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Gecko - Wikipedia

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Available in 25 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-25" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">25 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D8%A3%D8%A8%D9%88_%D8%A8%D8%B1%D9%8A%D8%B5" title="أبو بريص – Arabic" lang="ar" hreflang="ar" data-title="أبو بريص" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-ban mw-list-item"><a href="https://ban.wikipedia.org/wiki/Cekcek" title="Cekcek – Balinese" lang="ban" hreflang="ban" data-title="Cekcek" data-language-autonym="Basa Bali" data-language-local-name="Balinese" class="interlanguage-link-target"><span>Basa Bali</span></a></li><li class="interlanguage-link interwiki-bcl mw-list-item"><a href="https://bcl.wikipedia.org/wiki/Burubuaya" title="Burubuaya – Central Bikol" lang="bcl" hreflang="bcl" data-title="Burubuaya" data-language-autonym="Bikol Central" data-language-local-name="Central Bikol" class="interlanguage-link-target"><span>Bikol Central</span></a></li><li class="interlanguage-link interwiki-nv mw-list-item"><a href="https://nv.wikipedia.org/wiki/N%C3%A1%C3%A1%CA%BCdei%C5%82naadii_d%C3%B3%C3%B3_bik%CA%BC%C3%A9%C3%AD" title="Nááʼdeiłnaadii dóó bikʼéí – Navajo" lang="nv" hreflang="nv" data-title="Nááʼdeiłnaadii dóó bikʼéí" data-language-autonym="Diné bizaad" data-language-local-name="Navajo" class="interlanguage-link-target"><span>Diné bizaad</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Gekuloj" title="Gekuloj – Esperanto" lang="eo" hreflang="eo" data-title="Gekuloj" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%DA%AF%DA%A9%D9%88" title="گکو – Persian" lang="fa" hreflang="fa" data-title="گکو" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-guw mw-list-item"><a href="https://guw.wikipedia.org/wiki/H%E1%BB%8Dm%E1%BA%B9s%C3%AC" title="Họmẹsì – Gun" lang="guw" hreflang="guw" data-title="Họmẹsì" data-language-autonym="Gungbe" data-language-local-name="Gun" class="interlanguage-link-target"><span>Gungbe</span></a></li><li class="interlanguage-link interwiki-ha badge-Q17559452 badge-recommendedarticle mw-list-item" title="recommended article"><a href="https://ha.wikipedia.org/wiki/Tsaka" title="Tsaka – Hausa" lang="ha" hreflang="ha" data-title="Tsaka" data-language-autonym="Hausa" data-language-local-name="Hausa" class="interlanguage-link-target"><span>Hausa</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%97%E0%A5%87%E0%A4%95%E0%A5%8D%E0%A4%95%E0%A5%8B" title="गेक्को – Hindi" lang="hi" hreflang="hi" data-title="गेक्को" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Cecak" title="Cecak – Indonesian" lang="id" hreflang="id" data-title="Cecak" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Mjusi-kafiri" title="Mjusi-kafiri – Swahili" lang="sw" hreflang="sw" data-title="Mjusi-kafiri" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-mg mw-list-item"><a href="https://mg.wikipedia.org/wiki/Katsatsaka" title="Katsatsaka – Malagasy" lang="mg" hreflang="mg" data-title="Katsatsaka" data-language-autonym="Malagasy" data-language-local-name="Malagasy" class="interlanguage-link-target"><span>Malagasy</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%A8%D8%B1%D8%B5" title="برص – Egyptian Arabic" lang="arz" hreflang="arz" data-title="برص" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-ne mw-list-item"><a href="https://ne.wikipedia.org/wiki/%E0%A4%AE%E0%A4%BE%E0%A4%89%E0%A4%B8%E0%A5%81%E0%A4%B2%E0%A5%80" title="माउसुली – Nepali" lang="ne" hreflang="ne" data-title="माउसुली" data-language-autonym="नेपाली" data-language-local-name="Nepali" class="interlanguage-link-target"><span>नेपाली</span></a></li><li class="interlanguage-link interwiki-sq mw-list-item"><a href="https://sq.wikipedia.org/wiki/Gekoja" title="Gekoja – Albanian" lang="sq" hreflang="sq" data-title="Gekoja" data-language-autonym="Shqip" data-language-local-name="Albanian" class="interlanguage-link-target"><span>Shqip</span></a></li><li class="interlanguage-link interwiki-kab mw-list-item"><a href="https://kab.wikipedia.org/wiki/Tasallast" title="Tasallast – Kabyle" lang="kab" hreflang="kab" data-title="Tasallast" data-language-autonym="Taqbaylit" data-language-local-name="Kabyle" class="interlanguage-link-target"><span>Taqbaylit</span></a></li><li class="interlanguage-link interwiki-uk badge-Q70894304 mw-list-item" title=""><a href="https://uk.wikipedia.org/wiki/%D0%93%D0%B5%D0%BA%D0%BE%D0%BD" title="Гекон – Ukrainian" lang="uk" hreflang="uk" data-title="Гекон" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%DA%AF%DB%8C%DA%A9%D9%88" title="گیکو – Urdu" lang="ur" hreflang="ur" data-title="گیکو" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-war mw-list-item"><a href="https://war.wikipedia.org/wiki/Tuko" title="Tuko – Waray" lang="war" hreflang="war" data-title="Tuko" data-language-autonym="Winaray" data-language-local-name="Waray" class="interlanguage-link-target"><span>Winaray</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/%E1%BB%8Cm%E1%BB%8D_on%C3%ADl%C3%A9" title="Ọmọ onílé – Yoruba" lang="yo" hreflang="yo" data-title="Ọmọ onílé" data-language-autonym="Yorùbá" data-language-local-name="Yoruba" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E5%AE%88%E5%AE%AE" title="守宮 – Chinese" lang="zh" hreflang="zh" data-title="守宮" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li><li class="interlanguage-link interwiki-bbc mw-list-item"><a href="https://bbc.wikipedia.org/wiki/Boraspati" title="Boraspati – Batak Toba" lang="bbc" hreflang="bbc" data-title="Boraspati" data-language-autonym="Batak Toba" data-language-local-name="Batak Toba" class="interlanguage-link-target"><span>Batak Toba</span></a></li><li class="interlanguage-link interwiki-bew mw-list-item"><a href="https://bew.wikipedia.org/wiki/Cecek" title="Cecek – Betawi" lang="bew" hreflang="bew" data-title="Cecek" data-language-autonym="Betawi" data-language-local-name="Betawi" class="interlanguage-link-target"><span>Betawi</span></a></li><li class="interlanguage-link interwiki-fon mw-list-item"><a href="https://fon.wikipedia.org/wiki/X%C9%94m%C9%9Bsi" title="Xɔmɛsi – Fon" lang="fon" hreflang="fon" data-title="Xɔmɛsi" data-language-autonym="Fɔ̀ngbè" data-language-local-name="Fon" class="interlanguage-link-target"><span>Fɔ̀ngbè</span></a></li><li class="interlanguage-link interwiki-kge mw-list-item"><a href="https://kge.wikipedia.org/wiki/Kicak" title="Kicak – Komering" lang="kge" hreflang="kge" data-title="Kicak" data-language-autonym="Kumoring" data-language-local-name="Komering" class="interlanguage-link-target"><span>Kumoring</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a 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navigation-not-searchable">This article is about the type of reptile. For other uses, see <a href="/wiki/Gecko_(disambiguation)" class="mw-disambig" title="Gecko (disambiguation)">Gecko (disambiguation)</a>.