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Plastid – Wikipedia

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border: 1px solid #CCD2D9; width: 200px; text-align: center; border-collapse: collapse; font-size: 90%;"> <tbody><tr> <th style="background:#ABCDEF; color:#202122; border: 1px solid #ABCDEF;">Übergeordnet </th></tr> <tr> <td style="padding: 0 .5em;"><a href="/wiki/Organell" title="Organell">Organell</a> </td></tr> <tr> <th style="background:#ABCDEF; color:#202122; border: 1px solid #ABCDEF;">Untergeordnet </th></tr> <tr> <td style="padding: 0 .5em;"><a href="/wiki/Chloroplast" title="Chloroplast">Chloroplast</a><br /><a href="/wiki/Amyloplast" title="Amyloplast">Amyloplast</a><br /><a href="/wiki/Chromoplast" title="Chromoplast">Chromoplast</a><br /><a href="/wiki/Etioplast" title="Etioplast">Etioplast</a><br /><a href="/wiki/Leukoplasten" title="Leukoplasten">Leukoplasten</a><br /><a href="/wiki/Elaioplast" class="mw-redirect" title="Elaioplast">Elaioplast</a><br /><a href="/wiki/Proteinoplast" title="Proteinoplast">Proteinoplast</a><br /><a href="/w/index.php?title=Apicoplast&amp;action=edit&amp;redlink=1" class="new" title="Apicoplast (Seite nicht vorhanden)">Apicoplast</a> (Apikoplast)<br /><a href="/wiki/Gerontoplast" title="Gerontoplast">Gerontoplast</a><br /><a href="/w/index.php?title=Cyanelle&amp;action=edit&amp;redlink=1" class="new" title="Cyanelle (Seite nicht vorhanden)">Cyanelle</a> (Cyanoplast, Muroplast)<br /><a href="/wiki/Proplastid" title="Proplastid">Proplastid</a><br /><a href="/wiki/Rhodoplast" title="Rhodoplast">Rhodoplast</a><br />Phaeoplast<br /><a href="/wiki/Chromatophor" class="mw-redirect" title="Chromatophor">Chromatophor</a><sup id="cite_ref-BMCBiology-EvolvinganOrganelle_1-0" class="reference"><a href="#cite_note-BMCBiology-EvolvinganOrganelle-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-MolecularBiologyEvol-EndosymbioticGeneTransfer_2-0" class="reference"><a href="#cite_note-MolecularBiologyEvol-EndosymbioticGeneTransfer-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><br /><a href="/wiki/Kleptoplastid" title="Kleptoplastid">Kleptoplastid</a> </td></tr> <tr> <th style="background:#ABCDEF; color:ä202122; border: 1px solid #ABCDEF;"><a href="/wiki/Gene_Ontology" title="Gene Ontology">Gene Ontology</a> </th></tr> <tr> <td style="text-align:left; padding: 0 .5em;"><a rel="nofollow" class="external text" href="http://www.ebi.ac.uk/QuickGO/GTerm?id=GO:0009536">QuickGO</a> </td></tr></tbody></table> <p><b>Plastiden</b> (von <span style="font-style:normal;font-weight:normal"><a href="/wiki/Altgriechische_Sprache" title="Altgriechische Sprache">altgriechisch</a></span> <span lang="grc-Grek" class="Grek" style="font-style:normal">πλαστός</span> <i>plastós</i> „geformt“) sind die in <a href="/wiki/Pflanzen" class="mw-redirect" title="Pflanzen">Pflanzen</a> und <a href="/wiki/Alge" title="Alge">Algen</a> vorkommenden besonderen <a href="/wiki/Organell" title="Organell">Zellorganellen</a>, die aus <a href="/wiki/Endosymbiontentheorie" title="Endosymbiontentheorie">endosymbiontisch</a> lebenden Zellen hervorgegangen sind und unter anderem für die <a href="/wiki/Photosynthese" title="Photosynthese">Photosynthese</a> gebraucht werden, oder die aus solchen Organellen evolutionär hervorgegangen sind. </p> <div id="toc" class="toc" role="navigation" aria-labelledby="mw-toc-heading"><input type="checkbox" role="button" id="toctogglecheckbox" class="toctogglecheckbox" style="display:none" /><div class="toctitle" lang="de" dir="ltr"><h2 id="mw-toc-heading">Inhaltsverzeichnis</h2><span class="toctogglespan"><label class="toctogglelabel" for="toctogglecheckbox"></label></span></div> <ul> <li class="toclevel-1 tocsection-1"><a href="#Bau"><span class="tocnumber">1</span> <span class="toctext">Bau</span></a></li> <li class="toclevel-1 tocsection-2"><a href="#Evolution"><span class="tocnumber">2</span> <span class="toctext">Evolution</span></a></li> <li class="toclevel-1 tocsection-3"><a href="#Teilung"><span class="tocnumber">3</span> <span class="toctext">Teilung</span></a></li> <li class="toclevel-1 tocsection-4"><a href="#Sonstiges"><span class="tocnumber">4</span> <span class="toctext">Sonstiges</span></a></li> <li class="toclevel-1 tocsection-5"><a href="#Literatur"><span class="tocnumber">5</span> <span class="toctext">Literatur</span></a></li> <li class="toclevel-1 tocsection-6"><a href="#Weblinks"><span class="tocnumber">6</span> <span class="toctext">Weblinks</span></a></li> <li class="toclevel-1 tocsection-7"><a href="#Einzelnachweise"><span class="tocnumber">7</span> <span class="toctext">Einzelnachweise</span></a></li> </ul> </div> <div class="mw-heading mw-heading2"><h2 id="Bau">Bau</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plastid&amp;veaction=edit&amp;section=1" title="Abschnitt bearbeiten: Bau" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Plastid&amp;action=edit&amp;section=1" title="Quellcode des Abschnitts bearbeiten: Bau"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Plastiden verfügen (fast immer<sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup>) über ein eigenes ringförmiges <a href="/wiki/Genom" title="Genom">Genom</a> (plastidäres Genom oder auch <b><a href="/wiki/Chloroplasten-DNA" title="Chloroplasten-DNA">Plastom</a></b> genannt), sowie eigene <a href="/wiki/Ribosom" title="Ribosom">Ribosomen</a> (<i><a href="/w/index.php?title=Plastoribosomen&amp;action=edit&amp;redlink=1" class="new" title="Plastoribosomen (Seite nicht vorhanden)">Plastoribosomen</a></i> genannt), die in seine <a href="/wiki/Cytoplasma" title="Cytoplasma">plasmatische</a> Grundsubstanz (<i>Stroma</i>) eingebettet sind. Darüber hinaus liegen weitere plastidenspezifische Komponenten für die plastidäre <a href="/wiki/Replikation" title="Replikation">Replikation</a>, <a href="/wiki/Transkription_(Biologie)" title="Transkription (Biologie)">Transkription</a> und <a href="/wiki/Translation_(Biologie)" title="Translation (Biologie)">Translation</a> vor. </p><p>Nach den sie umhüllenden <a href="/wiki/Biomembran" title="Biomembran">Membranen</a> unterscheidet man einfache Plastiden, die auf ein primäres Endosymbioseereignis zurückzuführen sind und von zwei Hüllmembranen umgeben sind, und <a href="/wiki/Komplexe_Plastiden" title="Komplexe Plastiden">komplexe Plastiden</a>, die durch sekundäre oder tertiäre Endosymbiose entstanden und so drei oder vier Hüllmembranen besitzen. Kommen in einer Zelle mehrere Plastiden vor, so sind diese meist über <a href="/wiki/Stromuli" class="mw-redirect" title="Stromuli">Stromuli</a> miteinander verbunden. </p> <div class="mw-heading mw-heading2"><h2 id="Evolution">Evolution</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plastid&amp;veaction=edit&amp;section=2" title="Abschnitt bearbeiten: Evolution" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Plastid&amp;action=edit&amp;section=2" title="Quellcode des Abschnitts bearbeiten: Evolution"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/Datei:Plastids_types_en.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/88/Plastids_types_en.svg/220px-Plastids_types_en.svg.png" decoding="async" width="220" height="183" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/88/Plastids_types_en.svg/330px-Plastids_types_en.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/88/Plastids_types_en.svg/440px-Plastids_types_en.svg.png 2x" data-file-width="560" data-file-height="465" /></a><figcaption>Beziehungen zwischen den Typen von Plastiden in <a href="/wiki/Gef%C3%A4%C3%9Fpflanzen" title="Gefäßpflanzen">Gefäßpflanzen</a></figcaption></figure> <p>Die einfachen Plastiden der <a href="/wiki/Glaucocystaceae" class="mw-redirect" title="Glaucocystaceae">Glaucocystophyceen</a>, <a href="/wiki/Rotalgen" title="Rotalgen">Rotalgen</a>, <a href="/wiki/Gr%C3%BCnalge" title="Grünalge">Grünalgen</a> (Chlorophyta) und <a href="/wiki/Landpflanzen" class="mw-redirect" title="Landpflanzen">Landpflanzen</a> (Embryophyta) stammen wahrscheinlich aus einer primären <a href="/wiki/Endosymbiose" class="mw-redirect" title="Endosymbiose">Endosymbiose</a>. Sie sind monophyletisch, d.&#160;h. die drei Algengruppen und die Landpflanzen (Embryophyta) stammen von einem gemeinsamen einzelligen Vorfahren ab, dessen Nachkommen sich in drei Evolutionslinien aufspalteten: </p> <ul><li>Die Plastiden der Glaucocystophyceen werden <i>Cyanellen</i> (auch <i>Cyanoplasten</i> oder <i>Muroplasten</i>) genannt, verwenden als Lichtsammelkomplexe <a href="/wiki/Phycobilisom" title="Phycobilisom">Phycobilisomen</a> und sind noch von einem Rest einer bakteriellen Zellwand umgeben.