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Riboviria - Wikipedia
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<div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>Interactions with hosts</span> </div> </a> <button aria-controls="toc-Interactions_with_hosts-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Interactions with hosts subsection</span> </button> <ul id="toc-Interactions_with_hosts-sublist" class="vector-toc-list"> <li id="toc-Disease" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Disease"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.1</span> <span>Disease</span> </div> </a> <ul id="toc-Disease-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Endogenization" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Endogenization"> <div class="vector-toc-text"> <span class="vector-toc-numb">5.2</span> <span>Endogenization</span> </div> </a> <ul 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Available in 20 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-20" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">20 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%B1%D9%8A%D8%A8%D9%88%D9%81%D9%8A%D8%B1%D9%8A%D8%A7" title="ريبوفيريا – Arabic" lang="ar" hreflang="ar" data-title="ريبوفيريا" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Riboviria" title="Riboviria – German" lang="de" hreflang="de" data-title="Riboviria" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Riboviria" title="Riboviria – Spanish" lang="es" hreflang="es" data-title="Riboviria" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%B1%DB%8C%D8%A8%D9%88%D9%88%DB%8C%D8%B1%DB%8C%D8%A7" title="ریبوویریا – Persian" lang="fa" hreflang="fa" data-title="ریبوویریا" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Riboviria" title="Riboviria – French" lang="fr" hreflang="fr" data-title="Riboviria" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%B0%E0%A4%BE%E0%A4%87%E0%A4%AC%E0%A5%8B%E0%A4%B5%E0%A4%BF%E0%A4%B0%E0%A4%BF%E0%A4%AF%E0%A4%BE" title="राइबोविरिया – Hindi" lang="hi" hreflang="hi" data-title="राइबोविरिया" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Riboviria" title="Riboviria – Indonesian" lang="id" hreflang="id" data-title="Riboviria" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Riboviria" title="Riboviria – Italian" lang="it" hreflang="it" data-title="Riboviria" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ku mw-list-item"><a href="https://ku.wikipedia.org/wiki/Riboviria" title="Riboviria – Kurdish" lang="ku" hreflang="ku" data-title="Riboviria" data-language-autonym="Kurdî" data-language-local-name="Kurdish" class="interlanguage-link-target"><span>Kurdî</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Virus_ARN" title="Virus ARN – Latin" lang="la" hreflang="la" data-title="Virus ARN" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-lv mw-list-item"><a href="https://lv.wikipedia.org/wiki/Ribov%C4%ABrusi" title="Ribovīrusi – Latvian" lang="lv" hreflang="lv" data-title="Ribovīrusi" data-language-autonym="Latviešu" data-language-local-name="Latvian" class="interlanguage-link-target"><span>Latviešu</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Riboviria" title="Riboviria – Dutch" lang="nl" hreflang="nl" data-title="Riboviria" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%83%AA%E3%83%9C%E3%82%A6%E3%82%A3%E3%83%AA%E3%82%A2" title="リボウィリア – Japanese" lang="ja" hreflang="ja" data-title="リボウィリア" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Riboviria" title="Riboviria – Portuguese" lang="pt" hreflang="pt" data-title="Riboviria" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru badge-Q17559452 badge-recommendedarticle mw-list-item" title="recommended article"><a href="https://ru.wikipedia.org/wiki/Riboviria" title="Riboviria – Russian" lang="ru" hreflang="ru" data-title="Riboviria" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sco mw-list-item"><a href="https://sco.wikipedia.org/wiki/Riboviria" title="Riboviria – Scots" lang="sco" hreflang="sco" data-title="Riboviria" data-language-autonym="Scots" data-language-local-name="Scots" class="interlanguage-link-target"><span>Scots</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/Riboviria" title="Riboviria – Serbian" lang="sr" hreflang="sr" data-title="Riboviria" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Riboviria" title="Riboviria – Turkish" lang="tr" hreflang="tr" data-title="Riboviria" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/Riboviria" title="Riboviria – Ukrainian" lang="uk" hreflang="uk" data-title="Riboviria" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E6%A0%B8%E7%B3%96%E7%97%85%E6%AF%92%E5%9F%9F" title="核糖病毒域 – Chinese" lang="zh" hreflang="zh" data-title="核糖病毒域" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q62002503#sitelinks-wikipedia" title="Edit interlanguage links" class="wbc-editpage">Edit links</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Namespaces"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Riboviria" title="View 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style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Riboviria_collage.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/d/da/Riboviria_collage.png/220px-Riboviria_collage.png" decoding="async" width="220" height="188" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/da/Riboviria_collage.png/330px-Riboviria_collage.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/da/Riboviria_collage.png/440px-Riboviria_collage.png 2x" data-file-width="3264" data-file-height="2784" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%">Clockwise from top left: <a href="/wiki/Transmission_electron_microscopy" title="Transmission electron microscopy">TEM</a> of <i><a href="/wiki/Avian_coronavirus" title="Avian coronavirus">avian coronavirus</a></i>, <a href="/wiki/Polio_virus" class="mw-redirect" title="Polio virus">polio virus</a>, <a href="/wiki/Bacteriophage_Q%CE%B2" class="mw-redirect" title="Bacteriophage Qβ">bacteriophage Qβ</a>, <i><a href="/wiki/Ebolavirus" title="Ebolavirus">ebolavirus</a></i>, <i><a href="/wiki/Tobacco_mosaic_virus" title="Tobacco mosaic virus">tobacco mosaic virus</a></i>, <i><a href="/wiki/Influenzavirus_A" class="mw-redirect" title="Influenzavirus A">influenzavirus A</a></i>, <i><a href="/wiki/Rotavirus" title="Rotavirus">rotavirus</a></i>, <a href="/wiki/HIV-1" class="mw-redirect" title="HIV-1">HIV-1</a>. Center: <a href="/wiki/Homology_(biology)" title="Homology (biology)">homologous</a> <a href="/wiki/Reverse_transcriptase" title="Reverse transcriptase">RT</a> and <a href="/wiki/RNA-dependent_RNA_polymerase" title="RNA-dependent RNA polymerase">RdRps</a> with conserved palm domain. </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(250,250,190)"><a href="/wiki/Virus_classification" title="Virus classification">Virus classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Riboviria" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>(unranked): </td> <td><a href="/wiki/Virus" title="Virus">Virus</a> </td></tr> <tr> <td><i>Realm</i>: </td> <td><b><i>Riboviria</i></b> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(250,250,190)">Kingdoms </th></tr> <tr> <td colspan="2" style="text-align: left"> <ul><li><i><a href="/wiki/Orthornavirae" title="Orthornavirae">Orthornavirae</a></i></li> <li><i><a href="/wiki/Pararnavirae" class="mw-redirect" title="Pararnavirae">Pararnavirae</a></i></li></ul> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><i><b>Riboviria</b></i> is a <a href="/wiki/Realm_(virology)" title="Realm (virology)">realm</a> of <a href="/wiki/Virus" title="Virus">viruses</a> that includes all viruses that use a homologous RNA-dependent polymerase for replication. It includes <a href="/wiki/RNA_virus" title="RNA virus">RNA viruses</a> that <a href="/wiki/Genetic_code" title="Genetic code">encode</a> an <a href="/wiki/RNA-dependent_RNA_polymerase" title="RNA-dependent RNA polymerase">RNA-dependent RNA polymerase</a>, as well as <a href="/wiki/Pararnavirae" class="mw-redirect" title="Pararnavirae">reverse-transcribing viruses</a> (with either RNA or DNA genomes) that encode an <a href="/wiki/Reverse_transcriptase" title="Reverse transcriptase">RNA-dependent DNA polymerase</a>. RNA-dependent RNA polymerase (RdRp), also called RNA replicase, produces RNA (<a href="/wiki/Ribonucleic_acid" class="mw-redirect" title="Ribonucleic acid">ribonucleic acid</a>) from RNA. RNA-dependent DNA polymerase (RdDp), also called reverse transcriptase (RT), produces DNA (<a href="/wiki/Deoxyribonucleic_acid" class="mw-redirect" title="Deoxyribonucleic acid">deoxyribonucleic acid</a>) from RNA. These <a href="/wiki/Enzyme" title="Enzyme">enzymes</a> are essential for <a href="/wiki/Viral_replication" title="Viral replication">replicating</a> the viral genome and <a href="/wiki/Transcription_(biology)" title="Transcription (biology)">transcribing</a> viral genes into <a href="/wiki/Messenger_RNA" title="Messenger RNA">messenger RNA</a> (mRNA) for <a href="/wiki/Translation_(biology)" title="Translation (biology)">translation</a> of viral <a href="/wiki/Protein" title="Protein">proteins</a>. </p><p><i>Riboviria</i> was established in 2018 to accommodate all RdRp-encoding RNA viruses and was expanded a year later to also include RdDp-encoding <a href="/wiki/Retrovirus" title="Retrovirus">retroviruses</a>. These two groups of viruses are assigned to two separate kingdoms: <i>Orthornavirae</i> for RdRp-encoding RNA viruses, and <i>Pararnavirae</i> for RdDp-encoding viruses, i.e. all reverse-transcribing viruses. While the realm has few <a href="/wiki/Prokaryotic" class="mw-redirect" title="Prokaryotic">prokaryotic</a> viruses, it includes most <a href="/wiki/Eukaryotic" class="mw-redirect" title="Eukaryotic">eukaryotic</a> viruses, including most human, animal, and plant viruses, however, metagenomic studies are changing this perspective. </p><p>Many of the most widely known viral diseases are caused by viruses in <i>Riboviria</i>, which includes <a href="/wiki/Coronavirus" title="Coronavirus">coronaviruses</a>, <a href="/wiki/Ebola_virus" class="mw-redirect" title="Ebola virus">ebola virus</a>, <a href="/wiki/HIV" title="HIV">HIV</a>, <a href="/wiki/Influenza_virus" class="mw-redirect" title="Influenza virus">influenza viruses</a>, and the <a href="/wiki/Rabies_virus" title="Rabies virus">rabies virus</a>. These viruses and others have been prominent throughout history, including <i><a href="/wiki/Tobacco_mosaic_virus" title="Tobacco mosaic virus">Tobacco mosaic virus</a></i>, which was the first virus to be discovered. Many reverse transcribing viruses notably become integrated into the genome of their host as part of their replication cycle. As a result of that, it is estimated that about 6.5–8% of the human genome originates from these viruses. <sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (November 2024)">citation needed</span></a></i>]</sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Etymology">Etymology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Riboviria&action=edit&section=1" title="Edit section: Etymology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><i>Riboviria</i> is a <a href="/wiki/Portmanteau" class="mw-redirect" title="Portmanteau">portmanteau</a> of <i>ribo</i>, referencing ribonucleic acid, and the suffix -<i>viria</i>, which is the suffix used for virus realms.<sup id="cite_ref-2017a_1-0" class="reference"><a href="#cite_note-2017a-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Characteristics">Characteristics</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Riboviria&action=edit&section=2" title="Edit section: Characteristics"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>All members of <i>Riboviria</i> contain a gene that encodes for an RNA-dependent polymerase, also called RNA-directed polymerase. There are two types of RNA-dependent polymerases: RNA-dependent RNA polymerase (RdRp), also called RNA replicase, which synthesizes RNA from RNA, and RNA-dependent DNA polymerase (RdDp), also called reverse transcriptase (RT), which synthesizes DNA from RNA.