><a href="#cite_note-McNamara-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> Because of the checkered beetles predaceous nature and insatiable appetite, they are often key players in the <a href="/wiki/Biological_control" class="mw-redirect" title="Biological control">biological control</a> of other insects. The checkered beetles have also developed a unique adaptation to aid in their quest for prey. The beetles use <a href="/wiki/Pheromones" class="mw-redirect" title="Pheromones">pheromones</a> to help them locate, kill, and consume their prey.<sup id="cite_ref-Costello_8-0" class="reference"><a href="#cite_note-Costello-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Necrobia_rufipes.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/87/Necrobia_rufipes.jpg/170px-Necrobia_rufipes.jpg" decoding="async" width="170" height="257" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/87/Necrobia_rufipes.jpg/255px-Necrobia_rufipes.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/87/Necrobia_rufipes.jpg/340px-Necrobia_rufipes.jpg 2x" data-file-width="508" data-file-height="768" /></a><figcaption><i><a href="/wiki/Necrobia_rufipes" title="Necrobia rufipes">Necrobia rufipes</a></i> (<a href="/w/index.php?title=Korynetinae&action=edit&redlink=1" class="new" title="Korynetinae (page does not exist)">Korynetinae</a>)</figcaption></figure> <p>The diversity of checkered beetle's feeding habits is quite evident when different species are examined. <i><a href="/wiki/Necrobia" title="Necrobia">Necrobia</a></i> <a href="/wiki/Species" title="Species">spp.</a> are attracted to dry carrion and other decomposing animal matter such as bones and skin as well as various meat products.<sup id="cite_ref-Majka_9-0" class="reference"><a href="#cite_note-Majka-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> <i><a href="/wiki/Thanasimus" title="Thanasimus">Thanasimus</a></i> spp. are found in <a href="/wiki/Woodland" title="Woodland">woodland</a> areas where bark beetle species constitute their main source of <a href="/wiki/Prey" class="mw-redirect" title="Prey">prey</a>.<sup id="cite_ref-Majka_9-1" class="reference"><a href="#cite_note-Majka-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> The primary source of prey for <i><a href="/wiki/Phyllobaenus" title="Phyllobaenus">Phyllobaenus</a></i> spp. are <a href="/wiki/Wood_borer" class="mw-redirect" title="Wood borer">wood borers</a>, immature <a href="/wiki/Weevils" class="mw-redirect" title="Weevils">weevils</a>, and <a href="/wiki/Hymenoptera" title="Hymenoptera">hymenoptera</a> larvae.<sup id="cite_ref-Majka_9-2" class="reference"><a href="#cite_note-Majka-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> One of the more diverse <a href="/wiki/Genera" class="mw-redirect" title="Genera">genera</a> is <i><a href="/wiki/Trichodes" title="Trichodes">Trichodes</a></i>, in which larvae feed on the pollen of flowering plants and adults prey upon grasshoppers and wasps.<sup id="cite_ref-Majka_9-3" class="reference"><a href="#cite_note-Majka-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Life_cycle">Life cycle</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cleridae&action=edit&section=6" title="Edit section: Life cycle"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The general life cycle of clerids has been known to last anywhere from 35 days to more than 3 years, and is strongly dependent on the life cycle of their prey.<sup id="cite_ref-Gredilha_10-0" class="reference"><a href="#cite_note-Gredilha-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> While the life cycle can vary in length between genus and species, temperature is also a major determinant in the length of time spent in each stage of development. The warmer the temperature is, the quicker the life cycle, and the cooler the temperature is the slower the life cycle. If temperatures dip below a threshold temperature for an extended period of time, clerids and most other insects will have growth and developmental progress arrested. Like all beetles, Cleridae follow a <a href="/wiki/Holometabolism" title="Holometabolism">holometabolous</a> life cycle: the egg hatches into a <a href="/wiki/Larva" title="Larva">larva</a>, which grows and feeds, changing its skin to form a <a href="/wiki/Pupa" title="Pupa">pupa</a>, and the pupa shedding its skin to emerge as an adult. The larvae of the majority of the known species of Cleridae feed upon the eggs and young of <a href="/wiki/Wood-boring_beetle" class="mw-redirect" title="Wood-boring beetle">wood-boring beetles</a>, while the adults feed on the adult bark beetles.<sup id="cite_ref-Mahr_11-0" class="reference"><a href="#cite_note-Mahr-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> </p> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Thanasimus_dubius_larva.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/f7/Thanasimus_dubius_larva.jpg/220px-Thanasimus_dubius_larva.jpg" decoding="async" width="220" height="147" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/f7/Thanasimus_dubius_larva.jpg/330px-Thanasimus_dubius_larva.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/f7/Thanasimus_dubius_larva.jpg/440px-Thanasimus_dubius_larva.jpg 2x" data-file-width="768" data-file-height="512" /></a><figcaption>Larva of <i><a href="/wiki/Thanasimus_dubius" title="Thanasimus dubius">Thanasimus dubius</a></i></figcaption></figure> <p><a href="/wiki/Copulation_(zoology)" title="Copulation (zoology)">Copulation</a> takes place while the female feeds, because females need a large amount of food for egg development.<sup id="cite_ref-Gorton_12-0" class="reference"><a href="#cite_note-Gorton-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> The female lays her eggs 36–72 hours after copulation. The eggs are laid in between pieces of bark on wood-borer-infested trees or under stones in the soil.<sup id="cite_ref-Mahr_11-1" class="reference"><a href="#cite_note-Mahr-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> She may lay 28–42 eggs at a time.<sup id="cite_ref-Gorton_12-1" class="reference"><a href="#cite_note-Gorton-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> For the longer-living species such as <i><a href="/wiki/Thanasimus" title="Thanasimus">Thanasimus</a>,</i> this occurs in late summer or early fall to give the <a href="/wiki/Larva" title="Larva">larvae</a> enough time for proper growth before having to <a href="/wiki/Overwinter" class="mw-redirect" title="Overwinter">overwinter</a>.<sup id="cite_ref-Ye_13-0" class="reference"><a href="#cite_note-Ye-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p><p>When larvae hatch from their eggs, they are either red or yellow.<sup id="cite_ref-Mahr_11-2" class="reference"><a href="#cite_note-Mahr-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> Their bodies have a slender and flat appearance with short legs due to their minimal movement. The larvae are covered in hair and have two horn-type projections on the <a href="/wiki/Dorsum_(biology)" class="mw-redirect" title="Dorsum (biology)">dorsal</a> area of the last body segment.<sup id="cite_ref-Mahr_11-3" class="reference"><a href="#cite_note-Mahr-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> Immediately after birth, they start searching for food close to where they hatched. They feed on wood-borer insects on trees, or feed on their species' substrate or prey of choice.<sup id="cite_ref-Mahr_11-4" class="reference"><a href="#cite_note-Mahr-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> Feeding is the main purpose of the larval stage to prepare for pupation. Once their larval stage is complete the tree dwelling species make their way to the bottom of the tree to pupate.<sup id="cite_ref-Ye_13-1" class="reference"><a href="#cite_note-Ye-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> The <a href="/wiki/Pupa" title="Pupa">pupal</a> stage can last from 6 weeks to one year depending on the need to overwinter, and how short the overall life cycle is for a particular species. A majority of clerid species pupate in earthen cells which are made from soil and certain enzymes secreted from their mouths.<sup id="cite_ref-Mahr_11-5" class="reference"><a href="#cite_note-Mahr-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> The rest remain in pupal cells. Adult beetles emerge from pupation and spend a variable time of their life maturing, and eventually <a href="/wiki/Oviposition" class="mw-redirect" title="Oviposition">oviposit</a>. Sexually mature adults or <a href="/wiki/Imago" title="Imago">imagos</a> of <i>Thanasiumus</i> overwinter inside the wood-borer-infested trees and oviposit during the <a href="/wiki/Spring_(season)" title="Spring (season)">spring</a>.<sup id="cite_ref-Gorton_12-2" class="reference"><a href="#cite_note-Gorton-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Forensic_relevance">Forensic relevance</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cleridae&action=edit&section=7" title="Edit section: Forensic relevance"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Stored_product_entomology">Stored product entomology</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cleridae&action=edit&section=8" title="Edit section: Stored product entomology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><i><a href="/wiki/Necrobia_rufipes" title="Necrobia rufipes">Necrobia rufipes</a></i>, commonly known as the red-legged ham beetle, is of particular importance in <a href="/wiki/Stored_product_entomology" class="mw-redirect" title="Stored product entomology">stored product entomology</a>. It infests <a href="/wiki/Dried_meat" title="Dried meat">dried</a> or <a href="/wiki/Smoked_meats" class="mw-redirect" title="Smoked meats">smoked meats</a>, especially those products that are stored unwrapped for long periods of time. Adults feed on the surface of the products, while the larvae damage the meat by boring down usually in the fatty parts.<sup id="cite_ref-Ebeling_14-0" class="reference"><a href="#cite_note-Ebeling-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> <i>Necrobia rufipes</i> has been recorded feeding upon a large variety of items ranging from hides and dried figs to Egyptian mummies.<sup id="cite_ref-Ebeling_14-1" class="reference"><a href="#cite_note-Ebeling-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> In addition, products such as <a href="/wiki/Wool" title="Wool">wool</a> and <a href="/wiki/Silk" title="Silk">silk</a> can become infested, but not destroyed.<sup id="cite_ref-Ebeling_14-2" class="reference"><a href="#cite_note-Ebeling-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Medico-legal_entomology">Medico-legal entomology</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cleridae&action=edit&section=9" title="Edit section: Medico-legal entomology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Since clerids are predaceous in nature, they have been found feeding on fly larvae as well as the skin and bones of <a href="/wiki/Carrion" title="Carrion">carrion</a>.<sup id="cite_ref-Majka_9-4" class="reference"><a href="#cite_note-Majka-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> Most clerids are not useful in <a href="/wiki/Forensics" class="mw-redirect" title="Forensics">forensics</a> because of their food choice, but some species such as <i>Necrobia rufipes</i> can be useful. <i>Necrobia rufipes</i> is attracted towards carrion in the later stages of decomposition, so its arrival on carrion can help provide an estimate for the <a href="/wiki/Post-mortem_interval" title="Post-mortem interval">post-mortem interval</a> or PMI. Although the checkered beetle is not the most significant insect on carrion, the beetle's predaceous nature and its ability to reproduce in carrion that is exposed to the environment provides some forensic importance.<sup id="cite_ref-Souza_15-0" class="reference"><a href="#cite_note-Souza-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> </p> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Thanasimus_dubius_Attacking_Prey.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/80/Thanasimus_dubius_Attacking_Prey.jpg/220px-Thanasimus_dubius_Attacking_Prey.jpg" decoding="async" width="220" height="147" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/80/Thanasimus_dubius_Attacking_Prey.jpg/330px-Thanasimus_dubius_Attacking_Prey.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/80/Thanasimus_dubius_Attacking_Prey.jpg/440px-Thanasimus_dubius_Attacking_Prey.jpg 2x" data-file-width="768" data-file-height="512" /></a><figcaption><i><a href="/wiki/Thanasimus_dubius" title="Thanasimus dubius">Thanasimus dubius</a></i> attacking bark beetle prey</figcaption></figure> <div class="mw-heading mw-heading2"><h2 id="Ongoing_research">Ongoing research</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cleridae&action=edit&section=10" title="Edit section: Ongoing research"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>There is ongoing research with some clerid species. Forensic research is limited because of their late arrival on carrion, but members such as <i><a href="/wiki/Thanasimus_undatulus" title="Thanasimus undatulus">Thanasimus undatulus</a></i> have been researched as a possible role in <a href="/wiki/Integrated_pest_management" title="Integrated pest management">integrated pest management</a> or IPM. <i>Thanasimus undatulus</i> is a predator of bark beetles. Some species of bark beetles such as the <a href="/wiki/Southern_pine_beetle" class="mw-redirect" title="Southern pine beetle">southern pine beetle</a> and the <a href="/wiki/Mountain_pine_beetle" title="Mountain pine beetle">mountain pine beetle</a> can become pests to the <a href="/wiki/Lumber" title="Lumber">lumber</a> industry because in large numbers they can cause damage and kill live trees. <i>Thanasimus undatulus</i> has been researched as a possible <a href="/wiki/Biological_control" class="mw-redirect" title="Biological control">biological control</a> agent for these pests. Researchers and forestry officials have used bark beetle aggregation pheromones to attract the checkered beetle to specific trees. This causes the bark beetles to be overwhelmed, extensively preyed upon by the clerid beetles, and typically eliminated.