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title="建议你登录，尽管并非必须。[o]" accesskey="o"><span class="vector-icon mw-ui-icon-logIn mw-ui-icon-wikimedia-logIn"></span> <span>登录</span></a></li> </ul> </div> </div> <div id="p-user-menu-anon-editor" class="vector-menu mw-portlet mw-portlet-user-menu-anon-editor" > <div class="vector-menu-heading"> 未登录编辑者的页面 <a href="/wiki/Help:%E6%96%B0%E6%89%8B%E5%85%A5%E9%97%A8" aria-label="了解有关编辑的更多信息"><span>了解详情</span></a> </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-anoncontribs" class="mw-list-item"><a href="/wiki/Special:%E6%88%91%E7%9A%84%E8%B4%A1%E7%8C%AE" title="来自此IP地址的编辑列表[y]" accesskey="y"><span>贡献</span></a></li><li id="pt-anontalk" class="mw-list-item"><a href="/wiki/Special:%E6%88%91%E7%9A%84%E8%AE%A8%E8%AE%BA%E9%A1%B5" title="对于来自此IP地址编辑的讨论[n]" accesskey="n"><span>讨论</span></a></li> </ul> </div> </div> </div> </div> </nav> </div> </header> </div> <div class="mw-page-container"> <div class="mw-page-container-inner"> <div class="vector-sitenotice-container"> <div id="siteNotice"></div> </div> <div class="vector-column-start"> <div class="vector-main-menu-container"> <div id="mw-navigation"> <nav id="mw-panel" class="vector-main-menu-landmark" aria-label="站点"> <div id="vector-main-menu-pinned-container" class="vector-pinned-container"> </div> </nav> </div> </div> <div class="vector-sticky-pinned-container"> <nav id="mw-panel-toc" aria-label="目录" data-event-name="ui.sidebar-toc" class="mw-table-of-contents-container vector-toc-landmark"> <div id="vector-toc-pinned-container" class="vector-pinned-container"> <div id="vector-toc" class="vector-toc vector-pinnable-element"> <div class="vector-pinnable-header vector-toc-pinnable-header vector-pinnable-header-pinned" data-feature-name="toc-pinned" data-pinnable-element-id="vector-toc" > <h2 class="vector-pinnable-header-label">目录</h2> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-toc.pin">移至侧栏</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-toc.unpin">隐藏</button> </div> <ul class="vector-toc-contents" id="mw-panel-toc-list"> <li id="toc-mw-content-text" class="vector-toc-list-item vector-toc-level-1"> <a href="#" class="vector-toc-link"> <div class="vector-toc-text">序言</div> </a> </li> <li id="toc-参考文献" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#参考文献"> <div class="vector-toc-text"> <span class="vector-toc-numb">1</span> <span>参考文献</span> </div> </a> <ul id="toc-参考文献-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="目录" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="开关目录" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">开关目录</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">顺式作用元件</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="前往另一种语言写成的文章。12种语言可用" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-12" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">12种语言</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-bs mw-list-item"><a href="https://bs.wikipedia.org/wiki/Cis-regulatorni_element" title="Cis-regulatorni element – 波斯尼亚语" lang="bs" hreflang="bs" data-title="Cis-regulatorni element" data-language-autonym="Bosanski" data-language-local-name="波斯尼亚语" class="interlanguage-link-target"><span>Bosanski</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Cis-Element" title="Cis-Element – 德语" lang="de" hreflang="de" data-title="Cis-Element" data-language-autonym="Deutsch" data-language-local-name="德语" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Cis-regulatory_element" title="Cis-regulatory element – 英语" lang="en" hreflang="en" data-title="Cis-regulatory element" data-language-autonym="English" data-language-local-name="英语" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Genetika_elemento" title="Genetika elemento – 世界语" lang="eo" hreflang="eo" data-title="Genetika elemento" data-language-autonym="Esperanto" data-language-local-name="世界语" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Elemento_regulador_en_cis" title="Elemento regulador en cis – 西班牙语" lang="es" hreflang="es" data-title="Elemento regulador en cis" data-language-autonym="Español" data-language-local-name="西班牙语" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%B3%D8%A7%D9%85%D8%A7%D9%86%D8%AF%D9%87%D9%86%D8%AF%D9%87_%D9%87%D9%85%D8%B3%D9%88" title="ساماندهنده همسو – 波斯语" lang="fa" hreflang="fa" data-title="ساماندهنده همسو" data-language-autonym="فارسی" data-language-local-name="波斯语" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/%C3%89l%C3%A9ment_cis-r%C3%A9gulateur" title="Élément cis-régulateur – 法语" lang="fr" hreflang="fr" data-title="Élément cis-régulateur" data-language-autonym="Français" data-language-local-name="法语" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Elemento_regulador_en_cis" title="Elemento regulador en cis – 加利西亚语" lang="gl" hreflang="gl" data-title="Elemento regulador en cis" data-language-autonym="Galego" data-language-local-name="加利西亚语" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%B7%E3%82%B9%E3%82%A8%E3%83%AC%E3%83%A1%E3%83%B3%E3%83%88" title="シスエレメント – 日语" lang="ja" hreflang="ja" data-title="シスエレメント" data-language-autonym="日本語" data-language-local-name="日语" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Cis-element" title="Cis-element – 荷兰语" lang="nl" hreflang="nl" data-title="Cis-element" data-language-autonym="Nederlands" data-language-local-name="荷兰语" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%A6%D0%B8%D1%81-%D1%80%D0%B5%D0%B3%D1%83%D0%BB%D1%8F%D1%82%D0%BE%D1%80%D0%BD%D1%8B%D0%B5_%D1%8D%D0%BB%D0%B5%D0%BC%D0%B5%D0%BD%D1%82%D1%8B" title="Цис-регуляторные элементы – 俄语" lang="ru" hreflang="ru" data-title="Цис-регуляторные элементы" data-language-autonym="Русский" data-language-local-name="俄语" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Cis-regulativa_element" title="Cis-regulativa element – 瑞典语" lang="sv" hreflang="sv" data-title="Cis-regulativa element" data-language-autonym="Svenska" data-language-local-name="瑞典语" class="interlanguage-link-target"><span>Svenska</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q1093165#sitelinks-wikipedia" title="编辑跨语言链接" class="wbc-editpage">编辑链接</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="命名空间"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/%E9%A1%BA%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%85%83%E4%BB%B6" title="浏览条目正文[c]" accesskey="c"><span>条目</span></a></li><li id="ca-talk" class="vector-tab-noicon mw-list-item"><a href="/wiki/Talk:%E9%A1%BA%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%85%83%E4%BB%B6" rel="discussion" title="关于此页面的讨论[t]" accesskey="t"><span>讨论</span></a></li> </ul> </div> </div> <div id="vector-variants-dropdown" class="vector-dropdown " > <input type="checkbox" id="vector-variants-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-variants-dropdown" class="vector-dropdown-checkbox " aria-label="更改语言变体" > <label id="vector-variants-dropdown-label" for="vector-variants-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">不转换</span> </label> <div class="vector-dropdown-content"> <div id="p-variants" class="vector-menu mw-portlet mw-portlet-variants" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-varlang-0" class="selected ca-variants-zh