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Gyrus - Wikipedia
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Available in 22 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-22" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">22 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%AA%D9%84%D9%81%D9%8A%D9%81" title="تلفيف – Arabic" lang="ar" hreflang="ar" data-title="تلفيف" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-ast mw-list-item"><a href="https://ast.wikipedia.org/wiki/Xiros_cerebrales" title="Xiros cerebrales – Asturian" lang="ast" hreflang="ast" data-title="Xiros cerebrales" data-language-autonym="Asturianu" data-language-local-name="Asturian" class="interlanguage-link-target"><span>Asturianu</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Circumvoluci%C3%B3" title="Circumvolució – Catalan" lang="ca" hreflang="ca" data-title="Circumvolució" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Gyrus" title="Gyrus – German" lang="de" hreflang="de" data-title="Gyrus" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%95%CE%B3%CE%BA%CE%B5%CF%86%CE%B1%CE%BB%CE%B9%CE%BA%CE%AE_%CE%AD%CE%BB%CE%B9%CE%BA%CE%B1" title="Εγκεφαλική έλικα – Greek" lang="el" hreflang="el" data-title="Εγκεφαλική έλικα" data-language-autonym="Ελληνικά" data-language-local-name="Greek" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Giros_cerebrales" title="Giros cerebrales – Spanish" lang="es" hreflang="es" data-title="Giros cerebrales" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Cerba_giro" title="Cerba giro – Esperanto" lang="eo" hreflang="eo" data-title="Cerba giro" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Zirkunboluzio_zerebral" title="Zirkunboluzio zerebral – Basque" lang="eu" hreflang="eu" data-title="Zirkunboluzio zerebral" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%B4%DA%A9%D9%86%D8%AC_%D9%85%D8%BA%D8%B2" title="شکنج مغز – Persian" lang="fa" hreflang="fa" data-title="شکنج مغز" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Circonvolution_c%C3%A9r%C3%A9brale" title="Circonvolution cérébrale – French" lang="fr" hreflang="fr" data-title="Circonvolution cérébrale" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Circunvoluci%C3%B3n" title="Circunvolución – Galician" lang="gl" hreflang="gl" data-title="Circunvolución" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Gyrus" title="Gyrus – Indonesian" lang="id" hreflang="id" data-title="Gyrus" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Circonvoluzione" title="Circonvoluzione – Italian" lang="it" hreflang="it" data-title="Circonvoluzione" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Hersenwinding" title="Hersenwinding – Dutch" lang="nl" hreflang="nl" data-title="Hersenwinding" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E8%84%B3%E5%9B%9E" title="脳回 – Japanese" lang="ja" hreflang="ja" data-title="脳回" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Giro" title="Giro – Portuguese" lang="pt" hreflang="pt" data-title="Giro" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%98%D0%B7%D0%B2%D0%B8%D0%BB%D0%B8%D0%BD%D0%B0_(%D0%BC%D0%BE%D0%B7%D0%B3)" title="Извилина (мозг) – Russian" lang="ru" hreflang="ru" data-title="Извилина (мозг)" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Hj%C3%A4rnbark#Gyrus_och_sulcus" title="Hjärnbark – Swedish" lang="sv" hreflang="sv" data-title="Hjärnbark" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Girus" title="Girus – Turkish" lang="tr" hreflang="tr" data-title="Girus" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%97%D0%B2%D0%B8%D0%B2%D0%B8%D0%BD%D0%B0" title="Звивина – Ukrainian" lang="uk" hreflang="uk" data-title="Звивина" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E8%85%A6%E5%9B%9E" title="腦回 – Cantonese" lang="yue" hreflang="yue" data-title="腦回" data-language-autonym="粵語" data-language-local-name="Cantonese" class="interlanguage-link-target"><span>粵語</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E8%84%91%E5%9B%9E" title="脑回 – Chinese" lang="zh" hreflang="zh" data-title="脑回" data-language-autonym="中文" data-language-local-name="Chinese" 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title="Gyri">Gyri</a>)</span></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Ridge on the cerebral cortex of the brain</div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">For the video game, see <a href="/wiki/Gyruss" title="Gyruss">Gyruss</a>.</div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">"Gyral" redirects here. For the album, see <a href="/wiki/Gyral_(album)" title="Gyral (album)">Gyral (album)</a>.</div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Not to be confused with <a href="/wiki/Girus" class="mw-redirect" title="Girus">Girus</a>.</div> <style data-mw-deduplicate="TemplateStyles:r1257001546">.mw-parser-output .infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox-full-data:not(.notheme)>div:not(.notheme)[style]{background:#1f1f23!important;color:#f8f9fa}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox-full-data:not(.notheme) div:not(.notheme){background:#1f1f23!important;color:#f8f9fa}}@media(min-width:640px){body.skin--responsive .mw-parser-output .infobox-table{display:table!important}body.skin--responsive .mw-parser-output .infobox-table>caption{display:table-caption!important}body.skin--responsive .mw-parser-output .infobox-table>tbody{display:table-row-group}body.skin--responsive .mw-parser-output .infobox-table tr{display:table-row!important}body.skin--responsive .mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox"><tbody><tr><th colspan="2" class="infobox-above" style="background-color:dimgray; color: white">Gyrus</th></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Gyrus_sulcus-en.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/e/e6/Gyrus_sulcus-en.svg/250px-Gyrus_sulcus-en.svg.png" decoding="async" width="250" height="201" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/e6/Gyrus_sulcus-en.svg/375px-Gyrus_sulcus-en.