</div> <p class="mw-empty-elt"> </p> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">Gecko<br /><div style="font-size: 85%;">Temporal range: <span class="noprint"><span style="display:inline-block;"></span><span style="display:inline-block;">100–0&#160;<a href="/wiki/Megaannum" class="mw-redirect" title="Megaannum">Ma</a></span> <span style="display:inline-block;"></span><div id="Timeline-row" style="margin: 4px auto 0; clear:both; width:220px; padding:0px; height:18px; overflow:visible; white-space:nowrap; border:1px #666; border-style:solid none; position:relative; z-index:0; font-size:97%;"> <div style="position:absolute; height:100%; left:0px; width:207.23076923077px; padding-left:5px; text-align:left; background-color:rgb(254,217,106); background-image: linear-gradient(to right, rgba(255,255,255,1), rgba(254,217,106,1) 15%, rgba(254,217,106,1));"><a href="/wiki/Precambrian" title="Precambrian">PreꞒ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,160,86); left:37.636923076923px; width:17.583076923077px;"><a href="/wiki/Cambrian" title="Cambrian">Ꞓ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(0,146,112); left:55.22px; width:14.807692307692px;"><a href="/wiki/Ordovician" title="Ordovician">O</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(179,225,182); left:70.027692307692px; width:7.9470769230769px;"><a href="/wiki/Silurian" title="Silurian">S</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(203,140,55); left:77.974769230769px; width:20.564923076923px;"><a href="/wiki/Devonian" title="Devonian">D</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(103,165,153); left:98.539692307692px; width:20.294153846154px;"><a href="/wiki/Carboniferous" title="Carboniferous">C</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(240,64,40); left:118.83384615385px; width:15.907015384615px;"><a href="/wiki/Permian" title="Permian">P</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(129,43,146); left:134.74086153846px; width:17.092984615385px;"><a href="/wiki/Triassic" title="Triassic">T</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(52,178,201); left:151.83384615385px; width:19.732307692308px;"><a href="/wiki/Jurassic" title="Jurassic">J</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,198,78); left:171.56615384615px; width:26.095384615385px;"><a href="/wiki/Cretaceous" title="Cretaceous">K</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(253,154,82); left:197.66153846154px; width:14.540307692308px;"><a href="/wiki/Paleogene" title="Paleogene">Pg</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(255,230,25); left:212.20184615385px; width:6.9249230769231px;"><a href="/wiki/Neogene" title="Neogene">N</a></div> <div id="end-border" style="position:absolute; height:100%; background-color:#666; width:1px; left:219px"></div><div style="margin:0 auto; line-height:0; clear:both; width:220px; padding:0px; height:8px; overflow:visible; background-color:transparent; position:relative; top:-4px; z-index:100;"><div style="position:absolute; height:8px; left:181.75384615385px; width:38.246153846154px; background-color:#360; opacity:0.42;"></div> <div style="position:absolute; height:8px; left:186.15384615385px; width:33.846153846154px; background-color:#360; opacity:1;"></div> <div style="position:absolute; height:6px; top:1px; left:187.15384615385px; width:31.846153846154px; background-color:#6c3;"></div> </div> </div></span><a href="/wiki/Cenomanian" title="Cenomanian">Cenomanian</a> – <a href="/wiki/Holocene" title="Holocene">present</a><br />Stem-groups present since <a href="/wiki/Aptian" title="Aptian">Aptian</a>-<a href="/wiki/Albian" title="Albian">Albian</a><sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup></div> </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Phelsuma_l._laticauda.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c3/Phelsuma_l._laticauda.jpg/250px-Phelsuma_l._laticauda.jpg" decoding="async" width="220" height="270" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c3/Phelsuma_l._laticauda.jpg/330px-Phelsuma_l._laticauda.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c3/Phelsuma_l._laticauda.jpg/500px-Phelsuma_l._laticauda.jpg 2x" data-file-width="600" data-file-height="737" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%"><a href="/wiki/Gold_dust_day_gecko" title="Gold dust day gecko">Gold dust day gecko</a> </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Gekkota" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Chordate" title="Chordate">Chordata</a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Reptile" title="Reptile">Reptilia</a> </td></tr> <tr> <td>Order: </td> <td><a href="/wiki/Squamata" title="Squamata">Squamata</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Gekkonomorpha" title="Gekkonomorpha">Gekkonomorpha</a> </td></tr> <tr> <td>Infraorder: </td> <td><a class="mw-selflink selflink">Gekkota</a><br /><small><a href="/wiki/Georges_Cuvier" title="Georges Cuvier">Cuvier</a>, 1817</small> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)">Subgroups </th></tr> <tr> <td colspan="2" style="text-align: left"> <ul><li><b><a href="/wiki/Pygopodoidea" title="Pygopodoidea">Pygopodoidea</a></b> <ul><li><a href="/wiki/Carphodactylidae" title="Carphodactylidae">Carphodactylidae</a></li> <li><a href="/wiki/Diplodactylidae" title="Diplodactylidae">Diplodactylidae</a></li> <li><a href="/wiki/Pygopodidae" title="Pygopodidae">Pygopodidae</a></li></ul></li> <li><b>Gekkonoidea</b> <ul><li><a href="/wiki/Eublepharidae" title="Eublepharidae">Eublepharidae</a></li> <li><a href="/wiki/Gekkonidae" title="Gekkonidae">Gekkonidae</a></li> <li><a href="/wiki/Phyllodactylidae" title="Phyllodactylidae">Phyllodactylidae</a></li> <li><a href="/wiki/Sphaerodactylidae" title="Sphaerodactylidae">Sphaerodactylidae</a></li></ul></li></ul> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><b>Geckos</b> are small, mostly carnivorous <a href="/wiki/Lizard" title="Lizard">lizards</a> that have a wide distribution, found on every continent except <a href="/wiki/Antarctica" title="Antarctica">Antarctica</a>. Belonging to the <a href="/wiki/Infraorder#Hierarchy_of_ranks" class="mw-redirect" title="Infraorder">infraorder</a> <b>Gekkota</b>, geckos are found in warm <a href="/wiki/Climate" title="Climate">climates</a> throughout the world. They range from 1.6 to 60 <a href="/wiki/Centimetre" title="Centimetre">centimetres</a> (0.6 to 23.6 <a href="/wiki/Inch" title="Inch">inches</a>). </p><p>Geckos are unique among lizards for their <a href="/wiki/Animal_communication" title="Animal communication">vocalisations</a>, which differ from species to species. Most geckos in the family <a href="/wiki/Gekkonidae" title="Gekkonidae">Gekkonidae</a> use chirping or clicking sounds in their social interactions. <a href="/wiki/Tokay_gecko" title="Tokay gecko">Tokay geckos</a> (<i>Gekko gecko</i>) are known for their loud <a href="/wiki/Mating_call" title="Mating call">mating calls</a>, and some other species are capable of making hissing noises when alarmed or threatened. They are the most species-rich group of lizards, with about 1,500 different species worldwide.