</li> <li>Die Plastiden der Rotalgen heißen <i>Rhodoplasten</i>, enthalten ebenfalls noch Phycobilisomen, jedoch keine bakterielle Zellwand mehr.</li> <li>Die <a href="/wiki/Chloroplast" title="Chloroplast">Chloroplasten</a> der Grünalgen und höheren Pflanzen bilden keine Phycobilisomen mehr, enthalten Chlorophyll b und bilden <a href="/wiki/St%C3%A4rke" title="Stärke">Stärke</a> im Plastiden.</li> <li>Die Plastiden der <a href="/wiki/Braunalgen" title="Braunalgen">Braunalgen</a> nennt man auch <i>Phaeoplasten</i>.</li></ul> <p>Auch die Plastiden der <a href="/wiki/Am%C3%B6be" title="Amöbe">amöboiden</a> <i><a href="/wiki/Paulinella" title="Paulinella">Paulinella</a> chromatophora</i> (<a href="/wiki/Euglyphida" title="Euglyphida">Euglyphida</a>, s.&#160;u.), sowie einiger anderer Arten dieser Gattung, sind offenbar primäre Chloroplasten (<a href="/wiki/Chromatophor" class="mw-redirect" title="Chromatophor">Chromatophoren</a> oder <a href="/wiki/Cyanellen" class="mw-redirect" title="Cyanellen">Cyanellen</a> genannt). Diese <a href="/wiki/Klade" title="Klade">Klade</a> der Gattung <i>Paulinella</i> stammt offenbar von einem Vorfahren ab, der unabhängig und viel später ein Cyanobakterium aus der Verwandtschaft der Gattungen <i><a href="/wiki/Prochlorococcus" class="mw-redirect" title="Prochlorococcus">Prochlorococcus</a></i> und <i><a href="/wiki/Synechococcus" title="Synechococcus">Synechococcus</a></i> aufgenommen hatte.<sup id="cite_ref-paulinella_5-0" class="reference"><a href="#cite_note-paulinella-5"><span class="cite-bracket">&#91;</span>5<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-:1_6-0" class="reference"><a href="#cite_note-:1-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-PSanchezB2017_7-0" class="reference"><a href="#cite_note-PSanchezB2017-7"><span class="cite-bracket">&#91;</span>7<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">&#91;</span>8<span class="cite-bracket">&#93;</span></a></sup> </p> <div style="z-index: 0; border-radius: 2px; position: relative; height: 675px; float: left; width: 69%; margin: 10px; margin-bottom: 4px; overflow: hidden;"> <div style="z-index: 2; border-radius: 2px; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 0px; left: 0px; width: 100%; background: #00c9be; color: #00c9be;"><a href="/wiki/Cyanobacteria" class="mw-redirect" title="Cyanobacteria"><span style="background-color: rgba(255,255,255,0.8); padding-left: 4px; padding-right: 4px; color: #00c9be;">Cyanobacteria</span></a></div> <div style="position: absolute; line-height: 28px; height: 100px; top: 0px; margin-left: 15%; width: 4px; background: #00c9be; color: #00c9be;"></div> <div style="z-index: 3; border-radius: 2px 0 0 2px; position: absolute; line-height: 28px; height: 4px; top: 100px; left: 0px; width: 15%; background: #ddd; color: #ddd;"></div> <div style="z-index: 3; position: absolute; width: 0px; height: 0px; left: 15%; top: 100px;"> <div style="position: relative; line-height: 28px; height: 16px; top: -14px; left: -14px; width: 16px; border-radius: 16px; background: white; padding: 8px;"> <div style="position: relative; line-height: 28px; height: 16px; left: -4px; top: -4px; width: 16px; border-radius: 16px; background: white; border: 4px solid rgb(144,238,144); color: rgb(144,238,144);"></div></div> </div> <div style="z-index: 3; border-radius: 2px 0 0 2px; position: absolute; line-height: 20px; height: 40px; top: 104px; left: 0px; color: #ddd; background-color: rgba(255,255,255,0.8); padding-right: 2px;"><b><a href="/wiki/Archaeplastida" title="Archaeplastida"><span style="color: #ccc;">Archaeplastida</span></a></b></div> <div style="z-index: 2; position: absolute; line-height: 28px; height: 4px; top: 100px; margin-left: 15%; width: 15%; left: 2px; background: rgb(144,238,144); color: rgb(144,238,144);"></div> <div style="z-index: 2; position: absolute; line-height: 28px; height: 4px; top: 100px; margin-left: 30%; width: 5%; left: 2px; background: #24d12b; color: #24d12b;"></div> <div style="z-index: 2; border-radius: 0 2px 2px 0; z-index: 1; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 100px; margin-left: 35%; width: 65%; background: #24d12b; color: #24d12b;"><a href="/wiki/Landpflanzen" class="mw-redirect" title="Landpflanzen"><span style="background-color: rgba(255,255,255,0.8); padding-left: 4px; padding-right: 4px; color: #24d12b;">Landpflanzen</span></a></div> <div style="border-radius: 2px; position: absolute; line-height: 28px; height: 52px; top: 50px; margin-left: 25%; width: 4px; background: #00ca9b; color: #00c9be;"></div> <div style="z-index: 2; border-radius: 2px; position: absolute; line-height: 28px; height: 4px; top: 50px; right: 0px; width: 75%; background: #00ca9b; color: #00ca9b; text-align: right;"><a href="/wiki/Glaucophyta" title="Glaucophyta"><span style="background-color: rgba(255,255,255,0.8); padding-left: 4px; padding-right: 4px; color: #00ca9b;">Glaucophyta</span></a></div> <div style="z-index: 2; border-radius: 2px; position: absolute; line-height: 28px; height: 104px; top: 100px; margin-left: 35%; width: 4px; background: #24d14e; color: #24d14e;"></div> <div style="border-radius: 2px; z-index: 2; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 200px; margin-left: 35%; width: 65%; background: #24d14e; color: #24d14e;"><a href="/wiki/Gr%C3%BCnalgen" class="mw-redirect" title="Grünalgen"><span style="background-color: rgba(255,255,255,0.8); padding-left: 4px; padding-right: 4px; color: #24d14e;">Grünalgen</span></a></div> <div style="border-radius: 2px; position: absolute; line-height: 28px; height: 52px; top: 150px; margin-left: 70%; width: 4px; background: #24d14e; color: #24d14e;"></div> <div style="border-radius: 2px 0 0 2px; z-index: 0; text-align: left; position: absolute; line-height: 28px; height: 4px; top: 150px; width: 70%; background: #ddd; color: #ddd;"><b><a href="/wiki/Excavata" title="Excavata"><span style="color: #ccc;">Excavata</span></a></b></div> <div style="border-radius: 2px; z-index: 1; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 150px; margin-left: 70%; width: 30%; background: #24d16d; color: #24d16d;"><a href="/wiki/Euglenida" title="Euglenida"><span style="background-color: rgba(255,255,255,0.8)e; padding-left: 4px; padding-right: 4px; color: #24d16d;">Euglenida</span></a></div> <div style="z-index: 2; position: absolute; width: 0px; height: 0px; left: 70%; top: 150px;"> <div style="position: relative; line-height: 28px; height: 16px; top: -14px; left: -14px; width: 16px; border-radius: 16px; background: white; padding: 8px;"> <div style="position: relative; line-height: 28px; height: 16px; left: -4px; top: -4px; width: 16px; border-radius: 16px; background: white; border: 4px solid #24d16d; color: #24d16d;"></div></div> </div> <div style="z-index: 1; border-radius: 2px; position: absolute; line-height: 28px; height: 252px; top: 102px; margin-left: 29%; width: 2%; background: white; color: #ddd;"></div> <div style="border-radius: 2px; z-index: 1; position: absolute; height: 252px; width: 4px; top: 102px; margin-left: 30%; left: -2px; background: #dc003e; color: #dc003e;"></div> <div style="border-radius: 0 2px 2px 0; z-index: 1; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 350px; margin-left: 30%; width: 70%; background: #dc003e; color: #dc003e;"><a href="/wiki/Rhodophyta" class="mw-redirect" title="Rhodophyta"><span style="background: white; padding-left: 4px; padding-right: 4px; color: #dc003e;">Rhodophyta</span></a></div> <div style="z-index: 1; border-radius: 2px; position: absolute; line-height: 28px; height: 202px; top: 200px; margin-left: 37%; width: 2%; left: 2px; background: white; color: #ddd;"></div> <div style="border-radius: 2px; z-index: 1; position: absolute; height: 202px; top: 200px; width: 4px; margin-left: 38%; left: 0px; background: #24d14e; color: #24d14e;"></div> <div style="border-radius: 2px; z-index: 1; text-align: left; position: absolute; line-height: 28px; height: 4px; top: 400px; margin-left: 0%; width: 38%; background: #ddd; color: #ddd;"><b><a href="/wiki/Chromalveolata" title="Chromalveolata"><span style="color: #ccc;">Chromalveolata</span></a></b></div> <div style="z-index: 