CINXE.COM

<!DOCTYPE html> <html class="client-nojs vector-feature-language-in-header-enabled vector-feature-language-in-main-page-header-disabled vector-feature-sticky-header-disabled vector-feature-page-tools-pinned-disabled vector-feature-toc-pinned-clientpref-1 vector-feature-main-menu-pinned-disabled vector-feature-limited-width-clientpref-1 vector-feature-limited-width-content-enabled vector-feature-custom-font-size-clientpref-1 vector-feature-appearance-pinned-clientpref-1 vector-feature-night-mode-enabled skin-theme-clientpref-day vector-toc-available" lang="ja" dir="ltr"> <head> <meta charset="UTF-8"> <title>遺伝子導入 - Wikipedia</title> <script>(function(){var className="client-js vector-feature-language-in-header-enabled vector-feature-language-in-main-page-header-disabled vector-feature-sticky-header-disabled vector-feature-page-tools-pinned-disabled vector-feature-toc-pinned-clientpref-1 vector-feature-main-menu-pinned-disabled vector-feature-limited-width-clientpref-1 vector-feature-limited-width-content-enabled vector-feature-custom-font-size-clientpref-1 vector-feature-appearance-pinned-clientpref-1 vector-feature-night-mode-enabled skin-theme-clientpref-day vector-toc-available";var cookie=document.cookie.match(/(?:^|; )jawikimwclientpreferences=([^;]+)/);if(cookie){cookie[1].split('%2C').forEach(function(pref){className=className.replace(new RegExp('(^| )'+pref.replace(/-clientpref-\w+$|[^\w-]+/g,'')+'-clientpref-\\w+( |$)'),'$1'+pref+'$2');});}document.documentElement.className=className;}());RLCONF={"wgBreakFrames":false,"wgSeparatorTransformTable":["",""],"wgDigitTransformTable":["",""],"wgDefaultDateFormat":"ja", "wgMonthNames":["","1月","2月","3月","4月","5月","6月","7月","8月","9月","10月","11月","12月"],"wgRequestId":"b7de955d-f852-464b-8651-841d8aa56faa","wgCanonicalNamespace":"","wgCanonicalSpecialPageName":false,"wgNamespaceNumber":0,"wgPageName":"遺伝子導入","wgTitle":"遺伝子導入","wgCurRevisionId":102726399,"wgRevisionId":102726399,"wgArticleId":2398503,"wgIsArticle":true,"wgIsRedirect":false,"wgAction":"view","wgUserName":null,"wgUserGroups":["*"],"wgCategories":["遺伝子工学"],"wgPageViewLanguage":"ja","wgPageContentLanguage":"ja","wgPageContentModel":"wikitext","wgRelevantPageName":"遺伝子導入","wgRelevantArticleId":2398503,"wgIsProbablyEditable":true,"wgRelevantPageIsProbablyEditable":true,"wgRestrictionEdit":[],"wgRestrictionMove":[],"wgNoticeProject":"wikipedia","wgCiteReferencePreviewsActive":true,"wgMediaViewerOnClick":true,"wgMediaViewerEnabledByDefault":true,"wgPopupsFlags":0,"wgVisualEditor":{"pageLanguageCode":"ja","pageLanguageDir":"ltr", "pageVariantFallbacks":"ja"},"wgMFDisplayWikibaseDescriptions":{"search":true,"watchlist":true,"tagline":true,"nearby":true},"wgWMESchemaEditAttemptStepOversample":false,"wgWMEPageLength":4000,"wgRelatedArticlesCompat":[],"wgEditSubmitButtonLabelPublish":true,"wgULSPosition":"interlanguage","wgULSisCompactLinksEnabled":false,"wgVector2022LanguageInHeader":true,"wgULSisLanguageSelectorEmpty":false,"wgWikibaseItemId":"Q2305545","wgCheckUserClientHintsHeadersJsApi":["brands","architecture","bitness","fullVersionList","mobile","model","platform","platformVersion"],"GEHomepageSuggestedEditsEnableTopics":true,"wgGETopicsMatchModeEnabled":false,"wgGEStructuredTaskRejectionReasonTextInputEnabled":false,"wgGELevelingUpEnabledForUser":false};RLSTATE={"ext.gadget.PDFLinkIcon":"ready","ext.gadget.RedirectColor":"ready","ext.globalCssJs.user.styles":"ready","site.styles":"ready","user.styles":"ready","ext.globalCssJs.user":"ready","user":"ready","user.options":"loading","ext.cite.styles":"ready", "skins.vector.search.codex.styles":"ready","skins.vector.styles":"ready","skins.vector.icons":"ready","ext.wikimediamessages.styles":"ready","ext.visualEditor.desktopArticleTarget.noscript":"ready","ext.uls.interlanguage":"ready","wikibase.client.init":"ready","ext.wikimediaBadges":"ready"};RLPAGEMODULES=["ext.cite.ux-enhancements","site","mediawiki.page.ready","mediawiki.toc","skins.vector.js","ext.centralNotice.geoIP","ext.centralNotice.startUp","ext.gadget.NormalizeCharWidth","ext.gadget.suppressEnterAtSummaryBox","ext.gadget.checkSignature","ext.gadget.charinsert","ext.gadget.WikiMiniAtlas","ext.gadget.switcher","ext.gadget.protectionIndicator","ext.urlShortener.toolbar","ext.centralauth.centralautologin","ext.popups","ext.visualEditor.desktopArticleTarget.init","ext.visualEditor.targetLoader","ext.echo.centralauth","ext.eventLogging","ext.wikimediaEvents","ext.navigationTiming","ext.uls.interface","ext.cx.eventlogging.campaigns","ext.cx.uls.quick.actions", "wikibase.client.vector-2022","ext.checkUser.clientHints","ext.quicksurveys.init","ext.growthExperiments.SuggestedEditSession","wikibase.sidebar.tracking"];</script> <script>(RLQ=window.RLQ||[]).push(function(){mw.loader.impl(function(){return["user.options@12s5i",function($,jQuery,require,module){mw.user.tokens.set({"patrolToken":"+\\","watchToken":"+\\","csrfToken":"+\\"}); }];});});</script> <link rel="stylesheet" href="/w/load.php?lang=ja&modules=ext.cite.styles%7Cext.uls.interlanguage%7Cext.visualEditor.desktopArticleTarget.noscript%7Cext.wikimediaBadges%7Cext.wikimediamessages.styles%7Cskins.vector.icons%2Cstyles%7Cskins.vector.search.codex.styles%7Cwikibase.client.init&only=styles&skin=vector-2022"> <script async="" src="/w/load.php?lang=ja&modules=startup&only=scripts&raw=1&skin=vector-2022"></script> <meta name="ResourceLoaderDynamicStyles" content=""> <link rel="stylesheet" href="/w/load.php?lang=ja&modules=ext.gadget.PDFLinkIcon%2CRedirectColor&only=styles&skin=vector-2022"> <link rel="stylesheet" href="/w/load.php?lang=ja&modules=site.styles&only=styles&skin=vector-2022"> <meta name="generator" content="MediaWiki 1.44.0-wmf.5"> <meta name="referrer" content="origin"> <meta name="referrer" content="origin-when-cross-origin"> <meta name="robots" content="max-image-preview:standard"> <meta name="format-detection" content="telephone=no"> <meta name="viewport" content="width=1120"> <meta