mw-list-item"><a href="/zh/%E9%A1%BA%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%85%83%E4%BB%B6" lang="zh" hreflang="zh"><span>不转换</span></a></li><li id="ca-varlang-1" class="ca-variants-zh-Hans mw-list-item"><a href="/zh-hans/%E9%A1%BA%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%85%83%E4%BB%B6" lang="zh-Hans" hreflang="zh-Hans"><span>简体</span></a></li><li id="ca-varlang-2" class="ca-variants-zh-Hant mw-list-item"><a href="/zh-hant/%E9%A1%BA%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%85%83%E4%BB%B6" lang="zh-Hant" hreflang="zh-Hant"><span>繁體</span></a></li><li id="ca-varlang-3" class="ca-variants-zh-Hans-CN mw-list-item"><a href="/zh-cn/%E9%A1%BA%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%85%83%E4%BB%B6" lang="zh-Hans-CN" hreflang="zh-Hans-CN"><span>大陆简体</span></a></li><li id="ca-varlang-4" class="ca-variants-zh-Hant-HK mw-list-item"><a href="/zh-hk/%E9%A1%BA%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%85%83%E4%BB%B6" lang="zh-Hant-HK" hreflang="zh-Hant-HK"><span>香港繁體</span></a></li><li id="ca-varlang-5" class="ca-variants-zh-Hant-MO mw-list-item"><a href="/zh-mo/%E9%A1%BA%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%85%83%E4%BB%B6" lang="zh-Hant-MO" hreflang="zh-Hant-MO"><span>澳門繁體</span></a></li><li id="ca-varlang-6" class="ca-variants-zh-Hans-MY mw-list-item"><a href="/zh-my/%E9%A1%BA%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%85%83%E4%BB%B6" lang="zh-Hans-MY" hreflang="zh-Hans-MY"><span>大马简体</span></a></li><li id="ca-varlang-7" class="ca-variants-zh-Hans-SG mw-list-item"><a href="/zh-sg/%E9%A1%BA%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%85%83%E4%BB%B6" lang="zh-Hans-SG" hreflang="zh-Hans-SG"><span>新加坡简体</span></a></li><li id="ca-varlang-8" class="ca-variants-zh-Hant-TW mw-list-item"><a href="/zh-tw/%E9%A1%BA%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%85%83%E4%BB%B6" lang="zh-Hant-TW" hreflang="zh-Hant-TW"><span>臺灣正體</span></a></li> </ul> </div> </div> </div> </div> </nav> </div> <div id="right-navigation" class="vector-collapsible"> <nav aria-label="查看"> <div id="p-views" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-views" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-view" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/%E9%A1%BA%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%85%83%E4%BB%B6"><span>阅读</span></a></li><li id="ca-edit" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=%E9%A1%BA%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%85%83%E4%BB%B6&action=edit" title="编辑该页面[e]" accesskey="e"><span>编辑</span></a></li><li id="ca-history" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=%E9%A1%BA%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%85%83%E4%BB%B6&action=history" title="本页面的早前版本。[h]" accesskey="h"><span>查看历史</span></a></li> </ul> </div> </div> </nav> <nav class="vector-page-tools-landmark" aria-label="页面工具"> <div id="vector-page-tools-dropdown" class="vector-dropdown vector-page-tools-dropdown" > <input type="checkbox" id="vector-page-tools-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-tools-dropdown" class="vector-dropdown-checkbox " aria-label="工具" > <label id="vector-page-tools-dropdown-label" for="vector-page-tools-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">工具</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-tools-unpinned-container" class="vector-unpinned-container"> <div id="vector-page-tools" class="vector-page-tools vector-pinnable-element"> <div class="vector-pinnable-header vector-page-tools-pinnable-header vector-pinnable-header-unpinned" data-feature-name="page-tools-pinned" data-pinnable-element-id="vector-page-tools" data-pinned-container-id="vector-page-tools-pinned-container" data-unpinned-container-id="vector-page-tools-unpinned-container" > <div class="vector-pinnable-header-label">工具</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" 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class="vector-body-before-content"> <div class="mw-indicators"> <div id="mw-indicator-noteTA-65f4264b" class="mw-indicator"><div class="mw-parser-output"><span class="skin-invert" typeof="mw:File"><span title="本页使用了标题或全文手工转换"><img alt="本页使用了标题或全文手工转换" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Zh_conversion_icon_m.svg/35px-Zh_conversion_icon_m.svg.png" decoding="async" width="35" height="22" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Zh_conversion_icon_m.svg/53px-Zh_conversion_icon_m.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Zh_conversion_icon_m.svg/70px-Zh_conversion_icon_m.svg.png 2x" data-file-width="32" data-file-height="20" /></span></span></div></div> </div> <div id="siteSub" class="noprint">维基百科，自由的百科全书</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="zh" dir="ltr"><div id="noteTA-65f4264b" class="noteTA"><div class="noteTA-local"><div data-noteta-code="zh-hans:顺式;zh-tw:同側"></div><div data-noteta-code="zh-hans:反式;zh-tw:異側"></div></div></div> <style data-mw-deduplicate="TemplateStyles:r83732972">.mw-parser-output .ambox{border:1px solid #a2a9b1;border-left:10px solid #36c;background-color:#fbfbfb;box-sizing:border-box}.mw-parser-output .ambox+link+.ambox,.mw-parser-output .ambox+link+style+.ambox,.mw-parser-output .ambox+link+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+style+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+link+.ambox{margin-top:-1px}html body.mediawiki .mw-parser-output .ambox.mbox-small-left{margin:4px 1em 4px 0;overflow:hidden;width:238px;border-collapse:collapse;font-size:88%;line-height:1.25em}.mw-parser-output .ambox-speedy{border-left:10px solid #b32424;background-color:#fee7e6}.mw-parser-output .ambox-delete{border-left:10px solid #b32424}.mw-parser-output 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.mw-parser-output .ambox-delete{border-left-color:#b32424!important}html.skin-theme-clientpref-night .mw-parser-output .ambox-speedy{background-color:#300!important}html.skin-theme-clientpref-night .mw-parser-output .ambox-content{border-left-color:#f28500!important}html.skin-theme-clientpref-night .mw-parser-output .ambox-style{border-left-color:#fc3!important}html.skin-theme-clientpref-night .mw-parser-output .ambox-move{border-left-color:#9932cc!important}html.skin-theme-clientpref-night .mw-parser-output .ambox-protection{border-left-color:#a2a9b1!important}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .ambox{border-left-color:#36c!important}html.skin-theme-clientpref-os .mw-parser-output .ambox-speedy,html.skin-theme-clientpref-os .mw-parser-output .ambox-delete{border-left-color:#b32424!important}html.skin-theme-clientpref-os .mw-parser-output .ambox-speedy{background-color:#300!important}html.skin-theme-clientpref-os .mw-parser-output .ambox-content{border-left-color:#f28500!important}html.skin-theme-clientpref-os .mw-parser-output .ambox-style{border-left-color:#fc3!important}html.skin-theme-clientpref-os .mw-parser-output .ambox-move{border-left-color:#9932cc!important}html.skin-theme-clientpref-os .mw-parser-output .ambox-protection{border-left-color:#a2a9b1!important}}</style><table class="box-Unreferenced plainlinks metadata ambox ambox-content" role="presentation"><tbody><tr><td class="mbox-image"><div style="width:52px"><span typeof="mw:File"><a href="/wiki/File:Tango-nosources.