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Michael K Oliver | Peabody museum of Natural History - Academia.edu
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u-displayNone">less</div></div></div><div class="ri-section"><div class="ri-section-header"><span>Interests</span><a class="ri-more-link js-profile-ri-list-card" data-click-track="profile-user-info-primary-research-interest" data-has-card-for-ri-list="1462379">View All (13)</a></div><div class="ri-tags-container"><a data-click-track="profile-user-info-expand-research-interests" data-has-card-for-ri-list="1462379" href="https://www.academia.edu/Documents/in/Systematics_Taxonomy_"><div id="js-react-on-rails-context" style="display:none" data-rails-context="{"inMailer":false,"i18nLocale":"en","i18nDefaultLocale":"en","href":"https://99main.academia.edu/MichaelKOliver","location":"/MichaelKOliver","scheme":"https","host":"99main.academia.edu","port":null,"pathname":"/MichaelKOliver","search":null,"httpAcceptLanguage":null,"serverSide":false}"></div> <div class="js-react-on-rails-component" style="display:none" data-component-name="Pill" data-props="{"color":"gray","children":["Systematics 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Documents"><span>2</span> <span class="ds2-5-body-sm-bold">Teaching Documents</span></a></li></ul></div><div class="divider ds-divider-16" style="margin: 0px;"></div><div class="documents-container backbone-social-profile-documents" style="width: 100%;"><div class="u-taCenter"></div><div class="profile--tab_content_container js-tab-pane tab-pane active" id="all"><div class="profile--tab_heading_container js-section-heading" data-section="Papers" id="Papers"><h3 class="profile--tab_heading_container">Papers by Michael K Oliver</h3></div><div class="js-work-strip profile--work_container" data-work-id="124895759"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/124895759/African_cichlid_fishes_morphological_data_and_taxonomic_insights_from_a_genus_level_survey_of_supraneurals_pterygiophores_and_vertebral_counts_Ovalentaria_Blenniiformes_Cichlidae_Pseudocrenilabrinae"><img alt="Research paper thumbnail of African cichlid fishes: morphological data and taxonomic insights from a genus-level survey of supraneurals, pterygiophores, and vertebral counts (Ovalentaria, Blenniiformes, Cichlidae, Pseudocrenilabrinae" class="work-thumbnail" src="https://attachments.academia-assets.com/119035927/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/124895759/African_cichlid_fishes_morphological_data_and_taxonomic_insights_from_a_genus_level_survey_of_supraneurals_pterygiophores_and_vertebral_counts_Ovalentaria_Blenniiformes_Cichlidae_Pseudocrenilabrinae">African cichlid fishes: morphological data and taxonomic insights from a genus-level survey of supraneurals, pterygiophores, and vertebral counts (Ovalentaria, Blenniiformes, Cichlidae, Pseudocrenilabrinae</a></div><div class="wp-workCard_item"><span>Biodiversity Data Journal</span><span>, 2024</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Background The iconic freshwater cichlid fishes (Cichlidae) comprise about 1750 validly named sp...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Background <br />The iconic freshwater cichlid fishes (Cichlidae) comprise about 1750 validly named species and hundreds more that are known, but not yet described and named. Cichlids are an important source of protein for millions of people on several continents, are model organisms in studies of evolution, speciation, ecology, development, behaviour and physiology and are popular as aquarium fishes. Yet, comparative studies of cichlid internal anatomy are rare. Even their osteology has not been taxonomically surveyed. The cichlid postcranial skeleton has been especially neglected. <br /><br />New information<br />Here, I provide the first survey in cichlids of the considerable variation in numbers of vertebrae, supraneurals and dorsal- and anal-fin supports (pterygiophores), as well as the patterns with which the pterygiophores insert between the neural or haemal spines. The study includes some 1700 specimens of nearly 400 cichlid species. Focusing on the largest subfamily, the African cichlids or Pseudocrenilabrinae, the survey furnishes data from species in all but one of its 166 genera. Limited data from species in the other cichlid subfamilies (Etroplinae, Ptychochrominae and Cichlinae) and from the related leaffishes, Polycentridae, are also presented. Key examples of pterygiophore insertion patterns from<br />throughout the range of variation are illustrated and discussed. Detailed analytical tables and all raw data are provided in supplementary files.<br /><br />A bizarre specialisation in Cyprichromis is noted, evidently for the first time. Uniquely in this Lake Tanganyikan genus, five to seven anal pterygiophores are abdominal in position, located anterior to the anal fin and inserting toward or between successive pairs of pleural ribs.<br /><br />Taxonomic changes: The most speciose tribe of African cichlids, currently known as Haplochromini, is correctly called Pseudocrenilabrini. Based chiefly on the molecular phylogenetic findings of other workers, I propose four pseudocrenilabrine subtribes, one occurring in rivers and three endemic to Lake Malawi. I also re-assign the Lake Tanganyikan tribe Tropheini as another subtribe of Pseudocrenilabrini, in line with numerous molecular studies placing tropheines firmly within this tribe. The remaining genera of Pseudocrenilabrini remain incertae sedis in this tribe pending clarification of their phylogenetic relationships.<br /><br />The character complex here surveyed is a promising source of taxonomically and phylogenetically informative characteristics distinguishing or uniting cichlid taxa at multiple hierarchical levels, from species through subfamily. This reference set of novel character data can also provide information for palaeontological studies of African cichlids. These attributes are skeletal features potentially available for study in well<br />preserved fossils and may help determine their correct taxonomic placement.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="436067345e244c255c70e7bb499cde96" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":119035927,"asset_id":124895759,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/119035927/download_file?st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="124895759"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="124895759"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 124895759; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=124895759]").text(description); $(".js-view-count[data-work-id=124895759]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 124895759; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='124895759']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 124895759, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "436067345e244c255c70e7bb499cde96" } } $('.js-work-strip[data-work-id=124895759]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":124895759,"title":"African cichlid fishes: morphological data and taxonomic insights from a genus-level survey of supraneurals, pterygiophores, and vertebral counts (Ovalentaria, Blenniiformes, Cichlidae, Pseudocrenilabrinae","translated_title":"","metadata":{"doi":"10.3897/BDJ.12.e130707","abstract":"Background \nThe iconic freshwater cichlid fishes (Cichlidae) comprise about 1750 validly named species and hundreds more that are known, but not yet described and named. Cichlids are an important source of protein for millions of people on several continents, are model organisms in studies of evolution, speciation, ecology, development, behaviour and physiology and are popular as aquarium fishes. Yet, comparative studies of cichlid internal anatomy are rare. Even their osteology has not been taxonomically surveyed. The cichlid postcranial skeleton has been especially neglected. \n\nNew information\nHere, I provide the first survey in cichlids of the considerable variation in numbers of vertebrae, supraneurals and dorsal- and anal-fin supports (pterygiophores), as well as the patterns with which the pterygiophores insert between the neural or haemal spines. The study includes some 1700 specimens of nearly 400 cichlid species. Focusing on the largest subfamily, the African cichlids or Pseudocrenilabrinae, the survey furnishes data from species in all but one of its 166 genera. Limited data from species in the other cichlid subfamilies (Etroplinae, Ptychochrominae and Cichlinae) and from the related leaffishes, Polycentridae, are also presented. Key examples of pterygiophore insertion patterns from\nthroughout the range of variation are illustrated and discussed. Detailed analytical tables and all raw data are provided in supplementary files.\n\nA bizarre specialisation in Cyprichromis is noted, evidently for the first time. Uniquely in this Lake Tanganyikan genus, five to seven anal pterygiophores are abdominal in position, located anterior to the anal fin and inserting toward or between successive pairs of pleural ribs.\n\nTaxonomic changes: The most speciose tribe of African cichlids, currently known as Haplochromini, is correctly called Pseudocrenilabrini. Based chiefly on the molecular phylogenetic findings of other workers, I propose four pseudocrenilabrine subtribes, one occurring in rivers and three endemic to Lake Malawi. I also re-assign the Lake Tanganyikan tribe Tropheini as another subtribe of Pseudocrenilabrini, in line with numerous molecular studies placing tropheines firmly within this tribe. The remaining genera of Pseudocrenilabrini remain incertae sedis in this tribe pending clarification of their phylogenetic relationships.\n\nThe character complex here surveyed is a promising source of taxonomically and phylogenetically informative characteristics distinguishing or uniting cichlid taxa at multiple hierarchical levels, from species through subfamily. This reference set of novel character data can also provide information for palaeontological studies of African cichlids. These attributes are skeletal features potentially available for study in well\npreserved fossils and may help determine their correct taxonomic placement.","publication_date":{"day":null,"month":null,"year":2024,"errors":{}},"publication_name":"Biodiversity Data Journal"},"translated_abstract":"Background \nThe iconic freshwater cichlid fishes (Cichlidae) comprise about 1750 validly named species and hundreds more that are known, but not yet described and named. Cichlids are an important source of protein for millions of people on several continents, are model organisms in studies of evolution, speciation, ecology, development, behaviour and physiology and are popular as aquarium fishes. Yet, comparative studies of cichlid internal anatomy are rare. Even their osteology has not been taxonomically surveyed. The cichlid postcranial skeleton has been especially neglected. \n\nNew information\nHere, I provide the first survey in cichlids of the considerable variation in numbers of vertebrae, supraneurals and dorsal- and anal-fin supports (pterygiophores), as well as the patterns with which the pterygiophores insert between the neural or haemal spines. The study includes some 1700 specimens of nearly 400 cichlid species. Focusing on the largest subfamily, the African cichlids or Pseudocrenilabrinae, the survey furnishes data from species in all but one of its 166 genera. Limited data from species in the other cichlid subfamilies (Etroplinae, Ptychochrominae and Cichlinae) and from the related leaffishes, Polycentridae, are also presented. Key examples of pterygiophore insertion patterns from\nthroughout the range of variation are illustrated and discussed. Detailed analytical tables and all raw data are provided in supplementary files.\n\nA bizarre specialisation in Cyprichromis is noted, evidently for the first time. Uniquely in this Lake Tanganyikan genus, five to seven anal pterygiophores are abdominal in position, located anterior to the anal fin and inserting toward or between successive pairs of pleural ribs.\n\nTaxonomic changes: The most speciose tribe of African cichlids, currently known as Haplochromini, is correctly called Pseudocrenilabrini. Based chiefly on the molecular phylogenetic findings of other workers, I propose four pseudocrenilabrine subtribes, one occurring in rivers and three endemic to Lake Malawi. I also re-assign the Lake Tanganyikan tribe Tropheini as another subtribe of Pseudocrenilabrini, in line with numerous molecular studies placing tropheines firmly within this tribe. 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$(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") }); </script> <div class="js-work-strip profile--work_container" data-work-id="78685166"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/78685166/Carabids_previously_unreported_from_Connecticut_Coleoptera_Carabidae_"><img alt="Research paper thumbnail of Carabids previously unreported from Connecticut (Coleoptera: Carabidae)" class="work-thumbnail" src="https://a.academia-assets.com/images/blank-paper.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/78685166/Carabids_previously_unreported_from_Connecticut_Coleoptera_Carabidae_">Carabids previously unreported from Connecticut (Coleoptera: Carabidae)</a></div><div class="wp-workCard_item wp-workCard--coauthors"><span>by </span><span><a class="" data-click-track="profile-work-strip-authors" href="https://independent.academia.edu/WilliamKrinsky">William Krinsky</a> and <a class="" data-click-track="profile-work-strip-authors" href="https://99main.academia.edu/MichaelKOliver">Michael K Oliver</a></span></div><div class="wp-workCard_item"><span>The Coleopterists' Bulletin</span><span>, 1988</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">... Pk., W. shore L. Wintergreen, 12.IV. 1987, under rock on moist soil, MKO and Charles M. Olive...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">... Pk., W. shore L. Wintergreen, 12.IV. 1987, under rock on moist soil, MKO and Charles M. Oliver, 1 a (MKOC). ... LITERATURE CITED BELL, RT 1960. A revision of the genus Chlaenius Bonelli (Coleoptera: Carabidae) in North America. Misc. Publ. Entomol. Soc. 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This brings to 335 th...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Forty-six carabid species are reported from Connecticut for the first time. 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This brings to 335 th...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Forty-six carabid species are reported from Connecticut for the first time. This brings to 335 the number of carabid species known from Connecticut.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="76472234"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="76472234"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 76472234; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=76472234]").text(description); $(".js-view-count[data-work-id=76472234]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 76472234; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='76472234']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 76472234, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (false){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "-1" } } $('.js-work-strip[data-work-id=76472234]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":76472234,"title":"Connecticut (Coleoptera: Carabidae)","translated_title":"","metadata":{"abstract":"Forty-six carabid species are reported from Connecticut for the first time. 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This brings to 335 the number of carabid species known from Connecticut.","internal_url":"https://www.academia.edu/76472234/Connecticut_Coleoptera_Carabidae_","translated_internal_url":"","created_at":"2022-04-14T17:59:37.936-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":1462379,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[{"id":42576568,"work_id":76472234,"tagging_user_id":1462379,"tagged_user_id":70741506,"co_author_invite_id":null,"email":"w***y@gmail.com","display_order":0,"name":"William Krinsky","title":"Connecticut (Coleoptera: Carabidae)"}],"downloadable_attachments":[],"slug":"Connecticut_Coleoptera_Carabidae_","translated_slug":"","page_count":null,"language":"en","content_type":"Work","owner":{"id":1462379,"first_name":"Michael","middle_initials":"K","last_name":"Oliver","page_name":"MichaelKOliver","domain_name":"99main","created_at":"2012-04-07T13:04:06.491-07:00","display_name":"Michael K Oliver","url":"https://99main.academia.edu/MichaelKOliver"},"attachments":[],"research_interests":[{"id":261,"name":"Geography","url":"https://www.academia.edu/Documents/in/Geography"}],"urls":[]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") }); </script> <div class="js-work-strip profile--work_container" data-work-id="76472233"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/76472233/Hemitaeniochromis_brachyrhynchus_a_new_species_of_cichlid_fish_from_Lake_Mala%C5%B5i_with_comments_on_some_other_supposed_members_of_the_genus_Teleostei_Cichlidae_"><img alt="Research paper thumbnail of Hemitaeniochromis brachyrhynchus, a new species of cichlid fish from Lake Malaŵi, with comments on some other supposed members of the genus (Teleostei: Cichlidae)" class="work-thumbnail" src="https://attachments.academia-assets.com/84170289/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/76472233/Hemitaeniochromis_brachyrhynchus_a_new_species_of_cichlid_fish_from_Lake_Mala%C5%B5i_with_comments_on_some_other_supposed_members_of_the_genus_Teleostei_Cichlidae_">Hemitaeniochromis brachyrhynchus, a new species of cichlid fish from Lake Malaŵi, with comments on some other supposed members of the genus (Teleostei: Cichlidae)</a></div><div class="wp-workCard_item"><span>Zootaxa</span><span>, 2012</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Hemitaeniochromis brachyrhynchus, an anatomically distinctive and apparently rare new cichlid, is...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Hemitaeniochromis brachyrhynchus, an anatomically distinctive and apparently rare new cichlid, is described and illus-trated from specimens collected at two widely separated localities within Lake Malaŵi. It is easily distinguished from H.urotaenia, type species of the genus, by its narrow lacrimal bone which is only one-third of the orbit length, a characterthought to be unique not only in Hemitaeniochromis but among all known Lake Malaŵi cichlids. The genus Hemitaenio-chromis Eccles & Trewavas (1989) is redefined to allow provisional inclusion of this new species. Two species placed inProtomelas by Eccles & Trewavas (1989) [P. insignis (Trewavas) and P. spilopterus (Trewavas)] were recently transferredto Hemitaeniochromis by some authors, without much evidence. The generic placement of these controversial taxa, and of several undescribed species known only from underwater photographs, is briefly reconsidered.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="1c9ff539c05684f01602398e905d7412" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":84170289,"asset_id":76472233,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/84170289/download_file?st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="76472233"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="76472233"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 76472233; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=76472233]").text(description); $(".js-view-count[data-work-id=76472233]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 76472233; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='76472233']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 76472233, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "1c9ff539c05684f01602398e905d7412" } } $('.js-work-strip[data-work-id=76472233]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":76472233,"title":"Hemitaeniochromis brachyrhynchus, a new species of cichlid fish from Lake Malaŵi, with comments on some other supposed members of the genus (Teleostei: Cichlidae)","translated_title":"","metadata":{"abstract":"Hemitaeniochromis brachyrhynchus, an anatomically distinctive and apparently rare new cichlid, is described and illus-trated from specimens collected at two widely separated localities within Lake Malaŵi. 