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/e6/Gyrus_sulcus-en.svg/500px-Gyrus_sulcus-en.svg.png 2x" data-file-width="789" data-file-height="633" /></a></span><div class="infobox-caption">Gyrus and <a href="/wiki/Sulcus_(neuroanatomy)" title="Sulcus (neuroanatomy)">sulcus</a></div></td></tr><tr><th colspan="2" class="infobox-header" style="background-color: #efefef">Identifiers</th></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/Terminologia_Anatomica" title="Terminologia Anatomica">TA98</a></th><td class="infobox-data"><a rel="nofollow" class="external text" href="https://ifaa.unifr.ch/Public/EntryPage/TA98%20Tree/Entity%20TA98%20EN/14.1.09.004%20Entity%20TA98%20EN.htm">A14.1.09.004</a></td></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/Terminologia_Anatomica" title="Terminologia Anatomica">TA2</a></th><td class="infobox-data"><a rel="nofollow" class="external text" href="https://ta2viewer.openanatomy.org/?id=5432">5432</a></td></tr><tr><th scope="row" class="infobox-label" style="padding-right:0.25em"><a href="/wiki/Foundational_Model_of_Anatomy" title="Foundational Model of Anatomy">FMA</a></th><td class="infobox-data"><a rel="nofollow" class="external text" href="https://bioportal.bioontology.org/ontologies/FMA/?p=classes&conceptid=http%3A%2F%2Fpurl.org%2Fsig%2Font%2Ffma%2Ffma83874">83874</a></td></tr><tr><td colspan="2" class="infobox-below"><a href="/wiki/Anatomical_terminology" title="Anatomical terminology">Anatomical terminology</a><div style="text-align: right;"><small class="noprint">[<a href="https://www.wikidata.org/wiki/Q1758383" class="extiw" title="d:Q1758383">edit on Wikidata</a>]</small></div></td></tr></tbody></table> <figure typeof="mw:File/Thumb"><a href="/wiki/File:Gray726.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/35/Gray726.png/300px-Gray726.png" decoding="async" width="300" height="174" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/35/Gray726.png/450px-Gray726.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/35/Gray726.png/600px-Gray726.png 2x" data-file-width="700" data-file-height="405" /></a><figcaption><a href="/wiki/Gray%27s_Anatomy" title="Gray's Anatomy">Gray's</a> FIG. 726 – Lateral surface of left <a href="/wiki/Cerebral_hemisphere" title="Cerebral hemisphere">cerebral hemisphere</a>, viewed from the side</figcaption></figure> <figure typeof="mw:File/Thumb"><a href="/wiki/File:Gray727.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/fe/Gray727.svg/300px-Gray727.svg.png" decoding="async" width="300" height="175" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/fe/Gray727.svg/450px-Gray727.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/fe/Gray727.svg/600px-Gray727.svg.png 2x" data-file-width="1025" data-file-height="598" /></a><figcaption><a href="/wiki/Gray%27s_Anatomy" title="Gray's Anatomy">Gray's</a> Fig. 727 – Medial surface of left cerebral hemisphere</figcaption></figure> <p>In <a href="/wiki/Neuroanatomy" title="Neuroanatomy">neuroanatomy</a>, a <b>gyrus</b> (<abbr title="plural form">pl.</abbr>: <b>gyri</b>) is a ridge on the <a href="/wiki/Cerebral_cortex" title="Cerebral cortex">cerebral cortex</a>. It is generally surrounded by one or more <a href="/wiki/Sulcus_(neuroanatomy)" title="Sulcus (neuroanatomy)">sulci</a> (depressions or furrows; <abbr title="singular form">sg.</abbr>: <b>sulcus</b>).<sup id="cite_ref-DengJiang2013_1-0" class="reference"><a href="#cite_note-DengJiang2013-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> Gyri and sulci create the folded appearance of the <a href="/wiki/Brain" title="Brain">brain</a> in <a href="/wiki/Human" title="Human">humans</a> and other <a href="/wiki/Mammal" title="Mammal">mammals</a>. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Structure">Structure</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Gyrus&action=edit&section=1" title="Edit section: Structure"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The gyri are part of a system of folds and ridges that create a larger surface area for the human brain and other mammalian brains.<sup id="cite_ref-marieb_2-0" class="reference"><a href="#cite_note-marieb-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> Because the brain is confined to the skull, brain size is limited. Ridges and depressions create folds allowing a larger cortical surface area, and greater cognitive function, to exist in the confines of a smaller <a href="/wiki/Cranium" class="mw-redirect" title="Cranium">cranium</a>.<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Development">Development</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Gyrus&action=edit&section=2" title="Edit section: Development"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Gyrification" title="Gyrification">Gyrification</a></div> <p>The human brain undergoes <a href="/wiki/Gyrification" title="Gyrification">gyrification</a> during fetal and neonatal development. In <a href="/wiki/Embryo" title="Embryo">embryonic</a> development, all <a href="/wiki/Mammal" title="Mammal">mammalian</a> brains begin as smooth structures derived from the <a href="/wiki/Neural_tube" title="Neural tube">neural tube</a>. A <a href="/wiki/Cerebral_cortex" title="Cerebral cortex">cerebral cortex</a> without surface convolutions is <a href="/wiki/Lissencephaly" title="Lissencephaly">lissencephalic</a>, meaning 'smooth-brained'.<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> As development continues, gyri and <a href="/wiki/Sulcus_(neuroanatomy)" title="Sulcus (neuroanatomy)">sulci</a> begin to take shape on the fetal brain, with deepening indentations and ridges developing on the surface of the cortex.<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Clinical_significance">Clinical significance</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Gyrus&action=edit&section=3" title="Edit section: Clinical significance"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Changes in the structure of gyri in the cerebral cortex are associated with various diseases and disorders. <a href="/wiki/Pachygyria" title="Pachygyria">Pachygyria</a>, <a href="/wiki/Lissencephaly" title="Lissencephaly">lissencephaly</a>, and <a href="/wiki/Polymicrogyria" title="Polymicrogyria">polymicrogyria</a> are all the results of abnormal <a href="/wiki/Cell_migration" title="Cell migration">cell migration</a> associated with a disorganized cellular architecture, failure to form six layers of cortical neurons (a four-layer cortex is common), and functional problems.