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class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Effect_on_host_immunity" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Effect_on_host_immunity"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>Effect on host immunity</span> </div> </a> <ul id="toc-Effect_on_host_immunity-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Virus-like_particles" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Virus-like_particles"> <div class="vector-toc-text"> <span class="vector-toc-numb">8</span> <span>Virus-like particles</span> </div> </a> <ul id="toc-Virus-like_particles-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Micro-RNA" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Micro-RNA"> <div class="vector-toc-text"> <span class="vector-toc-numb">9</span> <span>Micro-RNA</span> </div> </a> <ul id="toc-Micro-RNA-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-See_also" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#See_also"> <div class="vector-toc-text"> <span class="vector-toc-numb">10</span> <span>See also</span> </div> </a> <ul id="toc-See_also-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">11</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span 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Please update your browser bookmarks and please use the default browser print function instead.</div></div><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Family of viruses</div> <p class="mw-empty-elt"> </p> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(250,250,190)">Polydnavirus </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:GiBV-em.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/6/6e/GiBV-em.jpg/220px-GiBV-em.jpg" decoding="async" width="220" height="201" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/6/6e/GiBV-em.jpg 1.5x" data-file-width="293" data-file-height="268" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%">Electron micrograph of a bracovirus </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(250,250,190)"><a href="/wiki/Virus_classification" title="Virus classification">Virus classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Polydnaviriformidae" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>(unranked): </td> <td><a href="/wiki/Viriform" class="mw-redirect" title="Viriform">Viriform</a> </td></tr> <tr> <td>Family: </td> <td><b><i>Polydnaviriformidae</i></b> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(250,250,190)">Genera </th></tr> <tr> <td colspan="2" style="text-align: left"> <ul><li><i><a href="/wiki/Bracovirus" title="Bracovirus">Bracoviriform</a></i></li> <li><i><a href="/wiki/Ichnovirus" title="Ichnovirus">Ichnoviriform</a></i></li></ul> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(250,250,190)"><a href="/wiki/Synonym_(taxonomy)" title="Synonym (taxonomy)">Synonyms</a> </th></tr> <tr> <td colspan="2" style="text-align: left"> <ul><li><i>Polydnaviriformidae</i> <small>ICTV 2021</small></li> <li><i>Polydnaviridae</i> <small>ICTV 1984</small></li></ul> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><i><b>Polydnaviriformidae</b></i> (<span class="rt-commentedText nowrap"><span class="IPA nopopups noexcerpt" lang="en-fonipa"><a href="/wiki/Help:IPA/English" title="Help:IPA/English">/<span style="border-bottom:1px dotted"><span title="'p' in 'pie'">p</span><span title="/ɒ/: 'o' in 'body'">ɒ</span><span title="/ˈ/: primary stress follows">ˈ</span><span title="'l' in 'lie'">l</span><span title="/ɪ/: 'i' in 'kit'">ɪ</span><span title="'d' in 'dye'">d</span><span title="'n' in 'nigh'">n</span><span title="/ə/: 'a' in 'about'">ə</span><span title="'v' in 'vie'">v</span><span title="/ɪ/: 'i' in 'kit'">ɪ</span><span title="'r' in 'rye'">r</span><span title="/ə/: 'a' in 'about'">ə</span><span title="/ˌ/: secondary stress follows">ˌ</span><span title="'f' in 'find'">f</span><span title="/ɔː/: 'au' in 'fraud'">ɔː</span><span title="'m' in 'my'">m</span><span title="/ɪ/: 'i' in 'kit'">ɪ</span><span title="'d' in 'dye'">d</span><span title="/ɛ/: 'e' in 'dress'">ɛ</span></span>/</a></span></span> <b>PDV</b>)<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> is a <a href="/wiki/Family_(biology)" title="Family (biology)">family</a> of <a href="/wiki/Insect" title="Insect">insect</a> <a href="/wiki/Viriform" class="mw-redirect" title="Viriform">viriforms</a>; members are known as <b>polydnaviruses</b>. There are two genera in the family: <i><a href="/wiki/Bracovirus" title="Bracovirus">Bracoform</a></i> and <i><a href="/wiki/Ichnovirus" title="Ichnovirus">Ichnoviriform</a></i>. Polydnaviruses form a <a href="/wiki/Symbiotic_relationship" class="mw-redirect" title="Symbiotic relationship">symbiotic relationship</a> with <a href="/wiki/Parasitoid_wasps" class="mw-redirect" title="Parasitoid wasps">parasitoid wasps</a>. <a href="/wiki/Ichnovirus" title="Ichnovirus">Ichnoviriforms</a> (IV) occur in <a href="/wiki/Ichneumonidae" title="Ichneumonidae">Ichneumonid wasps</a> and <a href="/wiki/Bracovirus" title="Bracovirus">Bracoviriforms</a> (BV) in <a href="/wiki/Braconidae" title="Braconidae">Braconid wasps</a>. The larvae of wasps in both of those groups are themselves parasitic on <a href="/wiki/Lepidoptera" title="Lepidoptera">Lepidoptera</a> (moths and butterflies), and the polydnaviruses are important in circumventing the immune response of their parasitized hosts.<sup id="cite_ref-ViralZone_2-0" class="reference"><a href="#cite_note-ViralZone-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-ICTV_3-0" class="reference"><a href="#cite_note-ICTV-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Little or no <a href="/wiki/Sequence_homology" title="Sequence homology">sequence homology</a> exists between BV and IV, suggesting that the two genera have been evolving independently for a long time. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Taxonomy">Taxonomy</h2></div> <p>Bracoviriform </p> <style data-mw-deduplicate="TemplateStyles:r1184024115">.mw-parser-output .div-col{margin-top:0.3em;column-width:30em}.mw-parser-output .div-col-small{font-size:90%}.mw-parser-output .div-col-rules{column-rule:1px solid #aaa}.mw-parser-output .div-col dl,.mw-parser-output .div-col ol,.mw-parser-output .div-col ul{margin-top:0}.mw-parser-output .div-col li,.mw-parser-output .div-col dd{page-break-inside:avoid;break-inside:avoid-column}</style><div class="div-col" style="column-width: 18em;"> <ul><li>Bracoviriform altitudinis</li> <li>Bracoviriform argentifrontis</li> <li>Bracoviriform blackburni</li> <li>Bracoviriform canadense</li> <li>Bracoviriform congregatae</li> <li>Bracoviriform crassicornis</li> <li>Bracoviriform croceipedis</li> <li>Bracoviriform curvimaculati</li> <li>Bracoviriform demolitoris</li> <li>Bracoviriform ectdytolophae</li> <li>Bracoviriform facetosae</li> <li>Bracoviriform flavicoxis</li> <li>Bracoviriform flavipedis</li> <li>Bracoviriform flavitestaceae</li> <li>Bracoviriform fumiferanae</li> <li>Bracoviriform glomeratae</li> <li>Bracoviriform hyphantriae</li> <li>Bracoviriform inaniti</li> <li>Bracoviriform indiense</li> <li>Bracoviriform insularis</li> <li>Bracoviriform kariyai</li> <li>Bracoviriform liparidis</li> <li>Bracoviriform marginiventris</li> <li>Bracoviriform melanoscelae</li> <li>Bracoviriform nigricipitis</li> <li>Bracoviriform ornigis</li> <li>Bracoviriform paleacritae</li> <li>Bracoviriform quadridentatae</li> <li>Bracoviriform rubeculae</li> <li>Bracoviriform schaeferi</li> <li>Bracoviriform texani</li></ul> </div> <p>Ichnoviriform </p> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1184024115"><div class="div-col" style="column-width: 18em;"> <ul><li>Ichnoviriform acronyctae</li> <li>Ichnoviriform annulipedis</li> <li>Ichnoviriform aprilis</li> <li>Ichnoviriform arjunae</li> <li>Ichnoviriform benefactoris</li> <li>Ichnoviriform eribori</li> <li>Ichnoviriform exiguae</li> <li>Ichnoviriform flavicinctae</li> <li>Ichnoviriform forcipatae</li> <li>Ichnoviriform fugitivi</li> <li>Ichnoviriform fumiferanae</li> <li>Ichnoviriform geniculatae</li> <li>Ichnoviriform infestae</li> <li>Ichnoviriform interrupti</li> <li>Ichnoviriform lymantriae</li> <li>Ichnoviriform montani</li> <li>Ichnoviriform pilosuli</li> <li>Ichnoviriform rivalis</li> <li>Ichnoviriform rostralis</li> <li>Ichnoviriform sonorense</li> <li>Ichnoviriform tenuifemoris</li> <li>Ichnoviriform terebrantis</li></ul> </div> <div class="mw-heading mw-heading2"><h2 id="Structure">Structure</h2></div> <p>Viruses in <i>Polydnaviridae</i> are <a href="/wiki/Viral_envelope" title="Viral envelope">enveloped</a>, with <a href="/wiki/Prolate" class="mw-redirect" title="Prolate">prolate</a> <a href="/wiki/Ellipsoid" title="Ellipsoid">ellipsoid</a> and cylindrical geometries. <a href="/wiki/Genomes" class="mw-redirect" title="Genomes">Genomes</a> are circular and segmented, composed of multiple segments of double-stranded, <a href="/wiki/Superhelical_DNA" class="mw-redirect" title="Superhelical DNA">superhelical DNA</a> packaged in <a href="/wiki/Capsid" title="Capsid">capsid</a> <a href="/wiki/Proteins" class="mw-redirect" title="Proteins">proteins</a>. They are around 2.0–31kb in length.