<sup id="cite_ref-2019a_2-0" class="reference"><a href="#cite_note-2019a-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> In a typical virus particle, called a virion, the RNA-dependent polymerase is bound to the viral genome in some manner and begins transcription of the viral genome after <a href="/wiki/Viral_entry" title="Viral entry">entering a cell</a>. As part of a <a href="/wiki/Viral_life_cycle" title="Viral life cycle">virus's life cycle</a>, the RNA-dependent polymerase also synthesizes copies of the viral genome as part of the process of creating new viruses. </p><p>Viruses that replicate via RdRp belong to three groups in the Baltimore classification system, all of which are in the kingdom <i>Orthornavirae</i>: single-stranded RNA (ssRNA) viruses, which are either positive (+) or negative (-) <a href="/wiki/Sense_(molecular_biology)" title="Sense (molecular biology)">sense</a>, and double-stranded RNA viruses (dsRNA). +ssRNA viruses have genomes that can functionally act as mRNA, and a negative sense strand can also be created to form dsRNA from which mRNA is transcribed from the negative strand.<sup id="cite_ref-pssrna_3-0" class="reference"><a href="#cite_note-pssrna-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> The genomes of -ssRNA viruses and dsRNA viruses act as templates from which RdRp creates mRNA.<sup id="cite_ref-dsrna_4-0" class="reference"><a href="#cite_note-dsrna-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-nssrna_5-0" class="reference"><a href="#cite_note-nssrna-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p><p>Viruses that replicate via reverse transcription belong to two Baltimore groups, both of which are in the kingdom <i>Pararnavirae</i>: single-stranded RNA (ssRNA-RT) viruses, all of which belong to the order <i><a href="/wiki/Ortervirales" title="Ortervirales">Ortervirales</a></i>, and double-stranded DNA (dsDNA-RT) viruses, which belong to the family <i><a href="/wiki/Caulimoviridae" title="Caulimoviridae">Caulimoviridae</a></i>, also in <i>Ortervirales</i>, and the family <i><a href="/wiki/Hepadnaviridae" title="Hepadnaviridae">Hepadnaviridae</a></i> of the order <i>Blubervirales</i>. ssRNA-RT viruses have their positive-sense genome transcribed by RdDp to synthesize a negative sense complementary DNA (-cDNA) strand. The +RNA strand is degraded and later replaced by RdDp with a +DNA strand to synthesize a linear dsDNA copy of the viral genome. This genome is then <a href="/wiki/Integrase" title="Integrase">integrated</a> into the host cell's DNA.<sup id="cite_ref-ssrnart_6-0" class="reference"><a href="#cite_note-ssrnart-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p><p>For dsDNA-RT viruses, a pregenomic +RNA strand is transcribed from the relaxed circular DNA (rcDNA), which is in turn used by RdDp to transcribe a -cDNA strand. The +RNA strand is degraded and replaced in a similar manner as +ssRNA-RT viruses to synthesize the rcDNA. The rcDNA genome is later repaired by the host cell's DNA repair mechanisms to synthesize a covalently closed circular DNA (cccDNA) genome.<sup id="cite_ref-dsdnart_7-0" class="reference"><a href="#cite_note-dsdnart-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> The integrated genome of +ssRNA-RT viruses and the cccDNA of dsDNA-RT viruses are then transcribed into mRNA by the host cell enzyme <a href="/wiki/RNA_polymerase_II" title="RNA polymerase II">RNA polymerase II</a>.<sup id="cite_ref-ssrnart_6-1" class="reference"><a href="#cite_note-ssrnart-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-dsdnart_7-1" class="reference"><a href="#cite_note-dsdnart-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p><p>Viral mRNA is <a href="/wiki/Translation_(biology)" title="Translation (biology)">translated</a> by the host cell's <a href="/wiki/Ribosome" title="Ribosome">ribosomes</a> to produce viral proteins. In order to produce more viruses, viral RNA-dependent polymerases use copies of the viral genome as templates to replicate the viral genome. For +ssRNA viruses, an intermediate dsRNA genome is created from which +ssRNA is synthesized from the negative strand.<sup id="cite_ref-pssrna_3-1" class="reference"><a href="#cite_note-pssrna-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> For -ssRNA viruses, genomes are synthesized from complementary positive sense strands.<sup id="cite_ref-nssrna_5-1" class="reference"><a href="#cite_note-nssrna-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> dsRNA viruses replicate their genomes from mRNA by synthesizing a complementary negative sense strand to form genomic dsRNA.<sup id="cite_ref-dsrna_4-1" class="reference"><a href="#cite_note-dsrna-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> For dsDNA-RT viruses, pregenomic RNA created from the cccDNA is retrotranscribed into new dsDNA genomes.<sup id="cite_ref-dsdnart_7-2" class="reference"><a href="#cite_note-dsdnart-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> For +ssRNA-RT viruses, the genome is replicated from the integrated genome.<sup id="cite_ref-ssrnart_6-2" class="reference"><a href="#cite_note-ssrnart-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> After replication and translation, the genome and the viral proteins are assembled into complete virions, which then <a href="/wiki/Viral_shedding" title="Viral shedding">leave the host cell</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Phylogenetics">Phylogenetics</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Riboviria&action=edit&section=3" title="Edit section: Phylogenetics"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:MBio.02329-18.F1.large.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/e/eb/MBio.02329-18.F1.large.jpg/220px-MBio.02329-18.F1.large.jpg" decoding="async" width="220" height="164" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/eb/MBio.02329-18.F1.large.jpg/330px-MBio.02329-18.F1.large.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/eb/MBio.02329-18.F1.large.jpg/440px-MBio.02329-18.F1.large.jpg 2x" data-file-width="1280" data-file-height="952" /></a><figcaption>Phylogeny of <i>Orthornavirae</i> RNA-dependent polymerase. The five colored branches are the five phyla of <i>Orthornavirae</i>.</figcaption></figure> <p>Both kingdoms in <i>Riboviria</i> show a relation to the reverse transcriptases of <a href="/wiki/Group_II_intron" title="Group II intron">group II introns</a> that encode RTs and <a href="/wiki/Retrotransposon" title="Retrotransposon">retrotransposons</a>, which are self-replicating DNA sequences, the latter of which self-replicate via reverse transcription and integrate themselves into other parts of the same DNA molecule. Reverse transcribing viruses, assigned to <i>Pararnavirae</i>, appear to have evolved from a retrotransposon on a single occasion. The origin of the RdRps of <i>Orthornavirae</i> is less clear due to a lack of information, that they originate from a reverse transcriptase from bacterial <a href="/wiki/Group_II_intron" title="Group II intron">group II intron</a> before the emergence of eukaryotes<sup id="cite_ref-2019a_2-1" class="reference"><a href="#cite_note-2019a-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-wolf_8-0" class="reference"><a href="#cite_note-wolf-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-orter_9-0" class="reference"><a href="#cite_note-orter-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> or originated before the <a href="/wiki/Last_universal_common_ancestor" title="Last universal common ancestor">last universal common ancestor</a> (LUCA) being descendants of the ancient <a href="/wiki/RNA_world" title="RNA world">RNA world</a> and that they preceded the retroelement reverse transcriptases.<sup id="cite_ref-Krupovic2020_10-0" class="reference"><a href="#cite_note-Krupovic2020-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> A larger study (2022) where new lineages (phyla) were described, was in favor of the hypothesis that RNA viruses descend from the RNA world, suggesting that retroelements originated from an ancestor related to the phylum <i><a href="/wiki/Lenarviricota" title="Lenarviricota">Lenarviricota</a></i> and that members of a newly discovered <i><a href="/w/index.php?title=Taraviricota&action=edit&redlink=1" class="new" title="Taraviricota (page does not exist)">Taraviricota</a></i> lineage (phylum) would be the ancestors of all RNA viruses.<sup id="cite_ref-Zayed_12-0" class="reference"><a href="#cite_note-Zayed-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Classification">Classification</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Riboviria&action=edit&section=4" title="Edit section: Classification"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><i>Riboviria</i> contains two kingdoms: <i>Orthornavirae</i> and <i>Pararnavirae</i>. <i>Orthornavirae</i> contains multiple phyla and unassigned taxa, whereas <i>Pararnavirae</i> is monotypic down to the rank of class. This taxonomy can be visualized hereafter.<sup id="cite_ref-ictv_13-0" class="reference"><a href="#cite_note-ictv-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p> <ul><li>Kingdom: <i><a href="/wiki/Orthornavirae" title="Orthornavirae">Orthornavirae</a></i>, which contains all RdRp-encoding RNA viruses, i.e. all dsRNA, +ssRNA, and -ssRNA viruses, often collectively called RNA viruses <ul><li>Phylum: <i><a href="/wiki/Ambiviricota" title="Ambiviricota">Ambiviricota</a></i></li> <li>Phylum: <i><a href="/wiki/Duplornaviricota" title="Duplornaviricota">Duplornaviricota</a></i></li> <li>Phylum: <i><a href="/wiki/Kitrinoviricota" title="Kitrinoviricota">Kitrinoviricota</a></i></li> <li>Phylum: <i><a href="/wiki/Lenarviricota" title="Lenarviricota">Lenarviricota</a></i></li> <li>Phylum: <i><a href="/wiki/Negarnaviricota" class="mw-redirect" title="Negarnaviricota">Negarnaviricota</a></i></li> <li>Phylum: <i><a href="/wiki/Pisuviricota" title="Pisuviricota">Pisuviricota</a></i></li> <li>Family <i><a href="/wiki/Incertae_sedis" title="Incertae sedis">incertae sedis</a></i>: <i><a href="/wiki/Birnaviridae" title="Birnaviridae">Birnaviridae</a></i></li> <li>Family <i><a href="/wiki/Incertae_sedis" title="Incertae sedis">incertae sedis</a></i>: <i><a href="/wiki/Permutotetraviridae" title="Permutotetraviridae">Permutotetraviridae</a></i></li></ul></li> <li>Kingdom: <i><a href="/wiki/Pararnavirae" class="mw-redirect" title="Pararnavirae">Pararnavirae</a></i>, which contains all RdDp-encoding viruses, i.e. all ssRNA-RT and dsDNA-RT viruses, collectively called reverse transcribing viruses <ul><li>Phylum: <i><a href="/wiki/Artverviricota" class="mw-redirect" title="Artverviricota">Artverviricota</a></i></li></ul></li></ul> <p>Additionally, <i>Riboviria</i> contains two <i><a href="/wiki/Incertae_sedis" title="Incertae sedis">incertae sedis</a></i> families and four <i>incertae sedis</i> genera. Additional information about them is needed to know their exact placement in higher taxa.<sup id="cite_ref-2019a_2-2" class="reference"><a href="#cite_note-2019a-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-ictv_13-1" class="reference"><a href="#cite_note-ictv-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p> <ul><li>Family <i><a href="/wiki/Incertae_sedis" title="Incertae sedis">incertae sedis</a></i>: <i><a href="/w/index.php?title=Polymycoviridae&action=edit&redlink=1" class="new" title="Polymycoviridae (page does not exist)">Polymycoviridae</a></i></li> <li>Family <i><a href="/wiki/Incertae_sedis" title="Incertae sedis">incertae sedis</a></i>: <i><a href="/wiki/Sarthroviridae" class="mw-redirect" title="Sarthroviridae">Sarthroviridae</a></i></li> <li>Genus <i><a href="/wiki/Incertae_sedis" title="Incertae sedis">incertae sedis</a></i>: <i><a href="/wiki/Albetovirus" title="Albetovirus">Albetovirus</a></i></li> <li>Genus <i><a href="/wiki/Incertae_sedis" title="Incertae sedis">incertae sedis</a></i>: <i><a href="/wiki/Aumaivirus" class="mw-redirect" title="Aumaivirus">Aumaivirus</a></i></li> <li>Genus <i><a href="/wiki/Incertae_sedis" title="Incertae sedis">incertae sedis</a></i>: <i><a href="/wiki/Papanivirus" class="mw-redirect" title="Papanivirus">Papanivirus</a></i></li> <li>Genus <i><a href="/wiki/Incertae_sedis" title="Incertae sedis">incertae sedis</a></i>: <i><a href="/wiki/Virtovirus" class="mw-redirect" title="Virtovirus">Virtovirus</a></i></li></ul> <p>Metagenomic studies have suggested the existence of six new phyla not in the ICTV: <i><a href="/w/index.php?title=Arctiviricota&action=edit&redlink=1" class="new" title="Arctiviricota (page does not exist)">Arctiviricota</a></i>, <i><a href="/w/index.php?title=Taraviricota&action=edit&redlink=1" class="new" title="Taraviricota (page does not exist)">Taraviricota</a></i>, <i><a href="/w/index.php?title=Pomiviricota&action=edit&redlink=1" class="new" title="Pomiviricota (page does not exist)">Pomiviricota</a></i>, <i><a href="/w/index.php?title=Paraxenoviricota&action=edit&redlink=1" class="new" title="Paraxenoviricota (page does not exist)">Paraxenoviricota</a></i>, <i><a href="/w/index.php?title=Wamoviricota&action=edit&redlink=1" class="new" title="Wamoviricota (page does not exist)">Wamoviricota</a></i><sup id="cite_ref-Zayed_12-1" class="reference"><a href="#cite_note-Zayed-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> and <i><a href="/w/index.php?title=Artimaviricota&action=edit&redlink=1" class="new" title="Artimaviricota (page does not exist)">Artimaviricota</a></i>.