property="og:title" content="遺伝子導入 - Wikipedia"> <meta property="og:type" content="website"> <link rel="alternate" media="only screen and (max-width: 640px)" href="//ja.m.wikipedia.org/wiki/%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5"> <link rel="alternate" type="application/x-wiki" title="編集" href="/w/index.php?title=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5&action=edit"> <link rel="apple-touch-icon" href="/static/apple-touch/wikipedia.png"> <link rel="icon" href="/static/favicon/wikipedia.ico"> <link rel="search" type="application/opensearchdescription+xml" href="/w/rest.php/v1/search" title="Wikipedia (ja)"> <link rel="EditURI" type="application/rsd+xml" href="//ja.wikipedia.org/w/api.php?action=rsd"> <link rel="canonical" href="https://ja.wikipedia.org/wiki/%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5"> <link rel="license" href="https://creativecommons.org/licenses/by-sa/4.0/deed.ja"> <link rel="alternate" type="application/atom+xml" title="WikipediaのAtomフィード" href="/w/index.php?title=%E7%89%B9%E5%88%A5:%E6%9C%80%E8%BF%91%E3%81%AE%E6%9B%B4%E6%96%B0&feed=atom"> <link rel="dns-prefetch" href="//meta.wikimedia.org" /> <link rel="dns-prefetch" href="//login.wikimedia.org"> </head> <body class="skin--responsive skin-vector skin-vector-search-vue mediawiki ltr sitedir-ltr mw-hide-empty-elt ns-0 ns-subject mw-editable page-遺伝子導入 rootpage-遺伝子導入 skin-vector-2022 action-view"><a class="mw-jump-link" href="#bodyContent">コンテンツにスキップ</a> <div class="vector-header-container"> <header class="vector-header mw-header"> <div class="vector-header-start"> <nav class="vector-main-menu-landmark" aria-label="サイト"> <div id="vector-main-menu-dropdown" class="vector-dropdown vector-main-menu-dropdown vector-button-flush-left vector-button-flush-right" > <input type="checkbox" id="vector-main-menu-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-main-menu-dropdown" class="vector-dropdown-checkbox " aria-label="メインメニュー" > <label id="vector-main-menu-dropdown-label" for="vector-main-menu-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-menu mw-ui-icon-wikimedia-menu"></span> <span class="vector-dropdown-label-text">メインメニュー</span> </label> <div class="vector-dropdown-content"> <div id="vector-main-menu-unpinned-container" class="vector-unpinned-container"> <div id="vector-main-menu" class="vector-main-menu vector-pinnable-element"> <div class="vector-pinnable-header vector-main-menu-pinnable-header vector-pinnable-header-unpinned" data-feature-name="main-menu-pinned" data-pinnable-element-id="vector-main-menu" data-pinned-container-id="vector-main-menu-pinned-container" data-unpinned-container-id="vector-main-menu-unpinned-container" > <div class="vector-pinnable-header-label">メインメニュー</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-main-menu.pin">サイドバーに移動</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-main-menu.unpin">非表示</button> </div> <div id="p-navigation" class="vector-menu mw-portlet mw-portlet-navigation" > <div class="vector-menu-heading"> 案内 </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="n-mainpage-description" class="mw-list-item"><a href="/wiki/%E3%83%A1%E3%82%A4%E3%83%B3%E3%83%9A%E3%83%BC%E3%82%B8" title="メインページに移動する [z]" accesskey="z"><span>メインページ</span></a></li><li id="n-portal" class="mw-list-item"><a href="/wiki/Wikipedia:%E3%82%B3%E3%83%9F%E3%83%A5%E3%83%8B%E3%83%86%E3%82%A3%E3%83%BB%E3%83%9D%E3%83%BC%E3%82%BF%E3%83%AB" title="このプロジェクトについて、できること、情報を入手する場所"><span>コミュニティ・ポータル</span></a></li><li id="n-currentevents" class="mw-list-item"><a href="/wiki/Portal:%E6%9C%80%E8%BF%91%E3%81%AE%E5%87%BA%E6%9D%A5%E4%BA%8B" title="最近の出来事の背景を知る"><span>最近の出来事</span></a></li><li id="n-newpages" class="mw-list-item"><a href="/wiki/%E7%89%B9%E5%88%A5:%E6%96%B0%E3%81%97%E3%81%84%E3%83%9A%E3%83%BC%E3%82%B8" title="最近新規に作成されたページの一覧"><span>新しいページ</span></a></li><li id="n-recentchanges" class="mw-list-item"><a href="/wiki/%E7%89%B9%E5%88%A5:%E6%9C%80%E8%BF%91%E3%81%AE%E6%9B%B4%E6%96%B0" title="このウィキにおける最近の更新の一覧 [r]" accesskey="r"><span>最近の更新</span></a></li><li id="n-randompage" class="mw-list-item"><a href="/wiki/%E7%89%B9%E5%88%A5:%E3%81%8A%E3%81%BE%E3%81%8B%E3%81%9B%E8%A1%A8%E7%A4%BA" title="無作為に選択されたページを読み込む [x]" accesskey="x"><span>おまかせ表示</span></a></li><li id="n-sandbox" class="mw-list-item"><a href="/wiki/Wikipedia:%E3%82%B5%E3%83%B3%E3%83%89%E3%83%9C%E3%83%83%E3%82%AF%E3%82%B9" title="練習用のページ"><span>練習用ページ</span></a></li><li id="n-commonsupload" class="mw-list-item"><a href="//commons.wikimedia.org/wiki/Special:UploadWizard?uselang=ja" title="画像やメディアファイルをウィキメディア・コモンズにアップロード"><span>アップロード (ウィキメディア・コモンズ)</span></a></li> </ul> </div> </div> <div id="p-help" class="vector-menu mw-portlet mw-portlet-help" > <div class="vector-menu-heading"> ヘルプ </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="n-help" class="mw-list-item"><a href="/wiki/Help:%E7%9B%AE%E6%AC%A1" title="情報を得る場所"><span>ヘルプ</span></a></li><li id="n-villagepump" class="mw-list-item"><a href="/wiki/Wikipedia:%E4%BA%95%E6%88%B8%E7%AB%AF" title="プロジェクトについての意見交換"><span>井戸端</span></a></li><li id="n-notice" class="mw-list-item"><a href="/wiki/Wikipedia:%E3%81%8A%E7%9F%A5%E3%82%89%E3%81%9B" title="プロジェクトについてのお知らせ"><span>お知らせ</span></a></li><li id="n-bugreportspage" class="mw-list-item"><a href="/wiki/Wikipedia:%E3%83%90%E3%82%B0%E3%81%AE%E5%A0%B1%E5%91%8A" title="ウィキペディア・ソフトウェアのバグ報告"><span>バグの報告</span></a></li><li id="n-contact" class="mw-list-item"><a href="/wiki/Wikipedia:%E9%80%A3%E7%B5%A1%E5%85%88" title="ウィキペディアやウィキメディア財団に関する連絡先"><span>ウィキペディアに関するお問い合わせ</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> <a href="/wiki/%E3%83%A1%E3%82%A4%E3%83%B3%E3%83%9A%E3%83%BC%E3%82%B8" class="mw-logo"> <img class="mw-logo-icon" src="/static/images/icons/wikipedia.png" alt="" aria-hidden="true" height="50" width="50"> <span class="mw-logo-container skin-invert"> <img class="mw-logo-wordmark" alt="Wikipedia" src="/static/images/mobile/copyright/wikipedia-wordmark-ja.svg" style="width: 7.5em; height: 