<sup id="cite_ref-ViralZone_2-1" class="reference"><a href="#cite_note-ViralZone-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p> <table class="wikitable sortable" style="text-align:center"> <tbody><tr> <th>Genus</th> <th>Structure</th> <th>Symmetry</th> <th>Capsid</th> <th>Genomic arrangement</th> <th>Genomic segmentation </th></tr> <tr> <td><i>Ichnoviriform</i></td> <td>Prolate ellipsoid</td> <td></td> <td>Enveloped</td> <td>Circular</td> <td>Segmented </td></tr> <tr> <td><i>Bracoviriform</i></td> <td>Prolate ellipsoid</td> <td></td> <td>Enveloped</td> <td>Circular</td> <td>Segmented </td></tr></tbody></table> <div class="mw-heading mw-heading2"><h2 id="Life_cycle">Life cycle</h2></div> <p>Viral replication is nuclear. DNA-templated <a href="/wiki/Transcription_(biology)" title="Transcription (biology)">transcription</a> is the method of transcription. The virus exits the host cell by <a href="/wiki/Nuclear_pore#Export_of_proteins" title="Nuclear pore">nuclear pore export</a>. </p><p>Parasitoid wasps serve as hosts for the virus, and Lepidoptera serve as hosts for these wasps. The female wasp injects one or more eggs into its host along with a quantity of virus. The virus and wasp are in a <a href="/wiki/Mutualism_(biology)" title="Mutualism (biology)">mutualistic</a> <a href="/wiki/Symbiotic" class="mw-redirect" title="Symbiotic">symbiotic</a> relationship: expression of viral genes prevents the wasp's host's immune system from killing the wasp's injected egg and causes other physiological alterations that ultimately cause the parasitized host to die. Transmission routes are parental.<sup id="cite_ref-ViralZone_2-2" class="reference"><a href="#cite_note-ViralZone-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p> <table class="wikitable sortable" style="text-align:center"> <tbody><tr> <th>Genus</th> <th>Host details</th> <th>Tissue tropism</th> <th>Entry details</th> <th>Release details</th> <th>Replication site</th> <th>Assembly site</th> <th>Transmission </th></tr> <tr> <td><i>Ichnoviriform</i></td> <td>Parasitoid wasps (Ichneumonidae)</td> <td>Hemocytes; fat bodies</td> <td>Unknown</td> <td>Budding through cell membrane<sup id="cite_ref-ViralZone_2-3" class="reference"><a href="#cite_note-ViralZone-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-webb_4-0" class="reference"><a href="#cite_note-webb-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup></td> <td>Nucleus</td> <td>Nucleus</td> <td>Unknown </td></tr> <tr> <td><i>Bracoviriform</i></td> <td>Parasitoid wasps (Braconidae)</td> <td>Hemocytes; fat bodies</td> <td>Unknown</td> <td><a href="/wiki/Lysis" title="Lysis">Lysis</a><sup id="cite_ref-ViralZone_2-4" class="reference"><a href="#cite_note-ViralZone-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup></td> <td>Nucleus</td> <td>Nucleus</td> <td>Unknown </td></tr></tbody></table> <div class="mw-heading mw-heading2"><h2 id="Biology">Biology</h2></div> <figure class="mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Pdv-host.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e6/Pdv-host.svg/288px-Pdv-host.svg.png" decoding="async" width="288" height="232" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/e6/Pdv-host.svg/432px-Pdv-host.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/e6/Pdv-host.svg/576px-Pdv-host.svg.png 2x" data-file-width="1150" data-file-height="925" /></a><figcaption>Diagram of a PDV host association</figcaption></figure> <p>These viruses are part of a unique biological system consisting of an endoparasitic wasp (<a href="/wiki/Parasitoid" title="Parasitoid">parasitoid</a>), a host (usually <a href="/wiki/Lepidoptera" title="Lepidoptera">lepidopteran</a>) larva, and the virus. The full <a href="/wiki/Genome" title="Genome">genome</a> of the virus is <a href="/wiki/Endogenous" class="mw-redirect" title="Endogenous">endogenous</a>, dispersed among the genome of the wasp. The virus only replicates in a particular part of the ovary, called the <a href="/wiki/Calyx_(anatomy)#Insects" title="Calyx (anatomy)">calyx</a>, of pupal and adult female wasps. The virus is injected along with the wasp egg into the body cavity of a lepidopteran host caterpillar and infects cells of the caterpillar. The infection does not lead to replication of new viruses; rather, it affects the caterpillar's <a href="/wiki/Immune_system" title="Immune system">immune system</a>, as the virion carries virulence genes instead of viral replication genes.<sup id="cite_ref-pmid25670535_5-0" class="reference"><a href="#cite_note-pmid25670535-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> It can be considered a type of <a href="/wiki/Viral_vector" title="Viral vector">viral vector</a>.<sup id="cite_ref-vec_6-0" class="reference"><a href="#cite_note-vec-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p><p>Without the virus infection, <a href="/wiki/Phagocyte" title="Phagocyte">phagocytic</a> <a href="/wiki/Hemocyte_(invertebrate_immune_system_cell)" title="Hemocyte (invertebrate immune system cell)">hemocytes</a> (blood cells) will encapsulate and kill the wasp egg and larvae, but the immune suppression caused by the virus allows survival of the wasp egg and larvae, leading to hatching and complete development of the immature wasp in the caterpillar. Additionally, genes expressed from the polydnavirus in the parasitised host alter host development and metabolism to be beneficial for the growth and survival of the parasitoid larva.<sup id="cite_ref-webb_4-1" class="reference"><a href="#cite_note-webb-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Potential_carrier_subfamilies">Potential carrier subfamilies</h3></div> <ul><li><a href="/wiki/Ichneumonoidea" title="Ichneumonoidea">Ichneumonoidea</a> <ul><li><a href="/wiki/Braconidae" title="Braconidae">Braconidae</a> <ul><li><a href="/wiki/Microgastrinae" title="Microgastrinae">Microgastrinae</a><sup id="cite_ref-webb_4-2" class="reference"><a href="#cite_note-webb-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup></li> <li><a href="/w/index.php?title=Miracinae&action=edit&redlink=1" class="new" title="Miracinae (page does not exist)">Miracinae</a></li> <li><a href="/wiki/Cheloninae" title="Cheloninae">Cheloninae</a><sup id="cite_ref-webb_4-3" class="reference"><a href="#cite_note-webb-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Cardiochilinae" title="Cardiochilinae">Cardiochilinae</a><sup id="cite_ref-webb_4-4" class="reference"><a href="#cite_note-webb-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup></li> <li><a href="/w/index.php?title=Mendeselinae&action=edit&redlink=1" class="new" title="Mendeselinae (page does not exist)">Mendeselinae</a></li> <li><a href="/w/index.php?title=Khoikhoiinae&action=edit&redlink=1" class="new" title="Khoikhoiinae (page does not exist)">Khoikhoiinae</a></li></ul></li> <li><a href="/wiki/Ichneumonidae" title="Ichneumonidae">Ichneumonidae</a> <ul><li><a href="/wiki/Campopleginae" title="Campopleginae">Campopleginae</a><sup id="cite_ref-webb_4-5" class="reference"><a href="#cite_note-webb-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Banchinae" title="Banchinae">Banchinae</a><sup id="cite_ref-webb_4-6" class="reference"><a href="#cite_note-webb-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-vec_6-1" class="reference"><a href="#cite_note-vec-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup></li></ul></li></ul></li></ul> <div class="mw-heading mw-heading2"><h2 id="Characteristics">Characteristics</h2></div> <p>Both genera of PDV share certain characteristics: </p> <ul><li>the virus particles of each contain multiple segments of dsDNA (double-strand, or "normal" DNA, as contrasted with positive- or negative-sense single-strand DNA or RNA, as found in some other viruses) with each segment containing only part of the full genome (much like chromosomes in <a href="/wiki/Eukaryote" title="Eukaryote">eukaryotic organisms</a>)</li> <li>the genome of the virus has eukaryotic characteristics such as the presence of <a href="/wiki/Intron" title="Intron">introns</a> (common for insect genes but rare for viruses) and a low coding density<sup id="cite_ref-virology_2006_9-0" class="reference"><a href="#cite_note-virology_2006-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup></li> <li>the genome of each virus is integrated into the host wasp genome<sup id="cite_ref-integrated_1991_10-0" class="reference"><a href="#cite_note-integrated_1991-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup></li> <li>the genome is organized in several multiple-member genes families (which differ between Bracoviruses and Ichnoviruses)</li> <li>the virus particles are only produced in specific cell types in the female wasp's reproductive organs</li></ul> <p>The morphologies of the two genera are different when observed by electron microscopy. Ichnoviruses tend to be ovoid while bracoviruses are short rods. The virions of Bracoviruses are released by <a href="/wiki/Lysis" title="Lysis">cell lysis</a>; the virions of Ichnoviruses are released by budding. </p> <div class="mw-heading mw-heading2"><h2 id="Evolution">Evolution</h2></div> <p><a href="/wiki/Nucleic_acid" title="Nucleic acid">Nucleic acid</a> analysis suggests a very long association of the viruses with the wasps (estimated 73.7 million years ± 10 million).