<sup id="cite_ref-Urayama_14-0" class="reference"><a href="#cite_note-Urayama-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> </p><p><i>Riboviria</i> partially merges <a href="/wiki/Baltimore_classification" title="Baltimore classification">Baltimore classification</a> with virus taxonomy, including the Baltimore groups for RNA viruses and reverse transcribing viruses in the realm. Baltimore classification is a classification system used for viruses based on their manner of mRNA production, often used alongside standard virus taxonomy, which is based on evolutionary history. All members of five Baltimore groups belong to <i>Riboviria</i>: Group III: dsRNA viruses, Group IV: +ssRNA viruses, Group V: -ssRNA viruses, Group VI: ssRNA-RT viruses, and Group VII: dsDNA-RT viruses. Realms are the highest level of taxonomy used for viruses and <i>Riboviria</i> is one of four, the other three being <i><a href="/wiki/Duplodnaviria" title="Duplodnaviria">Duplodnaviria</a></i>, <i><a href="/wiki/Monodnaviria" title="Monodnaviria">Monodnaviria</a></i>, and <i><a href="/wiki/Varidnaviria" title="Varidnaviria">Varidnaviria</a></i>.<sup id="cite_ref-wolf_8-1" class="reference"><a href="#cite_note-wolf-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-orter_9-1" class="reference"><a href="#cite_note-orter-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-ictv_13-2" class="reference"><a href="#cite_note-ictv-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p><p>Most identified eukaryotic viruses are RNA viruses, and for that reason most eukaryotic viruses belong to <i>Riboviria</i>, including most human, animal, and plant viruses. Other major branches of eukaryotic viruses include <a href="/wiki/Herpesvirus" class="mw-redirect" title="Herpesvirus">herpesviruses</a> in <i>Duplodnaviria</i>,<sup id="cite_ref-duplo_15-0" class="reference"><a href="#cite_note-duplo-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> the kingdom <i><a href="/wiki/Shotokuvirae" title="Shotokuvirae">Shotokuvirae</a></i> in <i>Monodnaviria</i>,<sup id="cite_ref-mono_16-0" class="reference"><a href="#cite_note-mono-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> and many viruses in <i>Varidnaviria</i>.<sup id="cite_ref-vari_17-0" class="reference"><a href="#cite_note-vari-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> In contrast, only three groups of prokaryotic RNA viruses have been identified: the class <i><a href="/wiki/Leviviricetes" title="Leviviricetes">Leviviricetes</a></i>,<sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> the family <i><a href="/wiki/Cystoviridae" class="mw-redirect" title="Cystoviridae">Cystoviridae</a></i><sup id="cite_ref-wolf_8-2" class="reference"><a href="#cite_note-wolf-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> and the phylum <i><a href="/w/index.php?title=Artimaviricota&action=edit&redlink=1" class="new" title="Artimaviricota (page does not exist)">Artimaviricota</a></i>.<sup id="cite_ref-Urayama_14-1" class="reference"><a href="#cite_note-Urayama-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> They also suggest that families <i><a href="/wiki/Picobirnaviridae" class="mw-redirect" title="Picobirnaviridae">Picobirnaviridae</a></i> and <i><a href="/wiki/Partitiviridae" title="Partitiviridae">Partitiviridae</a></i> previously associated with eukaryotes also infect prokaryotes<sup id="cite_ref-Neri_19-0" class="reference"><a href="#cite_note-Neri-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> and also the phylum <i><a href="/w/index.php?title=Taraviricota&action=edit&redlink=1" class="new" title="Taraviricota (page does not exist)">Taraviricota</a></i>.<sup id="cite_ref-Zayed_12-2" class="reference"><a href="#cite_note-Zayed-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> Studies of metagenomic samples have uncovered new prokaryotic RNA virus taxa including two new phyla that infect only prokaryotes, suggesting that their diversity is greater than previously thought and challenging the traditional view that RNA viruses only infect mostly eukaryotes.<sup id="cite_ref-Neri_19-1" class="reference"><a href="#cite_note-Neri-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Interactions_with_hosts">Interactions with hosts</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Riboviria&action=edit&section=5" title="Edit section: Interactions with hosts"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Disease">Disease</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Riboviria&action=edit&section=6" title="Edit section: Disease"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Viruses in <i>Riboviria</i> are associated with a wide range of diseases, including many of the most widely known viral diseases. Notable disease-causing viruses in the realm include:<sup id="cite_ref-ictv_13-3" class="reference"><a href="#cite_note-ictv-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p> <style data-mw-deduplicate="TemplateStyles:r1184024115">.mw-parser-output .div-col{margin-top:0.3em;column-width:30em}.mw-parser-output .div-col-small{font-size:90%}.mw-parser-output .div-col-rules{column-rule:1px solid #aaa}.mw-parser-output .div-col dl,.mw-parser-output .div-col ol,.mw-parser-output .div-col ul{margin-top:0}.mw-parser-output .div-col li,.mw-parser-output .div-col dd{page-break-inside:avoid;break-inside:avoid-column}</style><div class="div-col" style="column-width: 20em;"> <ul><li><a href="/wiki/Coronavirus" title="Coronavirus">coronaviruses</a></li> <li><i><a href="/wiki/Crimean-Congo_hemorrhagic_fever_orthonairovirus" class="mw-redirect" title="Crimean-Congo hemorrhagic fever orthonairovirus">Crimean-Congo hemorrhagic fever orthonairovirus</a></i></li> <li><i><a href="/wiki/Dengue_virus" title="Dengue virus">Dengue virus</a></i></li> <li><i><a href="/wiki/Ebolavirus" title="Ebolavirus">Ebolavirus</a></i></li> <li><a href="/wiki/Hantavirus" class="mw-redirect" title="Hantavirus">hantaviruses</a></li> <li>the <i><a href="/wiki/Viral_hepatitis#Viral_hepatitis_types" title="Viral hepatitis">Hepatitis viruses</a></i></li> <li>the <a href="/wiki/Human_immunodeficiency_virus" class="mw-redirect" title="Human immunodeficiency virus">human immunodeficiency viruses</a></li> <li><i><a href="/wiki/Human_orthopneumovirus" class="mw-redirect" title="Human orthopneumovirus">Human orthopneumovirus</a></i></li> <li><a href="/wiki/Influenza_virus" class="mw-redirect" title="Influenza virus">influenza viruses</a></li> <li><i><a href="/wiki/Japanese_encephalitis_virus" class="mw-redirect" title="Japanese encephalitis virus">Japanese encephalitis virus</a></i></li> <li><i><a href="/wiki/Lassa_mammarenavirus" title="Lassa mammarenavirus">Lassa mammarenavirus</a></i></li> <li><i><a href="/wiki/Measles_morbillivirus" class="mw-redirect" title="Measles morbillivirus">Measles morbillivirus</a></i></li> <li><i><a href="/wiki/Mumps_orthorubulavirus" class="mw-redirect" title="Mumps orthorubulavirus">Mumps orthorubulavirus</a></i></li> <li><i><a href="/wiki/Norovirus" title="Norovirus">Norovirus</a></i></li> <li><a href="/wiki/Poliovirus" title="Poliovirus">Poliovirus</a></li> <li><i><a href="/wiki/Rabies_lyssavirus" class="mw-redirect" title="Rabies lyssavirus">Rabies lyssavirus</a></i></li> <li><a href="/wiki/Rhinovirus" title="Rhinovirus">Rhinoviruses</a></li> <li><i><a href="/wiki/Rift_Valley_fever_phlebovirus" class="mw-redirect" title="Rift Valley fever phlebovirus">Rift Valley fever phlebovirus</a></i></li> <li><i><a href="/wiki/Rotavirus" title="Rotavirus">Rotavirus</a></i></li> <li><i><a href="/wiki/Rubella_virus" title="Rubella virus">Rubella virus</a></i></li> <li><i><a href="/wiki/Tick-borne_encephalitis_virus" title="Tick-borne encephalitis virus">Tick-borne encephalitis virus</a></i></li> <li><i><a href="/wiki/West_Nile_virus" title="West Nile virus">West Nile virus</a></i></li> <li><i><a href="/wiki/Yellow_fever_virus" class="mw-redirect" title="Yellow fever virus">Yellow fever virus</a></i></li> <li><i><a href="/wiki/Zika_virus" title="Zika virus">Zika virus</a></i></li></ul> </div> <p>Animal viruses in <i>Riboviria</i> include <a href="/wiki/Orbivirus" title="Orbivirus">orbiviruses</a>, which cause various diseases in ruminants and horses, including <i><a href="/wiki/Bluetongue_virus" title="Bluetongue virus">Bluetongue virus</a></i>, <i><a href="/wiki/African_horse_sickness_virus" class="mw-redirect" title="African horse sickness virus">African horse sickness virus</a></i>, <i><a href="/wiki/Equine_encephalosis_virus" title="Equine encephalosis virus">Equine encephalosis virus</a></i>, and <i><a href="/wiki/Epizootic_hemorrhagic_disease_virus" title="Epizootic hemorrhagic disease virus">epizootic hemorrhagic disease virus</a></i>.<sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> The <a href="/wiki/Vesicular_stomatitis_virus" class="mw-redirect" title="Vesicular stomatitis virus">vesicular stomatitis virus</a> causes disease in cattle, horses, and pigs.<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> Bats harbor many viruses including <a href="/wiki/Ebolavirus" title="Ebolavirus">ebolaviruses</a> and <a href="/wiki/Henipavirus" title="Henipavirus">henipaviruses</a>, which also can cause disease in humans.<sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup> Similarly, arthropod viruses in the <i><a href="/wiki/Flavivirus" title="Flavivirus">Flavivirus</a></i> and <i><a href="/wiki/Phlebovirus" title="Phlebovirus">Phlebovirus</a></i> genera are numerous and often transmitted to humans.<sup id="cite_ref-23" class="reference"><a href="#cite_note-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-24" class="reference"><a href="#cite_note-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup> Coronaviruses and influenza viruses cause disease in various vertebrates, including bats, birds, and pigs.<sup id="cite_ref-25" class="reference"><a href="#cite_note-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-26" class="reference"><a href="#cite_note-26"><span class="cite-bracket">[</span>26<span class="cite-bracket">]</span></a></sup> The family <i><a href="/wiki/Retroviridae" class="mw-redirect" title="Retroviridae">Retroviridae</a></i> contains many viruses that cause <a href="/wiki/Leukemia" title="Leukemia">leukemia</a>, <a href="/wiki/Immunodeficiency" title="Immunodeficiency">immunodeficiency</a>, and other cancers and immune system-related diseases in animals.<sup id="cite_ref-27" class="reference"><a href="#cite_note-27"><span class="cite-bracket">[</span>27<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-28" class="reference"><a href="#cite_note-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup> </p><p>Plant viruses in the realm are numerous and infect many economically important crops. <a href="/wiki/Tomato_spotted_wilt_virus" class="mw-redirect" title="Tomato spotted wilt virus">Tomato spotted wilt virus</a> is estimated to cause more than US$1 billion in damages annually, affecting more than 800 plant species including chrysanthemum, lettuce, peanut, pepper, and tomato. <i><a href="/wiki/Cucumber_mosaic_virus" title="Cucumber mosaic virus">Cucumber mosaic virus</a></i> infects more than 1,200 plant species and likewise causes significant crop losses. <i><a href="/wiki/Potato_virus_Y" title="Potato virus Y">Potato virus Y</a></i> causes significant reductions in yield and quality for pepper, potato, tobacco, and tomato, and <i><a href="/wiki/Plum_pox_virus" class="mw-redirect" title="Plum pox virus">Plum pox virus</a></i> is the most important virus among stone fruit crops. <i><a href="/wiki/Brome_mosaic_virus" title="Brome mosaic virus">Brome mosaic virus</a></i>, while not causing significant economic losses, is found throughout much of the world and primarily infects grasses, including cereals.