2; position: absolute; width: 0px; height: 0px; left: 38%; top: 400px;"> <div style="position: relative; line-height: 28px; height: 16px; top: -14px; left: -14px; width: 16px; border-radius: 16px; background: white; padding: 8px;"> <div style="position: relative; line-height: 28px; height: 16px; left: -4px; top: -4px; width: 16px; border-radius: 16px; background: white; border: 4px solid #24d16d; color: #24d16d;"></div></div> </div> <div style="border-radius: 2px; z-index: 1; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 400px; margin-left: 38%; width: 9%; background: #24d16d; color: #24d16d;"></div> <div style="z-index: 0; position: absolute; line-height: 28px; height: 52px; top: 350px; margin-left: 47%; width: 4px; background: #dc003e; color: #dc003e;"></div> <div style="z-index: 2; position: absolute; width: 0px; height: 0px; left: 47%; top: 400px;"> <div style="position: relative; line-height: 28px; height: 16px; top: -14px; left: -14px; width: 16px; border-radius: 16px; background: white; padding: 8px;"> <div style="position: relative; line-height: 28px; height: 16px; left: -4px; top: -4px; width: 16px; border-radius: 16px; background: white; border: 4px solid #bd3f00; color: #bd3f00;"></div></div> </div> <div style="border-radius: 2px; z-index: 1; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 400px; margin-left: 47%; width: 10%; left: 2px; background: #bd3f00; color: #bd3f00;"></div> <div style="z-index: 1; border-radius: 2px; position: absolute; line-height: 28px; height: 100px; top: 275px; margin-left: 59%; width: 2%; background: white; color: #ddd;"></div> <div style="border-radius: 2px; z-index: 1; position: absolute; height: 127px; width: 4px; top: 275px; margin-left: 60%; left: -2px; background: #ddd;"></div> <div style="border-radius: 2px; z-index: 1; text-align: left; position: absolute; line-height: 0px; height: 4px; top: 275px; margin-left: 60%; width: 5%; background: #ddd; color: #ccc;"><b><a href="/wiki/Rhizaria" title="Rhizaria"><span style="position: absolute; color: #ccc; right: 5px; bottom: 10px;">Rhizaria&#160;<sup>a</sup></span></a></b></div> <div style="z-index: 0; border-radius: 2px; position: absolute; line-height: 28px; height: 54px; top: 250px; margin-left: 65%; left: -2px; width: 4px; background: #ddd; color: #ddd;"></div> <div style="border-radius: 2px; z-index: 1; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 250px; margin-left: 65%; left: -2px; width: 5%; background: #ddd; color: #ddd;"></div> <div style="border-radius: 2px; z-index: 1; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 300px; margin-left: 65%; left: -2px; width: 20%; background: #ddd; color: #ddd;"></div> <div style="position: absolute; line-height: 28px; height: 45px; top: 0px; margin-left: 85%; width: 4px; background: #00c9be; color: #00c9be;"></div> <div style="position: absolute; line-height: 28px; height: 36px; top: 59px; margin-left: 85%; width: 4px; background: #00c9be; color: #00c9be;"></div> <div style="position: absolute; line-height: 28px; height: 36px; top: 109px; margin-left: 85%; width: 4px; background: #00c9be; color: #00c9be;"></div> <div style="position: absolute; line-height: 28px; height: 36px; top: 159px; margin-left: 85%; width: 4px; background: #00c9be; color: #00c9be;"></div> <div style="position: absolute; line-height: 28px; height: 36px; top: 209px; margin-left: 85%; width: 4px; background: #00c9be; color: #00c9be;"></div> <div style="position: absolute; line-height: 28px; height: 36px; top: 259px; margin-left: 85%; width: 4px; background: #00c9be; color: #00c9be;"></div> <div style="z-index: 2; position: absolute; width: 0px; height: 0px; left: 85%; top: 300px;"> <div style="position: relative; line-height: 28px; height: 16px; top: -14px; left: -14px; width: 16px; border-radius: 16px; background: white; padding: 8px;"> <div style="position: relative; line-height: 28px; height: 16px; left: -4px; top: -4px; width: 16px; border-radius: 16px; background: white; border: 4px solid #00c9be; color: #00c9be;"></div></div> </div> <div style="border-radius: 2px; z-index: 1; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 300px; margin-left: 85%; width: 15%; background: #00c9be; color: #00c9be;"><i><a href="/wiki/Paulinella" title="Paulinella"><span style="background: white; padding-left: 4px; padding-right: 4px; color: #00c9be;">Paulinella</span></a></i></div> <div style="border-radius: 2px; position: absolute; line-height: 28px; height: 50px; top: 200px; margin-left: 70%; width: 4px; background: #24d14e; color: #24d14e;"></div> <div style="z-index: 2; position: absolute; width: 0px; height: 0px; left: 70%; top: 250px;"> <div style="position: relative; line-height: 28px; height: 16px; top: -14px; left: -14px; width: 16px; border-radius: 16px; background: white; padding: 8px;"> <div style="position: relative; line-height: 28px; height: 16px; left: -4px; top: -4px; width: 16px; border-radius: 16px; background: white; border: 4px solid #24d150; color: #24d150;"></div></div> </div> <div style="border-radius: 2px; z-index: 1; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 250px; margin-left: 70%; width: 30%; background: #24d150; color: #24d150;"><a href="/wiki/Chlorarachniophyta" title="Chlorarachniophyta"><span style="background-color: rgba(255,255,255,0.8); padding-left: 4px; padding-right: 4px; color: #24d150;">Chlorarachniophyta</span></a></div> <div style="z-index: 0; border-radius: 2px; position: absolute; line-height: 28px; height: 229px; top: 400px; margin-left: 57%; left: 2px; width: 4px; background: #bd3f00; color: #bd3f00;"></div> <div style="border-radius: 2px; z-index: 1; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 625px; margin-left: 57%; width: 13%; left: 2px; background: #bd3f00; color: #bd3f00;"></div> <div style="z-index: 0; border-radius: 2px; position: absolute; line-height: 28px; height: 29px; top: 600px; margin-left: 70%; width: 4px; background: #b13b00; color: #b13b00;"></div> <div style="border-radius: 2px; z-index: 1; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 600px; margin-left: 70%; width: 30%; background: #b13b00; color: #b13b00;"><a href="/wiki/Haptophyta" title="Haptophyta"><span style="background: white; padding-left: 4px; padding-right: 4px; color: #b13b00;">Haptophyta</span></a></div> <div style="z-index: 0; border-radius: 2px; position: absolute; line-height: 28px; height: 29px; top: 625px; margin-left: 70%; width: 4px; background: #ca4300; color: #ca4300;"></div> <div style="border-radius: 2px; z-index: 1; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 650px; margin-left: 70%; width: 30%; background: #ca4300; color: #ca4300;"><a href="/wiki/Cryptomonaden" class="mw-redirect" title="Cryptomonaden"><span style="background: white; padding-left: 4px; padding-right: 4px; color: #ca4300;">Cryptophyta</span></a></div> <div style="border-radius: 2px; z-index: 1; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 400px; margin-left: 57%; width: 43%; background: #99602a; color: #99602a;"><a href="/wiki/Heterokontophyta" title="Heterokontophyta"><span style="background: white; padding-left: 4px; padding-right: 4px; color: #99602a;">Heterokontophyta</span></a></div> <div style="z-index: 0; border-radius: 2px; position: absolute; line-height: 28px; height: 54px; top: 400px; margin-left: 65%; left: -2px; width: 4px; background: #d96f02; color: #d96f02;"></div> <div style="border-radius: 2px; z-index: 1; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 450px; margin-left: 65%; width: 35%; background: #d96f02; color: #d96f02;"><a href="/wiki/Dinoflagellata" class="mw-redirect" title="Dinoflagellata"><span style="background-color: rgba(255,255,255,0.8); padding-left: 4px; padding-right: 4px; color: #d96f02;">Dinoflagellata</span></a></div> <div style="z-index: 0; border-radius: 2px; position: absolute; line-height: 28px; height: 54px; top: 450px; margin-left: 85%; left: -2px; width: 4px; background: #ccc; color: #ccc;"></div> <div style="border-radius: 2px; z-index: 1; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 500px; margin-left: 85%; width: 15%; background: #ccc; color: #aaa;"><a href="/wiki/Apicomplexa" title="Apicomplexa"><span style="background: white; padding-left: 0px; padding-right: 4px; color: #aaa;">Apicomplexa</span></a></div> <div style="z-index: 0; border-radius: 2px; position: absolute; line-height: 28px; height: 104px; top: 450px; margin-left: 75%; left: -2px; width: 4px; background: #ddd; color: #ddd;"></div> <div style="border-radius: 2px; z-index: 1; text-align: right; position: absolute; line-height: 28px; height: 4px; top: 550px; margin-left: 75%; width: 25%; background: #ddd; color: #bbb;"><a href="/wiki/Ciliophora" class="mw-redirect" title="Ciliophora"><span style="background: white; padding-left: 4px; padding-right: 4px; color: #bbb;">Ciliophora</span></a></div> <div style="position: absolute; line-height: 120%; width: 55%; bottom: 0; left: 0;"><b>Kladogramm der möglichen <span style="white-space:nowrap">Chloroplasten-Evolution</span></b> <p><small><sup><b>a</b></sup> Es hat sich mittlerweile herausgestellt, dass die <a href="/wiki/Chromalveolata" title="Chromalveolata">Chromalveolata</a> <a href="/wiki/Paraphyletisch" class="mw-redirect" title="Paraphyletisch">paraphyletisch</a> sind in Bezug auf die <a href="/wiki/Rhizaria" title="Rhizaria">Rhizaria</a>.