<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Older_wasp-derived_theory">Older wasp-derived theory</h3></div> <p>Two proposals have been advanced for how the wasp/virus association developed. The first suggests that the virus is derived from wasp genes. Many parasitoids that do not use PDVs inject proteins that provide many of the same functions, that is, a suppression of the immune response to the parasite egg. In this model, the braconid and ichneumonid wasps packaged genes for these functions into the viruses—essentially creating a gene-transfer system that results in the caterpillar producing the immune-suppressing factors. In this scenario, the PDV structural proteins (capsids) were probably "borrowed" from existing viruses.<sup id="cite_ref-Reineke2006_12-0" class="reference"><a href="#cite_note-Reineke2006-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Current_endogenous_virus_theory">Current endogenous virus theory</h3></div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">See also: <a href="/wiki/Endogenous_viral_element" title="Endogenous viral element">Endogenous viral element</a></div> <p>The alternative proposal suggests that ancestral wasps developed a beneficial association with an existing virus that eventually led to the integration of the virus into the wasp's genome. Following integration, the genes responsible for virus replication and the capsids were (eventually) no longer included in the PDV genome. This hypothesis is supported by the distinct morphology differences between IV and BV, suggesting different ancestral viruses for the two genera. BV has likely evolved from a <a href="/wiki/Nudivirus" title="Nudivirus">nudivirus</a>, specifically a betanudivirus,<sup id="cite_ref-pmid28728099_13-0" class="reference"><a href="#cite_note-pmid28728099-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> ~<span class="nourlexpansion plainlinks" style="white-space:nowrap;"><a rel="nofollow" class="external text" href="https://geoltime.github.io/?Ma=100">100</a> million years ago</span>.<sup id="cite_ref-Herniou2013_14-0" class="reference"><a href="#cite_note-Herniou2013-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> IV has a less clear origin: although earlier reports found a protein p44/p53 with structural similarities to <a href="/wiki/Ascovirus" class="mw-redirect" title="Ascovirus">ascovirus</a>, the link was not confirmed in later studies.<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> As a result, the current opinion is that IV originated from a yet-unidentified novel viral family,<sup id="cite_ref-pmid28728099_13-1" class="reference"><a href="#cite_note-pmid28728099-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> with a weak link to the <a href="/wiki/NCLDV" class="mw-redirect" title="NCLDV">NCLDVs</a>.<sup id="cite_ref-pmid26085165_16-0" class="reference"><a href="#cite_note-pmid26085165-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> In either case, both genera were formed through a single integration event in their respective wasp lineages.<sup id="cite_ref-pmid25670535_5-1" class="reference"><a href="#cite_note-pmid25670535-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p><p>The two groups of viruses in the family are not in fact phylogenetically related suggesting that this taxon may need revision.<sup id="cite_ref-Dupuy2006_17-0" class="reference"><a href="#cite_note-Dupuy2006-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Effect_on_host_immunity">Effect on host immunity</h2></div> <p>In the host, several mechanisms of the insect immune system can be triggered when the wasp lays its eggs and when the parasitic wasp is developing. When a large body (wasp egg or small particle used experimentally) is introduced into an insect's body, the classic immune reaction is the encapsulation by hematocytes. An encapsulated body can also be melanised in order to asphyxiate it, thanks to another type of hemocyte, which uses the <a href="/wiki/Phenoloxidase" class="mw-redirect" title="Phenoloxidase">phenoloxidase</a> pathway to produce melanin. Small particles can be phagocytosed, and macrophage cells can then be also melanised in a nodule. Finally, insects can also respond with production of <a href="/wiki/Antimicrobial_peptides" title="Antimicrobial peptides">antiviral peptides</a>.<sup id="cite_ref-Virologie_18-0" class="reference"><a href="#cite_note-Virologie-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> </p><p>PolyDNAvirus protect the hymenopteran larvae from the host immune system, acting at different levels. </p> <ul><li>First they can disable or destroy hematocytes. The polyDNAvirus associated with <i>Cotesia rubecula</i>, code for a protein CrV1 that denatures actin filaments in hematocytes, so those cells become less able to move and adhere to the larvae.<sup id="cite_ref-Virologie_18-1" class="reference"><a href="#cite_note-Virologie-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> <i>Microplitis demolitor</i> Bracovirus (MdBV) induce apoptosis of hematocytes, thanks to its gene PTP-H2.<sup id="cite_ref-Virologie_18-2" class="reference"><a href="#cite_note-Virologie-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> It also decreases the adhesion capacity of hematocytes, thanks to its gene Glc1.8. The gene also inhibits phagocytosis.<sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup></li> <li>PolyDNAvirus can also act on melanisation, MdBV interferes with the production of <a href="/wiki/Phenoloxidase" class="mw-redirect" title="Phenoloxidase">phenoloxidase</a>.<sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup></li> <li>Finally, polyDNAvirus can also produce viral <a href="/wiki/Ankyrin" title="Ankyrin">ankyrins</a>, that interfere with production of antiviral peptides.<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> In some Ichnoviruses, Vankyrin can also prevent apoptosis, the extreme reaction of a cell to block viral propagation.<sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Clavijo2011_23-0" class="reference"><a href="#cite_note-Clavijo2011-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup></li> <li>The Ichnoviruses produce some proteins called vinnexins which have been recognized as homologous to the <a href="/wiki/Innexin" title="Innexin">innexins</a> of insects. They are responsible for the encoding of the structural units of the gap-junctions. These proteins may alter the intercellular communication which could explain the disruption of the <a href="/wiki/Encapsidation" title="Encapsidation">encapsidation</a> process.<sup id="cite_ref-24" class="reference"><a href="#cite_note-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup></li></ul> <div class="mw-heading mw-heading2"><h2 id="Virus-like_particles">Virus-like particles</h2></div> <p>Another strategy used by parasitoid <a href="/wiki/Hymenoptera" title="Hymenoptera">Hymenoptera</a> to protect their offspring is production of <a href="/wiki/Virus-like_particle" title="Virus-like particle">virus-like particles</a>. VLPs are similar to viruses in their structure, but they don't carry any nucleic acid. For example, <i>Venturia canescens</i> (<a href="/wiki/Ichneumonidae" title="Ichneumonidae">Ichneumonidea</a>) and <i><a href="/wiki/Leptopilina" title="Leptopilina">Leptopilina</a> sp.</i> (<a href="/wiki/Figitidae" title="Figitidae">Figitidaea</a>) produce VLPs. </p><p>VLPs can be compared to PolyDNAvirus because they are secreted in the same way, and they both act to protect the larvae against the host's immune system. <i>V. canescens</i>-VLPs (VcVLP1, VcVLP2, VcNEP ...) are produced in the calyx cells before they go to the oviducts. Work in 2006 did not find their link to any viruses and assumed a cellular origin.<sup id="cite_ref-Reineke2006_12-1" class="reference"><a href="#cite_note-Reineke2006-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> More recent comparison links them to highly reshuffled domesticated <a href="/wiki/Nudivirus" title="Nudivirus">Nudivirus</a> sequences. This link produces the name <i>Venturia canescens endogenous nudivirus</i> (VcENV), an alphanudivirus closely related to NlENV found in <i><a href="/wiki/Nilaparvata_lugens" class="mw-redirect" title="Nilaparvata lugens">Nilaparvata lugens</a></i>.<sup id="cite_ref-nv_25-0" class="reference"><a href="#cite_note-nv-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup> </p><p>VLPs protect the Hymenoptera larvae locally, whereas polyDNAvirus can have a more global effect. VLPs allow the larvae to escape the immune system: the larva is not recognised as harmful by its host, or the immune cells can't interact with it thanks to the VLPs.<sup id="cite_ref-Reineke2006_12-2" class="reference"><a href="#cite_note-Reineke2006-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> <i>Venturia canescens</i> uses these instead of polydnaviruses because its ichnovirus has been deactivated.<sup id="cite_ref-nv_25-1" class="reference"><a href="#cite_note-nv-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup> </p><p>The wasp <i>Leptopilina heterotoma</i> secrete VLPs that are able to penetrate into the <a href="/wiki/Hemocyte_(invertebrate_immune_system_cell)" title="Hemocyte (invertebrate immune system cell)">lamellocytes</a>, thanks to specific receptors, and then modify the shape and surface properties of the lamellocytes so they become inefficient and the larvae are safe from encapsulation.