<sup id="cite_ref-ictv_13-4" class="reference"><a href="#cite_note-ictv-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-29" class="reference"><a href="#cite_note-29"><span class="cite-bracket">[</span>29<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Endogenization">Endogenization</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Riboviria&action=edit&section=7" title="Edit section: Endogenization"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Many reverse transcribing viruses, called retroviruses, in <i>Riboviria</i> are able to become integrated into the DNA of their host. These viruses become endogenized as part of their replication cycle. Namely, the viral genome is integrated into the host genome by the retroviral enzyme integrase, and viral mRNA is produced from that DNA. Endogenization is a form of <a href="/wiki/Horizontal_gene_transfer" title="Horizontal gene transfer">horizontal gene transfer</a> between unrelated organisms, and it is estimated that about 7–8% of the human genome consists of retroviral DNA. Endogenization can also be used to study the evolutionary history of viruses, showing an approximate time period when a virus first became endogenized into the host's genome as well as the rate of evolution for the viruses since endogenization first occurred.<sup id="cite_ref-aiewsakun_30-0" class="reference"><a href="#cite_note-aiewsakun-30"><span class="cite-bracket">[</span>30<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="History">History</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Riboviria&action=edit&section=8" title="Edit section: History"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Diseases caused by viruses in <i>Riboviria</i> have been known for much of recorded history, though their cause was only discovered in modern times. <i>Tobacco mosaic virus</i> was discovered in 1898 and was the first virus to be discovered.<sup id="cite_ref-harrison_31-0" class="reference"><a href="#cite_note-harrison-31"><span class="cite-bracket">[</span>31<span class="cite-bracket">]</span></a></sup> Viruses transmitted by arthropods have been central in the development of <a href="/wiki/Vector_control" title="Vector control">vector control</a>, which often aims to prevent viral infections.<sup id="cite_ref-wilson_32-0" class="reference"><a href="#cite_note-wilson-32"><span class="cite-bracket">[</span>32<span class="cite-bracket">]</span></a></sup> In modern history, numerous disease outbreaks have been caused by various members of the realm, including coronaviruses, ebola, and influenza.<sup id="cite_ref-norris_33-0" class="reference"><a href="#cite_note-norris-33"><span class="cite-bracket">[</span>33<span class="cite-bracket">]</span></a></sup> HIV especially has had dramatic effects on society as it causes a sharp decline in life expectancy and significant stigma for infected persons.<sup id="cite_ref-yaghoobi_34-0" class="reference"><a href="#cite_note-yaghoobi-34"><span class="cite-bracket">[</span>34<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-cernasev_35-0" class="reference"><a href="#cite_note-cernasev-35"><span class="cite-bracket">[</span>35<span class="cite-bracket">]</span></a></sup> </p><p>For a long time, the relation between many viruses in <i>Riboviria</i> could not be established due to the high amount of genetic divergence among RNA viruses. With the development of viral <a href="/wiki/Metagenomics" title="Metagenomics">metagenomics</a>, many additional RNA viruses were identified, helping to fill in the gaps of their relations.<sup id="cite_ref-wolf_8-3" class="reference"><a href="#cite_note-wolf-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> This led to the establishment of <i>Riboviria</i> in 2018 to accommodate all RdRp-encoding RNA viruses based on phylogenetic analysis that they were related.<sup id="cite_ref-2017a_1-1" class="reference"><a href="#cite_note-2017a-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </p><p>A year later, all reverse transcribing viruses were added to the realm. The kingdoms were also established in 2019, separating the two RNA-dependent polymerase branches.<sup id="cite_ref-2019a_2-3" class="reference"><a href="#cite_note-2019a-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> When the realm was founded, it mistakenly included two <a href="/wiki/Viroid" title="Viroid">viroid</a> families, <i><a href="/wiki/Avsunviroidae" title="Avsunviroidae">Avsunviroidae</a></i> and <i><a href="/wiki/Pospiviroidae" title="Pospiviroidae">Pospiviroidae</a></i>, and the genus <i><a href="/wiki/Deltavirus" class="mw-redirect" title="Deltavirus">Deltavirus</a></i>, which were promptly removed in 2019 because they use host cell enzymes for replication.<sup id="cite_ref-36" class="reference"><a href="#cite_note-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Riboviria&action=edit&section=9" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239009302">.mw-parser-output .portalbox{padding:0;margin:0.5em 0;display:table;box-sizing:border-box;max-width:175px;list-style:none}.mw-parser-output .portalborder{border:1px solid var(--border-color-base,#a2a9b1);padding:0.1em;background:var(--background-color-neutral-subtle,#f8f9fa)}.mw-parser-output .portalbox-entry{display:table-row;font-size:85%;line-height:110%;height:1.9em;font-style:italic;font-weight:bold}.mw-parser-output .portalbox-image{display:table-cell;padding:0.2em;vertical-align:middle;text-align:center}.mw-parser-output .portalbox-link{display:table-cell;padding:0.2em 0.2em 0.2em 0.3em;vertical-align:middle}@media(min-width:720px){.mw-parser-output .portalleft{clear:left;float:left;margin:0.5em 1em 0.5em 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taxa</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Riboviria&action=edit&section=10" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width reflist-columns-2"> <ol class="references"> <li id="cite_note-2017a-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-2017a_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-2017a_1-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output 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title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=mBio&rft.atitle=Origins+and+Evolution+of+the+Global+RNA+Virome&rft.volume=9&rft.issue=6&rft.pages=e02329-18&rft.date=2018-11-27&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC6282212%23id-name%3DPMC&rft_id=info%3Apmid%2F30482837&rft_id=info%3Adoi%2F10.1128%2FmBio.02329-18&rft.aulast=Wolf&rft.aufirst=YI&rft.au=Kazlauskas%2C+D&rft.au=Iranzo%2C+J&rft.au=Lucia-Sanz%2C+A&rft.au=Kuhn%2C+JH&rft.au=Krupovic%2C+M&rft.au=Dolja%2C+VV&rft.au=Kooning%2C+EV&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC6282212&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARiboviria" class="Z3988"></span></span> </li> <li id="cite_note-orter-9"><span class="mw-cite-backlink">^ <a href="#cite_ref-orter_9-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-orter_9-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" 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title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=J+Virol&rft.atitle=Ortervirales%3A+New+Virus+Order+Unifying+Five+Families+of+Reverse-Transcribing+Viruses&rft.volume=92&rft.issue=12&rft.pages=e00515-18&rft.date=2018-06-15&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5974489%23id-name%3DPMC&rft_id=info%3Apmid%2F29618642&rft_id=info%3Adoi%2F10.1128%2FJVI.00515-18&rft.aulast=Krupovic&rft.aufirst=M&rft.au=Blomberg%2C+J&rft.au=Coffin%2C+JM&rft.au=Dasgupta%2C+I&rft.au=Fan%2C+H&rft.au=Geering%2C+AD&rft.au=Gifford%2C+R&rft.au=Harrach%2C+B&rft.au=Hull%2C+R&rft.au=Johnson%2C+W&rft.au=Kreuze%2C+JF&rft.au=Lindemann%2C+D&rft.au=Llorens%2C+C&rft.au=Lockhart%2C+B&rft.au=Mayer%2C+J&rft.au=Muller%2C+E&rft.au=Olszewski%2C+NE&rft.au=Pappu%2C+HR&rft.au=Pooggin%2C+MM&rft.au=Richert-Poggeler%2C+KR&rft.au=Sabanadzovic%2C+S&rft.au=Sanfacon%2C+H&rft.au=Schoelz%2C+JE&rft.au=Seal%2C+S&rft.au=Stavolone%2C+L&rft.au=Stoye%2C+JP&rft.au=Teycheney%2C+PY&rft.au=Tristem%2C+M&rft.au=Koonin%2C+EV&rft.au=Kuhn%2C+JH&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5974489&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARiboviria" 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Retrieved <span class="nowrap">15 May</span> 2020</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=International+Committee+on+Taxonomy+of+Viruses+%28ICTV%29&rft.atitle=Correction+of+an+administrative+error+resulting+in+an+incorrect+taxonomy+of+the+realm+Riboviria&rft.date=2019-07&rft.aulast=Gorbalenya&rft.aufirst=AE&rft.au=Krupovic%2C+M&rft.au=Siddell%2C+S&rft.au=Varsani%2C+A&rft.au=Kuhn%2C+JH&rft_id=https%3A%2F%2Fictv.global%2Fictv%2Fproposals%2F2019.009G.zip&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARiboviria" class="Z3988"></span></span> </li> </ol></div> <div class="mw-heading mw-heading2"><h2 id="Further_reading">Further reading</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Riboviria&action=edit&section=11" title="Edit section: Further reading"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWard1993" class="citation journal cs1">Ward, C. W. (1993). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7135741">"Progress towards a higher taxonomy of viruses"</a>. <i>Research in Virology</i>. <b>144</b> (6): 419–53. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2FS0923-2516%2806%2980059-2">10.1016/S0923-2516(06)80059-2</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7135741">7135741</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/8140287">8140287</a>.</cite><span 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style="width:1%">Cellular life</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">"<a href="/wiki/Prokaryote" title="Prokaryote">Prokaryota</a>"</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Archaea" title="Archaea">Archaea</a></li> <li><a href="/wiki/Bacteria" title="Bacteria">Bacteria</a> <ul><li><a href="/wiki/Mitochondrion" title="Mitochondrion">Mitochondrion</a></li> <li><a href="/wiki/Plastid" title="Plastid">Plastid</a></li></ul></li> <li><a href="/wiki/LUCA" class="mw-redirect" title="LUCA">LUCA</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Animal" title="Animal">Animalia</a></li> <li><a href="/wiki/Fungus" title="Fungus">Fungi</a></li> <li><a href="/wiki/Plant" title="Plant">Plantae</a></li> <li>'<a href="/wiki/Protist" title="Protist">Protista</a>'</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Incertae_sedis" title="Incertae sedis">Incertae sedis</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Parakaryon" title="Parakaryon">Parakaryon</a></i></li> <li><a href="/wiki/Microbial_dark_matter#Microbes_with_highly_unusual_DNA" title="Microbial dark matter">Microbes with highly unusual DNA</a> (?)