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Phuwiangvenator - Wikipedia

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class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-See_also" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#See_also"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>See also</span> </div> </a> <ul id="toc-See_also-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc 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href="https://es.wikipedia.org/wiki/Phuwiangvenator_yaemniyomi" title="Phuwiangvenator yaemniyomi – Spanish" lang="es" hreflang="es" data-title="Phuwiangvenator yaemniyomi" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Phuwiangvenator" title="Phuwiangvenator – French" lang="fr" hreflang="fr" data-title="Phuwiangvenator" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Phuwiangvenator" title="Phuwiangvenator – Italian" lang="it" hreflang="it" data-title="Phuwiangvenator" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Phuwiangvenator" title="Phuwiangvenator – Dutch" lang="nl" hreflang="nl" data-title="Phuwiangvenator" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%83%97%E3%82%A6%E3%82%A3%E3%82%A2%E3%83%B3%E3%82%B0%E3%83%B4%E3%82%A7%E3%83%8A%E3%83%88%E3%83%AB" title="プウィアングヴェナトル – Japanese" lang="ja" hreflang="ja" data-title="プウィアングヴェナトル" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Phuwiangvenator" title="Phuwiangvenator – Polish" lang="pl" hreflang="pl" data-title="Phuwiangvenator" data-language-autonym="Polski" data-language-local-name="Polish" 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style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><i>Phuwiangvenator</i><br /><div style="font-size: 85%;">Temporal range: <a href="/wiki/Early_Cretaceous" title="Early Cretaceous">Early Cretaceous</a><sup id="cite_ref-newbones_1-0" class="reference"><a href="#cite_note-newbones-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup><br /> ~125 to 120 Ma - Late <span class="noprint"><span style="display:inline-block;"></span><span style="display:inline-block;">Barremian</span> <span style="display:inline-block;"></span><div id="Timeline-row" style="margin: 4px auto 0; clear:both; width:220px; padding:0px; height:18px; overflow:visible; white-space:nowrap; border:1px #666; border-style:solid none; position:relative; z-index:0; font-size:97%;"> <div style="position:absolute; height:100%; left:0px; width:207.23076923077px; padding-left:5px; text-align:left; background-color:rgb(254,217,106); background-image: linear-gradient(to right, rgba(255,255,255,1), rgba(254,217,106,1) 15%, rgba(254,217,106,1));"><a href="/wiki/Precambrian" title="Precambrian">PreꞒ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,160,86); left:37.636923076923px; width:18.073846153846px;"><a href="/wiki/Cambrian" title="Cambrian">Ꞓ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(0,146,112); left:55.710769230769px; width:14.08px;"><a href="/wiki/Ordovician" title="Ordovician">O</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(179,225,182); left:69.790769230769px; width:8.3261538461539px;"><a href="/wiki/Silurian" title="Silurian">S</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(203,140,55); left:78.116923076923px; width:20.409230769231px;"><a href="/wiki/Devonian" title="Devonian">D</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(103,165,153); left:98.526153846154px; width:20.307692307692px;"><a href="/wiki/Carboniferous" title="Carboniferous">C</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(240,64,40); left:118.83384615385px; width:15.907015384615px;"><a href="/wiki/Permian" title="Permian">P</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(129,43,146); left:134.74086153846px; width:17.092984615385px;"><a href="/wiki/Triassic" title="Triassic">T</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(52,178,201); left:151.83384615385px; width:19.089230769231px;"><a href="/wiki/Jurassic" title="Jurassic">J</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,198,78); left:170.92307692308px; width:26.738461538462px;"><a href="/wiki/Cretaceous" title="Cretaceous">K</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(253,154,82); left:197.66153846154px; width:14.543692307692px;"><a href="/wiki/Paleogene" title="Paleogene">Pg</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(255,230,25); left:212.20523076923px; width:6.9215384615385px;"><a href="/wiki/Neogene" title="Neogene">N</a></div> <div id="end-border" style="position:absolute; height:100%; background-color:#666; width:1px; left:219px"></div><div style="margin:0 auto; line-height:0; clear:both; width:220px; padding:0px; height:8px; overflow:visible; background-color:transparent; position:relative; top:-4px; z-index:100;"> <div style="position:absolute; left:177.43169230769px; font-size:50%"><div style="position:relative; left:-0.42em">&#8595;</div></div> </div> </div></span></div> </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Phuwiangvenator_Hands.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/Phuwiangvenator_Hands.png/220px-Phuwiangvenator_Hands.png" decoding="async" width="220" height="406" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/Phuwiangvenator_Hands.png/330px-Phuwiangvenator_Hands.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/Phuwiangvenator_Hands.png/440px-Phuwiangvenator_Hands.png 2x" data-file-width="730" data-file-height="1348" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%">The finger bones of the holotype of <i>Phuwiangvenator</i> </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Phuwiangvenator" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Chordate" title="Chordate">Chordata</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Dinosaur" title="Dinosaur">Dinosauria</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Saurischia" title="Saurischia">Saurischia</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Theropoda" title="Theropoda">Theropoda</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><span style="font-style:normal;font-weight:normal;">†</span><a href="/wiki/Megaraptora" title="Megaraptora">Megaraptora</a> </td></tr> <tr> <td>Genus: </td> <td><span style="font-style:normal;font-weight:normal;">†</span><a class="mw-selflink selflink"><i>Phuwiangvenator</i></a><br /><small>Samathi <i>et al.</i>, 2019</small> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Type_species" title="Type species">Type species</a> </th></tr> <tr> <td colspan="2" style="text-align: center"><abbr title="Extinct" aria-label="Extinct" style="border: none; text-decoration: none; cursor: inherit; font-weight: normal; font-style: normal;">†</abbr><i><b>Phuwiangvenator yaemniyomi</b></i><br /><div style="font-size: 85%;">Samathi <i>et al.</i>, 2019</div> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><i><b>Phuwiangvenator</b></i> (<span class="rt-commentedText nowrap"><span class="IPA nopopups noexcerpt" lang="en-fonipa"><a href="/wiki/Help:IPA/English" title="Help:IPA/English">/<span style="border-bottom:1px dotted"><span title="&#39;f&#39; in &#39;find&#39;">f</span><span title="/u/: &#39;u&#39; in &#39;influence&#39;">u</span><span title="/ˈ/: primary stress follows">ˈ</span><span title="&#39;w&#39; in &#39;wind&#39;">w</span><span title="/ɛ/: &#39;e&#39; in &#39;dress&#39;">ɛ</span><span title="/ŋ/: &#39;ng&#39; in &#39;sing&#39;">ŋ</span><span title="&#39;v&#39; in &#39;vie&#39;">v</span><span title="/ɪ/: &#39;i&#39; in &#39;kit&#39;">ɪ</span><span title="&#39;n&#39; in &#39;nigh&#39;">n</span><span title="/ɑː/: &#39;a&#39; in &#39;father&#39;">ɑː</span><span title="&#39;t&#39; in &#39;tie&#39;">t</span><span title="/ə/: &#39;a&#39; in &#39;about&#39;">ə</span><span title="&#39;r&#39; in &#39;rye&#39;">r</span></span>/</a></span></span>) is an extinct genus of <a href="/wiki/Megaraptora" title="Megaraptora">megaraptoran</a> <a href="/wiki/Theropoda" title="Theropoda">theropod</a> that lived during the <a href="/wiki/Early_Cretaceous" title="Early Cretaceous">Early Cretaceous</a> period in what is now <a href="/wiki/Thailand" title="Thailand">Thailand</a>. It contains only the type species, <i><b>P. yaemniyomi</b></i>. The <a href="/wiki/Genus" title="Genus">generic name</a> of <i>Phuwiangvenator</i> comes from the <a href="/wiki/Phu_Wiang" title="Phu Wiang">Phu Wiang</a> mountains, where the <a href="/wiki/Holotype" title="Holotype">holotype</a> was discovered, and the <a href="/wiki/Latin_(language)" class="mw-redirect" title="Latin (language)">Latin</a> word "<i>venator</i>" meaning hunter. The <a href="/wiki/Species" title="Species">specific name</a>, "<i>yaemniyomi</i>", is in honor of <a href="/wiki/Paleontology_in_Thailand" title="Paleontology in Thailand">Sudham Yaemniyom</a>, who was a historical <a href="/wiki/Paleontologist" class="mw-redirect" title="Paleontologist">paleontologist</a> from Thailand and the first person to discover fossils there.<sup id="cite_ref-description_2-0" class="reference"><a href="#cite_note-description-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Discovery">Discovery</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Phuwiangvenator&amp;action=edit&amp;section=1" title="Edit section: Discovery"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Phu_Wiang_Mountain_Locality.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/cb/Phu_Wiang_Mountain_Locality.png/220px-Phu_Wiang_Mountain_Locality.png" decoding="async" width="220" height="240" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/cb/Phu_Wiang_Mountain_Locality.png/330px-Phu_Wiang_Mountain_Locality.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/cb/Phu_Wiang_Mountain_Locality.png/440px-Phu_Wiang_Mountain_Locality.png 2x" data-file-width="1220" data-file-height="1332" /></a><figcaption>A map of Thailand with the location of the Phu Wiang locality in the inset</figcaption></figure> <p>The <a href="/wiki/Holotype" title="Holotype">holotype</a> specimen of <i>Phuwiangvenator</i>, designated <b> SM-PW9B</b>, is a partially-complete skeleton consisting of a <a href="/wiki/Dorsal_vertebra" class="mw-redirect" title="Dorsal vertebra">dorsal vertebra</a>, three <a href="/wiki/Sacral_vertebrae" class="mw-redirect" title="Sacral vertebrae">sacral vertebrae</a>, a right <a href="/wiki/Metacarpal" class="mw-redirect" title="Metacarpal">metacarpal</a>, right manual <a href="/wiki/Phalanges" class="mw-redirect" title="Phalanges">phalanges</a> and <a href="/wiki/Unguals" class="mw-redirect" title="Unguals">unguals</a>, both <a href="/wiki/Tibia" title="Tibia">tibiae</a>, the left <a href="/wiki/Astragalus_(bone)" class="mw-redirect" title="Astragalus (bone)">astragalocalcaneum</a>, several left <a href="/wiki/Metatarsal" class="mw-redirect" title="Metatarsal">metatarsals</a>, and several right pedal phalanges and unguals. It was discovered in 1993 by a team led by Preecha Sainongkham in the <a href="/wiki/Sao_Khua_Formation" title="Sao Khua Formation">Sao Khua Formation</a> of <a href="/wiki/Khon_Kaen_province" title="Khon Kaen province">Khon Kaen province</a> in <a href="/wiki/Thailand" title="Thailand">Thailand</a>. At the time of this discovery, Sainongkham was working at the <a href="/wiki/Phu_Wiang_Dinosaur_Museum" title="Phu Wiang Dinosaur Museum">Phu Wiang Dinosaur Museum</a>. The specimen was transported to the <a href="/wiki/Sirindhorn_Museum" title="Sirindhorn Museum">Sirindhorn Museum</a> in the town of Non Buri, which is near the locality where the fossils were discovered. The fossils were prepared and remained in the museum's collection until <a href="/wiki/2019_in_archosaur_paleontology" title="2019 in archosaur paleontology">2019</a> when Adun Samathi, Phornphen Chanthasit, and P. Martin Sander published a formal description, which was accompanied by a description of the closely related taxon, <i><a href="/wiki/Vayuraptor" title="Vayuraptor">Vayuraptor</a></i>.<sup id="cite_ref-description_2-1" class="reference"><a href="#cite_note-description-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> </p><p>A few years after the description of the holotype, another specimen was discovered in the collections at the Sirindhorn Museum. This was described in 2022 by Adun Samathi (one of the original describers), Suravech Suteethorn, Nakorn Pradit, and Varavudh Suteethorn, the latter of whom was part of the original team which uncovered the holotype. They concluded, based on the reported type locality from which the specimen was retrieved as well as several characteristics of the bones themselves, that these bones likely belonged to the same individual animal as the holotype specimen. These additional elements were named as the topotype and were given the designation <b>SM-PW9A</b>. This specimen included both ends of the left <a href="/wiki/Fibula" title="Fibula">fibula</a>, several of the left metatarsals, and the distal ends of the right metatarsals.<sup id="cite_ref-newbones_1-1" class="reference"><a href="#cite_note-newbones-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> One of the metatarsals was later discovered to contain a fracture, which was studied and published on in 2023.<sup id="cite_ref-pathology_3-0" class="reference"><a href="#cite_note-pathology-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> </p><p>The discovery of <i>Phuwiangvenator</i> in Thailand is used by its describers to suggest an Asian origin of megaraptora. They state that the existence of <i>Vayuraptor</i> in Thailand and <i><a href="/wiki/Fukuiraptor" title="Fukuiraptor">Fukuiraptor</a></i> in <a href="/wiki/Japan" title="Japan">Japan</a> are also evidence of this.<sup id="cite_ref-description_2-2" class="reference"><a href="#cite_note-description-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Description">Description</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Phuwiangvenator&amp;action=edit&amp;section=2" title="Edit section: Description"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The authors of its original description estimate that <i>Phuwiangvenator</i> was probably about 6 metres (20&#160;ft) long in life. From what is known of the skeleton, it likely had proportions similar to the related taxon <i><a href="/wiki/Australovenator" title="Australovenator">Australovenator</a></i>, which is known from slightly more complete remains. The authors name two <a href="/wiki/Autapomorphies" class="mw-redirect" title="Autapomorphies">autapomorphies</a> for <i>Phuwiangvenator</i>. The first of these are short furrows on the ventral side of the <a href="/wiki/Sacral_vertebrae" class="mw-redirect" title="Sacral vertebrae">sacral vertebrae</a> on the front and rear ends. The other autapomorphy is a sloping rim of the fourth <a href="/wiki/Metatarsal" class="mw-redirect" title="Metatarsal">metatarsal</a> which is angled away from the middle of the body at the dorsal end. This sloping fourth metatarsal is additionally unique in having the lowest corner relative to the third metatarsal of any known <a href="/wiki/Theropod" class="mw-redirect" title="Theropod">theropod</a>.