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The generic placement of these controversial taxa, and of several undescribed species known only from underwater photographs, is briefly reconsidered.","publisher":"Magnolia Press","publication_date":{"day":null,"month":null,"year":2012,"errors":{}},"publication_name":"Zootaxa"},"translated_abstract":"Hemitaeniochromis brachyrhynchus, an anatomically distinctive and apparently rare new cichlid, is described and illus-trated from specimens collected at two widely separated localities within Lake Malaŵi. It is easily distinguished from H.urotaenia, type species of the genus, by its narrow lacrimal bone which is only one-third of the orbit length, a characterthought to be unique not only in Hemitaeniochromis but among all known Lake Malaŵi cichlids. The genus Hemitaenio-chromis Eccles \u0026 Trewavas (1989) is redefined to allow provisional inclusion of this new species. 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The generic placement of these controversial taxa, and of several undescribed species known only from underwater photographs, is briefly reconsidered.","internal_url":"https://www.academia.edu/76472233/Hemitaeniochromis_brachyrhynchus_a_new_species_of_cichlid_fish_from_Lake_Mala%C5%B5i_with_comments_on_some_other_supposed_members_of_the_genus_Teleostei_Cichlidae_","translated_internal_url":"","created_at":"2022-04-14T17:59:37.767-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":1462379,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[{"id":84170289,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/84170289/thumbnails/1.jpg","file_name":"43377.pdf","download_url":"https://www.academia.edu/attachments/84170289/download_file?st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"Hemitaeniochromis_brachyrhynchus_a_new_s.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/84170289/43377-libre.pdf?1649985879=\u0026response-content-disposition=attachment%3B+filename%3DHemitaeniochromis_brachyrhynchus_a_new_s.pdf\u0026Expires=1732422414\u0026Signature=KP5kao0BLFwAvQM6eDS40LcjN1Ci4p9M~7ZYgVeWmo8VH51jLHfXYHkQFsW-6E0bxpEi2WTMgqOuJfZQAUBijqEZ-nVQbMBtUEQfK1-PN2OeSQNo5C9pS44gl2V1zRugtkX6SEN-eQq9FdrEge9md3xz3geBCFsY~VTDHQwMQUVJKR4~i~-eKmGWr~M2e~dF6Eq4zCqv2-Z582oKqsv5iI6eOzgVxgDgepDUHfD0sKaZSvs4x~57KvKkXGIDXzUnLCtLc-4MXObEJtyXrb6N6NZBxj4~-8v~bUNo5upfSO7wr9~mtEPUAzDuFxpqJ3FuDiM-mTi-VzaCmpA4GTHzHA__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"Hemitaeniochromis_brachyrhynchus_a_new_species_of_cichlid_fish_from_Lake_Malaŵi_with_comments_on_some_other_supposed_members_of_the_genus_Teleostei_Cichlidae_","translated_slug":"","page_count":16,"language":"en","content_type":"Work","owner":{"id":1462379,"first_name":"Michael","middle_initials":"K","last_name":"Oliver","page_name":"MichaelKOliver","domain_name":"99main","created_at":"2012-04-07T13:04:06.491-07:00","display_name":"Michael K Oliver","url":"https://99main.academia.edu/MichaelKOliver"},"attachments":[{"id":84170289,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/84170289/thumbnails/1.jpg","file_name":"43377.pdf","download_url":"https://www.academia.edu/attachments/84170289/download_file?st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"Hemitaeniochromis_brachyrhynchus_a_new_s.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/84170289/43377-libre.pdf?1649985879=\u0026response-content-disposition=attachment%3B+filename%3DHemitaeniochromis_brachyrhynchus_a_new_s.pdf\u0026Expires=1732422414\u0026Signature=KP5kao0BLFwAvQM6eDS40LcjN1Ci4p9M~7ZYgVeWmo8VH51jLHfXYHkQFsW-6E0bxpEi2WTMgqOuJfZQAUBijqEZ-nVQbMBtUEQfK1-PN2OeSQNo5C9pS44gl2V1zRugtkX6SEN-eQq9FdrEge9md3xz3geBCFsY~VTDHQwMQUVJKR4~i~-eKmGWr~M2e~dF6Eq4zCqv2-Z582oKqsv5iI6eOzgVxgDgepDUHfD0sKaZSvs4x~57KvKkXGIDXzUnLCtLc-4MXObEJtyXrb6N6NZBxj4~-8v~bUNo5upfSO7wr9~mtEPUAzDuFxpqJ3FuDiM-mTi-VzaCmpA4GTHzHA__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"},{"id":84170288,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/84170288/thumbnails/1.jpg","file_name":"43377.pdf","download_url":"https://www.academia.edu/attachments/84170288/download_file","bulk_download_file_name":"Hemitaeniochromis_brachyrhynchus_a_new_s.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/84170288/43377-libre.pdf?1649985879=\u0026response-content-disposition=attachment%3B+filename%3DHemitaeniochromis_brachyrhynchus_a_new_s.pdf\u0026Expires=1732422414\u0026Signature=Wl~Z8GYAh~iyBDhrDY~cvqnFUCb0~fnKwnMwqmhTJ3SCm9Lyf3n1wdvWtBH33zkDa1xR0JgGm9k-cl5crQSJn0Jwcxw-bu375zNslntrBHS~ND75fQEFFOwoj1E~FfW9NzxZJs7e7KXAZwur1uGG58EvekqufYpRov3XzgDlpkxmatIIujYec1nvFrs~0V-cGuqGMmgCndSbBk-tCLbCAPvmLqz2A6uDsT~IUQTHKpk1QOJoOFFjYGeljip8Tm6IwHR68P8-nahYpXRj2DVjzUQmwHjf2jAyl3GLpfiooskOdviFAoTbIz-yZSyo-XI2qj223CUlOSgSeAUlo4WihQ__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[{"id":155,"name":"Evolutionary Biology","url":"https://www.academia.edu/Documents/in/Evolutionary_Biology"},{"id":173,"name":"Zoology","url":"https://www.academia.edu/Documents/in/Zoology"},{"id":7076,"name":"Taxonomy","url":"https://www.academia.edu/Documents/in/Taxonomy"},{"id":7710,"name":"Biology","url":"https://www.academia.edu/Documents/in/Biology"},{"id":31192,"name":"Teleostei","url":"https://www.academia.edu/Documents/in/Teleostei"},{"id":216604,"name":"Cichlidae","url":"https://www.academia.edu/Documents/in/Cichlidae"}],"urls":[{"id":19491886,"url":"https://biotaxa.org/Zootaxa/article/viewFile/49993/43377"}]}, dispatcherData: dispatcherData }); 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$(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") }); </script> <div class="js-work-strip profile--work_container" data-work-id="76472231"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/76472231/A_Kink_in_the_Line_Does_a_Unique_Lateral_Line_Peculiarity_Really_Characterize_Lake_Mala%C5%B5is_Huge_Haplochromine_Species_Flock_Teleostei_Cichlidae_"><img alt="Research paper thumbnail of A Kink in the Line: Does a Unique Lateral-Line Peculiarity Really Characterize Lake Malaŵi's Huge Haplochromine Species Flock (Teleostei: Cichlidae)?" class="work-thumbnail" src="https://a.academia-assets.com/images/blank-paper.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/76472231/A_Kink_in_the_Line_Does_a_Unique_Lateral_Line_Peculiarity_Really_Characterize_Lake_Mala%C5%B5is_Huge_Haplochromine_Species_Flock_Teleostei_Cichlidae_">A Kink in the Line: Does a Unique Lateral-Line Peculiarity Really Characterize Lake Malaŵi's Huge Haplochromine Species Flock (Teleostei: Cichlidae)?</a></div><div class="wp-workCard_item"><span>Bulletin of the Peabody Museum of Natural History</span><span>, 2016</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Abstract A downward kink one or more scales in length at the posterior end of the upper lateral l...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Abstract A downward kink one or more scales in length at the posterior end of the upper lateral line, leaving only one scale row instead of two between it and the lower lateral line, has been considered a synapomorphy uniting as a monophyletic group all members of both clades of endemic haplochromine cichlids of Lake Malaŵi. Here, this claim is tested by surveying the character in representatives of all genera of Lake Malaŵi haplochromines (tribe Haplochromini of the subfamily Pseudocrenilabrinae). Presence of the kink is shown to be variable in most genera, and it was not documented at all in two (Abactochromis, Cyrtocara). Specimens with no kink occur in many genera, and kink presence and length frequently vary on the two sides of the body. Thus, it is difficult to justify the kink as a synapomorphy of the species flock. The two clades differ in kink frequency; a kink occurred on at least one side of the body in 79% of bilaterally assessable specimens in the non-mbuna clade, but in only 53% of those in the mbuna clade (p &lt; 0.0001). When a kink is present, its length is more variable in non-mbuna (1 to 8 scales, mode 2) than mbuna (1 to 4 scales, mode 1). Extension of the upper lateral line backward from the upper flank close to or onto the caudal peduncle, where no scale row is as dorsally situated, is identified as one cause of kink existence; additional factors likely await discovery. Contrary to published statements, the kink is expressed in some specimens of Astatotilapia calliptera, a phylogenetically important nonendemic haplochromine found in Lake Malaŵi and elsewhere, which may be the sister of the lake&#39;s endemic flock. An apparently identical lateral-line kink occurs sporadically in haplochromines in other lakes and in members of three other pseudocrenilabrine cichlid tribes, further refuting the hypothesis that this character is a synapomorphy of the Lake Malaŵi species flock.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="76472231"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="76472231"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 76472231; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=76472231]").text(description); $(".js-view-count[data-work-id=76472231]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 76472231; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='76472231']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 76472231, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (false){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "-1" } } $('.js-work-strip[data-work-id=76472231]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":76472231,"title":"A Kink in the Line: Does a Unique Lateral-Line Peculiarity Really Characterize Lake Malaŵi's Huge Haplochromine Species Flock (Teleostei: Cichlidae)?","translated_title":"","metadata":{"abstract":"Abstract A downward kink one or more scales in length at the posterior end of the upper lateral line, leaving only one scale row instead of two between it and the lower lateral line, has been considered a synapomorphy uniting as a monophyletic group all members of both clades of endemic haplochromine cichlids of Lake Malaŵi. Here, this claim is tested by surveying the character in representatives of all genera of Lake Malaŵi haplochromines (tribe Haplochromini of the subfamily Pseudocrenilabrinae). Presence of the kink is shown to be variable in most genera, and it was not documented at all in two (Abactochromis, Cyrtocara). Specimens with no kink occur in many genera, and kink presence and length frequently vary on the two sides of the body. Thus, it is difficult to justify the kink as a synapomorphy of the species flock. The two clades differ in kink frequency; a kink occurred on at least one side of the body in 79% of bilaterally assessable specimens in the non-mbuna clade, but in only 53% of those in the mbuna clade (p \u0026lt; 0.0001). When a kink is present, its length is more variable in non-mbuna (1 to 8 scales, mode 2) than mbuna (1 to 4 scales, mode 1). Extension of the upper lateral line backward from the upper flank close to or onto the caudal peduncle, where no scale row is as dorsally situated, is identified as one cause of kink existence; additional factors likely await discovery. Contrary to published statements, the kink is expressed in some specimens of Astatotilapia calliptera, a phylogenetically important nonendemic haplochromine found in Lake Malaŵi and elsewhere, which may be the sister of the lake\u0026#39;s endemic flock. An apparently identical lateral-line kink occurs sporadically in haplochromines in other lakes and in members of three other pseudocrenilabrine cichlid tribes, further refuting the hypothesis that this character is a synapomorphy of the Lake Malaŵi species flock.","publisher":"Bulletin of the Peabody Museum of Natural History","publication_date":{"day":null,"month":null,"year":2016,"errors":{}},"publication_name":"Bulletin of the Peabody Museum of Natural History"},"translated_abstract":"Abstract A downward kink one or more scales in length at the posterior end of the upper lateral line, leaving only one scale row instead of two between it and the lower lateral line, has been considered a synapomorphy uniting as a monophyletic group all members of both clades of endemic haplochromine cichlids of Lake Malaŵi. Here, this claim is tested by surveying the character in representatives of all genera of Lake Malaŵi haplochromines (tribe Haplochromini of the subfamily Pseudocrenilabrinae). Presence of the kink is shown to be variable in most genera, and it was not documented at all in two (Abactochromis, Cyrtocara). Specimens with no kink occur in many genera, and kink presence and length frequently vary on the two sides of the body. Thus, it is difficult to justify the kink as a synapomorphy of the species flock. The two clades differ in kink frequency; a kink occurred on at least one side of the body in 79% of bilaterally assessable specimens in the non-mbuna clade, but in only 53% of those in the mbuna clade (p \u0026lt; 0.0001). When a kink is present, its length is more variable in non-mbuna (1 to 8 scales, mode 2) than mbuna (1 to 4 scales, mode 1). Extension of the upper lateral line backward from the upper flank close to or onto the caudal peduncle, where no scale row is as dorsally situated, is identified as one cause of kink existence; additional factors likely await discovery. Contrary to published statements, the kink is expressed in some specimens of Astatotilapia calliptera, a phylogenetically important nonendemic haplochromine found in Lake Malaŵi and elsewhere, which may be the sister of the lake\u0026#39;s endemic flock. An apparently identical lateral-line kink occurs sporadically in haplochromines in other lakes and in members of three other pseudocrenilabrine cichlid tribes, further refuting the hypothesis that this character is a synapomorphy of the Lake Malaŵi species flock.","internal_url":"https://www.academia.edu/76472231/A_Kink_in_the_Line_Does_a_Unique_Lateral_Line_Peculiarity_Really_Characterize_Lake_Mala%C5%B5is_Huge_Haplochromine_Species_Flock_Teleostei_Cichlidae_","translated_internal_url":"","created_at":"2022-04-14T17:59:37.409-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":1462379,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[],"slug":"A_Kink_in_the_Line_Does_a_Unique_Lateral_Line_Peculiarity_Really_Characterize_Lake_Malaŵis_Huge_Haplochromine_Species_Flock_Teleostei_Cichlidae_","translated_slug":"","page_count":null,"language":"en","content_type":"Work","owner":{"id":1462379,"first_name":"Michael","middle_initials":"K","last_name":"Oliver","page_name":"MichaelKOliver","domain_name":"99main","created_at":"2012-04-07T13:04:06.491-07:00","display_name":"Michael K Oliver","url":"https://99main.academia.edu/MichaelKOliver"},"attachments":[],"research_interests":[{"id":135987,"name":"Hypothesis testing","url":"https://www.academia.edu/Documents/in/Hypothesis_testing"},{"id":216604,"name":"Cichlidae","url":"https://www.academia.edu/Documents/in/Cichlidae"},{"id":452259,"name":"Lake Malawi","url":"https://www.academia.edu/Documents/in/Lake_Malawi"},{"id":1418924,"name":"Lateral Line","url":"https://www.academia.edu/Documents/in/Lateral_Line"}],"urls":[]}, dispatcherData: dispatcherData }); 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Diverse systematic proble...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">&quot;The cichlids of Africa present many challenges to the taxonomist. Diverse systematic problems at three levels in cichlid phylogeny form the subject matter of this dissertation. At the most general level, I summarize current morphological evidence that supports cichlid monophyly and the most-inclusive groups of cichlids. With this review as a framework, I argue that the most phylogenetically primitive African cichlid is Heterochromis multidens, a little-known large cichlid from the Zaire River basin. I redescribe this monotypic genus and species and illustrate much of its osteology. It possesses a distinctive suite of derived characters, but is uniquely primitive among African cichlids in infraorbital morphology and other features. At a less inclusive phylogenetic level, I examine the controversy over the monophyletic status of the &quot;species flock&quot; of haplochromine cichlids in Lake Malawi. A derived type of anal-fin markings present in a subset of the Malawian fauna is shared with many non-Malawian haplochromines, indicating that the species flock of Lake Malawi is not monophyletic. The two (or more) faunal components of Lake Malawi are ecologically distinct. The remaining major parts of the study are concerned with particular assemblages of haplochromine cichlids in Lake Malawi. I present a detailed phylogenetic analysis of the Cyrtocara livingstonii species-group of ambush predators, and characterize a new species in the group. I also describe three new species of haplochromines with three lateral spots and present a key to the three-spot assemblage. Intralacustrine dispersion of Malawi cichlids is briefly addressed in appended material.&quot;</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="76472228"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="76472228"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 76472228; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=76472228]").text(description); $(".js-view-count[data-work-id=76472228]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 76472228; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='76472228']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 76472228, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (false){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "-1" } } $('.js-work-strip[data-work-id=76472228]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":76472228,"title":"Systematics of African Cichlid Fishes: Determination of the Most Primitive Taxon, and Studies on the Haplochromines of Lake Malawi (Teleostei: Cichlidae)","translated_title":"","metadata":{"abstract":"\u0026quot;The cichlids of Africa present many challenges to the taxonomist. 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Pk., W. shore L. Wintergreen, 12.IV. 1987, under rock on moist soil, MKO and Charles M. Olive...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">... Pk., W. shore L. Wintergreen, 12.IV. 1987, under rock on moist soil, MKO and Charles M. Oliver, 1 a (MKOC). ... LITERATURE CITED BELL, RT 1960. A revision of the genus Chlaenius Bonelli (Coleoptera: Carabidae) in North America. Misc. Publ. Entomol. Soc. 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He based one of them, L. joanjohnsonae, on a holotype and paratype belonging to two species that differ strikingly from each other in coloration and oral dentition. Although he gave conflicting statements both of standard length and total length of the paratype in different paragraphs, Johnson (1974a: 15) clearly designated the smaller of the two specimens as the holotype, and since they differ in standard length by nearly 10 mrn there is no doubt as to which this is. The number of dorsal fin spines (sixteen in the holotype, seventeen in the paratype) corroborates his designation. It might be noted, however, that Johnson's counts of segmented dorsal rays are inaccurate for both specimens. I have examined the types of all species described by Johnson (1974a and 1974b) and found that the holotype of L. joanjohnsonue Johnson is a specimen of L. fryeri Oliver in Davies (1973) ; L. ioanjohnsonae is therefore a junior synonym of that species. The manuscript name Labidochromis fryeri and a diagnosis of this taxon were unfortunately published in an aquarium magazine in a letter from T. E. (Peter) Davies, who specifically (and correctly) attributed to me the name, the diagnosis, and my intention eventually to publish them. Regrettably, by Articles 11 and 13 (a) (i) of the International Labidochromis textilis, new species Brocade Cichlid","publication_date":{"day":null,"month":null,"year":1975,"errors":{}},"grobid_abstract_attachment_id":79834848},"translated_abstract":null,"internal_url":"https://www.academia.edu/69923161/Labidochromis_textilis_a_new_cichlid_fish_Teleostei_Cichlidae_from_Lake_Malawi","translated_internal_url":"","created_at":"2022-01-29T07:22:34.262-08:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":1462379,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[{"id":79834848,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/79834848/thumbnails/1.jpg","file_name":"oliver_1975.pdf","download_url":"https://www.academia.edu/attachments/79834848/download_file?st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"Labidochromis_textilis_a_new_cichlid_fis.