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4e/Tango-nosources.svg/45px-Tango-nosources.svg.png" decoding="async" width="45" height="45" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4e/Tango-nosources.svg/68px-Tango-nosources.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4e/Tango-nosources.svg/90px-Tango-nosources.svg.png 2x" data-file-width="48" data-file-height="48" /></a></span></div></td><td class="mbox-text"><div class="mbox-text-span">此條目<b>没有列出任何<a href="/wiki/Wikipedia:%E5%88%97%E6%98%8E%E6%9D%A5%E6%BA%90" title="Wikipedia:列明来源">参考或来源</a></b>。<span class="hide-when-compact"></span> <small class="date-container"><i>(<span class="date">2021年5月7日</span>)</i></small><span class="hide-when-compact"><br /><small>維基百科所有的內容都應該<a href="/wiki/Wikipedia:%E5%8F%AF%E4%BE%9B%E6%9F%A5%E8%AD%89" title="Wikipedia:可供查證">可供查證</a>。请协助補充<a href="/wiki/Wikipedia:%E5%8F%AF%E9%9D%A0%E6%9D%A5%E6%BA%90" title="Wikipedia:可靠来源">可靠来源</a>以<a class="external text" href="https://zh.wikipedia.org/w/index.php?title=%E9%A1%BA%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%85%83%E4%BB%B6&action=edit">改善这篇条目</a>。无法查证的內容可能會因為異議提出而被移除。</small></span><span class="hide-when-compact"></span></div></td></tr></tbody></table> <p><b>顺式作用元件</b>（cis-regulatory element，CRE），是与结构基因串联的一段非编码DNA序列，一般位于转录点上游，作用是调节邻近基因的转录。CREs是遗传调控网络的重要组成部分，进而控制<a href="/wiki/%E5%BD%A2%E6%80%81%E5%8F%91%E7%94%9F" title="形态发生">形态发生</a>、解剖学的发展以及胚胎发育的其他方面。（顺式作用元件是DNA序列，而<a href="/wiki/%E5%8F%8D%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%9B%A0%E5%AD%90" title="反式作用因子">反式作用因子</a>是调控蛋白） </p><p>CREs存在于它们调节的基因附近。CRE通常通过与<a href="/wiki/%E8%BD%AC%E5%BD%95%E5%9B%A0%E5%AD%90" title="转录因子">转录因子</a>结合来调节基因转录。单个转录因子可以与许多CRE结合，因此控制许多基因的表达（多效性）。拉丁语前缀顺式意指“在这一侧”，即在与要转录的基因相同的DNA分子上。CRE通常但不总是位于转录位点的上游。<a href="/wiki/%E5%90%AF%E5%8A%A8%E5%AD%90" class="mw-redirect" title="启动子">启动子</a>和<a href="/wiki/%E7%B5%82%E6%AD%A2%E5%AD%90" title="終止子">终止子</a>是顺式作用位点的一个简单的例子，它们可以和反式作用的，如RNA聚合酶识别，从而达到调控基因表达的目的<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>。 </p><p>CRE与<span class="ilh-all" data-orig-title="反式调节元件" data-lang-code="en" data-lang-name="英语" data-foreign-title="Trans-regulatory element"><span class="ilh-page"><a href="/w/index.php?title=%E5%8F%8D%E5%BC%8F%E8%B0%83%E8%8A%82%E5%85%83%E4%BB%B6&action=edit&redlink=1" class="new" title="反式调节元件（页面不存在）">反式调节元件</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Trans-regulatory_element" class="extiw" title="en:Trans-regulatory element"><span lang="en" dir="auto">Trans-regulatory element</span></a></span>）</span></span>（trans-regulatory element，TRE）形成对比。 TRE编码转录因子。 </p> <div class="mw-heading mw-heading2"><h2 id="参考文献"><span id=".E5.8F.82.E8.80.83.E6.96.87.E7.8C.AE"></span>参考文献</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%E9%A1%BA%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%85%83%E4%BB%B6&action=edit&section=1" title="编辑章节：参考文献"><span>编辑</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="reflist" style="list-style-type: decimal;"> <ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text"><cite class="citation book">Krebs, Jocelyn E.; Goldstein, Elliott S.; Kilpatrick, Stephen T.; Lewin, Benjamin. Operon. <a rel="nofollow" class="external text" href="https://archive.org/details/lewinsgenesxii0000kreb">Lewin's genes XII</a> Twelfth edition. Burlington, Mass: Jones & Bartlett Learning. 2018. <a href="/wiki/Special:%E7%BD%91%E7%BB%9C%E4%B9%A6%E6%BA%90/978-1-284-10449-3" title="Special:网络书源/978-1-284-10449-3"><span title="国际标准书号">ISBN</span> 978-1-284-10449-3</a>.</cite><span title="ctx_ver=Z39.88-2004&rfr_id=info%3Asid%2Fzh.wikipedia.org%3A%E9%A1%BA%E5%BC%8F%E4%BD%9C%E7%94%A8%E5%85%83%E4%BB%B6&rft.atitle=Operon&rft.au=Goldstein%2C+Elliott+S.&rft.au=Kilpatrick%2C+Stephen+T.&rft.au=Lewin%2C+Benjamin&rft.aufirst=Jocelyn+E.&rft.aulast=Krebs&rft.btitle=Lewin%27s+genes+XII&rft.date=2018&rft.edition=Twelfth+edition&rft.genre=bookitem&rft.isbn=978-1-284-10449-3&rft.place=Burlington%2C+Mass&rft.pub=Jones+%26+Bartlett+Learning&rft_id=https%3A%2F%2Farchive.org%2Fdetails%2Flewinsgenesxii0000kreb&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook" class="Z3988"><span style="display:none;"> </span></span><span class="citation-comment" style="display:none; color:#33aa33"> 引文格式1维护：冗余文本 (<a href="/wiki/Category:%E5%BC%95%E6%96%87%E6%A0%BC%E5%BC%8F1%E7%BB%B4%E6%8A%A4%EF%BC%9A%E5%86%97%E4%BD%99%E6%96%87%E6%9C%AC" title="Category:引文格式1维护：冗余文本">link</a>)</span></span> </li> </ol></div> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r84265675">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol ol,.mw-parser-output .hlist ol ul,.mw-parser-output .hlist ul dl,.mw-parser-output .hlist ul ol,.mw-parser-output .hlist ul ul{display:inline}.mw-parser-output .hlist .mw-empty-li{display:none}.mw-parser-output .hlist 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.navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:110%;margin:0 8em}.mw-parser-output .navbar-ct-mini{font-size:110%;margin:0 5em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style></div><div role="navigation" class="navbox" aria-labelledby="基因表达" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="collapsible-title navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r84265675"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r84244141"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:%E5%9F%BA%E5%9B%A0%E8%A1%A8%E8%BE%BE" title="Template:基因表达"><abbr title="查看该模板">查</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:%E5%9F%BA%E5%9B%A0%E8%A1%A8%E8%BE%BE" title="Template talk:基因表达"><abbr title="讨论该模板">论</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:%E7%BC%96%E8%BE%91%E9%A1%B5%E9%9D%A2/Template:%E5%9F%BA%E5%9B%A0%E8%A1%A8%E8%BE%BE" title="Special:编辑页面/Template:基因表达"><abbr title="编辑该模板">编</abbr></a></li></ul></div><div id="基因表达" style="font-size:110%;margin:0 5em"><a href="/wiki/%E5%9F%BA%E5%9B%A0%E8%A1%A8%E7%8F%BE" title="基因表現">基因表达</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/%E9%81%97%E4%BC%A0%E5%AD%A6%E5%85%A5%E9%97%A8" title="遗传学入门">遗传学入门</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0px"><div style="padding:0em 0.25em"> <ul><li><a href="/wiki/%E9%81%97%E4%BC%A0%E5%AF%86%E7%A0%81" title="遗传密码">遗传密码</a></li> <li><a href="/wiki/%E4%B8%AD%E5%BF%83%E6%B3%95%E5%89%87" title="中心法則">经典中心法则</a>： <ul><li><a href="/wiki/%E8%84%B1%E6%B0%A7%E6%A0%B8%E7%B3%96%E6%A0%B8%E9%85%B8" title="脱氧核糖核酸">DNA</a> <a href="/wiki/%E8%BD%AC%E5%BD%95" title="转录">→</a> <a href="/wiki/%E4%BF%A1%E4%BD%BF%E6%A0%B8%E7%B3%96%E6%A0%B8%E9%85%B8" title="信使核糖核酸">RNA</a> <a href="/wiki/%E7%BF%BB%E8%AD%AF_(%E7%94%9F%E7%89%A9%E5%AD%B8)" title="翻譯 (生物學)">→</a> <a href="/wiki/%E8%9B%8B%E7%99%BD%E8%B4%A8" title="蛋白质">蛋白质</a></li></ul></li> <li><a href="/wiki/%E4%B8%AD%E5%BF%83%E6%B3%95%E5%89%87#中心法则的扩充" title="中心法則">扩充中心法则</a> <ul><li><a href="/wiki/RNA%E8%A4%87%E8%A3%BD%E9%85%B6" title="RNA複製酶">RNA → RNA</a></li> <li><a href="/wiki/%E9%80%86%E8%BD%AC%E5%BD%95" class="mw-redirect" title="逆转录">RNA → DNA</a></li> <li><a href="/wiki/%E8%9B%8B%E7%99%BD%E8%B4%A8" title="蛋白质">蛋白质 → 蛋白质</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/%E8%BD%AC%E5%BD%95" title="转录">转录</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0px"><div style="padding:0em 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8.0em;font-weight: normal;">类型</th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0px"><div style="padding:0em 0.25em"> <ul><li><span class="ilh-all" data-orig-title="细菌转录" data-lang-code="en" data-lang-name="英语" data-foreign-title="Bacterial transcription"><span class="ilh-page"><a href="/w/index.php?title=%E7%BB%86%E8%8F%8C%E8%BD%AC%E5%BD%95&action=edit&redlink=1" class="new" title="细菌转录（页面不存在）">细菌转录</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Bacterial_transcription" class="extiw" title="en:Bacterial transcription"><span lang="en" dir="auto">Bacterial transcription</span></a></span>）</span></span></li> <li><span class="ilh-all" data-orig-title="Archaeal transcription" data-lang-code="en" data-lang-name="英语" data-foreign-title="Archaeal transcription"><span class="ilh-page"><a href="/w/index.php?title=Archaeal_transcription&action=edit&redlink=1" class="new" title="Archaeal transcription（页面不存在）">古菌转录</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Archaeal_transcription" class="extiw" title="en:Archaeal transcription"><span lang="en" dir="auto">Archaeal transcription</span></a></span>）</span></span></li> <li><a href="/wiki/%E7%9C%9F%E6%A0%B8%E8%BD%AC%E5%BD%95" title="真核转录">真核转录</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.0em;font-weight: normal;">关键元素</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0px"><div style="padding:0em 0.25em"> <ul><li><a href="/wiki/%E8%BD%AC%E5%BD%95%E5%9B%A0%E5%AD%90" title="转录因子">转录因子</a></li> <li><a href="/wiki/RNA%E8%81%9A%E5%90%88%E9%85%B6" title="RNA聚合酶">RNA聚合酶</a></li> <li><a href="/wiki/%E5%90%AF%E5%8A%A8%E5%AD%90" class="mw-redirect" title="启动子">启动子</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.0em;font-weight: normal;"><a href="/wiki/%E8%BD%AC%E5%BD%95%E5%90%8E%E4%BF%AE%E9%A5%B0" title="转录后修饰">转录后修饰</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0px"><div style="padding:0em 0.25em"> <ul><li><a href="/wiki/%E5%89%8D%E4%BF%A1%E4%BD%BFRNA" class="mw-redirect" title="前信使RNA">hnRNA</a></li> <li><a href="/wiki/5%E2%80%B2%E7%AB%AF%E5%B8%BD" title="5′端帽">5'端加帽</a></li> <li><a href="/wiki/RNA%E5%89%AA%E6%8E%A5" title="RNA剪接">RNA剪接</a></li> <li><a href="/wiki/%E5%A4%9A%E8%85%BA%E8%8B%B7%E9%85%B8%E5%8C%96" title="多腺苷酸化">多腺苷酸化</a></li> <li><a href="/wiki/%E7%B5%84%E8%9B%8B%E7%99%BD%E4%B9%99%E9%86%AF%E5%8C%96" title="組蛋白乙醯化">組蛋白乙醯化</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/%E7%BF%BB%E8%AD%AF_(%E7%94%9F%E7%89%A9%E5%AD%B8)" title="翻譯 (生物學)">翻译</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0px"><div style="padding:0em 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:8.0em;font-weight: normal;">类型</th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0px"><div style="padding:0em 0.25em"> <ul><li><a href="/wiki/%E5%8E%9F%E6%A0%B8%E7%BF%BB%E8%AF%91" title="原核翻译">原核翻译</a></li> <li><span class="ilh-all" data-orig-title="Archaeal translation" data-lang-code="en" data-lang-name="英语" data-foreign-title="Archaeal translation"><span class="ilh-page"><a href="/w/index.php?title=Archaeal_translation&action=edit&redlink=1" class="new" title="Archaeal translation（页面不存在）">古菌翻译</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Archaeal_translation" class="extiw" title="en:Archaeal translation"><span lang="en" dir="auto">Archaeal translation</span></a></span>）</span></span></li> <li><a href="/wiki/%E7%9C%9F%E6%A0%B8%E7%BF%BB%E8%AF%91" title="真核翻译">真核翻译</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:8.0em;font-weight: normal;">关键元素</th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0px"><div style="padding:0em 0.25em"> <ul><li><a href="/wiki/%E6%A0%B8%E7%B3%96%E4%BD%93" title="核糖体">核糖体</a></li> <li><a href="/wiki/TRNA" class="mw-redirect" title="TRNA">tRNA</a></li> <li><a href="/wiki/%E7%BF%BB%E8%AF%91%E5%90%8E%E4%BF%AE%E9%A5%B0" title="翻译后修饰">翻译后修饰</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/%E5%9F%BA%E5%9B%A0%E8%A1%A8%E8%BE%BE%E8%B0%83%E6%8E%A7" title="基因表达调控">基因表达调控</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0px"><div style="padding:0em 0.25em"> <ul><li><a href="/wiki/%E8%A1%A8%E8%A7%80%E9%81%BA%E5%82%B3%E5%AD%B8" title="表觀遺傳學">表观遗传学调节</a> <ul><li><a href="/wiki/%E5%9F%BA%E5%9B%A0%E7%B5%84%E9%8A%98%E5%8D%B0" title="基因組銘印">基因組銘印</a> 等</li></ul></li> <li><a href="/wiki/%E8%BD%AC%E5%BD%95%E8%B0%83%E6%8E%A7" title="转录调控">转录调控</a> <ul><li><span class="ilh-all" data-orig-title="基因调控网络" data-lang-code="en" data-lang-name="英语" data-foreign-title="Gene regulatory network"><span class="ilh-page"><a href="/w/index.php?title=%E5%9F%BA%E5%9B%A0%E8%B0%83%E6%8E%A7%E7%BD%91%E7%BB%9C&action=edit&redlink=1" class="new" title="基因调控网络（页面不存在）">基因调控网络</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Gene_regulatory_network" class="extiw" title="en:Gene regulatory network"><span lang="en" dir="auto">Gene regulatory network</span></a></span>）</span></span></li> <li><a class="mw-selflink selflink">顺式作用元件</a></li></ul></li> <li><a href="/wiki/%E4%B9%B3%E7%B3%96%E6%93%8D%E7%B8%B1%E5%AD%90" title="乳糖操縱子">乳糖操縱子</a></li> <li><span class="ilh-all" data-orig-title="转录后调控" data-lang-code="en" data-lang-name="英语" data-foreign-title="Post-transcriptional regulation"><span class="ilh-page"><a href="/w/index.php?title=%E8%BD%AC%E5%BD%95%E5%90%8E%E8%B0%83%E6%8E%A7&action=edit&redlink=1" class="new" title="转录后调控（页面不存在）">转录后调控</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Post-transcriptional_regulation" class="extiw" title="en:Post-transcriptional regulation"><span lang="en" dir="auto">Post-transcriptional regulation</span></a></span>）</span></span> <ul><li><a href="/wiki/P%E4%BD%93" class="mw-redirect" title="P体">P体</a></li> <li><a href="/wiki/%E5%8F%AF%E5%8F%98%E5%89%AA%E6%8E%A5" class="mw-redirect" title="可变剪接">可变剪接</a></li> <li><a href="/wiki/MiRNA" class="mw-redirect" title="MiRNA">miRNA</a> 等</li></ul></li> <li><span class="ilh-all" data-orig-title="翻译调控" data-lang-code="en" data-lang-name="英语" data-foreign-title="Translational regulation"><span class="ilh-page"><a href="/w/index.php?title=%E7%BF%BB%E8%AF%91%E8%B0%83%E6%8E%A7&action=edit&redlink=1" class="new" title="翻译调控（页面不存在）">翻译调控</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Translational_regulation" class="extiw" title="en:Translational regulation"><span lang="en" dir="auto">Translational regulation</span></a></span>）</span></span></li> <li><span class="ilh-all" data-orig-title="翻译后调控" data-lang-code="en" data-lang-name="英语" data-foreign-title="Post-translational regulation"><span class="ilh-page"><a href="/w/index.php?title=%E7%BF%BB%E8%AF%91%E5%90%8E%E8%B0%83%E6%8E%A7&action=edit&redlink=1" class="new" title="翻译后调控（页面不存在）">翻译后调控</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Post-translational_regulation" class="extiw" title="en:Post-translational regulation"><span lang="en" dir="auto">Post-translational regulation</span></a></span>）</span></span> <ul><li><a href="/wiki/%E7%A3%B7%E9%85%B8%E5%8C%96" title="磷酸化">可逆的</a></li> <li><a