<sup id="cite_ref-description_2-3" class="reference"><a href="#cite_note-description-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> </p> <dl><dt>Vertebrae</dt></dl> <p><i>Phuwiangvenator</i> is only known from four vertebrae, one from the <a href="/wiki/Dorsal_vertebra" class="mw-redirect" title="Dorsal vertebra">dorsal</a> area and three from the <a href="/wiki/Sacrum" title="Sacrum">sacrum</a>. These vertebrae, as a whole, share numerous affinities with other large <a href="/wiki/Theropods" class="mw-redirect" title="Theropods">theropods</a>. The dorsal vertebra is hourglass-shaped when viewed from above (or below), which is similar to the vertebrae of <i><a href="/wiki/Fukuiraptor" title="Fukuiraptor">Fukuiraptor</a></i>, <i><a href="/wiki/Aerosteon" title="Aerosteon">Aerosteon</a></i>, and numerous other <a href="/wiki/Avetheropods" class="mw-redirect" title="Avetheropods">avetheropods</a>. In particular, the <a href="/wiki/Vertebral_centrum" class="mw-redirect" title="Vertebral centrum">centrum</a> of the only preserved dorsal vertebra does not have a <a href="/wiki/Pleurocoel" class="mw-redirect" title="Pleurocoel">pleurocoel</a> or any lateral ridges, which resembles the condition observed in <i><a href="/wiki/Gualicho" title="Gualicho">Gualicho</a></i>, a theropod of controversial affinities.<sup id="cite_ref-description_2-4" class="reference"><a href="#cite_note-description-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-maip_4-0" class="reference"><a href="#cite_note-maip-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> The three sacral vertebrae which are preserved are all fused and were damaged significantly during <a href="/wiki/Fossilization" class="mw-redirect" title="Fossilization">fossilization</a>. However, the centra are well-preserved enough to show one of the key autapomorphies of the genus, the saci (or furrows) on the anterior and posterior portion of the bone.<sup id="cite_ref-description_2-5" class="reference"><a href="#cite_note-description-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> </p> <dl><dt>Hands</dt></dl> <p>Most of the preserved finger bones have deep and well-defined attachment sites for <a href="/wiki/Ligaments" class="mw-redirect" title="Ligaments">ligaments</a>, which resemble those found in the theropods <i><a href="/wiki/Kileskus" title="Kileskus">Kileskus</a></i> and <i><a href="/wiki/Australovenator" title="Australovenator">Australovenator</a></i>.<sup id="cite_ref-description_2-6" class="reference"><a href="#cite_note-description-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> </p> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Phuwiangvenator_Feet.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Phuwiangvenator_Feet.png/220px-Phuwiangvenator_Feet.png" decoding="async" width="220" height="105" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Phuwiangvenator_Feet.png/330px-Phuwiangvenator_Feet.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Phuwiangvenator_Feet.png/440px-Phuwiangvenator_Feet.png 2x" data-file-width="2162" data-file-height="1032" /></a><figcaption>The preserved toe bones and claws of <i>Phuwiangvenator</i></figcaption></figure> <dl><dt>Legs</dt></dl> <p>The leg bones share numerous marked similarities with other <a href="/wiki/Megaraptorans" class="mw-redirect" title="Megaraptorans">megaraptorans</a>. The tibia is more than twelve times longer than its mid-length anteroposterior width, which is a <a href="/wiki/Synapomorphy" class="mw-redirect" title="Synapomorphy">synapomorphy</a> of megaraptorans and other <a href="/wiki/Coelurosaurs" class="mw-redirect" title="Coelurosaurs">coelurosaurs</a>. There are also several characteristics of the <a href="/wiki/Astragalus_(bone)" class="mw-redirect" title="Astragalus (bone)">astragalus</a> and the <a href="/wiki/Metatarsals" class="mw-redirect" title="Metatarsals">metatarsals</a> which resemble those of <i><a href="/wiki/Australovenator" title="Australovenator">Australovenator</a></i>. There is one major exception to the megaraptoran affinities of the holotype's leg bones; the anterolateral process of the lateral condyle is sharply curved downward, which is a condition seen in <i><a href="/wiki/Neovenator" title="Neovenator">Neovenator</a></i> and numerous <a href="/wiki/Tyrannosaurids" class="mw-redirect" title="Tyrannosaurids">tyrannosaurids</a>. This led the authors of the paper to suggest that this trait, and other characteristics of the leg bones may be more widespread among theropods than was previously thought.<sup id="cite_ref-description_2-7" class="reference"><a href="#cite_note-description-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> In their subsequent analysis of the topotype material, Samathi and colleagues draw comparisons between the <a href="/wiki/Metatarsal" class="mw-redirect" title="Metatarsal">metatarsals</a> of <i>Phuwiangvenator</i> with those of numerous <a href="/wiki/Theropods" class="mw-redirect" title="Theropods">theropods</a> including <i><a href="/wiki/Allosaurus" title="Allosaurus">Allosaurus</a></i>, <i><a href="/wiki/Concavenator" title="Concavenator">Concavenator</a></i>, <i><a href="/wiki/Neovenator" title="Neovenator">Neovenator</a></i>, and <i><a href="/wiki/Suchomimus" title="Suchomimus">Suchomimus</a></i>. They use this to suggest that the condition seen in the species represents the ancestral condition of all <a href="/wiki/Avetheropods" class="mw-redirect" title="Avetheropods">avetheropods</a>, rather than the derived condition seen in <a href="/wiki/Megaraptoridae" class="mw-redirect" title="Megaraptoridae">megaraptorids</a>.<sup id="cite_ref-newbones_1-2" class="reference"><a href="#cite_note-newbones-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Classification">Classification</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Phuwiangvenator&amp;action=edit&amp;section=3" title="Edit section: Classification"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Phylogeny">Phylogeny</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Phuwiangvenator&amp;action=edit&amp;section=4" title="Edit section: Phylogeny"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Samathi and colleagues performed a <a href="/wiki/Phylogenetic" class="mw-redirect" title="Phylogenetic">phylogenetic</a> analysis in their description of <i>Phuwiangvenator</i> and <i><a href="/wiki/Vayuraptor" title="Vayuraptor">Vayuraptor</a></i> based on the data set used by Apesteguía and colleagues in their 2016 description of <i><a href="/wiki/Gualicho" title="Gualicho">Gualicho</a></i>.<sup id="cite_ref-description_2-8" class="reference"><a href="#cite_note-description-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Gualicho_5-0" class="reference"><a href="#cite_note-Gualicho-5"><span class="cite-bracket">&#91;</span>5<span class="cite-bracket">&#93;</span></a></sup> In their analysis, they added the two new taxa from their description as well as the contemporaneous theropod <i><a href="/wiki/Siamotyrannus" title="Siamotyrannus">Siamotyrannus</a></i> and removed the problematic <i><a href="/wiki/Santanaraptor" title="Santanaraptor">Santanaraptor</a></i> from their data set. They recovered <i>Phuwiangvenator</i> as a basal member of <a href="/wiki/Megaraptora" title="Megaraptora">megaraptora</a> with the following synapomorphies for the group: a ventral groove on the first manual phalanx, a transversely compressed first manual ungual, a prominence on the anterior surface of the distal end of the <a href="/wiki/Tibia" title="Tibia">tibia</a>, a lateral extension of the astragalus, and a flattened <a href="/wiki/Sacrum" title="Sacrum">sacrum</a>. Notably, they recovered <i>Vayuraptor</i> as a non-megaraptoran <a href="/wiki/Coelurosaur" class="mw-redirect" title="Coelurosaur">coelurosaur</a>, which has not been recovered by subsequent analyses.<sup id="cite_ref-description_2-9" class="reference"><a href="#cite_note-description-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-maip_4-1" class="reference"><a href="#cite_note-maip-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> The strict consensus tree from their analysis is shown below.<sup id="cite_ref-description_2-10" class="reference"><a href="#cite_note-description-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="clade"><style data-mw-deduplicate="TemplateStyles:r1258728058">body.skin-vector-2022 .mw-parser-output div.clade,body.skin-minerva .mw-parser-output div.clade{overflow-x:auto;overflow-y:hidden}body.skin-minerva .mw-parser-output div.clade p{font-size:inherit}.mw-parser-output table.clade{border-spacing:0;margin:0;font-size:100%;line-height:100%;border-collapse:separate;width:auto}.mw-parser-output table.clade table.clade{width:100%;line-height:inherit}.mw-parser-output table.clade td.clade-label{min-width:0.2em;width:0.2em;padding:0.1em 0.25em;vertical-align:bottom;text-align:center;border-left:1px solid;border-bottom:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-label::before,.mw-parser-output table.clade td.clade-slabel::before{content:"\2060 "}.mw-parser-output table.clade td.clade-fixed-width{overflow:hidden;text-overflow:ellipsis}.mw-parser-output table.clade td.clade-fixed-width:hover{overflow:visible}.mw-parser-output table.clade td.clade-label.first{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-label.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-slabel{padding:0.1em 0.25em;vertical-align:top;text-align:center;border-left:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-slabel:hover{overflow:visible}.mw-parser-output table.clade td.clade-slabel.last{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-slabel.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-bar{vertical-align:middle;text-align:left;padding:0 0.5em;position:relative}.mw-parser-output table.clade td.clade-bar.reverse{text-align:right;position:relative}.mw-parser-output table.clade td.clade-leaf{border:0;padding:0;text-align:left}.mw-parser-output table.clade td.clade-leafR{border:0;padding:0;text-align:right}.mw-parser-output table.clade td.clade-leaf.reverse{text-align:right}.mw-parser-output table.clade:hover span.linkA{background-color:yellow}.mw-parser-output table.clade:hover span.linkB{background-color:green}</style> <table class="clade"> <tbody><tr> <td class="clade-label first"><b><a href="/wiki/Coelurosauria" title="Coelurosauria">Coelurosauria</a></b> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Siamotyrannus" title="Siamotyrannus">Siamotyrannus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Chilantaisaurus" title="Chilantaisaurus">Chilantaisaurus</a></i> <span typeof="mw:File"><a href="/wiki/File:Chilantaisaurus.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/ac/Chilantaisaurus.jpg/50px-Chilantaisaurus.jpg" decoding="async" width="50" height="18" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/ac/Chilantaisaurus.jpg/75px-Chilantaisaurus.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/ac/Chilantaisaurus.jpg/100px-Chilantaisaurus.jpg 2x" data-file-width="4197" data-file-height="1551" /></a></span> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <i><a href="/wiki/Gualicho" title="Gualicho">Gualicho</a></i> <div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Gualicho_NT_small.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/e/ed/Gualicho_NT_small.jpg/50px-Gualicho_NT_small.jpg" decoding="async" width="50" height="24" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/ed/Gualicho_NT_small.jpg/75px-Gualicho_NT_small.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/ed/Gualicho_NT_small.jpg/100px-Gualicho_NT_small.jpg 2x" data-file-width="583" data-file-height="278" /></a></span></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Vayuraptor" title="Vayuraptor">Vayuraptor</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Tyrannoraptora" class="mw-redirect" title="Tyrannoraptora">Tyrannoraptora</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Tanycolagreus" title="Tanycolagreus">Tanycolagreus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Compsognathidae" title="Compsognathidae">Compsognathidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Maniraptoriformes" class="mw-redirect" title="Maniraptoriformes">Maniraptoriformes</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Tyrannosauroidea" title="Tyrannosauroidea">Tyrannosauroidea</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Megaraptora" title="Megaraptora">Megaraptora</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><b>Phuwiangvenator</b></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Fukuiraptor" title="Fukuiraptor">Fukuiraptor</a></i> <span typeof="mw:File"><a href="/wiki/File:Fukuiraptor_BW.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Fukuiraptor_BW.jpg/50px-Fukuiraptor_BW.jpg" decoding="async" width="50" height="21" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Fukuiraptor_BW.jpg/75px-Fukuiraptor_BW.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Fukuiraptor_BW.jpg/100px-Fukuiraptor_BW.jpg 2x" data-file-width="789" data-file-height="325" /></a></span> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Megaraptoridae" class="mw-redirect" title="Megaraptoridae">Megaraptoridae</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Megaraptor" title="Megaraptor">Megaraptor</a></i> <span typeof="mw:File"><a href="/wiki/File:MegaraptorNV.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/MegaraptorNV.jpg/50px-MegaraptorNV.jpg" decoding="async" width="50" height="22" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/MegaraptorNV.jpg/75px-MegaraptorNV.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4a/MegaraptorNV.jpg/100px-MegaraptorNV.jpg 2x" data-file-width="800" data-file-height="348" /></a></span> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <i><a href="/wiki/Orkoraptor" title="Orkoraptor">Orkoraptor</a></i> <div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Orkoraptor_drawing.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/79/Orkoraptor_drawing.jpg/50px-Orkoraptor_drawing.jpg" decoding="async" width="50" height="19" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/79/Orkoraptor_drawing.jpg/75px-Orkoraptor_drawing.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/79/Orkoraptor_drawing.jpg/100px-Orkoraptor_drawing.jpg 2x" data-file-width="3492" data-file-height="1347" /></a></span></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Aerosteon" title="Aerosteon">Aerosteon</a></i> <span typeof="mw:File"><a href="/wiki/File:Aerosteon_BW.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/89/Aerosteon_BW.jpg/50px-Aerosteon_BW.jpg" decoding="async" width="50" height="25" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/89/Aerosteon_BW.jpg/75px-Aerosteon_BW.