<sup id="cite_ref-Poland_16-0" class="reference"><a href="#cite_note-Poland-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> There is also additional research being done pertaining to the impact of clerids on pollination in flowers.<sup id="cite_ref-Ecological_17-0" class="reference"><a href="#cite_note-Ecological-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Systematics">Systematics</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cleridae&action=edit&section=11" title="Edit section: Systematics"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The <a href="/wiki/Genera" class="mw-redirect" title="Genera">genera</a> of Cleridae are divided among several <a href="/wiki/Subfamilies" class="mw-redirect" title="Subfamilies">subfamilies</a>, though some genera still defy easy classification. Several <a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">taxonomic</a> schemes exist, recognizing for example a group around <i><a href="/wiki/Neorthopleura" title="Neorthopleura">Neorthopleura</a></i> as distinct subfamily <a href="/w/index.php?title=Neorthopleurinae&action=edit&redlink=1" class="new" title="Neorthopleurinae (page does not exist)">Neorthopleurinae</a>, or splitting off the Thaneroclerinae as distinct family, or circumscribing the Korynetinae <i><a href="/wiki/Sensu_stricto" class="mw-redirect" title="Sensu stricto">sensu stricto</a></i> or <i><a href="/wiki/Sensu_lato" class="mw-redirect" title="Sensu lato">sensu lato</a></i>. The following list of tribes and selected genera is thus preliminary. Some notable species are also listed. The oldest members of the family are <i><a href="/w/index.php?title=Protoclerus&action=edit&redlink=1" class="new" title="Protoclerus (page does not exist)">Protoclerus</a></i> and <i><a href="/w/index.php?title=Wangweiella&action=edit&redlink=1" class="new" title="Wangweiella (page does not exist)">Wangweiella</a></i> the late <a href="/wiki/Middle_Jurassic" title="Middle Jurassic">Middle Jurassic</a> (<a href="/wiki/Callovian" title="Callovian">Callovian</a>) <a href="/wiki/Tiaojishan_Formation" title="Tiaojishan Formation">Daohugou bed</a> in Inner Mongolia, China.<sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> </p> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Clerus_mutillarius01.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/0/0d/Clerus_mutillarius01.jpg/220px-Clerus_mutillarius01.jpg" decoding="async" width="220" height="293" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/0/0d/Clerus_mutillarius01.jpg 1.5x" data-file-width="278" data-file-height="370" /></a><figcaption><i><a href="/wiki/Clerus_mutillarius" title="Clerus mutillarius">Clerus mutillarius</a></i> (<a href="/wiki/Clerinae" title="Clerinae">Clerinae</a>)</figcaption></figure> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Phymatophaea_guttigera.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/6/65/Phymatophaea_guttigera.jpg/220px-Phymatophaea_guttigera.jpg" decoding="async" width="220" height="294" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/65/Phymatophaea_guttigera.jpg/330px-Phymatophaea_guttigera.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/65/Phymatophaea_guttigera.jpg/440px-Phymatophaea_guttigera.jpg 2x" data-file-width="1040" data-file-height="1392" /></a><figcaption><i><a href="/w/index.php?title=Phymatophaea_guttigera&action=edit&redlink=1" class="new" title="Phymatophaea guttigera (page does not exist)">Phymatophaea guttigera</a></i> (<a href="/wiki/Enopliinae" title="Enopliinae">Enopliinae</a>)</figcaption></figure> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Lemidia_aptera.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/38/Lemidia_aptera.jpg/220px-Lemidia_aptera.jpg" decoding="async" width="220" height="294" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/38/Lemidia_aptera.jpg/330px-Lemidia_aptera.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/38/Lemidia_aptera.jpg/440px-Lemidia_aptera.jpg 2x" data-file-width="1040" data-file-height="1392" /></a><figcaption><i><a href="/w/index.php?title=Lemidia_aptera&action=edit&redlink=1" class="new" title="Lemidia aptera (page does not exist)">Lemidia aptera</a></i> (<a href="/wiki/Hydnocerinae" title="Hydnocerinae">Hydnocerinae</a>)</figcaption></figure> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Necrobia_violacea.JPG" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e5/Necrobia_violacea.JPG/220px-Necrobia_violacea.JPG" decoding="async" width="220" height="323" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/e5/Necrobia_violacea.JPG/330px-Necrobia_violacea.JPG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/e5/Necrobia_violacea.JPG/440px-Necrobia_violacea.JPG 2x" data-file-width="570" data-file-height="838" /></a><figcaption><i><a href="/wiki/Necrobia_violacea" title="Necrobia violacea">Necrobia violacea</a></i> (<a href="/w/index.php?title=Korynetinae&action=edit&redlink=1" class="new" title="Korynetinae (page does not exist)">Korynetinae</a>)</figcaption></figure> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Tarsostenus_univittatus-Curtis.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/24/Tarsostenus_univittatus-Curtis.png/220px-Tarsostenus_univittatus-Curtis.png" decoding="async" width="220" height="237" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/24/Tarsostenus_univittatus-Curtis.png/330px-Tarsostenus_univittatus-Curtis.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/24/Tarsostenus_univittatus-Curtis.png/440px-Tarsostenus_univittatus-Curtis.png 2x" data-file-width="650" data-file-height="700" /></a><figcaption><i><a href="/w/index.php?title=Tarsostenus_univittatus&action=edit&redlink=1" class="new" title="Tarsostenus univittatus (page does not exist)">Tarsostenus univittatus</a></i> (<a href="/wiki/Tarsosteninae" title="Tarsosteninae">Tarsosteninae</a>)</figcaption></figure> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Diplocladus_kuwerti.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e7/Diplocladus_kuwerti.jpg/220px-Diplocladus_kuwerti.jpg" decoding="async" width="220" height="366" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/e7/Diplocladus_kuwerti.jpg/330px-Diplocladus_kuwerti.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/e7/Diplocladus_kuwerti.jpg/440px-Diplocladus_kuwerti.jpg 2x" data-file-width="1746" data-file-height="2907" /></a><figcaption><i><a href="/wiki/Diplocladus_kuwerti" title="Diplocladus kuwerti">Diplocladus kuwerti</a></i> (<a href="/wiki/Tillinae" title="Tillinae">Tillinae</a>)</figcaption></figure> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Dermestoides.sanguinicollis.-.calwer.28.14.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c8/Dermestoides.sanguinicollis.-.calwer.28.14.jpg/220px-Dermestoides.sanguinicollis.-.calwer.28.14.jpg" decoding="async" width="220" height="283" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/c/c8/Dermestoides.sanguinicollis.-.calwer.28.14.jpg 1.5x" data-file-width="261" data-file-height="336" /></a><figcaption><i><a href="/w/index.php?title=Dermestoides_sanguinicollis&action=edit&redlink=1" class="new" title="Dermestoides sanguinicollis (page does not exist)">Dermestoides sanguinicollis</a></i> (<i><a href="/wiki/Incertae_sedis" title="Incertae sedis">incertae sedis</a></i>)</figcaption></figure> <div> <style data-mw-deduplicate="TemplateStyles:r1216972533">.mw-parser-output .col-begin{border-collapse:collapse;padding:0;color:inherit;width:100%;border:0;margin:0}.mw-parser-output .col-begin-small{font-size:90%}.mw-parser-output .col-break{vertical-align:top;text-align:left}.mw-parser-output .col-break-2{width:50%}.mw-parser-output .col-break-3{width:33.3%}.mw-parser-output .col-break-4{width:25%}.mw-parser-output .col-break-5{width:20%}@media(max-width:720px){.mw-parser-output .col-begin,.mw-parser-output .col-begin>tbody,.mw-parser-output .col-begin>tbody>tr,.mw-parser-output .col-begin>tbody>tr>td{display:block!important;width:100%!important}.mw-parser-output .col-break{padding-left:0!important}}</style><div> <table class="col-begin" role="presentation" style="width: 75%;"> <tbody><tr> <td class="col-break col-break-3"> <p><b><a href="/wiki/Clerinae" title="Clerinae">Clerinae</a></b> </p> <ul><li><i><a href="/wiki/Allonyx" title="Allonyx">Allonyx</a></i> <small>Jacquelin du Val, 1860</small></li> <li><i><a href="/w/index.php?title=Anthicoclerus&action=edit&redlink=1" class="new" title="Anthicoclerus (page does not exist)">Anthicoclerus</a></i> <small>Schenkling, 1906</small></li> <li><i><a href="/wiki/Aphelochroa" title="Aphelochroa">Aphelochroa</a></i> <small>Quedenfeldt, 1885</small></li> <li><i><a href="/w/index.php?title=Apopempsis&action=edit&redlink=1" class="new" title="Apopempsis (page does not exist)">Apopempsis</a></i> <small>Schenkling, 1903</small></li> <li><i><a href="/w/index.php?title=Apteroclerus&action=edit&redlink=1" class="new" title="Apteroclerus (page does not exist)">Apteroclerus</a></i> <small>Wollaston, 1867</small></li> <li><i><a href="/w/index.php?title=Aptinoclerus&action=edit&redlink=1" class="new" title="Aptinoclerus (page does not exist)">Aptinoclerus</a></i> <small><a href="/wiki/August_Ferdinand_Kuwert" title="August Ferdinand Kuwert">Kuwert</a>, 1893</small></li> <li><i><a href="/w/index.php?title=Aradamicula&action=edit&redlink=1" class="new" title="Aradamicula (page does not exist)">Aradamicula</a></i> <small>Sedlacek & Winkler, 1975</small></li> <li>†<i><a href="/w/index.php?title=Arawakis&action=edit&redlink=1" class="new" title="Arawakis (page does not exist)">Arawakis</a></i> (<a href="/wiki/Fossil" title="Fossil">fossil</a>)</li> <li><i><a href="/w/index.php?title=Astigmus&action=edit&redlink=1" class="new" title="Astigmus (page does not exist)">Astigmus</a></i> <small>Kuwert, 1894</small></li> <li><i><a href="/wiki/Aulicus" title="Aulicus">Aulicus</a></i> <small>Spinola, 1841</small></li> <li><i><a href="/w/index.php?title=Axina&action=edit&redlink=1" class="new" title="Axina (page does not exist)">Axina</a></i> <small>Kirby, 1818</small></li> <li><i><a href="/w/index.php?title=Balcus&action=edit&redlink=1" class="new" title="Balcus (page does not exist)">Balcus</a></i></li> <li><i><a href="/w/index.php?title=Barriella&action=edit&redlink=1" class="new" title="Barriella (page does not exist)">Barriella</a></i> <small>Opitz, 2003</small></li> <li><i><a href="/w/index.php?title=Barrotillus&action=edit&redlink=1" class="new" title="Barrotillus (page does not exist)">Barrotillus</a></i> <small>Rifkind, 1996</small></li> <li><i><a href="/w/index.php?title=Blaxima&action=edit&redlink=1" class="new" title="Blaxima (page does not exist)">Blaxima</a></i> <small>Gorham, 1882</small></li> <li><i><a href="/w/index.php?title=Bousquetoclerus&action=edit&redlink=1" class="new" title="Bousquetoclerus (page does not exist)">Bousquetoclerus</a></i> <small>Menier, 1997</small></li> <li><i><a href="/w/index.php?title=Burgeoneus&action=edit&redlink=1" class="new" title="Burgeoneus (page does not exist)">Burgeoneus</a></i> <small>Pic, 1950</small></li> <li><i><a href="/w/index.php?title=Caestron&action=edit&redlink=1" class="new" title="Caestron (page does not exist)">Caestron</a></i> <small>Dupont in Spinola, 1844</small></li> <li><i><a href="/w/index.php?title=Calendyma&action=edit&redlink=1" class="new" title="Calendyma (page does not exist)">Calendyma</a></i> <small>Lacordaire, 1857</small></li> <li><i><a href="/w/index.php?title=Canariclerus&action=edit&redlink=1" class="new" title="Canariclerus (page does not exist)">Canariclerus</a></i> <small>Winkler, 1982</small></li> <li><i><a href="/w/index.php?title=Cardiostichus&action=edit&redlink=1" class="new" title="Cardiostichus (page does not exist)">Cardiostichus</a></i> <small>Quedenfeldt, 1885</small></li> <li><i><a href="/w/index.php?title=Caridopus&action=edit&redlink=1" class="new" title="Caridopus (page does not exist)">Caridopus</a></i> <small>Schenkling in Sjöstedt, 1908</small></li> <li><i><a href="/w/index.php?title=Cleromorpha&action=edit&redlink=1" class="new" title="Cleromorpha (page does not exist)">Cleromorpha</a></i> <small>Gorham, 1876</small></li> <li><i><a href="/w/index.php?title=Cleropiestus&action=edit&redlink=1" class="new" title="Cleropiestus (page does not exist)">Cleropiestus</a></i> <small>Fairmaire, 1889</small></li> <li><i><a href="/wiki/Clerus" title="Clerus">Clerus</a></i> <small>Fabricius, 1775</small></li> <li><i><a href="/w/index.php?title=Clytomadius&action=edit&redlink=1" class="new" title="Clytomadius (page does not exist)">Clytomadius</a></i> <small><a href="/wiki/Johannes_Bastiaan_Corporaal" title="Johannes Bastiaan Corporaal">Corporaal</a>, 1949</small></li> <li><i><a href="/w/index.php?title=Colyphus&action=edit&redlink=1" class="new" title="Colyphus (page does not exist)">Colyphus</a></i> <small>Spinola, 1841</small></li> <li><i><a href="/w/index.php?title=Coptoclerus&action=edit&redlink=1" class="new" title="Coptoclerus (page does not exist)">Coptoclerus</a></i> <small>Chapin, 1924</small></li> <li><i><a href="/w/index.php?title=Cormodes&action=edit&redlink=1" class="new" title="Cormodes (page does not exist)">Cormodes</a></i> <small>Pascoe, 1860</small></li> <li><i><a href="/w/index.php?title=Corynommadius&action=edit&redlink=1" class="new" title="Corynommadius (page does not exist)">Corynommadius</a></i> <small>Schenkling, 1899</small></li> <li><i><a href="/w/index.php?title=Ctenaxina&action=edit&redlink=1" class="new" title="Ctenaxina (page does not exist)">Ctenaxina</a></i> <small>Schenkling, 1906</small></li> <li><i><a href="/w/index.php?title=Ctenoclerus&action=edit&redlink=1" class="new" title="Ctenoclerus (page does not exist)">Ctenoclerus</a></i> <small>Solervicens, 1997</small></li> <li><i><a href="/w/index.php?title=Dasyceroclerus&action=edit&redlink=1" class="new" title="Dasyceroclerus (page does not exist)">Dasyceroclerus</a></i> <small>Kuwert, 1894</small></li> <li><i><a href="/w/index.php?title=Dasyteneclines&action=edit&redlink=1" class="new" title="Dasyteneclines (page does not exist)">Dasyteneclines</a></i> <small>Pic, 1941</small></li> <li><i><a href="/w/index.php?title=Dieropsis&action=edit&redlink=1" class="new" title="Dieropsis (page does not exist)">Dieropsis</a></i> <small>Gahan, 1908</small></li> <li><i><a href="/w/index.php?title=Dologenitus&action=edit&redlink=1" class="new" title="Dologenitus (page does not exist)">Dologenitus</a></i> <small>Opitz, 2009</small></li> <li><i><a href="/w/index.php?title=Dozocolletus&action=edit&redlink=1" class="new" title="Dozocolletus (page does not exist)">Dozocolletus</a></i> <small>Chevrolat, 1842</small></li> <li><i><a href="/w/index.php?title=Eburiphora&action=edit&redlink=1" class="new" title="Eburiphora (page does not exist)">Eburiphora</a></i> <small>Spinola, 1841</small></li> <li><i><a href="/w/index.php?title=Eburneoclerus&action=edit&redlink=1" class="new" title="Eburneoclerus (page does not exist)">Eburneoclerus</a></i> <small>Pic, 1950</small></li> <li><i><a href="/w/index.php?title=Ekisius&action=edit&redlink=1" class="new" title="Ekisius (page does not exist)">Ekisius</a></i> <small>Winkler, 1987</small></li> <li><i><a href="/w/index.php?title=Eleale&action=edit&redlink=1" class="new" title="Eleale (page does not exist)">Eleale</a></i> <small>Newman, 1840</small></li> <li><i><a href="/wiki/Enoclerus" title="Enoclerus">Enoclerus</a></i> <small>Gahan, 1910</small></li> <li><i><a href="/wiki/Epiclines" title="Epiclines">Epiclines</a></i> <small>Chevrolat in Guérin-Ménéville, 1839</small></li> <li><i><a href="/w/index.php?title=Eunatalis&action=edit&redlink=1" class="new" title="Eunatalis (page does not exist)">Eunatalis</a></i> <small>Schenkling, 1909</small> <ul><li><i><a href="/wiki/Eunatalis_porcata" title="Eunatalis porcata">Eunatalis porcata</a></i></li></ul></li> <li><i><a href="/wiki/Erymanthus_(beetle)" title="Erymanthus (beetle)">Erymanthus</a></i> <small>Spinola, 1841</small></li> <li><i><a href="/w/index.php?title=Eurymetomorphon&action=edit&redlink=1" class="new" title="Eurymetomorphon (page does not exist)">Eurymetomorphon</a></i> <small>Pic, 1950</small></li> <li><i><a href="/w/index.php?title=Falsomadius&action=edit&redlink=1" class="new" title="Falsomadius (page does not exist)">Falsomadius</a></i> <small>Gerstmeier, 2002</small></li> <li><i><a href="/w/index.php?title=Falsoorthrius&action=edit&redlink=1" class="new" title="Falsoorthrius (page does not exist)">Falsoorthrius</a></i> <small>Pic, 1940</small></li> <li><i><a href="/w/index.php?title=Graptoclerus&action=edit&redlink=1" class="new" title="Graptoclerus (page does not exist)">Graptoclerus</a></i> <small>Gorham, 1901</small></li> <li><i><a href="/wiki/Gyponyx" title="Gyponyx">Gyponyx</a></i> <small>Gorham, 1883</small></li> <li><i><a href="/w/index.php?title=Hemitrachys&action=edit&redlink=1" class="new" title="Hemitrachys (page does not exist)">Hemitrachys</a></i> <small>Gorham, 1876</small></li> <li><i><a href="/w/index.php?title=Homalopilo&action=edit&redlink=1" class="new" title="Homalopilo (page does not exist)">Homalopilo</a></i> <small>Schenkling, 1915</small></li> <li><i><a href="/w/index.php?title=Inhumeroclerus&action=edit&redlink=1" class="new" title="Inhumeroclerus (page does not exist)">Inhumeroclerus</a></i> <small>Pic, 1955</small></li> <li><i><a href="/w/index.php?title=Jenjouristia&action=edit&redlink=1" class="new" title="Jenjouristia (page does not exist)">Jenjouristia</a></i> <small>Fursov, 1936</small></li> <li><i><a href="/w/index.php?title=Languropilus&action=edit&redlink=1" class="new" title="Languropilus (page does not exist)">Languropilus</a></i> <small>Pic, 1940</small></li> <li><i><a href="/w/index.php?title=Lissaulicus&action=edit&redlink=1" class="new" title="Lissaulicus (page does not exist)">Lissaulicus</a></i> <small>C.O.Waterhouse, 1879</small></li> <li><i><a href="/w/index.php?title=Memorthrius&action=edit&redlink=1" class="new" title="Memorthrius (page does not exist)">Memorthrius</a></i> <small>Pic, 1940</small></li> <li><i><a href="/w/index.php?title=Metademius&action=edit&redlink=1" class="new" title="Metademius (page does not exist)">Metademius</a></i> <small>Schenkling, 1899</small></li> <li><i><a href="/w/index.php?title=Microclerus&action=edit&redlink=1" class="new" title="Microclerus (page does not exist)">Microclerus</a></i> <small>Wollaston, 1867</small></li> <li><i><a href="/w/index.php?title=Micropteroclerus&action=edit&redlink=1" class="new" title="Micropteroclerus (page does not exist)">Micropteroclerus</a></i> <small>Chapin, 1920</small></li> <li><i><a href="/w/index.php?title=Microstigmatium&action=edit&redlink=1" class="new" title="Microstigmatium (page does not exist)">Microstigmatium</a></i> <small>Kraatz, 1899</small></li> <li><i><a href="/w/index.php?title=Mimolesterus&action=edit&redlink=1" class="new" title="Mimolesterus (page does not exist)">Mimolesterus</a></i> <small>Gerstmeier, 1991</small></li> <li><i><a href="/w/index.php?title=Mitrandiria&action=edit&redlink=1" class="new" title="Mitrandiria (page does not exist)">Mitrandiria</a></i> <small>Kolibac, 1997</small></li> <li><i><a href="/w/index.php?title=Myrmecomaea&action=edit&redlink=1" class="new" title="Myrmecomaea (page does not exist)">Myrmecomaea</a></i> <small>Fairmaire, 1886</small></li> <li><i><a href="/w/index.php?title=Natalis_(beetle)&action=edit&redlink=1" class="new" title="Natalis (beetle) (page does not exist)">Natalis</a></i> <small>Laporte de Castelnau, 1836</small></li> <li><i><a href="/w/index.php?title=Neogyponyx&action=edit&redlink=1" class="new" title="Neogyponyx (page does not exist)">Neogyponyx</a></i> <small>Schenkling, 1906</small></li> <li><i><a href="/w/index.php?title=Neoscrobiger&action=edit&redlink=1" class="new" title="Neoscrobiger (page does not exist)">Neoscrobiger</a></i> <small>Blackburn, 1900</small></li> <li><i><a href="/w/index.php?title=Ohanlonella&action=edit&redlink=1" class="new" title="Ohanlonella (page does not exist)">Ohanlonella</a></i> <small>Rifkind, 2008</small></li> <li><i><a href="/w/index.php?title=Olesterus&action=edit&redlink=1" class="new" title="Olesterus (page does not exist)">Olesterus</a></i> <small>Spinola, 1841</small></li> <li><i><a href="/wiki/Omadius" title="Omadius">Omadius</a></i> <small>Laporte de Castelnau, 1836</small></li> <li><i><a href="/w/index.php?title=Oodontophlogistus&action=edit&redlink=1" class="new" title="Oodontophlogistus (page does not exist)">Oodontophlogistus</a></i> <small>Elston, 1923</small></li> <li><i><a href="/w/index.php?title=Operculiphorus&action=edit&redlink=1" class="new" title="Operculiphorus (page does not exist)">Operculiphorus</a></i> <small>Kuwert, 1894</small></li> <li><i><a href="/wiki/Opilo" title="Opilo">Opilo</a></i> <small>Latreille, 1802</small></li> <li><i><a href="/wiki/Orthrius" title="Orthrius">Orthrius</a></i> <small>Gorham, 1876</small></li> <li><i><a href="/w/index.php?title=Oxystigmatium&action=edit&redlink=1" class="new" title="Oxystigmatium (page does not exist)">Oxystigmatium</a></i> <small>Kraatz, 1899</small></li> <li><i><a href="/w/index.php?title=Phlogistomorpha&action=edit&redlink=1" class="new" title="Phlogistomorpha (page does not exist)">Phlogistomorpha</a></i> <small>Hintz, 1908</small></li> <li><i><a href="/w/index.php?title=Phlogistus&action=edit&redlink=1" class="new" title="Phlogistus (page does not exist)">Phlogistus</a></i> <small>Gorham, 1876</small></li> <li><i><a href="/w/index.php?title=Phloiocopus&action=edit&redlink=1" class="new" title="Phloiocopus (page does not exist)">Phloiocopus</a></i> <small>Spinola, 1841</small></li> <li><i><a href="/w/index.php?title=Phonius&action=edit&redlink=1" class="new" title="Phonius (page does not exist)">Phonius</a></i> <small>Chevrolat, 1843</small></li> <li><i><a href="/w/index.php?title=Pieleus&action=edit&redlink=1" class="new" title="Pieleus (page does not exist)">Pieleus</a></i> <small>Pic, 1940</small></li> <li><i><a href="/w/index.php?title=Placocerus&action=edit&redlink=1" class="new" title="Placocerus (page does not exist)">Placocerus</a></i> <small>Klug, 1837</small></li> <li><i><a href="/wiki/Placopterus" title="Placopterus">Placopterus</a></i> <small>Wolcott, 1910</small></li> <li><i><a href="/w/index.php?title=Plathanocera&action=edit&redlink=1" class="new" title="Plathanocera (page does not exist)">Plathanocera</a></i> <small>Schenkling, 1902</small></li> <li><i><a href="/w/index.php?title=Platyclerus&action=edit&redlink=1" class="new" title="Platyclerus (page does not exist)">Platyclerus</a></i> <small>Spinola, 1841</small></li> <li><i><a href="/wiki/Priocera" title="Priocera">Priocera</a></i> <small>Kirby, 1818</small></li> <li><i><a href="/w/index.php?title=Priocleromorphus&action=edit&redlink=1" class="new" title="Priocleromorphus (page does not exist)">Priocleromorphus</a></i> <small>Pic, 1950</small></li> <li><i><a href="/w/index.php?title=Prioclerus&action=edit&redlink=1" class="new" title="Prioclerus (page does not exist)">Prioclerus</a></i> <small>Hintz, 1902</small></li> <li><i><a href="/w/index.php?title=Pseudolesterus&action=edit&redlink=1" class="new" title="Pseudolesterus (page does not exist)">Pseudolesterus</a></i> <small>Miyatake, 1968</small></li> <li><i><a href="/w/index.php?title=Pseudomadius&action=edit&redlink=1" class="new" title="Pseudomadius (page does not exist)">Pseudomadius</a></i> <small>Chapin, 1924</small></li> <li><i><a href="/w/index.php?title=Pujoliclerus&action=edit&redlink=1" class="new" title="Pujoliclerus (page does not exist)">Pujoliclerus</a></i> <small>Pic, 1947</small></li> <li><i><a href="/w/index.php?title=Sallea&action=edit&redlink=1" class="new" title="Sallea (page does not exist)">Sallea</a></i> <small>Chevrolat, 1874</small></li> <li><i><a href="/w/index.php?title=Scrobiger&action=edit&redlink=1" class="new" title="Scrobiger (page does not exist)">Scrobiger</a></i> <small>Spinola, 1841</small></li> <li><i><a href="/w/index.php?title=Sedlacekius&action=edit&redlink=1" class="new" title="Sedlacekius (page does not exist)">Sedlacekius</a></i> <small>Winkler, 1972</small></li> <li><i><a href="/w/index.php?title=Sikorius&action=edit&redlink=1" class="new" title="Sikorius (page does not exist)">Sikorius</a></i> <small>Kuwert, 1893</small></li> <li><i><a href="/wiki/Stigmatium" title="Stigmatium">Stigmatium</a></i> <small>Gray in Griffith, 1832</small></li> <li><i><a href="/w/index.php?title=Systenoderes&action=edit&redlink=1" class="new" title="Systenoderes (page does not exist)">Systenoderes</a></i> <small>Spinola, 1841</small></li> <li><i><a href="/w/index.php?title=Tanocleria&action=edit&redlink=1" class="new" title="Tanocleria (page does not exist)">Tanocleria</a></i> <small>Hong, 2002</small></li> <li><i><a href="/w/index.php?title=Thalerocnemis&action=edit&redlink=1" class="new" title="Thalerocnemis (page does not exist)">Thalerocnemis</a></i> <small>Lohde, 1900</small></li> <li><i><a href="/wiki/Thanasimodes" title="Thanasimodes">Thanasimodes</a></i> <small>Murray, 1867</small></li> <li><i><a href="/wiki/Thanasimus" title="Thanasimus">Thanasimus</a></i> <small>Latreille, 1806</small> <ul><li><i><a href="/wiki/Thanasimus_formicarius" title="Thanasimus formicarius">Thanasimus formicarius</a></i> – <a href="/wiki/Ant_Beetle" class="mw-redirect" title="Ant Beetle">Ant Beetle</a></li></ul></li> <li><i><a href="/w/index.php?title=Tillicera&action=edit&redlink=1" class="new" title="Tillicera (page does not exist)">Tillicera</a></i> <small>Spinola, 1841</small></li> <li><i><a href="/wiki/Trichodes" title="Trichodes">Trichodes</a></i> <small>Herbst, 1792</small> <ul><li><i><a href="/wiki/Trichodes_alvearius" title="Trichodes alvearius">Trichodes alvearius</a></i></li> <li><i><a href="/wiki/Trichodes_apiarius" title="Trichodes apiarius">Trichodes apiarius</a></i></li> <li><i><a href="/wiki/Trichodes_leucopsideus" title="Trichodes leucopsideus">Trichodes leucopsideus</a></i></li></ul></li> <li><i><a href="/w/index.php?title=Trogodendron&action=edit&redlink=1" class="new" title="Trogodendron (page does not exist)">Trogodendron</a></i> <small>Spinola, 1841</small> <ul><li><i><a href="/wiki/Trogodendron_fasciculatum" title="Trogodendron fasciculatum">Trogodendron fasciculatum</a></i> – <a href="/wiki/Yellow-horned_Clerid" class="mw-redirect" title="Yellow-horned Clerid">Yellow-horned Clerid</a></li></ul></li> <li><i><a href="/w/index.php?title=Wilsonoclerus&action=edit&redlink=1" class="new" title="Wilsonoclerus (page does not exist)">Wilsonoclerus</a></i></li> <li><i><a href="/w/index.php?title=Winklerius&action=edit&redlink=1" class="new" title="Winklerius (page does not exist)">Winklerius</a></i> <small>Menier, 1986</small></li> <li><i><a href="/w/index.php?title=Wittmeridecus&action=edit&redlink=1" class="new" title="Wittmeridecus (page does not exist)">Wittmeridecus</a></i> <small>Winkler, 1981</small></li> <li><i><a href="/wiki/Xenorthrius" title="Xenorthrius">Xenorthrius</a></i> <small>Gorham, 1892</small></li> <li><i><a href="/w/index.php?title=Zahradnikius&action=edit&redlink=1" class="new" title="Zahradnikius (page does not exist)">Zahradnikius</a></i> <small>Winkler, 1992</small></li> <li><i><a href="/wiki/Zenithicola" title="Zenithicola">Zenithicola</a></i> <small>Spinola, 1841</small></li></ul> </td> <td class="col-break col-break-3"> <p><b><a