href="/wiki/%E8%9B%8B%E7%99%BD%E9%85%B6%E8%A7%A3" title="蛋白酶解">不可逆的</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">该领域名人</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0px"><div style="padding:0em 0.25em"> <ul><li><a href="/wiki/%E6%96%B9%E6%96%AF%E8%8F%AF%C2%B7%E8%B3%88%E5%85%8B%E6%9F%8F" title="方斯華·賈克柏">方斯華·賈克柏</a></li> <li><a href="/wiki/%E8%B3%88%E5%85%8B%C2%B7%E8%8E%AB%E8%AB%BE" title="賈克·莫諾">賈克·莫諾</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2"><div></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:%E8%9B%8B%E7%99%BD%E8%B4%A8%E7%94%9F%E7%89%A9%E5%90%88%E6%88%90%E5%AF%BC%E8%88%AA" title="Template:蛋白质生物合成导航"><abbr title="查看该模板">查</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:%E8%9B%8B%E7%99%BD%E8%B4%A8%E7%94%9F%E7%89%A9%E5%90%88%E6%88%90%E5%AF%BC%E8%88%AA" title="Template talk:蛋白质生物合成导航"><abbr title="讨论该模板">论</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:%E7%BC%96%E8%BE%91%E9%A1%B5%E9%9D%A2/Template:%E8%9B%8B%E7%99%BD%E8%B4%A8%E7%94%9F%E7%89%A9%E5%90%88%E6%88%90%E5%AF%BC%E8%88%AA" title="Special:编辑页面/Template:蛋白质生物合成导航"><abbr title="编辑该模板">编</abbr></a></li></ul></div><div id="遗传学索引" style="font-size:110%;margin:0 5em">遗传学索引</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">描述</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0px"><div style="padding:0em 0.25em"> <ul><li><a href="/wiki/Template:%E7%94%9F%E7%89%A9%E5%8C%96%E5%AD%A6%E6%A8%A1%E6%9D%BF%E6%80%BB%E8%A7%88" title="Template:生物化学模板总览">生化</a></li> <li><a href="/wiki/Template:%E5%9F%BA%E5%9B%A0%E8%A1%A8%E8%BE%BE" title="Template:基因表达">基因表达</a></li> <li>DNA <ul><li><a href="/wiki/Template:DNA%E5%A4%8D%E5%88%B6" title="Template:DNA复制">复制</a></li> <li><a href="/wiki/Template:%E7%BB%86%E8%83%9E%E5%91%A8%E6%9C%9F%E8%9B%8B%E7%99%BD%E8%B4%A8" title="Template:细胞周期蛋白质">周期蛋白</a></li> <li><a href="/wiki/Template:%E5%9F%BA%E5%9B%A0%E9%87%8D%E7%BB%84" title="Template:基因重组">基因重组</a></li> <li><a href="/wiki/Template:DNA%E4%BF%AE%E5%A4%8D" title="Template:DNA修复">修复</a></li></ul></li> <li><a href="/wiki/Template:%E8%BD%AC%E5%BD%95" title="Template:转录">转录</a> <ul><li><a href="/wiki/Template:%E8%BD%AC%E5%BD%95%E5%9B%A0%E5%AD%90%E4%B8%8E%E7%BB%86%E8%83%9E%E5%86%85%E5%8F%97%E4%BD%93" title="Template:转录因子与细胞内受体">转录因子与细胞内受体</a></li> <li><a href="/wiki/Template:Transcription_coregulators" title="Template:Transcription coregulators">辅调节因子</a></li> <li><a href="/wiki/Template:RNA%E7%B1%BB%E5%9E%8B" title="Template:RNA类型">RNA</a></li> <li><a href="/wiki/Template:%E9%87%8D%E5%A4%8D%E5%BA%8F%E5%88%97" title="Template:重复序列">重复序列</a></li> <li><a href="/wiki/Template:%E8%BD%AC%E5%BD%95%E5%90%8E%E4%BF%AE%E9%A5%B0" title="Template:转录后修饰">转录后修饰</a></li></ul></li> <li><a href="/wiki/Template:%E9%81%97%E4%BC%A0%E5%AD%A6/%E7%BF%BB%E8%AF%91" title="Template:遗传学/翻译">翻译</a> <ul><li><a href="/wiki/Template:%E6%A0%B8%E7%B3%96%E4%BD%93%E4%BA%9A%E5%9F%BA" title="Template:核糖体亚基">核糖体亚基</a></li> <li><a href="/wiki/Template:%E7%BF%BB%E8%AF%91%E5%90%8E%E4%BF%AE%E9%A5%B0" title="Template:翻译后修饰">翻译后修饰</a></li> <li><span class="ilh-all" data-orig-title="Template:Sorting nexin" data-lang-code="en" data-lang-name="英语" data-foreign-title="Template:Sorting nexin"><span class="ilh-page"><a href="/w/index.php?title=Template:Sorting_nexin&action=edit&redlink=1" class="new" title="Template:Sorting nexin（页面不存在）">分拣蛋白</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Template:Sorting_nexin" class="extiw" title="en:Template:Sorting nexin"><span lang="en" dir="auto">Template:Sorting nexin</span></a></span>）</span></span></li></ul></li> <li><span class="ilh-all" data-orig-title="Template:DNA-binding proteins" data-lang-code="en" data-lang-name="英语" data-foreign-title="Template:DNA-binding proteins"><span class="ilh-page"><a href="/w/index.php?title=Template:DNA-binding_proteins&action=edit&redlink=1" class="new" title="Template:DNA-binding proteins（页面不存在）">DNA结合蛋白</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Template:DNA-binding_proteins" class="extiw" title="en:Template:DNA-binding proteins"><span lang="en" dir="auto">Template:DNA-binding proteins</span></a></span>）</span></span>/<span class="ilh-all" data-orig-title="Template:RNA-binding proteins" data-lang-code="en" data-lang-name="英语" data-foreign-title="Template:RNA-binding proteins"><span class="ilh-page"><a href="/w/index.php?title=Template:RNA-binding_proteins&action=edit&redlink=1" class="new" title="Template:RNA-binding proteins（页面不存在）">RNA结合蛋白</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Template:RNA-binding_proteins" class="extiw" title="en:Template:RNA-binding proteins"><span lang="en" dir="auto">Template:RNA-binding proteins</span></a></span>）</span></span> <ul><li><a href="/wiki/Template:%E6%A0%B8%E7%B3%96%E6%A0%B8%E8%9B%8B%E7%99%BD" title="Template:核糖核蛋白">核糖核蛋白</a>)</li></ul></li> <li>蛋白质 <ul><li><a href="/wiki/Template:Protein_domains" title="Template:Protein domains">结构域</a></li> <li>结构 <ul><li><a href="/wiki/Template:%E8%9B%8B%E7%99%BD%E8%B4%A8%E4%B8%80%E7%BA%A7%E7%BB%93%E6%9E%84" title="Template:蛋白质一级结构">1°</a></li> <li><a href="/wiki/Template:%E8%9B%8B%E7%99%BD%E8%B4%A8%E4%BA%8C%E7%BA%A7%E7%BB%93%E6%9E%84" title="Template:蛋白质二级结构">2°</a></li> <li><a href="/wiki/Template:%E8%9B%8B%E7%99%BD%E8%B4%A8%E8%B6%85%E4%BA%8C%E7%BA%A7%E7%BB%93%E6%9E%84" title="Template:蛋白质超二级结构">2<sup>+</sup>°</a></li> <li><a href="/wiki/Template:%E8%9B%8B%E7%99%BD%E8%B4%A8%E4%B8%89%E7%BA%A7%E7%BB%93%E6%9E%84" title="Template:蛋白质三级结构">3°</a></li> <li><a href="/wiki/Template:%E8%9B%8B%E7%99%BD%E8%B4%A8%E5%9B%9B%E7%BA%A7%E7%BB%93%E6%9E%84" title="Template:蛋白质四级结构">4°</a></li></ul></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">疾病</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0px"><div style="padding:0em 0.25em"> <ul><li><span class="ilh-all" data-orig-title="Template:DNA replication and repair-deficiency disorder" data-lang-code="en" data-lang-name="英语" data-foreign-title="Template:DNA replication and repair-deficiency disorder"><span class="ilh-page"><a href="/w/index.php?title=Template:DNA_replication_and_repair-deficiency_disorder&action=edit&redlink=1" class="new" title="Template:DNA replication and repair-deficiency disorder（页面不存在）">Replication and repair</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Template:DNA_replication_and_repair-deficiency_disorder" class="extiw" title="en:Template:DNA replication and repair-deficiency disorder"><span lang="en" dir="auto">Template:DNA replication and repair-deficiency disorder</span></a></span>）</span></span></li> <li><span class="ilh-all" data-orig-title="Template:Transcription factor and coregulator deficiencies" data-lang-code="en" data-lang-name="英语" data-foreign-title="Template:Transcription factor and coregulator deficiencies"><span class="ilh-page"><a href="/w/index.php?title=Template:Transcription_factor_and_coregulator_deficiencies&action=edit&redlink=1" class="new" title="Template:Transcription factor and coregulator deficiencies（页面不存在）">Transcription factor</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Template:Transcription_factor_and_coregulator_deficiencies" class="extiw" title="en:Template:Transcription factor and coregulator deficiencies"><span