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/89/Aerosteon_BW.jpg/100px-Aerosteon_BW.jpg 2x" data-file-width="295" data-file-height="147" /></a></span> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Australovenator" title="Australovenator">Australovenator</a></i> <span typeof="mw:File"><a href="/wiki/File:Australovenator_reconstruction.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/44/Australovenator_reconstruction.jpg/50px-Australovenator_reconstruction.jpg" decoding="async" width="50" height="32" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/44/Australovenator_reconstruction.jpg/75px-Australovenator_reconstruction.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/44/Australovenator_reconstruction.jpg/100px-Australovenator_reconstruction.jpg 2x" data-file-width="2017" data-file-height="1311" /></a></span> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> <p>A more recent phylogenetic analysis to include <i>Phuwiangvenator</i> was the one which accompanied the description of <i><a href="/wiki/Maip" title="Maip">Maip</a></i> in <a href="/wiki/2022_in_archosaur_paleontology" title="2022 in archosaur paleontology">2022</a>. The analysis of the authors, led by Alexis Aranciaga Rolando, included every taxon confidently assigned to <a href="/wiki/Megaraptora" title="Megaraptora">megaraptora</a> including an unnamed specimen from the <a href="/wiki/Griman_Creek_Formation" title="Griman Creek Formation">Griman Creek Formation</a> (called the "Lightning Claw" by some publications) and another unnamed specimen from the <a href="/wiki/Bajo_Barreal_Formation" title="Bajo Barreal Formation">Bajo Barreal Formation</a>. Their analysis of Megaraptora differs from the initial only in the exact classification of <i><a href="/wiki/Vayuraptor" title="Vayuraptor">Vayuraptor</a></i> and the inclusion of more taxa, which the authors state led to a more resolved tree consistent with previous analyses.<sup id="cite_ref-maip_4-2" class="reference"><a href="#cite_note-maip-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="clade"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"><b><a href="/wiki/Coelurosauria" title="Coelurosauria">Coelurosauria</a></b> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Maniraptoriformes" class="mw-redirect" title="Maniraptoriformes">Maniraptoriformes</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Compsognathidae" title="Compsognathidae">Compsognathidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Tyrannosauroidea" title="Tyrannosauroidea">Tyrannosauroidea</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Megaraptora" title="Megaraptora">Megaraptora</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><b>Phuwiangvenator</b></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Vayuraptor" title="Vayuraptor">Vayuraptor</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Fukuiraptor" title="Fukuiraptor">Fukuiraptor</a></i> <span typeof="mw:File"><a href="/wiki/File:Fukuiraptor_BW.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Fukuiraptor_BW.jpg/50px-Fukuiraptor_BW.jpg" decoding="async" width="50" height="21" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Fukuiraptor_BW.jpg/75px-Fukuiraptor_BW.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Fukuiraptor_BW.jpg/100px-Fukuiraptor_BW.jpg 2x" data-file-width="789" data-file-height="325" /></a></span> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Megaraptoridae" class="mw-redirect" title="Megaraptoridae">Megaraptoridae</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Griman_Creek_Formation" title="Griman Creek Formation">Lightning Ridge</a> megaraptoran </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Australovenator" title="Australovenator">Australovenator</a></i> <span typeof="mw:File"><a href="/wiki/File:Australovenator_reconstruction.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/44/Australovenator_reconstruction.jpg/50px-Australovenator_reconstruction.jpg" decoding="async" width="50" height="32" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/44/Australovenator_reconstruction.jpg/75px-Australovenator_reconstruction.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/44/Australovenator_reconstruction.jpg/100px-Australovenator_reconstruction.jpg 2x" data-file-width="2017" data-file-height="1311" /></a></span> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label">"Clade&#160;A" </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Aoniraptor" title="Aoniraptor">Aoniraptor</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Bajo_Barreal_Formation" title="Bajo Barreal Formation">Bajo Barreal</a> megaraptoran </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Megaraptor" title="Megaraptor">Megaraptor</a></i> <span typeof="mw:File"><a href="/wiki/File:MegaraptorNV.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/MegaraptorNV.jpg/50px-MegaraptorNV.jpg" decoding="async" width="50" height="22" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/MegaraptorNV.jpg/75px-MegaraptorNV.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4a/MegaraptorNV.jpg/100px-MegaraptorNV.jpg 2x" data-file-width="800" data-file-height="348" /></a></span> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Murusraptor" title="Murusraptor">Murusraptor</a></i> <span typeof="mw:File"><a href="/wiki/File:Murusraptor_NT_small.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c4/Murusraptor_NT_small.jpg/50px-Murusraptor_NT_small.jpg" decoding="async" width="50" height="26" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c4/Murusraptor_NT_small.jpg/75px-Murusraptor_NT_small.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c4/Murusraptor_NT_small.jpg/100px-Murusraptor_NT_small.jpg 2x" data-file-width="907" data-file-height="474" /></a></span> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label">"Clade&#160;B" </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Aerosteon" title="Aerosteon">Aerosteon</a></i> <span typeof="mw:File"><a href="/wiki/File:Aerosteon_BW.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/89/Aerosteon_BW.jpg/50px-Aerosteon_BW.jpg" decoding="async" width="50" height="25" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/89/Aerosteon_BW.jpg/75px-Aerosteon_BW.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/89/Aerosteon_BW.jpg/100px-Aerosteon_BW.jpg 2x" data-file-width="295" data-file-height="147" /></a></span> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Tratayenia" title="Tratayenia">Tratayenia</a></i> <span typeof="mw:File"><a href="/wiki/File:Tratayenia.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/45/Tratayenia.jpg/50px-Tratayenia.jpg" decoding="async" width="50" height="29" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/45/Tratayenia.jpg/75px-Tratayenia.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/45/Tratayenia.jpg/100px-Tratayenia.jpg 2x" data-file-width="1016" data-file-height="584" /></a></span> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <i><a href="/wiki/Orkoraptor" title="Orkoraptor">Orkoraptor</a></i> <div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Orkoraptor_drawing.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/79/Orkoraptor_drawing.jpg/50px-Orkoraptor_drawing.jpg" decoding="async" width="50" height="19" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/79/Orkoraptor_drawing.jpg/75px-Orkoraptor_drawing.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/79/Orkoraptor_drawing.jpg/100px-Orkoraptor_drawing.jpg 2x" data-file-width="3492" data-file-height="1347" /></a></span></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Maip" title="Maip">Maip</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> <p>In 2024, Andrea Cau published a comprehensive analysis of theropod phylogeny using a novel analytical method to test the effects of <a href="/wiki/Ontogeny" title="Ontogeny">ontogeny</a> on the phylogenetic position of various taxa.<sup id="cite_ref-Cau2024_6-0" class="reference"><a href="#cite_note-Cau2024-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup> This was based on an earlier publication by Cau in which he hypothesized that the enigmatic European taxa <i><a href="/wiki/Juratyrant" title="Juratyrant">Juratyrant</a></i>, <i><a href="/wiki/Scipionyx" title="Scipionyx">Scipionyx</a></i>, and <i><a href="/wiki/Sciurumimus" title="Sciurumimus">Sciurumimus</a></i> (which are known to be juvenile specimens), may not be juvenile <a href="/wiki/Compsognathid" class="mw-redirect" title="Compsognathid">compsognathids</a>, but rather juvenile <a href="/wiki/Megalosauroid" class="mw-redirect" title="Megalosauroid">megalosauroids</a> or <a href="/wiki/Allosauroid" class="mw-redirect" title="Allosauroid">allosauroids</a>.<sup id="cite_ref-Cau2021_7-0" class="reference"><a href="#cite_note-Cau2021-7"><span class="cite-bracket">&#91;</span>7<span class="cite-bracket">&#93;</span></a></sup> The 2024 analysis recovered a novel result, which placed <i>Phuwiangvenator</i> and the closely-related <i><a href="/wiki/Vayuraptor" title="Vayuraptor">Vayuraptor</a></i> within the family <a href="/wiki/Coeluridae" title="Coeluridae">Coeluridae</a>, which had previously been considered a <a href="/wiki/Wastebasket_taxon" title="Wastebasket taxon">wastebasket taxon</a>. Cau also recovered a variety of disparate taxa to be a member of this clade including <i><a href="/wiki/Shishugounykus" title="Shishugounykus">Shishugounykus</a></i>, <i><a href="/wiki/Fukuivenator" title="Fukuivenator">Fukuivenator</a></i>, and <i><a href="/wiki/Migmanychion" title="Migmanychion">Migmanychion</a></i>, which had previously been recovered as <a href="/wiki/Maniraptora" title="Maniraptora">maniraptorans</a>. An abbreviated form of the relevant section of Cau's analysis is shown below.<sup id="cite_ref-Cau2024_6-1" class="reference"><a href="#cite_note-Cau2024-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="clade"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Tyrannoraptora" class="mw-redirect" title="Tyrannoraptora">Tyrannoraptora</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <a href="/wiki/Tyrannosauroidea" title="Tyrannosauroidea">Tyrannosauroidea</a> <div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Tyrannosaurus-rex-Profile-steveoc86_(coloured).png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c7/Tyrannosaurus-rex-Profile-steveoc86_%28coloured%29.png/70px-Tyrannosaurus-rex-Profile-steveoc86_%28coloured%29.png" decoding="async" width="70" height="21" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c7/Tyrannosaurus-rex-Profile-steveoc86_%28coloured%29.png/105px-Tyrannosaurus-rex-Profile-steveoc86_%28coloured%29.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c7/Tyrannosaurus-rex-Profile-steveoc86_%28coloured%29.png/140px-Tyrannosaurus-rex-Profile-steveoc86_%28coloured%29.png 2x" data-file-width="990" data-file-height="303" /></a></span></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Maniraptoromorpha" class="mw-redirect" title="Maniraptoromorpha">Maniraptoromorpha</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Ornitholestes" title="Ornitholestes">Ornitholestes</a></i> <span typeof="mw:File"><a href="/wiki/File:Ornitholestes_reconstruction.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/a9/Ornitholestes_reconstruction.png/60px-Ornitholestes_reconstruction.png" decoding="async" width="60" height="25" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/a9/Ornitholestes_reconstruction.png/90px-Ornitholestes_reconstruction.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/a9/Ornitholestes_reconstruction.png/120px-Ornitholestes_reconstruction.png 2x" data-file-width="4876" data-file-height="2060" /></a></span> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <a href="/wiki/Maniraptoriformes" class="mw-redirect" title="Maniraptoriformes">Maniraptoriformes</a> <div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Meyers_grosses_Konversations-Lexikon_-_ein_Nachschlagewerk_des_allgemeinen_Wissens_(1908)_(Antwerpener_Breiftaube).jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/41/Meyers_grosses_Konversations-Lexikon_-_ein_Nachschlagewerk_des_allgemeinen_Wissens_%281908%29_%28Antwerpener_Breiftaube%29.jpg/20px-Meyers_grosses_Konversations-Lexikon_-_ein_Nachschlagewerk_des_allgemeinen_Wissens_%281908%29_%28Antwerpener_Breiftaube%29.jpg" decoding="async" width="20" height="22" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/41/Meyers_grosses_Konversations-Lexikon_-_ein_Nachschlagewerk_des_allgemeinen_Wissens_%281908%29_%28Antwerpener_Breiftaube%29.jpg/30px-Meyers_grosses_Konversations-Lexikon_-_ein_Nachschlagewerk_des_allgemeinen_Wissens_%281908%29_%28Antwerpener_Breiftaube%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/41/Meyers_grosses_Konversations-Lexikon_-_ein_Nachschlagewerk_des_allgemeinen_Wissens_%281908%29_%28Antwerpener_Breiftaube%29.jpg/40px-Meyers_grosses_Konversations-Lexikon_-_ein_Nachschlagewerk_des_allgemeinen_Wissens_%281908%29_%28Antwerpener_Breiftaube%29.jpg 2x" data-file-width="638" data-file-height="698" /></a></span></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Coeluridae" title="Coeluridae">Coeluridae</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <i><a href="/wiki/Coelurus" title="Coelurus">Coelurus</a></i> <div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Coelurus_BW.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/5/50/Coelurus_BW.jpg/60px-Coelurus_BW.jpg" decoding="async" width="60" height="26" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/50/Coelurus_BW.jpg/90px-Coelurus_BW.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/50/Coelurus_BW.jpg/120px-Coelurus_BW.jpg 2x" data-file-width="800" data-file-height="347" /></a></span></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><b>Phuwiangvenator</b></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Shishugounykus" title="Shishugounykus">Shishugounykus</a></i> <span typeof="mw:File"><a href="/wiki/File:Shishugounykus_inexpectus_skeletal_reconstruction.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Shishugounykus_inexpectus_skeletal_reconstruction.png/60px-Shishugounykus_inexpectus_skeletal_reconstruction.png" decoding="async" width="60" height="24" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Shishugounykus_inexpectus_skeletal_reconstruction.png/90px-Shishugounykus_inexpectus_skeletal_reconstruction.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/2d/Shishugounykus_inexpectus_skeletal_reconstruction.png/120px-Shishugounykus_inexpectus_skeletal_reconstruction.png 2x" data-file-width="1566" data-file-height="630" /></a></span> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Vayuraptor" title="Vayuraptor">Vayuraptor</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Fukuivenator" title="Fukuivenator">Fukuivenator</a></i> <span typeof="mw:File"><a href="/wiki/File:Fukuivenator_(Therizinosauria).png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Fukuivenator_%28Therizinosauria%29.png/70px-Fukuivenator_%28Therizinosauria%29.png" decoding="async" width="70" height="32" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Fukuivenator_%28Therizinosauria%29.png/105px-Fukuivenator_%28Therizinosauria%29.