</small> </p> </div> </div><div style="clear:both;"></div> <p>Bei den <a href="/wiki/Gef%C3%A4%C3%9Fpflanzen" title="Gefäßpflanzen">Gefäßpflanzen</a> unterscheidet man neben den (photosynthetisch aktiven) Chloroplasten weitere Typen von Plastiden: die <a href="/wiki/Gerontoplast" title="Gerontoplast">Gerontoplasten</a> und <a href="/wiki/Etioplast" title="Etioplast">Etioplasten</a> als Entwicklungen der Chloroplasten, sowie den <a href="/wiki/Chromoplast" title="Chromoplast">Chromoplasten</a> und den <a href="/wiki/Leukoplasten" title="Leukoplasten">Leukoplasten</a>, aus dem <a href="/wiki/Amyloplast" title="Amyloplast">Amyloplasten</a>, <a href="/wiki/Elaioplast" class="mw-redirect" title="Elaioplast">Elaioplasten</a> und <a href="/wiki/Proteinoplast" title="Proteinoplast">Proteinoplasten</a> hervorgehen können. <a href="/wiki/Proplastid" title="Proplastid">Proplastid</a> wird jener Vorläufertyp genannt, aus dem sich die Plastiden entwickeln können. </p><p>Die übrigen Algen aus den Evolutionslinien der <a href="/wiki/Stramenopile" title="Stramenopile">Stramenopilen</a>, <a href="/wiki/Haptophyta" title="Haptophyta">Haptophyta</a>, <a href="/wiki/Cryptophyceae" title="Cryptophyceae">Cryptophyceae</a>, <a href="/wiki/Chlorarachniophyta" title="Chlorarachniophyta">Chlorarachniophyta</a> und <a href="/wiki/Euglenozoa" title="Euglenozoa">Euglenozoa</a> bilden komplexe Plastiden. Die Wirtszellen sind nicht mit den obengenannten der Plantae (Rotalgen, Grünalgen, sog. höhere Pflanzen und vermutlich auch Glaucocystophyceen) verwandt, jedoch ihre Plastiden, die höchstwahrscheinlich aus sekundären <a href="/wiki/Endosymbiontentheorie" title="Endosymbiontentheorie">Endosymbiosen</a> herstammen. </p><p>Die photosynthetisch aktiven Vertreter der Euglenozoa (= Euglenida) und die Chlorarachniophyta erhielten ihre Plastiden durch Aufnahme einer Grünalge, enthalten also komplexe Chloroplasten, alle übrigen sind auf Rotalgen zurückzuführen, also komplexe Rhodoplasten. Bei Dinoflagellaten finden sich verschiedene Endosymbiose-Ereignisse von sekundären Endosymbiosen mit Rotalgen, tertiären Endosymbiosen mit Haptophyceen und Cryptophyceen bis zu instabilen <a href="/wiki/Kleptoplastid" title="Kleptoplastid">Kleptoplastiden</a>, die wieder verdaut werden. </p><p>Seit den 1990er-Jahren hat man plastidenähnliche <a href="/wiki/Organell" title="Organell">Zellorganellen</a> auch in verschiedenen <a href="/wiki/Protozoen" title="Protozoen">Protozoen</a>, den <a href="/wiki/Apicomplexa" title="Apicomplexa">Apicomplexa</a>, gefunden. Über die „Apicoplasten“ genannten Zellbestandteile verfügen etwa auch die <a href="/wiki/Malaria" title="Malaria">Malaria</a>-Erreger aus der Gattung <a href="/wiki/Plasmodien" title="Plasmodien">Plasmodium</a>. Nach heutigem Stand des Wissens handelt es sich hierbei um komplexe <a href="/wiki/Rhodoplast" title="Rhodoplast">Rhodoplasten</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Teilung">Teilung</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plastid&amp;veaction=edit&amp;section=3" title="Abschnitt bearbeiten: Teilung" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Plastid&amp;action=edit&amp;section=3" title="Quellcode des Abschnitts bearbeiten: Teilung"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Lange Zeit war unbekannt, wie Plastiden sich teilen und ihre Form verändern. Heute weiß man, dass auch <a href="/wiki/Bakterien" title="Bakterien">Bakterien</a> ein <a href="/wiki/Zytoskelett" class="mw-redirect" title="Zytoskelett">Zytoskelett</a> besitzen, dessen <a href="/wiki/Protein" title="Protein">Proteine</a> <a href="/wiki/Evolution" title="Evolution">evolutionäre</a> Vorläufer des <a href="/wiki/Eukaryoten" title="Eukaryoten">eukaryotischen</a> Zytoskeletts sind. Aus Versuchen am <a href="/wiki/Laubmoos" class="mw-redirect" title="Laubmoos">Laubmoos</a> <i><a href="/wiki/Physcomitrella_patens" class="mw-redirect" title="Physcomitrella patens">Physcomitrella patens</a></i> (unter anderem mit <a href="/wiki/Knockout-Moos" title="Knockout-Moos">Knockout-Moosen</a>) ist bekannt, dass die <a href="/wiki/FtsZ" class="mw-redirect" title="FtsZ">FtsZ</a>-Proteine, <a href="/wiki/Tubulin" class="mw-redirect" title="Tubulin">Tubulin</a>-<a href="/wiki/Homologie_(Genetik)" title="Homologie (Genetik)">Homologe</a>, nicht nur an der Chloroplasten-Teilung beteiligt sind<sup id="cite_ref-René_Strepp(1998)_9-0" class="reference"><a href="#cite_note-René_Strepp(1998)-9"><span class="cite-bracket">&#91;</span>9<span class="cite-bracket">&#93;</span></a></sup>, sondern auch ein komplexes Netzwerk in den Plastiden ausbilden können. Sie erfüllen ähnliche Funktionen wie das Zytoskelett im <a href="/wiki/Cytoplasma" title="Cytoplasma">Cytoplasma</a>.<sup id="cite_ref-Ralf_Reski(2002)_10-0" class="reference"><a href="#cite_note-Ralf_Reski(2002)-10"><span class="cite-bracket">&#91;</span>10<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Kiessling,_J.(2000)_11-0" class="reference"><a href="#cite_note-Kiessling,_J.(2000)-11"><span class="cite-bracket">&#91;</span>11<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Sonstiges">Sonstiges</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plastid&amp;veaction=edit&amp;section=4" title="Abschnitt bearbeiten: Sonstiges" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Plastid&amp;action=edit&amp;section=4" title="Quellcode des Abschnitts bearbeiten: Sonstiges"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li>Ein weiteres primäres Endosymbiose-Ereignis ist mit einem <b>nicht</b>-photosynthetischen cyanobakteriellen Symbionten in der <a href="/wiki/Kieselalgen" title="Kieselalgen">Kieselalgen</a>familie <a href="/wiki/Rhopalodiaceae" title="Rhopalodiaceae">Rhopalodiaceae</a> (Ordnung Rhopalodiales) bekannt.<sup id="cite_ref-Nakayama2017_12-0" class="reference"><a href="#cite_note-Nakayama2017-12"><span class="cite-bracket">&#91;</span>12<span class="cite-bracket">&#93;</span></a></sup></li> <li>Ein weiterer Fall ist das <a href="/wiki/Wimpertierchen" title="Wimpertierchen">Wimpertierchen</a> <i><a href="/w/index.php?title=Pseudoblepharisma&amp;action=edit&amp;redlink=1" class="new" title="Pseudoblepharisma (Seite nicht vorhanden)">Pseudoblepharisma</a> tenue</i><sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">&#91;</span>13<span class="cite-bracket">&#93;</span></a></sup> (Heterotrichea), das neben einem <a href="/wiki/Gr%C3%BCnalgen" class="mw-redirect" title="Grünalgen">Grünalgen</a>-Endosymbionten (<i><a href="/wiki/Chlorella" title="Chlorella">Chlorella</a></i> sp. K10, sekundäre Endosymbiose), auch ein photosynthetisch aktives Bakterium als Endosymbionten hat – dies ist aber kein Cyanobakterium, sondern ein <a href="/wiki/Schwefelpurpurbakterien" class="mw-redirect" title="Schwefelpurpurbakterien">Schwefelpurpurbakterium</a> aus der Familie <a href="/wiki/Chromatiaceae" title="Chromatiaceae">Chromatiaceae</a> (Candidatus <i><a href="/w/index.php?title=Thiodictyon&amp;action=edit&amp;redlink=1" class="new" title="Thiodictyon (Seite nicht vorhanden)">Thiodictyon</a> intracellulare</i><sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">&#91;</span>14<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">&#91;</span>15<span class="cite-bracket">&#93;</span></a></sup>).