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/79834848/oliver_1975-libre.pdf?1643473844=\u0026response-content-disposition=attachment%3B+filename%3DLabidochromis_textilis_a_new_cichlid_fis.pdf\u0026Expires=1732422414\u0026Signature=HIor6lHUL4caezDsGyqzC27-7cX1E4hRTX69YR1AZM~ouf0pGRCWLcG-GozGURgcyQ7xADlDVebhkmTnoQdJNQE9beIDvEbxlD6VWbLXrAJt62U26DZAzFiGgvRVYDYghlvIXcNVvxR054HmxI-1sT-MToOOipvAp5jkUdb2wUznX9~kHuuLwUb3KJSN11Lyn6grjN6Wx62i0aHaU-OAlJxucKCaSupJFDv7ZSVrkOmMn8gASOcIPNcUKuQN5MVyA6fqr8Ls~Ly5IJiSfQnZZeY8RCZ6JXukWsBfAM8B68ycDIANJwoLsv90qMZQbjPODVtRMHISTdFELSYqR4B4mA__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"Labidochromis_textilis_a_new_cichlid_fish_Teleostei_Cichlidae_from_Lake_Malawi","translated_slug":"","page_count":12,"language":"en","content_type":"Work","owner":{"id":1462379,"first_name":"Michael","middle_initials":"K","last_name":"Oliver","page_name":"MichaelKOliver","domain_name":"99main","created_at":"2012-04-07T13:04:06.491-07:00","display_name":"Michael K Oliver","url":"https://99main.academia.edu/MichaelKOliver"},"attachments":[{"id":79834848,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/79834848/thumbnails/1.jpg","file_name":"oliver_1975.pdf","download_url":"https://www.academia.edu/attachments/79834848/download_file?st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"Labidochromis_textilis_a_new_cichlid_fis.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/79834848/oliver_1975-libre.pdf?1643473844=\u0026response-content-disposition=attachment%3B+filename%3DLabidochromis_textilis_a_new_cichlid_fis.pdf\u0026Expires=1732422414\u0026Signature=HIor6lHUL4caezDsGyqzC27-7cX1E4hRTX69YR1AZM~ouf0pGRCWLcG-GozGURgcyQ7xADlDVebhkmTnoQdJNQE9beIDvEbxlD6VWbLXrAJt62U26DZAzFiGgvRVYDYghlvIXcNVvxR054HmxI-1sT-MToOOipvAp5jkUdb2wUznX9~kHuuLwUb3KJSN11Lyn6grjN6Wx62i0aHaU-OAlJxucKCaSupJFDv7ZSVrkOmMn8gASOcIPNcUKuQN5MVyA6fqr8Ls~Ly5IJiSfQnZZeY8RCZ6JXukWsBfAM8B68ycDIANJwoLsv90qMZQbjPODVtRMHISTdFELSYqR4B4mA__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[{"id":169,"name":"Systematics (Taxonomy)","url":"https://www.academia.edu/Documents/in/Systematics_Taxonomy_"},{"id":7710,"name":"Biology","url":"https://www.academia.edu/Documents/in/Biology"},{"id":216604,"name":"Cichlidae","url":"https://www.academia.edu/Documents/in/Cichlidae"}],"urls":[]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") }); </script> <div class="js-work-strip profile--work_container" data-work-id="69922906"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/69922906/A_new_genus_for_Melanochromis_labrosus_a_problematic_Lake_Malawi_cichlid_with_hypertrophied_lips_Teleostei_Cichlidae_"><img alt="Research paper thumbnail of A new genus for Melanochromis labrosus, a problematic Lake Malawi cichlid with hypertrophied lips (Teleostei: Cichlidae)" class="work-thumbnail" src="https://attachments.academia-assets.com/79834585/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/69922906/A_new_genus_for_Melanochromis_labrosus_a_problematic_Lake_Malawi_cichlid_with_hypertrophied_lips_Teleostei_Cichlidae_">A new genus for Melanochromis labrosus, a problematic Lake Malawi cichlid with hypertrophied lips (Teleostei: Cichlidae)</a></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Melanochromis labrosus is the only one of the 800 or so endemic cichlid species in Lake Malawi wh...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Melanochromis labrosus is the only one of the 800 or so endemic cichlid species in Lake Malawi whose placement with mbuna or non-mbuna has been disputed on morphological grounds. Here, we redescribe M. labrosus using all obtainable specimens collected since its original description. In addition, we extensively survey vertebral counts in the mbuna. To determine if M. labrosus belongs with the mbuna, we review the morphological traits often considered to define the mbuna as a natural group, concluding that none is a uniquely shared specialization. Two features of body coloration (a relatively small number of ocellate spots in the anal fin and a possible polychromatism in one population) suggest that M. labrosus is best considered a species of mbuna. Although its placement within the mbuna group remains somewhat equivocal, we unambiguously demonstrate that M. labrosus is not a Melanochromis, since it lacks the synapomorphic striped color pattern typical of that genus. Melanochromis lab...</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="f670b6a1f7043405db9e0fc9aed1b3d0" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":79834585,"asset_id":69922906,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/79834585/download_file?st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="69922906"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="69922906"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 69922906; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=69922906]").text(description); $(".js-view-count[data-work-id=69922906]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 69922906; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='69922906']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 69922906, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "f670b6a1f7043405db9e0fc9aed1b3d0" } } $('.js-work-strip[data-work-id=69922906]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":69922906,"title":"A new genus for Melanochromis labrosus, a problematic Lake Malawi cichlid with hypertrophied lips (Teleostei: Cichlidae)","translated_title":"","metadata":{"abstract":"Melanochromis labrosus is the only one of the 800 or so endemic cichlid species in Lake Malawi whose placement with mbuna or non-mbuna has been disputed on morphological grounds. Here, we redescribe M. labrosus using all obtainable specimens collected since its original description. In addition, we extensively survey vertebral counts in the mbuna. To determine if M. labrosus belongs with the mbuna, we review the morphological traits often considered to define the mbuna as a natural group, concluding that none is a uniquely shared specialization. Two features of body coloration (a relatively small number of ocellate spots in the anal fin and a possible polychromatism in one population) suggest that M. labrosus is best considered a species of mbuna. Although its placement within the mbuna group remains somewhat equivocal, we unambiguously demonstrate that M. labrosus is not a Melanochromis, since it lacks the synapomorphic striped color pattern typical of that genus. Melanochromis lab...","publication_date":{"day":null,"month":null,"year":2010,"errors":{}}},"translated_abstract":"Melanochromis labrosus is the only one of the 800 or so endemic cichlid species in Lake Malawi whose placement with mbuna or non-mbuna has been disputed on morphological grounds. Here, we redescribe M. labrosus using all obtainable specimens collected since its original description. In addition, we extensively survey vertebral counts in the mbuna. To determine if M. labrosus belongs with the mbuna, we review the morphological traits often considered to define the mbuna as a natural group, concluding that none is a uniquely shared specialization. Two features of body coloration (a relatively small number of ocellate spots in the anal fin and a possible polychromatism in one population) suggest that M. labrosus is best considered a species of mbuna. Although its placement within the mbuna group remains somewhat equivocal, we unambiguously demonstrate that M. labrosus is not a Melanochromis, since it lacks the synapomorphic striped color pattern typical of that genus. Melanochromis lab...","internal_url":"https://www.academia.edu/69922906/A_new_genus_for_Melanochromis_labrosus_a_problematic_Lake_Malawi_cichlid_with_hypertrophied_lips_Teleostei_Cichlidae_","translated_internal_url":"","created_at":"2022-01-29T07:20:10.360-08:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":1462379,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[{"id":37626419,"work_id":69922906,"tagging_user_id":1462379,"tagged_user_id":null,"co_author_invite_id":7440491,"email":"m***r@fhcrc.org","display_order":0,"name":"Matthew Arnegard","title":"A new genus for Melanochromis labrosus, a problematic Lake Malawi cichlid with hypertrophied lips (Teleostei: Cichlidae)"}],"downloadable_attachments":[{"id":79834585,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/79834585/thumbnails/1.jpg","file_name":"oliver_arnegard_2010.pdf","download_url":"https://www.academia.edu/attachments/79834585/download_file?st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"A_new_genus_for_Melanochromis_labrosus_a.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/79834585/oliver_arnegard_2010-libre.pdf?1643469709=\u0026response-content-disposition=attachment%3B+filename%3DA_new_genus_for_Melanochromis_labrosus_a.pdf\u0026Expires=1732422414\u0026Signature=OVD1vxpl5~MKcfapqWws6pDmNQIfBi89a9olT62OPIjb24r2qCsxpkVq-cbBAl9CoYG4Mljbdbg3taNZPURjv2yGCrSlYeZGTSOBgzDj9hV9-0lzQRTVYPYe2m8VjWsMXODCh7KTbKeHWX6N7yOP2WCdJIaFvS7UhUCTyr1pqmNuLEpza6ijbS3RwlgC-6Plxgw2o9NA9zM-pGkX6i0KAGTU~BQpKg0C-CAkRA0w1VF2bMT4I25mgg1BzpXKG1OW8rOTlOT4yU1KELA1qXrMx7Ju-IK9FJLV1CBB9gXQUsdbx9QqE8Sg5lH-Oik0fF616vc0e68KTJSw-UrrTlI4vA__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"A_new_genus_for_Melanochromis_labrosus_a_problematic_Lake_Malawi_cichlid_with_hypertrophied_lips_Teleostei_Cichlidae_","translated_slug":"","page_count":24,"language":"en","content_type":"Work","owner":{"id":1462379,"first_name":"Michael","middle_initials":"K","last_name":"Oliver","page_name":"MichaelKOliver","domain_name":"99main","created_at":"2012-04-07T13:04:06.491-07:00","display_name":"Michael K Oliver","url":"https://99main.academia.edu/MichaelKOliver"},"attachments":[{"id":79834585,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/79834585/thumbnails/1.jpg","file_name":"oliver_arnegard_2010.pdf","download_url":"https://www.academia.edu/attachments/79834585/download_file?st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"A_new_genus_for_Melanochromis_labrosus_a.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/79834585/oliver_arnegard_2010-libre.pdf?1643469709=\u0026response-content-disposition=attachment%3B+filename%3DA_new_genus_for_Melanochromis_labrosus_a.pdf\u0026Expires=1732422414\u0026Signature=OVD1vxpl5~MKcfapqWws6pDmNQIfBi89a9olT62OPIjb24r2qCsxpkVq-cbBAl9CoYG4Mljbdbg3taNZPURjv2yGCrSlYeZGTSOBgzDj9hV9-0lzQRTVYPYe2m8VjWsMXODCh7KTbKeHWX6N7yOP2WCdJIaFvS7UhUCTyr1pqmNuLEpza6ijbS3RwlgC-6Plxgw2o9NA9zM-pGkX6i0KAGTU~BQpKg0C-CAkRA0w1VF2bMT4I25mgg1BzpXKG1OW8rOTlOT4yU1KELA1qXrMx7Ju-IK9FJLV1CBB9gXQUsdbx9QqE8Sg5lH-Oik0fF616vc0e68KTJSw-UrrTlI4vA__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"},{"id":79834587,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/79834587/thumbnails/1.jpg","file_name":"oliver_arnegard_2010.pdf","download_url":"https://www.academia.edu/attachments/79834587/download_file","bulk_download_file_name":"A_new_genus_for_Melanochromis_labrosus_a.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/79834587/oliver_arnegard_2010-libre.pdf?1643469708=\u0026response-content-disposition=attachment%3B+filename%3DA_new_genus_for_Melanochromis_labrosus_a.pdf\u0026Expires=1732422414\u0026Signature=EPOhYKz8hp4fH2pd8vlH56zOh4tBxgSpqXYPj-BrQigF3wti2miBYJv~KAIQpBLK5Ig3KjSNuMFwUTFl3Z03wV0WeOzWWGBSE38vKLo4YpLaKmf7ZBt8UYRbpuA7m9mNYQg2Adr6hZ5c4~qMkUNQfLJe9UaYCwdHl~lKj5gatBFvW6xjTpQvHNx6rHEA51uBzgBAjiAJySrWTeTIXZUGIWJ2A-~q02Eb5oDNRmPZCYPr~62by9034Mzk9s4sE4QIoEFXjfVMKMNQAVwqqTpOfRxcl9anukIdpcxmhrIzWgLIRZFeuYlahyhIb9ZuvQMZC8J4recRIfymYtmdCmURvg__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[{"id":7076,"name":"Taxonomy","url":"https://www.academia.edu/Documents/in/Taxonomy"},{"id":7710,"name":"Biology","url":"https://www.academia.edu/Documents/in/Biology"},{"id":10225,"name":"Agriculture","url":"https://www.academia.edu/Documents/in/Agriculture"},{"id":216604,"name":"Cichlidae","url":"https://www.academia.edu/Documents/in/Cichlidae"}],"urls":[{"id":17075021,"url":"http://malawicichlids.com/oliver_arnegard_2010.pdf"}]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") }); </script> <div class="js-work-strip profile--work_container" data-work-id="37695426"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/37695426/Six_New_Species_of_the_Cichlid_Genus_Otopharynx_from_Lake_Mala%C5%B5i_Teleostei_Cichlidae"><img alt="Research paper thumbnail of Six New Species of the Cichlid Genus Otopharynx from Lake Malaŵi (Teleostei: Cichlidae" class="work-thumbnail" src="https://attachments.academia-assets.com/57686794/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/37695426/Six_New_Species_of_the_Cichlid_Genus_Otopharynx_from_Lake_Mala%C5%B5i_Teleostei_Cichlidae">Six New Species of the Cichlid Genus Otopharynx from Lake Malaŵi (Teleostei: Cichlidae</a></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">The endemic Lake Malaŵi cichlid genus Otopharynx is increased from 14 species to 20 by the descri...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">The endemic Lake Malaŵi cichlid genus Otopharynx is increased from 14 species to 20 by the description of six new species collected from inshore sandy or intermediate rocky/sandy areas or trawled in depths to 59 m. Otopharynx alpha is distinguished from congeners by its unique melanin pattern with suprapectoral spot discrete but supraanal and precaudal spots connected to form a stripe, and by its distinctively shaped pharyngeal bone with numerous small, crowded teeth. Otopharynx mumboensis, thought to be endemic to Mumbo Island, is separated from most congeners by a suprapectoral spot placed entirely below the upper lateral line, and from all others by the combination of 13 gill rakers, pharyngeal bone with posterior margin nearly straight, crowns of all pharyngeal teeth in the median columns broadened and flattened, head length 30.9–32.2% SL, body depth 35.5–36.1% SL, and lower jaw 35.7–36.2% HL. Otopharynx styrax has a more elongate body (depth 24.3–28.6% SL) than any congener. Otopharynx aletes can be diagnosed by the combination of a suprapectoral spot overlapping the upper lateral line, seven or eight vertical bars below the dorsal-fin base, 10 or 11 gill rakers, 34–36 scales in the lateral line, lower pharyngeal bone with molariform posteromedian teeth, and caudal peduncle length 1.6–2.0 times its depth. Otopharynx panniculus is diagnosed by the combination of a prominent square suprapectoral spot spanning subdorsal bars 3 to 4, no dorsal midline spots, 13–15 gill rakers, 31–33 scales in the lateral line, 15 or 16 dorsal-fin spines, absence of a lacrimal notch, and slightly<br />enlarged teeth in the median columns of the lower pharyngeal bone. Otopharynx peridodeka most closely resembles O. panniculus and shares the loss of the lacrimal notch, but has 34–35 scales in the lateral line and 11–13 gill rakers. Photographs of living or freshly collected specimens of five of the new species are provided.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="dd20a3e5fea564f5b0028ad88c64cb9c" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":57686794,"asset_id":37695426,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/57686794/download_file?st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="37695426"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="37695426"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 37695426; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=37695426]").text(description); $(".js-view-count[data-work-id=37695426]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 37695426; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='37695426']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 37695426, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "dd20a3e5fea564f5b0028ad88c64cb9c" } } $('.js-work-strip[data-work-id=37695426]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":37695426,"title":"Six New Species of the Cichlid Genus Otopharynx from Lake Malaŵi (Teleostei: Cichlidae","translated_title":"","metadata":{"abstract":"The endemic Lake Malaŵi cichlid genus Otopharynx is increased from 14 species to 20 by the description of six new species collected from inshore sandy or intermediate rocky/sandy areas or trawled in depths to 59 m. Otopharynx alpha is distinguished from congeners by its unique melanin pattern with suprapectoral spot discrete but supraanal and precaudal spots connected to form a stripe, and by its distinctively shaped pharyngeal bone with numerous small, crowded teeth. Otopharynx mumboensis, thought to be endemic to Mumbo Island, is separated from most congeners by a suprapectoral spot placed entirely below the upper lateral line, and from all others by the combination of 13 gill rakers, pharyngeal bone with posterior margin nearly straight, crowns of all pharyngeal teeth in the median columns broadened and flattened, head length 30.9–32.2% SL, body depth 35.5–36.1% SL, and lower jaw 35.7–36.2% HL. Otopharynx styrax has a more elongate body (depth 24.3–28.6% SL) than any congener. Otopharynx aletes can be diagnosed by the combination of a suprapectoral spot overlapping the upper lateral line, seven or eight vertical bars below the dorsal-fin base, 10 or 11 gill rakers, 34–36 scales in the lateral line, lower pharyngeal bone with molariform posteromedian teeth, and caudal peduncle length 1.6–2.0 times its depth. Otopharynx panniculus is diagnosed by the combination of a prominent square suprapectoral spot spanning subdorsal bars 3 to 4, no dorsal midline spots, 13–15 gill rakers, 31–33 scales in the lateral line, 15 or 16 dorsal-fin spines, absence of a lacrimal notch, and slightly\nenlarged teeth in the median columns of the lower pharyngeal bone. Otopharynx peridodeka most closely resembles O. panniculus and shares the loss of the lacrimal notch, but has 34–35 scales in the lateral line and 11–13 gill rakers. Photographs of living or freshly collected specimens of five of the new species are provided."},"translated_abstract":"The endemic Lake Malaŵi cichlid genus Otopharynx is increased from 14 species to 20 by the description of six new species collected from inshore sandy or intermediate rocky/sandy areas or trawled in depths to 59 m. Otopharynx alpha is distinguished from congeners by its unique melanin pattern with suprapectoral spot discrete but supraanal and precaudal spots connected to form a stripe, and by its distinctively shaped pharyngeal bone with numerous small, crowded teeth. Otopharynx mumboensis, thought to be endemic to Mumbo Island, is separated from most congeners by a suprapectoral spot placed entirely below the upper lateral line, and from all others by the combination of 13 gill rakers, pharyngeal bone with posterior margin nearly straight, crowns of all pharyngeal teeth in the median columns broadened and flattened, head length 30.9–32.2% SL, body depth 35.5–36.1% SL, and lower jaw 35.7–36.2% HL. Otopharynx styrax has a more elongate body (depth 24.3–28.6% SL) than any congener. Otopharynx aletes can be diagnosed by the combination of a suprapectoral spot overlapping the upper lateral line, seven or eight vertical bars below the dorsal-fin base, 10 or 11 gill rakers, 34–36 scales in the lateral line, lower pharyngeal bone with molariform posteromedian teeth, and caudal peduncle length 1.6–2.0 times its depth. Otopharynx panniculus is diagnosed by the combination of a prominent square suprapectoral spot spanning subdorsal bars 3 to 4, no dorsal midline spots, 13–15 gill rakers, 31–33 scales in the lateral line, 15 or 16 dorsal-fin spines, absence of a lacrimal notch, and slightly\nenlarged teeth in the median columns of the lower pharyngeal bone. Otopharynx peridodeka most closely resembles O. panniculus and shares the loss of the lacrimal notch, but has 34–35 scales in the lateral line and 11–13 gill rakers. Photographs of living or freshly collected specimens of five of the new species are provided.","internal_url":"https://www.academia.edu/37695426/Six_New_Species_of_the_Cichlid_Genus_Otopharynx_from_Lake_Mala%C5%B5i_Teleostei_Cichlidae","translated_internal_url":"","created_at":"2018-11-03T07:44:53.145-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":1462379,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[{"id":57686794,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/57686794/thumbnails/1.jpg","file_name":"Oliver_2018_Six_New_Species_of_the_Cichlid_Genus_Otopharynx_from_Lake_Malawi_Teleostei-_Cichlidae.pdf","download_url":"https://www.academia.edu/attachments/57686794/download_file?st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"Six_New_Species_of_the_Cichlid_Genus_Oto.