<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> </p><p>All geckos, except species in the family <a href="/wiki/Eublepharidae" title="Eublepharidae">Eublepharidae</a> lack eyelids; instead, the outer surface of the eyeball has a <a href="/wiki/Transparency_and_translucency" title="Transparency and translucency">transparent</a> membrane, the <a href="/wiki/Brille" title="Brille">brille</a>. They have a fixed <a href="/wiki/Lens_(anatomy)" class="mw-redirect" title="Lens (anatomy)">lens</a> within each <a href="/wiki/Iris_(anatomy)" title="Iris (anatomy)">iris</a> that enlarges in darkness to let in more light. Since they cannot <a href="/wiki/Blink" class="mw-redirect" title="Blink">blink</a>, species without eyelids generally lick their own brilles when they need to clear them of dust and dirt, in order to keep them clean and moist.<sup id="cite_ref-Badger_Lizards_3-0" class="reference"><a href="#cite_note-Badger_Lizards-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> </p><p>Unlike most lizards, geckos are usually <a href="/wiki/Nocturnal" class="mw-redirect" title="Nocturnal">nocturnal</a><sup id="cite_ref-Into_the_light:_Diurnality_has_evol_4-0" class="reference"><a href="#cite_note-Into_the_light:_Diurnality_has_evol-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> and have excellent <a href="/wiki/Night_vision" title="Night vision">night vision</a>; their <a href="/wiki/Colour_vision" class="mw-redirect" title="Colour vision">colour vision</a> in low light is 350 times more sensitive than <a href="/wiki/Human_eye" title="Human eye">human eyes</a>.<sup id="cite_ref-Roth_Pupils_5-0" class="reference"><a href="#cite_note-Roth_Pupils-5"><span class="cite-bracket">&#91;</span>5<span class="cite-bracket">&#93;</span></a></sup> The nocturnal geckos evolved from <a href="/wiki/Diurnality" title="Diurnality">diurnal</a> species, which had lost the <a href="/wiki/Rod_cell" title="Rod cell">rod cells</a> from their eyes. The gecko eye, therefore, modified its <a href="/wiki/Cone_cell" title="Cone cell">cone cells</a> that increased in size into different types, both single and double. Three different photo-pigments have been retained, and are sensitive to ultraviolet, blue, and green. They also use a multifocal optical system that allows them to generate a sharp image for at least two different depths.<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">&#91;</span>7<span class="cite-bracket">&#93;</span></a></sup> While most gecko species are nocturnal, some species are diurnal and active during the day, which have evolved multiple times independently.<sup id="cite_ref-Into_the_light:_Diurnality_has_evol_4-1" class="reference"><a href="#cite_note-Into_the_light:_Diurnality_has_evol-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> </p><p>Many species are well known for their specialised toe pads, which enable them to grab and climb onto smooth and vertical surfaces, and even cross indoor ceilings with ease. Geckos are well known to people who live in warm regions of the world, where several species make their home inside human habitations. These, for example the <a href="/wiki/Hemidactylus" title="Hemidactylus">house gecko</a>, become part of the indoor menagerie and are often welcomed, as they feed on <a href="/wiki/Insect" title="Insect">insect</a> <a href="/wiki/Pest_(organism)" title="Pest (organism)">pests</a>; including <a href="/wiki/Moth" title="Moth">moths</a> and <a href="/wiki/Mosquito" title="Mosquito">mosquitoes</a>. Like most lizards, geckos can lose their tails in defence, a process called <a href="/wiki/Autotomy" title="Autotomy">autotomy</a>; the predator may attack the wriggling tail, allowing the gecko to escape.<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">&#91;</span>8<span class="cite-bracket">&#93;</span></a></sup> </p><p>The largest species, <i><a href="/wiki/Gigarcanum_delcourti" class="mw-redirect" title="Gigarcanum delcourti">Gigarcanum delcourti</a></i>, is only known from a single, stuffed specimen probably collected in the 19th century found in the basement of the <a href="/wiki/Mus%C3%A9um_d%27histoire_naturelle_de_Marseille" title="Muséum d&#39;histoire naturelle de Marseille">Natural History Museum of Marseille</a> in <a href="/wiki/Marseille" title="Marseille">Marseille</a>, France. This gecko was 600 <a href="/wiki/Millimetre" title="Millimetre">millimetres</a> (24 <a href="/wiki/Inch" title="Inch">inches</a>) long, and it was likely <a href="/wiki/Endemic" class="mw-redirect" title="Endemic">endemic</a> to <a href="/wiki/New_Caledonia" title="New Caledonia">New Caledonia</a>, where it lived in native forests.<sup id="cite_ref-:1_9-0" class="reference"><a href="#cite_note-:1-9"><span class="cite-bracket">&#91;</span>9<span class="cite-bracket">&#93;</span></a></sup> The smallest gecko, the <a href="/wiki/Sphaerodactylus_ariasae" title="Sphaerodactylus ariasae">Jaragua sphaero</a>, is a mere 16 millimetres (0.63 inches) long, and was discovered in 2001 on a small island off the coast of <a href="/wiki/Hispaniola" title="Hispaniola">Hispaniola</a>.<sup id="cite_ref-Piper_10-0" class="reference"><a href="#cite_note-Piper-10"><span class="cite-bracket">&#91;</span>10<span class="cite-bracket">&#93;</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Etymology">Etymology</h2></div> <p>The <a href="/wiki/Neo-Latin" title="Neo-Latin">Neo-Latin</a> <i>gekko</i> and <a href="/wiki/English_language" title="English language">English</a> 'gecko' stem from <a href="/wiki/Indonesian_language" title="Indonesian language">Indonesian</a>-<a href="/wiki/Malaysian_language" class="mw-redirect" title="Malaysian language">Malaysian</a> <i>gēkoq</i>,<sup id="cite_ref-OED_11-0" class="reference"><a href="#cite_note-OED-11"><span class="cite-bracket">&#91;</span>11<span class="cite-bracket">&#93;</span></a></sup> a <a href="/wiki/Malay_language" title="Malay language">Malay</a> word borrowed from <a href="/wiki/Javanese_language" title="Javanese language">Javanese</a>,<sup id="cite_ref-Wilkinson_12-0" class="reference"><a href="#cite_note-Wilkinson-12"><span class="cite-bracket">&#91;</span>12<span class="cite-bracket">&#93;</span></a></sup> from <i>tokek</i>, which <a href="/wiki/Onomatopoeia" title="Onomatopoeia">imitates</a> the sounds that some species like <i><a href="/wiki/Tokay_gecko" title="Tokay gecko">Tokay gecko</a></i> make.