<sup id="cite_ref-BarkovichGuerrini2012_6-0" class="reference"><a href="#cite_note-BarkovichGuerrini2012-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> The abnormal formation is commonly associated with <a href="/wiki/Epilepsy" title="Epilepsy">epilepsy</a> and <a href="/wiki/Intellectual_disability" title="Intellectual disability">mental dysfunctions</a>.<sup id="cite_ref-PangAtefy2008_7-0" class="reference"><a href="#cite_note-PangAtefy2008-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p><p><a href="/wiki/Pachygyria" title="Pachygyria">Pachygyria</a> (meaning "thick" or "fat" gyri) is a congenital malformation of the cerebral hemisphere, resulting in unusually thick gyri in the <a href="/wiki/Cerebral_cortex" title="Cerebral cortex">cerebral cortex</a>.<sup id="cite_ref-guerrini5_8-0" class="reference"><a href="#cite_note-guerrini5-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> Pachygyria is used to describe brain characteristics in association with several <a href="/wiki/Neuronal_migration_disorders" class="mw-redirect" title="Neuronal migration disorders">neuronal migration disorders</a>; most commonly relating to lissencephaly. </p><p><a href="/wiki/Lissencephaly" title="Lissencephaly">Lissencephaly</a> (<i>smooth brain</i>) is a rare congenital brain malformation caused by <a href="/wiki/Neuronal_migration_disorder" title="Neuronal migration disorder">defective neuronal migration</a> during the 12th to 24th weeks of fetal <a href="/wiki/Gestation" title="Gestation">gestation</a> resulting in a lack of development of gyri and sulci.<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> </p><p><a href="/wiki/Polymicrogyria" title="Polymicrogyria">Polymicrogyria</a> (meaning "many small gyri") is a developmental malformation of the human brain characterized by excessive folding of the <a href="/wiki/Gyri" class="mw-redirect" title="Gyri">gyri</a> and a thickening of the <a href="/wiki/Cerebral_cortex" title="Cerebral cortex">cerebral cortex</a>.<sup id="cite_ref-chang_10-0" class="reference"><a href="#cite_note-chang-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> It may be generalized, affecting the whole surface of the cerebral cortex or may be focal, affecting only parts of the surface. Polymicrogyria may be caused by mutations within several genes, including ion channels.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Notable_gyri">Notable gyri</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Gyrus&action=edit&section=4" title="Edit section: Notable gyri"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="/wiki/Superior_frontal_gyrus" title="Superior frontal gyrus">Superior frontal gyrus</a>, <i>lat</i>. gyrus frontalis superior</li> <li><a href="/wiki/Middle_frontal_gyrus" title="Middle frontal gyrus">Middle frontal gyrus</a>, <i>lat</i>. gyrus frontalis medius</li> <li><a href="/wiki/Inferior_frontal_gyrus" title="Inferior frontal gyrus">Inferior frontal gyrus</a>, <i>lat</i>. gyrus frontalis inferior with 3 parts: pars opercularis (<a href="/wiki/Brodmann_area_44" title="Brodmann area 44">Brodmann area 44</a>) pars triangularis (<a href="/wiki/Brodmann_area_45" title="Brodmann area 45">Brodmann area 45</a>), and pars orbitalis (<a href="/wiki/Orbital_part_of_inferior_frontal_gyrus" title="Orbital part of inferior frontal gyrus">orbital part of inferior frontal gyrus</a>)</li> <li><a href="/wiki/Superior_temporal_gyrus" title="Superior temporal gyrus">Superior temporal gyrus</a>, <i>lat</i>. gyrus temporalis superior</li> <li><a href="/wiki/Middle_temporal_gyrus" title="Middle temporal gyrus">Middle temporal gyrus</a>, <i>lat</i>. gyrus temporalis medius</li> <li><a href="/wiki/Inferior_temporal_gyrus" title="Inferior temporal gyrus">Inferior temporal gyrus</a>, <i>lat</i>. gyrus temporalis inferior</li> <li><a href="/wiki/Fusiform_gyrus" title="Fusiform gyrus">Fusiform gyrus</a>, <i>lat</i>. gyrus occipitotemporalis lateralis</li> <li><a href="/wiki/Parahippocampal_gyrus" title="Parahippocampal gyrus">Parahippocampal gyrus</a>, <i>lat</i>. gyrus parahippocampalis</li> <li><a href="/wiki/Transverse_temporal_gyrus" title="Transverse temporal gyrus">Transverse temporal gyrus</a></li> <li><a href="/wiki/Lingual_gyrus" title="Lingual gyrus">Lingual gyrus</a> <i>lat</i>. gyrus lingualis</li> <li><a href="/wiki/Precentral_gyrus" title="Precentral gyrus">Precentral gyrus</a>, <i>lat</i>. gyrus praecentralis</li> <li><a href="/wiki/Postcentral_gyrus" title="Postcentral gyrus">Postcentral gyrus</a>, <i>lat</i>. gyrus postcentralis</li> <li><a href="/wiki/Supramarginal_gyrus" title="Supramarginal gyrus">Supramarginal gyrus</a>, <i>lat</i>. gyrus supramarginalis</li> <li><a href="/wiki/Angular_gyrus" title="Angular gyrus">Angular gyrus</a>, <i>lat</i>. gyrus angularis</li> <li><a href="/wiki/Cingulate_gyrus" class="mw-redirect" title="Cingulate gyrus">Cingulate gyrus</a> <i>lat</i>. gyrus cinguli</li> <li><a href="/wiki/Fornicate_gyrus" class="mw-redirect" title="Fornicate gyrus">Fornicate gyrus</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Gyrus&action=edit&section=5" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="/wiki/Gyrification" title="Gyrification">Gyrification</a></li> <li><a href="/wiki/Sulcus_(neuroanatomy)" title="Sulcus (neuroanatomy)">Sulcus</a></li> <li><a href="/wiki/Ulegyria" title="Ulegyria">Ulegyria</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Gyrus&action=edit&section=6" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width" style="column-width: 30em;"> <ol class="references"> <li id="cite_note-DengJiang2013-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-DengJiang2013_1-0">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFDengJiangZhuZhang2013" class="citation journal cs1">Deng, Fan; Jiang, Xi; Zhu, Dajiang; Zhang, Tuo; Li, Kaiming; Guo, Lei; Liu, Tianming (2013). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3909019">"A functional model of cortical gyri and sulci"</a>. <i>Brain Structure and Function</i>. <b>219</b> (4): 1473–1491. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1007%2Fs00429-013-0581-z">10.1007/s00429-013-0581-z</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/1863-2653">1863-2653</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3909019">3909019</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/23689502">23689502</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Brain+Structure+and+Function&rft.atitle=A+functional+model+of+cortical+gyri+and+sulci&rft.volume=219&rft.issue=4&rft.pages=1473-1491&rft.date=2013&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3909019%23id-name%3DPMC&rft.issn=1863-2653&rft_id=info%3Apmid%2F23689502&rft_id=info%3Adoi%2F10.1007%2Fs00429-013-0581-z&rft.aulast=Deng&rft.aufirst=Fan&rft.au=Jiang%2C+Xi&rft.au=Zhu%2C+Dajiang&rft.au=Zhang%2C+Tuo&rft.au=Li%2C+Kaiming&rft.au=Guo%2C+Lei&rft.au=Liu%2C+Tianming&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3909019&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGyrus" class="Z3988"></span></span> </li> <li id="cite_note-marieb-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-marieb_2-0">^</a></b></span> <span class="reference-text"> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMariebHoehn2012" class="citation book cs1">Marieb, Elaine N.; Hoehn, Katja (2012). <i>Human Anatomy & Physiology</i> (9th ed.). Pearson. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0321852120" title="Special:BookSources/978-0321852120"><bdi>978-0321852120</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Human+Anatomy+%26+Physiology&rft.edition=9th&rft.pub=Pearson&rft.date=2012&rft.isbn=978-0321852120&rft.aulast=Marieb&rft.aufirst=Elaine+N.&rft.au=Hoehn%2C+Katja&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGyrus" class="Z3988"></span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-3">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCusack2005" class="citation journal cs1">Cusack, Rhodri (April 2005). "The Intraparietal Sulcus and Perceptual Organization". <i>Journal of Cognitive Neuroscience</i>. <b>17</b> (4): 641–651. <a href="/wiki/CiteSeerX_(identifier)" class="mw-redirect" title="CiteSeerX (identifier)">CiteSeerX</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://citeseerx.ist.psu.edu/viewdoc/summary?doi=10.1.1.452.462">10.1.1.452.462</a></span>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1162%2F0898929053467541">10.1162/0898929053467541</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/15829084">15829084</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:25453338">25453338</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Journal+of+Cognitive+Neuroscience&rft.atitle=The+Intraparietal+Sulcus+and+Perceptual+Organization&rft.volume=17&rft.issue=4&rft.pages=641-651&rft.date=2005-04&rft_id=https%3A%2F%2Fciteseerx.ist.psu.edu%2Fviewdoc%2Fsummary%3Fdoi%3D10.1.1.452.462%23id-name%3DCiteSeerX&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A25453338%23id-name%3DS2CID&rft_id=info%3Apmid%2F15829084&rft_id=info%3Adoi%2F10.1162%2F0898929053467541&rft.aulast=Cusack&rft.aufirst=Rhodri&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGyrus" class="Z3988"></span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFArmstrongSchleicherOmranCurtis1991" class="citation journal cs1">Armstrong, E; Schleicher, A; Omran, H; Curtis, M; Zilles, K (1991). "The ontogeny of human gyrification". <i>Cerebral Cortex</i>. <b>5</b> (1): 56–63. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fcercor%2F5.1.56">10.1093/cercor/5.1.56</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/7719130">7719130</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Cerebral+Cortex&rft.atitle=The+ontogeny+of+human+gyrification.&rft.volume=5&rft.issue=1&rft.pages=56-63&rft.date=1991&rft_id=info%3Adoi%2F10.1093%2Fcercor%2F5.1.56&rft_id=info%3Apmid%2F7719130&rft.aulast=Armstrong&rft.aufirst=E&rft.au=Schleicher%2C+A&rft.au=Omran%2C+H&rft.au=Curtis%2C+M&rft.au=Zilles%2C+K&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGyrus" class="Z3988"></span></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-5">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFRajagopalanScottHabasKim2011" class="citation journal cs1">Rajagopalan, V; Scott, J; Habas, PA; Kim, K; Corbett-Detig, J; Rousseau, F; Barkovich, AJ; Glenn, OA; Studholme, C (23 February 2011). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3093305">"Local tissue growth patterns underlying normal fetal human brain gyrification quantified in utero"</a>. <i>The Journal of Neuroscience</i>. <b>31</b> (8): 2878–87. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1523%2Fjneurosci.5458-10.2011">10.1523/jneurosci.5458-10.2011</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3093305">3093305</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/21414909">21414909</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+Journal+of+Neuroscience&rft.atitle=Local+tissue+growth+patterns+underlying+normal+fetal+human+brain+gyrification+quantified+in+utero.&rft.volume=31&rft.issue=8&rft.pages=2878-87&rft.date=2011-02-23&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3093305%23id-name%3DPMC&rft_id=info%3Apmid%2F21414909&rft_id=info%3Adoi%2F10.1523%2Fjneurosci.5458-10.2011&rft.aulast=Rajagopalan&rft.aufirst=V&rft.au=Scott%2C+J&rft.au=Habas%2C+PA&rft.au=Kim%2C+K&rft.au=Corbett-Detig%2C+J&rft.au=Rousseau%2C+F&rft.au=Barkovich%2C+AJ&rft.au=Glenn%2C+OA&rft.au=Studholme%2C+C&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3093305&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGyrus" class="Z3988"></span></span> </li> <li id="cite_note-BarkovichGuerrini2012-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-BarkovichGuerrini2012_6-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBarkovichGuerriniKuznieckyJackson2012" class="citation journal cs1">Barkovich, A. J.; Guerrini, R.; Kuzniecky, R. I.; Jackson, G. D.; Dobyns, W. B. (2012). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3338922">"A developmental and genetic classification for malformations of cortical development: update 2012"</a>. <i>Brain</i>. <b>135</b> (5): 1348–1369. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fbrain%2Faws019">10.1093/brain/aws019</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0006-8950">0006-8950</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3338922">3338922</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/22427329">22427329</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Brain&rft.atitle=A+developmental+and+genetic+classification+for+malformations+of+cortical+development%3A+update+2012&rft.volume=135&rft.issue=5&rft.pages=1348-1369&rft.date=2012&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3338922%23id-name%3DPMC&rft.issn=0006-8950&rft_id=info%3Apmid%2F22427329&rft_id=info%3Adoi%2F10.1093%2Fbrain%2Faws019&rft.aulast=Barkovich&rft.aufirst=A.+J.&rft.au=Guerrini%2C+R.&rft.au=Kuzniecky%2C+R.+I.&rft.au=Jackson%2C+G.+D.&rft.au=Dobyns%2C+W.+B.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3338922&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGyrus" class="Z3988"></span></span> </li> <li id="cite_note-PangAtefy2008-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-PangAtefy2008_7-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPangAtefySheen2008" class="citation journal cs1">Pang, Trudy; Atefy, Ramin; Sheen, Volney (2008). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3547618">"Malformations of Cortical Development"</a>. <i>The Neurologist</i>. <b>14</b> (3): 181–191. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1097%2FNRL.0b013e31816606b9">10.1097/NRL.0b013e31816606b9</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/1074-7931">1074-7931</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3547618">3547618</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/18469675">18469675</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+Neurologist&rft.atitle=Malformations+of+Cortical+Development&rft.volume=14&rft.issue=3&rft.pages=181-191&rft.date=2008&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3547618%23id-name%3DPMC&rft.issn=1074-7931&rft_id=info%3Apmid%2F18469675&rft_id=info%3Adoi%2F10.1097%2FNRL.0b013e31816606b9&rft.aulast=Pang&rft.aufirst=Trudy&rft.au=Atefy%2C+Ramin&rft.au=Sheen%2C+Volney&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3547618&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGyrus" class="Z3988"></span></span> </li> <li id="cite_note-guerrini5-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-guerrini5_8-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFGuerrini_R2005" class="citation journal cs1">Guerrini R (2005). <a rel="nofollow" class="external text" href="https://doi.org/10.1111%2Fj.0013-9580.2005.461010.x">"Genetic malformations of the cerebral cortex and epilepsy"</a>. <i>Epilepsia</i>. <b>46</b> (Suppl 1): 32–37. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1111%2Fj.0013-9580.2005.461010.x">10.1111/j.0013-9580.2005.461010.x</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/15816977">15816977</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:24119081">24119081</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Epilepsia&rft.atitle=Genetic+malformations+of+the+cerebral+cortex+and+epilepsy&rft.volume=46&rft.issue=Suppl+1&rft.pages=32-37&rft.date=2005&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A24119081%23id-name%3DS2CID&rft_id=info%3Apmid%2F15816977&rft_id=info%3Adoi%2F10.1111%2Fj.0013-9580.2005.461010.x&rft.au=Guerrini+R&rft_id=https%3A%2F%2Fdoi.org%2F10.1111%252Fj.0013-9580.2005.461010.x&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGyrus" class="Z3988"></span></span> </li> <li id="cite_note-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-9">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFDobyns_WB1987" class="citation journal cs1">Dobyns WB (1987). "Developmental aspects of lissencephaly and the lissencephaly syndromes". <i>Birth Defects Orig. Artic. Ser</i>. <b>23</b> (1): 225–41. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/3472611">3472611</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Birth+Defects+Orig.+Artic.+Ser.&rft.atitle=Developmental+aspects+of+lissencephaly+and+the+lissencephaly+syndromes&rft.volume=23&rft.issue=1&rft.pages=225-41&rft.date=1987&rft_id=info%3Apmid%2F3472611&rft.au=Dobyns+WB&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGyrus" class="Z3988"></span></span> </li> <li id="cite_note-chang-10"><span class="mw-cite-backlink"><b><a href="#cite_ref-chang_10-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFChangWalshApseBodell1993" class="citation journal cs1">Chang, B; Walsh, CA; Apse, K; Bodell, A; Pagon, RA; Adam, TD; Bird, CR; Dolan, K; Fong, MP; Stephens, K (1993). "Polymicrogyria Overview". <i>GeneReviews</i>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/20301504">20301504</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=GeneReviews&rft.atitle=Polymicrogyria+Overview&rft.date=1993&rft_id=info%3Apmid%2F20301504&rft.aulast=Chang&rft.aufirst=B&rft.au=Walsh%2C+CA&rft.au=Apse%2C+K&rft.au=Bodell%2C+A&rft.au=Pagon%2C+RA&rft.au=Adam%2C+TD&rft.au=Bird%2C+CR&rft.au=Dolan%2C+K&rft.au=Fong%2C+MP&rft.au=Stephens%2C+K&rfr_id=info%3Asid%2Fen.wikipedia.org%3AGyrus" class="Z3988"></span></span> </li> <li id="cite_note-11"><span class="mw-cite-backlink"><b><a href="#cite_ref-11">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFSmithWalsh2020" class="citation journal cs1">Smith, RS; Walsh, CA (February 2020). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7092371">"Ion Channel Functions in Early Brain Development"</a>. <i>Trends in Neurosciences</i>. <b>43</b> (2): 103–114. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.tins.2019.12.004">10.1016/j.tins.2019.12.004</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7092371">7092371</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/31959360">31959360</a>.