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/57686794/Oliver_2018_Six_New_Species_of_the_Cichlid_Genus_Otopharynx_from_Lake_Malawi_Teleostei-_Cichlidae-libre.pdf?1541256795=\u0026response-content-disposition=attachment%3B+filename%3DSix_New_Species_of_the_Cichlid_Genus_Oto.pdf\u0026Expires=1732422414\u0026Signature=aYy7EuFzebSn3VLr3B3tbQUXRc3H2RVaeEUeHflnvrdznvcCRHIzNp9d87YrVbm0If95hKz3I~T2OGfanClhQBBKEk9tC~Ks-dDPfoEMtu2ePqPCjboC0n88sXom8JZJ3fkGXH8k134ec48eT7kL3ala7qYrxE3EQIbQbxmSwscX3ff9tm5m72N1XJ5WD3rcFVRi1BBm2WEJzM4By~zV1U8tgH-CEz~-uJids7ZTQKGIJDgqWmZrxKpNqcxeekAW7aDDqKNqO3259ABvjuZhqM5IwhMC21j~RocYJEEzurfy2sc9n6Kx2GPkHzmuW3DnooSS~WVvQejF3jYelp5gTQ__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"Six_New_Species_of_the_Cichlid_Genus_Otopharynx_from_Lake_Malaŵi_Teleostei_Cichlidae","translated_slug":"","page_count":39,"language":"en","content_type":"Work","owner":{"id":1462379,"first_name":"Michael","middle_initials":"K","last_name":"Oliver","page_name":"MichaelKOliver","domain_name":"99main","created_at":"2012-04-07T13:04:06.491-07:00","display_name":"Michael K Oliver","url":"https://99main.academia.edu/MichaelKOliver"},"attachments":[{"id":57686794,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/57686794/thumbnails/1.jpg","file_name":"Oliver_2018_Six_New_Species_of_the_Cichlid_Genus_Otopharynx_from_Lake_Malawi_Teleostei-_Cichlidae.pdf","download_url":"https://www.academia.edu/attachments/57686794/download_file?st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"Six_New_Species_of_the_Cichlid_Genus_Oto.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/57686794/Oliver_2018_Six_New_Species_of_the_Cichlid_Genus_Otopharynx_from_Lake_Malawi_Teleostei-_Cichlidae-libre.pdf?1541256795=\u0026response-content-disposition=attachment%3B+filename%3DSix_New_Species_of_the_Cichlid_Genus_Oto.pdf\u0026Expires=1732422414\u0026Signature=aYy7EuFzebSn3VLr3B3tbQUXRc3H2RVaeEUeHflnvrdznvcCRHIzNp9d87YrVbm0If95hKz3I~T2OGfanClhQBBKEk9tC~Ks-dDPfoEMtu2ePqPCjboC0n88sXom8JZJ3fkGXH8k134ec48eT7kL3ala7qYrxE3EQIbQbxmSwscX3ff9tm5m72N1XJ5WD3rcFVRi1BBm2WEJzM4By~zV1U8tgH-CEz~-uJids7ZTQKGIJDgqWmZrxKpNqcxeekAW7aDDqKNqO3259ABvjuZhqM5IwhMC21j~RocYJEEzurfy2sc9n6Kx2GPkHzmuW3DnooSS~WVvQejF3jYelp5gTQ__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[{"id":151753,"name":"New Species","url":"https://www.academia.edu/Documents/in/New_Species"},{"id":211405,"name":"Endemic species","url":"https://www.academia.edu/Documents/in/Endemic_species"},{"id":243830,"name":"Bottom Trawling","url":"https://www.academia.edu/Documents/in/Bottom_Trawling"},{"id":452259,"name":"Lake Malawi","url":"https://www.academia.edu/Documents/in/Lake_Malawi"}],"urls":[]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") }); </script> <div class="js-work-strip profile--work_container" data-work-id="24075167"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/24075167/A_Kink_in_the_Line_Does_a_Unique_Lateral_Line_Peculiarity_Really_Characterize_Lake_Malawis_Huge_Haplochromine_Species_Flock_Teleostei_Cichlidae_"><img alt="Research paper thumbnail of A Kink in the Line: Does a Unique Lateral-Line Peculiarity Really Characterize Lake Malawi's Huge Haplochromine Species Flock (Teleostei: Cichlidae)?" class="work-thumbnail" src="https://attachments.academia-assets.com/44445264/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/24075167/A_Kink_in_the_Line_Does_a_Unique_Lateral_Line_Peculiarity_Really_Characterize_Lake_Malawis_Huge_Haplochromine_Species_Flock_Teleostei_Cichlidae_">A Kink in the Line: Does a Unique Lateral-Line Peculiarity Really Characterize Lake Malawi's Huge Haplochromine Species Flock (Teleostei: Cichlidae)?</a></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">A downward kink one or more scales in length at the posterior end of the upper lateral line, leav...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">A downward kink one or more scales in length at the posterior end of the upper lateral line, leaving only one scale row instead of two between it and the lower lateral line, has been considered a synapomorphy uniting as a monophyletic group all members of both clades of endemic haplochromine cichlids of Lake Malawi. Here, this claim is tested by surveying the character in representatives of all genera of Lake Malawi haplochromines (tribe Haplochromini of the subfamily Pseudocrenilabrinae). Presence of the kink is shown to be variable in most genera, and it was not documented at all in two (Abactochromis, Cyrtocara). Specimens with no kink occur in many genera, and kink presence and length frequently vary on the two sides of the body. Thus, it is difficult to justify the kink as a synapomorphy of the species flock. The two clades differ in kink frequency; a kink occurred on at least one side of the body in 79% of bilaterally assessable specimens in the non-mbuna clade, but in only 53% of those in the mbuna clade (p < 0.0001). When a kink is present, its length is more variable in non-mbuna (1 to 8 scales, mode 2) than mbuna (1 to 4 scales, mode 1). Extension of the upper lateral line backward from the upper flank close to or onto the caudal peduncle, where no scale row is as dorsally situated, is identified as one cause of kink existence; additional factors likely await discovery. Contrary to published statements, the kink is expressed in some specimens of Astatotilapia calliptera, a phylogenetically important nonendemic haplochromine found in Lake Malawi and elsewhere, which may be the sister of the lake's endemic flock. An apparently identical lateral-line kink occurs sporadically in haplochromines in other lakes and in members of three other pseudocrenilabrine cichlid tribes, further refuting the hypothesis that this character is a synapomorphy of the Lake Malawi species flock.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="0fddbcd241fc04aac1884328c968f993" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":44445264,"asset_id":24075167,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/44445264/download_file?st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="24075167"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="24075167"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 24075167; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=24075167]").text(description); $(".js-view-count[data-work-id=24075167]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 24075167; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='24075167']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 24075167, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "0fddbcd241fc04aac1884328c968f993" } } $('.js-work-strip[data-work-id=24075167]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":24075167,"title":"A Kink in the Line: Does a Unique Lateral-Line Peculiarity Really Characterize Lake Malawi's Huge Haplochromine Species Flock (Teleostei: Cichlidae)?","translated_title":"","metadata":{"abstract":"A downward kink one or more scales in length at the posterior end of the upper lateral line, leaving only one scale row instead of two between it and the lower lateral line, has been considered a synapomorphy uniting as a monophyletic group all members of both clades of endemic haplochromine cichlids of Lake Malawi. Here, this claim is tested by surveying the character in representatives of all genera of Lake Malawi haplochromines (tribe Haplochromini of the subfamily Pseudocrenilabrinae). Presence of the kink is shown to be variable in most genera, and it was not documented at all in two (Abactochromis, Cyrtocara). Specimens with no kink occur in many genera, and kink presence and length frequently vary on the two sides of the body. Thus, it is difficult to justify the kink as a synapomorphy of the species flock. The two clades differ in kink frequency; a kink occurred on at least one side of the body in 79% of bilaterally assessable specimens in the non-mbuna clade, but in only 53% of those in the mbuna clade (p \u003c 0.0001). When a kink is present, its length is more variable in non-mbuna (1 to 8 scales, mode 2) than mbuna (1 to 4 scales, mode 1). Extension of the upper lateral line backward from the upper flank close to or onto the caudal peduncle, where no scale row is as dorsally situated, is identified as one cause of kink existence; additional factors likely await discovery. Contrary to published statements, the kink is expressed in some specimens of Astatotilapia calliptera, a phylogenetically important nonendemic haplochromine found in Lake Malawi and elsewhere, which may be the sister of the lake's endemic flock. An apparently identical lateral-line kink occurs sporadically in haplochromines in other lakes and in members of three other pseudocrenilabrine cichlid tribes, further refuting the hypothesis that this character is a synapomorphy of the Lake Malawi species flock."},"translated_abstract":"A downward kink one or more scales in length at the posterior end of the upper lateral line, leaving only one scale row instead of two between it and the lower lateral line, has been considered a synapomorphy uniting as a monophyletic group all members of both clades of endemic haplochromine cichlids of Lake Malawi. Here, this claim is tested by surveying the character in representatives of all genera of Lake Malawi haplochromines (tribe Haplochromini of the subfamily Pseudocrenilabrinae). Presence of the kink is shown to be variable in most genera, and it was not documented at all in two (Abactochromis, Cyrtocara). Specimens with no kink occur in many genera, and kink presence and length frequently vary on the two sides of the body. Thus, it is difficult to justify the kink as a synapomorphy of the species flock. The two clades differ in kink frequency; a kink occurred on at least one side of the body in 79% of bilaterally assessable specimens in the non-mbuna clade, but in only 53% of those in the mbuna clade (p \u003c 0.0001). When a kink is present, its length is more variable in non-mbuna (1 to 8 scales, mode 2) than mbuna (1 to 4 scales, mode 1). Extension of the upper lateral line backward from the upper flank close to or onto the caudal peduncle, where no scale row is as dorsally situated, is identified as one cause of kink existence; additional factors likely await discovery. Contrary to published statements, the kink is expressed in some specimens of Astatotilapia calliptera, a phylogenetically important nonendemic haplochromine found in Lake Malawi and elsewhere, which may be the sister of the lake's endemic flock. An apparently identical lateral-line kink occurs sporadically in haplochromines in other lakes and in members of three other pseudocrenilabrine cichlid tribes, further refuting the hypothesis that this character is a synapomorphy of the Lake Malawi species flock.","internal_url":"https://www.academia.edu/24075167/A_Kink_in_the_Line_Does_a_Unique_Lateral_Line_Peculiarity_Really_Characterize_Lake_Malawis_Huge_Haplochromine_Species_Flock_Teleostei_Cichlidae_","translated_internal_url":"","created_at":"2016-04-05T12:30:18.606-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":1462379,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[{"id":44445264,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/44445264/thumbnails/1.jpg","file_name":"B57-1-01-Oliver_pp3-20.pdf","download_url":"https://www.academia.edu/attachments/44445264/download_file?st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"A_Kink_in_the_Line_Does_a_Unique_Lateral.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/44445264/B57-1-01-Oliver_pp3-20-libre.pdf?1459884527=\u0026response-content-disposition=attachment%3B+filename%3DA_Kink_in_the_Line_Does_a_Unique_Lateral.pdf\u0026Expires=1732422414\u0026Signature=U0MPLQBS8gsUJnMB-y8hbtAS-UmTCRN3hZMCkeFacacg6AyUz5DSaeESzhsLNwu0XJutHw0flM7roU0uzDx32L3EDk45Gk-Xc4WzbnbzeG9dohoxjv4s8prSsvNcWX942Ici1ZOlk8ItLDfcMM~zzTabbQhjMGXBIId60prNk3HTauttggS462CLvkYJoBJfAiB1An5rBSEGhEQlH93ImO-8yGSeCKtUOUkxF5t4wGmo0GtsGgcjsgNwbFPX5Q-dX9EQYCQ8JW-cgLbWBZxOxW10JLC~q~Bq8tGF60m8blbNu7ogeb8PFoTlYHSZz06mQNQbzRM97CjHs1ZVcdVN9Q__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"A_Kink_in_the_Line_Does_a_Unique_Lateral_Line_Peculiarity_Really_Characterize_Lake_Malawis_Huge_Haplochromine_Species_Flock_Teleostei_Cichlidae_","translated_slug":"","page_count":18,"language":"en","content_type":"Work","owner":{"id":1462379,"first_name":"Michael","middle_initials":"K","last_name":"Oliver","page_name":"MichaelKOliver","domain_name":"99main","created_at":"2012-04-07T13:04:06.491-07:00","display_name":"Michael K Oliver","url":"https://99main.academia.edu/MichaelKOliver"},"attachments":[{"id":44445264,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/44445264/thumbnails/1.jpg","file_name":"B57-1-01-Oliver_pp3-20.pdf","download_url":"https://www.academia.edu/attachments/44445264/download_file?st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"A_Kink_in_the_Line_Does_a_Unique_Lateral.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/44445264/B57-1-01-Oliver_pp3-20-libre.pdf?1459884527=\u0026response-content-disposition=attachment%3B+filename%3DA_Kink_in_the_Line_Does_a_Unique_Lateral.pdf\u0026Expires=1732422414\u0026Signature=U0MPLQBS8gsUJnMB-y8hbtAS-UmTCRN3hZMCkeFacacg6AyUz5DSaeESzhsLNwu0XJutHw0flM7roU0uzDx32L3EDk45Gk-Xc4WzbnbzeG9dohoxjv4s8prSsvNcWX942Ici1ZOlk8ItLDfcMM~zzTabbQhjMGXBIId60prNk3HTauttggS462CLvkYJoBJfAiB1An5rBSEGhEQlH93ImO-8yGSeCKtUOUkxF5t4wGmo0GtsGgcjsgNwbFPX5Q-dX9EQYCQ8JW-cgLbWBZxOxW10JLC~q~Bq8tGF60m8blbNu7ogeb8PFoTlYHSZz06mQNQbzRM97CjHs1ZVcdVN9Q__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[{"id":135987,"name":"Hypothesis testing","url":"https://www.academia.edu/Documents/in/Hypothesis_testing"},{"id":216604,"name":"Cichlidae","url":"https://www.academia.edu/Documents/in/Cichlidae"},{"id":452259,"name":"Lake Malawi","url":"https://www.academia.edu/Documents/in/Lake_Malawi"},{"id":1418924,"name":"Lateral Line","url":"https://www.academia.edu/Documents/in/Lateral_Line"}],"urls":[]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") }); </script> <div class="js-work-strip profile--work_container" data-work-id="2998160"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/2998160/Hemitaeniochromis_brachyrhynchus_a_new_species_of_cichlid_fish_from_Lake_Mala%C5%B5i_with_comments_on_some_other_supposed_members_of_the_genus_Teleostei_Cichlidae_"><img alt="Research paper thumbnail of Hemitaeniochromis brachyrhynchus, a new species of cichlid fish from Lake Malaŵi, with comments on some other supposed members of the genus (Teleostei: Cichlidae)" class="work-thumbnail" src="https://attachments.academia-assets.com/30947824/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/2998160/Hemitaeniochromis_brachyrhynchus_a_new_species_of_cichlid_fish_from_Lake_Mala%C5%B5i_with_comments_on_some_other_supposed_members_of_the_genus_Teleostei_Cichlidae_">Hemitaeniochromis brachyrhynchus, a new species of cichlid fish from Lake Malaŵi, with comments on some other supposed members of the genus (Teleostei: Cichlidae)</a></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Hemitaeniochromis brachyrhynchus, an anatomically distinctive and apparently rare new cichlid, is...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Hemitaeniochromis brachyrhynchus, an anatomically distinctive and apparently rare new cichlid, is described and illustrated <br />from specimens collected at two widely separated localities within Lake Malaŵi. It is easily distinguished from H. urotaenia, type species of the genus, by its narrow lacrimal bone which is only one-third of the orbit length, a character thought to be unique not only in Hemitaeniochromis but among all known Lake Malaŵi cichlids. The genus Hemitaeniochromis Eccles & Trewavas (1989) is redefined to allow provisional inclusion of this new species. Two species placed in Protomelas by Eccles & Trewavas (1989) [P. insignis (Trewavas) and P. spilopterus (Trewavas)] were recently transferred to Hemitaeniochromis by some authors, without much evidence. The generic placement of these controversial taxa, and <br />of several undescribed species known only from underwater photographs, is briefly reconsidered.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="04fefe6b247975531353cfff6668b3c3" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":30947824,"asset_id":2998160,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/30947824/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="2998160"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="2998160"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 2998160; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=2998160]").text(description); $(".js-view-count[data-work-id=2998160]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 2998160; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='2998160']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 2998160, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "04fefe6b247975531353cfff6668b3c3" } } $('.js-work-strip[data-work-id=2998160]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":2998160,"title":"Hemitaeniochromis brachyrhynchus, a new species of cichlid fish from Lake Malaŵi, with comments on some other supposed members of the genus (Teleostei: Cichlidae)","translated_title":"","metadata":{"abstract":"Hemitaeniochromis brachyrhynchus, an anatomically distinctive and apparently rare new cichlid, is described and illustrated\r\nfrom specimens collected at two widely separated localities within Lake Malaŵi. It is easily distinguished from H. urotaenia, type species of the genus, by its narrow lacrimal bone which is only one-third of the orbit length, a character thought to be unique not only in Hemitaeniochromis but among all known Lake Malaŵi cichlids. The genus Hemitaeniochromis Eccles \u0026 Trewavas (1989) is redefined to allow provisional inclusion of this new species. Two species placed in Protomelas by Eccles \u0026 Trewavas (1989) [P. insignis (Trewavas) and P. spilopterus (Trewavas)] were recently transferred to Hemitaeniochromis by some authors, without much evidence. The generic placement of these controversial taxa, and\r\nof several undescribed species known only from underwater photographs, is briefly reconsidered.","journal_name":"Zootaxa","publication_date":{"day":6,"month":8,"year":2012,"errors":{}}},"translated_abstract":"Hemitaeniochromis brachyrhynchus, an anatomically distinctive and apparently rare new cichlid, is described and illustrated\r\nfrom specimens collected at two widely separated localities within Lake Malaŵi. It is easily distinguished from H. urotaenia, type species of the genus, by its narrow lacrimal bone which is only one-third of the orbit length, a character thought to be unique not only in Hemitaeniochromis but among all known Lake Malaŵi cichlids. The genus Hemitaeniochromis Eccles \u0026 Trewavas (1989) is redefined to allow provisional inclusion of this new species. Two species placed in Protomelas by Eccles \u0026 Trewavas (1989) [P. insignis (Trewavas) and P. spilopterus (Trewavas)] were recently transferred to Hemitaeniochromis by some authors, without much evidence. The generic placement of these controversial taxa, and\r\nof several undescribed species known only from underwater photographs, is briefly reconsidered.","internal_url":"https://www.academia.edu/2998160/Hemitaeniochromis_brachyrhynchus_a_new_species_of_cichlid_fish_from_Lake_Mala%C5%B5i_with_comments_on_some_other_supposed_members_of_the_genus_Teleostei_Cichlidae_","translated_internal_url":"","created_at":"2013-03-14T11:34:51.595-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":1462379,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[{"id":30947824,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/30947824/thumbnails/1.jpg","file_name":"zt03410p050.pdf","download_url":"https://www.academia.edu/attachments/30947824/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"Hemitaeniochromis_brachyrhynchus_a_new_s.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/30947824/zt03410p050-libre.pdf?1392233686=\u0026response-content-disposition=attachment%3B+filename%3DHemitaeniochromis_brachyrhynchus_a_new_s.pdf\u0026Expires=1732422414\u0026Signature=Ju6mrcbHrmcBKXM-ihDlAggLC6CMtC9RpEitn1rF~gwXxdnMw7FjDz0n3OmeMDgpJ6JKV8nldsiOoJW9b32q3WnduKEhqw~MjKDRlCHd~lEZRPwPLJYxLrKzTKsY~RPZjr4i9ld-EFG3WfUEOr~7PoxkU0rCYWyaFvtpZWnTDjQ5JGzo5pjkQ3vBQJqIqk-jHaareAd2q5NOjpmtExcVkTlLXEKxZVl6tt83-vUslJPp-uHj9UtZ6BBmNOMem9A8t9CJyuUIEnM1RgGMpPADUgdyJKdQcyP7EGCxsNvCRzO8oQ2i9AhjZM7isaADdMcAQ2RNVaI8wuBUPs-UVGpXnw__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"Hemitaeniochromis_brachyrhynchus_a_new_species_of_cichlid_fish_from_Lake_Malaŵi_with_comments_on_some_other_supposed_members_of_the_genus_Teleostei_Cichlidae_","translated_slug":"","page_count":16,"language":"en","content_type":"Work","owner":{"id":1462379,"first_name":"Michael","middle_initials":"K","last_name":"Oliver","page_name":"MichaelKOliver","domain_name":"99main","created_at":"2012-04-07T13:04:06.491-07:00","display_name":"Michael K Oliver","url":"https://99main.academia.edu/MichaelKOliver"},"attachments":[{"id":30947824,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/30947824/thumbnails/1.jpg","file_name":"zt03410p050.pdf","download_url":"https://www.academia.edu/attachments/30947824/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"Hemitaeniochromis_brachyrhynchus_a_new_s.