<sup id="cite_ref-OED_11-1" class="reference"><a href="#cite_note-OED-11"><span class="cite-bracket">&#91;</span>11<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">&#91;</span>13<span class="cite-bracket">&#93;</span></a></sup><sup class="reference nowrap"><span title="Page / location: 120">&#58;&#8202;120&#8202;</span></sup><sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">&#91;</span>14<span class="cite-bracket">&#93;</span></a></sup><sup class="reference nowrap"><span title="Page / location: 253">&#58;&#8202;253&#8202;</span></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Common_traits">Common traits</h2></div> <p>Like other reptiles, geckos are <a href="/wiki/Ectotherm" title="Ectotherm">ectothermic</a>,<sup id="cite_ref-nzetc.victoria.ac.nz_15-0" class="reference"><a href="#cite_note-nzetc.victoria.ac.nz-15"><span class="cite-bracket">&#91;</span>15<span class="cite-bracket">&#93;</span></a></sup> producing very little metabolic heat. Essentially, a gecko's body temperature is dependent on its environment. Also, to accomplish their main functions; such as locomotion, feeding, reproduction, etc., geckos must have a relatively elevated temperature.<sup id="cite_ref-nzetc.victoria.ac.nz_15-1" class="reference"><a href="#cite_note-nzetc.victoria.ac.nz-15"><span class="cite-bracket">&#91;</span>15<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Shedding_or_molting">Shedding or molting</h3></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><span><video id="mwe_player_0" poster="//upload.wikimedia.org/wikipedia/commons/thumb/7/79/Leopard_Gecko_Shedding_Skin.ogv/220px--Leopard_Gecko_Shedding_Skin.ogv.jpg" controls="" preload="none" data-mw-tmh="" class="mw-file-element" width="220" height="165" data-durationhint="123" data-mwtitle="Leopard_Gecko_Shedding_Skin.ogv" data-mwprovider="wikimediacommons" resource="/wiki/File:Leopard_Gecko_Shedding_Skin.ogv"><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/7/79/Leopard_Gecko_Shedding_Skin.ogv/Leopard_Gecko_Shedding_Skin.ogv.480p.vp9.webm" type="video/webm; codecs=&quot;vp9, opus&quot;" data-transcodekey="480p.vp9.webm" data-width="640" data-height="480" /><source src="//upload.wikimedia.org/wikipedia/commons/7/79/Leopard_Gecko_Shedding_Skin.ogv" type="video/ogg; codecs=&quot;theora&quot;" data-width="640" data-height="480" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/7/79/Leopard_Gecko_Shedding_Skin.ogv/Leopard_Gecko_Shedding_Skin.ogv.144p.mjpeg.mov" type="video/quicktime" data-transcodekey="144p.mjpeg.mov" data-width="192" data-height="144" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/7/79/Leopard_Gecko_Shedding_Skin.ogv/Leopard_Gecko_Shedding_Skin.ogv.240p.vp9.webm" type="video/webm; codecs=&quot;vp9, opus&quot;" data-transcodekey="240p.vp9.webm" data-width="320" data-height="240" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/7/79/Leopard_Gecko_Shedding_Skin.ogv/Leopard_Gecko_Shedding_Skin.ogv.360p.vp9.webm" type="video/webm; codecs=&quot;vp9, opus&quot;" data-transcodekey="360p.vp9.webm" data-width="480" data-height="360" /><source src="//upload.wikimedia.org/wikipedia/commons/transcoded/7/79/Leopard_Gecko_Shedding_Skin.ogv/Leopard_Gecko_Shedding_Skin.ogv.360p.webm" type="video/webm; codecs=&quot;vp8, vorbis&quot;" data-transcodekey="360p.webm" data-width="480" data-height="360" /></video></span><figcaption>Video of leopard gecko shedding skin</figcaption></figure> <p>All geckos shed their skin at fairly regular intervals, with species differing in timing and method. <a href="/wiki/Leopard_gecko" title="Leopard gecko">Leopard geckos</a> shed at about two- to four-week intervals. The presence of moisture aids in the shedding. When shedding begins, the gecko speeds the process by detaching the loose skin from its body and eating it.<sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">&#91;</span>16<span class="cite-bracket">&#93;</span></a></sup> For young geckos, shedding occurs more frequently, once a week, but when they are fully grown, they shed once every one to two months.<sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">&#91;</span>17<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Adhesion_ability">Adhesion ability</h3></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951" /><div role="note" class="hatnote navigation-not-searchable">See also: <a href="/wiki/Synthetic_setae" title="Synthetic setae">Synthetic setae</a> and <a href="/wiki/Gecko_feet" title="Gecko feet">Gecko feet</a></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Gecko_foot_on_glass.JPG" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/b/b2/Gecko_foot_on_glass.JPG/220px-Gecko_foot_on_glass.JPG" decoding="async" width="220" height="230" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/b/b2/Gecko_foot_on_glass.JPG/330px-Gecko_foot_on_glass.JPG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/b/b2/Gecko_foot_on_glass.JPG/440px-Gecko_foot_on_glass.JPG 2x" data-file-width="1871" data-file-height="1955" /></a><figcaption>Close-up of the underside of a gecko's foot as it walks on vertical glass</figcaption></figure> <p>About 60% of gecko species have adhesive toe pads which allow them to <a href="/wiki/Adhesion" title="Adhesion">adhere</a> to most surfaces without the use of liquids or <a href="/wiki/Surface_tension" title="Surface tension">surface tension</a>. Such pads have been gained and lost repeatedly over the course of gecko evolution.<sup id="cite_ref-plosone.org_18-0" class="reference"><a href="#cite_note-plosone.org-18"><span class="cite-bracket">&#91;</span>18<span class="cite-bracket">&#93;</span></a></sup> Adhesive toepads evolved independently in about eleven different gecko lineages, and were lost in at least nine lineages.<sup id="cite_ref-plosone.org_18-1" class="reference"><a href="#cite_note-plosone.org-18"><span class="cite-bracket">&#91;</span>18<span class="cite-bracket">&#93;</span></a></sup> </p><p>It was previously thought that the spatula-shaped <a href="/wiki/Seta" title="Seta">setae</a> arranged in <a href="/wiki/Lamella_(anatomy)" class="mw-redirect" title="Lamella (anatomy)">lamellae</a> on gecko footpads enable attractive <a href="/wiki/Van_der_Waals%27_force" class="mw-redirect" title="Van der Waals&#39; force">van der Waals' forces</a> (the weakest of the weak chemical forces) between the <a href="/wiki/Beta-keratin" title="Beta-keratin">β-keratin</a> lamellae / setae / spatulae structures and the surface.<sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">&#91;</span>19<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Santos2007_20-0" class="reference"><a href="#cite_note-Santos2007-20"><span class="cite-bracket">&#91;</span>20<span class="cite-bracket">&#93;</span></a></sup> These van der Waals interactions involve no fluids; in theory, a boot made of <a href="/wiki/Synthetic_setae" title="Synthetic setae">synthetic setae</a> would adhere as easily to the surface of the <a href="/wiki/International_Space_Station" title="International Space Station">International Space Station</a> as it would to a living-room wall, although adhesion varies with humidity.<sup id="cite_ref-Puthoff_21-0" class="reference"><a href="#cite_note-Puthoff-21"><span class="cite-bracket">&#91;</span>21<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Prowse_22-0" class="reference"><a href="#cite_note-Prowse-22"><span class="cite-bracket">&#91;</span>22<span class="cite-bracket">&#93;</span></a></sup> However, a 2014 study suggests that gecko adhesion is in fact mainly determined by electrostatic interaction (caused by contact electrification), not van der Waals or capillary forces.<sup id="cite_ref-RSI-2014_23-0" class="reference"><a href="#cite_note-RSI-2014-23"><span class="cite-bracket">&#91;</span>23<span class="cite-bracket">&#93;</span></a></sup> </p><p>The setae on the feet of geckos are also self-cleaning, and usually remove any clogging dirt within a few steps.<sup id="cite_ref-Hansen_24-0" class="reference"><a href="#cite_note-Hansen-24"><span class="cite-bracket">&#91;</span>24<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-25" class="reference"><a href="#cite_note-25"><span class="cite-bracket">&#91;</span>25<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-SpatulaeNature_26-0" class="reference"><a href="#cite_note-SpatulaeNature-26"><span class="cite-bracket">&#91;</span>26<span class="cite-bracket">&#93;</span></a></sup> <a href="/wiki/Polytetrafluoroethylene" title="Polytetrafluoroethylene">Polytetrafluoroethylene</a> (PTFE), which has very low surface energy,<sup id="cite_ref-27" class="reference"><a href="#cite_note-27"><span class="cite-bracket">&#91;</span>27<span class="cite-bracket">&#93;</span></a></sup> is more difficult for geckos to adhere to than many other surfaces. </p><p>Gecko adhesion is typically improved by higher humidity,<sup id="cite_ref-Puthoff_21-1" class="reference"><a href="#cite_note-Puthoff-21"><span class="cite-bracket">&#91;</span>21<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Prowse_22-1" class="reference"><a href="#cite_note-Prowse-22"><span class="cite-bracket">&#91;</span>22<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Huber,_G.,_et_al._2005_16293–6_28-0" class="reference"><a href="#cite_note-Huber,_G.,_et_al._2005_16293–6-28"><span class="cite-bracket">&#91;</span>28<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Chen,_B._2010_1–9_29-0" class="reference"><a href="#cite_note-Chen,_B._2010_1–9-29"><span class="cite-bracket">&#91;</span>29<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Loskill,_P._20120587_30-0" class="reference"><a href="#cite_note-Loskill,_P._20120587-30"><span class="cite-bracket">&#91;</span>30<span class="cite-bracket">&#93;</span></a></sup> even on hydrophobic surfaces, yet is reduced under conditions of complete immersion in water. The role of water in that system is under discussion, yet recent experiments agree that the presence of molecular water layers (water molecules carry a very large dipole moment) on the setae, as well as on the surface, increase the surface energy of both, therefore the energy gain in getting these surfaces in contact is enlarged, which results in an increased gecko adhesion force.<sup id="cite_ref-Puthoff_21-2" class="reference"><a href="#cite_note-Puthoff-21"><span class="cite-bracket">&#91;</span>21<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Prowse_22-2" class="reference"><a href="#cite_note-Prowse-22"><span class="cite-bracket">&#91;</span>22<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Huber,_G.,_et_al._2005_16293–6_28-1" class="reference"><a href="#cite_note-Huber,_G.,_et_al._2005_16293–6-28"><span class="cite-bracket">&#91;</span>28<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Chen,_B._2010_1–9_29-1" class="reference"><a href="#cite_note-Chen,_B._2010_1–9-29"><span class="cite-bracket">&#91;</span>29<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Loskill,_P._20120587_30-1" class="reference"><a href="#cite_note-Loskill,_P._20120587-30"><span class="cite-bracket">&#91;</span>30<span class="cite-bracket">&#93;</span></a></sup> Moreover, the elastic properties of the b-keratin change with water uptake.<sup id="cite_ref-Puthoff_21-3" class="reference"><a href="#cite_note-Puthoff-21"><span class="cite-bracket">&#91;</span>21<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Prowse_22-3" class="reference"><a href="#cite_note-Prowse-22"><span class="cite-bracket">&#91;</span>22<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Huber,_G.,_et_al._2005_16293–6_28-2" class="reference"><a href="#cite_note-Huber,_G.,_et_al._2005_16293–6-28"><span class="cite-bracket">&#91;</span>28<span class="cite-bracket">&#93;</span></a></sup> </p><p>Gecko toes seem to be <a href="/wiki/Double_jointed" class="mw-redirect" title="Double jointed">double-jointed</a>, but this is a misnomer, and is properly called digital hyperextension.<sup id="cite_ref-Russell,_A.P._1975_31-0" class="reference"><a href="#cite_note-Russell,_A.P._1975-31"><span class="cite-bracket">&#91;</span>31<span class="cite-bracket">&#93;</span></a></sup> Gecko toes can hyperextend in the opposite direction from human fingers and toes. This allows them to overcome the van der Waals force by peeling their toes off surfaces from the tips inward. In essence, by this peeling action, the gecko separates spatula by spatula from the surface, so for each spatula separation, only some force necessary. (The process is similar to removing <a href="/wiki/Scotch_Tape" class="mw-redirect" title="Scotch Tape">Scotch Tape</a> from a surface.) </p><p>Geckos' toes operate well below their full attractive capabilities most of the time, because the margin for error is great depending upon the <a href="/wiki/Surface_roughness" title="Surface roughness">surface roughness</a>, and therefore the number of setae in contact with that surface. </p><p>Use of small van der Waals force requires very large surface areas; every square millimetre of a gecko's footpad contains about 14,000&#160;hair-like setae. Each seta has a diameter of 5&#160;<a href="/wiki/Micrometer_(unit)" class="mw-redirect" title="Micrometer (unit)">μm</a>. Human hair varies from 18 to 180&#160;μm, so the cross-sectional area of a human hair is equivalent to 12 to 1300 setae. Each seta is in turn tipped with between 100 and 1,000 spatulae.