<sup id="cite_ref-SAMG2021_16-0" class="reference"><a href="#cite_note-SAMG2021-16"><span class="cite-bracket">&#91;</span>16<span class="cite-bracket">&#93;</span></a></sup></li></ul> <div class="mw-heading mw-heading2"><h2 id="Literatur">Literatur</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plastid&amp;veaction=edit&amp;section=5" title="Abschnitt bearbeiten: Literatur" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Plastid&amp;action=edit&amp;section=5" title="Quellcode des Abschnitts bearbeiten: Literatur"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li>Patrick J. Keeling: <i>The endosymbiotic origin, diversification and fate of plastids</i>. In: <i>Philosophical Transactions of the Royal Society B: Biological Sciences.</i> Band 365, Nr. 1541, 2010, S. 729–748, <a class="external mw-magiclink-pmid" rel="nofollow" href="//www.ncbi.nlm.nih.gov/pubmed/20124341?dopt=Abstract">PMID 20124341</a>, <a rel="nofollow" class="external text" href="http://rstb.royalsocietypublishing.org/content/365/1541/729.full.pdf+html">PDF</a> (freier Volltextzugriff, englisch)</li> <li>Przemysław Gagat, Paweł Mackiewicz: <a rel="nofollow" class="external text" href="https://d-nb.info/1120829321/34"><i>Cymbomonas tetramitiformis</i> - a peculiar prasinophyte with a taste for bacteria sheds light on plastid evolution</a>, in: Symbiosis, 10. November 2016, <a href="//doi.org/10.1007/s13199-016-0464-1" class="extiw" title="doi:10.1007/s13199-016-0464-1">doi:10.1007/s13199-016-0464-1</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="Weblinks">Weblinks</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plastid&amp;veaction=edit&amp;section=6" title="Abschnitt bearbeiten: Weblinks" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Plastid&amp;action=edit&amp;section=6" title="Quellcode des Abschnitts bearbeiten: Weblinks"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="sisterproject" style="margin:0.1em 0 0 0;"><div class="noresize noviewer" style="display:inline-block; line-height:10px; min-width:1.6em; text-align:center;" aria-hidden="true" role="presentation"><span class="mw-default-size" typeof="mw:File"><span title="Commons"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/12px-Commons-logo.svg.png" decoding="async" width="12" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/18px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/24px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></span></span></div><b><span class="plainlinks"><a class="external text" href="https://commons.wikimedia.org/wiki/Category:Plastids?uselang=de"><span lang="en">Commons</span>: Plastids</a></span></b>&#160;– Sammlung von Bildern, Videos und Audiodateien</div> <ul><li>Wilfried Probst: <a rel="nofollow" class="external text" href="http://www.wilfried-probst.de/site/tag/exosymbiose/">Frühe Evolution und Symbiose</a>, Europa-Universität Flensburg, Institut für Biologie und Sachunterricht und ihre Didaktik: §Plastiden, abgerufen am 19. April 2019</li></ul> <div class="mw-heading mw-heading2"><h2 id="Einzelnachweise">Einzelnachweise</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Plastid&amp;veaction=edit&amp;section=7" title="Abschnitt bearbeiten: Einzelnachweise" class="mw-editsection-visualeditor"><span>Bearbeiten</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Plastid&amp;action=edit&amp;section=7" title="Quellcode des Abschnitts bearbeiten: Einzelnachweise"><span>Quelltext bearbeiten</span></a><span class="mw-editsection-bracket">]</span></span></div> <ol class="references"> <li id="cite_note-BMCBiology-EvolvinganOrganelle-1"><span class="mw-cite-backlink"><a href="#cite_ref-BMCBiology-EvolvinganOrganelle_1-0">↑</a></span> <span class="reference-text">Takuro&#32;Nakayama,&#32;John M&#32;Archibald&#58; <cite style="font-style:italic">Evolving a photosynthetic organelle</cite>. In: <cite style="font-style:italic">BMC Biology</cite>. 10. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em">&#160;</span>1</span>, 2012, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em">&#160;</span>35</span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1186/1741-7007-10-35">10.1186/1741-7007-10-35</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="//www.ncbi.nlm.nih.gov/pubmed/22531210?dopt=Abstract">PMID 22531210</a>, <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3337241/">PMC&#160;3337241</a> (freier Volltext).<span class="Z3988" title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rfr_id=info:sid/de.wikipedia.org:Plastid&amp;rft.atitle=Evolving+a+photosynthetic+organelle&amp;rft.au=Takuro%26%2332%3BNakayama%2C%26%2332%3BJohn+M%26%2332%3BArchibald&amp;rft.date=2012&amp;rft.doi=10.1186%2F1741-7007-10-35&amp;rft.genre=journal&amp;rft.issue=1&amp;rft.jtitle=BMC+Biology&amp;rft.pages=35&amp;rft.pmc=3337241&amp;rft.pmid=22531210&amp;rft.volume=10.+Jahrgang" style="display:none">&#160;</span></span> </li> <li id="cite_note-MolecularBiologyEvol-EndosymbioticGeneTransfer-2"><span class="mw-cite-backlink"><a href="#cite_ref-MolecularBiologyEvol-EndosymbioticGeneTransfer_2-0">↑</a></span> <span class="reference-text">E. C. M.&#32;Nowack,&#32;H.&#32;Vogel,&#32;M.&#32;Groth,&#32;A. R.&#32;Grossman,&#32;M.&#32;Melkonian,&#32;G.&#32;Glöckner&#58; <cite style="font-style:italic">Endosymbiotic Gene Transfer and Transcriptional Regulation of Transferred Genes in <i>Paulinella chromatophora</i></cite>. In: <cite style="font-style:italic">Molecular Biology and Evolution</cite>. 28. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em">&#160;</span>1</span>, 2010, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em">&#160;</span>407–22</span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1093/molbev%2Fmsq209">10.1093/molbev/msq209</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="//www.ncbi.nlm.nih.gov/pubmed/20702568?dopt=Abstract">PMID 20702568</a>.<span class="Z3988" title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rfr_id=info:sid/de.wikipedia.org:Plastid&amp;rft.atitle=Endosymbiotic+Gene+Transfer+and+Transcriptional+Regulation+of+Transferred+Genes+in+Paulinella+chromatophora&amp;rft.au=E.+C.+M.%26%2332%3BNowack%2C%26%2332%3BH.%26%2332%3BVogel%2C%26%2332%3BM.%26%2332%3BGroth%2C+...&amp;rft.date=2010&amp;rft.doi=10.1093%2Fmolbev%2Fmsq209&amp;rft.genre=journal&amp;rft.issue=1&amp;rft.jtitle=Molecular+Biology+and+Evolution&amp;rft.pages=407-22&amp;rft.pmid=20702568&amp;rft.volume=28.+Jahrgang" style="display:none">&#160;</span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><a href="#cite_ref-3">↑</a></span> <span class="reference-text">David Roy Smith, Robert W. Lee&#58; <cite style="font-style:italic">A Plastid without a Genome: Evidence from the Nonphotosynthetic Green Algal Genus <i>Polytomella</i></cite>. In: <cite style="font-style:italic">Plant Physiology</cite>. <span style="white-space:nowrap">Band<span style="display:inline-block;width:.2em">&#160;</span>164</span>, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em">&#160;</span>4</span>, 1.&#160;April 2014, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em">&#160;</span>1812–1819</span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1104/pp.113.233718">10.1104/pp.113.233718</a></span>, <a class="external mw-magiclink-pmid" rel="nofollow" href="//www.ncbi.nlm.nih.gov/pubmed/24563281?dopt=Abstract">PMID 24563281</a>, <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3982744/">PMC&#160;3982744</a> (freier Volltext).<span class="Z3988" title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rfr_id=info:sid/de.wikipedia.org:Plastid&amp;rft.atitle=A+Plastid+without+a+Genome%3A+Evidence+from+the+Nonphotosynthetic+Green+Algal+Genus+Polytomella&amp;rft.au=David+Roy+Smith%2C+Robert+W.+Lee&amp;rft.date=2014-04-01&amp;rft.doi=10.1104%2Fpp.113.233718&amp;rft.genre=journal&amp;rft.issue=4&amp;rft.jtitle=Plant+Physiology&amp;rft.pages=1812-1819&amp;rft.pmc=3982744&amp;rft.pmid=24563281&amp;rft.volume=164" style="display:none">&#160;</span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><a href="#cite_ref-4">↑</a></span> <span class="reference-text">Jake Buehler: <a rel="nofollow" class="external text" href="https://www.sciencenews.org/article/reeking-parasitic-sapria-plant-lost-body-much-genetic-code">A reeking, parasitic plant lost its body and much of its genetic blueprint</a>, auf: ScienceNews vom 10. Februar 2021.</span> </li> <li id="cite_note-paulinella-5"><span class="mw-cite-backlink"><a href="#cite_ref-paulinella_5-0">↑</a></span> <span class="reference-text"><span class="cite">Luis&#32;Delaye,&#32;Cecilio&#32;Valadez-Cano,&#32;Bernardo&#32;Pérez-Zamorano:&#32;<a rel="nofollow" class="external text" href="http://currents.plos.org/treeoflife/article/how-really-ancient-is-paulinella-chromatophora/"><i>How Really Ancient Is Paulinella Chromatophora?