1.25em;"> <img class="mw-logo-tagline" alt="" src="/static/images/mobile/copyright/wikipedia-tagline-ja.svg" width="100" height="14" style="width: 6.25em; height: 0.875em;"> </span> </a> </div> <div class="vector-header-end"> <div id="p-search" role="search" class="vector-search-box-vue vector-search-box-collapses vector-search-box-show-thumbnail vector-search-box-auto-expand-width vector-search-box"> <a href="/wiki/%E7%89%B9%E5%88%A5:%E6%A4%9C%E7%B4%A2" class="cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only search-toggle" title="Wikipedia内を検索 [f]" accesskey="f"><span class="vector-icon mw-ui-icon-search mw-ui-icon-wikimedia-search"></span> <span>検索</span> </a> <div class="vector-typeahead-search-container"> <div class="cdx-typeahead-search cdx-typeahead-search--show-thumbnail cdx-typeahead-search--auto-expand-width"> <form action="/w/index.php" id="searchform" class="cdx-search-input cdx-search-input--has-end-button"> <div id="simpleSearch" class="cdx-search-input__input-wrapper" data-search-loc="header-moved"> <div class="cdx-text-input cdx-text-input--has-start-icon"> <input class="cdx-text-input__input" type="search" name="search" placeholder="Wikipedia内を検索" aria-label="Wikipedia内を検索" autocapitalize="sentences" title="Wikipedia内を検索 [f]" accesskey="f" id="searchInput" > <span class="cdx-text-input__icon cdx-text-input__start-icon"></span> </div> <input type="hidden" name="title" value="特別:検索"> </div> <button class="cdx-button cdx-search-input__end-button">検索</button> </form> </div> </div> </div> <nav class="vector-user-links vector-user-links-wide" aria-label="個人用ツール"> <div class="vector-user-links-main"> <div id="p-vector-user-menu-preferences" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <div id="p-vector-user-menu-userpage" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <nav class="vector-appearance-landmark" aria-label="表示"> <div id="vector-appearance-dropdown" class="vector-dropdown " title="ページのフォントサイズ、幅、色の外観を変更する" > <input type="checkbox" id="vector-appearance-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-appearance-dropdown" class="vector-dropdown-checkbox " aria-label="表示" > <label id="vector-appearance-dropdown-label" for="vector-appearance-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-appearance mw-ui-icon-wikimedia-appearance"></span> <span class="vector-dropdown-label-text">表示</span> </label> <div class="vector-dropdown-content"> <div id="vector-appearance-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <div id="p-vector-user-menu-notifications" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <div id="p-vector-user-menu-overflow" class="vector-menu mw-portlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-sitesupport-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="//donate.wikimedia.org/wiki/Special:FundraiserRedirector?utm_source=donate&utm_medium=sidebar&utm_campaign=C13_ja.wikipedia.org&uselang=ja" class=""><span>寄付</span></a> </li> <li id="pt-createaccount-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="/w/index.php?title=%E7%89%B9%E5%88%A5:%E3%82%A2%E3%82%AB%E3%82%A6%E3%83%B3%E3%83%88%E4%BD%9C%E6%88%90&returnto=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5" title="アカウントを作成してログインすることをお勧めしますが、必須ではありません" class=""><span>アカウント作成</span></a> </li> <li id="pt-login-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="/w/index.php?title=%E7%89%B9%E5%88%A5:%E3%83%AD%E3%82%B0%E3%82%A4%E3%83%B3&returnto=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5" title="ログインすることを推奨します。ただし、必須ではありません。 [o]" accesskey="o" class=""><span>ログイン</span></a> </li> </ul> </div> </div> </div> <div id="vector-user-links-dropdown" class="vector-dropdown vector-user-menu vector-button-flush-right vector-user-menu-logged-out" title="その他の操作" > <input type="checkbox" id="vector-user-links-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-user-links-dropdown" class="vector-dropdown-checkbox " aria-label="個人用ツール" > <label id="vector-user-links-dropdown-label" for="vector-user-links-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-ellipsis mw-ui-icon-wikimedia-ellipsis"></span> <span class="vector-dropdown-label-text">個人用ツール</span> </label> <div class="vector-dropdown-content"> <div id="p-personal" class="vector-menu mw-portlet mw-portlet-personal user-links-collapsible-item" title="利用者メニュー" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-sitesupport" class="user-links-collapsible-item mw-list-item"><a href="//donate.wikimedia.org/wiki/Special:FundraiserRedirector?utm_source=donate&utm_medium=sidebar&utm_campaign=C13_ja.wikipedia.org&uselang=ja"><span>寄付</span></a></li><li id="pt-createaccount" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=%E7%89%B9%E5%88%A5:%E3%82%A2%E3%82%AB%E3%82%A6%E3%83%B3%E3%83%88%E4%BD%9C%E6%88%90&returnto=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5" title="アカウントを作成してログインすることをお勧めしますが、必須ではありません"><span class="vector-icon mw-ui-icon-userAdd mw-ui-icon-wikimedia-userAdd"></span> <span>アカウント作成</span></a></li><li id="pt-login" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=%E7%89%B9%E5%88%A5:%E3%83%AD%E3%82%B0%E3%82%A4%E3%83%B3&returnto=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5" title="ログインすることを推奨します。ただし、必須ではありません。 [o]" accesskey="o"><span class="vector-icon mw-ui-icon-logIn mw-ui-icon-wikimedia-logIn"></span> <span>ログイン</span></a></li> </ul> </div> </div> <div id="p-user-menu-anon-editor" class="vector-menu mw-portlet mw-portlet-user-menu-anon-editor" > <div class="vector-menu-heading"> ログアウトした編集者のページ <a href="/wiki/Wikipedia:%E3%82%A6%E3%82%A3%E3%82%AD%E3%83%9A%E3%83%87%E3%82%A3%E3%82%A2%E3%81%B8%E3%82%88%E3%81%86%E3%81%93%E3%81%9D" aria-label="編集の詳細"><span>もっと詳しく</span></a> </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-anoncontribs" class="mw-list-item"><a href="/wiki/%E7%89%B9%E5%88%A5:%E8%87%AA%E5%88%86%E3%81%AE%E6%8A%95%E7%A8%BF%E8%A8%98%E9%8C%B2" title="このIPアドレスからなされた編集の一覧 [y]" accesskey="y"><span>投稿記録</span></a></li><li id="pt-anontalk" class="mw-list-item"><a href="/wiki/%E7%89%B9%E5%88%A5:%E3%83%88%E3%83%BC%E3%82%AF%E3%83%9A%E3%83%BC%E3%82%B8" title="このIPアドレスからなされた編集についての議論 [n]" accesskey="n"><span>トーク</span></a></li> </ul> </div> </div> </div> </div> </nav> </div> </header> </div> <div class="mw-page-container"> <div class="mw-page-container-inner"> <div class="vector-sitenotice-container"> <div id="siteNotice"></div> </div> <div class="vector-column-start"> <div class="vector-main-menu-container"> <div id="mw-navigation"> <nav id="mw-panel" class="vector-main-menu-landmark" aria-label="サイト"> <div id="vector-main-menu-pinned-container" class="vector-pinned-container"> </div> </nav> </div> </div> <div class="vector-sticky-pinned-container"> <nav id="mw-panel-toc" aria-label="目次" data-event-name="ui.sidebar-toc" class="mw-table-of-contents-container vector-toc-landmark"> <div id="vector-toc-pinned-container" class="vector-pinned-container"> <div id="vector-toc" class="vector-toc vector-pinnable-element"> <div class="vector-pinnable-header vector-toc-pinnable-header vector-pinnable-header-pinned" data-feature-name="toc-pinned" data-pinnable-element-id="vector-toc" > <h2 class="vector-pinnable-header-label">目次</h2> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-toc.pin">サイドバーに移動</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-toc.unpin">非表示</button> </div> <ul class="vector-toc-contents" id="mw-panel-toc-list"> <li id="toc-mw-content-text" class="vector-toc-list-item vector-toc-level-1"> <a href="#" class="vector-toc-link"> <div class="vector-toc-text">ページ先頭</div> </a> </li> <li id="toc-遺伝子導入の手段" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#遺伝子導入の手段"> <div class="vector-toc-text"> <span class="vector-toc-numb">1</span> <span>遺伝子導入の手段</span> </div> </a> <ul id="toc-遺伝子導入の手段-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-遺伝子導入の方法" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#遺伝子導入の方法"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>遺伝子導入の方法</span> </div> </a> <ul id="toc-遺伝子導入の方法-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-遺伝子導入の応用" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#遺伝子導入の応用"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>遺伝子導入の応用</span> </div> </a> <ul id="toc-遺伝子導入の応用-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-脚注・出典" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#脚注・出典"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>脚注・出典</span> </div> </a> <ul id="toc-脚注・出典-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-外部リンク" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#外部リンク"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>外部リンク</span> </div> </a> <ul id="toc-外部リンク-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="目次" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="目次の表示・非表示を切り替え" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">目次の表示・非表示を切り替え</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">遺伝子導入</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="特定の記事の別の言語版に移動します。利用可能な言語13件" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-13" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">13の言語版</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D9%86%D9%82%D9%84_%D9%88%D8%B1%D8%A7%D8%AB%D9%8A" title="アラビア語: نقل وراثي" lang="ar" hreflang="ar" data-title="نقل وراثي" data-language-autonym="العربية" data-language-local-name="アラビア語" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Transg%C3%A8nesi" title="カタロニア語: Transgènesi" lang="ca" hreflang="ca" data-title="Transgènesi" data-language-autonym="Català" data-language-local-name="カタロニア語" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-en badge-Q70893996 mw-list-item" title=""><a href="https://en.wikipedia.org/wiki/Transgenesis" title="英語: Transgenesis" lang="en" hreflang="en" data-title="Transgenesis" data-language-autonym="English" data-language-local-name="英語" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Transg%C3%A9nesis" title="スペイン語: Transgénesis" lang="es" hreflang="es" data-title="Transgénesis" data-language-autonym="Español" data-language-local-name="スペイン語" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%AA%D8%B1%D8%A7%D8%B1%DB%8C%D8%AE%D8%AA%D9%87" title="ペルシア語: تراریخته" lang="fa" hreflang="fa" data-title="تراریخته" data-language-autonym="فارسی" data-language-local-name="ペルシア語" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Transg%C3%A9n%C3%A8se" title="フランス語: Transgénèse" lang="fr" hreflang="fr" data-title="Transgénèse" data-language-autonym="Français" data-language-local-name="フランス語" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Trasgh%C3%A9ineach" title="アイルランド語: Trasghéineach" lang="ga" hreflang="ga" data-title="Trasghéineach" data-language-autonym="Gaeilge" data-language-local-name="アイルランド語" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Transgenesi" title="イタリア語: Transgenesi" lang="it" hreflang="it" data-title="Transgenesi" data-language-autonym="Italiano" data-language-local-name="イタリア語" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Transgenez%C4%97" title="リトアニア語: Transgenezė" lang="lt" hreflang="lt" data-title="Transgenezė" data-language-autonym="Lietuvių" data-language-local-name="リトアニア語" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Transgen%C3%A8si" title="オック語: Transgenèsi" lang="oc" hreflang="oc" data-title="Transgenèsi" data-language-autonym="Occitan" data-language-local-name="オック語" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Transg%C3%AAnese" title="ポルトガル語: Transgênese" lang="pt" hreflang="pt" data-title="Transgênese" data-language-autonym="Português" data-language-local-name="ポルトガル語" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%A2%D1%80%D0%B0%D0%BD%D1%81%D0%B3%D0%B5%D0%BD%D0%B5%D0%B7" title="ロシア語: Трансгенез" lang="ru" hreflang="ru" data-title="Трансгенез" data-language-autonym="Русский" data-language-local-name="ロシア語" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-scn mw-list-item"><a href="https://scn.wikipedia.org/wiki/Transg%C3%A8nisi" title="シチリア語: Transgènisi" lang="scn" hreflang="scn" data-title="Transgènisi" data-language-autonym="Sicilianu" data-language-local-name="シチリア語" class="interlanguage-link-target"><span>Sicilianu</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q2305545#sitelinks-wikipedia" title="言語間リンクを編集" class="wbc-editpage">リンクを編集</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="名前空間"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5" title="本文を閲覧 [c]" accesskey="c"><span>ページ</span></a></li><li id="ca-talk" class="new vector-tab-noicon mw-list-item"><a href="/w/index.php?title=%E3%83%8E%E3%83%BC%E3%83%88:%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5&action=edit&redlink=1" rel="discussion" class="new" title="「本文ページについての議論」 (存在しないページ) [t]" accesskey="t"><span>ノート</span></a></li> </ul> </div> </div> <div id="vector-variants-dropdown" class="vector-dropdown emptyPortlet" > <input type="checkbox" id="vector-variants-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-variants-dropdown" class="vector-dropdown-checkbox " aria-label="別の言語に切り替える" > <label id="vector-variants-dropdown-label" for="vector-variants-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">日本語</span> </label> <div class="vector-dropdown-content"> <div id="p-variants" class="vector-menu mw-portlet mw-portlet-variants emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> </div> </div> </nav> </div> <div id="right-navigation" class="vector-collapsible"> <nav aria-label="表示"> <div id="p-views" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-views" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-view" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5"><span>閲覧</span></a></li><li id="ca-edit" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5&action=edit" title="このページのソースコードを編集する [e]" accesskey="e"><span>編集</span></a></li><li id="ca-history" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5&action=history" title="このページの過去の版 [h]" accesskey="h"><span>履歴表示</span></a></li> </ul> </div> </div> </nav> <nav class="vector-page-tools-landmark" aria-label="ページツール"> <div id="vector-page-tools-dropdown" class="vector-dropdown vector-page-tools-dropdown" > <input type="checkbox" id="vector-page-tools-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-tools-dropdown" class="vector-dropdown-checkbox " aria-label="ツール" > <label id="vector-page-tools-dropdown-label" for="vector-page-tools-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">ツール</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-tools-unpinned-container" class="vector-unpinned-container"> <div id="vector-page-tools" class="vector-page-tools vector-pinnable-element"> <div class="vector-pinnable-header vector-page-tools-pinnable-header vector-pinnable-header-unpinned" data-feature-name="page-tools-pinned" data-pinnable-element-id="vector-page-tools" data-pinned-container-id="vector-page-tools-pinned-container" data-unpinned-container-id="vector-page-tools-unpinned-container" > <div class="vector-pinnable-header-label">ツール</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-page-tools.pin">サイドバーに移動</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-page-tools.unpin">非表示</button> </div> <div id="p-cactions" class="vector-menu mw-portlet mw-portlet-cactions emptyPortlet vector-has-collapsible-items" title="その他の操作" > <div class="vector-menu-heading"> 操作 </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-more-view" class="selected vector-more-collapsible-item mw-list-item"><a href="/wiki/%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5"><span>閲覧</span></a></li><li id="ca-more-edit" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5&action=edit" title="このページのソースコードを編集する [e]" accesskey="e"><span>編集</span></a></li><li id="ca-more-history" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5&action=history"><span>履歴表示</span></a></li> </ul> </div> </div> <div id="p-tb" class="vector-menu mw-portlet mw-portlet-tb" > <div class="vector-menu-heading"> 全般 </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="t-whatlinkshere" class="mw-list-item"><a href="/wiki/%E7%89%B9%E5%88%A5:%E3%83%AA%E3%83%B3%E3%82%AF%E5%85%83/%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5" title="ここにリンクしている全ウィキページの一覧 [j]" accesskey="j"><span>リンク元</span></a></li><li id="t-recentchangeslinked" class="mw-list-item"><a href="/wiki/%E7%89%B9%E5%88%A5:%E9%96%A2%E9%80%A3%E3%83%9A%E3%83%BC%E3%82%B8%E3%81%AE%E6%9B%B4%E6%96%B0%E7%8A%B6%E6%B3%81/%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5" rel="nofollow" title="このページからリンクしているページの最近の更新 [k]" accesskey="k"><span>関連ページの更新状況</span></a></li><li id="t-upload" class="mw-list-item"><a href="/wiki/Wikipedia:%E3%83%95%E3%82%A1%E3%82%A4%E3%83%AB%E3%81%AE%E3%82%A2%E3%83%83%E3%83%97%E3%83%AD%E3%83%BC%E3%83%89" title="ファイルをアップロードする [u]" accesskey="u"><span>ファイルをアップロード</span></a></li><li id="t-specialpages" class="mw-list-item"><a href="/wiki/%E7%89%B9%E5%88%A5:%E7%89%B9%E5%88%A5%E3%83%9A%E3%83%BC%E3%82%B8%E4%B8%80%E8%A6%A7" title="特別ページの一覧 [q]" accesskey="q"><span>特別ページ</span></a></li><li id="t-permalink" class="mw-list-item"><a href="/w/index.php?title=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5&oldid=102726399" title="このページのこの版への固定リンク"><span>この版への固定リンク</span></a></li><li id="t-info" class="mw-list-item"><a href="/w/index.php?title=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5&action=info" title="このページについての詳細情報"><span>ページ情報</span></a></li><li id="t-cite" class="mw-list-item"><a href="/w/index.php?title=%E7%89%B9%E5%88%A5:%E3%81%93%E3%81%AE%E3%83%9A%E3%83%BC%E3%82%B8%E3%82%92%E5%BC%95%E7%94%A8&page=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5&id=102726399&wpFormIdentifier=titleform" title="このページの引用方法"><span>このページを引用</span></a></li><li id="t-urlshortener" class="mw-list-item"><a href="/w/index.php?title=%E7%89%B9%E5%88%A5:UrlShortener&url=https%3A%2F%2Fja.wikipedia.org%2Fwiki%2F%25E9%2581%25BA%25E4%25BC%259D%25E5%25AD%2590%25E5%25B0%258E%25E5%2585%25A5"><span>短縮URLを取得する</span></a></li><li