href="/wiki/Enopliinae" title="Enopliinae">Enopliinae</a></b> (sometimes in Korynetinae) </p> <ul><li><i><a href="/w/index.php?title=Antygodera&action=edit&redlink=1" class="new" title="Antygodera (page does not exist)">Antygodera</a></i></li> <li><i><a href="/w/index.php?title=Apolopha&action=edit&redlink=1" class="new" title="Apolopha (page does not exist)">Apolopha</a></i> <small>Spinola, 1841</small></li> <li><i><a href="/wiki/Corinthiscus" title="Corinthiscus">Corinthiscus</a></i> <small>Fairmaire & Germain, 1861</small></li> <li><i><a href="/wiki/Cregya" title="Cregya">Cregya</a></i> <small>LeConte, 1861</small></li> <li><i><a href="/w/index.php?title=Curacavi_(beetle)&action=edit&redlink=1" class="new" title="Curacavi (beetle) (page does not exist)">Curacavi</a></i></li> <li><i><a href="/w/index.php?title=Enoplium&action=edit&redlink=1" class="new" title="Enoplium (page does not exist)">Enoplium</a></i> <small>Latreille, 1802</small></li> <li><i><a href="/w/index.php?title=Exochonotus&action=edit&redlink=1" class="new" title="Exochonotus (page does not exist)">Exochonotus</a></i></li> <li><i><a href="/w/index.php?title=Hublella&action=edit&redlink=1" class="new" title="Hublella (page does not exist)">Hublella</a></i></li> <li><i><a href="/w/index.php?title=Lasiodera&action=edit&redlink=1" class="new" title="Lasiodera (page does not exist)">Lasiodera</a></i> <small>Gray in Griffith, 1832</small></li> <li><i><a href="/w/index.php?title=Neopylus&action=edit&redlink=1" class="new" title="Neopylus (page does not exist)">Neopylus</a></i> <small>Solervicens, 1989</small></li> <li><i><a href="/w/index.php?title=Paracregya&action=edit&redlink=1" class="new" title="Paracregya (page does not exist)">Paracregya</a></i></li> <li><i><a href="/wiki/Pelonium" title="Pelonium">Pelonium</a></i></li> <li><i><a href="/wiki/Phymatophaea" title="Phymatophaea">Phymatophaea</a></i> <small>Pascoe, 1876</small></li> <li><i><a href="/w/index.php?title=Platynoptera&action=edit&redlink=1" class="new" title="Platynoptera (page does not exist)">Platynoptera</a></i> <small>Chevrolat, 1834</small></li> <li><i><a href="/w/index.php?title=Pseudichnea&action=edit&redlink=1" class="new" title="Pseudichnea (page does not exist)">Pseudichnea</a></i> <small>Schenkling, 1900</small></li> <li><i><a href="/w/index.php?title=Pylus_(beetle)&action=edit&redlink=1" class="new" title="Pylus (beetle) (page does not exist)">Pylus</a></i> <small>Newman, 1840</small></li> <li><i><a href="/w/index.php?title=Pyticara&action=edit&redlink=1" class="new" title="Pyticara (page does not exist)">Pyticara</a></i> <small>Spinola, 1841</small> (including <i>Pelonides</i>)</li> <li><i><a href="/w/index.php?title=Teneroides&action=edit&redlink=1" class="new" title="Teneroides (page does not exist)">Teneroides</a></i> <small>Gahan, 1910</small></li> <li><i><a href="/wiki/Tenerus_(beetle)" title="Tenerus (beetle)">Tenerus</a></i> <small>Laporte de Castelnau, 1836</small></li> <li><i><a href="/w/index.php?title=Thriocerodes&action=edit&redlink=1" class="new" title="Thriocerodes (page does not exist)">Thriocerodes</a></i> <small>Wolcott & Dybas, 1947</small></li></ul> <p><b><a href="/wiki/Epiphloeinae" title="Epiphloeinae">Epiphloeinae</a></b> (sometimes in Korynetinae) </p> <ul><li><i><a href="/w/index.php?title=Acanthocollum&action=edit&redlink=1" class="new" title="Acanthocollum (page does not exist)">Acanthocollum</a></i></li> <li><i><a href="/w/index.php?title=Amboakis&action=edit&redlink=1" class="new" title="Amboakis (page does not exist)">Amboakis</a></i></li> <li><i><a href="/w/index.php?title=Decaphloeus&action=edit&redlink=1" class="new" title="Decaphloeus (page does not exist)">Decaphloeus</a></i></li> <li><i><a href="/w/index.php?title=Decorosa&action=edit&redlink=1" class="new" title="Decorosa (page does not exist)">Decorosa</a></i> <small>Opitz, 2008</small></li> <li><i><a href="/w/index.php?title=Diapromeces&action=edit&redlink=1" class="new" title="Diapromeces (page does not exist)">Diapromeces</a></i> <small>Opitz, 1997</small></li> <li><i><a href="/w/index.php?title=Ellipotoma&action=edit&redlink=1" class="new" title="Ellipotoma (page does not exist)">Ellipotoma</a></i> <small>Spinola, 1844</small></li> <li><i><a href="/w/index.php?title=Epiphloeus&action=edit&redlink=1" class="new" title="Epiphloeus (page does not exist)">Epiphloeus</a></i> <small>Spinola, 1841</small></li> <li><i><a href="/w/index.php?title=Hapsidopteris&action=edit&redlink=1" class="new" title="Hapsidopteris (page does not exist)">Hapsidopteris</a></i> <small>Opitz, 1997</small></li> <li><i><a href="/wiki/Ichnea" title="Ichnea">Ichnea</a></i> <small>Laporte de Castelnau, 1836</small></li> <li><i><a href="/w/index.php?title=Iontoclerus&action=edit&redlink=1" class="new" title="Iontoclerus (page does not exist)">Iontoclerus</a></i> <small>Opitz, 1997</small></li> <li><i><a href="/w/index.php?title=Katamyurus&action=edit&redlink=1" class="new" title="Katamyurus (page does not exist)">Katamyurus</a></i> <small>Opitz, 1997</small></li> <li><i><a href="/wiki/Madoniella" title="Madoniella">Madoniella</a></i> <small>Pic, 1935</small></li> <li><i><a href="/w/index.php?title=Megaphloeus&action=edit&redlink=1" class="new" title="Megaphloeus (page does not exist)">Megaphloeus</a></i></li> <li><i><a href="/w/index.php?title=Megatrachys&action=edit&redlink=1" class="new" title="Megatrachys (page does not exist)">Megatrachys</a></i> <small>Opitz, 1997</small></li> <li><i><a href="/w/index.php?title=Opitzius&action=edit&redlink=1" class="new" title="Opitzius (page does not exist)">Opitzius</a></i> <small>Barr, 2006</small></li> <li><i><a href="/w/index.php?title=Parvochaetus&action=edit&redlink=1" class="new" title="Parvochaetus (page does not exist)">Parvochaetus</a></i> <small>Opitz, 2006</small></li> <li><i><a href="/wiki/Pennasolis" title="Pennasolis">Pennasolis</a></i> <small>Opitz, 2008</small></li> <li><i><a href="/w/index.php?title=Pericales&action=edit&redlink=1" class="new" title="Pericales (page does not exist)">Pericales</a></i></li> <li><i><a href="/w/index.php?title=Pilosirus&action=edit&redlink=1" class="new" title="Pilosirus (page does not exist)">Pilosirus</a></i> <small>Opitz, 1997</small></li> <li><i><a href="/w/index.php?title=Plocamocera&action=edit&redlink=1" class="new" title="Plocamocera (page does not exist)">Plocamocera</a></i> <small>Spinola, 1844</small></li> <li><i><a href="/w/index.php?title=Pteroferus&action=edit&redlink=1" class="new" title="Pteroferus (page does not exist)">Pteroferus</a></i></li> <li><i><a href="/wiki/Pyticeroides" title="Pyticeroides">Pyticeroides</a></i> <small>Kuwert, 1894</small></li> <li><i><a href="/w/index.php?title=Silveirasia&action=edit&redlink=1" class="new" title="Silveirasia (page does not exist)">Silveirasia</a></i></li> <li><i><a href="/w/index.php?title=Stegnoclava&action=edit&redlink=1" class="new" title="Stegnoclava (page does not exist)">Stegnoclava</a></i></li> <li><i><a href="/w/index.php?title=Turbophloeus&action=edit&redlink=1" class="new" title="Turbophloeus (page does not exist)">Turbophloeus</a></i></li></ul> <p><b><a href="/wiki/Hydnocerinae" title="Hydnocerinae">Hydnocerinae</a></b> (including Phyllobaeninae) </p> <ul><li><i><a href="/w/index.php?title=Abrosius_(beetle)&action=edit&redlink=1" class="new" title="Abrosius (beetle) (page does not exist)">Abrosius</a></i> <small>Fairmaire, 1902</small></li> <li><i><a href="/w/index.php?title=Achlamys&action=edit&redlink=1" class="new" title="Achlamys (page does not exist)">Achlamys</a></i> <small>C.O.Waterhouse, 1879</small></li> <li><i><a href="/w/index.php?title=Allelidea&action=edit&redlink=1" class="new" title="Allelidea (page does not exist)">Allelidea</a></i> <small>G.R.Waterhouse, 1839</small></li> <li><i><a href="/w/index.php?title=Blaesiophthalmus&action=edit&redlink=1" class="new" title="Blaesiophthalmus (page does not exist)">Blaesiophthalmus</a></i> <small>Schenkling, 1903</small></li> <li><i><a href="/w/index.php?title=Brachycallimerus&action=edit&redlink=1" class="new" title="Brachycallimerus (page does not exist)">Brachycallimerus</a></i> <small>Chapin, 1924</small></li> <li><i><a href="/w/index.php?title=Brachyptevenus&action=edit&redlink=1" class="new" title="Brachyptevenus (page does not exist)">Brachyptevenus</a></i></li> <li><i><a href="/wiki/Callimerus" title="Callimerus">Callimerus</a></i> <small>Gorham, 1876</small></li> <li><i><a href="/w/index.php?title=Cephaloclerus&action=edit&redlink=1" class="new" title="Cephaloclerus (page does not exist)">Cephaloclerus</a></i> <small>Kuwert, 1893</small></li> <li><i><a href="/w/index.php?title=Cucujocallimerus&action=edit&redlink=1" class="new" title="Cucujocallimerus (page does not exist)">Cucujocallimerus</a></i> <small>Pic, 1929</small></li> <li><i><a href="/w/index.php?title=Emmepus&action=edit&redlink=1" class="new" title="Emmepus (page does not exist)">Emmepus</a></i> <small>Motschulsky, 1845</small></li> <li><i><a href="/w/index.php?title=Eurymetopum&action=edit&redlink=1" class="new" title="Eurymetopum (page does not exist)">Eurymetopum</a></i> <small>Blanchard, 1842</small></li> <li><i><a href="/wiki/Isohydnocera" title="Isohydnocera">Isohydnocera</a></i> <small>Chapin, 1917</small></li> <li><i><a href="/w/index.php?title=Isolemidia&action=edit&redlink=1" class="new" title="Isolemidia (page does not exist)">Isolemidia</a></i> <small>Gorham, 1877</small></li> <li><i><a href="/w/index.php?title=Laiomorphus&action=edit&redlink=1" class="new" title="Laiomorphus (page does not exist)">Laiomorphus</a></i> <small>Pic, 1927</small></li> <li><i><a href="/w/index.php?title=Lasiocallimerus&action=edit&redlink=1" class="new" title="Lasiocallimerus (page does not exist)">Lasiocallimerus</a></i> <small><a href="/wiki/Johannes_Bastiaan_Corporaal" title="Johannes Bastiaan Corporaal">Corporaal</a>, 1939</small></li> <li><i><a href="/w/index.php?title=Lemidia&action=edit&redlink=1" class="new" title="Lemidia (page does not exist)">Lemidia</a></i> <small>Spinola, 1841</small></li> <li><i><a href="/w/index.php?title=Neohydnus&action=edit&redlink=1" class="new" title="Neohydnus (page does not exist)">Neohydnus</a></i> <small>Gorham, 1892</small></li> <li><i><a href="/w/index.php?title=Parmius&action=edit&redlink=1" class="new" title="Parmius (page does not exist)">Parmius</a></i> <small>Sharp, 1877</small></li> <li><i><a href="/w/index.php?title=Paupris&action=edit&redlink=1" class="new" title="Paupris (page does not exist)">Paupris</a></i> <small>Sharp, 1877</small></li> <li><i><a href="/wiki/Phyllobaenus" title="Phyllobaenus">Phyllobaenus</a></i> <small>Dejean, 1837</small></li> <li><i><a href="/w/index.php?title=Silviella_(beetle)&action=edit&redlink=1" class="new" title="Silviella (beetle) (page does not exist)">Silviella</a></i> <small>Solervicens, 1987</small></li> <li><i><a href="/w/index.php?title=Solemidia&action=edit&redlink=1" class="new" title="Solemidia (page does not exist)">Solemidia</a></i></li> <li><i><a href="/w/index.php?title=Stenocallimerus&action=edit&redlink=1" class="new" title="Stenocallimerus (page does not exist)">Stenocallimerus</a></i> <small><a href="/wiki/Johannes_Bastiaan_Corporaal" title="Johannes Bastiaan Corporaal">Corporaal</a>& Pic, 1940</small></li> <li><i><a href="/w/index.php?title=Theano_(beetle)&action=edit&redlink=1" class="new" title="Theano (beetle) (page does not exist)">Theano</a></i> <small>Laporte de Castelnau, 1836</small></li> <li><i><a href="/wiki/Wolcottia" title="Wolcottia">Wolcottia</a></i> <small>Chapin, 1917</small></li></ul> <p><b><a href="/w/index.php?title=Korynetinae&action=edit&redlink=1" class="new" title="Korynetinae (page does not exist)">Korynetinae</a></b> </p> <ul><li><i><a href="/wiki/Chariessa" title="Chariessa">Chariessa</a></i> <small>Perty in Spix & Martius, 1830</small></li> <li><i><a href="/wiki/Korynetes" title="Korynetes">Korynetes</a></i> <small>Herbst, 1792</small> <ul><li><i><a href="/wiki/Korynetes_caeruleus" title="Korynetes caeruleus">Korynetes caeruleus</a></i> – <a href="/wiki/Steely_blue_beetle" class="mw-redirect" title="Steely blue beetle">steely blue beetle</a></li></ul></li> <li><i><a href="/wiki/Lebasiella" title="Lebasiella">Lebasiella</a></i> <small>Spinola, 1844</small></li> <li><i><a href="/wiki/Loedelia" title="Loedelia">Loedelia</a></i> <small>R.Lucas, 1918</small></li> <li><i><a href="/wiki/Necrobia" title="Necrobia">Necrobia</a></i> <small>Latreille, 1797</small><sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> <ul><li><i><a href="/wiki/Necrobia_ruficollis" title="Necrobia ruficollis">Necrobia ruficollis</a></i> – <a href="/wiki/Red-shouldered_ham_beetle" class="mw-redirect" title="Red-shouldered ham beetle">red-shouldered ham beetle</a></li> <li><i><a href="/wiki/Necrobia_rufipes" title="Necrobia rufipes">Necrobia rufipes</a></i> – <a href="/wiki/Red-legged_ham_beetle" class="mw-redirect" title="Red-legged ham beetle">red-legged ham beetle</a></li></ul></li> <li><i><a href="/wiki/Neorthopleura" title="Neorthopleura">Neorthopleura</a></i> <small>Barr, 1976</small></li> <li><i><a href="/wiki/Opetiopalpus" title="Opetiopalpus">Opetiopalpus</a></i> <small>Spinola, 1844</small></li> <li><i><a href="/w/index.php?title=Romanaeclerus&action=edit&redlink=1" class="new" title="Romanaeclerus (page does not exist)">Romanaeclerus</a></i> <small>Winkler, 1960</small></li></ul> <p><b><a href="/wiki/Tarsosteninae" title="Tarsosteninae">Tarsosteninae</a></b> (sometimes in Korynetinae) </p> <ul><li><i><a href="/wiki/Paratillus" title="Paratillus">Paratillus</a></i> <small>Gorham, 1876</small></li> <li><i><a href="/w/index.php?title=Tarsostenodes&action=edit&redlink=1" class="new" title="Tarsostenodes (page does not exist)">Tarsostenodes</a></i> <small>Blackburn, 1900</small></li> <li><i><a href="/wiki/Tarsostenus" title="Tarsostenus">Tarsostenus</a></i> <small>Spinola, 