lang="en" dir="auto">Template:Transcription factor and coregulator deficiencies</span></a></span>）</span></span></li> <li><span class="ilh-all" data-orig-title="Template:Disorders of transcription and post transcriptional modification" data-lang-code="en" data-lang-name="英语" data-foreign-title="Template:Disorders of transcription and post transcriptional modification"><span class="ilh-page"><a href="/w/index.php?title=Template:Disorders_of_transcription_and_post_transcriptional_modification&action=edit&redlink=1" class="new" title="Template:Disorders of transcription and post transcriptional modification（页面不存在）">转录相关疾病</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Template:Disorders_of_transcription_and_post_transcriptional_modification" class="extiw" title="en:Template:Disorders of transcription and post transcriptional modification"><span lang="en" dir="auto">Template:Disorders of transcription and post transcriptional modification</span></a></span>）</span></span></li> <li><a href="/wiki/Template:Disorders_of_translation_and_posttranslational_modification" title="Template:Disorders of translation and posttranslational modification">轉譯相關疾病</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r84265675"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r84261037"></div><div role="navigation" class="navbox" aria-labelledby="表型发育" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="collapsible-title navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r84265675"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r84244141"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Genarch" title="Template:Genarch"><abbr title="查看该模板">查</abbr></a></li><li class="nv-talk"><a href="/w/index.php?title=Template_talk:Genarch&action=edit&redlink=1" class="new" title="Template talk:Genarch（页面不存在）"><abbr title="讨论该模板">论</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:%E7%BC%96%E8%BE%91%E9%A1%B5%E9%9D%A2/Template:Genarch" title="Special:编辑页面/Template:Genarch"><abbr title="编辑该模板">编</abbr></a></li></ul></div><div id="表型发育" style="font-size:110%;margin:0 5em"><a href="/wiki/%E8%A1%A8%E7%8F%BE%E5%9E%8B" title="表現型">表型</a><a href="/wiki/%E5%8F%91%E8%82%B2%E7%94%9F%E7%89%A9%E5%AD%A6" title="发育生物学">发育</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%">基础概念</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0px"><div style="padding:0em 0.25em"> <ul><li><span class="ilh-all" data-orig-title="基因型-表型的区别" data-lang-code="en" data-lang-name="英语" data-foreign-title="Genotype–phenotype distinction"><span class="ilh-page"><a href="/w/index.php?title=%E5%9F%BA%E5%9B%A0%E5%9E%8B-%E8%A1%A8%E5%9E%8B%E7%9A%84%E5%8C%BA%E5%88%AB&action=edit&redlink=1" class="new" title="基因型-表型的区别（页面不存在）">基因型-表型的区别</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Genotype%E2%80%93phenotype_distinction" class="extiw" title="en:Genotype–phenotype distinction"><span lang="en" dir="auto">Genotype–phenotype distinction</span></a></span>）</span></span></li> <li><span class="ilh-all" data-orig-title="反应规范" data-lang-code="en" data-lang-name="英语" data-foreign-title="Reaction norm"><span class="ilh-page"><a href="/w/index.php?title=%E5%8F%8D%E5%BA%94%E8%A7%84%E8%8C%83&action=edit&redlink=1" class="new" title="反应规范（页面不存在）">反应规范</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Reaction_norm" class="extiw" title="en:Reaction norm"><span lang="en" dir="auto">Reaction norm</span></a></span>）</span></span></li> <li><span class="ilh-all" data-orig-title="基因-環境相互作用" data-lang-code="en" data-lang-name="英语" data-foreign-title="Gene–environment interaction"><span class="ilh-page"><a href="/w/index.php?title=%E5%9F%BA%E5%9B%A0-%E7%92%B0%E5%A2%83%E7%9B%B8%E4%BA%92%E4%BD%9C%E7%94%A8&action=edit&redlink=1" class="new" title="基因-環境相互作用（页面不存在）">基因-環境相互作用</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Gene%E2%80%93environment_interaction" class="extiw" title="en:Gene–environment interaction"><span lang="en" dir="auto">Gene–environment interaction</span></a></span>）</span></span></li> <li><span class="ilh-all" data-orig-title="基因和环境之间的关联" data-lang-code="en" data-lang-name="英语" data-foreign-title="Gene–environment correlation"><span class="ilh-page"><a href="/w/index.php?title=%E5%9F%BA%E5%9B%A0%E5%92%8C%E7%8E%AF%E5%A2%83%E4%B9%8B%E9%97%B4%E7%9A%84%E5%85%B3%E8%81%94&action=edit&redlink=1" class="new" title="基因和环境之间的关联（页面不存在）">基因和环境之间的关联</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Gene%E2%80%93environment_correlation" class="extiw" title="en:Gene–environment correlation"><span lang="en" dir="auto">Gene–environment correlation</span></a></span>）</span></span></li> <li><a href="/wiki/%E6%93%8D%E7%B8%B1%E5%AD%90" title="操縱子">操作子</a></li> <li><a href="/wiki/%E9%81%97%E4%BC%A0%E5%BA%A6" title="遗传度">遗传度</a></li> <li><a href="/wiki/%E5%AE%9A%E9%87%8F%E9%81%97%E4%BC%A0%E5%AD%A6" title="定量遗传学">定量遗传学</a></li> <li><span class="ilh-all" data-orig-title="异时发生" data-lang-code="en" data-lang-name="英语" data-foreign-title="Heterochrony"><span class="ilh-page"><a href="/w/index.php?title=%E5%BC%82%E6%97%B6%E5%8F%91%E7%94%9F&action=edit&redlink=1" class="new" title="异时发生（页面不存在）">异时发生</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Heterochrony" class="extiw" title="en:Heterochrony"><span lang="en" dir="auto">Heterochrony</span></a></span>）</span></span> <ul><li><a href="/wiki/%E5%B9%BC%E6%80%81%E5%BB%B6%E7%BB%AD" title="幼态延续">幼态延续</a></li></ul></li> <li><span class="ilh-all" data-orig-title="异位性" data-lang-code="en" data-lang-name="英语" data-foreign-title="Heterotopy"><span class="ilh-page"><a href="/w/index.php?title=%E5%BC%82%E4%BD%8D%E6%80%A7&action=edit&redlink=1" class="new" title="异位性（页面不存在）">异位性</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Heterotopy" class="extiw" title="en:Heterotopy"><span lang="en" dir="auto">Heterotopy</span></a></span>）</span></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><span class="ilh-all" data-orig-title="基因架构" data-lang-code="en" data-lang-name="英语" data-foreign-title="Genetic architecture"><span class="ilh-page"><a href="/w/index.php?title=%E5%9F%BA%E5%9B%A0%E6%9E%B6%E6%9E%84&action=edit&redlink=1" class="new" title="基因架构（页面不存在）">基因架构</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Genetic_architecture" class="extiw" title="en:Genetic architecture"><span lang="en" dir="auto">Genetic architecture</span></a></span>）</span></span></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0px"><div style="padding:0em 0.25em"> <ul><li><span class="ilh-all" data-orig-title="遗传渠化" data-lang-code="en" data-lang-name="英语" data-foreign-title="Canalisation (genetics)"><span class="ilh-page"><a href="/w/index.php?title=%E9%81%97%E4%BC%A0%E6%B8%A0%E5%8C%96&action=edit&redlink=1" class="new" title="遗传渠化（页面不存在）">遗传渠化</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Canalisation_(genetics)" class="extiw" title="en:Canalisation (genetics)"><span lang="en" dir="auto">Canalisation (genetics)</span></a></span>）</span></span></li> <li><span class="ilh-all" data-orig-title="遗传同化" data-lang-code="en" data-lang-name="英语" data-foreign-title="Genetic assimilation"><span class="ilh-page"><a