id="t-urlshortener-qrcode" class="mw-list-item"><a href="/w/index.php?title=%E7%89%B9%E5%88%A5:QrCode&url=https%3A%2F%2Fja.wikipedia.org%2Fwiki%2F%25E9%2581%25BA%25E4%25BC%259D%25E5%25AD%2590%25E5%25B0%258E%25E5%2585%25A5"><span>QRコードをダウンロード</span></a></li> </ul> </div> </div> <div id="p-coll-print_export" class="vector-menu mw-portlet mw-portlet-coll-print_export" > <div class="vector-menu-heading"> 印刷/書き出し </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="coll-create_a_book" class="mw-list-item"><a href="/w/index.php?title=%E7%89%B9%E5%88%A5:%E3%83%96%E3%83%83%E3%82%AF&bookcmd=book_creator&referer=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5"><span>ブックの新規作成</span></a></li><li id="coll-download-as-rl" class="mw-list-item"><a href="/w/index.php?title=%E7%89%B9%E5%88%A5:DownloadAsPdf&page=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5&action=show-download-screen"><span>PDF 形式でダウンロード</span></a></li><li id="t-print" class="mw-list-item"><a href="/w/index.php?title=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5&printable=yes" title="このページの印刷用ページ [p]" accesskey="p"><span>印刷用バージョン</span></a></li> </ul> </div> </div> <div id="p-wikibase-otherprojects" class="vector-menu mw-portlet mw-portlet-wikibase-otherprojects" > <div class="vector-menu-heading"> 他のプロジェクト </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="t-wikibase" class="wb-otherproject-link wb-otherproject-wikibase-dataitem mw-list-item"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q2305545" title="関連付けられたデータリポジトリ項目へのリンク [g]" accesskey="g"><span>ウィキデータ項目</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> </div> </div> </div> <div class="vector-column-end"> <div class="vector-sticky-pinned-container"> <nav class="vector-page-tools-landmark" aria-label="ページツール"> <div id="vector-page-tools-pinned-container" class="vector-pinned-container"> </div> </nav> <nav class="vector-appearance-landmark" aria-label="表示"> <div id="vector-appearance-pinned-container" class="vector-pinned-container"> <div id="vector-appearance" class="vector-appearance vector-pinnable-element"> <div class="vector-pinnable-header vector-appearance-pinnable-header vector-pinnable-header-pinned" data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">表示</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">サイドバーに移動</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">非表示</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">出典: フリー百科事典『ウィキペディア（Wikipedia）』</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="ja" dir="ltr"><p><b>遺伝子導入</b>（いでんしどうにゅう、transgenesis）は、外来<a href="/wiki/%E9%81%BA%E4%BC%9D%E5%AD%90" title="遺伝子">遺伝子</a>を<a href="/wiki/%E7%94%9F%E7%89%A9" title="生物">生物</a>に導入する過程を指し、外来遺伝子の導入によって、その生物の<a href="/wiki/%E5%BD%A2%E8%B3%AA" title="形質">形質</a>が変わり、新たな能力を獲得する場合がある。 </p><p><a href="/wiki/%E7%B4%B0%E8%8F%8C" title="細菌">細菌</a>、<a href="/wiki/%E9%85%B5%E6%AF%8D" title="酵母">酵母</a>や<a href="/wiki/%E6%A4%8D%E7%89%A9" title="植物">植物</a><a href="/wiki/%E7%B4%B0%E8%83%9E" title="細胞">細胞</a>への遺伝子導入は<b><a href="/wiki/%E5%BD%A2%E8%B3%AA%E8%BB%A2%E6%8F%9B" title="形質転換">形質転換</a></b>（transformation）、<a href="/wiki/%E5%8B%95%E7%89%A9" title="動物">動物</a>細胞への遺伝子導入は<b><a href="/wiki/%E3%83%88%E3%83%A9%E3%83%B3%E3%82%B9%E3%83%95%E3%82%A7%E3%82%AF%E3%82%B7%E3%83%A7%E3%83%B3" title="トランスフェクション">トランスフェクション</a></b>（transfection）、<a href="/wiki/%E3%83%95%E3%82%A1%E3%83%BC%E3%82%B8" title="ファージ">ファージ</a>や<a href="/wiki/%E3%82%A6%E3%82%A4%E3%83%AB%E3%82%B9" title="ウイルス">ウイルス</a>を用いた遺伝子導入は<b><a href="/wiki/%E5%BD%A2%E8%B3%AA%E5%B0%8E%E5%85%A5" title="形質導入">形質導入</a></b>（transduction）と一般的に呼ばれる。 </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="遺伝子導入の手段"><span id=".E9.81.BA.E4.BC.9D.E5.AD.90.E5.B0.8E.E5.85.A5.E3.81.AE.E6.89.8B.E6.AE.B5"></span>遺伝子導入の手段</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5&action=edit&section=1" title="節を編集: 遺伝子導入の手段"><span>編集</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>遺伝子導入を促進する生物学的、化学的または物理的手法があり<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>、<a href="/wiki/%E7%B4%B0%E8%83%9E%E6%A0%AA" class="mw-redirect" title="細胞株">細胞株</a>、初代培養系、動物、植物や細菌への遺伝子導入に利用されている。生物学的手法にはウイルスを用いた方法が広く行われている<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup>。化学的方法には<a href="/wiki/%E3%83%AA%E3%83%9D%E3%83%95%E3%82%A7%E3%82%AF%E3%82%B7%E3%83%A7%E3%83%B3" title="リポフェクション">リポフェクション</a>法やポリマーやペプチドを使用する方法<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup>等がある。物理的手法には<a href="/wiki/%E9%9B%BB%E6%B0%97%E7%A9%BF%E5%AD%94%E6%B3%95" title="電気穿孔法">電気穿孔法</a>や<a href="/wiki/%E3%83%91%E3%83%BC%E3%83%86%E3%82%A3%E3%82%AF%E3%83%AB%E3%83%BB%E3%82%AC%E3%83%B3%E6%B3%95" title="パーティクル・ガン法">パーティクル・ガン法</a>や超音波を使用する方法（sonoporation）<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup>等がある。また細菌の場合、<a href="/wiki/%E5%A1%A9%E5%8C%96%E3%82%AB%E3%83%AB%E3%82%B7%E3%82%A6%E3%83%A0" title="塩化カルシウム">塩化カルシウム</a>存在下で<a href="/wiki/%E3%82%B3%E3%83%B3%E3%83%94%E3%83%86%E3%83%B3%E3%83%88%E3%82%BB%E3%83%AB" title="コンピテントセル">コンピテントセル</a>化した菌にヒートショックで <a href="/wiki/DNA" class="mw-redirect" title="DNA">DNA</a> を形質転換する方法が広く行われている。 </p> <div class="mw-heading mw-heading2"><h2 id="遺伝子導入の方法"><span id=".E9.81.BA.E4.BC.9D.E5.AD.90.E5.B0.8E.E5.85.A5.E3.81.AE.E6.96.B9.E6.B3.95"></span>遺伝子導入の方法</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5&action=edit&section=2" title="節を編集: 遺伝子導入の方法"><span>編集</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>遺伝子導入が一過的なものか<a href="/wiki/%E3%82%B2%E3%83%8E%E3%83%A0" title="ゲノム">ゲノム</a>内に組み込む安定的なものかの方法の違いがあり<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup>、<a href="/wiki/%E3%83%88%E3%83%A9%E3%83%B3%E3%82%B9%E3%83%9D%E3%82%BE%E3%83%B3" title="トランスポゾン">トランスポゾン</a>を用いてゲノムに挿入する方法<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup>やリコンビナーゼを用いて特定のゲノムの位置に挿入する方法<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup>がある。 </p> <div class="mw-heading mw-heading2"><h2 id="遺伝子導入の応用"><span id=".E9.81.BA.E4.BC.9D.E5.AD.90.E5.B0.8E.E5.85.A5.E3.81.AE.E5.BF.9C.E7.94.A8"></span>遺伝子導入の応用</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5&action=edit&section=3" title="節を編集: 遺伝子導入の応用"><span>編集</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>遺伝子導入は<a href="/wiki/%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B7%A5%E5%AD%A6" title="遺伝子工学">遺伝子工学</a>の<a href="/wiki/%E5%9F%BA%E7%A4%8E" title="基礎">基礎</a>となる技術である。この技術は<a href="/wiki/%E9%81%BA%E4%BC%9D%E5%AD%90%E7%B5%84%E3%81%BF%E6%8F%9B%E3%81%88%E4%BD%9C%E7%89%A9" title="遺伝子組み換え作物">遺伝子組み換え作物</a>や<a href="/wiki/%E9%81%BA%E4%BC%9D%E5%AD%90%E6%B2%BB%E7%99%82" title="遺伝子治療">遺伝子治療</a>等に応用されている。また遺伝子導入は<a href="/wiki/%E3%82%B2%E3%83%8E%E3%83%A0%E8%A7%A3%E6%9E%90" class="mw-redirect" title="ゲノム解析">ゲノム解析</a>、<a href="/wiki/%E9%81%BA%E4%BC%9D%E5%AD%90%E7%A0%B4%E5%A3%8A" title="遺伝子破壊">遺伝子破壊</a>や<a href="/wiki/%E9%81%BA%E4%BC%9D%E5%AD%90%E3%83%8E%E3%83%83%E3%82%AF%E3%83%80%E3%82%A6%E3%83%B3" title="遺伝子ノックダウン">遺伝子ノックダウン</a>技術の基礎になっており、<a href="/wiki/%E4%BA%BA%E5%B7%A5%E5%A4%9A%E8%83%BD%E6%80%A7%E5%B9%B9%E7%B4%B0%E8%83%9E" title="人工多能性幹細胞">iPS 細胞</a>作成に必要な基礎技術である。 </p> <div class="mw-heading mw-heading2"><h2 id="脚注・出典"><span id=".E8.84.9A.E6.B3.A8.E3.83.BB.E5.87.BA.E5.85.B8"></span>脚注・出典</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5&action=edit&section=4" title="節を編集: 脚注・出典"><span>編集</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="noprint" style="float:right; font-size:90%;">[<a href="/wiki/Help:%E8%84%9A%E6%B3%A8/%E8%AA%AD%E8%80%85%E5%90%91%E3%81%91" title="Help:脚注/読者向け"><span title="この欄の操作法">脚注の使い方</span></a>]</div> <ol class="references"> <li id="cite_note-1"><b><a href="#cite_ref-1">^</a></b> <span class="reference-text">Kim TK, Eberwine JH. "Mammalian cell transfection: the present and the future." <i>Anal Bioanal Chem.</i> 2010 Aug;397(8):3173-8. <style data-mw-deduplicate="TemplateStyles:r101121245">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation.cs-ja1 q,.mw-parser-output .citation.cs-ja2 q{quotes:"「""」""『""』"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free a,.mw-parser-output .citation .cs1-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited a,.mw-parser-output .id-lock-registration a,.mw-parser-output .citation .cs1-lock-limited a,.mw-parser-output .citation .cs1-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription a,.mw-parser-output .citation .cs1-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:var(--color-success,#3a3);margin-left:0.3em}.mw-parser-output .cs1-format{font-size:95%}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}</style><a href="/wiki/PMID_(%E8%AD%98%E5%88%A5%E5%AD%90)" class="mw-redirect" title="PMID (識別子)">PMID</a> <a rel="nofollow" class="external text" href="//pubmed.ncbi.nlm.nih.gov/20549496/">20549496</a></span> </li> <li id="cite_note-2"><b><a href="#cite_ref-2">^</a></b> <span class="reference-text">Howarth JL, Lee YB, Uney JB. "Using viral vectors as gene transfer tools (Cell Biology and Toxicology Special Issue: ETCS-UK 1 day meeting on genetic manipulation of cells)." <i>Cell Biol Toxicol.</i> 2010 Feb;26(1):1-20. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r101121245"><a href="/wiki/PMID_(%E8%AD%98%E5%88%A5%E5%AD%90)" class="mw-redirect" title="PMID (識別子)">PMID</a> <a rel="nofollow" class="external text" href="//pubmed.ncbi.nlm.nih.gov/19830583/">19830583</a></span> </li> <li id="cite_note-3"><b><a href="#cite_ref-3">^</a></b> <span class="reference-text">Schaffer DV, Koerber JT, Lim KI. "Molecular engineering of viral gene delivery vehicles." <i>Annu Rev Biomed Eng.</i> 2008;10:169-94. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r101121245"><a href="/wiki/PMID_(%E8%AD%98%E5%88%A5%E5%AD%90)" class="mw-redirect" title="PMID (識別子)">PMID</a> <a rel="nofollow" class="external text" href="//pubmed.ncbi.nlm.nih.gov/18647114/">18647114</a></span> </li> <li id="cite_note-4"><b><a href="#cite_ref-4">^</a></b> <span class="reference-text">Midoux P, Pichon C, Yaouanc JJ, Jaffrès PA. "Chemical vectors for gene delivery: a current review on polymers, peptides and lipids containing histidine or imidazole as nucleic acids carriers." <i>Br J Pharmacol.</i> 2009 May;157(2):166-78. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r101121245"><a href="/wiki/PMID_(%E8%AD%98%E5%88%A5%E5%AD%90)" class="mw-redirect" title="PMID (識別子)">PMID</a> <a rel="nofollow" class="external text" href="//pubmed.ncbi.nlm.nih.gov/19459843/">19459843</a></span> </li> <li id="cite_note-5"><b><a href="#cite_ref-5">^</a></b> <span class="reference-text">Al-Dosari MS, Gao X. "Nonviral gene delivery: principle, limitations, and recent progress." <i>AAPS J.</i> 2009 Dec;11(4):671-81. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r101121245"><a href="/wiki/PMID_(%E8%AD%98%E5%88%A5%E5%AD%90)" class="mw-redirect" title="PMID (識別子)">PMID</a> <a rel="nofollow" class="external text" href="//pubmed.ncbi.nlm.nih.gov/19834816/">19834816</a></span> </li> <li id="cite_note-6"><b><a href="#cite_ref-6">^</a></b> <span class="reference-text">Adamson AD, Jackson D, Davis JR. "Novel approaches to in vitro transgenesis." <i>J Endocrinol.</i> 2011 Mar;208(3):193-206. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r101121245"><a href="/wiki/PMID_(%E8%AD%98%E5%88%A5%E5%AD%90)" class="mw-redirect" title="PMID (識別子)">PMID</a> <a rel="nofollow" class="external text" href="//pubmed.ncbi.nlm.nih.gov/21134989/">21134989</a></span> </li> <li id="cite_note-7"><b><a href="#cite_ref-7">^</a></b> <span class="reference-text">Muñoz-López M, García-Pérez JL. "DNA transposons: nature and applications in genomics." <i>Curr Genomics.</i> 2010 Apr;11(2):115-28. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r101121245"><a href="/wiki/PMID_(%E8%AD%98%E5%88%A5%E5%AD%90)" class="mw-redirect" title="PMID (識別子)">PMID</a> <a rel="nofollow" class="external text" href="//pubmed.ncbi.nlm.nih.gov/20885819/">20885819</a></span> </li> <li id="cite_note-8"><b><a href="#cite_ref-8">^</a></b> <span class="reference-text">Wang Y, Yau YY, Perkins-Balding D, Thomson JG. "Recombinase technology: applications and possibilities." <i>Plant Cell Rep.