<sup id="cite_ref-Hansen_24-1" class="reference"><a href="#cite_note-Hansen-24"><span class="cite-bracket">&#91;</span>24<span class="cite-bracket">&#93;</span></a></sup> Each spatula is 0.2&#160;μm long<sup id="cite_ref-Hansen_24-2" class="reference"><a href="#cite_note-Hansen-24"><span class="cite-bracket">&#91;</span>24<span class="cite-bracket">&#93;</span></a></sup> (one five-millionth of a metre), or just below the wavelength of visible light.<sup id="cite_ref-Autumn_32-0" class="reference"><a href="#cite_note-Autumn-32"><span class="cite-bracket">&#91;</span>32<span class="cite-bracket">&#93;</span></a></sup> </p><p>The setae of a typical mature 70-<a href="/wiki/Gram" title="Gram">gram</a> (2.5-<a href="/wiki/Ounce" title="Ounce">ounce</a>) gecko would be capable of supporting a weight of 133 kilograms (293 pounds):<sup id="cite_ref-33" class="reference"><a href="#cite_note-33"><span class="cite-bracket">&#91;</span>33<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Autumn_SciAmer_34-0" class="reference"><a href="#cite_note-Autumn_SciAmer-34"><span class="cite-bracket">&#91;</span>34<span class="cite-bracket">&#93;</span></a></sup> each spatula could exert an adhesive force of 5 to 25&#160;nN.<sup id="cite_ref-Huber,_G.,_et_al._2005_16293–6_28-3" class="reference"><a href="#cite_note-Huber,_G.,_et_al._2005_16293–6-28"><span class="cite-bracket">&#91;</span>28<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Lee_35-0" class="reference"><a href="#cite_note-Lee-35"><span class="cite-bracket">&#91;</span>35<span class="cite-bracket">&#93;</span></a></sup> The exact value of the adhesion force of a spatula varies with the surface energy of the substrate to which it adheres. Recent studies<sup id="cite_ref-Loskill,_P._20120587_30-2" class="reference"><a href="#cite_note-Loskill,_P._20120587-30"><span class="cite-bracket">&#91;</span>30<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-36" class="reference"><a href="#cite_note-36"><span class="cite-bracket">&#91;</span>36<span class="cite-bracket">&#93;</span></a></sup> have moreover shown that the component of the surface energy derived from long-range forces, such as van der Waals forces, depends on the material's structure below the outermost atomic layers (up to 100&#160;nm beneath the surface); taking that into account, the adhesive strength can be inferred. </p><p>Apart from the <a href="/wiki/Seta" title="Seta">setae</a>, <a href="/wiki/Phospholipids" class="mw-redirect" title="Phospholipids">phospholipids</a>; fatty substances produced naturally in their bodies, also come into play.<sup id="cite_ref-Hsu_Phospholipids_37-0" class="reference"><a href="#cite_note-Hsu_Phospholipids-37"><span class="cite-bracket">&#91;</span>37<span class="cite-bracket">&#93;</span></a></sup> These lipids lubricate the setae and allow the gecko to detach its foot before the next step. </p><p>The origin of gecko adhesion likely started as simple modifications to the epidermis on the underside of the toes. This was recently discovered in the genus <i><a href="/wiki/Gonatodes" title="Gonatodes">Gonatodes</a></i> from South America.<sup id="cite_ref-Higham,_T.E._2016_38-0" class="reference"><a href="#cite_note-Higham,_T.E._2016-38"><span class="cite-bracket">&#91;</span>38<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-39" class="reference"><a href="#cite_note-39"><span class="cite-bracket">&#91;</span>39<span class="cite-bracket">&#93;</span></a></sup> Simple elaborations of the epidermal spinules into setae have enabled <i>Gonatodes humeralis</i> to climb smooth surfaces and sleep on smooth leaves. </p><p><a href="/wiki/Biomimetic" class="mw-redirect" title="Biomimetic">Biomimetic</a> technologies designed to <a href="/wiki/Synthetic_setae" title="Synthetic setae">mimic gecko adhesion</a> could produce reusable self-cleaning dry adhesives with many applications. Development effort is being put into these technologies, but manufacturing synthetic setae is not a trivial material design task. </p> <div class="mw-heading mw-heading2"><h2 id="Skin">Skin</h2></div> <p>Gecko skin does not generally bear scales, but appears at a macro scale as a papillose surface, which is made from hair-like protuberances developed across the entire body. These confer <a href="/wiki/Ultrahydrophobicity" title="Ultrahydrophobicity">superhydrophobicity</a>, and the unique design of the hair confers a profound antimicrobial action. These protuberances are very small, up to 4&#160;microns in length, and tapering to a point.<sup id="cite_ref-40" class="reference"><a href="#cite_note-40"><span class="cite-bracket">&#91;</span>40<span class="cite-bracket">&#93;</span></a></sup> Gecko skin has been observed to have an anti-bacterial property, killing <a href="/wiki/Gram-negative_bacteria" title="Gram-negative bacteria">gram-negative bacteria</a> when they come in contact with the skin.<sup id="cite_ref-41" class="reference"><a href="#cite_note-41"><span class="cite-bracket">&#91;</span>41<span class="cite-bracket">&#93;</span></a></sup> </p><p>The <a href="/wiki/Uroplatus_sikorae" title="Uroplatus sikorae">mossy leaf-tailed gecko</a> of Madagascar, <i>U. sikorae</i>, has coloration developed as <a href="/wiki/Camouflage" title="Camouflage">camouflage</a>, most being greyish brown to black, or greenish brown, with various markings meant to resemble <a href="/wiki/Tree" title="Tree">tree</a> bark; down to the <a href="/wiki/Lichen" title="Lichen">lichens</a> and moss found on the bark. It also has flaps of skin, running the length of its body, head and limbs, known as the <i>dermal flap</i>, which it can lay against the tree during the day, scattering shadows, and making its outline practically invisible.<sup id="cite_ref-42" class="reference"><a href="#cite_note-42"><span class="cite-bracket">&#91;</span>42<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Teeth">Teeth</h2></div> <p>As <a href="/wiki/Polyphyodont" title="Polyphyodont">polyphyodonts</a>, geckos can replace each of their 100&#160;teeth every 3 to 4 months.