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/30947824/zt03410p050-libre.pdf?1392233686=\u0026response-content-disposition=attachment%3B+filename%3DHemitaeniochromis_brachyrhynchus_a_new_s.pdf\u0026Expires=1732422415\u0026Signature=ZCCw7-sko3Cs3gDgX1iBGh4HeNPINtnh4qEwWa78vSY~zvSy1IrGxX64tRSOrC-eINcyPY-nj1f7lwF7DCJefk7nYyhWBjcDqHpjcdapJhtSyqlG5gRN82nSjcwMu8h25IPO8R3tEMF4J4i3aMi31i0JcEjh6kbT6NFbT5HppAjy~h2SRIOycd~mQkkflClQqmcwU1Me1zDF5Goj6faDg-PjBP05Nv225TTwdOHKJH1Pa1UNJSl15Hpm4kOnpztCzA9kV0yGx5ncG8av5beJIAdlqWe9mxMPHiCEzbVLnt0i76eCvZciWJj12Oh7AOu~R2Hc9KqKrRRKNsP9uHkfow__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[{"id":7076,"name":"Taxonomy","url":"https://www.academia.edu/Documents/in/Taxonomy"},{"id":216604,"name":"Cichlidae","url":"https://www.academia.edu/Documents/in/Cichlidae"}],"urls":[{"id":841538,"url":"http://www.mapress.com/zootaxa/2012/f/zt03410p050.pdf"}]}, dispatcherData: dispatcherData }); 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$(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") }); </script> </div><div class="profile--tab_content_container js-tab-pane tab-pane" data-section-id="414235" id="papers"><div class="js-work-strip profile--work_container" data-work-id="124895759"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/124895759/African_cichlid_fishes_morphological_data_and_taxonomic_insights_from_a_genus_level_survey_of_supraneurals_pterygiophores_and_vertebral_counts_Ovalentaria_Blenniiformes_Cichlidae_Pseudocrenilabrinae"><img alt="Research paper thumbnail of African cichlid fishes: morphological data and taxonomic insights from a genus-level survey of supraneurals, pterygiophores, and vertebral counts (Ovalentaria, Blenniiformes, Cichlidae, Pseudocrenilabrinae" class="work-thumbnail" src="https://attachments.academia-assets.com/119035927/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/124895759/African_cichlid_fishes_morphological_data_and_taxonomic_insights_from_a_genus_level_survey_of_supraneurals_pterygiophores_and_vertebral_counts_Ovalentaria_Blenniiformes_Cichlidae_Pseudocrenilabrinae">African cichlid fishes: morphological data and taxonomic insights from a genus-level survey of supraneurals, pterygiophores, and vertebral counts (Ovalentaria, Blenniiformes, Cichlidae, Pseudocrenilabrinae</a></div><div class="wp-workCard_item"><span>Biodiversity Data Journal</span><span>, 2024</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Background The iconic freshwater cichlid fishes (Cichlidae) comprise about 1750 validly named sp...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Background <br />The iconic freshwater cichlid fishes (Cichlidae) comprise about 1750 validly named species and hundreds more that are known, but not yet described and named. Cichlids are an important source of protein for millions of people on several continents, are model organisms in studies of evolution, speciation, ecology, development, behaviour and physiology and are popular as aquarium fishes. Yet, comparative studies of cichlid internal anatomy are rare. Even their osteology has not been taxonomically surveyed. The cichlid postcranial skeleton has been especially neglected. <br /><br />New information<br />Here, I provide the first survey in cichlids of the considerable variation in numbers of vertebrae, supraneurals and dorsal- and anal-fin supports (pterygiophores), as well as the patterns with which the pterygiophores insert between the neural or haemal spines. The study includes some 1700 specimens of nearly 400 cichlid species. Focusing on the largest subfamily, the African cichlids or Pseudocrenilabrinae, the survey furnishes data from species in all but one of its 166 genera. Limited data from species in the other cichlid subfamilies (Etroplinae, Ptychochrominae and Cichlinae) and from the related leaffishes, Polycentridae, are also presented. Key examples of pterygiophore insertion patterns from<br />throughout the range of variation are illustrated and discussed. Detailed analytical tables and all raw data are provided in supplementary files.<br /><br />A bizarre specialisation in Cyprichromis is noted, evidently for the first time. Uniquely in this Lake Tanganyikan genus, five to seven anal pterygiophores are abdominal in position, located anterior to the anal fin and inserting toward or between successive pairs of pleural ribs.<br /><br />Taxonomic changes: The most speciose tribe of African cichlids, currently known as Haplochromini, is correctly called Pseudocrenilabrini. Based chiefly on the molecular phylogenetic findings of other workers, I propose four pseudocrenilabrine subtribes, one occurring in rivers and three endemic to Lake Malawi. I also re-assign the Lake Tanganyikan tribe Tropheini as another subtribe of Pseudocrenilabrini, in line with numerous molecular studies placing tropheines firmly within this tribe. The remaining genera of Pseudocrenilabrini remain incertae sedis in this tribe pending clarification of their phylogenetic relationships.<br /><br />The character complex here surveyed is a promising source of taxonomically and phylogenetically informative characteristics distinguishing or uniting cichlid taxa at multiple hierarchical levels, from species through subfamily. This reference set of novel character data can also provide information for palaeontological studies of African cichlids. These attributes are skeletal features potentially available for study in well<br />preserved fossils and may help determine their correct taxonomic placement.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="436067345e244c255c70e7bb499cde96" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":119035927,"asset_id":124895759,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/119035927/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="124895759"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="124895759"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 124895759; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=124895759]").text(description); $(".js-view-count[data-work-id=124895759]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 124895759; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='124895759']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 124895759, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "436067345e244c255c70e7bb499cde96" } } $('.js-work-strip[data-work-id=124895759]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":124895759,"title":"African cichlid fishes: morphological data and taxonomic insights from a genus-level survey of supraneurals, pterygiophores, and vertebral counts (Ovalentaria, Blenniiformes, Cichlidae, Pseudocrenilabrinae","translated_title":"","metadata":{"doi":"10.3897/BDJ.12.e130707","abstract":"Background \nThe iconic freshwater cichlid fishes (Cichlidae) comprise about 1750 validly named species and hundreds more that are known, but not yet described and named. Cichlids are an important source of protein for millions of people on several continents, are model organisms in studies of evolution, speciation, ecology, development, behaviour and physiology and are popular as aquarium fishes. Yet, comparative studies of cichlid internal anatomy are rare. Even their osteology has not been taxonomically surveyed. The cichlid postcranial skeleton has been especially neglected. \n\nNew information\nHere, I provide the first survey in cichlids of the considerable variation in numbers of vertebrae, supraneurals and dorsal- and anal-fin supports (pterygiophores), as well as the patterns with which the pterygiophores insert between the neural or haemal spines. The study includes some 1700 specimens of nearly 400 cichlid species. Focusing on the largest subfamily, the African cichlids or Pseudocrenilabrinae, the survey furnishes data from species in all but one of its 166 genera. Limited data from species in the other cichlid subfamilies (Etroplinae, Ptychochrominae and Cichlinae) and from the related leaffishes, Polycentridae, are also presented. Key examples of pterygiophore insertion patterns from\nthroughout the range of variation are illustrated and discussed. Detailed analytical tables and all raw data are provided in supplementary files.\n\nA bizarre specialisation in Cyprichromis is noted, evidently for the first time. Uniquely in this Lake Tanganyikan genus, five to seven anal pterygiophores are abdominal in position, located anterior to the anal fin and inserting toward or between successive pairs of pleural ribs.\n\nTaxonomic changes: The most speciose tribe of African cichlids, currently known as Haplochromini, is correctly called Pseudocrenilabrini. Based chiefly on the molecular phylogenetic findings of other workers, I propose four pseudocrenilabrine subtribes, one occurring in rivers and three endemic to Lake Malawi. I also re-assign the Lake Tanganyikan tribe Tropheini as another subtribe of Pseudocrenilabrini, in line with numerous molecular studies placing tropheines firmly within this tribe. The remaining genera of Pseudocrenilabrini remain incertae sedis in this tribe pending clarification of their phylogenetic relationships.\n\nThe character complex here surveyed is a promising source of taxonomically and phylogenetically informative characteristics distinguishing or uniting cichlid taxa at multiple hierarchical levels, from species through subfamily. This reference set of novel character data can also provide information for palaeontological studies of African cichlids. These attributes are skeletal features potentially available for study in well\npreserved fossils and may help determine their correct taxonomic placement.","publication_date":{"day":null,"month":null,"year":2024,"errors":{}},"publication_name":"Biodiversity Data Journal"},"translated_abstract":"Background \nThe iconic freshwater cichlid fishes (Cichlidae) comprise about 1750 validly named species and hundreds more that are known, but not yet described and named. Cichlids are an important source of protein for millions of people on several continents, are model organisms in studies of evolution, speciation, ecology, development, behaviour and physiology and are popular as aquarium fishes. Yet, comparative studies of cichlid internal anatomy are rare. Even their osteology has not been taxonomically surveyed. The cichlid postcranial skeleton has been especially neglected. \n\nNew information\nHere, I provide the first survey in cichlids of the considerable variation in numbers of vertebrae, supraneurals and dorsal- and anal-fin supports (pterygiophores), as well as the patterns with which the pterygiophores insert between the neural or haemal spines. The study includes some 1700 specimens of nearly 400 cichlid species. Focusing on the largest subfamily, the African cichlids or Pseudocrenilabrinae, the survey furnishes data from species in all but one of its 166 genera. Limited data from species in the other cichlid subfamilies (Etroplinae, Ptychochrominae and Cichlinae) and from the related leaffishes, Polycentridae, are also presented. Key examples of pterygiophore insertion patterns from\nthroughout the range of variation are illustrated and discussed. Detailed analytical tables and all raw data are provided in supplementary files.\n\nA bizarre specialisation in Cyprichromis is noted, evidently for the first time. Uniquely in this Lake Tanganyikan genus, five to seven anal pterygiophores are abdominal in position, located anterior to the anal fin and inserting toward or between successive pairs of pleural ribs.\n\nTaxonomic changes: The most speciose tribe of African cichlids, currently known as Haplochromini, is correctly called Pseudocrenilabrini. Based chiefly on the molecular phylogenetic findings of other workers, I propose four pseudocrenilabrine subtribes, one occurring in rivers and three endemic to Lake Malawi. I also re-assign the Lake Tanganyikan tribe Tropheini as another subtribe of Pseudocrenilabrini, in line with numerous molecular studies placing tropheines firmly within this tribe. The remaining genera of Pseudocrenilabrini remain incertae sedis in this tribe pending clarification of their phylogenetic relationships.\n\nThe character complex here surveyed is a promising source of taxonomically and phylogenetically informative characteristics distinguishing or uniting cichlid taxa at multiple hierarchical levels, from species through subfamily. This reference set of novel character data can also provide information for palaeontological studies of African cichlids. 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$(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") }); </script> <div class="js-work-strip profile--work_container" data-work-id="78685166"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/78685166/Carabids_previously_unreported_from_Connecticut_Coleoptera_Carabidae_"><img alt="Research paper thumbnail of Carabids previously unreported from Connecticut (Coleoptera: Carabidae)" class="work-thumbnail" src="https://a.academia-assets.com/images/blank-paper.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/78685166/Carabids_previously_unreported_from_Connecticut_Coleoptera_Carabidae_">Carabids previously unreported from Connecticut (Coleoptera: Carabidae)</a></div><div class="wp-workCard_item wp-workCard--coauthors"><span>by </span><span><a class="" data-click-track="profile-work-strip-authors" href="https://independent.academia.edu/WilliamKrinsky">William Krinsky</a> and <a class="" data-click-track="profile-work-strip-authors" href="https://99main.academia.edu/MichaelKOliver">Michael K Oliver</a></span></div><div class="wp-workCard_item"><span>The Coleopterists' Bulletin</span><span>, 1988</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">... Pk., W. shore L. Wintergreen, 12.IV. 1987, under rock on moist soil, MKO and Charles M. Olive...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">... Pk., W. shore L. Wintergreen, 12.IV. 1987, under rock on moist soil, MKO and Charles M. Oliver, 1 a (MKOC). ... LITERATURE CITED BELL, RT 1960. A revision of the genus Chlaenius Bonelli (Coleoptera: Carabidae) in North America. Misc. Publ. Entomol. Soc. 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This brings to 335 th...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Forty-six carabid species are reported from Connecticut for the first time. 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This brings to 335 th...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Forty-six carabid species are reported from Connecticut for the first time. This brings to 335 the number of carabid species known from Connecticut.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="76472234"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="76472234"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 76472234; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=76472234]").text(description); $(".js-view-count[data-work-id=76472234]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 76472234; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='76472234']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 76472234, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (false){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "-1" } } $('.js-work-strip[data-work-id=76472234]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":76472234,"title":"Connecticut (Coleoptera: Carabidae)","translated_title":"","metadata":{"abstract":"Forty-six carabid species are reported from Connecticut for the first time. 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This brings to 335 the number of carabid species known from Connecticut.","internal_url":"https://www.academia.edu/76472234/Connecticut_Coleoptera_Carabidae_","translated_internal_url":"","created_at":"2022-04-14T17:59:37.936-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":1462379,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[{"id":42576568,"work_id":76472234,"tagging_user_id":1462379,"tagged_user_id":70741506,"co_author_invite_id":null,"email":"w***y@gmail.com","display_order":0,"name":"William Krinsky","title":"Connecticut (Coleoptera: Carabidae)"}],"downloadable_attachments":[],"slug":"Connecticut_Coleoptera_Carabidae_","translated_slug":"","page_count":null,"language":"en","content_type":"Work","owner":{"id":1462379,"first_name":"Michael","middle_initials":"K","last_name":"Oliver","page_name":"MichaelKOliver","domain_name":"99main","created_at":"2012-04-07T13:04:06.491-07:00","display_name":"Michael K Oliver","url":"https://99main.academia.edu/MichaelKOliver"},"attachments":[],"research_interests":[{"id":261,"name":"Geography","url":"https://www.academia.edu/Documents/in/Geography"}],"urls":[]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") }); </script> <div class="js-work-strip profile--work_container" data-work-id="76472233"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/76472233/Hemitaeniochromis_brachyrhynchus_a_new_species_of_cichlid_fish_from_Lake_Mala%C5%B5i_with_comments_on_some_other_supposed_members_of_the_genus_Teleostei_Cichlidae_"><img alt="Research paper thumbnail of Hemitaeniochromis brachyrhynchus, a new species of cichlid fish from Lake Malaŵi, with comments on some other supposed members of the genus (Teleostei: Cichlidae)" class="work-thumbnail" src="https://attachments.academia-assets.com/84170289/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/76472233/Hemitaeniochromis_brachyrhynchus_a_new_species_of_cichlid_fish_from_Lake_Mala%C5%B5i_with_comments_on_some_other_supposed_members_of_the_genus_Teleostei_Cichlidae_">Hemitaeniochromis brachyrhynchus, a new species of cichlid fish from Lake Malaŵi, with comments on some other supposed members of the genus (Teleostei: Cichlidae)</a></div><div class="wp-workCard_item"><span>Zootaxa</span><span>, 2012</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Hemitaeniochromis brachyrhynchus, an anatomically distinctive and apparently rare new cichlid, is...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Hemitaeniochromis brachyrhynchus, an anatomically distinctive and apparently rare new cichlid, is described and illus-trated from specimens collected at two widely separated localities within Lake Malaŵi. It is easily distinguished from H.urotaenia, type species of the genus, by its narrow lacrimal bone which is only one-third of the orbit length, a characterthought to be unique not only in Hemitaeniochromis but among all known Lake Malaŵi cichlids. The genus Hemitaenio-chromis Eccles & Trewavas (1989) is redefined to allow provisional inclusion of this new species. Two species placed inProtomelas by Eccles & Trewavas (1989) [P. insignis (Trewavas) and P. spilopterus (Trewavas)] were recently transferredto Hemitaeniochromis by some authors, without much evidence. The generic placement of these controversial taxa, and of several undescribed species known only from underwater photographs, is briefly reconsidered.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="1c9ff539c05684f01602398e905d7412" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":84170289,"asset_id":76472233,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/84170289/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="76472233"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="76472233"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 76472233; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=76472233]").text(description); $(".js-view-count[data-work-id=76472233]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 76472233; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='76472233']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 76472233, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "1c9ff539c05684f01602398e905d7412" } } $('.js-work-strip[data-work-id=76472233]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":76472233,"title":"Hemitaeniochromis brachyrhynchus, a new species of cichlid fish from Lake Malaŵi, with comments on some other supposed members of the genus (Teleostei: Cichlidae)","translated_title":"","metadata":{"abstract":"Hemitaeniochromis brachyrhynchus, an anatomically distinctive and apparently rare new cichlid, is described and illus-trated from specimens collected at two widely separated localities within Lake Malaŵi. 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$(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") }); </script> <div class="js-work-strip profile--work_container" data-work-id="76472231"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/76472231/A_Kink_in_the_Line_Does_a_Unique_Lateral_Line_Peculiarity_Really_Characterize_Lake_Mala%C5%B5is_Huge_Haplochromine_Species_Flock_Teleostei_Cichlidae_"><img alt="Research paper thumbnail of A Kink in the Line: Does a Unique Lateral-Line Peculiarity Really Characterize Lake Malaŵi's Huge Haplochromine Species Flock (Teleostei: Cichlidae)?" class="work-thumbnail" src="https://a.academia-assets.com/images/blank-paper.