1844</small></li></ul> <p><b><a href="/wiki/Thaneroclerinae" class="mw-redirect" title="Thaneroclerinae">Thaneroclerinae</a></b> (tentatively placed here) </p> <ul><li><i><a href="/w/index.php?title=Cleridopsis&action=edit&redlink=1" class="new" title="Cleridopsis (page does not exist)">Cleridopsis</a></i> <small>Champion, 1913</small></li> <li><i><a href="/w/index.php?title=Compactoclerus&action=edit&redlink=1" class="new" title="Compactoclerus (page does not exist)">Compactoclerus</a></i> <small>Pic, 1939</small></li> <li><i><a href="/w/index.php?title=Cyrtinoclerus&action=edit&redlink=1" class="new" title="Cyrtinoclerus (page does not exist)">Cyrtinoclerus</a></i> <small>Chapin, 1924</small></li> <li><i><a href="/wiki/Isoclerus" title="Isoclerus">Isoclerus</a></i> <small>Lewis, 1892</small></li> <li><i><a href="/w/index.php?title=Meprinogenus&action=edit&redlink=1" class="new" title="Meprinogenus (page does not exist)">Meprinogenus</a></i> <small>Kolibáč, 1992</small></li> <li><i><a href="/w/index.php?title=Neoclerus&action=edit&redlink=1" class="new" title="Neoclerus (page does not exist)">Neoclerus</a></i> <small>Lewis, 1892</small></li> <li><i><a href="/w/index.php?title=Onerunka&action=edit&redlink=1" class="new" title="Onerunka (page does not exist)">Onerunka</a></i> <small>Kolibáč</small></li> <li><i><a href="/wiki/Thaneroclerus" title="Thaneroclerus">Thaneroclerus</a></i> <small>Lefebvre, 1838</small></li> <li><i><a href="/w/index.php?title=Viticlerus&action=edit&redlink=1" class="new" title="Viticlerus (page does not exist)">Viticlerus</a></i></li> <li><i><a href="/wiki/Zenodosus" title="Zenodosus">Zenodosus</a></i> <small>Wolcott, 1910</small></li></ul> </td> <td class="col-break col-break-3"> <p><b><a href="/wiki/Tillinae" title="Tillinae">Tillinae</a></b> </p> <ul><li><i><a href="/w/index.php?title=Antenius&action=edit&redlink=1" class="new" title="Antenius (page does not exist)">Antenius</a></i> <small>Fairmaire, 1903</small></li> <li><i><a href="/w/index.php?title=Arachnoclerus&action=edit&redlink=1" class="new" title="Arachnoclerus (page does not exist)">Arachnoclerus</a></i> <small>Fairmaire, 1902</small></li> <li><i><a href="/wiki/Araeodontia" title="Araeodontia">Araeodontia</a></i> <small>Barr, 1952</small></li> <li><i><a href="/w/index.php?title=Archalius&action=edit&redlink=1" class="new" title="Archalius (page does not exist)">Archalius</a></i> <small>Fairmaire, 1903</small></li> <li><i><a href="/w/index.php?title=Aroterus&action=edit&redlink=1" class="new" title="Aroterus (page does not exist)">Aroterus</a></i> <small>Schenkling, 1906</small></li> <li><i><a href="/w/index.php?title=Basilewskyus&action=edit&redlink=1" class="new" title="Basilewskyus (page does not exist)">Basilewskyus</a></i> <small>Pic, 1950</small></li> <li><i><a href="/w/index.php?title=Biflabellotillus&action=edit&redlink=1" class="new" title="Biflabellotillus (page does not exist)">Biflabellotillus</a></i> <small>Pic, 1949</small></li> <li><i><a href="/w/index.php?title=Bilbotillus&action=edit&redlink=1" class="new" title="Bilbotillus (page does not exist)">Bilbotillus</a></i> <small>Kolibac, 1997</small></li> <li><i><a href="/w/index.php?title=Bogcia&action=edit&redlink=1" class="new" title="Bogcia (page does not exist)">Bogcia</a></i> <small>Barr, 1978</small></li> <li><i><a href="/w/index.php?title=Bostrichoclerus&action=edit&redlink=1" class="new" title="Bostrichoclerus (page does not exist)">Bostrichoclerus</a></i> <small>Van Dyke, 1938</small></li> <li><i><a href="/wiki/Callotillus" title="Callotillus">Callotillus</a></i> <small>Wolcott, 1911</small></li> <li><i><a href="/w/index.php?title=Ceratocopus&action=edit&redlink=1" class="new" title="Ceratocopus (page does not exist)">Ceratocopus</a></i> <small>Hintz, 1902</small></li> <li><i><a href="/w/index.php?title=Chilioclerus&action=edit&redlink=1" class="new" title="Chilioclerus (page does not exist)">Chilioclerus</a></i> <small>Solervicens, 1976</small></li> <li><i><a href="/w/index.php?title=Cladiscopallenis&action=edit&redlink=1" class="new" title="Cladiscopallenis (page does not exist)">Cladiscopallenis</a></i> <small>Pic, 1949</small></li> <li><i><a href="/w/index.php?title=Cladiscus&action=edit&redlink=1" class="new" title="Cladiscus (page does not exist)">Cladiscus</a></i> <small>Chevrolat, 1843</small></li> <li><i><a href="/w/index.php?title=Cladomorpha&action=edit&redlink=1" class="new" title="Cladomorpha (page does not exist)">Cladomorpha</a></i> <small>Pic, 1949</small></li> <li><i><a href="/w/index.php?title=Cteniopachys&action=edit&redlink=1" class="new" title="Cteniopachys (page does not exist)">Cteniopachys</a></i> <small>Fairmaire, 1889</small></li> <li><i><a href="/w/index.php?title=Cylidroctenus&action=edit&redlink=1" class="new" title="Cylidroctenus (page does not exist)">Cylidroctenus</a></i> <small>Kraatz, 1899</small></li> <li><i><a href="/w/index.php?title=Cylidrus&action=edit&redlink=1" class="new" title="Cylidrus (page does not exist)">Cylidrus</a></i> <small>Latreille, 1825</small></li> <li><i><a href="/wiki/Cymatodera" title="Cymatodera">Cymatodera</a></i> <small>Gray in Griffith, 1832</small></li> <li><i><a href="/wiki/Cymatoderella" title="Cymatoderella">Cymatoderella</a></i> <small>Barr, 1962</small></li> <li><i><a href="/w/index.php?title=Dedana&action=edit&redlink=1" class="new" title="Dedana (page does not exist)">Dedana</a></i> <small>Fairmaire, 1888</small></li> <li><i><a href="/w/index.php?title=Denops&action=edit&redlink=1" class="new" title="Denops (page does not exist)">Denops</a></i> <small>Fischer von Waldheim, 1829</small></li> <li><i><a href="/wiki/Diplocladus" title="Diplocladus">Diplocladus</a></i> <small>Fairmaire, 1885</small></li> <li><i><a href="/w/index.php?title=Diplopherusa&action=edit&redlink=1" class="new" title="Diplopherusa (page does not exist)">Diplopherusa</a></i> <small>Heller, 1921</small></li> <li><i><a href="/w/index.php?title=Eburneocladiscus&action=edit&redlink=1" class="new" title="Eburneocladiscus (page does not exist)">Eburneocladiscus</a></i> <small>Pic, 1955</small></li> <li><i><a href="/w/index.php?title=Egenocladiscus&action=edit&redlink=1" class="new" title="Egenocladiscus (page does not exist)">Egenocladiscus</a></i> <small><a href="/wiki/Johannes_Bastiaan_Corporaal" title="Johannes Bastiaan Corporaal">Corporaal</a>& van der Wiel, 1949</small></li> <li><i><a href="/w/index.php?title=Elasmocylidrus&action=edit&redlink=1" class="new" title="Elasmocylidrus (page does not exist)">Elasmocylidrus</a></i> <small><a href="/wiki/Johannes_Bastiaan_Corporaal" title="Johannes Bastiaan Corporaal">Corporaal</a>, 1939</small></li> <li><i><a href="/w/index.php?title=Enoploclerus&action=edit&redlink=1" class="new" title="Enoploclerus (page does not exist)">Enoploclerus</a></i> <small>Hintz, 1902</small></li> <li><i><a href="/w/index.php?title=Eucymatodera&action=edit&redlink=1" class="new" title="Eucymatodera (page does not exist)">Eucymatodera</a></i> <small>Schenkling, 1899</small></li> <li><i><a href="/w/index.php?title=Falsopallenis&action=edit&redlink=1" class="new" title="Falsopallenis (page does not exist)">Falsopallenis</a></i> <small>Pic, 1926</small></li> <li><i><a href="/w/index.php?title=Falsotillus&action=edit&redlink=1" class="new" title="Falsotillus (page does not exist)">Falsotillus</a></i> <small>Gerstmeier & Kuff, 1992</small></li> <li><i><a href="/w/index.php?title=Flabellotilloidea&action=edit&redlink=1" class="new" title="Flabellotilloidea (page does not exist)">Flabellotilloidea</a></i> <small>Gerstmeier & Kuff, 1992</small></li> <li><i><a href="/w/index.php?title=Gastrocentrum&action=edit&redlink=1" class="new" title="Gastrocentrum (page does not exist)">Gastrocentrum</a></i> <small>Gorham, 1876</small></li> <li><i><a href="/w/index.php?title=Gracilotillus&action=edit&redlink=1" class="new" title="Gracilotillus (page does not exist)">Gracilotillus</a></i> <small>Pic, 1933</small></li> <li><i><a href="/w/index.php?title=Impressopallenis&action=edit&redlink=1" class="new" title="Impressopallenis (page does not exist)">Impressopallenis</a></i> <small>Pic, 1953</small></li> <li><i><a href="/w/index.php?title=Isocymatodera&action=edit&redlink=1" class="new" title="Isocymatodera (page does not exist)">Isocymatodera</a></i> <small>Hintz, 1902</small></li> <li><i><a href="/wiki/Lecontella" title="Lecontella">Lecontella</a></i> <small>Wolcott & Chapin, 1918</small></li> <li><i><a href="/w/index.php?title=Leptoclerus&action=edit&redlink=1" class="new" title="Leptoclerus (page does not exist)">Leptoclerus</a></i> <small>Kraatz, 1899</small></li> <li><i><a href="/w/index.php?title=Liostylus&action=edit&redlink=1" class="new" title="Liostylus (page does not exist)">Liostylus</a></i> <small>Fairmaire, 1886</small></li> <li><i><a href="/w/index.php?title=Macroliostylus&action=edit&redlink=1" class="new" title="Macroliostylus (page does not exist)">Macroliostylus</a></i> <small>Pic, 1939</small></li> <li><i><a href="/w/index.php?title=Magnotillus&action=edit&redlink=1" class="new" title="Magnotillus (page does not exist)">Magnotillus</a></i> <small>Pic, 1936</small></li> <li><i><a href="/w/index.php?title=Melanoclerus&action=edit&redlink=1" class="new" title="Melanoclerus (page does not exist)">Melanoclerus</a></i> <small>Chapin, 1919</small></li> <li><i><a href="/w/index.php?title=Microtillus&action=edit&redlink=1" class="new" title="Microtillus (page does not exist)">Microtillus</a></i> <small>Pic, 1950</small></li> <li><i><a href="/wiki/Monophylla" title="Monophylla">Monophylla</a></i> <small>Spinola, 1841</small></li> <li><i><a href="/w/index.php?title=Neocallotillus&action=edit&redlink=1" class="new" title="Neocallotillus (page does not exist)">Neocallotillus</a></i> <small>Burke, 2016</small></li> <li><i><a href="/w/index.php?title=Nodepus&action=edit&redlink=1" class="new" title="Nodepus (page does not exist)">Nodepus</a></i> <small>Gorham, 1892</small></li> <li><i><a href="/w/index.php?title=Notocymatodera&action=edit&redlink=1" class="new" title="Notocymatodera (page does not exist)">Notocymatodera</a></i> <small>Schenkling, 1907</small></li> <li><i><a href="/w/index.php?title=Onychotillus&action=edit&redlink=1" class="new" title="Onychotillus (page does not exist)">Onychotillus</a></i> <small>Chapin, 1945</small></li> <li><i><a href="/w/index.php?title=Orthocladiscus&action=edit&redlink=1" class="new" title="Orthocladiscus (page does not exist)">Orthocladiscus</a></i> <small><a href="/wiki/Johannes_Bastiaan_Corporaal" title="Johannes Bastiaan Corporaal">Corporaal</a>& van der Wiel, 1949</small></li> <li><i><a href="/w/index.php?title=Pallenis_(beetle)&action=edit&redlink=1" class="new" title="Pallenis (beetle) (page does not exist)">Pallenis</a></i> <small>Laporte de Castelnau, 1836</small></li> <li><i><a href="/w/index.php?title=Paracladiscus&action=edit&redlink=1" class="new" title="Paracladiscus (page does not exist)">Paracladiscus</a></i> <small>Miyatake, 1965</small></li> <li><i><a href="/w/index.php?title=Paradoxocerus&action=edit&redlink=1" class="new" title="Paradoxocerus (page does not exist)">Paradoxocerus</a></i> <small>Kraatz, 1899</small></li> <li><i><a href="/w/index.php?title=Paraspinoza&action=edit&redlink=1" class="new" title="Paraspinoza (page does not exist)">Paraspinoza</a></i> <small><a href="/wiki/Johannes_Bastiaan_Corporaal" title="Johannes Bastiaan Corporaal">Corporaal</a>, 1942</small></li> <li><i><a href="/w/index.php?title=Philocalus_(beetle)&action=edit&redlink=1" class="new" title="Philocalus (beetle) (page does not exist)">Philocalus</a></i> <small>Klug, 1842</small></li> <li><i><a href="/w/index.php?title=Picoclerus&action=edit&redlink=1" class="new" title="Picoclerus (page does not exist)">Picoclerus</a></i> <small><a href="/wiki/Johannes_Bastiaan_Corporaal" title="Johannes Bastiaan Corporaal">Corporaal</a>, 1936</small></li> <li>†<i><a href="/w/index.php?title=Prospinoza&action=edit&redlink=1" class="new" title="Prospinoza (page does not exist)">Prospinoza</a></i> (fossil)</li> <li><i><a href="/w/index.php?title=Pseudachlamys&action=edit&redlink=1" class="new" title="Pseudachlamys (page does not exist)">Pseudachlamys</a></i> <small>Duvivier, 1892</small></li> <li><i><a href="/w/index.php?title=Pseudogyponix&action=edit&redlink=1" class="new" title="Pseudogyponix (page does not exist)">Pseudogyponix</a></i> <small>Pic, 1939</small></li> <li><i><a href="/w/index.php?title=Pseudopallenis&action=edit&redlink=1" class="new" title="Pseudopallenis (page does not exist)">Pseudopallenis</a></i> <small>Kuwert, 1893</small></li> <li><i><a href="/w/index.php?title=Pseudoteloclerus&action=edit&redlink=1" class="new" title="Pseudoteloclerus (page does not exist)">Pseudoteloclerus</a></i> <small>Pic, 1932</small></li> <li><i><a href="/w/index.php?title=Rhopaloclerus&action=edit&redlink=1" class="new" title="Rhopaloclerus (page does not exist)">Rhopaloclerus</a></i> <small>Fairmaire, 1886</small></li> <li><i><a href="/w/index.php?title=Smudlotillus&action=edit&redlink=1" class="new" title="Smudlotillus (page does not exist)">Smudlotillus</a></i> <small>Kolibac, 1997</small></li> <li><i><a href="/w/index.php?title=Spinoza_(beetle)&action=edit&redlink=1" class="new" title="Spinoza (beetle) (page does not exist)">Spinoza</a></i> <small>Lewis, 1892</small></li> <li><i><a href="/w/index.php?title=Stenocylidrus&action=edit&redlink=1" class="new" title="Stenocylidrus (page does not exist)">Stenocylidrus</a></i> <small>Spinola, 1844</small></li> <li><i><a href="/w/index.php?title=Strotocera&action=edit&redlink=1" class="new" title="Strotocera (page does not exist)">Strotocera</a></i> <small>Schenkling, 1902</small></li> <li><i><a href="/w/index.php?title=Synellapotillus&action=edit&redlink=1" class="new" title="Synellapotillus (page does not exist)">Synellapotillus</a></i> <small>Pic, 1939</small></li> <li><i><a href="/w/index.php?title=Synellapus&action=edit&redlink=1" class="new" title="Synellapus (page does not exist)">Synellapus</a></i> <small>Fairmaire, 1903</small></li> <li><i><a href="/w/index.php?title=Teloclerus&action=edit&redlink=1" class="new" title="Teloclerus (page does not exist)">Teloclerus</a></i> <small>Schenkling, 1903</small></li> <li><i><a href="/w/index.php?title=Tilloclerus&action=edit&redlink=1" class="new" title="Tilloclerus (page does not exist)">Tilloclerus</a></i> <small>White, 1849</small></li> <li><i><a href="/w/index.php?title=Tillodadiscus&action=edit&redlink=1" class="new" title="Tillodadiscus (page does not exist)">Tillodadiscus</a></i> <small>Pic, 1953</small></li> <li><i><a href="/w/index.php?title=Tillodenops&action=edit&redlink=1" class="new" title="Tillodenops (page does not exist)">Tillodenops</a></i> <small>Hintz, 1905</small></li> <li><i><a href="/wiki/Tilloidea" title="Tilloidea">Tilloidea</a></i> <small>Laporte de Castelnau, 1832</small></li> <li><i><a href="/wiki/Tillus" title="Tillus">Tillus</a></i> <small>Olivier, 1790</small></li> <li><i><a href="/w/index.php?title=Tylotosoma&action=edit&redlink=1" class="new" title="Tylotosoma (page does not exist)">Tylotosoma</a></i> <small>Hintz, 1902</small></li></ul> <p><i><b><a href="/wiki/Incertae_sedis" title="Incertae sedis">Incertae sedis</a></b></i> </p> <ul><li><i><a href="/w/index.php?title=Aphelocerus&action=edit&redlink=1" class="new" title="Aphelocerus (page does not exist)">Aphelocerus</a></i> <small>Kirsch, 1871</small> (Clerinae? Tillinae?)