</li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Non-cellular life</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Virus" title="Virus">Virus</a> <hr /><a href="/wiki/Viroid" title="Viroid">Viroid</a> <hr /><a href="/wiki/Satellite_(biology)" title="Satellite (biology)">Satellite</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Realms</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Adnaviria" title="Adnaviria">Adnaviria</a></i></li> <li><i><a href="/wiki/Duplodnaviria" title="Duplodnaviria">Duplodnaviria</a></i></li> <li><i><a href="/wiki/Monodnaviria" title="Monodnaviria">Monodnaviria</a></i></li> <li><i><a class="mw-selflink selflink">Riboviria</a></i></li> <li><i><a href="/wiki/Ribozyviria" title="Ribozyviria">Ribozyviria</a></i></li> <li><i><a href="/wiki/Varidnaviria" title="Varidnaviria">Varidnaviria</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Classes</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><i><a href="/wiki/Naldaviricetes" title="Naldaviricetes">Naldaviricetes</a></i></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Families</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alphasatellitidae" class="mw-redirect" title="Alphasatellitidae">Alphasatellitidae</a></i></li> <li><i><a href="/wiki/Ampullaviridae" title="Ampullaviridae">Ampullaviridae</a></i></li> <li><i><a href="/wiki/Anelloviridae" title="Anelloviridae">Anelloviridae</a></i></li> <li><i><a href="/wiki/Avsunviroidae" title="Avsunviroidae">Avsunviroidae</a></i></li> <li><i><a href="/wiki/Bicaudaviridae" title="Bicaudaviridae">Bicaudaviridae</a></i></li> <li><i><a href="/wiki/Clavaviridae" title="Clavaviridae">Clavaviridae</a></i></li> <li><i><a href="/wiki/Finnlakeviridae" title="Finnlakeviridae">Finnlakeviridae</a></i></li> <li><i><a href="/wiki/Fuselloviridae" title="Fuselloviridae">Fuselloviridae</a></i></li> <li><i><a href="/wiki/Globuloviridae" title="Globuloviridae">Globuloviridae</a></i></li> <li><i><a href="/wiki/Guttaviridae" title="Guttaviridae">Guttaviridae</a></i></li> <li><i><a href="/wiki/Halspiviridae" title="Halspiviridae">Halspiviridae</a></i></li> <li><i><a href="/wiki/Ovaliviridae" title="Ovaliviridae">Ovaliviridae</a></i></li> <li><i><a href="/wiki/Plasmaviridae" title="Plasmaviridae">Plasmaviridae</a></i></li> <li><i><a href="/wiki/Polydnaviridae" class="mw-redirect" title="Polydnaviridae">Polydnaviridae</a></i></li> <li><i><a href="/wiki/Portogloboviridae" title="Portogloboviridae">Portogloboviridae</a></i></li> <li><i><a href="/wiki/Pospiviroidae" title="Pospiviroidae">Pospiviroidae</a></i></li> <li><i><a href="/wiki/Spiraviridae" title="Spiraviridae">Spiraviridae</a></i></li> <li><i><a href="/wiki/Thaspiviridae" title="Thaspiviridae">Thaspiviridae</a></i></li> <li><i><a href="/wiki/Tolecusatellitidae" title="Tolecusatellitidae">Tolecusatellitidae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Genera</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Dinodnavirus" title="Dinodnavirus">Dinodnavirus</a></i></li> <li><i><a href="/wiki/Rhizidiovirus" title="Rhizidiovirus">Rhizidiovirus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Nanobe" title="Nanobe">Nanobe</a> (?)</li> <li><a href="/wiki/Obelisk_(biology)" title="Obelisk (biology)">Obelisk</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Comparable<br />structures</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bio-like_structure" title="Bio-like structure">Bio-like structure</a></li> <li><a href="/wiki/Cancer_cell" title="Cancer cell">Cancer cell</a> <ul><li><a href="/wiki/HeLa" title="HeLa">HeLa</a></li> <li><a href="/wiki/Clonally_transmissible_cancer" title="Clonally transmissible cancer">Clonally transmissible cancer</a></li></ul></li> <li><a href="/wiki/Cosmid" title="Cosmid">Cosmid</a></li> <li><a href="/wiki/Defective_interfering_particle" title="Defective interfering particle">Defective interfering particle</a></li> <li><a href="/wiki/Endogenous_viral_element" title="Endogenous viral element">Endogenous viral element</a></li> <li><a href="/wiki/Fosmid" title="Fosmid">Fosmid</a></li> <li><a href="/wiki/Integrative_and_conjugative_element" title="Integrative and conjugative element">Integrative and conjugative element</a></li> <li><a href="/wiki/Jeewanu" title="Jeewanu">Jeewanu</a></li> <li>"<i><a href="/wiki/Nanobacterium" title="Nanobacterium">Nanobacterium</a></i>"</li> <li><a href="/wiki/Phagemid" title="Phagemid">Phagemid</a></li> <li><a href="/wiki/Plasmid" title="Plasmid">Plasmid</a></li> <li><a href="/wiki/Prion" title="Prion">Prion</a> <ul><li><a href="/wiki/Fungal_prion" title="Fungal prion">Fungal prion</a></li></ul></li> <li><a href="/wiki/Proteinoid" title="Proteinoid">Proteinoid</a> <a href="/wiki/Microparticle#Biological_protocells" title="Microparticle">microsphere</a></li> <li>Retroelements not elsewhere classified <ul><li><a href="/wiki/Retron" title="Retron">Retron</a></li> <li><a href="/wiki/Diversity-generating_retroelement" title="Diversity-generating retroelement">Diversity-generating retroelement</a></li> <li><a href="/wiki/Telomerase_reverse_transcriptase" title="Telomerase reverse transcriptase">Telomerase reverse transcriptase</a></li> <li><a href="/wiki/Reverse_transcriptase-related_cellular_gene" class="mw-redirect" title="Reverse transcriptase-related cellular gene">Reverse transcriptase-related cellular gene</a></li></ul></li> <li><a href="/wiki/Ribozyme" title="Ribozyme">Ribozyme</a></li> <li><a href="/wiki/Spiegelman%27s_Monster" title="Spiegelman's Monster">Spiegelman's Monster</a></li> <li><a href="/wiki/Tandem_repeat" title="Tandem repeat">Tandem repeat</a></li> <li><a href="/wiki/Transposable_element" title="Transposable element">Transposable element</a> <ul><li><a href="/wiki/Retroposon" title="Retroposon">Retroposon</a></li></ul></li> <li><a href="/wiki/Transpoviron" title="Transpoviron">Transpoviron</a></li> <li><a href="/wiki/Xenobot" title="Xenobot">Xenobot</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Baltimore_(virus_classification)" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse 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style="font-size:114%;margin:0 4em"><a href="/wiki/Baltimore_classification" title="Baltimore classification">Baltimore</a> (<a href="/wiki/Virus_classification" title="Virus classification">virus classification</a>)</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/DNA_virus" title="DNA virus">DNA</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="I:_dsDNA_viruses" style="font-size:114%;margin:0 4em">I: <a href="/wiki/DsDNA_virus" class="mw-redirect" title="DsDNA virus">dsDNA viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Adnaviria" title="Adnaviria">Adnaviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Zilligvirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Zilligvirae" class="mw-redirect" title="Zilligvirae">Zilligvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Taleaviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Taleaviricota" class="mw-redirect" title="Taleaviricota">Taleaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Tokiviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tokiviricetes" class="mw-redirect" title="Tokiviricetes">Tokiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ligamenvirales" title="Ligamenvirales">Ligamenvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Lipothrixviridae" title="Lipothrixviridae">Lipothrixviridae</a></i></li> <li><i><a href="/wiki/Rudiviridae" title="Rudiviridae">Rudiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Primavirales" class="mw-redirect" title="Primavirales">Primavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Tristromaviridae" title="Tristromaviridae">Tristromaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Duplodnaviria" title="Duplodnaviria">Duplodnaviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Heunggongvirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Heunggongvirae" class="mw-redirect" title="Heunggongvirae">Heunggongvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Peploviricota" class="mw-redirect" title="Peploviricota">Peploviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Herviviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Herviviricetes" class="mw-redirect" title="Herviviricetes">Herviviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Herpesvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Herpesvirales" title="Herpesvirales">Herpesvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alloherpesviridae" title="Alloherpesviridae">Alloherpesviridae</a></i></li> <li><i><a href="/wiki/Herpesviridae" title="Herpesviridae">Herpesviridae</a></i></li> <li><i><a href="/wiki/Malacoherpesviridae" title="Malacoherpesviridae">Malacoherpesviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Uroviricota" class="mw-redirect" title="Uroviricota">Uroviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Caudoviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Caudoviricetes" title="Caudoviricetes">Caudoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Caudovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Caudovirales" class="mw-redirect" title="Caudovirales">Caudovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Ackermannviridae" title="Ackermannviridae">Ackermannviridae</a></i></li> <li><i><a href="/wiki/Autographiviridae" title="Autographiviridae">Autographiviridae</a></i></li> <li><i><a href="/w/index.php?title=Chaseviridae&action=edit&redlink=1" class="new" title="Chaseviridae (page does not exist)">Chaseviridae</a></i></li> <li><i><a href="/w/index.php?title=Demerecviridae&action=edit&redlink=1" class="new" title="Demerecviridae (page does not exist)">Demerecviridae</a></i></li> <li><i><a href="/w/index.php?title=Drexlerviridae&action=edit&redlink=1" class="new" title="Drexlerviridae (page does not exist)">Drexlerviridae</a></i></li> <li><i><a href="/w/index.php?title=Guelinviridae&action=edit&redlink=1" class="new" title="Guelinviridae (page does not exist)">Guelinviridae</a></i></li> <li><i><a href="/wiki/Herelleviridae" title="Herelleviridae">Herelleviridae</a></i></li> <li><i><a href="/wiki/Myoviridae" title="Myoviridae">Myoviridae</a></i></li> <li><i><a href="/wiki/Podoviridae" title="Podoviridae">Podoviridae</a></i></li> <li><i><a href="/w/index.php?title=Rountreeviridae&action=edit&redlink=1" class="new" title="Rountreeviridae (page does not exist)">Rountreeviridae</a></i></li> <li><i><a href="/w/index.php?title=Salasmaviridae&action=edit&redlink=1" class="new" title="Salasmaviridae (page does not exist)">Salasmaviridae</a></i></li> <li><i><a href="/w/index.php?title=Schitoviridae&action=edit&redlink=1" class="new" title="Schitoviridae (page does not exist)">Schitoviridae</a></i></li> <li><i><a href="/wiki/Siphoviridae" title="Siphoviridae">Siphoviridae</a></i></li> <li><i><a href="/w/index.php?title=Zobellviridae&action=edit&redlink=1" class="new" title="Zobellviridae (page does not exist)">Zobellviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Monodnaviria" title="Monodnaviria">Monodnaviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Shotokuvirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Shotokuvirae" title="Shotokuvirae">Shotokuvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Cossaviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cossaviricota" title="Cossaviricota">Cossaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Papovaviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Papovaviricetes" title="Papovaviricetes">Papovaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Sepolyvirales" class="mw-redirect" title="Sepolyvirales">Sepolyvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Polyomaviridae" title="Polyomaviridae">Polyomaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Zurhausenvirales" class="mw-redirect" title="Zurhausenvirales">Zurhausenvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Papillomaviridae" title="Papillomaviridae">Papillomaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Varidnaviria" title="Varidnaviria">Varidnaviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Bamfordvirae" title="Bamfordvirae">Bamfordvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Nucleocytoviricota" title="Nucleocytoviricota">Nucleocytoviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pokkesviricetes" title="Pokkesviricetes">Pokkesviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Asfuvirales" class="mw-redirect" title="Asfuvirales">Asfuvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Asfarviridae" title="Asfarviridae">Asfarviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Chitovirales" class="mw-redirect" title="Chitovirales">Chitovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Poxviridae" title="Poxviridae">Poxviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Megaviricetes" title="Megaviricetes">Megaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Algavirales" class="mw-redirect" title="Algavirales">Algavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Phycodnaviridae" title="Phycodnaviridae">Phycodnaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Imitervirales" class="mw-redirect" title="Imitervirales">Imitervirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Mimiviridae" title="Mimiviridae">Mimiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pimascovirales" title="Pimascovirales">Pimascovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Ascoviridae" title="Ascoviridae">Ascoviridae</a></i></li> <li><i><a