<sup id="cite_ref-26" class="reference"><a href="#cite_note-26"><span class="cite-bracket">[</span>26<span class="cite-bracket">]</span></a></sup> The <i>Leptopilina</i> VLPs or mixed-strategy extracellular vesicles (MSEVs) contain some secretion systems. Their evolutionary picture is less clear,<sup id="cite_ref-pmid28889977_27-0" class="reference"><a href="#cite_note-pmid28889977-27"><span class="cite-bracket">[</span>27<span class="cite-bracket">]</span></a></sup> but a recently reported virus, <i>L. boulardi</i> Filamentous Virus (LbFV), shows significant similarities.<sup id="cite_ref-pmid32080746_28-0" class="reference"><a href="#cite_note-pmid32080746-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Micro-RNA">Micro-RNA</h2></div> <p>MicroRNA are small RNA fragments produced in the host cells thanks to a specific enzymatic mechanism. They promote viral RNA destruction. MicroRNA attach to viral-RNA because they are complementary. Then the complex is recognised by an enzyme that destroys it. This phenomenon is known as <a href="/wiki/PTGS" class="mw-redirect" title="PTGS">PTGS</a> (for post transcriptional gene silencing)<sup id="cite_ref-29" class="reference"><a href="#cite_note-29"><span class="cite-bracket">[</span>29<span class="cite-bracket">]</span></a></sup> or RNAi (<a href="/wiki/RNA_interference" title="RNA interference">RNA interference</a>.) </p><p>It is interesting to consider the microRNA phenomenon in the polyDNAvirus context. Many hypotheses can be formulated: </p> <ul><li>Braconidae carry <a href="/wiki/Nudivirus" title="Nudivirus">nudivirus</a>-related genes in their genome,<sup id="cite_ref-30" class="reference"><a href="#cite_note-30"><span class="cite-bracket">[</span>30<span class="cite-bracket">]</span></a></sup> so they may be able to produce microRNA against nudivirus, as an innate immunity.</li> <li>Wasps perhaps use microRNA to control the viral genes they carry.</li> <li>PolyDNAvirus can also use PTGS to interfere with the host's gene expression.</li> <li>PTGS is also used for organisms' development, using the same enzymes as antiviral gene silencing, so we can imagine that if the host uses PTGS against polyDNAvirus, perhaps it also affects its development.</li></ul> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2></div> <ul><li><a href="/wiki/Mutualism_(biology)" title="Mutualism (biology)">Mutualism</a></li></ul> <div class="mw-heading mw-heading2"><h2 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href="http://research.biology.arizona.edu/mosquito/willott/507/polydnaviruses.html">http://research.biology.arizona.edu/mosquito/willott/507/polydnaviruses.html</a></li> <li><a rel="nofollow" class="external text" href="https://viralzone.expasy.org/147"><b>Viralzone</b>: Polydnaviridae</a></li> <li><a rel="nofollow" class="external text" href="https://ictv.global/taxonomy"><b>ICTV</b></a></li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol 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class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="I:_dsDNA_viruses" style="font-size:114%;margin:0 4em">I: <a href="/wiki/DsDNA_virus" class="mw-redirect" title="DsDNA virus">dsDNA viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Adnaviria" title="Adnaviria">Adnaviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Zilligvirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Zilligvirae" class="mw-redirect" title="Zilligvirae">Zilligvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Taleaviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Taleaviricota" class="mw-redirect" title="Taleaviricota">Taleaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Tokiviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tokiviricetes" class="mw-redirect" title="Tokiviricetes">Tokiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ligamenvirales" title="Ligamenvirales">Ligamenvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Lipothrixviridae" title="Lipothrixviridae">Lipothrixviridae</a></i></li> <li><i><a href="/wiki/Rudiviridae" title="Rudiviridae">Rudiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Primavirales" class="mw-redirect" title="Primavirales">Primavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Tristromaviridae" title="Tristromaviridae">Tristromaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Duplodnaviria" title="Duplodnaviria">Duplodnaviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Heunggongvirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Heunggongvirae" class="mw-redirect" title="Heunggongvirae">Heunggongvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Peploviricota" class="mw-redirect" title="Peploviricota">Peploviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Herviviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Herviviricetes" class="mw-redirect" title="Herviviricetes">Herviviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Herpesvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Herpesvirales" title="Herpesvirales">Herpesvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alloherpesviridae" title="Alloherpesviridae">Alloherpesviridae</a></i></li> <li><i><a href="/wiki/Herpesviridae" title="Herpesviridae">Herpesviridae</a></i></li> <li><i><a href="/wiki/Malacoherpesviridae" title="Malacoherpesviridae">Malacoherpesviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Uroviricota" class="mw-redirect" title="Uroviricota">Uroviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Caudoviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Caudoviricetes" title="Caudoviricetes">Caudoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Caudovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Caudovirales" class="mw-redirect" title="Caudovirales">Caudovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Ackermannviridae" title="Ackermannviridae">Ackermannviridae</a></i></li> <li><i><a href="/wiki/Autographiviridae" title="Autographiviridae">Autographiviridae</a></i></li> <li><i><a href="/w/index.php?title=Chaseviridae&action=edit&redlink=1" class="new" title="Chaseviridae (page does not exist)">Chaseviridae</a></i></li> <li><i><a href="/w/index.php?title=Demerecviridae&action=edit&redlink=1" class="new" title="Demerecviridae (page does not exist)">Demerecviridae</a></i></li> <li><i><a href="/w/index.php?title=Drexlerviridae&action=edit&redlink=1" class="new" title="Drexlerviridae (page does not exist)">Drexlerviridae</a></i></li> <li><i><a href="/w/index.php?title=Guelinviridae&action=edit&redlink=1" class="new" title="Guelinviridae (page does not exist)">Guelinviridae</a></i></li> <li><i><a href="/wiki/Herelleviridae" title="Herelleviridae">Herelleviridae</a></i></li> <li><i><a href="/wiki/Myoviridae" title="Myoviridae">Myoviridae</a></i></li> <li><i><a href="/wiki/Podoviridae" title="Podoviridae">Podoviridae</a></i></li> <li><i><a href="/w/index.php?title=Rountreeviridae&action=edit&redlink=1" class="new" title="Rountreeviridae (page does not exist)">Rountreeviridae</a></i></li> <li><i><a href="/w/index.php?title=Salasmaviridae&action=edit&redlink=1" class="new" title="Salasmaviridae (page does not exist)">Salasmaviridae</a></i></li> <li><i><a href="/w/index.php?title=Schitoviridae&action=edit&redlink=1" class="new" title="Schitoviridae (page does not exist)">Schitoviridae</a></i></li> <li><i><a href="/wiki/Siphoviridae" title="Siphoviridae">Siphoviridae</a></i></li> <li><i><a href="/w/index.php?title=Zobellviridae&action=edit&redlink=1" class="new" title="Zobellviridae (page does not exist)">Zobellviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Monodnaviria" title="Monodnaviria">Monodnaviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Shotokuvirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Shotokuvirae" title="Shotokuvirae">Shotokuvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Cossaviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cossaviricota" title="Cossaviricota">Cossaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Papovaviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Papovaviricetes" title="Papovaviricetes">Papovaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Sepolyvirales" class="mw-redirect" title="Sepolyvirales">Sepolyvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Polyomaviridae" title="Polyomaviridae">Polyomaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Zurhausenvirales" class="mw-redirect" title="Zurhausenvirales">Zurhausenvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Papillomaviridae" title="Papillomaviridae">Papillomaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Varidnaviria" title="Varidnaviria">Varidnaviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Bamfordvirae" title="Bamfordvirae">Bamfordvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Nucleocytoviricota" title="Nucleocytoviricota">Nucleocytoviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pokkesviricetes" title="Pokkesviricetes">Pokkesviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Asfuvirales" class="mw-redirect" title="Asfuvirales">Asfuvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Asfarviridae" title="Asfarviridae">Asfarviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Chitovirales" class="mw-redirect" title="Chitovirales">Chitovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Poxviridae" title="Poxviridae">Poxviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Megaviricetes" title="Megaviricetes">Megaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Algavirales" class="mw-redirect" title="Algavirales">Algavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Phycodnaviridae" title="Phycodnaviridae">Phycodnaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Imitervirales" class="mw-redirect" title="Imitervirales">Imitervirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Mimiviridae" title="Mimiviridae">Mimiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pimascovirales" title="Pimascovirales">Pimascovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Ascoviridae" title="Ascoviridae">Ascoviridae</a></i></li> <li><i><a