<sup id="cite_ref-43" class="reference"><a href="#cite_note-43"><span class="cite-bracket">&#91;</span>43<span class="cite-bracket">&#93;</span></a></sup> Next to the full grown tooth there is a small replacement tooth developing from the <a href="/wiki/Animal_tooth_development" title="Animal tooth development">odontogenic</a> <a href="/wiki/Stem_cell" title="Stem cell">stem cell</a> in the <a href="/wiki/Dental_lamina" title="Dental lamina">dental lamina</a>.<sup id="cite_ref-44" class="reference"><a href="#cite_note-44"><span class="cite-bracket">&#91;</span>44<span class="cite-bracket">&#93;</span></a></sup> The formation of the teeth is <a href="/wiki/Pleurodont" title="Pleurodont">pleurodont</a>; they are fused (ankylosed) by their sides to the inner surface of the jaw bones. This formation is common in all species in the order <a href="/wiki/Squamata" title="Squamata">Squamata</a>. </p> <div style="clear:left;" class=""></div> <div class="mw-heading mw-heading2"><h2 id="Taxonomy_and_classification">Taxonomy and classification</h2></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:LizardFemoralPoresRooij.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/4/48/LizardFemoralPoresRooij.png/250px-LizardFemoralPoresRooij.png" decoding="async" width="220" height="192" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/48/LizardFemoralPoresRooij.png/330px-LizardFemoralPoresRooij.png 1.5x, //upload.wikimedia.org/wikipedia/commons/4/48/LizardFemoralPoresRooij.png 2x" data-file-width="393" data-file-height="343" /></a><figcaption>Pores on the skin are often used in classification.</figcaption></figure> <p>The <a href="/wiki/Infraorder" class="mw-redirect" title="Infraorder">infraorder</a> <a href="/wiki/Gekkota" class="mw-redirect" title="Gekkota">Gekkota</a> is divided into seven families, containing about 125 <a href="/wiki/Genus" title="Genus">genera</a> of geckos, including the snake-like (legless) pygopods.<sup id="cite_ref-plosone.org_18-2" class="reference"><a href="#cite_note-plosone.org-18"><span class="cite-bracket">&#91;</span>18<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-45" class="reference"><a href="#cite_note-45"><span class="cite-bracket">&#91;</span>45<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-46" class="reference"><a href="#cite_note-46"><span class="cite-bracket">&#91;</span>46<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Gamble_Gondwana_47-0" class="reference"><a href="#cite_note-Gamble_Gondwana-47"><span class="cite-bracket">&#91;</span>47<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-48" class="reference"><a href="#cite_note-48"><span class="cite-bracket">&#91;</span>48<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-Into_the_light:_Diurnality_has_evol_4-2" class="reference"><a href="#cite_note-Into_the_light:_Diurnality_has_evol-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-49" class="reference"><a href="#cite_note-49"><span class="cite-bracket">&#91;</span>49<span class="cite-bracket">&#93;</span></a></sup> </p> <ul><li>Family <a href="/wiki/Carphodactylidae" title="Carphodactylidae">Carphodactylidae</a></li> <li>Family <a href="/wiki/Diplodactylidae" title="Diplodactylidae">Diplodactylidae</a></li> <li>Family <a href="/wiki/Eublepharidae" title="Eublepharidae">Eublepharidae</a></li> <li>Family <a href="/wiki/Gekkonidae" title="Gekkonidae">Gekkonidae</a></li> <li>Family <a href="/wiki/Phyllodactylidae" title="Phyllodactylidae">Phyllodactylidae</a></li> <li>Family <a href="/wiki/Pygopodidae" title="Pygopodidae">Pygopodidae</a></li> <li>Family <a href="/wiki/Sphaerodactylidae" title="Sphaerodactylidae">Sphaerodactylidae</a></li></ul> <p>Legless lizards of the family <a href="/wiki/Dibamidae" title="Dibamidae">Dibamidae</a>, also referred to as blind lizards,<sup id="cite_ref-50" class="reference"><a href="#cite_note-50"><span class="cite-bracket">&#91;</span>50<span class="cite-bracket">&#93;</span></a></sup> have occasionally been counted as gekkotans, but recent molecular phylogenies suggest otherwise.<sup id="cite_ref-51" class="reference"><a href="#cite_note-51"><span class="cite-bracket">&#91;</span>51<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-52" class="reference"><a href="#cite_note-52"><span class="cite-bracket">&#91;</span>52<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="clade"><style data-mw-deduplicate="TemplateStyles:r1261294616">body.skin-vector-2022 .mw-parser-output div.clade,body.skin-minerva .mw-parser-output div.clade{overflow-x:auto;overflow-y:hidden}body.skin-minerva .mw-parser-output div.clade p{font-size:inherit}.mw-parser-output table.clade{border-spacing:0;margin:0;font-size:100%;line-height:100%;border-collapse:separate;width:auto;display:table}.mw-parser-output table.clade table.clade{width:100%;line-height:inherit}.mw-parser-output table.clade td.clade-label{min-width:0.2em;width:0.2em;padding:0.1em 0.25em;vertical-align:bottom;text-align:center;border-left:1px solid;border-bottom:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-label::before,.mw-parser-output table.clade td.clade-slabel::before{content:"\2060 "}.mw-parser-output table.clade td.clade-fixed-width{overflow:hidden;text-overflow:ellipsis}.mw-parser-output table.clade td.clade-fixed-width:hover{overflow:visible}.mw-parser-output table.clade td.clade-label.first{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-label.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-slabel{padding:0.1em 0.25em;vertical-align:top;text-align:center;border-left:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-slabel:hover{overflow:visible}.mw-parser-output table.clade td.clade-slabel.last{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-slabel.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-bar{vertical-align:middle;text-align:left;padding:0 0.5em;position:relative}.mw-parser-output table.clade td.clade-bar.reverse{text-align:right;position:relative}.mw-parser-output table.clade td.clade-leaf{border:0;padding:0;text-align:left}.mw-parser-output table.clade td.clade-leaf p{padding-right:5px;padding-left:2px}.mw-parser-output table.clade td.clade-leafR{border:0;padding:0;text-align:right}.mw-parser-output table.clade td.clade-leafR p{padding-left:5px;padding-right:2px}.mw-parser-output table.clade td.clade-leaf.reverse{text-align:right}.mw-parser-output table.clade td.clade-leaf.reverse p{padding-left:5px;padding-right:2px}.mw-parser-output table.clade:hover span.linkA{background-color:yellow}.mw-parser-output table.clade:hover span.linkB{background-color:green}</style> <table class="clade"> <tbody><tr> <td class="clade-label