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/76472231/A_Kink_in_the_Line_Does_a_Unique_Lateral_Line_Peculiarity_Really_Characterize_Lake_Mala%C5%B5is_Huge_Haplochromine_Species_Flock_Teleostei_Cichlidae_">A Kink in the Line: Does a Unique Lateral-Line Peculiarity Really Characterize Lake Malaŵi's Huge Haplochromine Species Flock (Teleostei: Cichlidae)?</a></div><div class="wp-workCard_item"><span>Bulletin of the Peabody Museum of Natural History</span><span>, 2016</span></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Abstract A downward kink one or more scales in length at the posterior end of the upper lateral l...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Abstract A downward kink one or more scales in length at the posterior end of the upper lateral line, leaving only one scale row instead of two between it and the lower lateral line, has been considered a synapomorphy uniting as a monophyletic group all members of both clades of endemic haplochromine cichlids of Lake Malaŵi. Here, this claim is tested by surveying the character in representatives of all genera of Lake Malaŵi haplochromines (tribe Haplochromini of the subfamily Pseudocrenilabrinae). Presence of the kink is shown to be variable in most genera, and it was not documented at all in two (Abactochromis, Cyrtocara). Specimens with no kink occur in many genera, and kink presence and length frequently vary on the two sides of the body. Thus, it is difficult to justify the kink as a synapomorphy of the species flock. The two clades differ in kink frequency; a kink occurred on at least one side of the body in 79% of bilaterally assessable specimens in the non-mbuna clade, but in only 53% of those in the mbuna clade (p &lt; 0.0001). When a kink is present, its length is more variable in non-mbuna (1 to 8 scales, mode 2) than mbuna (1 to 4 scales, mode 1). Extension of the upper lateral line backward from the upper flank close to or onto the caudal peduncle, where no scale row is as dorsally situated, is identified as one cause of kink existence; additional factors likely await discovery. Contrary to published statements, the kink is expressed in some specimens of Astatotilapia calliptera, a phylogenetically important nonendemic haplochromine found in Lake Malaŵi and elsewhere, which may be the sister of the lake&#39;s endemic flock. An apparently identical lateral-line kink occurs sporadically in haplochromines in other lakes and in members of three other pseudocrenilabrine cichlid tribes, further refuting the hypothesis that this character is a synapomorphy of the Lake Malaŵi species flock.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="76472231"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="76472231"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 76472231; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=76472231]").text(description); $(".js-view-count[data-work-id=76472231]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 76472231; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='76472231']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 76472231, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (false){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "-1" } } $('.js-work-strip[data-work-id=76472231]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":76472231,"title":"A Kink in the Line: Does a Unique Lateral-Line Peculiarity Really Characterize Lake Malaŵi's Huge Haplochromine Species Flock (Teleostei: Cichlidae)?","translated_title":"","metadata":{"abstract":"Abstract A downward kink one or more scales in length at the posterior end of the upper lateral line, leaving only one scale row instead of two between it and the lower lateral line, has been considered a synapomorphy uniting as a monophyletic group all members of both clades of endemic haplochromine cichlids of Lake Malaŵi. Here, this claim is tested by surveying the character in representatives of all genera of Lake Malaŵi haplochromines (tribe Haplochromini of the subfamily Pseudocrenilabrinae). Presence of the kink is shown to be variable in most genera, and it was not documented at all in two (Abactochromis, Cyrtocara). Specimens with no kink occur in many genera, and kink presence and length frequently vary on the two sides of the body. Thus, it is difficult to justify the kink as a synapomorphy of the species flock. The two clades differ in kink frequency; a kink occurred on at least one side of the body in 79% of bilaterally assessable specimens in the non-mbuna clade, but in only 53% of those in the mbuna clade (p \u0026lt; 0.0001). When a kink is present, its length is more variable in non-mbuna (1 to 8 scales, mode 2) than mbuna (1 to 4 scales, mode 1). Extension of the upper lateral line backward from the upper flank close to or onto the caudal peduncle, where no scale row is as dorsally situated, is identified as one cause of kink existence; additional factors likely await discovery. Contrary to published statements, the kink is expressed in some specimens of Astatotilapia calliptera, a phylogenetically important nonendemic haplochromine found in Lake Malaŵi and elsewhere, which may be the sister of the lake\u0026#39;s endemic flock. An apparently identical lateral-line kink occurs sporadically in haplochromines in other lakes and in members of three other pseudocrenilabrine cichlid tribes, further refuting the hypothesis that this character is a synapomorphy of the Lake Malaŵi species flock.","publisher":"Bulletin of the Peabody Museum of Natural History","publication_date":{"day":null,"month":null,"year":2016,"errors":{}},"publication_name":"Bulletin of the Peabody Museum of Natural History"},"translated_abstract":"Abstract A downward kink one or more scales in length at the posterior end of the upper lateral line, leaving only one scale row instead of two between it and the lower lateral line, has been considered a synapomorphy uniting as a monophyletic group all members of both clades of endemic haplochromine cichlids of Lake Malaŵi. Here, this claim is tested by surveying the character in representatives of all genera of Lake Malaŵi haplochromines (tribe Haplochromini of the subfamily Pseudocrenilabrinae). Presence of the kink is shown to be variable in most genera, and it was not documented at all in two (Abactochromis, Cyrtocara). Specimens with no kink occur in many genera, and kink presence and length frequently vary on the two sides of the body. Thus, it is difficult to justify the kink as a synapomorphy of the species flock. The two clades differ in kink frequency; a kink occurred on at least one side of the body in 79% of bilaterally assessable specimens in the non-mbuna clade, but in only 53% of those in the mbuna clade (p \u0026lt; 0.0001). When a kink is present, its length is more variable in non-mbuna (1 to 8 scales, mode 2) than mbuna (1 to 4 scales, mode 1). Extension of the upper lateral line backward from the upper flank close to or onto the caudal peduncle, where no scale row is as dorsally situated, is identified as one cause of kink existence; additional factors likely await discovery. Contrary to published statements, the kink is expressed in some specimens of Astatotilapia calliptera, a phylogenetically important nonendemic haplochromine found in Lake Malaŵi and elsewhere, which may be the sister of the lake\u0026#39;s endemic flock. An apparently identical lateral-line kink occurs sporadically in haplochromines in other lakes and in members of three other pseudocrenilabrine cichlid tribes, further refuting the hypothesis that this character is a synapomorphy of the Lake Malaŵi species flock.","internal_url":"https://www.academia.edu/76472231/A_Kink_in_the_Line_Does_a_Unique_Lateral_Line_Peculiarity_Really_Characterize_Lake_Mala%C5%B5is_Huge_Haplochromine_Species_Flock_Teleostei_Cichlidae_","translated_internal_url":"","created_at":"2022-04-14T17:59:37.409-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":1462379,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[],"slug":"A_Kink_in_the_Line_Does_a_Unique_Lateral_Line_Peculiarity_Really_Characterize_Lake_Malaŵis_Huge_Haplochromine_Species_Flock_Teleostei_Cichlidae_","translated_slug":"","page_count":null,"language":"en","content_type":"Work","owner":{"id":1462379,"first_name":"Michael","middle_initials":"K","last_name":"Oliver","page_name":"MichaelKOliver","domain_name":"99main","created_at":"2012-04-07T13:04:06.491-07:00","display_name":"Michael K Oliver","url":"https://99main.academia.edu/MichaelKOliver"},"attachments":[],"research_interests":[{"id":135987,"name":"Hypothesis testing","url":"https://www.academia.edu/Documents/in/Hypothesis_testing"},{"id":216604,"name":"Cichlidae","url":"https://www.academia.edu/Documents/in/Cichlidae"},{"id":452259,"name":"Lake Malawi","url":"https://www.academia.edu/Documents/in/Lake_Malawi"},{"id":1418924,"name":"Lateral Line","url":"https://www.academia.edu/Documents/in/Lateral_Line"}],"urls":[]}, dispatcherData: dispatcherData }); 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Diverse systematic proble...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">&quot;The cichlids of Africa present many challenges to the taxonomist. Diverse systematic problems at three levels in cichlid phylogeny form the subject matter of this dissertation. At the most general level, I summarize current morphological evidence that supports cichlid monophyly and the most-inclusive groups of cichlids. With this review as a framework, I argue that the most phylogenetically primitive African cichlid is Heterochromis multidens, a little-known large cichlid from the Zaire River basin. I redescribe this monotypic genus and species and illustrate much of its osteology. It possesses a distinctive suite of derived characters, but is uniquely primitive among African cichlids in infraorbital morphology and other features. At a less inclusive phylogenetic level, I examine the controversy over the monophyletic status of the &quot;species flock&quot; of haplochromine cichlids in Lake Malawi. A derived type of anal-fin markings present in a subset of the Malawian fauna is shared with many non-Malawian haplochromines, indicating that the species flock of Lake Malawi is not monophyletic. The two (or more) faunal components of Lake Malawi are ecologically distinct. The remaining major parts of the study are concerned with particular assemblages of haplochromine cichlids in Lake Malawi. I present a detailed phylogenetic analysis of the Cyrtocara livingstonii species-group of ambush predators, and characterize a new species in the group. I also describe three new species of haplochromines with three lateral spots and present a key to the three-spot assemblage. Intralacustrine dispersion of Malawi cichlids is briefly addressed in appended material.&quot;</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="76472228"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="76472228"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 76472228; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=76472228]").text(description); $(".js-view-count[data-work-id=76472228]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 76472228; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='76472228']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 76472228, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (false){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "-1" } } $('.js-work-strip[data-work-id=76472228]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":76472228,"title":"Systematics of African Cichlid Fishes: Determination of the Most Primitive Taxon, and Studies on the Haplochromines of Lake Malawi (Teleostei: Cichlidae)","translated_title":"","metadata":{"abstract":"\u0026quot;The cichlids of Africa present many challenges to the taxonomist. 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Pk., W. shore L. Wintergreen, 12.IV. 1987, under rock on moist soil, MKO and Charles M. Olive...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">... Pk., W. shore L. Wintergreen, 12.IV. 1987, under rock on moist soil, MKO and Charles M. Oliver, 1 a (MKOC). ... LITERATURE CITED BELL, RT 1960. A revision of the genus Chlaenius Bonelli (Coleoptera: Carabidae) in North America. Misc. Publ. Entomol. Soc. 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He based one of them, L. joanjohnsonae, on a holotype and paratype belonging to two species that differ strikingly from each other in coloration and oral dentition. Although he gave conflicting statements both of standard length and total length of the paratype in different paragraphs, Johnson (1974a: 15) clearly designated the smaller of the two specimens as the holotype, and since they differ in standard length by nearly 10 mrn there is no doubt as to which this is. The number of dorsal fin spines (sixteen in the holotype, seventeen in the paratype) corroborates his designation. It might be noted, however, that Johnson's counts of segmented dorsal rays are inaccurate for both specimens. I have examined the types of all species described by Johnson (1974a and 1974b) and found that the holotype of L. joanjohnsonue Johnson is a specimen of L. fryeri Oliver in Davies (1973) ; L. ioanjohnsonae is therefore a junior synonym of that species. The manuscript name Labidochromis fryeri and a diagnosis of this taxon were unfortunately published in an aquarium magazine in a letter from T. E. (Peter) Davies, who specifically (and correctly) attributed to me the name, the diagnosis, and my intention eventually to publish them. Regrettably, by Articles 11 and 13 (a) (i) of the International Labidochromis textilis, new species Brocade Cichlid","publication_date":{"day":null,"month":null,"year":1975,"errors":{}},"grobid_abstract_attachment_id":79834848},"translated_abstract":null,"internal_url":"https://www.academia.edu/69923161/Labidochromis_textilis_a_new_cichlid_fish_Teleostei_Cichlidae_from_Lake_Malawi","translated_internal_url":"","created_at":"2022-01-29T07:22:34.262-08:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":1462379,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[{"id":79834848,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/79834848/thumbnails/1.jpg","file_name":"oliver_1975.pdf","download_url":"https://www.academia.edu/attachments/79834848/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"Labidochromis_textilis_a_new_cichlid_fis.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/79834848/oliver_1975-libre.pdf?1643473844=\u0026response-content-disposition=attachment%3B+filename%3DLabidochromis_textilis_a_new_cichlid_fis.pdf\u0026Expires=1732422414\u0026Signature=HIor6lHUL4caezDsGyqzC27-7cX1E4hRTX69YR1AZM~ouf0pGRCWLcG-GozGURgcyQ7xADlDVebhkmTnoQdJNQE9beIDvEbxlD6VWbLXrAJt62U26DZAzFiGgvRVYDYghlvIXcNVvxR054HmxI-1sT-MToOOipvAp5jkUdb2wUznX9~kHuuLwUb3KJSN11Lyn6grjN6Wx62i0aHaU-OAlJxucKCaSupJFDv7ZSVrkOmMn8gASOcIPNcUKuQN5MVyA6fqr8Ls~Ly5IJiSfQnZZeY8RCZ6JXukWsBfAM8B68ycDIANJwoLsv90qMZQbjPODVtRMHISTdFELSYqR4B4mA__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"Labidochromis_textilis_a_new_cichlid_fish_Teleostei_Cichlidae_from_Lake_Malawi","translated_slug":"","page_count":12,"language":"en","content_type":"Work","owner":{"id":1462379,"first_name":"Michael","middle_initials":"K","last_name":"Oliver","page_name":"MichaelKOliver","domain_name":"99main","created_at":"2012-04-07T13:04:06.491-07:00","display_name":"Michael K Oliver","url":"https://99main.academia.edu/MichaelKOliver"},"attachments":[{"id":79834848,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/79834848/thumbnails/1.jpg","file_name":"oliver_1975.pdf","download_url":"https://www.academia.edu/attachments/79834848/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"Labidochromis_textilis_a_new_cichlid_fis.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/79834848/oliver_1975-libre.pdf?1643473844=\u0026response-content-disposition=attachment%3B+filename%3DLabidochromis_textilis_a_new_cichlid_fis.pdf\u0026Expires=1732422414\u0026Signature=HIor6lHUL4caezDsGyqzC27-7cX1E4hRTX69YR1AZM~ouf0pGRCWLcG-GozGURgcyQ7xADlDVebhkmTnoQdJNQE9beIDvEbxlD6VWbLXrAJt62U26DZAzFiGgvRVYDYghlvIXcNVvxR054HmxI-1sT-MToOOipvAp5jkUdb2wUznX9~kHuuLwUb3KJSN11Lyn6grjN6Wx62i0aHaU-OAlJxucKCaSupJFDv7ZSVrkOmMn8gASOcIPNcUKuQN5MVyA6fqr8Ls~Ly5IJiSfQnZZeY8RCZ6JXukWsBfAM8B68ycDIANJwoLsv90qMZQbjPODVtRMHISTdFELSYqR4B4mA__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[{"id":169,"name":"Systematics (Taxonomy)","url":"https://www.academia.edu/Documents/in/Systematics_Taxonomy_"},{"id":7710,"name":"Biology","url":"https://www.academia.edu/Documents/in/Biology"},{"id":216604,"name":"Cichlidae","url":"https://www.academia.edu/Documents/in/Cichlidae"}],"urls":[]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") }); </script> <div class="js-work-strip profile--work_container" data-work-id="69922906"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/69922906/A_new_genus_for_Melanochromis_labrosus_a_problematic_Lake_Malawi_cichlid_with_hypertrophied_lips_Teleostei_Cichlidae_"><img alt="Research paper thumbnail of A new genus for Melanochromis labrosus, a problematic Lake Malawi cichlid with hypertrophied lips (Teleostei: Cichlidae)" class="work-thumbnail" src="https://attachments.academia-assets.com/79834585/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/69922906/A_new_genus_for_Melanochromis_labrosus_a_problematic_Lake_Malawi_cichlid_with_hypertrophied_lips_Teleostei_Cichlidae_">A new genus for Melanochromis labrosus, a problematic Lake Malawi cichlid with hypertrophied lips (Teleostei: Cichlidae)</a></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Melanochromis labrosus is the only one of the 800 or so endemic cichlid species in Lake Malawi wh...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Melanochromis labrosus is the only one of the 800 or so endemic cichlid species in Lake Malawi whose placement with mbuna or non-mbuna has been disputed on morphological grounds. Here, we redescribe M. labrosus using all obtainable specimens collected since its original description. In addition, we extensively survey vertebral counts in the mbuna. To determine if M. labrosus belongs with the mbuna, we review the morphological traits often considered to define the mbuna as a natural group, concluding that none is a uniquely shared specialization. Two features of body coloration (a relatively small number of ocellate spots in the anal fin and a possible polychromatism in one population) suggest that M. labrosus is best considered a species of mbuna. Although its placement within the mbuna group remains somewhat equivocal, we unambiguously demonstrate that M. labrosus is not a Melanochromis, since it lacks the synapomorphic striped color pattern typical of that genus. Melanochromis lab...