href="/wiki/Iridoviridae" title="Iridoviridae">Iridoviridae</a></i></li> <li><i><a href="/wiki/Marseilleviridae" title="Marseilleviridae">Marseilleviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Preplasmiviricota" title="Preplasmiviricota">Preplasmiviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Maveriviricetes" class="mw-redirect" title="Maveriviricetes">Maveriviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Priklausovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Priklausovirales" class="mw-redirect" title="Priklausovirales">Priklausovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Lavidaviridae" class="mw-redirect" title="Lavidaviridae">Lavidaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Polintoviricetes" class="mw-redirect" title="Polintoviricetes">Polintoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Orthopolintovirales" class="mw-redirect" title="Orthopolintovirales">Orthopolintovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Adintoviridae" title="Adintoviridae">Adintoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tectiliviricetes" title="Tectiliviricetes">Tectiliviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Belfryvirales" class="mw-redirect" title="Belfryvirales">Belfryvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Turriviridae" title="Turriviridae">Turriviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Kalamavirales" class="mw-redirect" title="Kalamavirales">Kalamavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Tectiviridae" class="mw-redirect" title="Tectiviridae">Tectiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Rowavirales" class="mw-redirect" title="Rowavirales">Rowavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Adenoviridae" title="Adenoviridae">Adenoviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Vinavirales" class="mw-redirect" title="Vinavirales">Vinavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Corticoviridae" class="mw-redirect" title="Corticoviridae">Corticoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Helvetiavirae" class="mw-redirect" title="Helvetiavirae">Helvetiavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Dividoviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Dividoviricota" class="mw-redirect" title="Dividoviricota">Dividoviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Laserviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Laserviricetes" class="mw-redirect" title="Laserviricetes">Laserviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Halopanivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Halopanivirales" title="Halopanivirales">Halopanivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Matshushitaviridae" class="mw-redirect" title="Matshushitaviridae">Matshushitaviridae</a></i></li> <li><i><a href="/wiki/Simuloviridae" class="mw-redirect" title="Simuloviridae">Simuloviridae</a></i></li> <li><i><a href="/wiki/Sphaerolipoviridae" class="mw-redirect" title="Sphaerolipoviridae">Sphaerolipoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Naldaviricetes" title="Naldaviricetes">Naldaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Lefavirales" title="Lefavirales">Lefavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Baculoviridae" title="Baculoviridae">Baculoviridae</a></i></li> <li><i><a href="/wiki/Hytrosaviridae" title="Hytrosaviridae">Hytrosaviridae</a></i></li> <li><i><a href="/wiki/Nudiviridae" class="mw-redirect" title="Nudiviridae">Nudiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Nimaviridae" class="mw-redirect" title="Nimaviridae">Nimaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><b>Families</b>: <i><a href="/wiki/Ampullaviridae" title="Ampullaviridae">Ampullaviridae</a></i></li> <li><i><a href="/wiki/Bicaudaviridae" title="Bicaudaviridae">Bicaudaviridae</a></i></li> <li><i><a href="/wiki/Clavaviridae" title="Clavaviridae">Clavaviridae</a></i></li> <li><i><a href="/wiki/Fuselloviridae" title="Fuselloviridae">Fuselloviridae</a></i></li> <li><i><a href="/wiki/Globuloviridae" title="Globuloviridae">Globuloviridae</a></i></li> <li><i><a href="/wiki/Guttaviridae" title="Guttaviridae">Guttaviridae</a></i></li> <li><i><a href="/wiki/Halspiviridae" title="Halspiviridae">Halspiviridae</a></i></li> <li><i><a href="/wiki/Ovaliviridae" title="Ovaliviridae">Ovaliviridae</a></i></li> <li><i><a href="/wiki/Plasmaviridae" title="Plasmaviridae">Plasmaviridae</a></i></li> <li><i><a href="/wiki/Polydnaviridae" class="mw-redirect" title="Polydnaviridae">Polydnaviridae</a></i></li> <li><i><a href="/wiki/Portogloboviridae" title="Portogloboviridae">Portogloboviridae</a></i></li> <li><i><a href="/wiki/Thaspiviridae" title="Thaspiviridae">Thaspiviridae</a></i></li> <li><b>Genera</b>: <i><a href="/wiki/Dinodnavirus" title="Dinodnavirus">Dinodnavirus</a></i></li> <li><i><a href="/wiki/Rhizidiovirus" title="Rhizidiovirus">Rhizidiovirus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="II:_ssDNA_viruses" style="font-size:114%;margin:0 4em">II: <a href="/wiki/SsDNA_virus" class="mw-redirect" title="SsDNA virus">ssDNA viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Monodnaviria" title="Monodnaviria">Monodnaviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Loebvirae" class="mw-redirect" title="Loebvirae">Loebvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Hofneiviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Hofneiviricota" class="mw-redirect" title="Hofneiviricota">Hofneiviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Faserviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Faserviricetes" class="mw-redirect" title="Faserviricetes">Faserviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Tubulavirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tubulavirales" title="Tubulavirales">Tubulavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Inoviridae" class="mw-redirect" title="Inoviridae">Inoviridae</a></i></li> <li><i><a href="/w/index.php?title=Paulinoviridae&action=edit&redlink=1" class="new" title="Paulinoviridae (page does not exist)">Paulinoviridae</a></i></li> <li><i><a href="/w/index.php?title=Plectroviridae&action=edit&redlink=1" class="new" title="Plectroviridae (page does not exist)">Plectroviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Sangervirae" class="mw-redirect" title="Sangervirae">Sangervirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Phixviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Phixviricota" class="mw-redirect" title="Phixviricota">Phixviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Malgrandaviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Malgrandaviricetes" class="mw-redirect" title="Malgrandaviricetes">Malgrandaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Petitvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Petitvirales" class="mw-redirect" title="Petitvirales">Petitvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Microviridae" title="Microviridae">Microviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Shotokuvirae" title="Shotokuvirae">Shotokuvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cossaviricota" title="Cossaviricota">Cossaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Mouviricetes" class="mw-redirect" title="Mouviricetes">Mouviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Polivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Polivirales" class="mw-redirect" title="Polivirales">Polivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Bidnaviridae" class="mw-redirect" title="Bidnaviridae">Bidnaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Quintoviricetes" class="mw-redirect" title="Quintoviricetes">Quintoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Piccovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Piccovirales" class="mw-redirect" title="Piccovirales">Piccovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Parvoviridae" title="Parvoviridae">Parvoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cressdnaviricota" title="Cressdnaviricota">Cressdnaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Arfiviricetes" title="Arfiviricetes">Arfiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Baphyvirales" class="mw-redirect" title="Baphyvirales">Baphyvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Bacilladnaviridae" title="Bacilladnaviridae">Bacilladnaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cirlivirales" class="mw-redirect" title="Cirlivirales">Cirlivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Circoviridae" title="Circoviridae">Circoviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cremevirales" class="mw-redirect" title="Cremevirales">Cremevirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Smacoviridae" title="Smacoviridae">Smacoviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Mulpavirales" title="Mulpavirales">Mulpavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Metaxyviridae&action=edit&redlink=1" class="new" title="Metaxyviridae (page does not exist)">Metaxyviridae</a></i></li> <li><i><a href="/wiki/Nanoviridae" title="Nanoviridae">Nanoviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Recrevirales" class="mw-redirect" title="Recrevirales">Recrevirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Redondoviridae" title="Redondoviridae">Redondoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Repensiviricetes" class="mw-redirect" title="Repensiviricetes">Repensiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Geplafuvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Geplafuvirales" title="Geplafuvirales">Geplafuvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Geminiviridae" title="Geminiviridae">Geminiviridae</a></i></li> <li><i><a href="/wiki/Genomoviridae" title="Genomoviridae">Genomoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Trapavirae" class="mw-redirect" title="Trapavirae">Trapavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Saleviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Saleviricota" class="mw-redirect" title="Saleviricota">Saleviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Huolimaviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Huolimaviricetes" class="mw-redirect" title="Huolimaviricetes">Huolimaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Haloruvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Haloruvirales" class="mw-redirect" title="Haloruvirales">Haloruvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Pleolipoviridae" title="Pleolipoviridae">Pleolipoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><b>Families</b>: <i><a href="/wiki/Anelloviridae" title="Anelloviridae">Anelloviridae</a></i></li> <li><i><a href="/wiki/Finnlakeviridae" title="Finnlakeviridae">Finnlakeviridae</a></i></li> <li><i><a href="/wiki/Spiraviridae" title="Spiraviridae">Spiraviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/RNA_virus" title="RNA virus">RNA</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="III:_dsRNA_viruses" style="font-size:114%;margin:0 4em">III: <a href="/wiki/Double-stranded_RNA_viruses" title="Double-stranded RNA viruses">dsRNA viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Riboviria" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Riboviria" title="Riboviria">Riboviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Orthornavirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Orthornavirae" title="Orthornavirae">Orthornavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Duplornaviricota" title="Duplornaviricota">Duplornaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Chrymotiviricetes" class="mw-redirect" title="Chrymotiviricetes">Chrymotiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Ghabrivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ghabrivirales" title="Ghabrivirales">Ghabrivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Chrysoviridae" title="Chrysoviridae">Chrysoviridae</a></i></li> <li><i><a href="/wiki/Megabirnaviridae" title="Megabirnaviridae">Megabirnaviridae</a></i></li> <li><i><a href="/wiki/Quadriviridae" title="Quadriviridae">Quadriviridae</a></i></li> <li><i><a href="/wiki/Totiviridae" title="Totiviridae">Totiviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Resentoviricetes" class="mw-redirect" title="Resentoviricetes">Resentoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Reovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Reovirales" class="mw-redirect" title="Reovirales">Reovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Reoviridae" class="mw-redirect" title="Reoviridae">Reoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Vidaverviricetes" class="mw-redirect" title="Vidaverviricetes">Vidaverviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Mindivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Mindivirales" class="mw-redirect" title="Mindivirales">Mindivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Cystoviridae" class="mw-redirect" title="Cystoviridae">Cystoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pisuviricota" title="Pisuviricota">Pisuviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Duplopiviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Duplopiviricetes" class="mw-redirect" title="Duplopiviricetes">Duplopiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Durnavirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Durnavirales" title="Durnavirales">Durnavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Amalgaviridae" title="Amalgaviridae">Amalgaviridae</a></i></li> <li><i><a href="/w/index.php?title=Curvulaviridae&action=edit&redlink=1" class="new" title="Curvulaviridae (page does not exist)">Curvulaviridae</a></i></li> <li><i><a href="/wiki/Hypoviridae" class="mw-redirect" title="Hypoviridae">Hypoviridae</a></i></li> <li><i><a href="/wiki/Picobirnaviridae" class="mw-redirect" title="Picobirnaviridae">Picobirnaviridae</a></i></li> <li><i><a href="/wiki/Partitiviridae" title="Partitiviridae">Partitiviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><b>Families</b>: <i><a href="/wiki/Birnaviridae" title="Birnaviridae">Birnaviridae</a></i></li> <li><i><a href="/w/index.php?title=Polymycoviridae&action=edit&redlink=1" class="new" title="Polymycoviridae (page does not exist)">Polymycoviridae</a></i></li> <li><b>Genera</b>: <i><a href="/w/index.php?title=Botybirnavirus&action=edit&redlink=1" class="new" title="Botybirnavirus (page does not exist)">Botybirnavirus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="IV:_(+)ssRNA_viruses" style="font-size:114%;margin:0 4em">IV: <a href="/wiki/Positive-sense_ssRNA_virus" class="mw-redirect" title="Positive-sense ssRNA virus">(+)ssRNA viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Riboviria" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Riboviria" title="Riboviria">Riboviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Orthornavirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Orthornavirae" title="Orthornavirae">Orthornavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Kitrinoviricota" title="Kitrinoviricota">Kitrinoviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Alsuviricetes" title="Alsuviricetes">Alsuviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Hepelivirales" title="Hepelivirales">Hepelivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alphatetraviridae" title="Alphatetraviridae">Alphatetraviridae</a></i></li> <li><i><a href="/wiki/Benyviridae" class="mw-redirect" title="Benyviridae">Benyviridae</a></i></li> <li><i><a href="/wiki/Hepeviridae" title="Hepeviridae">Hepeviridae</a></i></li> <li><i><a href="/wiki/Matonaviridae" class="mw-redirect" title="Matonaviridae">Matonaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Martellivirales" title="Martellivirales">Martellivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Bromoviridae" title="Bromoviridae">Bromoviridae</a></i></li> <li><i><a href="/wiki/Closteroviridae" title="Closteroviridae">Closteroviridae</a></i></li> <li><i><a href="/wiki/Endornaviridae" title="Endornaviridae">Endornaviridae</a></i></li> <li><i><a href="/w/index.php?title=Kitaviridae&action=edit&redlink=1" class="new" title="Kitaviridae (page does not exist)">Kitaviridae</a></i></li> <li><i><a href="/w/index.php?title=Mayoviridae&action=edit&redlink=1" class="new" title="Mayoviridae (page does not exist)">Mayoviridae</a></i></li> <li><i><a href="/wiki/Togaviridae" class="mw-redirect" title="Togaviridae">Togaviridae</a></i></li> <li><i><a href="/wiki/Virgaviridae" title="Virgaviridae">Virgaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tymovirales" title="Tymovirales">Tymovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alphaflexiviridae" title="Alphaflexiviridae">Alphaflexiviridae</a></i></li> <li><i><a href="/wiki/Betaflexiviridae" title="Betaflexiviridae">Betaflexiviridae</a></i></li> <li><i><a href="/w/index.php?title=Deltaflexiviridae&action=edit&redlink=1" class="new" title="Deltaflexiviridae (page does not exist)">Deltaflexiviridae</a></i></li> <li><i><a href="/wiki/Gammaflexiviridae" title="Gammaflexiviridae">Gammaflexiviridae</a></i></li> <li><i><a href="/wiki/Tymoviridae" title="Tymoviridae">Tymoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Flasuviricetes" class="mw-redirect" title="Flasuviricetes">Flasuviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Amarillovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Amarillovirales" class="mw-redirect" title="Amarillovirales">Amarillovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Flaviviridae" title="Flaviviridae">Flaviviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Magsaviricetes" class="mw-redirect" title="Magsaviricetes">Magsaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Nodamuvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Nodamuvirales" title="Nodamuvirales">Nodamuvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Nodaviridae" title="Nodaviridae">Nodaviridae</a></i></li> <li><i><a