href="/w/index.php?title=%E9%81%97%E4%BC%A0%E5%90%8C%E5%8C%96&action=edit&redlink=1" class="new" title="遗传同化（页面不存在）">遗传同化</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Genetic_assimilation" class="extiw" title="en:Genetic assimilation"><span lang="en" dir="auto">Genetic assimilation</span></a></span>）</span></span></li> <li><a href="/wiki/%E6%98%BE%E6%80%A7" title="显性">显性</a></li> <li><a href="/wiki/%E5%BC%82%E4%BD%8D%E6%98%BE%E6%80%A7" title="异位显性">异位显性</a></li> <li><span class="ilh-all" data-orig-title="适应度地形" data-lang-code="en" data-lang-name="英语" data-foreign-title="Fitness landscape"><span class="ilh-page"><a href="/w/index.php?title=%E9%80%82%E5%BA%94%E5%BA%A6%E5%9C%B0%E5%BD%A2&action=edit&redlink=1" class="new" title="适应度地形（页面不存在）">适应度地形</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Fitness_landscape" class="extiw" title="en:Fitness landscape"><span lang="en" dir="auto">Fitness landscape</span></a></span>）</span></span> / <span class="ilh-all" data-orig-title="进化景观" data-lang-code="en" data-lang-name="英语" data-foreign-title="Evolutionary landscape"><span class="ilh-page"><a href="/w/index.php?title=%E8%BF%9B%E5%8C%96%E6%99%AF%E8%A7%82&action=edit&redlink=1" class="new" title="进化景观（页面不存在）">进化景观</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Evolutionary_landscape" class="extiw" title="en:Evolutionary landscape"><span lang="en" dir="auto">Evolutionary landscape</span></a></span>）</span></span></li> <li><a href="/wiki/%E5%9F%BA%E5%9B%A0%E5%A4%9A%E6%95%88%E6%80%A7" title="基因多效性">基因多效性</a></li> <li><a href="/wiki/%E8%A1%A8%E5%9E%8B%E5%8F%AF%E5%A1%91%E6%80%A7" title="表型可塑性">表型可塑性</a></li> <li><a href="/w/index.php?title=%E5%A4%9A%E5%9F%BA%E5%9B%A0%E9%81%97%E4%BC%A0&action=edit&redlink=1" class="new" title="多基因遗传（页面不存在）">多基因遗传</a></li> <li><span class="ilh-all" data-orig-title="超亲遗传分离" data-lang-code="en" data-lang-name="英语" data-foreign-title="Transgressive segregation"><span class="ilh-page"><a href="/w/index.php?title=%E8%B6%85%E4%BA%B2%E9%81%97%E4%BC%A0%E5%88%86%E7%A6%BB&action=edit&redlink=1" class="new" title="超亲遗传分离（页面不存在）">超亲遗传分离</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Transgressive_segregation" class="extiw" title="en:Transgressive segregation"><span lang="en" dir="auto">Transgressive segregation</span></a></span>）</span></span></li> <li><span class="ilh-all" data-orig-title="序列空间" data-lang-code="en" data-lang-name="英语" data-foreign-title="Sequence space (evolution)"><span class="ilh-page"><a href="/w/index.php?title=%E5%BA%8F%E5%88%97%E7%A9%BA%E9%97%B4&action=edit&redlink=1" class="new" title="序列空间（页面不存在）">序列空间</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Sequence_space_(evolution)" class="extiw" title="en:Sequence space (evolution)"><span lang="en" dir="auto">Sequence space (evolution)</span></a></span>）</span></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">非遗传因素的影响</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0px"><div style="padding:0em 0.25em"> <ul><li><a href="/wiki/%E8%A1%A8%E8%A7%80%E9%81%BA%E5%82%B3%E5%AD%B8" title="表觀遺傳學">表觀遺傳學</a> <ul><li><a href="/wiki/%E8%A1%A8%E8%A7%80%E5%9F%BA%E5%9B%A0%E7%B5%84" title="表觀基因組">表觀基因組</a></li></ul></li> <li><a href="/wiki/%E6%AF%8D%E4%BD%93%E6%95%88%E5%BA%94" title="母体效应">母体效应</a></li> <li><a href="/wiki/%E5%9F%BA%E5%9B%A0%E7%B5%84%E9%8A%98%E5%8D%B0" title="基因組銘印">基因組銘印</a></li> <li><a href="/wiki/%E5%8F%8C%E7%BB%A7%E6%89%BF%E7%90%86%E8%AE%BA" title="双继承理论">双继承理论</a></li> <li><a href="/wiki/%E9%9D%9E%E9%81%97%E4%BC%A0%E5%A4%9A%E5%9E%8B%E6%80%A7" title="非遗传多型性">非遗传多型性</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">发育架构</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0px"><div style="padding:0em 0.25em"> <ul><li><a href="/wiki/%E5%8F%91%E8%82%B2%E7%94%9F%E7%89%A9%E5%AD%A6" title="发育生物学">发育生物学</a></li> <li><a href="/wiki/%E5%BD%A2%E6%80%81%E5%8F%91%E7%94%9F" title="形态发生">形态发生</a> <ul><li><a href="/wiki/%E7%9C%BC%E7%8B%80%E6%96%91%E9%BB%9E" title="眼狀斑點">眼狀斑點</a></li> <li><span class="ilh-all" data-orig-title="模式形成" data-lang-code="en" data-lang-name="英语" data-foreign-title="Pattern formation"><span class="ilh-page"><a href="/w/index.php?title=%E6%A8%A1%E5%BC%8F%E5%BD%A2%E6%88%90&action=edit&redlink=1" class="new" title="模式形成（页面不存在）">模式形成</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Pattern_formation" class="extiw" title="en:Pattern formation"><span lang="en" dir="auto">Pattern formation</span></a></span>）</span></span></li></ul></li> <li><span class="ilh-all" data-orig-title="分節" data-lang-code="en" data-lang-name="英语" data-foreign-title="Segmentation (biology)"><span class="ilh-page"><a href="/w/index.php?title=%E5%88%86%E7%AF%80&action=edit&redlink=1" class="new" title="分節（页面不存在）">分節</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Segmentation_(biology)" class="extiw" title="en:Segmentation (biology)"><span lang="en" dir="auto">Segmentation (biology)</span></a></span>）</span></span></li> <li><span class="ilh-all" data-orig-title="分节" data-lang-code="en" data-lang-name="英语" data-foreign-title="Metamerism (biology)"><span class="ilh-page"><a href="/w/index.php?title=%E5%88%86%E8%8A%82&action=edit&redlink=1" class="new" title="分节（页面不存在）">分节</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Metamerism_(biology)" class="extiw" title="en:Metamerism (biology)"><span lang="en" dir="auto">Metamerism (biology)</span></a></span>）</span></span></li> <li><span class="ilh-all" data-orig-title="生物模块化" data-lang-code="en" data-lang-name="英语" data-foreign-title="Modularity (biology)"><span class="ilh-page"><a href="/w/index.php?title=%E7%94%9F%E7%89%A9%E6%A8%A1%E5%9D%97%E5%8C%96&action=edit&redlink=1" class="new" title="生物模块化（页面不存在）">生物模块化</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Modularity_(biology)" class="extiw" title="en:Modularity (biology)"><span lang="en" dir="auto">Modularity (biology)</span></a></span>）</span></span></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">遗传系统的进化</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0px"><div style="padding:0em 0.25em"> <ul><li><span class="ilh-all" data-orig-title="演化能力" data-lang-code="en" data-lang-name="英语" data-foreign-title="Evolvability"><span class="ilh-page"><a href="/w/index.php?title=%E6%BC%94%E5%8C%96%E8%83%BD%E5%8A%9B&action=edit&redlink=1" class="new" title="演化能力（页面不存在）">演化能力</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Evolvability" class="extiw" title="en:Evolvability"><span lang="en" dir="auto">Evolvability</span></a></span>）</span></span></li> <li><span class="ilh-all" data-orig-title="进化鲁棒性" data-lang-code="en" data-lang-name="英语" data-foreign-title="Robustness (evolution)"><span class="ilh-page"><a href="/w/index.php?title=%E8%BF%9B%E5%8C%96%E9%B2%81%E6%A3%92%E6%80%A7&action=edit&redlink=1" class="new" title="进化鲁棒性（页面不存在）">进化鲁棒性</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Robustness_(evolution)" class="extiw" title="en:Robustness (evolution)"><span lang="en" dir="auto">Robustness (evolution)</span></a></span>）</span></span></li> <li><span class="ilh-all" data-orig-title="中性网络" data-lang-code="en" data-lang-name="英语" data-foreign-title="Neutral network (evolution)"><span class="ilh-page"><a