href="/wiki/Iridoviridae" title="Iridoviridae">Iridoviridae</a></i></li> <li><i><a href="/wiki/Marseilleviridae" title="Marseilleviridae">Marseilleviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Preplasmiviricota" title="Preplasmiviricota">Preplasmiviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Maveriviricetes" class="mw-redirect" title="Maveriviricetes">Maveriviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Priklausovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Priklausovirales" class="mw-redirect" title="Priklausovirales">Priklausovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Lavidaviridae" class="mw-redirect" title="Lavidaviridae">Lavidaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Polintoviricetes" class="mw-redirect" title="Polintoviricetes">Polintoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Orthopolintovirales" class="mw-redirect" title="Orthopolintovirales">Orthopolintovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Adintoviridae" title="Adintoviridae">Adintoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tectiliviricetes" title="Tectiliviricetes">Tectiliviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Belfryvirales" class="mw-redirect" title="Belfryvirales">Belfryvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Turriviridae" title="Turriviridae">Turriviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Kalamavirales" class="mw-redirect" title="Kalamavirales">Kalamavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Tectiviridae" class="mw-redirect" title="Tectiviridae">Tectiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Rowavirales" class="mw-redirect" title="Rowavirales">Rowavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Adenoviridae" title="Adenoviridae">Adenoviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Vinavirales" class="mw-redirect" title="Vinavirales">Vinavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Corticoviridae" class="mw-redirect" title="Corticoviridae">Corticoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Helvetiavirae" class="mw-redirect" title="Helvetiavirae">Helvetiavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Dividoviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Dividoviricota" class="mw-redirect" title="Dividoviricota">Dividoviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Laserviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Laserviricetes" class="mw-redirect" title="Laserviricetes">Laserviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Halopanivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Halopanivirales" title="Halopanivirales">Halopanivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Matshushitaviridae" class="mw-redirect" title="Matshushitaviridae">Matshushitaviridae</a></i></li> <li><i><a href="/wiki/Simuloviridae" class="mw-redirect" title="Simuloviridae">Simuloviridae</a></i></li> <li><i><a href="/wiki/Sphaerolipoviridae" class="mw-redirect" title="Sphaerolipoviridae">Sphaerolipoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Naldaviricetes" title="Naldaviricetes">Naldaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Lefavirales" title="Lefavirales">Lefavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Baculoviridae" title="Baculoviridae">Baculoviridae</a></i></li> <li><i><a href="/wiki/Hytrosaviridae" title="Hytrosaviridae">Hytrosaviridae</a></i></li> <li><i><a href="/wiki/Nudiviridae" class="mw-redirect" title="Nudiviridae">Nudiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Nimaviridae" class="mw-redirect" title="Nimaviridae">Nimaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><b>Families</b>: <i><a href="/wiki/Ampullaviridae" title="Ampullaviridae">Ampullaviridae</a></i></li> <li><i><a href="/wiki/Bicaudaviridae" title="Bicaudaviridae">Bicaudaviridae</a></i></li> <li><i><a href="/wiki/Clavaviridae" title="Clavaviridae">Clavaviridae</a></i></li> <li><i><a href="/wiki/Fuselloviridae" title="Fuselloviridae">Fuselloviridae</a></i></li> <li><i><a href="/wiki/Globuloviridae" title="Globuloviridae">Globuloviridae</a></i></li> <li><i><a href="/wiki/Guttaviridae" title="Guttaviridae">Guttaviridae</a></i></li> <li><i><a href="/wiki/Halspiviridae" title="Halspiviridae">Halspiviridae</a></i></li> <li><i><a href="/wiki/Ovaliviridae" title="Ovaliviridae">Ovaliviridae</a></i></li> <li><i><a href="/wiki/Plasmaviridae" title="Plasmaviridae">Plasmaviridae</a></i></li> <li><i><a href="/wiki/Polydnaviridae" class="mw-redirect" title="Polydnaviridae">Polydnaviridae</a></i></li> <li><i><a href="/wiki/Portogloboviridae" title="Portogloboviridae">Portogloboviridae</a></i></li> <li><i><a href="/wiki/Thaspiviridae" title="Thaspiviridae">Thaspiviridae</a></i></li> <li><b>Genera</b>: <i><a href="/wiki/Dinodnavirus" title="Dinodnavirus">Dinodnavirus</a></i></li> <li><i><a href="/wiki/Rhizidiovirus" title="Rhizidiovirus">Rhizidiovirus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="II:_ssDNA_viruses" style="font-size:114%;margin:0 4em">II: <a href="/wiki/SsDNA_virus" class="mw-redirect" title="SsDNA virus">ssDNA viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Monodnaviria" title="Monodnaviria">Monodnaviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Loebvirae" class="mw-redirect" title="Loebvirae">Loebvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Hofneiviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Hofneiviricota" class="mw-redirect" title="Hofneiviricota">Hofneiviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Faserviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Faserviricetes" class="mw-redirect" title="Faserviricetes">Faserviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Tubulavirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tubulavirales" title="Tubulavirales">Tubulavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Inoviridae" class="mw-redirect" title="Inoviridae">Inoviridae</a></i></li> <li><i><a href="/w/index.php?title=Paulinoviridae&action=edit&redlink=1" class="new" title="Paulinoviridae (page does not exist)">Paulinoviridae</a></i></li> <li><i><a href="/w/index.php?title=Plectroviridae&action=edit&redlink=1" class="new" title="Plectroviridae (page does not exist)">Plectroviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Sangervirae" class="mw-redirect" title="Sangervirae">Sangervirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Phixviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Phixviricota" class="mw-redirect" title="Phixviricota">Phixviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Malgrandaviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Malgrandaviricetes" class="mw-redirect" title="Malgrandaviricetes">Malgrandaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Petitvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Petitvirales" class="mw-redirect" title="Petitvirales">Petitvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Microviridae" title="Microviridae">Microviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Shotokuvirae" title="Shotokuvirae">Shotokuvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cossaviricota" title="Cossaviricota">Cossaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Mouviricetes" class="mw-redirect" title="Mouviricetes">Mouviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Polivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Polivirales" class="mw-redirect" title="Polivirales">Polivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Bidnaviridae" class="mw-redirect" title="Bidnaviridae">Bidnaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Quintoviricetes" class="mw-redirect" title="Quintoviricetes">Quintoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Piccovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Piccovirales" class="mw-redirect" title="Piccovirales">Piccovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Parvoviridae" title="Parvoviridae">Parvoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cressdnaviricota" title="Cressdnaviricota">Cressdnaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Arfiviricetes" title="Arfiviricetes">Arfiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Baphyvirales" class="mw-redirect" title="Baphyvirales">Baphyvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Bacilladnaviridae" title="Bacilladnaviridae">Bacilladnaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cirlivirales" class="mw-redirect" title="Cirlivirales">Cirlivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Circoviridae" title="Circoviridae">Circoviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cremevirales" class="mw-redirect" title="Cremevirales">Cremevirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Smacoviridae" title="Smacoviridae">Smacoviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Mulpavirales" title="Mulpavirales">Mulpavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Metaxyviridae&action=edit&redlink=1" class="new" title="Metaxyviridae (page does not exist)">Metaxyviridae</a></i></li> <li><i><a href="/wiki/Nanoviridae" title="Nanoviridae">Nanoviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Recrevirales" class="mw-redirect" title="Recrevirales">Recrevirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Redondoviridae" title="Redondoviridae">Redondoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Repensiviricetes" class="mw-redirect" title="Repensiviricetes">Repensiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Geplafuvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Geplafuvirales" title="Geplafuvirales">Geplafuvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Geminiviridae" title="Geminiviridae">Geminiviridae</a></i></li> <li><i><a href="/wiki/Genomoviridae" title="Genomoviridae">Genomoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Trapavirae" class="mw-redirect" title="Trapavirae">Trapavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Saleviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Saleviricota" class="mw-redirect" title="Saleviricota">Saleviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Huolimaviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Huolimaviricetes" class="mw-redirect" title="Huolimaviricetes">Huolimaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Haloruvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Haloruvirales" class="mw-redirect" title="Haloruvirales">Haloruvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Pleolipoviridae" title="Pleolipoviridae">Pleolipoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><b>Families</b>: <i><a href="/wiki/Anelloviridae" title="Anelloviridae">Anelloviridae</a></i></li> <li><i><a href="/wiki/Finnlakeviridae" title="Finnlakeviridae">Finnlakeviridae</a></i></li> <li><i><a href="/wiki/Spiraviridae" title="Spiraviridae">Spiraviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/RNA_virus" title="RNA virus">RNA</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="III:_dsRNA_viruses" style="font-size:114%;margin:0 4em">III: <a href="/wiki/Double-stranded_RNA_viruses" title="Double-stranded RNA viruses">dsRNA viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Riboviria" scope="row" class="navbox-group" style="width:1%"><i><a class="mw-selflink selflink">Riboviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Orthornavirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Orthornavirae" title="Orthornavirae">Orthornavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Duplornaviricota" title="Duplornaviricota">Duplornaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Chrymotiviricetes" class="mw-redirect" title="Chrymotiviricetes">Chrymotiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Ghabrivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ghabrivirales" title="Ghabrivirales">Ghabrivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Chrysoviridae" title="Chrysoviridae">Chrysoviridae</a></i></li> <li><i><a