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/ff/Fukuivenator_%28Therizinosauria%29.png/140px-Fukuivenator_%28Therizinosauria%29.png 2x" data-file-width="3180" data-file-height="1440" /></a></span> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <i><a href="/wiki/Migmanychion" title="Migmanychion">Migmanychion</a></i> <div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Migmanychion_UDL.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Migmanychion_UDL.png/70px-Migmanychion_UDL.png" decoding="async" width="70" height="25" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Migmanychion_UDL.png/105px-Migmanychion_UDL.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Migmanychion_UDL.png/140px-Migmanychion_UDL.png 2x" data-file-width="2800" data-file-height="1000" /></a></span></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> <div class="mw-heading mw-heading3"><h3 id="Possible_synonymity">Possible synonymity</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Phuwiangvenator&amp;action=edit&amp;section=5" title="Edit section: Possible synonymity"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Siamotyrannus_pelvis_01.JPG" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/9/9f/Siamotyrannus_pelvis_01.JPG/220px-Siamotyrannus_pelvis_01.JPG" decoding="async" width="220" height="161" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/9f/Siamotyrannus_pelvis_01.JPG/330px-Siamotyrannus_pelvis_01.JPG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/9f/Siamotyrannus_pelvis_01.JPG/440px-Siamotyrannus_pelvis_01.JPG 2x" data-file-width="1196" data-file-height="874" /></a><figcaption>The known material of <i>Siamotyrannus</i>, which coexisted with <i>Phuwiangvenator</i></figcaption></figure> <p>There are three mid-sized <a href="/wiki/Theropod" class="mw-redirect" title="Theropod">theropods</a> which have been discovered so far in the <a href="/wiki/Sao_Khua_Formation" title="Sao Khua Formation">Sao Khua Formation</a>, all of which are known from relatively incomplete remains. These are: <i>Phuwiangvenator</i>, <i><a href="/wiki/Vayuraptor" title="Vayuraptor">Vayuraptor</a></i>, and <i><a href="/wiki/Siamotyrannus" title="Siamotyrannus">Siamotyrannus</a></i>. The authors who described the former two were careful to draw numerous distinctions between all three taxa to ensure that they none of the three are junior synonyms of the others. The distinctions they draw between <i>Phuwiangvenator</i> and <i>Siamotyrannus</i> are as follows: the dorsal vertebra of the former is longer than it is tall, whereas in the latter they are much taller in relation to their length and the rims of the <a href="/wiki/Sacrum" title="Sacrum">sacral</a> <a href="/wiki/Vertebral_centrum" class="mw-redirect" title="Vertebral centrum">centra</a> of <i>Siamotyrannus</i> are much more convex than in <i>Phuwiangvenator</i>.<sup id="cite_ref-description_2-11" class="reference"><a href="#cite_note-description-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> </p><p>The distinctions between <i>Phuwiangvenator</i> and <i>Vayuraptor</i> which they outline are as follows: the cranioproximate process of the astragalus is much more pronounced in <i>Vayuraptor</i> and they note that <i>Vayuraptor</i> had two grooves on the body of the astragalus whereas <i>Phuwiangvenator</i> only had one. The final distinction between the two are the lack of an upward curving process of the lateral condyle in <i>Vayuraptor</i>, which is one of the autapomorphies of <i>Phuwiangvenator</i>. This last trait was also observed in all <a href="/wiki/Ontogeny" title="Ontogeny">ontogenetic</a> stages of <i><a href="/wiki/Allosaurus" title="Allosaurus">Allosaurus</a></i>, which the authors state implies that the absence of this trait in <i>Vayuraptor</i> means that the distinction is probably not the result of ontogeny.<sup id="cite_ref-description_2-12" class="reference"><a href="#cite_note-description-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Paleobiology">Paleobiology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Phuwiangvenator&amp;action=edit&amp;section=6" title="Edit section: Paleobiology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Arm_musculature">Arm musculature</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Phuwiangvenator&amp;action=edit&amp;section=7" title="Edit section: Arm musculature"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>In 2023, Alexis M. Aranciaga Rolando and colleagues published an analysis of the inferred forelimb musculature of <a href="/wiki/Megaraptorans" class="mw-redirect" title="Megaraptorans">megaraptorans</a>. Their reasoning for pursuing this analysis was the uniquely large forearms of megaraptorans that were not seen in any other group of large <a href="/wiki/Theropod" class="mw-redirect" title="Theropod">theropods</a> in the <a href="/wiki/Late_Cretaceous" title="Late Cretaceous">Late Cretaceous</a>. The analysis focuses mostly on the derived members of megaraptoridae, but they comment on the <a href="/wiki/Muscle" title="Muscle">muscular</a> and <a href="/wiki/Tendon" title="Tendon">tendinous</a> attachment sites on the bones of <i>Phuwiangvenator</i>. They outline several key differences between the manual <a href="/wiki/Finger_bones" class="mw-redirect" title="Finger bones">phalanges</a> and <a href="/wiki/Claws" class="mw-redirect" title="Claws">unguals</a> of <i>Phuwiangvenator</i> and those of younger megaraptorans like <i><a href="/wiki/Megaraptor" title="Megaraptor">Megaraptor</a></i>, <i><a href="/wiki/Maip" title="Maip">Maip</a></i>, and <i><a href="/wiki/Australovenator" title="Australovenator">Australovenator</a></i>. These differences are concentrated in the first finger of the hand, which in derived megaraptorans hosts a single giant claw which is presumed to have served a purpose in hunting or feeding. The first claw of <i>Phuwiangvenator</i> lacks the well-developed furrows which are present on the proximal extension of the claw. In other species, these are thought to have been attachment sites for powerful muscles, enabling greater control and flexibility of the finger movements of these animals.<sup id="cite_ref-arms_8-0" class="reference"><a href="#cite_note-arms-8"><span class="cite-bracket">&#91;</span>8<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Pathology">Pathology</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Phuwiangvenator&amp;action=edit&amp;section=8" title="Edit section: Pathology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Phuwiangvenator_Metatarsals.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/7c/Phuwiangvenator_Metatarsals.png/220px-Phuwiangvenator_Metatarsals.png" decoding="async" width="220" height="227" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/7c/Phuwiangvenator_Metatarsals.png/330px-Phuwiangvenator_Metatarsals.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/7c/Phuwiangvenator_Metatarsals.png/440px-Phuwiangvenator_Metatarsals.png 2x" data-file-width="1154" data-file-height="1192" /></a><figcaption>The metatarsals of the holotype, with the fracture visible (B5)</figcaption></figure> <p>One of the <a href="/wiki/Metatarsals" class="mw-redirect" title="Metatarsals">metatarsals</a> of the holotype of <i>Phuwiangvenator</i> was found to have signs of a <a href="/wiki/Greenstick_fracture" title="Greenstick fracture">greenstick fracture</a> when analyzed by Samathi and colleagues in 2022. They inferred that this fracture is not taphonomic because of the texture of the bone, which is smooth on the medial side, but is very calloused on the lateral side. These are interpreted as signs of the fracture being healed, which implies that the animal survived this injury. The bone itself is also twisted, which would not be the normal condition because the second metatarsal would have been in the center of the foot in life. Samathi and colleagues compare this fracture to other known pathologies on theropod bones, including those seen in specimens <i><a href="/wiki/Allosaurus" title="Allosaurus">Allosaurus</a></i> and <i><a href="/wiki/Yangchuanosaurus" title="Yangchuanosaurus">Yangchuanosaurus</a></i>. From these comparisons, they conclude that the injury was likely caused by an external blunt force. Other features of the bone determined by <a href="/wiki/Computer_tomography" class="mw-redirect" title="Computer tomography">computer tomography</a> led the authors to conclude that the animal was not yet fully grown when it died.<sup id="cite_ref-pathology_3-1" class="reference"><a href="#cite_note-pathology-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> </p><p>Samathi and colleagues suggest several possible causes of the fracture. They first suggest a blow from the tail club of an <a href="/wiki/Ankylosaur" class="mw-redirect" title="Ankylosaur">ankylosaur</a>, but they discard this hypothesis because no ankylosaur remains are known from the <a href="/wiki/Early_Cretaceous" title="Early Cretaceous">Early Cretaceous</a> of <a href="/wiki/Thailand" title="Thailand">Thailand</a>. Possible competition or combat with another large theropod is also suggested, because there are several large theropods known from nearby localities, but they state that there is no direct evidence for this and it is impossible to determine without additional evidence. They conclude that the most likely explanation is that the animal hit its foot into a hard object such as a rock or possibly fell over to cause the type of incomplete fracture which is observed.<sup id="cite_ref-pathology_3-2" class="reference"><a href="#cite_note-pathology-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> </p><p>Evidence for severe pathologies being healed has been used by some authors to infer <a href="/wiki/Herd" title="Herd">herding</a> behavior in extinct animals. There are known associations of large theropods (i.e. <i><a href="/wiki/Albertosaurus" title="Albertosaurus">Albertosaurus</a></i> and <i><a href="/wiki/Mapusaurus" title="Mapusaurus">Mapusaurus</a></i>), so the authors state that it is possible that <i>Phuwiangvenator</i> exhibited similar behaviors. However, they also suggest that the animal may have simply survived for a short time after the injury and lived with a limp.<sup id="cite_ref-pathology_3-3" class="reference"><a href="#cite_note-pathology-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Paleoecology">Paleoecology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Phuwiangvenator&amp;action=edit&amp;section=9" title="Edit section: Paleoecology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Diet">Diet</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Phuwiangvenator&amp;action=edit&amp;section=10" title="Edit section: Diet"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>None of the skull of <i>Phuwiangvenator</i> has been preserved, but the animal is presumed to have been <a href="/wiki/Carnivore" title="Carnivore">carnivorous</a>. However, it does not have the same specialized arm musculature to aid in feeding as other megaraptorans.<sup id="cite_ref-arms_8-1" class="reference"><a href="#cite_note-arms-8"><span class="cite-bracket">&#91;</span>8<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Paleoenvironment">Paleoenvironment</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Phuwiangvenator&amp;action=edit&amp;section=11" title="Edit section: Paleoenvironment"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Spinosaurid_and_Kinnareemimus.PNG" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/0/06/Spinosaurid_and_Kinnareemimus.PNG/220px-Spinosaurid_and_Kinnareemimus.PNG" decoding="async" width="220" height="118" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/06/Spinosaurid_and_Kinnareemimus.PNG/330px-Spinosaurid_and_Kinnareemimus.PNG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/06/Spinosaurid_and_Kinnareemimus.PNG/440px-Spinosaurid_and_Kinnareemimus.PNG 2x" data-file-width="2250" data-file-height="1205" /></a><figcaption>A life reconstruction of some animals which coexisted with <i>Phuwiangvenator</i></figcaption></figure> <p>The <a href="/wiki/Sao_Khua_Formation" title="Sao Khua Formation">Sao Khua Formation</a> has thus far only yielded continental fossils, with no evidence of marine fossils or marine <a href="/wiki/Sedimentary_rock" title="Sedimentary rock">sediments</a>.<sup id="cite_ref-environment_9-0" class="reference"><a href="#cite_note-environment-9"><span class="cite-bracket">&#91;</span>9<span class="cite-bracket">&#93;</span></a></sup> There have historically been reports of <a href="/wiki/Ichthyosaur" class="mw-redirect" title="Ichthyosaur">ichthyosaur</a> and <a href="/wiki/Plesiosaur" title="Plesiosaur">plesiosaur</a> teeth from the Sao Khua Formation, but these have now been identified as belonging to <a href="/wiki/Crocodyliformes" title="Crocodyliformes">crocodyliformes</a>, which are mostly non-marine during the <a href="/wiki/Cretaceous" title="Cretaceous">Cretaceous</a>.<sup id="cite_ref-dinodiversity_10-0" class="reference"><a href="#cite_note-dinodiversity-10"><span class="cite-bracket">&#91;</span>10<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">&#91;</span>11<span class="cite-bracket">&#93;</span></a></sup> The sediments of the Sao Khua Formation are composed of <a href="/wiki/Red_clay" class="mw-redirect" title="Red clay">red clays</a>, <a href="/wiki/Mudstone" title="Mudstone">mudstones</a>, <a href="/wiki/Sandstone" title="Sandstone">sandstones</a>, <a href="/wiki/Siltstone" title="Siltstone">siltstones</a>, and <a href="/wiki/Conglomerate_(geology)" title="Conglomerate (geology)">conglomerate</a> rocks, which indicate a <a href="/wiki/Fluvial" class="mw-redirect" title="Fluvial">fluvial</a> <a href="/wiki/Depositional_environment" title="Depositional environment">environment</a> which also possessed lakes, <a href="/wiki/Floodplain" title="Floodplain">floodplains</a>, and <a href="/wiki/Braided_channel" class="mw-redirect" title="Braided channel">braided channels</a>.<sup id="cite_ref-environment_9-1" class="reference"><a href="#cite_note-environment-9"><span class="cite-bracket">&#91;</span>9<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-critters_12-0" class="reference"><a href="#cite_note-critters-12"><span class="cite-bracket">&#91;</span>12<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Contemporary_fauna">Contemporary fauna</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Phuwiangvenator&amp;action=edit&amp;section=12" title="Edit section: Contemporary fauna"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Of all the <a href="/wiki/Mesozoic" title="Mesozoic">Mesozoic</a> formations in northeastern Thailand, the Sao Khua has the most abundant and diverse fossils.