</i></a>&#32;In:&#32;<i>PLoS Currents.</i>&#32;2016<span style="display:none">;</span><span class="Abrufdatum" style="display:none">&#32;abgerufen im 1.&#160;Januar 1</span>&#32;(10.1371/currents.tol.e68a099364bb1a1e129a17b4e06b0c6b).</span><span style="display: none;" class="Z3988" title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Adc&amp;rfr_id=info%3Asid%2Fde.wikipedia.org%3APlastid&amp;rft.title=How+Really+Ancient+Is+Paulinella+Chromatophora%3F&amp;rft.description=How+Really+Ancient+Is+Paulinella+Chromatophora%3F&amp;rft.identifier=&amp;rft.creator=Luis%26%2332%3BDelaye%2C%26%2332%3BCecilio%26%2332%3BValadez-Cano%2C%26%2332%3BBernardo%26%2332%3BP%C3%A9rez-Zamorano&amp;rft.date=2016">&#160;</span></span> </li> <li id="cite_note-:1-6"><span class="mw-cite-backlink"><a href="#cite_ref-:1_6-0">↑</a></span> <span class="reference-text">Purificación&#32;López-García,&#32;Laura&#32;Eme,&#32;David&#32;Moreira&#58; <cite style="font-style:italic">Symbiosis in eukaryotic evolution</cite>. In: <cite style="font-style:italic">Journal of Theoretical Biology</cite>. 434. Jahrgang, 2017, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em">&#160;</span>20–33</span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1016/j.jtbi.2017.02.031">10.1016/j.jtbi.2017.02.031</a></span> (<a rel="nofollow" class="external text" href="http://linkinghub.elsevier.com/retrieve/pii/S0022519317300954">elsevier.com</a>).<span class="Z3988" title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&amp;rfr_id=info:sid/de.wikipedia.org:Plastid&amp;rft.atitle=Symbiosis+in+eukaryotic+evolution&amp;rft.au=Purificaci%C3%B3n%26%2332%3BL%C3%B3pez-Garc%C3%ADa%2C%26%2332%3BLaura%26%2332%3BEme%2C%26%2332%3BDavid%26%2332%3BMoreira&amp;rft.btitle=Journal+of+Theoretical+Biology&amp;rft.date=2017&amp;rft.doi=10.1016%2Fj.jtbi.2017.02.031&amp;rft.genre=book&amp;rft.pages=20-33&amp;rft.volume=434.+Jahrgang" style="display:none">&#160;</span></span> </li> <li id="cite_note-PSanchezB2017-7"><span class="mw-cite-backlink"><a href="#cite_ref-PSanchezB2017_7-0">↑</a></span> <span class="reference-text">Patricia&#32;Sánchez-Baracaldo,&#32;John A.&#32;Raven,&#32;Davide&#32;Pisani,&#32;Andrew H.&#32;Knoll&#58; <cite style="font-style:italic">Early photosynthetic eukaryotes inhabited low-salinity habitats</cite>. In: <cite style="font-style:italic">Proceedings of the National Academy of Sciences</cite>. 114. Jahrgang, <span style="white-space:nowrap">Nr.<span style="display:inline-block;width:.2em">&#160;</span>37</span>, 12.&#160;September 2017, <span style="white-space:nowrap">S.<span style="display:inline-block;width:.2em">&#160;</span>E7737–E7745</span>, <a href="/wiki/Digital_Object_Identifier" title="Digital Object Identifier">doi</a>:<span class="uri-handle" style="white-space:nowrap"><a rel="nofollow" class="external text" href="https://doi.org/10.1073/pnas.1620089114">10.1073/pnas.1620089114</a></span> (<a rel="nofollow" class="external text" href="http://www.pnas.org/lookup/doi/10.1073/pnas.1620089114">pnas.org</a>).<span class="Z3988" title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rfr_id=info:sid/de.wikipedia.org:Plastid&amp;rft.atitle=Early+photosynthetic+eukaryotes+inhabited+low-salinity+habitats&amp;rft.au=Patricia%26%2332%3BS%C3%A1nchez-Baracaldo%2C%26%2332%3BJohn+A.%26%2332%3BRaven%2C%26%2332%3BDavide%26%2332%3BPisani%2C+...&amp;rft.date=2017-09-12&amp;rft.doi=10.1073%2Fpnas.1620089114&amp;rft.genre=journal&amp;rft.issue=37&amp;rft.jtitle=Proceedings+of+the+National+Academy+of+Sciences&amp;rft.pages=E7737-E7745&amp;rft.volume=114.+Jahrgang" style="display:none">&#160;</span></span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><a href="#cite_ref-8">↑</a></span> <span class="reference-text">W Probst, Europa-Universität Flensburg: §Auf dem Weg zur Chloroplastenbildung</span> </li> <li id="cite_note-René_Strepp(1998)-9"><span class="mw-cite-backlink"><a href="#cite_ref-René_Strepp(1998)_9-0">↑</a></span> <span class="reference-text">René Strepp, Sirkka Scholz, Sven Kruse, Volker Speth, <a href="/wiki/Ralf_Reski" title="Ralf Reski">Ralf Reski</a>: <i>Plant nuclear gene knockout reveals a role in plastid division for the homolog of the bacterial cell division protein FtsZ, an ancestral tubulin.</i> In: <i><a href="/wiki/Proceedings_of_the_National_Academy_of_Sciences" class="mw-redirect" title="Proceedings of the National Academy of Sciences">Proceedings of the National Academy of Sciences</a>.</i> Band 95, 1998, S.&#160;4368–4373 <a rel="nofollow" class="external text" href="http://www.pnas.org/content/95/8/4368.abstract">(Abstract)</a>.</span> </li> <li id="cite_note-Ralf_Reski(2002)-10"><span class="mw-cite-backlink"><a href="#cite_ref-Ralf_Reski(2002)_10-0">↑</a></span> <span class="reference-text">Ralf Reski: <i>Rings and networks: the amazing complexity of FtsZ in chloroplasts.</i> In: <i>Trends in Plant Science.</i> Band 7, 2002, S.&#160;103–105 <a rel="nofollow" class="external text" href="http://www.cell.com/trends/plant-science/abstract/S1360-1385(02)02232-X">(Abstract)</a>.</span> </li> <li id="cite_note-Kiessling,_J.(2000)-11"><span class="mw-cite-backlink"><a href="#cite_ref-Kiessling,_J.(2000)_11-0">↑</a></span> <span class="reference-text">J. Kiessling, S. Kruse, S.&#160;A. Rensing, K. Harter, E.&#160;L. Decker, R. Reski: <i>Visualization of a cytoskeleton-like FtsZ network in chloroplasts.</i> In: <i><a href="/wiki/Journal_of_Cell_Biology" title="Journal of Cell Biology">Journal of Cell Biology</a>.</i> Band 151, 2000, S.&#160;945–950 <a rel="nofollow" class="external text" href="http://jcb.rupress.org/cgi/content/abstract/151/4/945">(Abstract)</a>.</span> </li> <li id="cite_note-Nakayama2017-12"><span class="mw-cite-backlink"><a href="#cite_ref-Nakayama2017_12-0">↑</a></span> <span class="reference-text">Takuro Nakayama, Yuji Inagaki: <a rel="nofollow" class="external free" href="https://www.nature.com/articles/s41598-017-13578-8">https://www.nature.com/articles/s41598-017-13578-8</a> Genomic divergence within non-photosynthetic cyanobacterial endosymbionts in rhopalodiacean diatoms], in: <a href="/wiki/Nature" title="Nature">Nature</a> Scientific Reports, Band 7, Nr.&#160;13075, 12. Oktober 2017, <a href="//doi.org/10.1038/s41598-017-13578-8" class="extiw" title="doi:10.1038/s41598-017-13578-8">doi:10.1038/s41598-017-13578-8</a></span> </li> <li id="cite_note-13"><span class="mw-cite-backlink"><a href="#cite_ref-13">↑</a></span> <span class="reference-text">NCBI: <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?id=2528046"><i>Pseudoblepharisma tenue</i> Kahl, 1926</a> (species)</span> </li> <li id="cite_note-14"><span class="mw-cite-backlink"><a href="#cite_ref-14">↑</a></span> <span class="reference-text">NCBI: <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?id=2784338">Thiodictyon endosymbiont of Pseudoblepharisma tenue</a> (species)</span> </li> <li id="cite_note-15"><span class="mw-cite-backlink"><a href="#cite_ref-15">↑</a></span> <span class="reference-text">LPSN: [<a rel="nofollow" class="external free" href="https://lpsn.dsmz.de/species/thiodictyon-syntrophicum">https://lpsn.dsmz.de/species/thiodictyon-syntrophicum</a> "<i>Candidatus</i> Thiodictyon syntrophicum" Peduzzi et al. 2012</span> </li> <li id="cite_note-SAMG2021-16"><span class="mw-cite-backlink"><a href="#cite_ref-SAMG2021_16-0">↑</a></span> <span class="reference-text">Sergio A. Muñoz-Gómez, Martin Kreutz, Sebastian Hess: <a rel="nofollow" class="external text" href="https://advances.sciencemag.org/content/7/24/eabg4102">A microbial eukaryote with a unique combination of purple bacteria and green algae as endosymbionts</a>, in: Science Advances, Band 7, Nr.&#160;24, 11. Juni 2021, eabg4102, <a href="//doi.org/10.1126/sciadv.abg4102" class="extiw" title="doi:10.1126/sciadv.abg4102">doi:10.1126/sciadv.abg4102</a>. Dazu: <ul><li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20210709182319/https://www.eurekalert.org/pub_releases_ml/2021-06/uoc-v061421.php">Ungewöhnliche Symbiose aus Wimpertierchen, Grünalgen und Purpurbakterien</a>, auf: EurekAlert! vom 14. 