href="/wiki/Megabirnaviridae" title="Megabirnaviridae">Megabirnaviridae</a></i></li> <li><i><a href="/wiki/Quadriviridae" title="Quadriviridae">Quadriviridae</a></i></li> <li><i><a href="/wiki/Totiviridae" title="Totiviridae">Totiviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Resentoviricetes" class="mw-redirect" title="Resentoviricetes">Resentoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Reovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Reovirales" class="mw-redirect" title="Reovirales">Reovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Reoviridae" class="mw-redirect" title="Reoviridae">Reoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Vidaverviricetes" class="mw-redirect" title="Vidaverviricetes">Vidaverviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Mindivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Mindivirales" class="mw-redirect" title="Mindivirales">Mindivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Cystoviridae" class="mw-redirect" title="Cystoviridae">Cystoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pisuviricota" title="Pisuviricota">Pisuviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Duplopiviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Duplopiviricetes" class="mw-redirect" title="Duplopiviricetes">Duplopiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Durnavirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Durnavirales" title="Durnavirales">Durnavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Amalgaviridae" title="Amalgaviridae">Amalgaviridae</a></i></li> <li><i><a href="/w/index.php?title=Curvulaviridae&action=edit&redlink=1" class="new" title="Curvulaviridae (page does not exist)">Curvulaviridae</a></i></li> <li><i><a href="/wiki/Hypoviridae" class="mw-redirect" title="Hypoviridae">Hypoviridae</a></i></li> <li><i><a href="/wiki/Picobirnaviridae" class="mw-redirect" title="Picobirnaviridae">Picobirnaviridae</a></i></li> <li><i><a href="/wiki/Partitiviridae" title="Partitiviridae">Partitiviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><b>Families</b>: <i><a href="/wiki/Birnaviridae" title="Birnaviridae">Birnaviridae</a></i></li> <li><i><a href="/w/index.php?title=Polymycoviridae&action=edit&redlink=1" class="new" title="Polymycoviridae (page does not exist)">Polymycoviridae</a></i></li> <li><b>Genera</b>: <i><a href="/w/index.php?title=Botybirnavirus&action=edit&redlink=1" class="new" title="Botybirnavirus (page does not exist)">Botybirnavirus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="IV:_(+)ssRNA_viruses" style="font-size:114%;margin:0 4em">IV: <a href="/wiki/Positive-sense_ssRNA_virus" class="mw-redirect" title="Positive-sense ssRNA virus">(+)ssRNA viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Riboviria" scope="row" class="navbox-group" style="width:1%"><i><a class="mw-selflink selflink">Riboviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Orthornavirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Orthornavirae" title="Orthornavirae">Orthornavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Kitrinoviricota" title="Kitrinoviricota">Kitrinoviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Alsuviricetes" title="Alsuviricetes">Alsuviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Hepelivirales" title="Hepelivirales">Hepelivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alphatetraviridae" title="Alphatetraviridae">Alphatetraviridae</a></i></li> <li><i><a href="/wiki/Benyviridae" class="mw-redirect" title="Benyviridae">Benyviridae</a></i></li> <li><i><a href="/wiki/Hepeviridae" title="Hepeviridae">Hepeviridae</a></i></li> <li><i><a href="/wiki/Matonaviridae" class="mw-redirect" title="Matonaviridae">Matonaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Martellivirales" title="Martellivirales">Martellivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Bromoviridae" title="Bromoviridae">Bromoviridae</a></i></li> <li><i><a href="/wiki/Closteroviridae" title="Closteroviridae">Closteroviridae</a></i></li> <li><i><a href="/wiki/Endornaviridae" title="Endornaviridae">Endornaviridae</a></i></li> <li><i><a href="/w/index.php?title=Kitaviridae&action=edit&redlink=1" class="new" title="Kitaviridae (page does not exist)">Kitaviridae</a></i></li> <li><i><a href="/w/index.php?title=Mayoviridae&action=edit&redlink=1" class="new" title="Mayoviridae (page does not exist)">Mayoviridae</a></i></li> <li><i><a href="/wiki/Togaviridae" class="mw-redirect" title="Togaviridae">Togaviridae</a></i></li> <li><i><a href="/wiki/Virgaviridae" title="Virgaviridae">Virgaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tymovirales" title="Tymovirales">Tymovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alphaflexiviridae" title="Alphaflexiviridae">Alphaflexiviridae</a></i></li> <li><i><a href="/wiki/Betaflexiviridae" title="Betaflexiviridae">Betaflexiviridae</a></i></li> <li><i><a href="/w/index.php?title=Deltaflexiviridae&action=edit&redlink=1" class="new" title="Deltaflexiviridae (page does not exist)">Deltaflexiviridae</a></i></li> <li><i><a href="/wiki/Gammaflexiviridae" title="Gammaflexiviridae">Gammaflexiviridae</a></i></li> <li><i><a href="/wiki/Tymoviridae" title="Tymoviridae">Tymoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Flasuviricetes" class="mw-redirect" title="Flasuviricetes">Flasuviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Amarillovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Amarillovirales" class="mw-redirect" title="Amarillovirales">Amarillovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Flaviviridae" title="Flaviviridae">Flaviviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Magsaviricetes" class="mw-redirect" title="Magsaviricetes">Magsaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Nodamuvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Nodamuvirales" title="Nodamuvirales">Nodamuvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Nodaviridae" title="Nodaviridae">Nodaviridae</a></i></li> <li><i><a href="/w/index.php?title=Sinhaliviridae&action=edit&redlink=1" class="new" title="Sinhaliviridae (page does not exist)">Sinhaliviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tolucaviricetes" class="mw-redirect" title="Tolucaviricetes">Tolucaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Tolivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tolivirales" title="Tolivirales">Tolivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Carmotetraviridae" title="Carmotetraviridae">Carmotetraviridae</a></i></li> <li><i><a href="/wiki/Luteoviridae" title="Luteoviridae">Luteoviridae</a></i></li> <li><i><a href="/wiki/Tombusviridae" title="Tombusviridae">Tombusviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Lenarviricota" title="Lenarviricota">Lenarviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Leviviricetes" title="Leviviricetes">Leviviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Norzivirales" title="Norzivirales">Norzivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Atkinsviridae" title="Atkinsviridae">Atkinsviridae</a></i></li> <li><i><a href="/wiki/Duinviridae" title="Duinviridae">Duinviridae</a></i></li> <li><i><a href="/wiki/Fiersviridae" title="Fiersviridae">Fiersviridae</a></i></li> <li><i><a href="/wiki/Solspiviridae" title="Solspiviridae">Solspiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Timlovirales" title="Timlovirales">Timlovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Blumeviridae" title="Blumeviridae">Blumeviridae</a></i></li> <li><i><a href="/wiki/Steitzviridae" title="Steitzviridae">Steitzviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Amabiliviricetes" class="mw-redirect" title="Amabiliviricetes">Amabiliviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Wolframvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Wolframvirales" class="mw-redirect" title="Wolframvirales">Wolframvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Narnaviridae" class="mw-redirect" title="Narnaviridae">Narnaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Howeltoviricetes" class="mw-redirect" title="Howeltoviricetes">Howeltoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Cryppavirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cryppavirales" class="mw-redirect" title="Cryppavirales">Cryppavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Mitoviridae" class="mw-redirect" title="Mitoviridae">Mitoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Miaviricetes" class="mw-redirect" title="Miaviricetes">Miaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Ourlivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ourlivirales" class="mw-redirect" title="Ourlivirales">Ourlivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Botourmiaviridae" title="Botourmiaviridae">Botourmiaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pisuviricota" title="Pisuviricota">Pisuviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pisoniviricetes" title="Pisoniviricetes">Pisoniviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Nidovirales" title="Nidovirales">Nidovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Abyssoviridae" class="mw-redirect" title="Abyssoviridae">Abyssoviridae</a></i></li> <li><i><a href="/wiki/Arteriviridae" title="Arteriviridae">Arteriviridae</a></i></li> <li><i><a href="/w/index.php?title=Cremegaviridae&action=edit&redlink=1" class="new" title="Cremegaviridae (page does not exist)">Cremegaviridae</a></i></li> <li><i><a href="/wiki/Coronaviridae" title="Coronaviridae">Coronaviridae</a></i></li> <li><i><a href="/w/index.php?title=Euroniviridae&action=edit&redlink=1" class="new" title="Euroniviridae (page does not exist)">Euroniviridae</a></i></li> <li><i><a href="/w/index.php?title=Gresnaviridae&action=edit&redlink=1" class="new" title="Gresnaviridae (page does not exist)">Gresnaviridae</a></i></li> <li><i><a href="/w/index.php?title=Medioniviridae&action=edit&redlink=1" class="new" title="Medioniviridae (page does not exist)">Medioniviridae</a></i></li> <li><i><a href="/wiki/Mesoniviridae" title="Mesoniviridae">Mesoniviridae</a></i></li> <li><i><a href="/wiki/Mononiviridae" class="mw-redirect" title="Mononiviridae">Mononiviridae</a></i></li> <li><i><a href="/w/index.php?title=Nanghoshaviridae&action=edit&redlink=1" class="new" title="Nanghoshaviridae (page does not exist)">Nanghoshaviridae</a></i></li> <li><i><a href="/w/index.php?title=Nanhypoviridae&action=edit&redlink=1" class="new" title="Nanhypoviridae (page does not exist)">Nanhypoviridae</a></i></li> <li><i><a href="/w/index.php?title=Olifoviridae&action=edit&redlink=1" class="new" title="Olifoviridae (page does not exist)">Olifoviridae</a></i></li> <li><i><a href="/wiki/Roniviridae" class="mw-redirect" title="Roniviridae">Roniviridae</a></i></li> <li><i><a href="/wiki/Tobaniviridae" title="Tobaniviridae">Tobaniviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Picornavirales" title="Picornavirales">Picornavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Picornaviridae" class="mw-redirect" title="Picornaviridae">Picornaviridae</a></i></li> <li><i><a href="/wiki/Marnaviridae" title="Marnaviridae">Marnaviridae</a></i></li> <li><i><a href="/w/index.php?title=Solinviviridae&action=edit&redlink=1" class="new" title="Solinviviridae (page does not exist)">Solinviviridae</a></i></li> <li><i><a href="/wiki/Caliciviridae" title="Caliciviridae">Caliciviridae</a></i></li> <li><i><a href="/wiki/Iflaviridae" title="Iflaviridae">Iflaviridae</a></i></li> <li><i><a href="/wiki/Secoviridae" title="Secoviridae">Secoviridae</a></i></li> <li><i><a href="/wiki/Dicistroviridae" title="Dicistroviridae">Dicistroviridae</a></i></li> <li><i><a href="/w/index.php?title=Polycipiviridae&action=edit&redlink=1" class="new" title="Polycipiviridae (page does not exist)">Polycipiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Sobelivirales" title="Sobelivirales">Sobelivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alvernaviridae" title="Alvernaviridae">Alvernaviridae</a></i></li> <li><i><a