<sup id="cite_ref-spino_13-0" class="reference"><a href="#cite_note-spino-13"><span class="cite-bracket">&#91;</span>13<span class="cite-bracket">&#93;</span></a></sup> In addition to <i>Phuwiangvenator</i> and <i><a href="/wiki/Vayuraptor" title="Vayuraptor">Vayuraptor</a></i>,<sup id="cite_ref-description_2-13" class="reference"><a href="#cite_note-description-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> the region was host to numerous other theropods including the <a href="/wiki/Spinosaurid" class="mw-redirect" title="Spinosaurid">spinosaurid</a> <i><a href="/wiki/Siamosaurus" title="Siamosaurus">Siamosaurus</a></i>,<sup id="cite_ref-spino_13-1" class="reference"><a href="#cite_note-spino-13"><span class="cite-bracket">&#91;</span>13<span class="cite-bracket">&#93;</span></a></sup> the large theropod <i><a href="/wiki/Siamotyrannus" title="Siamotyrannus">Siamotyrannus</a></i>, an unnamed spinosaurid, and an unnamed <a href="/wiki/Carcharodontosaurid" class="mw-redirect" title="Carcharodontosaurid">carcharodontosaurid</a>.<sup id="cite_ref-dinodiversity_10-1" class="reference"><a href="#cite_note-dinodiversity-10"><span class="cite-bracket">&#91;</span>10<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-:15_14-0" class="reference"><a href="#cite_note-:15-14"><span class="cite-bracket">&#91;</span>14<span class="cite-bracket">&#93;</span></a></sup> The area was also home to numerous small theropods including the <a href="/wiki/Ornithomimosaur" class="mw-redirect" title="Ornithomimosaur">ornithomimosaur</a> <i><a href="/wiki/Kinnareemimus" title="Kinnareemimus">Kinnareemimus</a></i> and an unnamed <a href="/wiki/Compsognathid" class="mw-redirect" title="Compsognathid">compsognathid</a>.<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">&#91;</span>15<span class="cite-bracket">&#93;</span></a></sup> Numerous <a href="/wiki/Sauropod" class="mw-redirect" title="Sauropod">sauropods</a> are also known from the area including <i><a href="/wiki/Phuwiangosaurus" title="Phuwiangosaurus">Phuwiangosaurus</a></i> and several unnamed forms. However, there have yet to be any <a href="/wiki/Ornithischian" class="mw-redirect" title="Ornithischian">ornithischians</a> discovered from the formation.<sup id="cite_ref-dinodiversity_10-2" class="reference"><a href="#cite_note-dinodiversity-10"><span class="cite-bracket">&#91;</span>10<span class="cite-bracket">&#93;</span></a></sup> </p><p>Freshwater life is abundant in the Sao Khua Formation. These include the <a href="/wiki/Neosuchian" class="mw-redirect" title="Neosuchian">neosuchians</a> <i><a href="/wiki/Siamosuchus" title="Siamosuchus">Siamosuchus</a></i>, <i><a href="/wiki/Sunosuchus" title="Sunosuchus">Sunosuchus</a></i>, and <i><a href="/wiki/Theriosuchus" title="Theriosuchus">Theriosuchus</a></i>, numerous species of <a href="/wiki/Turtles" class="mw-redirect" title="Turtles">turtles</a>, <a href="/wiki/Amiiform" class="mw-redirect" title="Amiiform">amiiform</a> fish, several <a href="/wiki/Hybodont" class="mw-redirect" title="Hybodont">hybodontiform</a> sharks, and freshwater <a href="/wiki/Bivalves" class="mw-redirect" title="Bivalves">bivalves</a>.<sup id="cite_ref-critters_12-1" class="reference"><a href="#cite_note-critters-12"><span class="cite-bracket">&#91;</span>12<span class="cite-bracket">&#93;</span></a></sup> There is also an indeterminate <a href="/wiki/Pterosaur" title="Pterosaur">pterosaur</a> known from the formation.<sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">&#91;</span>16<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Phuwiangvenator&amp;action=edit&amp;section=13" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="/wiki/2019_in_archosaur_paleontology" title="2019 in archosaur paleontology">2019 in archosaur paleontology</a></li> <li><a href="/wiki/Aptian_extinction" title="Aptian extinction">Aptian extinction</a></li> <li><a href="/wiki/Khorat_Group" title="Khorat Group">Khorat Group</a></li> <li><a href="/wiki/List_of_Asian_dinosaurs" title="List of Asian dinosaurs">List of Asian dinosaurs</a></li> <li><a href="/wiki/Paleontology_in_Thailand" title="Paleontology in Thailand">Paleontology in Thailand</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Phuwiangvenator&amp;action=edit&amp;section=14" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output 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.navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Theropoda" title="Template:Theropoda"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Theropoda" title="Template talk:Theropoda"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Theropoda" title="Special:EditPage/Template:Theropoda"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Theropoda" style="font-size:114%;margin:0 4em"><a href="/wiki/Theropoda" title="Theropoda">Theropoda</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="*_Kingdom:_Animalia_*_Phylum:_Chordata_*_Class:_Sauropsida_*_Clade:_Avemetatarsalia_*_Clade:_Dinosauria" style="font-size:114%;margin:0 4em"><span style="font-weight:normal; font-style:normal"><span style="font-size:85%;"> <ul><li>Kingdom: <a href="/wiki/Animal" title="Animal">Animalia</a></li> <li>Phylum: <a href="/wiki/Chordate" title="Chordate">Chordata</a></li> <li>Class: <a href="/wiki/Sauropsida" title="Sauropsida">Sauropsida</a></li> <li><i>Clade</i>: <a href="/wiki/Avemetatarsalia" title="Avemetatarsalia">Avemetatarsalia</a></li> <li><i>Clade</i>: <a href="/wiki/Dinosaur" title="Dinosaur">Dinosauria</a></li></ul> </span></span></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Avemetatarsalia" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Avemetatarsalia" title="Avemetatarsalia">Avemetatarsalia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Avemetatarsalia" title="Template:Avemetatarsalia">Avemetatarsalia</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Theropoda" title="Theropoda">Theropoda</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><div id="Theropoda" style="font-size:114%;margin:0 4em"><a href="/wiki/Theropoda" title="Theropoda">Theropoda</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Theropoda" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Theropoda" title="Theropoda">Theropoda</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Chindesaurus" title="Chindesaurus">Chindesaurus</a></i>?</li> <li><i><a href="/wiki/Daemonosaurus" title="Daemonosaurus">Daemonosaurus</a></i>?</li> <li><i><a href="/wiki/Eodromaeus" title="Eodromaeus">Eodromaeus</a></i>?</li> <li><i><a href="/wiki/Erythrovenator" title="Erythrovenator">Erythrovenator</a></i></li> <li><i><a href="/wiki/Guaibasaurus" title="Guaibasaurus">Guaibasaurus</a></i>?</li> <li><i><a href="/wiki/Nhandumirim" title="Nhandumirim">Nhandumirim</a></i>?</li> <li><i><a href="/wiki/Tawa_hallae" title="Tawa hallae">Tawa</a></i>?</li> <li><b><a href="/wiki/Herrerasauridae" title="Herrerasauridae">Herrerasauria</a></b>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Neotheropoda" class="mw-redirect" title="Neotheropoda">Neotheropoda</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Dilophosaurus" title="Dilophosaurus">Dilophosaurus</a></i></li> <li><i><a href="/wiki/Dolichosuchus" title="Dolichosuchus">Dolichosuchus</a></i>?</li> <li><i><a href="/wiki/Dracovenator" title="Dracovenator">Dracovenator</a></i></li> <li><i><a href="/wiki/Gojirasaurus" title="Gojirasaurus">Gojirasaurus</a></i></li> <li><i><a href="/wiki/Lepidus_praecisio" title="Lepidus praecisio">Lepidus</a></i></li> <li><i><a href="/wiki/Liliensternus" title="Liliensternus">Liliensternus</a></i></li> <li><i><a href="/wiki/Lophostropheus" title="Lophostropheus">Lophostropheus</a></i></li> <li><i><a href="/wiki/Notatesseraeraptor" title="Notatesseraeraptor">Notatesseraeraptor</a></i></li> <li><i><a href="/wiki/Sarcosaurus" title="Sarcosaurus">Sarcosaurus</a></i></li> <li><i><a href="/wiki/Shuangbaisaurus" title="Shuangbaisaurus">Shuangbaisaurus</a></i></li> <li><i><a href="/wiki/Sinosaurus" title="Sinosaurus">Sinosaurus</a></i></li> <li><i><a href="/wiki/Tachiraptor" title="Tachiraptor">Tachiraptor</a></i></li> <li><i><a href="/wiki/Velocipes" title="Velocipes">Velocipes</a></i>?</li> <li><i><a href="/wiki/Zupaysaurus" title="Zupaysaurus">Zupaysaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Coelophysoidea" title="Coelophysoidea">Coelophysoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Dracoraptor" title="Dracoraptor">Dracoraptor</a></i>?</li> <li><i><a href="/wiki/Panguraptor" title="Panguraptor">Panguraptor</a></i></li> <li><i><a href="/wiki/Pendraig" title="Pendraig">Pendraig</a></i></li> <li><i><a href="/wiki/Podokesaurus" title="Podokesaurus">Podokesaurus</a></i></li> <li><i><a href="/wiki/Powellvenator" title="Powellvenator">Powellvenator</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Coelophysidae" class="mw-redirect" title="Coelophysidae">Coelophysidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Camposaurus" title="Camposaurus">Camposaurus</a></i></li> <li><i><a href="/wiki/Coelophysis" title="Coelophysis">Coelophysis</a></i></li> <li><i><a href="/wiki/Lucianovenator" title="Lucianovenator">Lucianovenator</a></i></li> <li><i><a href="/wiki/Megapnosaurus" title="Megapnosaurus">Megapnosaurus</a></i>?</li> <li><i><a href="/wiki/Panguraptor" title="Panguraptor">Panguraptor</a></i>?</li> <li><i><a href="/wiki/Procompsognathus" title="Procompsognathus">Procompsognathus</a></i></li> <li><i><a href="/wiki/Pterospondylus" title="Pterospondylus">Pterospondylus</a></i>?</li> <li><i><a href="/wiki/Segisaurus" title="Segisaurus">Segisaurus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Averostra" class="mw-redirect" title="Averostra">Averostra</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Dubious neotheropods</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Teinurosaurus" title="Teinurosaurus">Teinurosaurus</a></i></li> <li><i><a href="/wiki/Halticosaurus" title="Halticosaurus">Halticosaurus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Coelophysis_Jeff_Martz_(flipped).jpg" class="mw-file-description" title="Coelophysis bauri"><img alt="Coelophysis bauri" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/d8/Coelophysis_Jeff_Martz_%28flipped%29.jpg/140px-Coelophysis_Jeff_Martz_%28flipped%29.jpg" decoding="async" width="140" height="49" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/d8/Coelophysis_Jeff_Martz_%28flipped%29.jpg/210px-Coelophysis_Jeff_Martz_%28flipped%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/d8/Coelophysis_Jeff_Martz_%28flipped%29.jpg/280px-Coelophysis_Jeff_Martz_%28flipped%29.jpg 2x" data-file-width="6000" data-file-height="2105" /></a></span></div> <span typeof="mw:File"><a href="/wiki/File:Dilophosaurus_wetherilli.PNG" class="mw-file-description" title="Dilophosaurus wetherilli"><img alt="Dilophosaurus wetherilli" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/d5/Dilophosaurus_wetherilli.PNG/140px-Dilophosaurus_wetherilli.PNG" decoding="async" width="140" height="49" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/d5/Dilophosaurus_wetherilli.PNG/210px-Dilophosaurus_wetherilli.PNG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/d5/Dilophosaurus_wetherilli.PNG/280px-Dilophosaurus_wetherilli.PNG 2x" data-file-width="1098" data-file-height="385" /></a></span></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><div id="Averostra" style="font-size:114%;margin:0 4em"><a href="/wiki/Averostra" class="mw-redirect" title="Averostra">Averostra</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Averostra" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Averostra" class="mw-redirect" title="Averostra">Averostra</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Dornraptor" title="Dornraptor">Dornraptor</a></i></li> <li><i><a href="/wiki/Lophostropheus" title="Lophostropheus">Lophostropheus</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Ceratosauria" title="Ceratosauria">Ceratosauria</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Berberosaurus" title="Berberosaurus">Berberosaurus</a></i></li> <li><i><a href="/wiki/Fosterovenator" title="Fosterovenator">Fosterovenator</a></i>?</li> <li><i><a href="/wiki/Jubbulpuria" title="Jubbulpuria">Jubbulpuria</a></i></li> <li><i><a href="/wiki/Lukousaurus" title="Lukousaurus">Lukousaurus</a></i>?</li> <li><i><a href="/wiki/Saltriovenator" title="Saltriovenator">Saltriovenator</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ceratosauridae" class="mw-redirect" title="Ceratosauridae">Ceratosauridae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Ceratosaurus" title="Ceratosaurus">Ceratosaurus</a></i></li> <li><i><a href="/wiki/Genyodectes" title="Genyodectes">Genyodectes</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Abelisauroidea" class="mw-redirect" title="Abelisauroidea">Abelisauroidea</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Betasuchus" title="Betasuchus">Betasuchus</a></i>?</li> <li><i><a href="/wiki/Lametasaurus" title="Lametasaurus">Lametasaurus</a></i>?</li> <li><i><a href="/wiki/Ornithomimoides" title="Ornithomimoides">Ornithomimoides</a></i>?</li> <li><i><a href="/wiki/Ozraptor" title="Ozraptor">Ozraptor</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Noasauridae" title="Noasauridae">Noasauridae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Afromimus" title="Afromimus">Afromimus</a></i>?</li> <li><i><a href="/wiki/Austrocheirus" title="Austrocheirus">Austrocheirus</a></i>?</li> <li><i><a href="/wiki/Berthasaura" title="Berthasaura">Berthasaura</a></i></li> <li><i><a href="/wiki/Compsosuchus" title="Compsosuchus">Compsosuchus</a></i>?</li> <li><i><a href="/wiki/Dahalokely" title="Dahalokely">Dahalokely</a></i>?</li> <li><i><a href="/wiki/Deltadromeus" title="Deltadromeus">Deltadromeus</a></i>?</li> <li><i><a href="/wiki/Genusaurus" title="Genusaurus">Genusaurus</a></i>?</li> <li><i><a href="/wiki/Kiyacursor" title="Kiyacursor">Kiyacursor</a></i></li> <li><i><a href="/wiki/Ligabueino" title="Ligabueino">Ligabueino</a></i></li> <li><i><a href="/wiki/Spinostropheus" title="Spinostropheus">Spinostropheus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Elaphrosaurinae" class="mw-redirect" title="Elaphrosaurinae">Elaphrosaurinae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Elaphrosaurus" title="Elaphrosaurus">Elaphrosaurus</a></i></li> <li><i><a href="/wiki/Huinculsaurus" title="Huinculsaurus">Huinculsaurus</a></i></li> <li><i><a href="/wiki/Limusaurus" title="Limusaurus">Limusaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Noasaurinae" class="mw-redirect" title="Noasaurinae">Noasaurinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Laevisuchus" title="Laevisuchus">Laevisuchus</a></i></li> <li><i><a href="/wiki/Masiakasaurus" title="Masiakasaurus">Masiakasaurus</a></i></li> <li><i><a href="/wiki/Noasaurus" title="Noasaurus">Noasaurus</a></i></li> <li><i><a href="/wiki/Velocisaurus" title="Velocisaurus">Velocisaurus</a></i></li> <li><i><a href="/wiki/Vespersaurus" title="Vespersaurus">Vespersaurus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Abelisauridae" title="Abelisauridae">Abelisauridae</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Chenanisaurus" title="Chenanisaurus">Chenanisaurus</a></i></li> <li><i><a href="/wiki/Dryptosauroides" title="Dryptosauroides">Dryptosauroides</a></i>?