</i> 2011 Mar;30(3):267-85. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r101121245"><a href="/wiki/PMID_(%E8%AD%98%E5%88%A5%E5%AD%90)" class="mw-redirect" title="PMID (識別子)">PMID</a> <a rel="nofollow" class="external text" href="//pubmed.ncbi.nlm.nih.gov/20972794/">20972794</a></span> </li> </ol> <div class="mw-heading mw-heading2"><h2 id="外部リンク"><span id=".E5.A4.96.E9.83.A8.E3.83.AA.E3.83.B3.E3.82.AF"></span>外部リンク</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5&action=edit&section=5" title="節を編集: 外部リンク"><span>編集</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a rel="nofollow" class="external text" href="https://bsd.neuroinf.jp/wiki/%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5">遺伝子導入</a> - <a href="/wiki/%E8%84%B3%E7%A7%91%E5%AD%A6%E8%BE%9E%E5%85%B8" title="脳科学辞典">脳科学辞典</a></li></ul>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1&useformat=desktop" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">「<a dir="ltr" href="https://ja.wikipedia.org/w/index.php?title=遺伝子導入&oldid=102726399">https://ja.wikipedia.org/w/index.php?title=遺伝子導入&oldid=102726399</a>」から取得</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/%E7%89%B9%E5%88%A5:%E3%82%AB%E3%83%86%E3%82%B4%E3%83%AA" title="特別:カテゴリ">カテゴリ</a>: <ul><li><a href="/wiki/Category:%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B7%A5%E5%AD%A6" title="Category:遺伝子工学">遺伝子工学</a></li></ul></div></div> </div> </main> </div> <div class="mw-footer-container"> <footer id="footer" class="mw-footer" > <ul id="footer-info"> <li id="footer-info-lastmod"> 最終更新 2024年11月27日 (水) 11:23 （日時は<a href="/wiki/%E7%89%B9%E5%88%A5:%E5%80%8B%E4%BA%BA%E8%A8%AD%E5%AE%9A#mw-prefsection-rendering" title="特別:個人設定">個人設定</a>で未設定ならば<a href="/wiki/%E5%8D%94%E5%AE%9A%E4%B8%96%E7%95%8C%E6%99%82" title="協定世界時">UTC</a>）。</li> <li id="footer-info-copyright">テキストは<a rel="nofollow" class="external text" href="https://creativecommons.org/licenses/by-sa/4.0/deed.ja">クリエイティブ・コモンズ表示-継承ライセンス</a>のもとで利用できます。追加の条件が適用される場合があります。詳細については<a class="external text" href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Terms_of_Use">利用規約</a>を参照してください。</li> </ul> <ul id="footer-places"> <li id="footer-places-privacy"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Privacy_policy/ja">プライバシー・ポリシー</a></li> <li id="footer-places-about"><a href="/wiki/Wikipedia:%E3%82%A6%E3%82%A3%E3%82%AD%E3%83%9A%E3%83%87%E3%82%A3%E3%82%A2%E3%81%AB%E3%81%A4%E3%81%84%E3%81%A6">ウィキペディアについて</a></li> <li id="footer-places-disclaimers"><a href="/wiki/Wikipedia:%E5%85%8D%E8%B2%AC%E4%BA%8B%E9%A0%85">免責事項</a></li> <li id="footer-places-wm-codeofconduct"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Universal_Code_of_Conduct">行動規範</a></li> <li id="footer-places-developers"><a href="https://developer.wikimedia.org">開発者</a></li> <li id="footer-places-statslink"><a href="https://stats.wikimedia.org/#/ja.wikipedia.org">統計</a></li> <li id="footer-places-cookiestatement"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Cookie_statement">Cookieに関する声明</a></li> <li id="footer-places-mobileview"><a href="//ja.m.wikipedia.org/w/index.php?title=%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5&mobileaction=toggle_view_mobile" class="noprint stopMobileRedirectToggle">モバイルビュー</a></li> </ul> <ul id="footer-icons" class="noprint"> <li id="footer-copyrightico"><a href="https://wikimediafoundation.org/" class="cdx-button cdx-button--fake-button cdx-button--size-large cdx-button--fake-button--enabled"><img src="/static/images/footer/wikimedia-button.svg" width="84" height="29" alt="Wikimedia Foundation" loading="lazy"></a></li> <li id="footer-poweredbyico"><a href="https://www.mediawiki.org/" class="cdx-button cdx-button--fake-button cdx-button--size-large cdx-button--fake-button--enabled"><img src="/w/resources/assets/poweredby_mediawiki.svg" alt="Powered by MediaWiki" width="88" height="31" loading="lazy"></a></li> </ul> </footer> </div> </div> </div> <div class="vector-settings" id="p-dock-bottom"> <ul></ul> </div><script>(RLQ=window.RLQ||[]).push(function(){mw.config.set({"wgHostname":"mw-web.codfw.main-5c59558b9d-tb2bj","wgBackendResponseTime":134,"wgPageParseReport":{"limitreport":{"cputime":"0.063","walltime":"0.075","ppvisitednodes":{"value":879,"limit":1000000},"postexpandincludesize":{"value":4191,"limit":2097152},"templateargumentsize":{"value":190,"limit":2097152},"expansiondepth":{"value":14,"limit":100},"expensivefunctioncount":{"value":0,"limit":500},"unstrip-depth":{"value":1,"limit":20},"unstrip-size":{"value":18892,"limit":5000000},"entityaccesscount":{"value":0,"limit":400},"timingprofile":["100.00% 56.293 1 -total"," 83.63% 47.078 8 Template:PMID"," 79.95% 45.005 8 Template:Catalog_lookup_link"," 9.91% 5.578 16 Template:Yesno-no"," 7.75% 4.364 1 Template:脳科学事典"," 5.97% 3.359 16 Template:Yesno"," 2.27% 1.279 1 Template:脚注ヘルプ"]},"scribunto":{"limitreport-timeusage":{"value":"0.006","limit":"10.000"},"limitreport-memusage":{"value":898412,"limit":52428800}},"cachereport":{"origin":"mw-api-ext.codfw.main-75c5757d46-hs7gl","timestamp":"20241127112338","ttl":2592000,"transientcontent":false}}});});</script> <script type="application/ld+json">{"@context":"https:\/\/schema.org","@type":"Article","name":"\u907a\u4f1d\u5b50\u5c0e\u5165","url":"https:\/\/ja.wikipedia.org\/wiki\/%E9%81%BA%E4%BC%9D%E5%AD%90%E5%B0%8E%E5%85%A5","sameAs":"http:\/\/www.wikidata.org\/entity\/Q2305545","mainEntity":"http:\/\/www.wikidata.org\/entity\/Q2305545","author":{"@type":"Organization","name":"\u30a6\u30a3\u30ad\u30e1\u30c7\u30a3\u30a2\u30d7\u30ed\u30b8\u30a7\u30af\u30c8\u3078\u306e\u8ca2\u732e\u8005"},"publisher":{"@type":"Organization","name":"\u30a6\u30a3\u30ad\u30e1\u30c7\u30a3\u30a2\u8ca1\u56e3","logo":{"@type":"ImageObject","url":"https:\/\/www.wikimedia.org\/static\/images\/wmf-hor-googpub.png"}},"datePublished":"2011-07-27T20:43:31Z","dateModified":"2024-11-27T11:23:38Z"}</script> </body> </html>