href="/w/index.php?title=Sinhaliviridae&action=edit&redlink=1" class="new" title="Sinhaliviridae (page does not exist)">Sinhaliviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tolucaviricetes" class="mw-redirect" title="Tolucaviricetes">Tolucaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Tolivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tolivirales" title="Tolivirales">Tolivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Carmotetraviridae" title="Carmotetraviridae">Carmotetraviridae</a></i></li> <li><i><a href="/wiki/Luteoviridae" title="Luteoviridae">Luteoviridae</a></i></li> <li><i><a href="/wiki/Tombusviridae" title="Tombusviridae">Tombusviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Lenarviricota" title="Lenarviricota">Lenarviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Leviviricetes" title="Leviviricetes">Leviviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Norzivirales" title="Norzivirales">Norzivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Atkinsviridae" title="Atkinsviridae">Atkinsviridae</a></i></li> <li><i><a href="/wiki/Duinviridae" title="Duinviridae">Duinviridae</a></i></li> <li><i><a href="/wiki/Fiersviridae" title="Fiersviridae">Fiersviridae</a></i></li> <li><i><a href="/wiki/Solspiviridae" title="Solspiviridae">Solspiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Timlovirales" title="Timlovirales">Timlovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Blumeviridae" title="Blumeviridae">Blumeviridae</a></i></li> <li><i><a href="/wiki/Steitzviridae" title="Steitzviridae">Steitzviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Amabiliviricetes" class="mw-redirect" title="Amabiliviricetes">Amabiliviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Wolframvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Wolframvirales" class="mw-redirect" title="Wolframvirales">Wolframvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Narnaviridae" class="mw-redirect" title="Narnaviridae">Narnaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Howeltoviricetes" class="mw-redirect" title="Howeltoviricetes">Howeltoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Cryppavirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cryppavirales" class="mw-redirect" title="Cryppavirales">Cryppavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Mitoviridae" class="mw-redirect" title="Mitoviridae">Mitoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Miaviricetes" class="mw-redirect" title="Miaviricetes">Miaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Ourlivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ourlivirales" class="mw-redirect" title="Ourlivirales">Ourlivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Botourmiaviridae" title="Botourmiaviridae">Botourmiaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pisuviricota" title="Pisuviricota">Pisuviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pisoniviricetes" title="Pisoniviricetes">Pisoniviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Nidovirales" title="Nidovirales">Nidovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Abyssoviridae" class="mw-redirect" title="Abyssoviridae">Abyssoviridae</a></i></li> <li><i><a href="/wiki/Arteriviridae" title="Arteriviridae">Arteriviridae</a></i></li> <li><i><a href="/w/index.php?title=Cremegaviridae&action=edit&redlink=1" class="new" title="Cremegaviridae (page does not exist)">Cremegaviridae</a></i></li> <li><i><a href="/wiki/Coronaviridae" title="Coronaviridae">Coronaviridae</a></i></li> <li><i><a href="/w/index.php?title=Euroniviridae&action=edit&redlink=1" class="new" title="Euroniviridae (page does not exist)">Euroniviridae</a></i></li> <li><i><a href="/w/index.php?title=Gresnaviridae&action=edit&redlink=1" class="new" title="Gresnaviridae (page does not exist)">Gresnaviridae</a></i></li> <li><i><a href="/w/index.php?title=Medioniviridae&action=edit&redlink=1" class="new" title="Medioniviridae (page does not exist)">Medioniviridae</a></i></li> <li><i><a href="/wiki/Mesoniviridae" title="Mesoniviridae">Mesoniviridae</a></i></li> <li><i><a href="/wiki/Mononiviridae" class="mw-redirect" title="Mononiviridae">Mononiviridae</a></i></li> <li><i><a href="/w/index.php?title=Nanghoshaviridae&action=edit&redlink=1" class="new" title="Nanghoshaviridae (page does not exist)">Nanghoshaviridae</a></i></li> <li><i><a href="/w/index.php?title=Nanhypoviridae&action=edit&redlink=1" class="new" title="Nanhypoviridae (page does not exist)">Nanhypoviridae</a></i></li> <li><i><a href="/w/index.php?title=Olifoviridae&action=edit&redlink=1" class="new" title="Olifoviridae (page does not exist)">Olifoviridae</a></i></li> <li><i><a href="/wiki/Roniviridae" class="mw-redirect" title="Roniviridae">Roniviridae</a></i></li> <li><i><a href="/wiki/Tobaniviridae" title="Tobaniviridae">Tobaniviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Picornavirales" title="Picornavirales">Picornavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Picornaviridae" class="mw-redirect" title="Picornaviridae">Picornaviridae</a></i></li> <li><i><a href="/wiki/Marnaviridae" title="Marnaviridae">Marnaviridae</a></i></li> <li><i><a href="/w/index.php?title=Solinviviridae&action=edit&redlink=1" class="new" title="Solinviviridae (page does not exist)">Solinviviridae</a></i></li> <li><i><a href="/wiki/Caliciviridae" title="Caliciviridae">Caliciviridae</a></i></li> <li><i><a href="/wiki/Iflaviridae" title="Iflaviridae">Iflaviridae</a></i></li> <li><i><a href="/wiki/Secoviridae" title="Secoviridae">Secoviridae</a></i></li> <li><i><a href="/wiki/Dicistroviridae" title="Dicistroviridae">Dicistroviridae</a></i></li> <li><i><a href="/w/index.php?title=Polycipiviridae&action=edit&redlink=1" class="new" title="Polycipiviridae (page does not exist)">Polycipiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Sobelivirales" title="Sobelivirales">Sobelivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alvernaviridae" title="Alvernaviridae">Alvernaviridae</a></i></li> <li><i><a href="/wiki/Barnaviridae" title="Barnaviridae">Barnaviridae</a></i></li> <li><i><a href="/wiki/Solemoviridae" title="Solemoviridae">Solemoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Stelpaviricetes" title="Stelpaviricetes">Stelpaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Patatavirales" class="mw-redirect" title="Patatavirales">Patatavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Potyviridae" title="Potyviridae">Potyviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Stellavirales" class="mw-redirect" title="Stellavirales">Stellavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Astroviridae" class="mw-redirect" title="Astroviridae">Astroviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><b>Families</b>: <i><a href="/wiki/Permutotetraviridae" title="Permutotetraviridae">Permutotetraviridae</a></i></li> <li><i><a href="/wiki/Sarthroviridae" class="mw-redirect" title="Sarthroviridae">Sarthroviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="V:_(–)ssRNA_viruses" style="font-size:114%;margin:0 4em">V: <a href="/wiki/Negative-sense_ssRNA_virus" class="mw-redirect" title="Negative-sense ssRNA virus">(–)ssRNA viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Riboviria" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Riboviria" title="Riboviria">Riboviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Orthornavirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Orthornavirae" title="Orthornavirae">Orthornavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Negarnaviricota" class="mw-redirect" title="Negarnaviricota">Negarnaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Chunqiuviricetes" class="mw-redirect" title="Chunqiuviricetes">Chunqiuviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Muvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Muvirales" class="mw-redirect" title="Muvirales">Muvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Qinviridae" class="mw-redirect" title="Qinviridae">Qinviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ellioviricetes" class="mw-redirect" title="Ellioviricetes">Ellioviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Bunyavirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Bunyavirales" title="Bunyavirales">Bunyavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Cruliviridae" title="Cruliviridae">Cruliviridae</a></i></li> <li><i><a href="/wiki/Arenaviridae" class="mw-redirect" title="Arenaviridae">Arenaviridae</a></i></li> <li><i><a href="/wiki/Fimoviridae" class="mw-redirect" title="Fimoviridae">Fimoviridae</a></i></li> <li><i><a href="/wiki/Hantaviridae" title="Hantaviridae">Hantaviridae</a></i></li> <li><i><a href="/w/index.php?title=Leishbuviridae&action=edit&redlink=1" class="new" title="Leishbuviridae (page does not exist)">Leishbuviridae</a></i></li> <li><i><a href="/w/index.php?title=Mypoviridae&action=edit&redlink=1" class="new" title="Mypoviridae (page does not exist)">Mypoviridae</a></i></li> <li><i><a href="/wiki/Nairoviridae" title="Nairoviridae">Nairoviridae</a></i></li> <li><i><a href="/wiki/Peribunyaviridae" title="Peribunyaviridae">Peribunyaviridae</a></i></li> <li><i><a