</li> <li><i><a href="/wiki/Apteropilo" title="Apteropilo">Apteropilo</a></i> <small>Lea, 1908</small> (Clerinae? Enopliinae?)</li> <li><i><a href="/w/index.php?title=Cleropiestus&action=edit&redlink=1" class="new" title="Cleropiestus (page does not exist)">Cleropiestus</a></i> <small>Fairmaire, 1889</small> (Clerinae? Hydnocerinae?)</li> <li><i><a href="/w/index.php?title=Dermestoides&action=edit&redlink=1" class="new" title="Dermestoides (page does not exist)">Dermestoides</a></i> <small>Schaeffer, 1771</small> (Korynetinae <i><a href="/wiki/Sensu_lato" class="mw-redirect" title="Sensu lato">s.l.</a></i>?)</li> <li><i><a href="/w/index.php?title=Evenoclerus&action=edit&redlink=1" class="new" title="Evenoclerus (page does not exist)">Evenoclerus</a></i> <small><a href="/wiki/Johannes_Bastiaan_Corporaal" title="Johannes Bastiaan Corporaal">Corporaal</a>, 1950</small> (Clerinae? Hydnocerinae?)</li> <li><i><a href="/wiki/Muisca_(beetle)" title="Muisca (beetle)">Muisca</a></i> <small>Spinola, 1844</small> (Clerinae? Enopliinae?)</li> <li><i><a href="/w/index.php?title=Parapelonides&action=edit&redlink=1" class="new" title="Parapelonides (page does not exist)">Parapelonides</a></i> <small>Barr, 1980</small> (Korynetinae <i>s.l.</i>?)</li> <li><i><a href="/w/index.php?title=Perilypus&action=edit&redlink=1" class="new" title="Perilypus (page does not exist)">Perilypus</a></i> <small>Spinola, 1841</small> (Clerinae? Tillinae?)</li> <li><i><a href="/w/index.php?title=Syriopelta&action=edit&redlink=1" class="new" title="Syriopelta (page does not exist)">Syriopelta</a></i> <small>Winkler, 1984</small> (Korynetinae <i>s.l.</i>?)</li></ul> <p>  </p> </td></tr></tbody></table></div> </div> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cleridae&action=edit&section=12" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width" style="column-width: 30em;"> <ol class="references"> <li id="cite_note-Johnson-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-Johnson_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Johnson_1-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Johnson_1-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-Johnson_1-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-Johnson_1-4"><sup><i><b>e</b></i></sup></a> <a href="#cite_ref-Johnson_1-5"><sup><i><b>f</b></i></sup></a> <a href="#cite_ref-Johnson_1-6"><sup><i><b>g</b></i></sup></a> <a href="#cite_ref-Johnson_1-7"><sup><i><b>h</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFJohnsonTriplehorn2004" class="citation book cs1">Johnson, Norman F.; Triplehorn, Charles A. (2004). <i>Borror and DeLong's Introduction to the Study of Insects</i> (7th ed.). Belmont: Brooks/Cole. pp. 365–400, 428–429. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/0-03-096835-6" title="Special:BookSources/0-03-096835-6"><bdi>0-03-096835-6</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Borror+and+DeLong%27s+Introduction+to+the+Study+of+Insects&rft.place=Belmont&rft.pages=365-400%2C+428-429&rft.edition=7th&rft.pub=Brooks%2FCole&rft.date=2004&rft.isbn=0-03-096835-6&rft.aulast=Johnson&rft.aufirst=Norman+F.&rft.au=Triplehorn%2C+Charles+A.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ACleridae" class="Z3988"></span></span> </li> <li id="cite_note-White-2"><span class="mw-cite-backlink">^ <a href="#cite_ref-White_2-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-White_2-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-White_2-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWhite1998" class="citation book cs1">White, Richard E. (1998). <a rel="nofollow" class="external text" href="https://books.google.com/books?id=S9IK9OMjuNEC"><i>A Field Guide to the Beetles of North America</i></a> (illustrated and revised ed.). New York: Houghton Mifflin Harcourt. pp. 208–209. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-395-91089-4" title="Special:BookSources/978-0-395-91089-4"><bdi>978-0-395-91089-4</bdi></a><span class="reference-accessdate">. Retrieved <span class="nowrap">22 March</span> 2009</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=A+Field+Guide+to+the+Beetles+of+North+America&rft.place=New+York&rft.pages=208-209&rft.edition=illustrated+and+revised&rft.pub=Houghton+Mifflin+Harcourt&rft.date=1998&rft.isbn=978-0-395-91089-4&rft.aulast=White&rft.aufirst=Richard+E.&rft_id=https%3A%2F%2Fbooks.google.com%2Fbooks%3Fid%3DS9IK9OMjuNEC&rfr_id=info%3Asid%2Fen.wikipedia.org%3ACleridae" class="Z3988"></span></span> </li> <li id="cite_note-Byrd-3"><span class="mw-cite-backlink">^ <a href="#cite_ref-Byrd_3-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Byrd_3-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFByrd2001" class="citation book cs1">Byrd, Jason H. (2001). James L., Castner (ed.). <span class="id-lock-registration" title="Free registration required"><a rel="nofollow" class="external text" href="https://archive.org/details/forensicentomolo0000unse"><i>Forensic Entomology: The Utility of Arthropods in Legal Investigations</i></a></span> (2nd ed.). Boca Raton: CRC Press. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/0-8493-8120-7" title="Special:BookSources/0-8493-8120-7"><bdi>0-8493-8120-7</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Forensic+Entomology%3A+The+Utility+of+Arthropods+in+Legal+Investigations&rft.place=Boca+Raton&rft.edition=2nd&rft.pub=CRC+Press&rft.date=2001&rft.isbn=0-8493-8120-7&rft.aulast=Byrd&rft.aufirst=Jason+H.&rft_id=https%3A%2F%2Farchive.org%2Fdetails%2Fforensicentomolo0000unse&rfr_id=info%3Asid%2Fen.wikipedia.org%3ACleridae" class="Z3988"></span></span> </li> <li id="cite_note-Gerstmeier-4"><span class="mw-cite-backlink">^ <a href="#cite_ref-Gerstmeier_4-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Gerstmeier_4-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Gerstmeier_4-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-Gerstmeier_4-3"><sup><i><b>d</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFGerstmeier1998" class="citation book cs1">Gerstmeier, Ronald (1998). <i>Checkered Beetles: Illustrated Key to the Cleridae and Thanerocleridae of the Western Palaearctic</i>. Weikersheim, Germany: Margraf Verlag. pp. 12–15.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Checkered+Beetles%3A+Illustrated+Key+to+the+Cleridae+and+Thanerocleridae+of+the+Western+Palaearctic&rft.place=Weikersheim%2C+Germany&rft.pages=12-15&rft.pub=Margraf+Verlag&rft.date=1998&rft.aulast=Gerstmeier&rft.aufirst=Ronald&rfr_id=info%3Asid%2Fen.wikipedia.org%3ACleridae" class="Z3988"></span></span> </li> <li id="cite_note-Bellows-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-Bellows_5-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBellowsFisherCaltagirone1999" class="citation book cs1">Bellows, Thomas S.; Fisher, T. W.; Caltagirone, L. E. (1999). <a rel="nofollow" class="external text" href="https://books.google.com/books?id=u2X-rfgU0ewC&q=cleridae+feeding&pg=PA429"><i>Handbook of biological control</i></a>. San Diego: Academic Press. pp. 429–430. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/0-12-257305-6" title="Special:BookSources/0-12-257305-6"><bdi>0-12-257305-6</bdi></a><span class="reference-accessdate">. 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(1991) <a rel="nofollow" class="external text" href="http://www.canacoll.org/Coleo/Checklist/PDF%20files/CLERIDAE.pdf">"Family Cleridae: checkered beetles"</a> (PDF). <i>In</i>: Bousquet, Y. (Ed.). <i>Checklist of the Beetles of Canada and Alaska</i>. Agriculture Canada Publication 1861/E. 208–211.</span> </li> <li id="cite_note-Costello-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-Costello_8-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCostello2003" class="citation web cs1">Costello, Sheryl (2003). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20110724193406/http://www.colostate.edu/Depts/Entomology/courses/en507/papers_2003/costello.pdf">"Clerid Beetles- Voracious Predators"</a> <span class="cs1-format">(PDF)</span>. Colorado State University Department of Entomology. pp. 1–15. Archived from <a rel="nofollow" class="external text" href="http://www.colostate.edu/Depts/Entomology/courses/en507/papers_2003/costello.pdf">the original</a> <span class="cs1-format">(PDF)</span> on 24 July 2011<span class="reference-accessdate">. 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href="https://pubmed.ncbi.nlm.nih.gov/29674858">29674858</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=ZooKeys&rft.atitle=The+dating+of+the+fourth+volume+of+Guillaume-Antoine+Olivier%27s+%22Entomologie%2C+ou+histoire+naturelle+des+insectes%22&rft.issue=734&rft.pages=137-148&rft.date=2018&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5904342%23id-name%3DPMC&rft_id=info%3Apmid%2F29674858&rft_id=info%3Adoi%2F10.3897%2Fzookeys.734.22901&rft.aulast=Bousquet&rft.aufirst=Y.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5904342&rfr_id=info%3Asid%2Fen.wikipedia.org%3ACleridae" class="Z3988"></span></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Cleridae&action=edit&section=13" title="Edit section: External 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print{body.ns-0 .mw-parser-output .sistersitebox{display:none!important}}@media screen{html.skin-theme-clientpref-night .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .sistersitebox img[src*="Wiktionary-logo-en-v2.svg"]{background-color:white}}</style><div class="side-box side-box-right plainlinks sistersitebox"><style data-mw-deduplicate="TemplateStyles:r1126788409">.mw-parser-output .plainlist ol,.mw-parser-output .plainlist ul{line-height:inherit;list-style:none;margin:0;padding:0}.mw-parser-output .plainlist ol li,.mw-parser-output .plainlist ul li{margin-bottom:0}</style> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/34px-Wikispecies-logo.svg.png" decoding="async" width="34" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/51px-Wikispecies-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/df/Wikispecies-logo.svg/68px-Wikispecies-logo.svg.png 2x" data-file-width="941" data-file-height="1103" /></span></span></div> <div class="side-box-text plainlist"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a> has information related to <i><b><a href="https://species.wikimedia.org/wiki/Cleridae" class="extiw" title="wikispecies:Cleridae">Cleridae</a></b></i>.</div></div> </div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1235681985"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1237033735"><div class="side-box side-box-right plainlinks sistersitebox"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"> <div class="side-box-flex"> <div class="side-box-image"><span class="noviewer" typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/30px-Commons-logo.svg.png" decoding="async" width="30" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/45px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/4/4a/Commons-logo.svg/59px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></span></span></div> <div class="side-box-text plainlist">Wikimedia Commons has media related to <span style="font-weight: bold; font-style: italic;"><a href="https://commons.wikimedia.org/wiki/Category:Cleridae" class="extiw" title="commons:Category:Cleridae">Cleridae</a></span>.