</cite><span 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aria-labelledby="Anatomy_of_the_cerebral_cortex_of_the_human_brain" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar 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href="/wiki/Special:EditPage/Template:Cerebral_cortex" title="Special:EditPage/Template:Cerebral cortex"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Anatomy_of_the_cerebral_cortex_of_the_human_brain" style="font-size:114%;margin:0 4em">Anatomy of the <a href="/wiki/Cerebral_cortex" title="Cerebral cortex">cerebral cortex</a> of the <a href="/wiki/Human_brain" title="Human brain">human brain</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Frontal_lobe" title="Frontal lobe">Frontal lobe</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Superolateral</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Prefrontal_cortex" title="Prefrontal cortex">Prefrontal</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Superior_frontal_gyrus" title="Superior frontal gyrus">Superior frontal gyrus</a> <ul><li><a href="/wiki/Brodmann_area_4" title="Brodmann area 4">4</a></li> <li><a href="/wiki/Brodmann_area_6" title="Brodmann area 6">6</a></li> <li><a href="/wiki/Brodmann_area_8" title="Brodmann area 8">8</a></li></ul></li> <li><a href="/wiki/Middle_frontal_gyrus" title="Middle frontal gyrus">Middle frontal gyrus</a> <ul><li><a href="/wiki/Brodmann_area_9" title="Brodmann area 9">9</a></li> <li><a href="/wiki/Brodmann_area_10" title="Brodmann area 10">10</a></li> <li><a href="/wiki/Brodmann_area_46" title="Brodmann area 46">46</a></li></ul></li></ul> <ul><li><a href="/wiki/Inferior_frontal_gyrus" title="Inferior frontal gyrus">Inferior frontal gyrus</a>: <a href="/wiki/Brodmann_area_11" title="Brodmann area 11">11</a></li> <li><a href="/wiki/Brodmann_area_47" title="Brodmann area 47">47</a>-<a href="/wiki/Orbital_part_of_inferior_frontal_gyrus" title="Orbital part of inferior frontal gyrus">Pars orbitalis</a></li> <li><a href="/wiki/Broca%27s_area" title="Broca's area">Broca's area</a> <ul><li><a href="/wiki/Brodmann_area_44" title="Brodmann area 44">44</a>-<a href="/wiki/Opercular_part_of_inferior_frontal_gyrus" class="mw-redirect" title="Opercular part of inferior frontal gyrus">Pars opercularis</a></li> <li><a href="/wiki/Brodmann_area_45" title="Brodmann area 45">45</a>-<a href="/wiki/Triangular_part_of_inferior_frontal_gyrus" class="mw-redirect" title="Triangular part of inferior frontal gyrus">Pars triangularis</a></li></ul></li></ul> <ul><li><a href="/wiki/Superior_frontal_sulcus" title="Superior frontal sulcus">Superior frontal sulcus</a></li> <li><a href="/wiki/Inferior_frontal_sulcus" title="Inferior frontal sulcus">Inferior frontal sulcus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Precentral_gyrus" title="Precentral gyrus">Precentral</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Precentral_gyrus" title="Precentral gyrus">Precentral gyrus</a></li> <li><a href="/wiki/Precentral_sulcus" title="Precentral sulcus">Precentral sulcus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Medial/inferior</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Prefrontal_cortex" title="Prefrontal cortex">Prefrontal</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Superior_frontal_gyrus" title="Superior frontal gyrus">Superior frontal gyrus</a> <ul><li><a href="/wiki/Brodmann_area_4" title="Brodmann area 4">4</a></li> <li><a href="/wiki/Brodmann_area_6" title="Brodmann area 6">6</a></li></ul></li> <li><a href="/wiki/Medial_frontal_gyrus" title="Medial frontal gyrus">Medial frontal gyrus</a> <ul><li><a href="/wiki/Brodmann_area_8" title="Brodmann area 8">8</a></li> <li><a href="/wiki/Brodmann_area_9" title="Brodmann area 9">9</a></li></ul></li></ul> <ul><li><a href="/wiki/Paraterminal_gyrus" class="mw-redirect" title="Paraterminal gyrus">Paraterminal gyrus</a>/<a href="/wiki/Paraolfactory_area" class="mw-redirect" title="Paraolfactory area">Paraolfactory area</a> <ul><li><a href="/wiki/Brodmann_area_12" title="Brodmann area 12">12</a></li></ul></li> <li><a href="/wiki/Straight_gyrus" title="Straight gyrus">Straight gyrus</a> <ul><li><a href="/wiki/Brodmann_area_11" title="Brodmann area 11">11</a></li></ul></li> <li><a href="/wiki/Orbital_gyri" title="Orbital gyri">Orbital gyri</a>/<a href="/wiki/Orbitofrontal_cortex" title="Orbitofrontal cortex">Orbitofrontal cortex</a> <ul><li><a href="/wiki/Brodmann_area_10" title="Brodmann area 10">10</a></li> <li><a href="/wiki/Brodmann_area_11" title="Brodmann area 11">11</a></li> <li><a href="/wiki/Brodmann_area_12" title="Brodmann area 12">12</a></li></ul></li> <li><a href="/wiki/Ventromedial_prefrontal_cortex" title="Ventromedial prefrontal cortex">Ventromedial prefrontal cortex</a> <ul><li><a href="/wiki/Brodmann_area_10" title="Brodmann area 10">10</a></li></ul></li> <li><a href="/wiki/Subcallosal_area" title="Subcallosal area">Subcallosal area</a> <ul><li><a href="/wiki/Brodmann_area_25" title="Brodmann area 25">25</a></li></ul></li></ul> <ul><li><a href="/wiki/Olfactory_sulcus" class="mw-redirect" title="Olfactory sulcus">Olfactory sulcus</a></li> <li><a href="/wiki/Orbital_sulcus" title="Orbital sulcus">Orbital sulcus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Precentral_gyrus" title="Precentral gyrus">Precentral</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Paracentral_lobule" title="Paracentral lobule">Paracentral lobule</a> <ul><li><a href="/wiki/Brodmann_area_4" title="Brodmann area 4">4</a></li></ul></li> <li><a href="/wiki/Paracentral_sulcus" title="Paracentral sulcus">Paracentral sulcus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Both</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Primary_motor_cortex" title="Primary motor cortex">Primary motor cortex</a> <ul><li><a href="/wiki/Brodmann_area_4" title="Brodmann area 4">4</a></li></ul></li> <li><a href="/wiki/Premotor_cortex" title="Premotor cortex">Premotor cortex</a> <ul><li><a href="/wiki/Brodmann_area_6" title="Brodmann area 6">6</a></li></ul></li> <li><a href="/wiki/Supplementary_motor_area" title="Supplementary motor area">Supplementary motor area</a> <ul><li><a href="/wiki/Brodmann_area_6" title="Brodmann area 6">6</a></li></ul></li> <li><a href="/wiki/Supplementary_eye_field" title="Supplementary eye