</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="f670b6a1f7043405db9e0fc9aed1b3d0" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":79834585,"asset_id":69922906,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/79834585/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="69922906"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="69922906"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 69922906; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=69922906]").text(description); $(".js-view-count[data-work-id=69922906]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 69922906; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='69922906']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 69922906, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "f670b6a1f7043405db9e0fc9aed1b3d0" } } $('.js-work-strip[data-work-id=69922906]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":69922906,"title":"A new genus for Melanochromis labrosus, a problematic Lake Malawi cichlid with hypertrophied lips (Teleostei: Cichlidae)","translated_title":"","metadata":{"abstract":"Melanochromis labrosus is the only one of the 800 or so endemic cichlid species in Lake Malawi whose placement with mbuna or non-mbuna has been disputed on morphological grounds. Here, we redescribe M. labrosus using all obtainable specimens collected since its original description. In addition, we extensively survey vertebral counts in the mbuna. To determine if M. labrosus belongs with the mbuna, we review the morphological traits often considered to define the mbuna as a natural group, concluding that none is a uniquely shared specialization. Two features of body coloration (a relatively small number of ocellate spots in the anal fin and a possible polychromatism in one population) suggest that M. labrosus is best considered a species of mbuna. Although its placement within the mbuna group remains somewhat equivocal, we unambiguously demonstrate that M. labrosus is not a Melanochromis, since it lacks the synapomorphic striped color pattern typical of that genus. Melanochromis lab...","publication_date":{"day":null,"month":null,"year":2010,"errors":{}}},"translated_abstract":"Melanochromis labrosus is the only one of the 800 or so endemic cichlid species in Lake Malawi whose placement with mbuna or non-mbuna has been disputed on morphological grounds. Here, we redescribe M. labrosus using all obtainable specimens collected since its original description. In addition, we extensively survey vertebral counts in the mbuna. To determine if M. labrosus belongs with the mbuna, we review the morphological traits often considered to define the mbuna as a natural group, concluding that none is a uniquely shared specialization. Two features of body coloration (a relatively small number of ocellate spots in the anal fin and a possible polychromatism in one population) suggest that M. labrosus is best considered a species of mbuna. Although its placement within the mbuna group remains somewhat equivocal, we unambiguously demonstrate that M. labrosus is not a Melanochromis, since it lacks the synapomorphic striped color pattern typical of that genus. Melanochromis lab...","internal_url":"https://www.academia.edu/69922906/A_new_genus_for_Melanochromis_labrosus_a_problematic_Lake_Malawi_cichlid_with_hypertrophied_lips_Teleostei_Cichlidae_","translated_internal_url":"","created_at":"2022-01-29T07:20:10.360-08:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":1462379,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[{"id":37626419,"work_id":69922906,"tagging_user_id":1462379,"tagged_user_id":null,"co_author_invite_id":7440491,"email":"m***r@fhcrc.org","display_order":0,"name":"Matthew Arnegard","title":"A new genus for Melanochromis labrosus, a problematic Lake Malawi cichlid with hypertrophied lips (Teleostei: Cichlidae)"}],"downloadable_attachments":[{"id":79834585,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/79834585/thumbnails/1.jpg","file_name":"oliver_arnegard_2010.pdf","download_url":"https://www.academia.edu/attachments/79834585/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"A_new_genus_for_Melanochromis_labrosus_a.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/79834585/oliver_arnegard_2010-libre.pdf?1643469709=\u0026response-content-disposition=attachment%3B+filename%3DA_new_genus_for_Melanochromis_labrosus_a.pdf\u0026Expires=1732422414\u0026Signature=OVD1vxpl5~MKcfapqWws6pDmNQIfBi89a9olT62OPIjb24r2qCsxpkVq-cbBAl9CoYG4Mljbdbg3taNZPURjv2yGCrSlYeZGTSOBgzDj9hV9-0lzQRTVYPYe2m8VjWsMXODCh7KTbKeHWX6N7yOP2WCdJIaFvS7UhUCTyr1pqmNuLEpza6ijbS3RwlgC-6Plxgw2o9NA9zM-pGkX6i0KAGTU~BQpKg0C-CAkRA0w1VF2bMT4I25mgg1BzpXKG1OW8rOTlOT4yU1KELA1qXrMx7Ju-IK9FJLV1CBB9gXQUsdbx9QqE8Sg5lH-Oik0fF616vc0e68KTJSw-UrrTlI4vA__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"A_new_genus_for_Melanochromis_labrosus_a_problematic_Lake_Malawi_cichlid_with_hypertrophied_lips_Teleostei_Cichlidae_","translated_slug":"","page_count":24,"language":"en","content_type":"Work","owner":{"id":1462379,"first_name":"Michael","middle_initials":"K","last_name":"Oliver","page_name":"MichaelKOliver","domain_name":"99main","created_at":"2012-04-07T13:04:06.491-07:00","display_name":"Michael K Oliver","url":"https://99main.academia.edu/MichaelKOliver"},"attachments":[{"id":79834585,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/79834585/thumbnails/1.jpg","file_name":"oliver_arnegard_2010.pdf","download_url":"https://www.academia.edu/attachments/79834585/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"A_new_genus_for_Melanochromis_labrosus_a.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/79834585/oliver_arnegard_2010-libre.pdf?1643469709=\u0026response-content-disposition=attachment%3B+filename%3DA_new_genus_for_Melanochromis_labrosus_a.pdf\u0026Expires=1732422414\u0026Signature=OVD1vxpl5~MKcfapqWws6pDmNQIfBi89a9olT62OPIjb24r2qCsxpkVq-cbBAl9CoYG4Mljbdbg3taNZPURjv2yGCrSlYeZGTSOBgzDj9hV9-0lzQRTVYPYe2m8VjWsMXODCh7KTbKeHWX6N7yOP2WCdJIaFvS7UhUCTyr1pqmNuLEpza6ijbS3RwlgC-6Plxgw2o9NA9zM-pGkX6i0KAGTU~BQpKg0C-CAkRA0w1VF2bMT4I25mgg1BzpXKG1OW8rOTlOT4yU1KELA1qXrMx7Ju-IK9FJLV1CBB9gXQUsdbx9QqE8Sg5lH-Oik0fF616vc0e68KTJSw-UrrTlI4vA__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"},{"id":79834587,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/79834587/thumbnails/1.jpg","file_name":"oliver_arnegard_2010.pdf","download_url":"https://www.academia.edu/attachments/79834587/download_file","bulk_download_file_name":"A_new_genus_for_Melanochromis_labrosus_a.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/79834587/oliver_arnegard_2010-libre.pdf?1643469708=\u0026response-content-disposition=attachment%3B+filename%3DA_new_genus_for_Melanochromis_labrosus_a.pdf\u0026Expires=1732422414\u0026Signature=EPOhYKz8hp4fH2pd8vlH56zOh4tBxgSpqXYPj-BrQigF3wti2miBYJv~KAIQpBLK5Ig3KjSNuMFwUTFl3Z03wV0WeOzWWGBSE38vKLo4YpLaKmf7ZBt8UYRbpuA7m9mNYQg2Adr6hZ5c4~qMkUNQfLJe9UaYCwdHl~lKj5gatBFvW6xjTpQvHNx6rHEA51uBzgBAjiAJySrWTeTIXZUGIWJ2A-~q02Eb5oDNRmPZCYPr~62by9034Mzk9s4sE4QIoEFXjfVMKMNQAVwqqTpOfRxcl9anukIdpcxmhrIzWgLIRZFeuYlahyhIb9ZuvQMZC8J4recRIfymYtmdCmURvg__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[{"id":7076,"name":"Taxonomy","url":"https://www.academia.edu/Documents/in/Taxonomy"},{"id":7710,"name":"Biology","url":"https://www.academia.edu/Documents/in/Biology"},{"id":10225,"name":"Agriculture","url":"https://www.academia.edu/Documents/in/Agriculture"},{"id":216604,"name":"Cichlidae","url":"https://www.academia.edu/Documents/in/Cichlidae"}],"urls":[{"id":17075021,"url":"http://malawicichlids.com/oliver_arnegard_2010.pdf"}]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") }); </script> <div class="js-work-strip profile--work_container" data-work-id="37695426"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/37695426/Six_New_Species_of_the_Cichlid_Genus_Otopharynx_from_Lake_Mala%C5%B5i_Teleostei_Cichlidae"><img alt="Research paper thumbnail of Six New Species of the Cichlid Genus Otopharynx from Lake Malaŵi (Teleostei: Cichlidae" class="work-thumbnail" src="https://attachments.academia-assets.com/57686794/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/37695426/Six_New_Species_of_the_Cichlid_Genus_Otopharynx_from_Lake_Mala%C5%B5i_Teleostei_Cichlidae">Six New Species of the Cichlid Genus Otopharynx from Lake Malaŵi (Teleostei: Cichlidae</a></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">The endemic Lake Malaŵi cichlid genus Otopharynx is increased from 14 species to 20 by the descri...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">The endemic Lake Malaŵi cichlid genus Otopharynx is increased from 14 species to 20 by the description of six new species collected from inshore sandy or intermediate rocky/sandy areas or trawled in depths to 59 m. Otopharynx alpha is distinguished from congeners by its unique melanin pattern with suprapectoral spot discrete but supraanal and precaudal spots connected to form a stripe, and by its distinctively shaped pharyngeal bone with numerous small, crowded teeth. Otopharynx mumboensis, thought to be endemic to Mumbo Island, is separated from most congeners by a suprapectoral spot placed entirely below the upper lateral line, and from all others by the combination of 13 gill rakers, pharyngeal bone with posterior margin nearly straight, crowns of all pharyngeal teeth in the median columns broadened and flattened, head length 30.9–32.2% SL, body depth 35.5–36.1% SL, and lower jaw 35.7–36.2% HL. Otopharynx styrax has a more elongate body (depth 24.3–28.6% SL) than any congener. Otopharynx aletes can be diagnosed by the combination of a suprapectoral spot overlapping the upper lateral line, seven or eight vertical bars below the dorsal-fin base, 10 or 11 gill rakers, 34–36 scales in the lateral line, lower pharyngeal bone with molariform posteromedian teeth, and caudal peduncle length 1.6–2.0 times its depth. Otopharynx panniculus is diagnosed by the combination of a prominent square suprapectoral spot spanning subdorsal bars 3 to 4, no dorsal midline spots, 13–15 gill rakers, 31–33 scales in the lateral line, 15 or 16 dorsal-fin spines, absence of a lacrimal notch, and slightly<br />enlarged teeth in the median columns of the lower pharyngeal bone. Otopharynx peridodeka most closely resembles O. panniculus and shares the loss of the lacrimal notch, but has 34–35 scales in the lateral line and 11–13 gill rakers. Photographs of living or freshly collected specimens of five of the new species are provided.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="dd20a3e5fea564f5b0028ad88c64cb9c" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":57686794,"asset_id":37695426,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/57686794/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="37695426"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="37695426"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 37695426; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=37695426]").text(description); $(".js-view-count[data-work-id=37695426]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 37695426; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='37695426']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 37695426, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "dd20a3e5fea564f5b0028ad88c64cb9c" } } $('.js-work-strip[data-work-id=37695426]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":37695426,"title":"Six New Species of the Cichlid Genus Otopharynx from Lake Malaŵi (Teleostei: Cichlidae","translated_title":"","metadata":{"abstract":"The endemic Lake Malaŵi cichlid genus Otopharynx is increased from 14 species to 20 by the description of six new species collected from inshore sandy or intermediate rocky/sandy areas or trawled in depths to 59 m. Otopharynx alpha is distinguished from congeners by its unique melanin pattern with suprapectoral spot discrete but supraanal and precaudal spots connected to form a stripe, and by its distinctively shaped pharyngeal bone with numerous small, crowded teeth. Otopharynx mumboensis, thought to be endemic to Mumbo Island, is separated from most congeners by a suprapectoral spot placed entirely below the upper lateral line, and from all others by the combination of 13 gill rakers, pharyngeal bone with posterior margin nearly straight, crowns of all pharyngeal teeth in the median columns broadened and flattened, head length 30.9–32.2% SL, body depth 35.5–36.1% SL, and lower jaw 35.7–36.2% HL. Otopharynx styrax has a more elongate body (depth 24.3–28.6% SL) than any congener. Otopharynx aletes can be diagnosed by the combination of a suprapectoral spot overlapping the upper lateral line, seven or eight vertical bars below the dorsal-fin base, 10 or 11 gill rakers, 34–36 scales in the lateral line, lower pharyngeal bone with molariform posteromedian teeth, and caudal peduncle length 1.6–2.0 times its depth. Otopharynx panniculus is diagnosed by the combination of a prominent square suprapectoral spot spanning subdorsal bars 3 to 4, no dorsal midline spots, 13–15 gill rakers, 31–33 scales in the lateral line, 15 or 16 dorsal-fin spines, absence of a lacrimal notch, and slightly\nenlarged teeth in the median columns of the lower pharyngeal bone. Otopharynx peridodeka most closely resembles O. panniculus and shares the loss of the lacrimal notch, but has 34–35 scales in the lateral line and 11–13 gill rakers. Photographs of living or freshly collected specimens of five of the new species are provided."},"translated_abstract":"The endemic Lake Malaŵi cichlid genus Otopharynx is increased from 14 species to 20 by the description of six new species collected from inshore sandy or intermediate rocky/sandy areas or trawled in depths to 59 m. Otopharynx alpha is distinguished from congeners by its unique melanin pattern with suprapectoral spot discrete but supraanal and precaudal spots connected to form a stripe, and by its distinctively shaped pharyngeal bone with numerous small, crowded teeth. Otopharynx mumboensis, thought to be endemic to Mumbo Island, is separated from most congeners by a suprapectoral spot placed entirely below the upper lateral line, and from all others by the combination of 13 gill rakers, pharyngeal bone with posterior margin nearly straight, crowns of all pharyngeal teeth in the median columns broadened and flattened, head length 30.9–32.2% SL, body depth 35.5–36.1% SL, and lower jaw 35.7–36.2% HL. Otopharynx styrax has a more elongate body (depth 24.3–28.6% SL) than any congener. Otopharynx aletes can be diagnosed by the combination of a suprapectoral spot overlapping the upper lateral line, seven or eight vertical bars below the dorsal-fin base, 10 or 11 gill rakers, 34–36 scales in the lateral line, lower pharyngeal bone with molariform posteromedian teeth, and caudal peduncle length 1.6–2.0 times its depth. Otopharynx panniculus is diagnosed by the combination of a prominent square suprapectoral spot spanning subdorsal bars 3 to 4, no dorsal midline spots, 13–15 gill rakers, 31–33 scales in the lateral line, 15 or 16 dorsal-fin spines, absence of a lacrimal notch, and slightly\nenlarged teeth in the median columns of the lower pharyngeal bone. Otopharynx peridodeka most closely resembles O. panniculus and shares the loss of the lacrimal notch, but has 34–35 scales in the lateral line and 11–13 gill rakers. Photographs of living or freshly collected specimens of five of the new species are provided.","internal_url":"https://www.academia.edu/37695426/Six_New_Species_of_the_Cichlid_Genus_Otopharynx_from_Lake_Mala%C5%B5i_Teleostei_Cichlidae","translated_internal_url":"","created_at":"2018-11-03T07:44:53.145-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":1462379,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[{"id":57686794,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/57686794/thumbnails/1.jpg","file_name":"Oliver_2018_Six_New_Species_of_the_Cichlid_Genus_Otopharynx_from_Lake_Malawi_Teleostei-_Cichlidae.pdf","download_url":"https://www.academia.edu/attachments/57686794/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"Six_New_Species_of_the_Cichlid_Genus_Oto.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/57686794/Oliver_2018_Six_New_Species_of_the_Cichlid_Genus_Otopharynx_from_Lake_Malawi_Teleostei-_Cichlidae-libre.pdf?1541256795=\u0026response-content-disposition=attachment%3B+filename%3DSix_New_Species_of_the_Cichlid_Genus_Oto.pdf\u0026Expires=1732422414\u0026Signature=aYy7EuFzebSn3VLr3B3tbQUXRc3H2RVaeEUeHflnvrdznvcCRHIzNp9d87YrVbm0If95hKz3I~T2OGfanClhQBBKEk9tC~Ks-dDPfoEMtu2ePqPCjboC0n88sXom8JZJ3fkGXH8k134ec48eT7kL3ala7qYrxE3EQIbQbxmSwscX3ff9tm5m72N1XJ5WD3rcFVRi1BBm2WEJzM4By~zV1U8tgH-CEz~-uJids7ZTQKGIJDgqWmZrxKpNqcxeekAW7aDDqKNqO3259ABvjuZhqM5IwhMC21j~RocYJEEzurfy2sc9n6Kx2GPkHzmuW3DnooSS~WVvQejF3jYelp5gTQ__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"Six_New_Species_of_the_Cichlid_Genus_Otopharynx_from_Lake_Malaŵi_Teleostei_Cichlidae","translated_slug":"","page_count":39,"language":"en","content_type":"Work","owner":{"id":1462379,"first_name":"Michael","middle_initials":"K","last_name":"Oliver","page_name":"MichaelKOliver","domain_name":"99main","created_at":"2012-04-07T13:04:06.491-07:00","display_name":"Michael K Oliver","url":"https://99main.academia.edu/MichaelKOliver"},"attachments":[{"id":57686794,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/57686794/thumbnails/1.jpg","file_name":"Oliver_2018_Six_New_Species_of_the_Cichlid_Genus_Otopharynx_from_Lake_Malawi_Teleostei-_Cichlidae.pdf","download_url":"https://www.academia.edu/attachments/57686794/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"Six_New_Species_of_the_Cichlid_Genus_Oto.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/57686794/Oliver_2018_Six_New_Species_of_the_Cichlid_Genus_Otopharynx_from_Lake_Malawi_Teleostei-_Cichlidae-libre.pdf?1541256795=\u0026response-content-disposition=attachment%3B+filename%3DSix_New_Species_of_the_Cichlid_Genus_Oto.pdf\u0026Expires=1732422414\u0026Signature=aYy7EuFzebSn3VLr3B3tbQUXRc3H2RVaeEUeHflnvrdznvcCRHIzNp9d87YrVbm0If95hKz3I~T2OGfanClhQBBKEk9tC~Ks-dDPfoEMtu2ePqPCjboC0n88sXom8JZJ3fkGXH8k134ec48eT7kL3ala7qYrxE3EQIbQbxmSwscX3ff9tm5m72N1XJ5WD3rcFVRi1BBm2WEJzM4By~zV1U8tgH-CEz~-uJids7ZTQKGIJDgqWmZrxKpNqcxeekAW7aDDqKNqO3259ABvjuZhqM5IwhMC21j~RocYJEEzurfy2sc9n6Kx2GPkHzmuW3DnooSS~WVvQejF3jYelp5gTQ__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[{"id":151753,"name":"New Species","url":"https://www.academia.edu/Documents/in/New_Species"},{"id":211405,"name":"Endemic species","url":"https://www.academia.edu/Documents/in/Endemic_species"},{"id":243830,"name":"Bottom Trawling","url":"https://www.academia.edu/Documents/in/Bottom_Trawling"},{"id":452259,"name":"Lake Malawi","url":"https://www.academia.edu/Documents/in/Lake_Malawi"}],"urls":[]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") }); </script> <div class="js-work-strip profile--work_container" data-work-id="24075167"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/24075167/A_Kink_in_the_Line_Does_a_Unique_Lateral_Line_Peculiarity_Really_Characterize_Lake_Malawis_Huge_Haplochromine_Species_Flock_Teleostei_Cichlidae_"><img alt="Research paper thumbnail of A Kink in the Line: Does a Unique Lateral-Line Peculiarity Really Characterize Lake Malawi's Huge Haplochromine Species Flock (Teleostei: Cichlidae)?" class="work-thumbnail" src="https://attachments.academia-assets.com/44445264/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/24075167/A_Kink_in_the_Line_Does_a_Unique_Lateral_Line_Peculiarity_Really_Characterize_Lake_Malawis_Huge_Haplochromine_Species_Flock_Teleostei_Cichlidae_">A Kink in the Line: Does a Unique Lateral-Line Peculiarity Really Characterize Lake Malawi's Huge Haplochromine Species Flock (Teleostei: Cichlidae)?</a></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">A downward kink one or more scales in length at the posterior end of the upper lateral line, leav...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">A downward kink one or more scales in length at the posterior end of the upper lateral line, leaving only one scale row instead of two between it and the lower lateral line, has been considered a synapomorphy uniting as a monophyletic group all members of both clades of endemic haplochromine cichlids of Lake Malawi. Here, this claim is tested by surveying the character in representatives of all genera of Lake Malawi haplochromines (tribe Haplochromini of the subfamily Pseudocrenilabrinae). Presence of the kink is shown to be variable in most genera, and it was not documented at all in two (Abactochromis, Cyrtocara). Specimens with no kink occur in many genera, and kink presence and length frequently vary on the two sides of the body. Thus, it is difficult to justify the kink as a synapomorphy of the species flock. The two clades differ in kink frequency; a kink occurred on at least one side of the body in 79% of bilaterally assessable specimens in the non-mbuna clade, but in only 53% of those in the mbuna clade (p < 0.0001). When a kink is present, its length is more variable in non-mbuna (1 to 8 scales, mode 2) than mbuna (1 to 4 scales, mode 1). Extension of the upper lateral line backward from the upper flank close to or onto the caudal peduncle, where no scale row is as dorsally situated, is identified as one cause of kink existence; additional factors likely await discovery. Contrary to published statements, the kink is expressed in some specimens of Astatotilapia calliptera, a phylogenetically important nonendemic haplochromine found in Lake Malawi and elsewhere, which may be the sister of the lake's endemic flock. An apparently identical lateral-line kink occurs sporadically in haplochromines in other lakes and in members of three other pseudocrenilabrine cichlid tribes, further refuting the hypothesis that this character is a synapomorphy of the Lake Malawi species flock.