</div></div> </div> <ul><li><a rel="nofollow" class="external text" href="http://www.zin.ru/ANIMALIA/COLEOPTERA/eng/atl_cl.htm">Atlas of checkered beetles (Cleridae) of Russia</a></li> <li><a rel="nofollow" class="external text" href="http://www.coleoptera.org/p1684.htm">Roland Gerstmeier publications on Cleridae</a></li> <li><a rel="nofollow" class="external text" href="http://www.chebucto.ns.ca/environment/NHR/Cleridae.html">Cleridae of Atlantic Canada</a></li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol ol,.mw-parser-output .hlist ol ul,.mw-parser-output .hlist ul dl,.mw-parser-output .hlist ul ol,.mw-parser-output .hlist ul ul{display:inline}.mw-parser-output .hlist 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4em">Extant <a href="/wiki/Beetle" title="Beetle">Coleoptera</a> families</div></th></tr><tr><td class="navbox-abovebelow" colspan="2"><div> <ul><li>Kingdom: <a href="/wiki/Animal" title="Animal">Animalia</a></li> <li>Phylum: <a href="/wiki/Arthropod" title="Arthropod">Arthropoda</a></li> <li>Class: <a href="/wiki/Insect" title="Insect">Insecta</a></li> <li>Subclass: <a href="/wiki/Pterygota" title="Pterygota">Pterygota</a></li> <li>Infraclass: <a href="/wiki/Neoptera" title="Neoptera">Neoptera</a></li> <li>Superorder: <a href="/wiki/Holometabola" title="Holometabola">Holometabola</a></li></ul> </div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Archostemata" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Archostemata" title="Archostemata">Archostemata</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Crowsoniella" title="Crowsoniella">Crowsoniellidae <small>(<i>Crowsoniella relicta</i>)</small></a></li> <li><a href="/wiki/Cupedidae" title="Cupedidae">Cupedidae <small>(reticulated beetles)</small></a></li> <li><a href="/wiki/Jurodidae" title="Jurodidae">Jurodidae <small>(<i>Sikhotealinia zhiltzovae</i>)</small></a></li> <li><a href="/wiki/Micromalthidae" title="Micromalthidae">Micromalthidae</a></li> <li><a href="/wiki/Ommatidae" title="Ommatidae">Ommatidae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Adephaga" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Adephaga" title="Adephaga">Adephaga</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Extant_families" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Extant_taxon" class="mw-redirect" title="Extant taxon">Extant</a> families</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Amphizoa" title="Amphizoa">Amphizoidae <small>(trout-stream beetles)</small></a></li> <li><a href="/wiki/Aspidytidae" title="Aspidytidae">Aspidytidae</a></li> <li><a href="/wiki/Ground_beetle" title="Ground beetle">Carabidae <small>(ground beetles)</small></a></li> <li><a href="/wiki/Tiger_beetle" title="Tiger beetle">Cicindelidae <small>(tiger beetles)</small></a></li> <li><a href="/wiki/Dytiscidae" title="Dytiscidae">Dytiscidae <small>(predaceous diving beetles)</small></a></li> <li><a href="/wiki/Whirligig_beetle" title="Whirligig beetle">Gyrinidae <small>(whirligig beetles)</small></a></li> <li><a href="/wiki/Haliplidae" title="Haliplidae">Haliplidae <small>(crawling water beetles)</small></a></li> <li><a href="/wiki/Hygrobia" title="Hygrobia">Hygrobiidae</a></li> <li><a href="/wiki/Meru_phyllisae" title="Meru phyllisae">Meruidae <small>(<i>Meru phyllisae</i>)</small></a></li> <li><a href="/wiki/Noteridae" title="Noteridae">Noteridae <small>(burrowing water beetles)</small></a></li> <li><a href="/wiki/Trachypachidae" title="Trachypachidae">Trachypachidae <small>(false ground beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Myxophaga" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Myxophaga" title="Myxophaga">Myxophaga</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Hydroscaphidae" title="Hydroscaphidae">Hydroscaphidae <small>(skiff beetles)</small></a></li> <li><a href="/wiki/Lepicerus" title="Lepicerus">Lepiceridae</a></li> <li><a href="/wiki/Sphaerius" title="Sphaerius">Sphaeriusidae</a></li> <li><a href="/wiki/Torridincolidae" title="Torridincolidae">Torridincolidae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible uncollapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Suborder_Polyphaga" style="font-size:114%;margin:0 4em">Suborder <a href="/wiki/Polyphaga" title="Polyphaga">Polyphaga</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Bostrichiformia" title="Bostrichiformia">Bostrichiformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Bostrichoidea" title="Bostrichoidea">Bostrichoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bostrichidae" title="Bostrichidae">Bostrichidae <small>(auger beetles)</small></a></li> <li><a href="/wiki/Dermestidae" title="Dermestidae">Dermestidae <small>(skin beetles)</small></a></li> <li><a href="/wiki/Endecatomus" title="Endecatomus">Endecatomidae</a></li> <li><a href="/wiki/Jacobsoniidae" title="Jacobsoniidae">Jacobsoniidae <small>(Jacobson's beetles)</small></a></li> <li><a href="/wiki/Nosodendridae" title="Nosodendridae">Nosodendridae <small>(wounded-tree beetles)</small></a></li> <li><a href="/wiki/Ptinidae" title="Ptinidae">Ptiniidae <small>(furniture beetles, death watch beetles, spider beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Derodontidae" title="Derodontidae">Derodontoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Derodontidae" title="Derodontidae">Derodontidae <small>(tooth-necked fungus beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cucujiformia" title="Cucujiformia">Cucujiformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Chrysomeloidea" title="Chrysomeloidea">Chrysomeloidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Longhorn_beetle" title="Longhorn beetle">Cerambycidae <small>(longhorn beetles)</small></a></li> <li><a href="/wiki/Leaf_beetle" title="Leaf beetle">Chrysomelidae <small>(leaf beetles)</small></a></li> <li><a href="/wiki/Disteniidae" title="Disteniidae">Disteniidae</a></li> <li><a href="/wiki/Megalopodidae" title="Megalopodidae">Megalopodidae</a></li> <li><a href="/wiki/Orsodacnidae" title="Orsodacnidae">Orsodacnidae</a></li> <li><a href="/wiki/Oxypeltidae" title="Oxypeltidae">Oxypeltidae</a></li> <li><a href="/wiki/Vesperidae" title="Vesperidae">Vesperidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Cleroidea" title="Cleroidea">Cleroidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Acanthocnemus" title="Acanthocnemus">Acanthocnemidae <small>(<i>Acanthocnemus nigricans</i>)</small></a></li> <li><a href="/wiki/Biphyllidae" title="Biphyllidae">Biphyllidae <small>(false skin beetles)</small></a></li> <li><a href="/wiki/Byturidae" title="Byturidae">Byturidae <small>(fruitworm beetles)</small></a></li> <li><a href="/wiki/Chaetosomatidae" title="Chaetosomatidae">Chaetosomatidae</a></li> <li><a class="mw-selflink selflink">Cleridae <small>(checkered beetles)</small></a></li> <li><a href="/wiki/Lophocateridae" title="Lophocateridae">Lophocateridae</a></li> <li><a href="/wiki/Mauroniscidae" title="Mauroniscidae">Mauroniscidae</a></li> <li><a href="/wiki/Melyridae" title="Melyridae">Melyridae <small>(soft-wing flower beetles)</small></a></li> <li><a href="/wiki/Metaxina" title="Metaxina">Metaxinidae <small>(<i>Metaxina ornata</i>)</small></a></li> <li><a href="/wiki/Peltis" title="Peltis">Peltidae</a></li> <li><a href="/wiki/Phloiophilus" title="Phloiophilus">Phloiophilidae <small>(<i>Phloiophilus edwardsi</i>)</small></a></li> <li><a href="/wiki/Phycosecidae" title="Phycosecidae">Phycosecidae</a></li> <li><a href="/wiki/Prionoceridae" title="Prionoceridae">Prionoceridae</a></li> <li><a href="/wiki/Protopeltis" title="Protopeltis">Protopeltidae</a></li> <li><a href="/wiki/Rentoniidae" title="Rentoniidae">Rentoniidae</a></li> <li><a href="/wiki/Rhadalidae" title="Rhadalidae">Rhadalidae</a></li> <li><a href="/wiki/Thanerocleridae" title="Thanerocleridae">Thanerocleridae</a></li> <li><a href="/wiki/Thymalidae" title="Thymalidae">Thymalidae</a></li> <li><a href="/wiki/Trogossitidae" title="Trogossitidae">Trogossitidae <small>(bark-gnawing beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Coccinelloidea" title="Coccinelloidea">Coccinelloidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Akalyptoischion" title="Akalyptoischion">Akalyptoischiidae</a></li> <li><a href="/wiki/Alexiidae" title="Alexiidae">Alexiidae</a></li> <li><a href="/wiki/Anamorphidae" title="Anamorphidae">Anamorphidae</a></li> <li><a href="/wiki/Bothrideridae" title="Bothrideridae">Bothrideridae <small>(dry bark beetles)</small></a></li> <li><a href="/wiki/Cerylonidae" title="Cerylonidae">Cerylonidae <small>(minute bark beetles)</small></a></li> <li><a href="/wiki/Coccinellidae" title="Coccinellidae">Coccinellidae <small>(lady beetles, or God's cows)</small></a></li> <li><a href="/wiki/Corylophidae" title="Corylophidae">Corylophidae <small>(minute fungus beetles)</small></a></li> <li><a href="/wiki/Discolomatidae" title="Discolomatidae">Discolomatidae</a></li> <li><a href="/wiki/Endomychidae" title="Endomychidae">Endomychidae <small>(handsome fungus beetles)</small></a></li> <li><a href="/wiki/Eupsilobiidae" title="Eupsilobiidae">Eupsilobiidae</a></li> <li><a href="/wiki/Euxestidae" title="Euxestidae">Euxestidae <small>(well polished beetles)</small></a></li> <li><a href="/wiki/Latridiidae" title="Latridiidae">Latridiidae <small>(minute brown scavenger beetles)</small></a></li> <li><a href="/wiki/Murmidiidae" title="Murmidiidae">Murmidiidae</a></li> <li><a href="/wiki/Mycetaeidae" title="Mycetaeidae">Mycetaeidae</a></li> <li><a href="/wiki/Teredidae" title="Teredidae">Teredidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Cucujoidea" title="Cucujoidea">Cucujoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agapytho" title="Agapytho">Agapythidae <small>(<i>Agapytho foveicollis</i>)</small></a></li> <li><a href="/wiki/Boganiidae" title="Boganiidae">Boganiidae</a></li> <li><a href="/wiki/Cavognathidae" title="Cavognathidae">Cavognathidae</a></li> <li><a href="/wiki/Cryptophagidae" title="Cryptophagidae">Cryptophagidae <small>(silken fungus beetles)</small></a></li> <li><a href="/wiki/Cucujidae" title="Cucujidae">Cucujidae <small>(flat bark beetles)</small></a></li> <li><a href="/wiki/Cybocephalidae" title="Cybocephalidae">Cybocephalidae</a></li> <li><a href="/wiki/Cyclaxyridae" title="Cyclaxyridae">Cyclaxyridae</a></li> <li><a href="/wiki/Erotylidae" title="Erotylidae">Erotylidae <small>(pleasing fungus beetles)</small></a></li> <li><a href="/wiki/Helotidae" title="Helotidae">Helotidae</a></li> <li><a href="/wiki/Hobartiidae" title="Hobartiidae">Hobartiidae</a></li> <li><a href="/wiki/Kateretidae" title="Kateretidae">Kateretidae <small>(short-winged flower beetles)</small></a></li> <li><a href="/wiki/Laemophloeidae" title="Laemophloeidae">Laemophloeidae <small>(lined flat bark beetles)</small></a></li> <li><a href="/wiki/Lamingtonium" title="Lamingtonium">Lamingtoniidae <small>(<i>Lamingtonium binnaberrense</i>)</small></a></li> <li><a href="/wiki/Monotomidae" title="Monotomidae">Monotomidae <small>(root-eating beetles)</small></a></li> <li><a href="/wiki/Myraboliidae" class="mw-redirect" title="Myraboliidae">Myraboliidae</a></li> <li><a href="/wiki/Sap_beetle" title="Sap beetle">Nitidulidae <small>(sap beetles)</small></a></li> <li><a href="/wiki/Passandridae" title="Passandridae">Passandridae <small>(parasitic flat bark beetles)</small></a></li> <li><a href="/wiki/Phalacridae" title="Phalacridae">Phalacridae <small>(shining flower beetles)</small></a></li> <li><a href="/wiki/Phloeostichidae" title="Phloeostichidae">Phloeostichidae</a></li> <li><a href="/wiki/Priasilphidae" title="Priasilphidae">Priasilphidae</a></li> <li><a href="/wiki/Propalticidae" title="Propalticidae">Propalticidae</a></li> <li><a href="/wiki/Protocucujidae" title="Protocucujidae">Protocucujidae</a></li> <li><a href="/wiki/Silvanidae" title="Silvanidae">Silvanidae <small>(silvanid flat bark beetles)</small></a></li> <li><a href="/wiki/Smicripidae" title="Smicripidae">Smicripidae <small>(palmetto beetles)</small></a></li> <li><a href="/wiki/Sphindidae" title="Sphindidae">Sphindidae <small>(dry-fungus beetles)</small></a></li> <li><a href="/wiki/Tasmosalpingus" title="Tasmosalpingus">Tasmosalpingidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Weevil" title="Weevil">Curculionoidea</a><br /><small><span style="color:#696969"><span class="nobold">(weevils)</span></span></small></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Anthribidae" title="Anthribidae">Anthribidae <small>(fungus weevils)</small></a></li> <li><a href="/wiki/Attelabidae" title="Attelabidae">Attelabidae <small>(leaf-rolling weevils)</small></a></li> <li><a href="/wiki/Belidae" title="Belidae">Belidae <small>(primitive weevils)</small></a></li> <li><a href="/wiki/Brentidae" title="Brentidae">Brentidae <small>(straight snout weevils, New York weevil)</small></a></li> <li><a href="/wiki/Caridae" title="Caridae">Caridae</a></li> <li><a href="/wiki/Curculionidae" title="Curculionidae">Curculionidae <small>(true weevils, bark beetles, ambrosia beetles)</small></a></li> <li><a href="/wiki/Nemonychidae" title="Nemonychidae">Nemonychidae <small>(pine flower weevils)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Lymexylidae" title="Lymexylidae">Lymexyloidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Lymexylidae" title="Lymexylidae">Lymexylidae <small>(ship-timber beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Tenebrionoidea" title="Tenebrionoidea">Tenebrionoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Aderidae" title="Aderidae">Aderidae <small>(ant-like leaf beetles)</small></a></li> <li><a href="/wiki/Anthicidae" title="Anthicidae">Anthicidae <small>(ant-like flower beetles)</small></a></li> <li><a href="/wiki/Archeocrypticidae" title="Archeocrypticidae">Archeocrypticidae <small>(cryptic fungus beetles)</small></a></li> <li><a href="/wiki/Boridae" title="Boridae">Boridae <small>(conifer bark beetles)</small></a></li> <li><a href="/wiki/Chalcodryidae" title="Chalcodryidae">Chalcodryidae</a></li> <li><a href="/wiki/Ciidae" title="Ciidae">Ciidae <small>(minute tree-fungus beetles)</small></a></li> <li><a href="/wiki/Melandryidae" title="Melandryidae">Melandryidae <small>(false darkling beetles)</small></a></li> <li><a href="/wiki/Blister_beetle" title="Blister beetle">Meloidae <small>(blister beetles)</small></a></li> <li><a href="/wiki/Mordellidae" title="Mordellidae">Mordellidae <small>(tumbling flower beetles)</small></a></li> <li><a href="/wiki/Mycetophagidae" title="Mycetophagidae">Mycetophagidae <small>(hairy fungus beetles)</small></a></li> <li><a href="/wiki/Mycteridae" title="Mycteridae">Mycteridae <small>(palm and flower beetles)</small></a></li> <li><a href="/wiki/Oedemeridae" title="Oedemeridae">Oedemeridae <small>(false blister beetle)</small></a></li> <li><a href="/wiki/Perimylopidae" class="mw-redirect" title="Perimylopidae">Perimylopidae, or