field">Supplementary eye field</a> <ul><li><a href="/wiki/Brodmann_area_6" title="Brodmann area 6">6</a></li></ul></li> <li><a href="/wiki/Frontal_eye_fields" title="Frontal eye fields">Frontal eye fields</a> <ul><li><a href="/wiki/Brodmann_area_8" title="Brodmann area 8">8</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Parietal_lobe" title="Parietal lobe">Parietal lobe</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Superolateral</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Superior_parietal_lobule" title="Superior parietal lobule">Superior parietal lobule</a> <ul><li><a href="/wiki/Brodmann_area_5" title="Brodmann area 5">5</a></li> <li><a href="/wiki/Brodmann_area_7" title="Brodmann area 7">7</a></li></ul></li> <li><a href="/wiki/Inferior_parietal_lobule" title="Inferior parietal lobule">Inferior parietal lobule</a> <ul><li><a href="/wiki/Brodmann_area_40" title="Brodmann area 40">40</a>-<a href="/wiki/Supramarginal_gyrus" title="Supramarginal gyrus">Supramarginal gyrus</a></li> <li><a href="/wiki/Brodmann_area_39" title="Brodmann area 39">39</a>-<a href="/wiki/Angular_gyrus" title="Angular gyrus">Angular gyrus</a></li></ul></li> <li><a href="/wiki/Parietal_operculum" class="mw-redirect" title="Parietal operculum">Parietal operculum</a> <ul><li><a href="/wiki/Brodmann_area_43" title="Brodmann area 43">43</a></li></ul></li></ul> <ul><li><a href="/wiki/Intraparietal_sulcus" title="Intraparietal sulcus">Intraparietal sulcus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Medial/inferior</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Paracentral_lobule" title="Paracentral lobule">Paracentral lobule</a> <ul><li><a href="/wiki/Brodmann_area_1" class="mw-redirect" title="Brodmann area 1">1</a></li> <li><a href="/wiki/Brodmann_area_2" class="mw-redirect" title="Brodmann area 2">2</a></li> <li><a href="/wiki/Brodmann_area_3" class="mw-redirect" title="Brodmann area 3">3</a></li> <li><a href="/wiki/Brodmann_area_5" title="Brodmann area 5">5</a></li></ul></li> <li><a href="/wiki/Precuneus" title="Precuneus">Precuneus</a> <ul><li><a href="/wiki/Brodmann_area_7" title="Brodmann area 7">7</a></li></ul></li> <li><a href="/wiki/Marginal_sulcus" title="Marginal sulcus">Marginal sulcus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Both</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Postcentral_gyrus" title="Postcentral gyrus">Postcentral gyrus</a>/<a href="/wiki/Primary_somatosensory_cortex" title="Primary somatosensory cortex">Primary somatosensory cortex</a> <ul><li><a href="/wiki/Brodmann_areas_3,_1_and_2" class="mw-redirect" title="Brodmann areas 3, 1 and 2">3, 1 and 2</a></li></ul></li> <li><a href="/wiki/Secondary_somatosensory_cortex" title="Secondary somatosensory cortex">Secondary somatosensory cortex</a> <ul><li><a href="/wiki/Brodmann_area_5" title="Brodmann area 5">5</a></li></ul></li> <li><a href="/wiki/Posterior_parietal_cortex" title="Posterior parietal cortex">Posterior parietal cortex</a> <ul><li><a href="/wiki/Brodmann_area_7" title="Brodmann area 7">7</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Occipital_lobe" title="Occipital lobe">Occipital lobe</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Superolateral</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Occipital_pole_of_cerebrum" class="mw-redirect" title="Occipital pole of cerebrum">Occipital pole of cerebrum</a></li> <li><a href="/wiki/Occipital_gyri" title="Occipital gyri">Occipital gyri</a></li> <li><a href="/wiki/Lateral_occipital_sulcus" title="Lateral occipital sulcus">Lateral occipital gyrus</a> <ul><li><a href="/wiki/Brodmann_area_18" title="Brodmann area 18">18</a></li> <li><a href="/wiki/Brodmann_area_19" title="Brodmann area 19">19</a></li></ul></li> <li><a href="/wiki/Lunate_sulcus" title="Lunate sulcus">Lunate sulcus</a></li> <li><a href="/wiki/Transverse_occipital_sulcus" title="Transverse occipital sulcus">Transverse occipital sulcus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Medial/inferior</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Visual_cortex" title="Visual cortex">Visual cortex</a> <ul><li><a href="/wiki/Brodmann_area_17" class="mw-redirect" title="Brodmann area 17">17</a></li></ul></li> <li><a href="/wiki/Cuneus" title="Cuneus">Cuneus</a></li> <li><a href="/wiki/Lingual_gyrus" title="Lingual gyrus">Lingual gyrus</a></li></ul> <ul><li><a href="/wiki/Calcarine_sulcus" title="Calcarine sulcus">Calcarine sulcus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Temporal_lobe" title="Temporal lobe">Temporal lobe</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Superolateral</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Transverse_temporal_gyrus" title="Transverse temporal gyrus">Transverse temporal gyrus</a>/<a href="/wiki/Auditory_cortex" title="Auditory cortex">Auditory cortex</a> <ul><li><a href="/wiki/Brodmann_areas_41_and_42" title="Brodmann areas 41 and 42">41 and 42</a></li></ul></li> <li><a href="/wiki/Superior_temporal_gyrus" title="Superior temporal gyrus">Superior temporal gyrus</a> <ul><li><a href="/wiki/Brodmann_area_38" title="Brodmann area 38">38</a></li> <li><a href="/wiki/Brodmann_area_22" title="Brodmann area 22">22</a>/<a href="/wiki/Wernicke%27s_area" title="Wernicke's area">Wernicke's area</a> (<a href="/wiki/Planum_temporale" title="Planum temporale">Planum temporale</a>)</li></ul></li> <li><a href="/wiki/Superior_temporal_sulcus" title="Superior temporal sulcus">Superior temporal sulcus</a></li> <li><a href="/wiki/Middle_temporal_gyrus" title="Middle temporal gyrus">Middle temporal gyrus</a> <ul><li><a href="/wiki/Brodmann_area_21" title="Brodmann area 21">21</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Medial/inferior</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Occipitotemporal_sulcus" title="Occipitotemporal sulcus">Occipitotemporal sulcus</a></li> <li><a href="/wiki/Fusiform_gyrus" title="Fusiform gyrus">Fusiform gyrus</a> <ul><li><a href="/wiki/Brodmann_area_37" title="Brodmann area 37">37</a></li></ul></li> <li><a href="/wiki/Medial_temporal_lobe" class="mw-redirect" title="Medial temporal lobe">Medial