</li> <li><i><a href="/wiki/Eoabelisaurus" title="Eoabelisaurus">Eoabelisaurus</a></i></li> <li><i><a href="/wiki/Indosuchus" title="Indosuchus">Indosuchus</a></i></li> <li><i><a href="/wiki/Kryptops" title="Kryptops">Kryptops</a></i></li> <li><i><a href="/wiki/Kurupi_itaata" title="Kurupi itaata">Kurupi</a></i></li> <li><i><a href="/wiki/Rugops" title="Rugops">Rugops</a></i></li> <li><i><a href="/wiki/Spectrovenator" title="Spectrovenator">Spectrovenator</a></i></li> <li><i><a href="/wiki/Tarascosaurus" title="Tarascosaurus">Tarascosaurus</a></i></li> <li><i><a href="/wiki/Tralkasaurus" title="Tralkasaurus">Tralkasaurus</a></i></li> <li><i><a href="/wiki/Xenotarsosaurus" title="Xenotarsosaurus">Xenotarsosaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Majungasaurinae" title="Majungasaurinae">Majungasaurinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Arcovenator" title="Arcovenator">Arcovenator</a></i></li> <li><i><a href="/wiki/Dahalokely" title="Dahalokely">Dahalokely</a></i>?</li> <li><i><a href="/wiki/Genusaurus" title="Genusaurus">Genusaurus</a></i></li> <li><i><a href="/wiki/Indosaurus" title="Indosaurus">Indosaurus</a></i></li> <li><i><a href="/wiki/Majungasaurus" title="Majungasaurus">Majungasaurus</a></i></li> <li><i><a href="/wiki/Rahiolisaurus" title="Rahiolisaurus">Rahiolisaurus</a></i>?</li> <li><i><a href="/wiki/Rajasaurus" title="Rajasaurus">Rajasaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Carnotaurinae" class="mw-redirect" title="Carnotaurinae">Carnotaurinae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><b><a href="/wiki/Majungasaurinae" title="Majungasaurinae">Majungasaurinae</a></b>?</li></ul> </div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Brachyrostra" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Brachyrostra" title="Brachyrostra">Brachyrostra</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Dahalokely" title="Dahalokely">Dahalokely</a></i>?</li> <li><i><a href="/wiki/Ekrixinatosaurus" title="Ekrixinatosaurus">Ekrixinatosaurus</a></i></li> <li><i><a href="/wiki/Elemgasem" title="Elemgasem">Elemgasem</a></i></li> <li><i><a href="/wiki/Guemesia" title="Guemesia">Guemesia</a></i></li> <li><i><a href="/wiki/Ilokelesia" title="Ilokelesia">Ilokelesia</a></i></li> <li><i><a href="/wiki/Llukalkan" title="Llukalkan">Llukalkan</a></i></li> <li><i><a href="/wiki/Niebla_antiqua" title="Niebla antiqua">Niebla</a></i></li> <li><i><a href="/wiki/Rahiolisaurus" title="Rahiolisaurus">Rahiolisaurus</a></i>?</li> <li><i><a href="/wiki/Skorpiovenator" title="Skorpiovenator">Skorpiovenator</a></i></li> <li><i><a href="/wiki/Thanos_simonattoi" title="Thanos simonattoi">Thanos</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Furileusauria" class="mw-redirect" title="Furileusauria">Furileusauria</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Abelisaurus" title="Abelisaurus">Abelisaurus</a></i></li> <li><i><a href="/wiki/Aucasaurus" title="Aucasaurus">Aucasaurus</a></i></li> <li><i><a href="/wiki/Caletodraco" title="Caletodraco">Caletodraco</a></i></li> <li><i><a href="/wiki/Carnotaurus" title="Carnotaurus">Carnotaurus</a></i></li> <li><i><a href="/wiki/Elemgasem" title="Elemgasem">Elemgasem</a></i>?</li> <li><i><a href="/wiki/Koleken" title="Koleken">Koleken</a></i></li> <li><i><a href="/wiki/Pycnonemosaurus" title="Pycnonemosaurus">Pycnonemosaurus</a></i></li> <li><i><a href="/wiki/Quilmesaurus" title="Quilmesaurus">Quilmesaurus</a></i></li> <li><i><a href="/wiki/Viavenator" title="Viavenator">Viavenator</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Tetanurae" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Tetanurae" title="Tetanurae">Tetanurae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Ceratosaurus_nasicornis_DB.jpg" class="mw-file-description" title="Ceratosaurus nasicornis"><img alt="Ceratosaurus nasicornis" src="//upload.wikimedia.org/wikipedia/commons/thumb/2/23/Ceratosaurus_nasicornis_DB.jpg/140px-Ceratosaurus_nasicornis_DB.jpg" decoding="async" width="140" height="50" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/23/Ceratosaurus_nasicornis_DB.jpg/210px-Ceratosaurus_nasicornis_DB.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/23/Ceratosaurus_nasicornis_DB.jpg/280px-Ceratosaurus_nasicornis_DB.jpg 2x" data-file-width="939" data-file-height="334" /></a></span></div> <div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Limusaurus_runner_(flipped).jpg" class="mw-file-description" title="Limusaurus inextricabilis"><img alt="Limusaurus inextricabilis" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/96/Limusaurus_runner_%28flipped%29.jpg/140px-Limusaurus_runner_%28flipped%29.jpg" decoding="async" width="140" height="60" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/96/Limusaurus_runner_%28flipped%29.jpg/210px-Limusaurus_runner_%28flipped%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/96/Limusaurus_runner_%28flipped%29.jpg/280px-Limusaurus_runner_%28flipped%29.jpg 2x" data-file-width="3474" data-file-height="1496" /></a></span></div> <div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Rajasaurus_restoration.jpg" class="mw-file-description" title="Rajasaurus narmadensis"><img alt="Rajasaurus narmadensis" src="//upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Rajasaurus_restoration.jpg/140px-Rajasaurus_restoration.jpg" decoding="async" width="140" height="48" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Rajasaurus_restoration.jpg/210px-Rajasaurus_restoration.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Rajasaurus_restoration.jpg/280px-Rajasaurus_restoration.jpg 2x" data-file-width="2000" data-file-height="680" /></a></span></div> <div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Aucasaurus_garridoi_by_Paleocolour.jpg" class="mw-file-description" title="Aucasaurus garridoi"><img alt="Aucasaurus garridoi" src="//upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Aucasaurus_garridoi_by_Paleocolour.jpg/140px-Aucasaurus_garridoi_by_Paleocolour.jpg" decoding="async" width="140" height="53" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Aucasaurus_garridoi_by_Paleocolour.jpg/210px-Aucasaurus_garridoi_by_Paleocolour.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Aucasaurus_garridoi_by_Paleocolour.jpg/280px-Aucasaurus_garridoi_by_Paleocolour.jpg 2x" data-file-width="1500" data-file-height="564" /></a></span></div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><div id="Tetanurae" style="font-size:114%;margin:0 4em"><a href="/wiki/Tetanurae" title="Tetanurae">Tetanurae</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Tetanurae" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Tetanurae" title="Tetanurae">Tetanurae</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Calamospondylus" title="Calamospondylus">Calamospondylus</a></i>?</li> <li><i><a href="/wiki/Chienkosaurus" title="Chienkosaurus">Chienkosaurus</a></i>?</li> <li><i><a href="/wiki/Chilesaurus" title="Chilesaurus">Chilesaurus</a></i>?</li> <li><i><a href="/wiki/Chuandongocoelurus" title="Chuandongocoelurus">Chuandongocoelurus</a></i></li> <li><i><a href="/wiki/Cruxicheiros" title="Cruxicheiros">Cruxicheiros</a></i>?</li> <li><i><a href="/wiki/Cryolophosaurus" title="Cryolophosaurus">Cryolophosaurus</a></i></li> <li><i><a href="/wiki/Kaijiangosaurus" title="Kaijiangosaurus">Kaijiangosaurus</a></i></li> <li><i><a href="/wiki/Kayentavenator" title="Kayentavenator">Kayentavenator</a></i></li> <li><i><a href="/wiki/Monolophosaurus" title="Monolophosaurus">Monolophosaurus</a></i></li> <li><i><a href="/wiki/Pandoravenator" title="Pandoravenator">Pandoravenator</a></i></li> <li><i><a href="/wiki/Sinosaurus" title="Sinosaurus">Sinosaurus</a></i>?</li> <li><i><a href="/wiki/Szechuanosaurus" title="Szechuanosaurus">Szechuanosaurus</a></i></li> <li><i><a href="/wiki/Vectaerovenator" title="Vectaerovenator">Vectaerovenator</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Orionides" class="mw-redirect" title="Orionides">Orionides</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Megalosauroidea" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Megalosauroidea" title="Megalosauroidea">Megalosauroidea</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Yunyangosaurus" title="Yunyangosaurus">Yunyangosaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Piatnitzkysauridae" title="Piatnitzkysauridae">Piatnitzkysauridae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Condorraptor" title="Condorraptor">Condorraptor</a></i></li> <li><i><a href="/wiki/Marshosaurus" title="Marshosaurus">Marshosaurus</a></i></li> <li><i><a href="/wiki/Piatnitzkysaurus" title="Piatnitzkysaurus">Piatnitzkysaurus</a></i></li> <li><i><a href="/wiki/Xuanhanosaurus" title="Xuanhanosaurus">Xuanhanosaurus</a></i>?</li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Megalosauria" class="mw-redirect" title="Megalosauria">Megalosauria</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Streptospondylus" title="Streptospondylus">Streptospondylus</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Megalosauridae" title="Megalosauridae">Megalosauridae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Eustreptospondylus" title="Eustreptospondylus">Eustreptospondylus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Megalosaurinae" class="mw-redirect" title="Megalosaurinae">Megalosaurinae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Duriavenator" title="Duriavenator">Duriavenator</a></i></li> <li><i><a href="/wiki/Megalosaurus" title="Megalosaurus">Megalosaurus</a></i></li> <li><i><a href="/wiki/Torvosaurus" title="Torvosaurus">Torvosaurus</a></i></li> <li><i><a href="/wiki/Wiehenvenator" title="Wiehenvenator">Wiehenvenator</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Afrovenatorinae" class="mw-redirect" title="Afrovenatorinae">Afrovenatorinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Afrovenator" title="Afrovenator">Afrovenator</a></i></li> <li><i><a href="/wiki/Dubreuillosaurus" title="Dubreuillosaurus">Dubreuillosaurus</a></i></li> <li><i><a href="/wiki/Leshansaurus" title="Leshansaurus">Leshansaurus</a></i></li> <li><i><a href="/wiki/Magnosaurus" title="Magnosaurus">Magnosaurus</a></i></li> <li><i><a href="/wiki/Piveteausaurus" title="Piveteausaurus">Piveteausaurus</a></i></li> <li><i><a href="/wiki/Poekilopleuron" title="Poekilopleuron">Poekilopleuron</a></i>?</li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Spinosauridae" title="Spinosauridae">Spinosauridae</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Camarillasaurus" title="Camarillasaurus">Camarillasaurus</a></i>?</li> <li><i><a href="/wiki/Cristatusaurus" title="Cristatusaurus">Cristatusaurus</a></i></li> <li><i><a href="/wiki/Iberospinus" title="Iberospinus">Iberospinus</a></i></li> <li><i><a href="/wiki/Ostafrikasaurus" title="Ostafrikasaurus">Ostafrikasaurus</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Baryonychinae" title="Baryonychinae">Baryonychinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Baryonyx" title="Baryonyx">Baryonyx</a></i></li> <li><i><a href="/wiki/Protathlitis" title="Protathlitis">Protathlitis</a></i></li> <li><i><a href="/wiki/Riojavenatrix" title="Riojavenatrix">Riojavenatrix</a></i></li> <li><i><a href="/wiki/Suchosaurus" title="Suchosaurus">Suchosaurus</a></i>?</li> <li><i><a href="/wiki/Vallibonavenatrix" title="Vallibonavenatrix">Vallibonavenatrix</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ceratosuchopsini" class="mw-redirect" title="Ceratosuchopsini">Ceratosuchopsini</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Ceratosuchops" title="Ceratosuchops">Ceratosuchops</a></i></li> <li><i><a href="/wiki/Riparovenator" title="Riparovenator">Riparovenator</a></i></li> <li><i><a href="/wiki/Suchomimus" title="Suchomimus">Suchomimus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Spinosaurinae" class="mw-redirect" title="Spinosaurinae">Spinosaurinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Camarillasaurus" title="Camarillasaurus">Camarillasaurus</a></i>?</li> <li><i><a href="/wiki/Ichthyovenator" title="Ichthyovenator">Ichthyovenator</a></i></li> <li><i><a href="/wiki/Irritator" title="Irritator">Irritator</a></i></li> <li><i><a href="/wiki/Oxalaia" title="Oxalaia">Oxalaia</a></i></li> <li><i><a href="/wiki/Siamosaurus" title="Siamosaurus">Siamosaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Spinosaurini" class="mw-redirect" title="Spinosaurini">Spinosaurini</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Sigilmassasaurus" title="Sigilmassasaurus">Sigilmassasaurus</a></i></li> <li><i><a href="/wiki/Spinosaurus" title="Spinosaurus">Spinosaurus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Avetheropoda" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Avetheropoda" class="mw-redirect" title="Avetheropoda">Avetheropoda</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Piatnitzkysaurus_floresi_by_Paleocolour.jpg" class="mw-file-description" title="Piatnitzkysaurus floresi"><img alt="Piatnitzkysaurus floresi" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/d8/Piatnitzkysaurus_floresi_by_Paleocolour.jpg/140px-Piatnitzkysaurus_floresi_by_Paleocolour.jpg" decoding="async" width="140" height="55" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/d8/Piatnitzkysaurus_floresi_by_Paleocolour.jpg/210px-Piatnitzkysaurus_floresi_by_Paleocolour.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/d8/Piatnitzkysaurus_floresi_by_Paleocolour.jpg/280px-Piatnitzkysaurus_floresi_by_Paleocolour.jpg 2x" data-file-width="1500" data-file-height="589" /></a></span></div> <p><span typeof="mw:File"><a href="/wiki/File:Torvosaurus_tanneri_Reconstruction_(cropped).png" class="mw-file-description" title="Torvosaurus tanneri"><img alt="Torvosaurus tanneri" src="//upload.wikimedia.org/wikipedia/commons/thumb/5/54/Torvosaurus_tanneri_Reconstruction_%28cropped%29.png/140px-Torvosaurus_tanneri_Reconstruction_%28cropped%29.png" decoding="async" width="140" height="44" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/54/Torvosaurus_tanneri_Reconstruction_%28cropped%29.png/210px-Torvosaurus_tanneri_Reconstruction_%28cropped%29.