Promecheilidae</a></li> <li><a href="/wiki/Prostomidae" title="Prostomidae">Prostomidae <small>(jugular-horned beetles)</small></a></li> <li><a href="/wiki/Pterogeniidae" title="Pterogeniidae">Pterogeniidae</a></li> <li><a href="/wiki/Fire-coloured_beetle" title="Fire-coloured beetle">Pyrochroidae <small>(fire-coloured beetles)</small></a></li> <li><a href="/wiki/Pythidae" title="Pythidae">Pythidae <small>(dead log bark beetles)</small></a></li> <li><a href="/wiki/Ripiphoridae" title="Ripiphoridae">Ripiphoridae <small>(wedge-shaped beetles)</small></a></li> <li><a href="/wiki/Salpingidae" title="Salpingidae">Salpingidae <small>(narrow-waisted bark beetles)</small></a></li> <li><a href="/wiki/Scraptiidae" title="Scraptiidae">Scraptiidae <small>(false flower beetles)</small></a></li> <li><a href="/wiki/Stenotrachelidae" title="Stenotrachelidae">Stenotrachelidae <small>(false longhorn beetles)</small></a></li> <li><a href="/wiki/Synchroidae" title="Synchroidae">Synchroidae <small>(synchroa bark beetles)</small></a></li> <li><a href="/wiki/Darkling_beetle" title="Darkling beetle">Tenebrionidae <small>(darkling beetles)</small></a></li> <li><a href="/wiki/Tetratomidae" title="Tetratomidae">Tetratomidae <small>(polypore fungus beetles)</small></a></li> <li><a href="/wiki/Trictenotomidae" title="Trictenotomidae">Trictenotomidae</a></li> <li><a href="/wiki/Ulodidae" title="Ulodidae">Ulodidae</a></li> <li><a href="/wiki/Zopheridae" title="Zopheridae">Zopheridae <small>(ironclad beetles, cylindrical bark beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Elateriformia" title="Elateriformia">Elateriformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Buprestoidea" title="Buprestoidea">Buprestoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Buprestidae" title="Buprestidae">Buprestidae <small>(jewel beetles, or metallic wood-boring beetles)</small></a></li> <li><a href="/wiki/Schizopodidae" title="Schizopodidae">Schizopodidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Byrrhoidea" title="Byrrhoidea">Byrrhoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Byrrhidae" title="Byrrhidae">Byrrhidae <small>(pill beetles)</small></a></li> <li><a href="/wiki/Callirhipidae" title="Callirhipidae">Callirhipidae <small>(cedar beetles)</small></a></li> <li><a href="/wiki/Chelonariidae" title="Chelonariidae">Chelonariidae <small>(turtle beetles)</small></a></li> <li><a href="/wiki/Cneoglossidae" title="Cneoglossidae">Cneoglossidae</a></li> <li><a href="/wiki/Dryopidae" title="Dryopidae">Dryopidae <small>(long-toed water beetles)</small></a></li> <li><a href="/wiki/Elmidae" title="Elmidae">Elmidae <small>(riffle beetles)</small></a></li> <li><a href="/wiki/Eulichadidae" title="Eulichadidae">Eulichadidae <small>(forest stream beetles)</small></a></li> <li><a href="/wiki/Heteroceridae" title="Heteroceridae">Heteroceridae <small>(variegated mud-loving beetles)</small></a></li> <li><a href="/wiki/Limnichidae" title="Limnichidae">Limnichidae <small>(minute mud beetles)</small></a></li> <li><a href="/wiki/Lutrochidae" class="mw-redirect" title="Lutrochidae">Lutrochidae <small>(travertine beetles)</small></a></li> <li><a href="/wiki/Water-penny_beetles" class="mw-redirect" title="Water-penny beetles">Psephenidae <small>(water-penny beetles)</small></a></li> <li><a href="/wiki/Ptilodactylidae" title="Ptilodactylidae">Ptilodactylidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Dascilloidea" title="Dascilloidea">Dascilloidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Dascillidae" title="Dascillidae">Dascillidae <small>(soft bodied plant beetles)</small></a></li> <li><a href="/wiki/Rhipiceridae" title="Rhipiceridae">Rhipiceridae <small>(cicada beetle, cicada parasite beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Elateroidea" title="Elateroidea">Elateroidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Artematopodidae" title="Artematopodidae">Artematopodidae <small>(soft-bodied plant beetles)</small></a></li> <li><a href="/wiki/Brachypsectridae" title="Brachypsectridae">Brachypsectridae <small>(Texas beetles)</small></a></li> <li><a href="/wiki/Soldier_beetle" title="Soldier beetle">Cantharidae <small>(soldier beetles)</small></a></li> <li><a href="/wiki/Cerophytidae" title="Cerophytidae">Cerophytidae <small>(rare click beetles)</small></a></li> <li><a href="/wiki/Click_beetle" title="Click beetle">Elateridae <small>(click beetles)</small></a></li> <li><a href="/wiki/Eucnemidae" title="Eucnemidae">Eucnemidae <small>(false click beetles)</small></a></li> <li><a href="/wiki/Jurasaidae" title="Jurasaidae">Jurasaidae</a></li> <li><a href="/wiki/Firefly" title="Firefly">Lampyridae <small>(fireflies)</small></a></li> <li><a href="/wiki/Lycidae" title="Lycidae">Lycidae <small>(net-winged beetles)</small></a></li> <li><a href="/wiki/Omethidae" title="Omethidae">Omethidae <small>(false fireflies, long-lipped beetles)</small></a></li> <li><a href="/wiki/Phengodidae" title="Phengodidae">Phengodidae <small>(glowworm beetles)</small></a></li> <li><a href="/wiki/Rhagophthalmidae" title="Rhagophthalmidae">Rhagophthalmidae</a></li> <li><a href="/wiki/Sinopyrophoridae" class="mw-redirect" title="Sinopyrophoridae">Sinopyrophoridae</a></li> <li><a href="/wiki/Throscidae" title="Throscidae">Throscidae <small>(false metallic wood-boring beetles)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Rhinorhipus" title="Rhinorhipus">Rhinorhipoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Rhinorhipus" title="Rhinorhipus">Rhinorhipidae <small>(<i>Rhinorhipus tamborinensis</i>)</small></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Scirtoidea" title="Scirtoidea">Scirtoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Clambidae" title="Clambidae">Clambidae</a></li> <li><a href="/wiki/Declinia" title="Declinia">Decliniidae <small>(<i>Declinia relicta</i>)</small></a></li> <li><a href="/wiki/Eucinetidae" title="Eucinetidae">Eucinetidae <small>(plate-thigh beetles)</small></a></li> <li><a href="/wiki/Scirtidae" title="Scirtidae">Scirtidae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Scarabaeoidea" title="Scarabaeoidea">Scarabaeiformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Scarabaeoidea" title="Scarabaeoidea">Scarabaeoidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Belohina" title="Belohina">Belohinidae <small>(<i>Belohina inexpectata</i>)</small></a></li> <li><a href="/wiki/Bolboceratidae" title="Bolboceratidae">Bolboceratidae</a></li> <li><a href="/wiki/False_stag_beetle" title="False stag beetle">Diphyllostomatidae <small>(false stag beetles)</small></a></li> <li><a href="/wiki/Geotrupidae" title="Geotrupidae">Geotrupidae <small>(dor beetles)</small></a></li> <li><a href="/wiki/Glaphyridae" title="Glaphyridae">Glaphyridae <small>(bumble bee scarab beetles)</small></a></li> <li><a href="/wiki/Glaresis" title="Glaresis">Glaresidae <small>(enigmatic scarab beetles)</small></a></li> <li><a href="/wiki/Hybosoridae" title="Hybosoridae">Hybosoridae <small>(scavenger scarab beetles)</small></a></li> <li><a href="/wiki/Stag_beetle" title="Stag beetle">Lucanidae <small>(stag beetles)</small></a></li> <li><a href="/wiki/Ochodaeidae" title="Ochodaeidae">Ochodaeidae <small>(sand-loving scarab beetles)</small></a></li> <li><a href="/wiki/Passalidae" title="Passalidae">Passalidae <small>(betsy beetles)</small></a></li> <li><a href="/wiki/Rain_beetle" title="Rain beetle">Pleocomidae <small>(rain beetles)</small></a></li> <li><a href="/wiki/Scarabaeidae" title="Scarabaeidae">Scarabaeidae <small>(scarabs)</small></a></li> <li><a href="/wiki/Trogidae" title="Trogidae">Trogidae <small>(hide beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Staphyliniformia" title="Staphyliniformia">Staphyliniformia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Histeroidea" title="Histeroidea">Histeroidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Histeridae" title="Histeridae">Histeridae <small>(clown beetles)</small></a></li> <li><a href="/wiki/Sphaerites" title="Sphaerites">Sphaeritidae <small>(false clown beetles)</small></a></li> <li><a href="/wiki/Syntelia" title="Syntelia">Synteliidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Hydrophiloidea" title="Hydrophiloidea">Hydrophiloidea</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Epimetopidae" title="Epimetopidae">Epimetopidae</a></li> <li><a href="/wiki/Georissus" title="Georissus">Georissidae <small>(minute mud-loving beetles)</small></a></li> <li><a href="/wiki/Helophorus" title="Helophorus">Helophoridae</a></li> <li><a href="/wiki/Hydrochidae" class="mw-redirect" title="Hydrochidae">Hydrochidae</a></li> <li><a href="/wiki/Hydrophilidae" title="Hydrophilidae">Hydrophilidae <small>(water scavenger beetles)</small></a></li> <li><a href="/wiki/Spercheidae" class="mw-redirect" title="Spercheidae">Spercheidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:10em"><a href="/wiki/Staphylinoidea" title="Staphylinoidea">Staphylinoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agyrtidae" title="Agyrtidae">Agyrtidae <small>(primitive carrion beetles)</small></a></li> <li><a href="/wiki/Hydraenidae" title="Hydraenidae">Hydraenidae</a></li> <li><a href="/wiki/Leiodidae" title="Leiodidae">Leiodidae <small>(round fungus beetles)</small></a></li> <li><a href="/wiki/Ptiliidae" title="Ptiliidae">Ptiliidae <small>(feather-winged beetles)</small></a></li> <li><a href="/wiki/Silphidae" title="Silphidae">Silphidae <small>(carrion beetles)</small></a></li> <li><a href="/wiki/Rove_beetle" title="Rove beetle">Staphylinidae <small>(rove beetles)</small></a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div><a href="/wiki/List_of_subgroups_of_the_order_Coleoptera" title="List of subgroups of the order Coleoptera">List of subgroups of the order Coleoptera</a></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Cleridae</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q377988" class="extiw" title="wikidata:Q377988">Q377988</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Cleridae" class="extiw" title="wikispecies:Cleridae">Cleridae</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Australian_Faunal_Directory" title="Australian Faunal Directory">AFD</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://biodiversity.org.au/afd/taxa/Cleridae">Cleridae</a></span></span></li> <li><span style="white-space:nowrap;">BioLib: <span class="uid"><a rel="nofollow" class="external text" href="https://www.biolib.cz/en/taxon/id9979">9979</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Barcode_of_Life_Data_System" title="Barcode of Life Data System">BOLD</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://www.boldsystems.org/index.php/TaxBrowser_TaxonPage?taxid=24402">24402</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/BugGuide" title="BugGuide">BugGuide</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://bugguide.net/node/view/113">113</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Catalogue_of_Life" title="Catalogue of Life">CoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.catalogueoflife.org/data/taxon/8BP">8BP</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/7460">7460</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/EPPO_Code" title="EPPO Code">EPPO</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://gd.eppo.int/taxon/1CLERF">1CLERF</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Fauna_Europaea" title="Fauna Europaea">Fauna Europaea</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://www.eu-nomen.eu/portal/taxon.php?GUID=urn:lsid:faunaeur.org:taxname:11033">11033</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Fauna_Europaea" title="Fauna Europaea">Fauna Europaea (new)</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://fauna-eu.org/cdm_dataportal/taxon/960dc211-c8d1-4463-a368-4664a3a8697e">960dc211-c8d1-4463-a368-4664a3a8697e</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/3795">3795</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/INaturalist" title="INaturalist">iNaturalist</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://inaturalist.org/taxa/55051">55051</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&id=115923">115923</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Integrated_Taxonomic_Information_System" title="Integrated Taxonomic Information System">ITIS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=113852">113852</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Biodiversity_Network" title="National Biodiversity Network">NBN</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://data.nbn.org.uk/Taxa/NHMSYS0001717699">NHMSYS0001717699</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=186087">186087</a></span></span></li> <li><span style="white-space:nowrap;">NZOR: <span class="uid"><a rel="nofollow" class="external text" href="https://www.nzor.org.nz/names/c93ca142-b3f9-46f4-a408-3911cdf55287">c93ca142-b3f9-46f4-a408-3911cdf55287</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=192254">192254</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=69399">69399</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Plazi" title="Plazi">Plazi</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://treatment.plazi.org/id/396787A7-5C40-1B48-FF1D-FD7FFD62F14C">396787A7-5C40-1B48-FF1D-FD7FFD62F14C</a></span></span></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"><style data-mw-deduplicate="TemplateStyles:r1038841319">.mw-parser-output .tooltip-dotted{border-bottom:1px dotted;cursor:help}</style></div><div role="navigation" class="navbox authority-control" aria-label="Navbox" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Help:Authority_control" title="Help:Authority control">Authority control databases</a>: National <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a 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