href="/w/index.php?title=%E4%B8%AD%E6%80%A7%E7%BD%91%E7%BB%9C&action=edit&redlink=1" class="new" title="中性网络（页面不存在）">中性网络</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Neutral_network_(evolution)" class="extiw" title="en:Neutral network (evolution)"><span lang="en" dir="auto">Neutral network (evolution)</span></a></span>）</span></span></li> <li><a href="/wiki/%E6%BC%94%E5%8C%96%E7%99%BC%E8%82%B2%E7%94%9F%E7%89%A9%E5%AD%B8" title="演化發育生物學">有性繁殖的进化</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">对发育的控制</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0px"><div style="padding:0em 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">系统</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0px"><div style="padding:0em 0.25em"> <ul><li><a href="/wiki/%E5%9F%BA%E5%9B%A0%E8%A1%A8%E8%BE%BE%E8%B0%83%E6%8E%A7" title="基因表达调控">基因表达调控</a></li> <li><span class="ilh-all" data-orig-title="基因調控網路" data-lang-code="en" data-lang-name="英语" data-foreign-title="Gene regulatory network"><span class="ilh-page"><a href="/w/index.php?title=%E5%9F%BA%E5%9B%A0%E8%AA%BF%E6%8E%A7%E7%B6%B2%E8%B7%AF&action=edit&redlink=1" class="new" title="基因調控網路（页面不存在）">基因調控網路</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Gene_regulatory_network" class="extiw" title="en:Gene regulatory network"><span lang="en" dir="auto">Gene regulatory network</span></a></span>）</span></span></li> <li><a href="/wiki/%E5%B7%A5%E5%85%B7%E5%8C%85%E5%9F%BA%E5%9B%A0" title="工具包基因">工具包基因</a></li> <li><a href="/wiki/%E6%BC%94%E5%8C%96%E7%99%BC%E8%82%B2%E7%94%9F%E7%89%A9%E5%AD%B8" title="演化發育生物學">演化發育生物學</a></li> <li><a href="/wiki/%E5%90%8C%E6%BA%90%E6%A1%86" title="同源框">同源框</a></li> <li><a href="/wiki/%E5%88%BA%E7%8C%AC%E4%BF%A1%E5%8F%B7%E9%80%9A%E8%B7%AF" title="刺猬信号通路">刺猬信号通路</a></li> <li><a href="/wiki/Notch%E4%BF%A1%E5%8F%B7%E9%80%9A%E8%B7%AF" title="Notch信号通路">Notch信号通路</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">组件</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0px"><div style="padding:0em 0.25em"> <ul><li><a href="/wiki/%E5%90%8C%E6%BA%90%E5%9F%BA%E5%9B%A0" title="同源基因">同源基因</a></li> <li><a href="/wiki/%E5%90%8C%E6%BA%90%E5%BC%82%E5%BD%A2%E5%9F%BA%E5%9B%A0" title="同源异形基因">同源异形基因</a></li> <li><span class="ilh-all" data-orig-title="Pax基因" data-lang-code="en" data-lang-name="英语" data-foreign-title="Pax genes"><span class="ilh-page"><a href="/w/index.php?title=Pax%E5%9F%BA%E5%9B%A0&action=edit&redlink=1" class="new" title="Pax基因（页面不存在）">Pax基因</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Pax_genes" class="extiw" title="en:Pax genes"><span lang="en" dir="auto">Pax genes</span></a></span>）</span></span> <ul><li><span class="ilh-all" data-orig-title="PAX6" data-lang-code="en" data-lang-name="英语" data-foreign-title="PAX6"><span class="ilh-page"><a href="/w/index.php?title=PAX6&action=edit&redlink=1" class="new" title="PAX6（页面不存在）">PAX6</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/PAX6" class="extiw" title="en:PAX6"><span lang="en" dir="auto">PAX6</span></a></span>）</span></span></li></ul></li> <li><a href="/w/index.php?title=DLX%E5%9F%BA%E5%9B%A0%E5%AE%B6%E6%97%8F&action=edit&redlink=1" class="new" title="DLX基因家族（页面不存在）">DLX基因家族</a></li> <li><a href="/w/index.php?title=Engrailed(gene)&action=edit&redlink=1" class="new" title="Engrailed(gene)（页面不存在）">Engrailed</a></li> <li><a class="mw-selflink selflink">顺式作用元件</a></li> <li><a href="/wiki/%E9%85%8D%E9%AB%94_(%E7%94%9F%E7%89%A9%E5%8C%96%E5%AD%B8)" title="配體 (生物化學)">配体</a></li> <li><span class="ilh-all" data-orig-title="形態發生" data-lang-code="en" data-lang-name="英语" data-foreign-title="Morphogen"><span class="ilh-page"><a href="/wiki/%E5%BD%A2%E6%80%81%E5%8F%91%E7%94%9F" title="形态发生">形态素</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Morphogen" class="extiw" title="en:Morphogen"><span lang="en" dir="auto">Morphogen</span></a></span>）</span></span></li> <li><a href="/wiki/%E5%8F%97%E4%BD%93_(%E7%94%9F%E7%89%A9%E5%8C%96%E5%AD%A6)" title="受体 (生物化学)">细胞表面受体</a></li> <li><a href="/wiki/%E8%BD%AC%E5%BD%95%E5%9B%A0%E5%AD%90" title="转录因子">转录因子</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">有影响力的人物</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0px"><div style="padding:0em 0.25em"> <ul><li><a href="/wiki/%E5%BA%B7%E6%8B%89%E5%BE%B7%C2%B7%E5%93%88%E5%B0%94%C2%B7%E6%B2%83%E4%B8%81%E9%A1%BF" title="康拉德·哈尔·沃丁顿">康拉德·哈尔·沃丁顿</a></li> <li><a href="/wiki/%E7%90%86%E6%9F%A5%E5%BE%B7%C2%B7%E9%99%86%E6%96%87%E9%A1%BF" title="理查德·陆文顿">理查德·陆文顿</a></li> <li><a href="/wiki/%E6%96%B9%E6%96%AF%E8%8F%AF%C2%B7%E8%B3%88%E5%85%8B%E6%9F%8F" title="方斯華·賈克柏">方斯華·賈克柏</a> + <a href="/wiki/%E8%B3%88%E5%85%8B%C2%B7%E8%8E%AB%E8%AB%BE" title="賈克·莫諾">賈克·莫諾</a> <ul><li><a href="/wiki/%E4%B9%B3%E7%B3%96%E6%93%8D%E7%B8%B1%E5%AD%90" title="乳糖操縱子">乳糖操縱子</a></li></ul></li> <li><a href="/wiki/%E8%89%BE%E7%91%9E%E5%85%8B%C2%B7%E5%A8%81%E6%96%AF%E5%96%AC%E6%96%AF" title="艾瑞克·威斯喬斯">艾瑞克·威斯喬斯</a></li> <li><a href="/wiki/%E5%85%8B%E9%87%8C%E6%96%AF%E6%B1%80%C2%B7%E7%B4%90%E6%96%AF%E6%9E%97-%E6%B2%83%E7%88%BE%E5%93%88%E5%BE%B7" title="克里斯汀·紐斯林-沃爾哈德">克里斯汀·紐斯林-沃爾哈德</a></li> <li><a href="/w/index.php?title=William_McGinnis&action=edit&redlink=1" class="new" title="William McGinnis（页面不存在）">William McGinnis</a></li> <li><a href="/w/index.php?title=Michael_Levine_(biologist)&action=edit&redlink=1" class="new" title="Michael Levine (biologist)（页面不存在）">Mike Levine</a></li> <li><a href="/wiki/%E8%82%96%E6%81%A9%C2%B7B%C2%B7%E5%8D%A1%E7%BD%97%E5%B0%94" title="肖恩·B·卡罗尔">肖恩·B·卡罗尔</a> <ul><li>《<span class="ilh-all" data-orig-title="无尽之形最美" data-lang-code="en" data-lang-name="英语" data-foreign-title="Endless Forms Most Beautiful (book)"><span class="ilh-page"><a href="/w/index.php?title=%E6%97%A0%E5%B0%BD%E4%B9%8B%E5%BD%A2%E6%9C%80%E7%BE%8E&action=edit&redlink=1" class="new" title="无尽之形最美（页面不存在）">无尽之形最美——动物建造和演化的奥秘</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Endless_Forms_Most_Beautiful_(book)" class="extiw" title="en:Endless Forms Most Beautiful (book)"><span lang="en" dir="auto">Endless Forms Most Beautiful (book)</span></a></span>）</span></span>》</li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">争议</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0px"><div style="padding:0em 0.25em"> <ul><li><a href="/wiki/%E5%85%88%E5%A4%A9%E4%B8%8E%E5%90%8E%E5%A4%A9" title="先天与后天">先天与后天</a></li> <li><span class="ilh-all" data-orig-title="形态形成场" data-lang-code="en" data-lang-name="英语" data-foreign-title="Morphogenetic field"><span class="ilh-page"><a href="/w/index.php?title=%E5%BD%A2%E6%80%81%E5%BD%A2%E6%88%90%E5%9C%BA&action=edit&redlink=1" class="new" title="形态形成场（页面不存在）">形态形成场</a></span><span class="noprint ilh-comment">（<span class="ilh-lang">英语</span><span class="ilh-colon">：</span><span class="ilh-link"><a href="https://en.wikipedia.org/wiki/Morphogenetic_field" class="extiw" title="en:Morphogenetic field"><span lang="en" dir="auto">Morphogenetic field</span></a></span>）</span></span></li></ul> </div></td></tr></tbody></table></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">检索自“<a dir="ltr" href="https://zh.wikipedia.org/w/index.php?title=顺式作用元件&oldid=84161769">https://zh.wikipedia.org/w/index.php?title=顺式作用元件&oldid=84161769</a>”</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div 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