temporal lobe</a> <ul><li><a href="/wiki/Brodmann_area_27" title="Brodmann area 27">27</a></li> <li><a href="/wiki/Brodmann_area_28" title="Brodmann area 28">28</a></li> <li><a href="/wiki/Brodmann_area_34" title="Brodmann area 34">34</a></li> <li><a href="/wiki/Brodmann_area_35" class="mw-redirect" title="Brodmann area 35">35</a></li> <li><a href="/wiki/Brodmann_area_36" class="mw-redirect" title="Brodmann area 36">36</a></li></ul></li></ul> <ul><li><a href="/wiki/Inferior_temporal_sulcus" class="mw-redirect" title="Inferior temporal sulcus">Inferior temporal sulcus</a></li> <li><a href="/wiki/Inferior_temporal_gyrus" title="Inferior temporal gyrus">Inferior temporal gyrus</a> <ul><li><a href="/wiki/Brodmann_area_20" title="Brodmann area 20">20</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Sulcus_(neuroanatomy)" title="Sulcus (neuroanatomy)">Interlobar<br />sulci/fissures</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Superolateral</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Central_sulcus" title="Central sulcus">Central (frontal+parietal)</a></li> <li><a href="/wiki/Lateral_sulcus" title="Lateral sulcus">Lateral (frontal+parietal+temporal)</a></li> <li><a href="/wiki/Parieto-occipital_sulcus" title="Parieto-occipital sulcus">Parieto-occipital</a></li> <li><a href="/wiki/Preoccipital_notch" title="Preoccipital notch">Preoccipital notch</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Medial/inferior</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Longitudinal_fissure" title="Longitudinal fissure">Longitudinal fissure</a></li> <li><a href="/wiki/Cingulate_sulcus" title="Cingulate sulcus">Cingulate (frontal+cingulate)</a></li> <li><a href="/wiki/Collateral_fissure" title="Collateral fissure">Collateral (temporal+occipital)</a></li> <li><a href="/wiki/Callosal_sulcus" class="mw-redirect" title="Callosal sulcus">Callosal sulcus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Limbic_lobe" title="Limbic lobe">Limbic lobe</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Parahippocampal_gyrus" title="Parahippocampal gyrus">Parahippocampal gyrus</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>anterior</i> <ul><li><a href="/wiki/Entorhinal_cortex" title="Entorhinal cortex">Entorhinal cortex</a></li> <li><a href="/wiki/Perirhinal_cortex" title="Perirhinal cortex">Perirhinal cortex</a></li> <li><a href="/wiki/Postrhinal_cortex" title="Postrhinal cortex">Postrhinal cortex</a></li></ul></li> <li><a href="/wiki/Posterior_parahippocampal_gyrus" title="Posterior parahippocampal gyrus">Posterior parahippocampal gyrus</a></li> <li><a href="/wiki/Prepyriform_area" title="Prepyriform area">Prepyriform area</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Cingulate_cortex" title="Cingulate cortex">Cingulate cortex</a>/<a href="/wiki/Cingulate_gyrus" class="mw-redirect" title="Cingulate gyrus">gyrus</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Subgenual_area_25" class="mw-redirect" title="Subgenual area 25">Subgenual area</a> <ul><li><a href="/wiki/Brodmann_area_25" title="Brodmann area 25">25</a></li></ul></li> <li><a href="/wiki/Anterior_cingulate_cortex" title="Anterior cingulate cortex">Anterior cingulate</a> <ul><li><a href="/wiki/Brodmann_area_24" title="Brodmann area 24">24</a></li> <li><a href="/wiki/Brodmann_area_32" title="Brodmann area 32">32</a></li> <li><a href="/wiki/Brodmann_area_33" title="Brodmann area 33">33</a></li></ul></li> <li><a href="/wiki/Posterior_cingulate" class="mw-redirect" title="Posterior cingulate">Posterior cingulate</a> <ul><li><a href="/wiki/Brodmann_area_23" title="Brodmann area 23">23</a></li> <li><a href="/wiki/Brodmann_area_31" title="Brodmann area 31">31</a></li></ul></li></ul> <ul><li><a href="/wiki/Isthmus_of_cingulate_gyrus" title="Isthmus of cingulate gyrus">Isthmus of cingulate gyrus</a>: <a href="/wiki/Retrosplenial_region" class="mw-redirect" title="Retrosplenial region">Retrosplenial cortex</a> <ul><li><a href="/wiki/Brodmann_area_26" title="Brodmann area 26">26</a></li> <li><a href="/wiki/Brodmann_area_29" title="Brodmann area 29">29</a></li> <li><a href="/wiki/Brodmann_area_30" title="Brodmann area 30">30</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Hippocampal_formation" title="Hippocampal formation">Hippocampal formation</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Hippocampal_sulcus" title="Hippocampal sulcus">Hippocampal sulcus</a></li> <li><a href="/wiki/Fimbria_of_hippocampus" class="mw-redirect" title="Fimbria of hippocampus">Fimbria of hippocampus</a></li> <li><a href="/wiki/Dentate_gyrus" title="Dentate gyrus">Dentate gyrus</a></li> <li><a href="/wiki/Rhinal_sulcus" title="Rhinal sulcus">Rhinal sulcus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Indusium_griseum" title="Indusium griseum">Indusium griseum</a></li> <li><a href="/wiki/Uncus" title="Uncus">Uncus</a></li> <li><a href="/wiki/Amygdala" title="Amygdala">Amygdala</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Insular_cortex" title="Insular cortex">Insular cortex</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Insular_cortex" title="Insular cortex">Insular cortex</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">General</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Operculum_(brain)" title="Operculum (brain)">Operculum</a></li> <li><a href="/wiki/Poles_of_cerebral_hemispheres" class="mw-redirect" title="Poles of cerebral hemispheres">Poles of cerebral hemispheres</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2" style="background: transparent; padding: 0px;"><div><i>Some categorizations are approximations, and some <a href="/wiki/Brodmann_area" title="Brodmann area">Brodmann areas</a> span gyri.</i></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox authority-control" aria-label="Navbox" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Help:Authority_control" title="Help:Authority control">Authority control databases</a> <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q1758383#identifiers" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" 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