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/54/Torvosaurus_tanneri_Reconstruction_%28cropped%29.png/280px-Torvosaurus_tanneri_Reconstruction_%28cropped%29.png 2x" data-file-width="1842" data-file-height="576" /></a></span> </p> <div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Spinosaurus_aegyptiacus.png" class="mw-file-description" title="Spinosaurus aegyptiacus"><img alt="Spinosaurus aegyptiacus" src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e0/Spinosaurus_aegyptiacus.png/140px-Spinosaurus_aegyptiacus.png" decoding="async" width="140" height="86" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/e0/Spinosaurus_aegyptiacus.png/210px-Spinosaurus_aegyptiacus.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/e0/Spinosaurus_aegyptiacus.png/280px-Spinosaurus_aegyptiacus.png 2x" data-file-width="4485" data-file-height="2764" /></a></span></div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible uncollapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><div id="Avetheropoda" style="font-size:114%;margin:0 4em"><a href="/wiki/Avetheropoda" class="mw-redirect" title="Avetheropoda">Avetheropoda</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Avetheropoda" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Avetheropoda" class="mw-redirect" title="Avetheropoda">Avetheropoda</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Gasosaurus" title="Gasosaurus">Gasosaurus</a></i>?</li> <li><i><a href="/wiki/Lourinhanosaurus" title="Lourinhanosaurus">Lourinhanosaurus</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Carnosauria" title="Carnosauria">Carnosauria</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Altispinax" title="Altispinax">Altispinax</a></i></li> <li><i><a href="/wiki/Monolophosaurus" title="Monolophosaurus">Monolophosaurus</a></i>?</li> <li><b><a href="/wiki/Megalosauridae" title="Megalosauridae">Megalosauridae</a></b>?</li> <li><b><a href="/wiki/Spinosauridae" title="Spinosauridae">Spinosauridae</a></b>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Allosauroidea" class="mw-redirect" title="Allosauroidea">Allosauroidea</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Asfaltovenator" title="Asfaltovenator">Asfaltovenator</a></i></li> <li><i><a href="/wiki/Erectopus" title="Erectopus">Erectopus</a></i></li> <li><i><a href="/wiki/Xuanhanosaurus" title="Xuanhanosaurus">Xuanhanosaurus</a></i>?</li> <li><b><a href="/wiki/Piatnitzkysauridae" title="Piatnitzkysauridae">Piatnitzkysauridae</a></b>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Metriacanthosauridae" title="Metriacanthosauridae">Metriacanthosauridae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Xuanhanosaurus" title="Xuanhanosaurus">Xuanhanosaurus</a></i>?</li> <li><i><a href="/wiki/Yangchuanosaurus" title="Yangchuanosaurus">Yangchuanosaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Metriacanthosaurinae" class="mw-redirect" title="Metriacanthosaurinae">Metriacanthosaurinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alpkarakush" title="Alpkarakush">Alpkarakush</a></i></li> <li><i><a href="/wiki/Metriacanthosaurus" title="Metriacanthosaurus">Metriacanthosaurus</a></i></li> <li><i><a href="/wiki/Shidaisaurus" title="Shidaisaurus">Shidaisaurus</a></i></li> <li><i><a href="/wiki/Siamotyrannus" title="Siamotyrannus">Siamotyrannus</a></i></li> <li><i><a href="/wiki/Sinraptor" title="Sinraptor">Sinraptor</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Allosauria" class="mw-redirect" title="Allosauria">Allosauria</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Allosauridae" title="Allosauridae">Allosauridae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Allosaurus" title="Allosaurus">Allosaurus</a></i></li> <li><i><a href="/wiki/Antrodemus" title="Antrodemus">Antrodemus</a></i>?</li> <li><i><a href="/wiki/Epanterias" title="Epanterias">Epanterias</a></i>?</li> <li><i><a href="/wiki/Saurophaganax" title="Saurophaganax">Saurophaganax</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Carcharodontosauria" class="mw-redirect" title="Carcharodontosauria">Carcharodontosauria</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Datanglong" title="Datanglong">Datanglong</a></i></li> <li><i><a href="/wiki/Lusovenator" title="Lusovenator">Lusovenator</a></i></li> <li><i><a href="/wiki/Siamraptor" title="Siamraptor">Siamraptor</a></i></li> <li><i><a href="/wiki/Siats" title="Siats">Siats</a></i>?</li> <li><i><a href="/wiki/Ulughbegsaurus" title="Ulughbegsaurus">Ulughbegsaurus</a></i>?</li> <li><i><a href="/wiki/Veterupristisaurus" title="Veterupristisaurus">Veterupristisaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Neovenatoridae" class="mw-redirect" title="Neovenatoridae">Neovenatoridae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Chilantaisaurus" title="Chilantaisaurus">Chilantaisaurus</a></i>?</li> <li><i><a href="/wiki/Gualicho" title="Gualicho">Gualicho</a></i>?</li> <li><i><a href="/wiki/Neovenator" title="Neovenator">Neovenator</a></i></li> <li><i><a href="/wiki/Siats" title="Siats">Siats</a></i>?</li> <li><b><a href="/wiki/Megaraptora" title="Megaraptora">Megaraptora</a></b>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Carcharodontosauridae" title="Carcharodontosauridae">Carcharodontosauridae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Acrocanthosaurus" title="Acrocanthosaurus">Acrocanthosaurus</a></i></li> <li><i><a href="/wiki/Altispinax" title="Altispinax">Altispinax</a></i>?</li> <li><i><a href="/wiki/Concavenator" title="Concavenator">Concavenator</a></i></li> <li><i><a href="/wiki/Eocarcharia" title="Eocarcharia">Eocarcharia</a></i></li> <li><i><a href="/wiki/Kelmayisaurus" title="Kelmayisaurus">Kelmayisaurus</a></i></li> <li><i><a href="/wiki/Lajasvenator" title="Lajasvenator">Lajasvenator</a></i></li> <li><i><a href="/wiki/Sauroniops" title="Sauroniops">Sauroniops</a></i></li> <li><i><a href="/wiki/Shaochilong" title="Shaochilong">Shaochilong</a></i></li> <li><i><a href="/wiki/Ulughbegsaurus" title="Ulughbegsaurus">Ulughbegsaurus</a></i>?</li> <li><i><a href="/wiki/Veterupristisaurus" title="Veterupristisaurus">Veterupristisaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Carcharodontosaurinae" class="mw-redirect" title="Carcharodontosaurinae">Carcharodontosaurinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Carcharodontosaurus" title="Carcharodontosaurus">Carcharodontosaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Giganotosaurini" class="mw-redirect" title="Giganotosaurini">Giganotosaurini</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Giganotosaurus" title="Giganotosaurus">Giganotosaurus</a></i></li> <li><i><a href="/wiki/Mapusaurus" title="Mapusaurus">Mapusaurus</a></i></li> <li><i><a href="/wiki/Meraxes" title="Meraxes">Meraxes</a></i></li> <li><i><a href="/wiki/Taurovenator" title="Taurovenator">Taurovenator</a></i></li> <li><i><a href="/wiki/Tyrannotitan" title="Tyrannotitan">Tyrannotitan</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Megaraptora" title="Megaraptora">Megaraptora</a>?</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Aoniraptor" title="Aoniraptor">Aoniraptor</a></i></li> <li><i><a href="/wiki/Bahariasaurus" title="Bahariasaurus">Bahariasaurus</a></i>?</li> <li><i><a href="/wiki/Deltadromeus" title="Deltadromeus">Deltadromeus</a></i>?</li> <li><i><a href="/wiki/Fukuiraptor" title="Fukuiraptor">Fukuiraptor</a></i></li> <li><i><a class="mw-selflink selflink">Phuwiangvenator</a></i></li> <li><i><a href="/wiki/Rapator" title="Rapator">Rapator</a></i></li> <li><i><a href="/wiki/Siats" title="Siats">Siats</a>?</i></li> <li><i><a href="/wiki/Chilantaisaurus" title="Chilantaisaurus">Chilantaisaurus</a>?</i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Megaraptoridae" class="mw-redirect" title="Megaraptoridae">Megaraptoridae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Aerosteon" title="Aerosteon">Aerosteon</a></i></li> <li><i><a href="/wiki/Australovenator" title="Australovenator">Australovenator</a></i></li> <li><i><a href="/wiki/Maip" title="Maip">Maip</a></i></li> <li><i><a href="/wiki/Megaraptor" title="Megaraptor">Megaraptor</a></i></li> <li><i><a href="/wiki/Murusraptor" title="Murusraptor">Murusraptor</a></i></li> <li><i><a href="/wiki/Orkoraptor" title="Orkoraptor">Orkoraptor</a></i></li> <li><i><a href="/wiki/Tratayenia" title="Tratayenia">Tratayenia</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Coelurosauria" title="Coelurosauria">Coelurosauria</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Xuanhanosaurus_qilixiaensis.png" class="mw-file-description" title="Xuanhanosaurus qilixiaensis"><img alt="Xuanhanosaurus qilixiaensis" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/9b/Xuanhanosaurus_qilixiaensis.png/140px-Xuanhanosaurus_qilixiaensis.png" decoding="async" width="140" height="58" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/9b/Xuanhanosaurus_qilixiaensis.png/210px-Xuanhanosaurus_qilixiaensis.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/9b/Xuanhanosaurus_qilixiaensis.png/280px-Xuanhanosaurus_qilixiaensis.png 2x" data-file-width="5602" data-file-height="2311" /></a></span> <div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Allosaurus_Revised.jpg" class="mw-file-description" title="Allosaurus fragilis"><img alt="Allosaurus fragilis" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/3d/Allosaurus_Revised.jpg/140px-Allosaurus_Revised.jpg" decoding="async" width="140" height="64" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/3d/Allosaurus_Revised.jpg/210px-Allosaurus_Revised.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/3d/Allosaurus_Revised.jpg/280px-Allosaurus_Revised.jpg 2x" data-file-width="4570" data-file-height="2093" /></a></span></div> <p><span typeof="mw:File"><a href="/wiki/File:Neovenator.png" class="mw-file-description" title="Neovenator salerii"><img alt="Neovenator salerii" src="//upload.wikimedia.org/wikipedia/commons/thumb/6/61/Neovenator.png/140px-Neovenator.png" decoding="async" width="140" height="84" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/61/Neovenator.png/210px-Neovenator.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/61/Neovenator.png/280px-Neovenator.png 2x" data-file-width="5000" data-file-height="3000" /></a></span> <span typeof="mw:File"><a href="/wiki/File:Carcharodontosaurus.png" class="mw-file-description" title="Carcharodontosaurus saharicus"><img alt="Carcharodontosaurus saharicus" src="//upload.wikimedia.org/wikipedia/commons/thumb/5/52/Carcharodontosaurus.png/140px-Carcharodontosaurus.png" decoding="async" width="140" height="56" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/52/Carcharodontosaurus.png/210px-Carcharodontosaurus.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/52/Carcharodontosaurus.png/280px-Carcharodontosaurus.png 2x" data-file-width="4057" data-file-height="1609" /></a></span> </p> <div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Australovenator.jpg" class="mw-file-description" title="Australovenator wintonensis"><img alt="Australovenator wintonensis" src="//upload.wikimedia.org/wikipedia/commons/thumb/7/7a/Australovenator.jpg/140px-Australovenator.jpg" decoding="async" width="140" height="49" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/7a/Australovenator.jpg/210px-Australovenator.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/7a/Australovenator.jpg/280px-Australovenator.jpg 2x" data-file-width="937" data-file-height="328" /></a></span></div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><div id="Coelurosauria" style="font-size:114%;margin:0 4em"><a href="/wiki/Coelurosauria" title="Coelurosauria">Coelurosauria</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Coelurosauria" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Coelurosauria" title="Coelurosauria">Coelurosauria</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Aratasaurus" title="Aratasaurus">Aratasaurus</a></i></li> <li><i><a href="/wiki/Asiamericana" title="Asiamericana">Asiamericana</a></i></li> <li><i><a href="/wiki/Bahariasaurus" title="Bahariasaurus">Bahariasaurus</a></i>?</li> <li><i><a href="/wiki/Bicentenaria" title="Bicentenaria">Bicentenaria</a></i></li> <li><i><a href="/wiki/Chilantaisaurus" title="Chilantaisaurus">Chilantaisaurus</a></i>?</li> <li><i><a href="/wiki/Gualicho" title="Gualicho">Gualicho</a></i>?</li> <li><i><a href="/wiki/Richardoestesia" title="Richardoestesia">Richardoestesia</a></i></li> <li><i><a href="/wiki/Sciurumimus" title="Sciurumimus">Sciurumimus</a></i>?</li> <li><i><a href="/wiki/Vayuraptor" title="Vayuraptor">Vayuraptor</a></i></li> <li><i><a href="/wiki/Xinjiangovenator" title="Xinjiangovenator">Xinjiangovenator</a></i></li> <li><i><a href="/wiki/Zuolong" title="Zuolong">Zuolong</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Tyrannoraptora" class="mw-redirect" title="Tyrannoraptora">Tyrannoraptora</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Tyrannosauroidea" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Tyrannosauroidea" title="Tyrannosauroidea">Tyrannosauroidea</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Chingkankousaurus" title="Chingkankousaurus">Chingkankousaurus</a></i></li> <li><i><a href="/wiki/Labocania" title="Labocania">Labocania</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Coeluridae" title="Coeluridae">Coeluridae</a>?</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Coelurus" title="Coelurus">Coelurus</a></i></li> <li><i><a href="/wiki/Tanycolagreus" title="Tanycolagreus">Tanycolagreus</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Proceratosauridae" title="Proceratosauridae">Proceratosauridae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Guanlong" title="Guanlong">Guanlong</a></i></li> <li><i><a href="/wiki/Kileskus" title="Kileskus">Kileskus</a></i></li> <li><i><a href="/wiki/Proceratosaurus" title="Proceratosaurus">Proceratosaurus</a></i></li> <li><i><a href="/wiki/Sinotyrannus" title="Sinotyrannus">Sinotyrannus</a></i></li> <li><i><a href="/wiki/Yutyrannus" title="Yutyrannus">Yutyrannus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Pantyrannosauria" class="mw-redirect" title="Pantyrannosauria">Pantyrannosauria</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alectrosaurus" title="Alectrosaurus">Alectrosaurus</a></i></li> <li><i><a href="/wiki/Aviatyrannis" title="Aviatyrannis">Aviatyrannis</a></i></li> <li><i><a href="/wiki/Bagaraatan" title="Bagaraatan">Bagaraatan</a></i>?</li> <li><i><a href="/wiki/Dilong_paradoxus" title="Dilong paradoxus">Dilong</a></i></li> <li><i><a href="/wiki/Eotyrannus" title="Eotyrannus">Eotyrannus</a></i></li> <li><i><a href="/wiki/Jinbeisaurus" title="Jinbeisaurus">Jinbeisaurus</a></i></li> <li><i><a href="/wiki/Juratyrant" title="Juratyrant">Juratyrant</a></i></li> <li><i><a href="/wiki/Moros_intrepidus" title="Moros intrepidus">Moros</a></i></li> <li><i><a href="/wiki/Santanaraptor" title="Santanaraptor">Santanaraptor</a></i></li> <li><i><a href="/wiki/Stokesosaurus" title="Stokesosaurus">Stokesosaurus</a></i></li> <li><i><a href="/wiki/Suskityrannus" title="Suskityrannus">Suskityrannus</a></i></li> <li><i><a href="/wiki/Timimus" title="Timimus">Timimus</a></i>?