href="/wiki/Phasmaviridae" title="Phasmaviridae">Phasmaviridae</a></i></li> <li><i><a href="/wiki/Phenuiviridae" title="Phenuiviridae">Phenuiviridae</a></i></li> <li><i><a href="/wiki/Tospoviridae" class="mw-redirect" title="Tospoviridae">Tospoviridae</a></i></li> <li><i><a href="/w/index.php?title=Wupedeviridae&action=edit&redlink=1" class="new" title="Wupedeviridae (page does not exist)">Wupedeviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Insthoviricetes" class="mw-redirect" title="Insthoviricetes">Insthoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Articulavirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Articulavirales" title="Articulavirales">Articulavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Amnoonviridae" class="mw-redirect" title="Amnoonviridae">Amnoonviridae</a></i></li> <li><i><a href="/wiki/Orthomyxoviridae" title="Orthomyxoviridae">Orthomyxoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Milneviricetes" class="mw-redirect" title="Milneviricetes">Milneviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Serpentovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Serpentovirales" class="mw-redirect" title="Serpentovirales">Serpentovirales </a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Aspiviridae" title="Aspiviridae">Aspiviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Monjiviricetes" title="Monjiviricetes">Monjiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Jingchuvirales" title="Jingchuvirales">Jingchuvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Aliusviridae&action=edit&redlink=1" class="new" title="Aliusviridae (page does not exist)">Aliusviridae</a></i></li> <li><i><a href="/wiki/Chuviridae" title="Chuviridae">Chuviridae</a></i></li> <li><i><a href="/w/index.php?title=Crepuscuviridae&action=edit&redlink=1" class="new" title="Crepuscuviridae (page does not exist)">Crepuscuviridae</a></i></li> <li><i><a href="/w/index.php?title=Myriaviridae&action=edit&redlink=1" class="new" title="Myriaviridae (page does not exist)">Myriaviridae</a></i></li> <li><i><a href="/w/index.php?title=Natareviridae&action=edit&redlink=1" class="new" title="Natareviridae (page does not exist)">Natareviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Mononegavirales" title="Mononegavirales">Mononegavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Artoviridae" title="Artoviridae">Artoviridae</a></i></li> <li><i><a href="/wiki/Bornaviridae" title="Bornaviridae">Bornaviridae</a></i></li> <li><i><a href="/wiki/Filoviridae" title="Filoviridae">Filoviridae</a></i></li> <li><i><a href="/w/index.php?title=Lispiviridae&action=edit&redlink=1" class="new" title="Lispiviridae (page does not exist)">Lispiviridae</a></i></li> <li><i><a href="/wiki/Mymonaviridae" title="Mymonaviridae">Mymonaviridae</a></i></li> <li><i><a href="/wiki/Nyamiviridae" title="Nyamiviridae">Nyamiviridae</a></i></li> <li><i><a href="/wiki/Paramyxoviridae" title="Paramyxoviridae">Paramyxoviridae</a></i></li> <li><i><a href="/wiki/Pneumoviridae" title="Pneumoviridae">Pneumoviridae</a></i></li> <li><i><a href="/wiki/Rhabdoviridae" title="Rhabdoviridae">Rhabdoviridae</a></i></li> <li><i><a href="/wiki/Sunviridae" title="Sunviridae">Sunviridae</a></i></li> <li><i><a href="/w/index.php?title=Xinmoviridae&action=edit&redlink=1" class="new" title="Xinmoviridae (page does not exist)">Xinmoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Yunchangviricetes" class="mw-redirect" title="Yunchangviricetes">Yunchangviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Goujianvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Goujianvirales" class="mw-redirect" title="Goujianvirales">Goujianvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Yueviridae" class="mw-redirect" title="Yueviridae">Yueviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ambiviricota" title="Ambiviricota">Ambiviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Suforviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Suforviricetes" class="mw-redirect" title="Suforviricetes">Suforviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Crytulvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Crytulvirales" class="mw-redirect" title="Crytulvirales">Crytulvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Dumbiviridae&action=edit&redlink=1" class="new" title="Dumbiviridae (page does not exist)">Dumbiviridae</a></i></li> <li><i><a href="/w/index.php?title=Quambiviridae&action=edit&redlink=1" class="new" title="Quambiviridae (page does not exist)">Quambiviridae</a></i></li> <li><i><a href="/w/index.php?title=Trimbiviridae&action=edit&redlink=1" class="new" title="Trimbiviridae (page does not exist)">Trimbiviridae</a></i></li> <li><i><a href="/w/index.php?title=Unambiviridae&action=edit&redlink=1" class="new" title="Unambiviridae (page does not exist)">Unambiviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Reverse_transcribing_virus" title="Reverse transcribing virus">RT</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="VI:_ssRNA-RT_viruses" style="font-size:114%;margin:0 4em">VI: <a href="/wiki/SsRNA-RT_virus" class="mw-redirect" title="SsRNA-RT virus">ssRNA-RT viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Riboviria" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Riboviria" title="Riboviria">Riboviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Pararnavirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pararnavirae" class="mw-redirect" title="Pararnavirae">Pararnavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Artverviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Artverviricota" class="mw-redirect" title="Artverviricota">Artverviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Revtraviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Revtraviricetes" title="Revtraviricetes">Revtraviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Ortervirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ortervirales" title="Ortervirales">Ortervirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Belpaoviridae" class="mw-redirect" title="Belpaoviridae">Belpaoviridae</a></i></li> <li><i><a href="/wiki/Metaviridae" title="Metaviridae">Metaviridae</a></i></li> <li><i><a href="/wiki/Pseudoviridae" title="Pseudoviridae">Pseudoviridae</a></i></li> <li><i><a href="/wiki/Retroviridae" class="mw-redirect" title="Retroviridae">Retroviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="VII:_dsDNA-RT_viruses" style="font-size:114%;margin:0 4em">VII: <a href="/wiki/DsDNA-RT_virus" class="mw-redirect" title="DsDNA-RT virus">dsDNA-RT viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Riboviria" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Riboviria" title="Riboviria">Riboviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Pararnavirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pararnavirae" class="mw-redirect" title="Pararnavirae">Pararnavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Artverviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Artverviricota" class="mw-redirect" title="Artverviricota">Artverviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Revtraviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Revtraviricetes" title="Revtraviricetes">Revtraviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Blubervirales" class="mw-redirect" title="Blubervirales">Blubervirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Hepadnaviridae" title="Hepadnaviridae">Hepadnaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ortervirales" title="Ortervirales">Ortervirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Caulimoviridae" title="Caulimoviridae">Caulimoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" 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style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Polydnaviriformidae" class="extiw" title="wikispecies:Polydnaviriformidae">Polydnaviriformidae</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Catalogue_of_Life" title="Catalogue of Life">CoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.catalogueoflife.org/data/taxon/627KB">627KB</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/8046">8046</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/1933">1933</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&id=106550">106550</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=10482">10482</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/World_Register_of_Marine_Species" title="World Register of Marine Species">WoRMS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.marinespecies.org/aphia.php?p=taxdetails&id=600029">600029</a></span></span></li></ul> </div></td></tr></tbody></table></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Retrieved from "<a dir="ltr" href="https://en.wikipedia.org/w/index.php?title=Polydnaviriformidae&oldid=1248163986">https://en.wikipedia.org/w/index.php?title=Polydnaviriformidae&oldid=1248163986</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Help:Category" title="Help:Category">Categories</a>: <ul><li><a href="/wiki/Category:Polydnaviridae" title="Category:Polydnaviridae">Polydnaviridae</a></li><li><a href="/wiki/Category:Symbiosis" title="Category:Symbiosis">Symbiosis</a></li></ul></div><div id="mw-hidden-catlinks" 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