href="/wiki/Barnaviridae" title="Barnaviridae">Barnaviridae</a></i></li> <li><i><a href="/wiki/Solemoviridae" title="Solemoviridae">Solemoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Stelpaviricetes" title="Stelpaviricetes">Stelpaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Patatavirales" class="mw-redirect" title="Patatavirales">Patatavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Potyviridae" title="Potyviridae">Potyviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Stellavirales" class="mw-redirect" title="Stellavirales">Stellavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Astroviridae" class="mw-redirect" title="Astroviridae">Astroviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><b>Families</b>: <i><a href="/wiki/Permutotetraviridae" title="Permutotetraviridae">Permutotetraviridae</a></i></li> <li><i><a href="/wiki/Sarthroviridae" class="mw-redirect" title="Sarthroviridae">Sarthroviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="V:_(–)ssRNA_viruses" style="font-size:114%;margin:0 4em">V: <a href="/wiki/Negative-sense_ssRNA_virus" class="mw-redirect" title="Negative-sense ssRNA virus">(–)ssRNA viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Riboviria" scope="row" class="navbox-group" style="width:1%"><i><a class="mw-selflink selflink">Riboviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Orthornavirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Orthornavirae" title="Orthornavirae">Orthornavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Negarnaviricota" class="mw-redirect" title="Negarnaviricota">Negarnaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Chunqiuviricetes" class="mw-redirect" title="Chunqiuviricetes">Chunqiuviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Muvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Muvirales" class="mw-redirect" title="Muvirales">Muvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Qinviridae" class="mw-redirect" title="Qinviridae">Qinviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ellioviricetes" class="mw-redirect" title="Ellioviricetes">Ellioviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Bunyavirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Bunyavirales" title="Bunyavirales">Bunyavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Cruliviridae" title="Cruliviridae">Cruliviridae</a></i></li> <li><i><a href="/wiki/Arenaviridae" class="mw-redirect" title="Arenaviridae">Arenaviridae</a></i></li> <li><i><a href="/wiki/Fimoviridae" class="mw-redirect" title="Fimoviridae">Fimoviridae</a></i></li> <li><i><a href="/wiki/Hantaviridae" title="Hantaviridae">Hantaviridae</a></i></li> <li><i><a href="/w/index.php?title=Leishbuviridae&action=edit&redlink=1" class="new" title="Leishbuviridae (page does not exist)">Leishbuviridae</a></i></li> <li><i><a href="/w/index.php?title=Mypoviridae&action=edit&redlink=1" class="new" title="Mypoviridae (page does not exist)">Mypoviridae</a></i></li> <li><i><a href="/wiki/Nairoviridae" title="Nairoviridae">Nairoviridae</a></i></li> <li><i><a href="/wiki/Peribunyaviridae" title="Peribunyaviridae">Peribunyaviridae</a></i></li> <li><i><a href="/wiki/Phasmaviridae" title="Phasmaviridae">Phasmaviridae</a></i></li> <li><i><a href="/wiki/Phenuiviridae" title="Phenuiviridae">Phenuiviridae</a></i></li> <li><i><a href="/wiki/Tospoviridae" class="mw-redirect" title="Tospoviridae">Tospoviridae</a></i></li> <li><i><a href="/w/index.php?title=Wupedeviridae&action=edit&redlink=1" class="new" title="Wupedeviridae (page does not exist)">Wupedeviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Insthoviricetes" class="mw-redirect" title="Insthoviricetes">Insthoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Articulavirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Articulavirales" title="Articulavirales">Articulavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Amnoonviridae" class="mw-redirect" title="Amnoonviridae">Amnoonviridae</a></i></li> <li><i><a href="/wiki/Orthomyxoviridae" title="Orthomyxoviridae">Orthomyxoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Milneviricetes" class="mw-redirect" title="Milneviricetes">Milneviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Serpentovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Serpentovirales" class="mw-redirect" title="Serpentovirales">Serpentovirales </a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Aspiviridae" title="Aspiviridae">Aspiviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Monjiviricetes" title="Monjiviricetes">Monjiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Jingchuvirales" title="Jingchuvirales">Jingchuvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Aliusviridae&action=edit&redlink=1" class="new" title="Aliusviridae (page does not exist)">Aliusviridae</a></i></li> <li><i><a href="/wiki/Chuviridae" title="Chuviridae">Chuviridae</a></i></li> <li><i><a href="/w/index.php?title=Crepuscuviridae&action=edit&redlink=1" class="new" title="Crepuscuviridae (page does not exist)">Crepuscuviridae</a></i></li> <li><i><a href="/w/index.php?title=Myriaviridae&action=edit&redlink=1" class="new" title="Myriaviridae (page does not exist)">Myriaviridae</a></i></li> <li><i><a href="/w/index.php?title=Natareviridae&action=edit&redlink=1" class="new" title="Natareviridae (page does not exist)">Natareviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Mononegavirales" title="Mononegavirales">Mononegavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Artoviridae" title="Artoviridae">Artoviridae</a></i></li> <li><i><a href="/wiki/Bornaviridae" title="Bornaviridae">Bornaviridae</a></i></li> <li><i><a href="/wiki/Filoviridae" title="Filoviridae">Filoviridae</a></i></li> <li><i><a href="/w/index.php?title=Lispiviridae&action=edit&redlink=1" class="new" title="Lispiviridae (page does not exist)">Lispiviridae</a></i></li> <li><i><a href="/wiki/Mymonaviridae" title="Mymonaviridae">Mymonaviridae</a></i></li> <li><i><a href="/wiki/Nyamiviridae" title="Nyamiviridae">Nyamiviridae</a></i></li> <li><i><a href="/wiki/Paramyxoviridae" title="Paramyxoviridae">Paramyxoviridae</a></i></li> <li><i><a href="/wiki/Pneumoviridae" title="Pneumoviridae">Pneumoviridae</a></i></li> <li><i><a href="/wiki/Rhabdoviridae" title="Rhabdoviridae">Rhabdoviridae</a></i></li> <li><i><a href="/wiki/Sunviridae" title="Sunviridae">Sunviridae</a></i></li> <li><i><a href="/w/index.php?title=Xinmoviridae&action=edit&redlink=1" class="new" title="Xinmoviridae (page does not exist)">Xinmoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Yunchangviricetes" class="mw-redirect" title="Yunchangviricetes">Yunchangviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Goujianvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Goujianvirales" class="mw-redirect" title="Goujianvirales">Goujianvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Yueviridae" class="mw-redirect" title="Yueviridae">Yueviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ambiviricota" title="Ambiviricota">Ambiviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Suforviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Suforviricetes" class="mw-redirect" title="Suforviricetes">Suforviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Crytulvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Crytulvirales" class="mw-redirect" title="Crytulvirales">Crytulvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Dumbiviridae&action=edit&redlink=1" class="new" title="Dumbiviridae (page does not exist)">Dumbiviridae</a></i></li> <li><i><a href="/w/index.php?title=Quambiviridae&action=edit&redlink=1" class="new" title="Quambiviridae (page does not exist)">Quambiviridae</a></i></li> <li><i><a href="/w/index.php?title=Trimbiviridae&action=edit&redlink=1" class="new" title="Trimbiviridae (page does not exist)">Trimbiviridae</a></i></li> <li><i><a href="/w/index.php?title=Unambiviridae&action=edit&redlink=1" class="new" title="Unambiviridae (page does not exist)">Unambiviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Reverse_transcribing_virus" title="Reverse transcribing virus">RT</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="VI:_ssRNA-RT_viruses" style="font-size:114%;margin:0 4em">VI: <a href="/wiki/SsRNA-RT_virus" class="mw-redirect" title="SsRNA-RT virus">ssRNA-RT viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Riboviria" scope="row" class="navbox-group" style="width:1%"><i><a class="mw-selflink selflink">Riboviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Pararnavirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pararnavirae" class="mw-redirect" title="Pararnavirae">Pararnavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Artverviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Artverviricota" class="mw-redirect" title="Artverviricota">Artverviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Revtraviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Revtraviricetes" title="Revtraviricetes">Revtraviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Ortervirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ortervirales" title="Ortervirales">Ortervirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Belpaoviridae" class="mw-redirect" title="Belpaoviridae">Belpaoviridae</a></i></li> <li><i><a href="/wiki/Metaviridae" title="Metaviridae">Metaviridae</a></i></li> <li><i><a href="/wiki/Pseudoviridae" title="Pseudoviridae">Pseudoviridae</a></i></li> <li><i><a href="/wiki/Retroviridae" class="mw-redirect" title="Retroviridae">Retroviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="VII:_dsDNA-RT_viruses" style="font-size:114%;margin:0 4em">VII: <a href="/wiki/DsDNA-RT_virus" class="mw-redirect" title="DsDNA-RT virus">dsDNA-RT viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Riboviria" scope="row" class="navbox-group" style="width:1%"><i><a class="mw-selflink selflink">Riboviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Pararnavirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pararnavirae" class="mw-redirect" title="Pararnavirae">Pararnavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Artverviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Artverviricota" class="mw-redirect" title="Artverviricota">Artverviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Revtraviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Revtraviricetes" title="Revtraviricetes">Revtraviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Blubervirales" class="mw-redirect" title="Blubervirales">Blubervirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Hepadnaviridae" title="Hepadnaviridae">Hepadnaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ortervirales" title="Ortervirales">Ortervirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Caulimoviridae" title="Caulimoviridae">Caulimoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Riboviria</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q62002503" class="extiw" title="wikidata:Q62002503">Q62002503</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Riboviria" class="extiw" title="wikispecies:Riboviria">Riboviria</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Barcode_of_Life_Data_System" title="Barcode of Life Data System">BOLD</a>: <span class="uid"><a rel="nofollow" class="external text" href="http://www.boldsystems.org/index.php/TaxBrowser_TaxonPage?taxid=980903">980903</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/58840353">58840353</a></span></span></li></ul> </div></td></tr></tbody></table></div> <!-- NewPP limit report Parsed by mw‐web.codfw.main‐f69cdc8f6‐dhbcv Cached time: 20241124120212 Cache expiry: 2592000 Reduced expiry: false Complications: [vary‐revision‐sha1, show‐toc] CPU time usage: 1.160 seconds Real time usage: 1.403 seconds Preprocessor visited node count: 5675/1000000 Post‐expand include size: 800830/2097152 bytes Template argument size: 2457/2097152 bytes Highest expansion depth: 16/100 Expensive parser function count: 5/500 Unstrip recursion depth: 1/20 Unstrip post‐expand size: 176639/5000000 bytes Lua time usage: 0.717/10.000 seconds Lua memory usage: 8070254/52428800 bytes Number of Wikibase entities loaded: 3/400 --> <!-- Transclusion expansion time report 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