</li> <li><i><a href="/wiki/Timurlengia" title="Timurlengia">Timurlengia</a></i></li> <li><i><a href="/wiki/Xiongguanlong" title="Xiongguanlong">Xiongguanlong</a></i></li> <li><b><a href="/wiki/Megaraptora" title="Megaraptora">Megaraptora</a></b>?</li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Eutyrannosauria" class="mw-redirect" title="Eutyrannosauria">Eutyrannosauria</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Appalachiosaurus" title="Appalachiosaurus">Appalachiosaurus</a></i></li> <li><i><a href="/wiki/Bistahieversor" title="Bistahieversor">Bistahieversor</a></i>?</li> <li><i><a href="/wiki/Dryptosaurus" title="Dryptosaurus">Dryptosaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Tyrannosauridae" title="Tyrannosauridae">Tyrannosauridae</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Albertosaurinae" title="Albertosaurinae">Albertosaurinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Albertosaurus" title="Albertosaurus">Albertosaurus</a></i></li> <li><i><a href="/wiki/Gorgosaurus" title="Gorgosaurus">Gorgosaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Tyrannosaurinae" title="Tyrannosaurinae">Tyrannosaurinae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Asiatyrannus" title="Asiatyrannus">Asiatyrannus</a></i></li> <li><i><a href="/wiki/Nanuqsaurus" title="Nanuqsaurus">Nanuqsaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Alioramini" title="Alioramini">Alioramini</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alioramus" title="Alioramus">Alioramus</a></i></li> <li><i><a href="/wiki/Qianzhousaurus" title="Qianzhousaurus">Qianzhousaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Daspletosaurini" title="Daspletosaurini">Daspletosaurini</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Daspletosaurus" title="Daspletosaurus">Daspletosaurus</a></i></li> <li><i><a href="/wiki/Thanatotheristes" title="Thanatotheristes">Thanatotheristes</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Teratophoneini" class="mw-redirect" title="Teratophoneini">Teratophoneini</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Bistahieversor" title="Bistahieversor">Bistahieversor</a></i>?</li> <li><i><a href="/wiki/Dynamoterror" title="Dynamoterror">Dynamoterror</a></i></li> <li><i><a href="/wiki/Labocania" title="Labocania">Labocania</a></i>?</li> <li><i><a href="/wiki/Lythronax" title="Lythronax">Lythronax</a></i></li> <li><i><a href="/wiki/Teratophoneus" title="Teratophoneus">Teratophoneus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Tyrannosaurini" class="mw-redirect" title="Tyrannosaurini">Tyrannosaurini</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Tarbosaurus" title="Tarbosaurus">Tarbosaurus</a></i></li> <li><i><a href="/wiki/Tyrannosaurus" title="Tyrannosaurus">Tyrannosaurus</a></i></li> <li><i><a href="/wiki/Zhuchengtyrannus" title="Zhuchengtyrannus">Zhuchengtyrannus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Maniraptoromorpha" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Maniraptoromorpha" class="mw-redirect" title="Maniraptoromorpha">Maniraptoromorpha</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Dubious_coelurosaurs" scope="row" class="navbox-group" style="width:1%">Dubious coelurosaurs</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Iliosuchus" title="Iliosuchus">Iliosuchus</a></i></li> <li><i><a href="/wiki/Kakuru" title="Kakuru">Kakuru</a></i></li> <li><i><a href="/wiki/Phaedrolosaurus" title="Phaedrolosaurus">Phaedrolosaurus</a></i></li> <li><i><a href="/wiki/Shanyangosaurus" title="Shanyangosaurus">Shanyangosaurus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Zuolong_salleei.jpg" class="mw-file-description" title="Zuolong salleei"><img alt="Zuolong salleei" src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e6/Zuolong_salleei.jpg/140px-Zuolong_salleei.jpg" decoding="async" width="140" height="73" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/e6/Zuolong_salleei.jpg/210px-Zuolong_salleei.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/e6/Zuolong_salleei.jpg/280px-Zuolong_salleei.jpg 2x" data-file-width="2735" data-file-height="1431" /></a></span> <div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Stokesosaurus_by_Tom_Parker.png" class="mw-file-description" title="Stokesosaurus clevelandi"><img alt="Stokesosaurus clevelandi" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4b/Stokesosaurus_by_Tom_Parker.png/140px-Stokesosaurus_by_Tom_Parker.png" decoding="async" width="140" height="54" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4b/Stokesosaurus_by_Tom_Parker.png/210px-Stokesosaurus_by_Tom_Parker.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4b/Stokesosaurus_by_Tom_Parker.png/280px-Stokesosaurus_by_Tom_Parker.png 2x" data-file-width="2262" data-file-height="867" /></a></span></div> <p><span typeof="mw:File"><a href="/wiki/File:Alioramus_Life_Restoration.jpg" class="mw-file-description" title="Alioramus remotus"><img alt="Alioramus remotus" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/da/Alioramus_Life_Restoration.jpg/140px-Alioramus_Life_Restoration.jpg" decoding="async" width="140" height="79" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/da/Alioramus_Life_Restoration.jpg/210px-Alioramus_Life_Restoration.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/da/Alioramus_Life_Restoration.jpg/280px-Alioramus_Life_Restoration.jpg 2x" data-file-width="3548" data-file-height="2009" /></a></span> </p> <div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Tarbosaurus_Steveoc86_flipped.jpg" class="mw-file-description" title="Tarbosaurus bataar"><img alt="Tarbosaurus bataar" src="//upload.wikimedia.org/wikipedia/commons/thumb/2/20/Tarbosaurus_Steveoc86_flipped.jpg/140px-Tarbosaurus_Steveoc86_flipped.jpg" decoding="async" width="140" height="50" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/20/Tarbosaurus_Steveoc86_flipped.jpg/210px-Tarbosaurus_Steveoc86_flipped.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/20/Tarbosaurus_Steveoc86_flipped.jpg/280px-Tarbosaurus_Steveoc86_flipped.jpg 2x" data-file-width="1024" data-file-height="366" /></a></span></div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><div id="Maniraptoromorpha" style="font-size:114%;margin:0 4em"><a href="/wiki/Maniraptoromorpha" class="mw-redirect" title="Maniraptoromorpha">Maniraptoromorpha</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Maniraptoromorpha" scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Maniraptoromorpha" class="mw-redirect" title="Maniraptoromorpha">Maniraptoromorpha</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Coelurus" title="Coelurus">Coelurus</a></i>?</li> <li><i><a href="/wiki/Juravenator" title="Juravenator">Juravenator</a></i></li> <li><i><a href="/wiki/Ornitholestes" title="Ornitholestes">Ornitholestes</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Neocoelurosauria" class="mw-redirect" title="Neocoelurosauria">Neocoelurosauria</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Aniksosaurus" title="Aniksosaurus">Aniksosaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Compsognathidae" title="Compsognathidae">Compsognathidae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Aniksosaurus" title="Aniksosaurus">Aniksosaurus</a></i>?</li> <li><i><a href="/wiki/Aristosuchus" title="Aristosuchus">Aristosuchus</a></i></li> <li><i><a href="/wiki/Beipiaognathus" title="Beipiaognathus">Beipiaognathus</a></i>?</li> <li><i><a href="/wiki/Compsognathus" title="Compsognathus">Compsognathus</a></i></li> <li><i><a href="/wiki/Huaxiagnathus" title="Huaxiagnathus">Huaxiagnathus</a></i></li> <li><i><a href="/wiki/Mirischia" title="Mirischia">Mirischia</a></i></li> <li><i><a href="/wiki/Scipionyx" title="Scipionyx">Scipionyx</a></i>?</li> <li><i><a href="/wiki/Sinocalliopteryx" title="Sinocalliopteryx">Sinocalliopteryx</a></i></li> <li><i><a href="/wiki/Sinosauropteryx" title="Sinosauropteryx">Sinosauropteryx</a></i></li> <li><i><a href="/wiki/Xunmenglong" title="Xunmenglong">Xunmenglong</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position:absolute;-webkit-transform:translate(-50%,-50%) rotate(270deg);-moz-transform:translate(-50%,-50%) rotate(270deg);-ms-transform:translate(-50%,-50%) rotate(270deg);-o-transform:translate(-50%,-50%) rotate(270deg);transform:translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Maniraptoriformes" class="mw-redirect" title="Maniraptoriformes">Maniraptoriformes</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><b><a href="/wiki/Compsognathidae" title="Compsognathidae">Compsognathidae</a></b>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ornithomimosauria" title="Ornithomimosauria">Ornithomimosauria</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Arkansaurus" title="Arkansaurus">Arkansaurus</a></i></li> <li><i><a href="/wiki/Aviatyrannis" title="Aviatyrannis">Aviatyrannis</a></i>?</li> <li><i><a href="/wiki/Calamosaurus" title="Calamosaurus">Calamosaurus</a></i>?</li> <li><i><a href="/wiki/Hexing" title="Hexing">Hexing</a></i></li> <li><i><a href="/wiki/Kinnareemimus" title="Kinnareemimus">Kinnareemimus</a></i></li> <li><i><a href="/wiki/Nedcolbertia" title="Nedcolbertia">Nedcolbertia</a></i></li> <li><i><a href="/wiki/Nqwebasaurus" title="Nqwebasaurus">Nqwebasaurus</a></i></li> <li><i><a href="/wiki/Thecocoelurus" title="Thecocoelurus">Thecocoelurus</a></i>?</li> <li><i><a href="/wiki/Valdoraptor" title="Valdoraptor">Valdoraptor</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Macrocheiriformes" class="mw-redirect" title="Macrocheiriformes">Macrocheiriformes</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Harpymimus" title="Harpymimus">Harpymimus</a></i></li> <li><i><a href="/wiki/Pelecanimimus" title="Pelecanimimus">Pelecanimimus</a></i></li> <li><i><a href="/wiki/Shenzhousaurus" title="Shenzhousaurus">Shenzhousaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Deinocheiridae" title="Deinocheiridae">Deinocheiridae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Beishanlong" title="Beishanlong">Beishanlong</a></i></li> <li><i><a href="/wiki/Deinocheirus" title="Deinocheirus">Deinocheirus</a></i></li> <li><i><a href="/wiki/Garudimimus" title="Garudimimus">Garudimimus</a></i></li> <li><i><a href="/wiki/Harpymimus" title="Harpymimus">Harpymimus</a></i>?</li> <li><i><a href="/wiki/Paraxenisaurus" title="Paraxenisaurus">Paraxenisaurus</a></i>?</li> <li><i><a href="/wiki/Tyrannomimus" title="Tyrannomimus">Tyrannomimus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ornithomimidae" title="Ornithomimidae">Ornithomimidae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Aepyornithomimus" title="Aepyornithomimus">Aepyornithomimus</a></i></li> <li><i><a href="/wiki/Anserimimus" title="Anserimimus">Anserimimus</a></i></li> <li><i><a href="/wiki/Archaeornithomimus" title="Archaeornithomimus">Archaeornithomimus</a></i></li> <li><i><a href="/wiki/%22Coelosaurus%22_antiquus" title="&quot;Coelosaurus&quot; antiquus">"Coelosaurus"</a></i></li> <li><i><a href="/wiki/Dromiceiomimus" title="Dromiceiomimus">Dromiceiomimus</a></i></li> <li><i><a href="/wiki/Gallimimus" title="Gallimimus">Gallimimus</a></i></li> <li><i><a href="/wiki/Ornithomimus" title="Ornithomimus">Ornithomimus</a></i></li> <li><i><a href="/wiki/Qiupalong" title="Qiupalong">Qiupalong</a></i></li> <li><i><a href="/wiki/Rativates" title="Rativates">Rativates</a></i></li> <li><i><a href="/wiki/Sinornithomimus" title="Sinornithomimus">Sinornithomimus</a></i></li> <li><i><a href="/wiki/Struthiomimus" title="Struthiomimus">Struthiomimus</a></i></li> <li><i><a href="/wiki/Tototlmimus" title="Tototlmimus">Tototlmimus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Maniraptora" title="Maniraptora">Maniraptora</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Maniraptora" title="Template:Maniraptora">Maniraptora</a></li> <li>includes <a href="/wiki/Bird" title="Bird">birds</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><div style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Sinosauropteryx_color_(without_scale_bar).jpg" class="mw-file-description" title="Sinosauropteryx prima"><img alt="Sinosauropteryx prima" src="//upload.wikimedia.org/wikipedia/commons/thumb/e/ed/Sinosauropteryx_color_%28without_scale_bar%29.jpg/140px-Sinosauropteryx_color_%28without_scale_bar%29.jpg" decoding="async" width="140" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/ed/Sinosauropteryx_color_%28without_scale_bar%29.jpg/210px-Sinosauropteryx_color_%28without_scale_bar%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/ed/Sinosauropteryx_color_%28without_scale_bar%29.jpg/280px-Sinosauropteryx_color_%28without_scale_bar%29.jpg 2x" data-file-width="1345" data-file-height="380" /></a></span></div> <p><span typeof="mw:File"><a href="/wiki/File:Deinocheirus_NT.png" class="mw-file-description" title="Deinocheirus mirificus"><img alt="Deinocheirus mirificus" src="//upload.wikimedia.org/wikipedia/commons/thumb/2/25/Deinocheirus_NT.png/130px-Deinocheirus_NT.png" decoding="async" width="130" height="66" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/25/Deinocheirus_NT.png/195px-Deinocheirus_NT.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/25/Deinocheirus_NT.png/260px-Deinocheirus_NT.png 2x" data-file-width="753" data-file-height="382" /></a></span> </p> <span typeof="mw:File"><a href="/wiki/File:Qiupalong_Restoration.png" class="mw-file-description" title="Qiupalong henanensis"><img alt="Qiupalong henanensis" src="//upload.wikimedia.org/wikipedia/commons/thumb/5/57/Qiupalong_Restoration.png/130px-Qiupalong_Restoration.png" decoding="async" width="130" height="71" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/5/57/Qiupalong_Restoration.png/195px-Qiupalong_Restoration.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/5/57/Qiupalong_Restoration.png/260px-Qiupalong_Restoration.png 2x" data-file-width="5177" data-file-height="2829" /></a></span></div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Phuwiangvenator</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q63981558" class="extiw" title="wikidata:Q63981558">Q63981558</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/10357221">10357221</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=387579">387579</a></span></span></li></ul> </div></td></tr></tbody></table></div> <!-- NewPP limit report Parsed by mw‐web.eqiad.main‐5dc468848‐hnpjs Cached time: 20241124121954 Cache expiry: 2592000 Reduced expiry: false Complications: [vary‐revision‐sha1, show‐toc] CPU time usage: 1.185 seconds Real time usage: 1.478 seconds Preprocessor visited node count: 20108/1000000 Post‐expand include size: 753859/2097152 bytes Template argument size: 22588/2097152 bytes Highest expansion depth: 30/100 Expensive parser function count: 3/500 Unstrip recursion depth: 1/20 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