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href="https://az.wikipedia.org/wiki/Plastidl%C9%99r" title="Plastidlər – Aserbaidschanisch" lang="az" hreflang="az" data-title="Plastidlər" data-language-autonym="Azərbaycanca" data-language-local-name="Aserbaidschanisch" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%9F%D0%BB%D0%B0%D1%81%D1%82%D1%8B%D0%B4%D1%8B" title="Пластыды – Belarussisch" lang="be" hreflang="be" data-title="Пластыды" data-language-autonym="Беларуская" data-language-local-name="Belarussisch" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%9F%D0%BB%D0%B0%D1%81%D1%82%D0%B8%D0%B4" title="Пластид – Bulgarisch" lang="bg" hreflang="bg" data-title="Пластид" data-language-autonym="Български" data-language-local-name="Bulgarisch" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%AA%E0%A7%8D%E0%A6%B2%E0%A6%BE%E0%A6%B8%E0%A7%8D%E0%A6%9F%E0%A6%BF%E0%A6%A1" title="প্লাস্টিড – Bengalisch" lang="bn" hreflang="bn" data-title="প্লাস্টিড" data-language-autonym="বাংলা" data-language-local-name="Bengalisch" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-bs mw-list-item"><a href="https://bs.wikipedia.org/wiki/Plastid" title="Plastid – Bosnisch" lang="bs" hreflang="bs" data-title="Plastid" data-language-autonym="Bosanski" data-language-local-name="Bosnisch" class="interlanguage-link-target"><span>Bosanski</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Plastidi" title="Plastidi – Katalanisch" lang="ca" hreflang="ca" data-title="Plastidi" data-language-autonym="Català" data-language-local-name="Katalanisch" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-ckb mw-list-item"><a href="https://ckb.wikipedia.org/wiki/%D9%BE%D9%84%D8%A7%D8%B3%D8%AA%DB%8C%D8%AF" title="پلاستید – Zentralkurdisch" lang="ckb" hreflang="ckb" data-title="پلاستید" data-language-autonym="کوردی" data-language-local-name="Zentralkurdisch" class="interlanguage-link-target"><span>کوردی</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Plastid" title="Plastid – Tschechisch" lang="cs" hreflang="cs" data-title="Plastid" data-language-autonym="Čeština" data-language-local-name="Tschechisch" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Plastid" title="Plastid – Dänisch" lang="da" hreflang="da" data-title="Plastid" data-language-autonym="Dansk" data-language-local-name="Dänisch" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%A0%CE%BB%CE%B1%CF%83%CF%84%CE%AF%CE%B4%CE%B9%CE%BF" title="Πλαστίδιο – Griechisch" lang="el" hreflang="el" data-title="Πλαστίδιο" data-language-autonym="Ελληνικά" data-language-local-name="Griechisch" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Plastid" title="Plastid – Englisch" lang="en" hreflang="en" data-title="Plastid" data-language-autonym="English" data-language-local-name="Englisch" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Plastido" title="Plastido – Esperanto" lang="eo" hreflang="eo" data-title="Plastido" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Plasto" title="Plasto – Spanisch" lang="es" hreflang="es" data-title="Plasto" data-language-autonym="Español" data-language-local-name="Spanisch" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Plastiidid" title="Plastiidid – Estnisch" lang="et" hreflang="et" data-title="Plastiidid" data-language-autonym="Eesti" data-language-local-name="Estnisch" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Plastidio" title="Plastidio – Baskisch" lang="eu" hreflang="eu" data-title="Plastidio" data-language-autonym="Euskara" data-language-local-name="Baskisch" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%AF%DB%8C%D8%B3%D9%87" title="دیسه – Persisch" lang="fa" hreflang="fa" data-title="دیسه" data-language-autonym="فارسی" data-language-local-name="Persisch" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Plastidi" title="Plastidi – Finnisch" lang="fi" hreflang="fi" data-title="Plastidi" data-language-autonym="Suomi" data-language-local-name="Finnisch" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Plaste" title="Plaste – Französisch" lang="fr" hreflang="fr" data-title="Plaste" data-language-autonym="Français" data-language-local-name="Französisch" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Plasto" title="Plasto – Galicisch" lang="gl" hreflang="gl" data-title="Plasto" data-language-autonym="Galego" data-language-local-name="Galicisch" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-gv mw-list-item"><a href="https://gv.wikipedia.org/wiki/Plastaid" title="Plastaid – Manx" lang="gv" hreflang="gv" data-title="Plastaid" data-language-autonym="Gaelg" data-language-local-name="Manx" class="interlanguage-link-target"><span>Gaelg</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%A4%D7%9C%D7%A1%D7%98%D7%99%D7%93%D7%94" title="פלסטידה – Hebräisch" lang="he" hreflang="he" data-title="פלסטידה" data-language-autonym="עברית" data-language-local-name="Hebräisch" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%B2%E0%A4%B5%E0%A4%95" title="लवक – Hindi" lang="hi" hreflang="hi" data-title="लवक" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Plastid" title="Plastid – Kroatisch" lang="hr" hreflang="hr" data-title="Plastid" data-language-autonym="Hrvatski" data-language-local-name="Kroatisch" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Sz%C3%ADntest" title="Színtest – Ungarisch" lang="hu" hreflang="hu" data-title="Színtest" data-language-autonym="Magyar" data-language-local-name="Ungarisch" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-hy mw-list-item"><a href="https://hy.wikipedia.org/wiki/%D5%8A%D5%AC%D5%A1%D5%BD%D5%BF%D5%AB%D5%A4%D5%B6%D5%A5%D6%80" title="Պլաստիդներ – Armenisch" lang="hy" hreflang="hy" data-title="Պլաստիդներ" data-language-autonym="Հայերեն" data-language-local-name="Armenisch" class="interlanguage-link-target"><span>Հայերեն</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Plastid" title="Plastid – Indonesisch" lang="id" hreflang="id" data-title="Plastid" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesisch" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Plastido" title="Plastido – Ido" lang="io" hreflang="io" data-title="Plastido" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-is mw-list-item"><a href="https://is.wikipedia.org/wiki/Plast%C3%AD%C3%B0" title="Plastíð – Isländisch" lang="is" hreflang="is" data-title="Plastíð" data-language-autonym="Íslenska" data-language-local-name="Isländisch" class="interlanguage-link-target"><span>Íslenska</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Plastidio" title="Plastidio – Italienisch" lang="it" hreflang="it" data-title="Plastidio" data-language-autonym="Italiano" data-language-local-name="Italienisch" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E8%89%B2%E7%B4%A0%E4%BD%93" title="色素体 – Japanisch" lang="ja" hreflang="ja" data-title="色素体" data-language-autonym="日本語" data-language-local-name="Japanisch" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-jv mw-list-item"><a href="https://jv.wikipedia.org/wiki/Plastida" title="Plastida – Javanisch" lang="jv" hreflang="jv" data-title="Plastida" data-language-autonym="Jawa" data-language-local-name="Javanisch" class="interlanguage-link-target"><span>Jawa</span></a></li><li class="interlanguage-link interwiki-ka mw-list-item"><a href="https://ka.wikipedia.org/wiki/%E1%83%9E%E1%83%9A%E1%83%90%E1%83%A1%E1%83%A2%E1%83%98%E1%83%93%E1%83%98" title="პლასტიდი – Georgisch" lang="ka" hreflang="ka" data-title="პლასტიდი" data-language-autonym="ქართული" data-language-local-name="Georgisch" class="interlanguage-link-target"><span>ქართული</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%9F%D0%BB%D0%B0%D1%81%D1%82%D0%B8%D0%B4%D1%82%D0%B5%D1%80" title="Пластидтер – Kasachisch" lang="kk" hreflang="kk" data-title="Пластидтер" data-language-autonym="Қазақша" data-language-local-name="Kasachisch" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-kn mw-list-item"><a href="https://kn.wikipedia.org/wiki/%E0%B2%AA%E0%B3%8D%E0%B2%B2%E0%B2%BE%E0%B2%B8%E0%B3%8D%E0%B2%9F%E0%B2%BF%E0%B2%A1%E0%B3%8D" title="ಪ್ಲಾಸ್ಟಿಡ್ – Kannada" lang="kn" hreflang="kn" data-title="ಪ್ಲಾಸ್ಟಿಡ್" data-language-autonym="ಕನ್ನಡ" data-language-local-name="Kannada" class="interlanguage-link-target"><span>ಕನ್ನಡ</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EC%83%89%EC%86%8C%EC%B2%B4" title="색소체 – Koreanisch" lang="ko" hreflang="ko" data-title="색소체" data-language-autonym="한국어" data-language-local-name="Koreanisch" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-ku mw-list-item"><a href="https://ku.wikipedia.org/wiki/Plast%C3%AEd" title="Plastîd – Kurdisch" lang="ku" hreflang="ku" data-title="Plastîd" data-language-autonym="Kurdî" data-language-local-name="Kurdisch" class="interlanguage-link-target"><span>Kurdî</span></a></li><li class="interlanguage-link interwiki-ky mw-list-item"><a href="https://ky.wikipedia.org/wiki/%D0%9F%D0%BB%D0%B0%D1%81%D1%82%D0%B8%D0%B4%D0%B0%D0%BB%D0%B0%D1%80" title="Пластидалар – Kirgisisch" lang="ky" hreflang="ky" data-title="Пластидалар" data-language-autonym="Кыргызча" data-language-local-name="Kirgisisch" class="interlanguage-link-target"><span>Кыргызча</span></a></li><li class="interlanguage-link interwiki-lb mw-list-item"><a href="https://lb.wikipedia.org/wiki/Plast" title="Plast – Luxemburgisch" lang="lb" hreflang="lb" data-title="Plast" data-language-autonym="Lëtzebuergesch" data-language-local-name="Luxemburgisch" class="interlanguage-link-target"><span>Lëtzebuergesch</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Plastid%C4%97" title="Plastidė – Litauisch" lang="lt" hreflang="lt" data-title="Plastidė" data-language-autonym="Lietuvių" data-language-local-name="Litauisch" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-lv mw-list-item"><a href="https://lv.wikipedia.org/wiki/Plast%C4%ABdas" title="Plastīdas – Lettisch" lang="lv" hreflang="lv" data-title="Plastīdas" data-language-autonym="Latviešu" data-language-local-name="Lettisch" class="interlanguage-link-target"><span>Latviešu</span></a></li><li class="interlanguage-link interwiki-mk mw-list-item"><a href="https://mk.wikipedia.org/wiki/%D0%9F%D0%BB%D0%B0%D1%81%D1%82%D0%B8%D0%B4" title="Пластид – Mazedonisch" lang="mk" hreflang="mk" data-title="Пластид" data-language-autonym="Македонски" data-language-local-name="Mazedonisch" class="interlanguage-link-target"><span>Македонски</span></a></li><li class="interlanguage-link interwiki-ml mw-list-item"><a href="https://ml.wikipedia.org/wiki/%E0%B4%AA%E0%B5%8D%E0%B4%B2%E0%B4%BE%E0%B4%B8%E0%B5%8D%E0%B4%B1%E0%B5%8D%E0%B4%B1%E0%B4%BF%E0%B4%A1%E0%B5%8D" title="പ്ലാസ്റ്റിഡ് – Malayalam" lang="ml" hreflang="ml" data-title="പ്ലാസ്റ്റിഡ്" data-language-autonym="മലയാളം" data-language-local-name="Malayalam" class="interlanguage-link-target"><span>മലയാളം</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Plastiden" title="Plastiden – Niederdeutsch" lang="nds" hreflang="nds" data-title="Plastiden" data-language-autonym="Plattdüütsch" data-language-local-name="Niederdeutsch" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Plastide" title="Plastide – Niederländisch" lang="nl" hreflang="nl" data-title="Plastide" data-language-autonym="Nederlands" data-language-local-name="Niederländisch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-om mw-list-item"><a href="https://om.wikipedia.org/wiki/Pilaastiidii" title="Pilaastiidii – Oromo" lang="om" hreflang="om" data-title="Pilaastiidii" data-language-autonym="Oromoo" data-language-local-name="Oromo" class="interlanguage-link-target"><span>Oromoo</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Plastyd" title="Plastyd – Polnisch" lang="pl" hreflang="pl" data-title="Plastyd" data-language-autonym="Polski" data-language-local-name="Polnisch" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%D9%BE%D9%84%D8%A7%D8%B3%D9%B9%DA%88%D8%B2" title="پلاسٹڈز – Westliches Panjabi" lang="pnb" hreflang="pnb" data-title="پلاسٹڈز" data-language-autonym="پنجابی" data-language-local-name="Westliches Panjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Plast%C3%ADdeo" title="Plastídeo – Portugiesisch" lang="pt" hreflang="pt" data-title="Plastídeo" data-language-autonym="Português" data-language-local-name="Portugiesisch" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Plastid%C4%83" title="Plastidă – Rumänisch" lang="ro" hreflang="ro" data-title="Plastidă" data-language-autonym="Română" data-language-local-name="Rumänisch" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9F%D0%BB%D0%B0%D1%81%D1%82%D0%B8%D0%B4%D1%8B" title="Пластиды – Russisch" lang="ru" hreflang="ru" data-title="Пластиды" data-language-autonym="Русский" data-language-local-name="Russisch" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/Plastid" title="Plastid – Serbokroatisch" lang="sh" hreflang="sh" data-title="Plastid" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="Serbokroatisch" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Plastid" title="Plastid – einfaches Englisch" lang="en-simple" hreflang="en-simple" data-title="Plastid" data-language-autonym="Simple English" data-language-local-name="einfaches Englisch" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Plastid" title="Plastid – Slowakisch" lang="sk" hreflang="sk" data-title="Plastid" data-language-autonym="Slovenčina" data-language-local-name="Slowakisch" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Plastid" title="Plastid – Slowenisch" lang="sl" hreflang="sl" data-title="Plastid" data-language-autonym="Slovenščina" data-language-local-name="Slowenisch" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%9F%D0%BB%D0%B0%D1%81%D1%82%D0%B8%D0%B4" title="Пластид – Serbisch" lang="sr" hreflang="sr" data-title="Пластид" data-language-autonym="Српски / srpski" data-language-local-name="Serbisch" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Plastid" title="Plastid – Schwedisch" lang="sv" hreflang="sv" data-title="Plastid" data-language-autonym="Svenska" data-language-local-name="Schwedisch" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%95%E0%AE%A3%E0%AE%BF%E0%AE%95%E0%AE%99%E0%AF%8D%E0%AE%95%E0%AE%B3%E0%AF%8D" title="கணிகங்கள் – Tamil" lang="ta" hreflang="ta" data-title="கணிகங்கள்" data-language-autonym="தமிழ்" data-language-local-name="Tamil" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-th mw-list-item"><a href="https://th.wikipedia.org/wiki/%E0%B8%9E%E0%B8%A5%E0%B8%B2%E0%B8%AA%E0%B8%95%E0%B8%B4%E0%B8%94" title="พลาสติด – Thailändisch" lang="th" hreflang="th" data-title="พลาสติด" data-language-autonym="ไทย" data-language-local-name="Thailändisch" class="interlanguage-link-target"><span>ไทย</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Plastit" title="Plastit – Türkisch" lang="tr" hreflang="tr" data-title="Plastit" data-language-autonym="Türkçe" data-language-local-name="Türkisch" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%9F%D0%BB%D0%B0%D1%81%D1%82%D0%B8%D0%B4%D0%B0" title="Пластида – Ukrainisch" lang="uk" hreflang="uk" data-title="Пластида" data-language-autonym="Українська" data-language-local-name="Ukrainisch" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/Plastidalar" title="Plastidalar – Usbekisch" lang="uz" hreflang="uz" data-title="Plastidalar" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Usbekisch" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/L%E1%BA%A1p_th%E1%BB%83" title="Lạp thể – Vietnamesisch" lang="vi" hreflang="vi" data-title="Lạp thể" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamesisch" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-wuu mw-list-item"><a href="https://wuu.wikipedia.org/wiki/%E8%89%B2%E7%B4%A0%E4%BD%93" title="色素体 – Wu" lang="wuu" hreflang="wuu" data-title="色素体" data-language-autonym="吴语" data-language-local-name="Wu" class="interlanguage-link-target"><span>吴语</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E8%89%B2%E7%B4%A0%E9%AB%94" title="色素體 – Chinesisch" lang="zh" hreflang="zh" data-title="色素體" data-language-autonym="中文" data-language-local-name="Chinesisch" class="interlanguage-link-target"><span>中文</span></a></li><li class="interlanguage-link interwiki-zh-min-nan mw-list-item"><a href="https://zh-min-nan.wikipedia.org/wiki/Sek-s%C3%B2%CD%98-th%C3%A9" title="Sek-sò͘-thé – Min Nan" lang="nan" hreflang="nan" data-title="Sek-sò͘-thé" data-language-autonym="閩南語 / Bân-lâm-gú" data-language-local-name="Min Nan" class="interlanguage-link-target"><span>閩南語 / Bân-lâm-gú</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E8%89%B2%E7%B4%A0%E9%AB%94" title="色素體 – Kantonesisch" lang="yue" hreflang="yue" data-title="色素體" data-language-autonym="粵語" data-language-local-name="Kantonesisch" class="interlanguage-link-target"><span>粵語</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q165963#sitelinks-wikipedia" title="Links auf Artikel in anderen Sprachen bearbeiten" class="wbc-editpage">Links bearbeiten</a></span></div> </div> </nav> </div> </div> <footer id="footer" class="mw-footer" > 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