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="0fddbcd241fc04aac1884328c968f993" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":44445264,"asset_id":24075167,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/44445264/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="24075167"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="24075167"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 24075167; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=24075167]").text(description); $(".js-view-count[data-work-id=24075167]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 24075167; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='24075167']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 24075167, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "0fddbcd241fc04aac1884328c968f993" } } $('.js-work-strip[data-work-id=24075167]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":24075167,"title":"A Kink in the Line: Does a Unique Lateral-Line Peculiarity Really Characterize Lake Malawi's Huge Haplochromine Species Flock (Teleostei: Cichlidae)?","translated_title":"","metadata":{"abstract":"A downward kink one or more scales in length at the posterior end of the upper lateral line, leaving only one scale row instead of two between it and the lower lateral line, has been considered a synapomorphy uniting as a monophyletic group all members of both clades of endemic haplochromine cichlids of Lake Malawi. Here, this claim is tested by surveying the character in representatives of all genera of Lake Malawi haplochromines (tribe Haplochromini of the subfamily Pseudocrenilabrinae). Presence of the kink is shown to be variable in most genera, and it was not documented at all in two (Abactochromis, Cyrtocara). Specimens with no kink occur in many genera, and kink presence and length frequently vary on the two sides of the body. Thus, it is difficult to justify the kink as a synapomorphy of the species flock. The two clades differ in kink frequency; a kink occurred on at least one side of the body in 79% of bilaterally assessable specimens in the non-mbuna clade, but in only 53% of those in the mbuna clade (p \u003c 0.0001). When a kink is present, its length is more variable in non-mbuna (1 to 8 scales, mode 2) than mbuna (1 to 4 scales, mode 1). Extension of the upper lateral line backward from the upper flank close to or onto the caudal peduncle, where no scale row is as dorsally situated, is identified as one cause of kink existence; additional factors likely await discovery. Contrary to published statements, the kink is expressed in some specimens of Astatotilapia calliptera, a phylogenetically important nonendemic haplochromine found in Lake Malawi and elsewhere, which may be the sister of the lake's endemic flock. An apparently identical lateral-line kink occurs sporadically in haplochromines in other lakes and in members of three other pseudocrenilabrine cichlid tribes, further refuting the hypothesis that this character is a synapomorphy of the Lake Malawi species flock."},"translated_abstract":"A downward kink one or more scales in length at the posterior end of the upper lateral line, leaving only one scale row instead of two between it and the lower lateral line, has been considered a synapomorphy uniting as a monophyletic group all members of both clades of endemic haplochromine cichlids of Lake Malawi. Here, this claim is tested by surveying the character in representatives of all genera of Lake Malawi haplochromines (tribe Haplochromini of the subfamily Pseudocrenilabrinae). Presence of the kink is shown to be variable in most genera, and it was not documented at all in two (Abactochromis, Cyrtocara). Specimens with no kink occur in many genera, and kink presence and length frequently vary on the two sides of the body. Thus, it is difficult to justify the kink as a synapomorphy of the species flock. The two clades differ in kink frequency; a kink occurred on at least one side of the body in 79% of bilaterally assessable specimens in the non-mbuna clade, but in only 53% of those in the mbuna clade (p \u003c 0.0001). When a kink is present, its length is more variable in non-mbuna (1 to 8 scales, mode 2) than mbuna (1 to 4 scales, mode 1). Extension of the upper lateral line backward from the upper flank close to or onto the caudal peduncle, where no scale row is as dorsally situated, is identified as one cause of kink existence; additional factors likely await discovery. Contrary to published statements, the kink is expressed in some specimens of Astatotilapia calliptera, a phylogenetically important nonendemic haplochromine found in Lake Malawi and elsewhere, which may be the sister of the lake's endemic flock. An apparently identical lateral-line kink occurs sporadically in haplochromines in other lakes and in members of three other pseudocrenilabrine cichlid tribes, further refuting the hypothesis that this character is a synapomorphy of the Lake Malawi species flock.","internal_url":"https://www.academia.edu/24075167/A_Kink_in_the_Line_Does_a_Unique_Lateral_Line_Peculiarity_Really_Characterize_Lake_Malawis_Huge_Haplochromine_Species_Flock_Teleostei_Cichlidae_","translated_internal_url":"","created_at":"2016-04-05T12:30:18.606-07:00","preview_url":null,"current_user_can_edit":null,"current_user_is_owner":null,"owner_id":1462379,"coauthors_can_edit":true,"document_type":"paper","co_author_tags":[],"downloadable_attachments":[{"id":44445264,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/44445264/thumbnails/1.jpg","file_name":"B57-1-01-Oliver_pp3-20.pdf","download_url":"https://www.academia.edu/attachments/44445264/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"A_Kink_in_the_Line_Does_a_Unique_Lateral.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/44445264/B57-1-01-Oliver_pp3-20-libre.pdf?1459884527=\u0026response-content-disposition=attachment%3B+filename%3DA_Kink_in_the_Line_Does_a_Unique_Lateral.pdf\u0026Expires=1732422414\u0026Signature=U0MPLQBS8gsUJnMB-y8hbtAS-UmTCRN3hZMCkeFacacg6AyUz5DSaeESzhsLNwu0XJutHw0flM7roU0uzDx32L3EDk45Gk-Xc4WzbnbzeG9dohoxjv4s8prSsvNcWX942Ici1ZOlk8ItLDfcMM~zzTabbQhjMGXBIId60prNk3HTauttggS462CLvkYJoBJfAiB1An5rBSEGhEQlH93ImO-8yGSeCKtUOUkxF5t4wGmo0GtsGgcjsgNwbFPX5Q-dX9EQYCQ8JW-cgLbWBZxOxW10JLC~q~Bq8tGF60m8blbNu7ogeb8PFoTlYHSZz06mQNQbzRM97CjHs1ZVcdVN9Q__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"slug":"A_Kink_in_the_Line_Does_a_Unique_Lateral_Line_Peculiarity_Really_Characterize_Lake_Malawis_Huge_Haplochromine_Species_Flock_Teleostei_Cichlidae_","translated_slug":"","page_count":18,"language":"en","content_type":"Work","owner":{"id":1462379,"first_name":"Michael","middle_initials":"K","last_name":"Oliver","page_name":"MichaelKOliver","domain_name":"99main","created_at":"2012-04-07T13:04:06.491-07:00","display_name":"Michael K Oliver","url":"https://99main.academia.edu/MichaelKOliver"},"attachments":[{"id":44445264,"title":"","file_type":"pdf","scribd_thumbnail_url":"https://attachments.academia-assets.com/44445264/thumbnails/1.jpg","file_name":"B57-1-01-Oliver_pp3-20.pdf","download_url":"https://www.academia.edu/attachments/44445264/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&st=MTczMjQxODgxNCw4LjIyMi4yMDguMTQ2&","bulk_download_file_name":"A_Kink_in_the_Line_Does_a_Unique_Lateral.pdf","bulk_download_url":"https://d1wqtxts1xzle7.cloudfront.net/44445264/B57-1-01-Oliver_pp3-20-libre.pdf?1459884527=\u0026response-content-disposition=attachment%3B+filename%3DA_Kink_in_the_Line_Does_a_Unique_Lateral.pdf\u0026Expires=1732422414\u0026Signature=U0MPLQBS8gsUJnMB-y8hbtAS-UmTCRN3hZMCkeFacacg6AyUz5DSaeESzhsLNwu0XJutHw0flM7roU0uzDx32L3EDk45Gk-Xc4WzbnbzeG9dohoxjv4s8prSsvNcWX942Ici1ZOlk8ItLDfcMM~zzTabbQhjMGXBIId60prNk3HTauttggS462CLvkYJoBJfAiB1An5rBSEGhEQlH93ImO-8yGSeCKtUOUkxF5t4wGmo0GtsGgcjsgNwbFPX5Q-dX9EQYCQ8JW-cgLbWBZxOxW10JLC~q~Bq8tGF60m8blbNu7ogeb8PFoTlYHSZz06mQNQbzRM97CjHs1ZVcdVN9Q__\u0026Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA"}],"research_interests":[{"id":135987,"name":"Hypothesis testing","url":"https://www.academia.edu/Documents/in/Hypothesis_testing"},{"id":216604,"name":"Cichlidae","url":"https://www.academia.edu/Documents/in/Cichlidae"},{"id":452259,"name":"Lake Malawi","url":"https://www.academia.edu/Documents/in/Lake_Malawi"},{"id":1418924,"name":"Lateral Line","url":"https://www.academia.edu/Documents/in/Lateral_Line"}],"urls":[]}, dispatcherData: dispatcherData }); $(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") }); </script> <div class="js-work-strip profile--work_container" data-work-id="2998160"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/2998160/Hemitaeniochromis_brachyrhynchus_a_new_species_of_cichlid_fish_from_Lake_Mala%C5%B5i_with_comments_on_some_other_supposed_members_of_the_genus_Teleostei_Cichlidae_"><img alt="Research paper thumbnail of Hemitaeniochromis brachyrhynchus, a new species of cichlid fish from Lake Malaŵi, with comments on some other supposed members of the genus (Teleostei: Cichlidae)" class="work-thumbnail" src="https://attachments.academia-assets.com/30947824/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/2998160/Hemitaeniochromis_brachyrhynchus_a_new_species_of_cichlid_fish_from_Lake_Mala%C5%B5i_with_comments_on_some_other_supposed_members_of_the_genus_Teleostei_Cichlidae_">Hemitaeniochromis brachyrhynchus, a new species of cichlid fish from Lake Malaŵi, with comments on some other supposed members of the genus (Teleostei: Cichlidae)</a></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">Hemitaeniochromis brachyrhynchus, an anatomically distinctive and apparently rare new cichlid, is...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">Hemitaeniochromis brachyrhynchus, an anatomically distinctive and apparently rare new cichlid, is described and illustrated <br />from specimens collected at two widely separated localities within Lake Malaŵi. It is easily distinguished from H. urotaenia, type species of the genus, by its narrow lacrimal bone which is only one-third of the orbit length, a character thought to be unique not only in Hemitaeniochromis but among all known Lake Malaŵi cichlids. The genus Hemitaeniochromis Eccles & Trewavas (1989) is redefined to allow provisional inclusion of this new species. Two species placed in Protomelas by Eccles & Trewavas (1989) [P. insignis (Trewavas) and P. spilopterus (Trewavas)] were recently transferred to Hemitaeniochromis by some authors, without much evidence. The generic placement of these controversial taxa, and <br />of several undescribed species known only from underwater photographs, is briefly reconsidered.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="04fefe6b247975531353cfff6668b3c3" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":30947824,"asset_id":2998160,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/30947824/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="2998160"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="2998160"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 2998160; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=2998160]").text(description); $(".js-view-count[data-work-id=2998160]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 2998160; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='2998160']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 2998160, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "04fefe6b247975531353cfff6668b3c3" } } $('.js-work-strip[data-work-id=2998160]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":2998160,"title":"Hemitaeniochromis brachyrhynchus, a new species of cichlid fish from Lake Malaŵi, with comments on some other supposed members of the genus (Teleostei: Cichlidae)","translated_title":"","metadata":{"abstract":"Hemitaeniochromis brachyrhynchus, an anatomically distinctive and apparently rare new cichlid, is described and illustrated\r\nfrom specimens collected at two widely separated localities within Lake Malaŵi. 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$(this).data('initialized', true); } }); $a.trackClickSource(".js-work-strip-work-link", "profile_work_strip") }); </script> </div><div class="profile--tab_content_container js-tab-pane tab-pane" data-section-id="5968147" id="thesischapters"><div class="js-work-strip profile--work_container" data-work-id="29111715"><div class="profile--work_thumbnail hidden-xs"><a class="js-work-strip-work-link" data-click-track="profile-work-strip-thumbnail" href="https://www.academia.edu/29111715/Systematics_of_African_cichlid_fishes_Determination_of_the_most_primitive_taxon_and_studies_on_the_haplochromines_of_Lake_Malawi_Teleostei_Cichlidae_"><img alt="Research paper thumbnail of Systematics of African cichlid fishes: Determination of the most primitive taxon, and studies on the haplochromines of Lake Malawi (Teleostei: Cichlidae)" class="work-thumbnail" src="https://attachments.academia-assets.com/49559788/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/29111715/Systematics_of_African_cichlid_fishes_Determination_of_the_most_primitive_taxon_and_studies_on_the_haplochromines_of_Lake_Malawi_Teleostei_Cichlidae_">Systematics of African cichlid fishes: Determination of the most primitive taxon, and studies on the haplochromines of Lake Malawi (Teleostei: Cichlidae)</a></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">The cichlids of Africa present many challenges to the taxonomist. Diverse systematic problems at ...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">The cichlids of Africa present many challenges to the taxonomist. Diverse systematic problems at three levels in cichlid phylogeny form the subject matter of this dissertation. At the most general level, I summarize current morphological evidence that supports cichlid monophyly and the most-inclusive groups of cichlids. With this review as a framework, I argue that the most phylogenetically primitive African cichlid is Heterochromis multidens, a little-known large cichlid from the Zaire River basin. I redescribe this monotypic genus and species and illustrate much of its osteology. It possesses a distinctive suite of derived characters, but is uniquely primitive among African cichlids in infraorbital morphology and other features.<br /><br />At a less inclusive phylogenetic level, I examine the controversy over the monophyletic status of the "species flock" of haplochromine cichlids in Lake Malawi. A derived type of anal-fin markings present in a subset of the Malawian fauna is shared with many non-Malawian haplochromines, indicating that the species flock of Lake Malawi is not monophyletic. The two (or more) faunal components of Lake Malawi are ecologically distinct.<br /><br />The remaining major parts of the study are concerned with particular assemblages of haplochromine cichlids in Lake Malawi. I present a detailed phylogenetic analysis of the Cyrtocara livingstonii species-group of ambush predators, and characterize a new species in the group. I also describe three new species of haplochromines with three lateral spots and present a key to the three-spot assemblage. Intralacustrine dispersion of Malawi cichlids is briefly addressed in appended material.</span></div><div class="wp-workCard_item wp-workCard--actions"><span class="work-strip-bookmark-button-container"></span><a id="d053b380692dd1b9669646a8aefbb3e3" class="wp-workCard--action" rel="nofollow" data-click-track="profile-work-strip-download" data-download="{"attachment_id":49559788,"asset_id":29111715,"asset_type":"Work","button_location":"profile"}" href="https://www.academia.edu/attachments/49559788/download_file?st=MTczMjQxODgxNSw4LjIyMi4yMDguMTQ2&s=profile"><span><i class="fa fa-arrow-down"></i></span><span>Download</span></a><span class="wp-workCard--action visible-if-viewed-by-owner inline-block" style="display: none;"><span class="js-profile-work-strip-edit-button-wrapper profile-work-strip-edit-button-wrapper" data-work-id="29111715"><a class="js-profile-work-strip-edit-button" tabindex="0"><span><i class="fa fa-pencil"></i></span><span>Edit</span></a></span></span><span id="work-strip-rankings-button-container"></span></div><div class="wp-workCard_item wp-workCard--stats"><span><span><span class="js-view-count view-count u-mr2x" data-work-id="29111715"><i class="fa fa-spinner fa-spin"></i></span><script>$(function () { var workId = 29111715; window.Academia.workViewCountsFetcher.queue(workId, function (count) { var description = window.$h.commaizeInt(count) + " " + window.$h.pluralize(count, 'View'); $(".js-view-count[data-work-id=29111715]").text(description); $(".js-view-count[data-work-id=29111715]").attr('title', description).tooltip(); }); });</script></span></span><span><span class="percentile-widget hidden"><span class="u-mr2x work-percentile"></span></span><script>$(function () { var workId = 29111715; window.Academia.workPercentilesFetcher.queue(workId, function (percentileText) { var container = $(".js-work-strip[data-work-id='29111715']"); container.find('.work-percentile').text(percentileText.charAt(0).toUpperCase() + percentileText.slice(1)); container.find('.percentile-widget').show(); container.find('.percentile-widget').removeClass('hidden'); }); });</script></span><span><script>$(function() { new Works.PaperRankView({ workId: 29111715, container: "", }); });</script></span></div><div id="work-strip-premium-row-container"></div></div></div><script> require.config({ waitSeconds: 90 })(["https://a.academia-assets.com/assets/wow_profile-f77ea15d77ce96025a6048a514272ad8becbad23c641fc2b3bd6e24ca6ff1932.js","https://a.academia-assets.com/assets/work_edit-ad038b8c047c1a8d4fa01b402d530ff93c45fee2137a149a4a5398bc8ad67560.js"], function() { // from javascript_helper.rb var dispatcherData = {} if (true){ window.WowProfile.dispatcher = window.WowProfile.dispatcher || _.clone(Backbone.Events); dispatcherData = { dispatcher: window.WowProfile.dispatcher, downloadLinkId: "d053b380692dd1b9669646a8aefbb3e3" } } $('.js-work-strip[data-work-id=29111715]').each(function() { if (!$(this).data('initialized')) { new WowProfile.WorkStripView({ el: this, workJSON: {"id":29111715,"title":"Systematics of African cichlid fishes: Determination of the most primitive taxon, and studies on the haplochromines of Lake Malawi (Teleostei: Cichlidae)","translated_title":"","metadata":{"abstract":"The cichlids of Africa present many challenges to the taxonomist. 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At the most general level, I summarize current morphological evidence that supports cichlid monophyly and the most-inclusive groups of cichlids. With this review as a framework, I argue that the most phylogenetically primitive African cichlid is Heterochromis multidens, a little-known large cichlid from the Zaire River basin. I redescribe this monotypic genus and species and illustrate much of its osteology. It possesses a distinctive suite of derived characters, but is uniquely primitive among African cichlids in infraorbital morphology and other features.\n\nAt a less inclusive phylogenetic level, I examine the controversy over the monophyletic status of the \"species flock\" of haplochromine cichlids in Lake Malawi. A derived type of anal-fin markings present in a subset of the Malawian fauna is shared with many non-Malawian haplochromines, indicating that the species flock of Lake Malawi is not monophyletic. The two (or more) faunal components of Lake Malawi are ecologically distinct.\n\nThe remaining major parts of the study are concerned with particular assemblages of haplochromine cichlids in Lake Malawi. I present a detailed phylogenetic analysis of the Cyrtocara livingstonii species-group of ambush predators, and characterize a new species in the group. I also describe three new species of haplochromines with three lateral spots and present a key to the three-spot assemblage. 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Nine new species are added: Lethrinops atrilabris, six new Labeotropheus...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">... Latest changes: (1.) Nine new species are added: Lethrinops atrilabris, six new Labeotropheus, Maylandia melissa, and Pseudotropheus likomae; (2.) Rhamphochromis macrophthalmus is relegated to synonymy with R. longiceps as in ECoF; (3.) 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Version of 3-Jan-2024" class="work-thumbnail" src="https://attachments.academia-assets.com/111013687/thumbnails/1.jpg" /></a></div><div class="wp-workCard wp-workCard_itemContainer"><div class="wp-workCard_item wp-workCard--title"><a class="js-work-strip-work-link text-gray-darker" data-click-track="profile-work-strip-title" href="https://www.academia.edu/114271808/Checklist_of_the_Cichlid_Fishes_of_Lake_Malawi_Lake_Nyasa_Niassa_Version_of_3_Jan_2024">Checklist of the Cichlid Fishes of Lake Malawi (Lake Nyasa/Niassa). Version of 3-Jan-2024</a></div><div class="wp-workCard_item"><span class="js-work-more-abstract-truncated">... Latest changes: (1.) Nine new species are added: Lethrinops atrilabris, six new Labeotropheus...</span><a class="js-work-more-abstract" data-broccoli-component="work_strip.more_abstract" data-click-track="profile-work-strip-more-abstract" href="javascript:;"><span> more </span><span><i class="fa fa-caret-down"></i></span></a><span class="js-work-more-abstract-untruncated hidden">... Latest changes: (1.) Nine new species are added: Lethrinops atrilabris, six new Labeotropheus, Maylandia melissa, and Pseudotropheus likomae; (2.) Rhamphochromis macrophthalmus is relegated to synonymy with R. longiceps as in ECoF; (3.) 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