first"><b>Gekkota</b> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Diplodactylidae" title="Diplodactylidae">Diplodactylidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Carphodactylidae" title="Carphodactylidae">Carphodactylidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Pygopodidae" title="Pygopodidae">Pygopodidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Eublepharidae" title="Eublepharidae">Eublepharidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Sphaerodactylidae" title="Sphaerodactylidae">Sphaerodactylidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1261294616" /> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Phyllodactylidae" title="Phyllodactylidae">Phyllodactylidae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Gekkonidae" title="Gekkonidae">Gekkonidae</a> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> <div class="mw-heading mw-heading3"><h3 id="Evolutionary_history">Evolutionary history</h3></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Eichstaettisaurus_schroederi_398858_(cropped).jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/e/ed/Eichstaettisaurus_schroederi_398858_%28cropped%29.jpg/220px-Eichstaettisaurus_schroederi_398858_%28cropped%29.jpg" decoding="async" width="220" height="268" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/e/ed/Eichstaettisaurus_schroederi_398858_%28cropped%29.jpg/330px-Eichstaettisaurus_schroederi_398858_%28cropped%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/e/ed/Eichstaettisaurus_schroederi_398858_%28cropped%29.jpg/440px-Eichstaettisaurus_schroederi_398858_%28cropped%29.jpg 2x" data-file-width="1445" data-file-height="1757" /></a><figcaption>Skeleton of <i><a href="/wiki/Eichstaettisaurus" title="Eichstaettisaurus">Eichstaettisaurus</a>,</i> thought to be an early member of the gecko lineage</figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Deutsches_Bernsteinmuseum_04_(cropped).jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/0/0b/Deutsches_Bernsteinmuseum_04_%28cropped%29.jpg/220px-Deutsches_Bernsteinmuseum_04_%28cropped%29.jpg" decoding="async" width="220" height="131" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/0b/Deutsches_Bernsteinmuseum_04_%28cropped%29.jpg/330px-Deutsches_Bernsteinmuseum_04_%28cropped%29.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/0b/Deutsches_Bernsteinmuseum_04_%28cropped%29.jpg/440px-Deutsches_Bernsteinmuseum_04_%28cropped%29.jpg 2x" data-file-width="3000" data-file-height="1786" /></a><figcaption>Fossil of <i><a href="/wiki/Yantarogekko" title="Yantarogekko">Yantarogekko</a></i> preserved in Baltic amber</figcaption></figure> <p>Several species of lizard from the Late Jurassic have been considered early relatives of geckos, the most prominent and most well supported being the arboreal <i><a href="/wiki/Eichstaettisaurus" title="Eichstaettisaurus">Eichstaettisaurus</a></i> from the Late Jurassic of Germany. <i><a href="/wiki/Norellius" title="Norellius">Norellius</a></i> from the Early Cretaceous of Mongolia is also usually placed as a close relative of geckos.<sup id="cite_ref-53" class="reference"><a href="#cite_note-53"><span class="cite-bracket">&#91;</span>53<span class="cite-bracket">&#93;</span></a></sup> The oldest known fossils of modern geckos are from the mid-Cretaceous <a href="/wiki/Burmese_amber" title="Burmese amber">Burmese amber</a> of Myanmar (including <i><a href="/wiki/Cretaceogekko" title="Cretaceogekko">Cretaceogekko</a></i>), around 100 million years old, which have adhesive pads on the feet similar to those of living geckos.<sup id="cite_ref-54" class="reference"><a href="#cite_note-54"><span class="cite-bracket">&#91;</span>54<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-55" class="reference"><a href="#cite_note-55"><span class="cite-bracket">&#91;</span>55<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-56" class="reference"><a href="#cite_note-56"><span class="cite-bracket">&#91;</span>56<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Species">Species</h2></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Mediterranean_house_gecko.JPG" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/ae/Mediterranean_house_gecko.JPG/220px-Mediterranean_house_gecko.JPG" decoding="async" width="220" height="180" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/ae/Mediterranean_house_gecko.JPG/330px-Mediterranean_house_gecko.JPG 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/ae/Mediterranean_house_gecko.JPG/440px-Mediterranean_house_gecko.JPG 2x" data-file-width="2465" data-file-height="2018" /></a><figcaption><a href="/wiki/Mediterranean_house_gecko" title="Mediterranean house gecko">Mediterranean house gecko</a></figcaption></figure><p>More than 1,850 species of geckos occur worldwide,<sup id="cite_ref-57" class="reference"><a href="#cite_note-57"><span class="cite-bracket">&#91;</span>57<span class="cite-bracket">&#93;</span></a></sup> including these familiar species: </p><ul><li><i><a href="/wiki/Coleonyx_variegatus" class="mw-redirect" title="Coleonyx variegatus">Coleonyx variegatus</a></i>, the western banded gecko, is native to the southwestern <a href="/wiki/United_States" title="United States">United States</a> and northwest <a href="/wiki/Mexico" title="Mexico">Mexico</a>.</li> <li><i><a href="/wiki/Cyrtopodion_brachykolon" class="mw-redirect" title="Cyrtopodion brachykolon">Cyrtopodion brachykolon</a></i>, the bent-toed gecko, is found in northwestern <a href="/wiki/Pakistan" title="Pakistan">Pakistan</a>; it was first described in 2007.</li> <li><i><a href="/wiki/Leopard_gecko" title="Leopard gecko">Eublepharis macularius</a></i>, the leopard gecko, is the most common gecko kept as a pet; it does not have adhesive toe pads and cannot climb the glass of a <a href="/wiki/Vivarium" title="Vivarium">vivarium</a>.</li> <li><i><a href="/wiki/Gehyra_mutilata" title="Gehyra mutilata">Gehyra mutilata</a></i> (<i>Pteropus mutilatus</i>), the stump-toed gecko, is able to vary its color from very light to very dark to camouflage itself; this gecko is at home in the wild, as well as in residential areas.</li> <li><i><a href="/wiki/Tokay_gecko" title="Tokay gecko">Gekko gecko</a></i>, the Tokay gecko, is a large, common, <a href="/wiki/Southeast_Asia" title="Southeast Asia">Southeast Asian</a> gecko known for its aggressive temperament, loud mating calls, and bright markings.</li> <li><i><a href="/wiki/Hemidactylus" title="Hemidactylus">Hemidactylus</a></i> is genus of geckos with many varieties. <ul><li><i><a href="/wiki/Common_house_gecko" title="Common house gecko">Hemidactylus frenatus</a></i>, the common house gecko, thrives around people and human habitation structures in the tropics and subtropics worldwide.</li> <li><i><a href="/wiki/Indo-Pacific_gecko" title="Indo-Pacific gecko">Hemidactylus garnotii</a></i>, the Indo-Pacific gecko, is found in houses throughout the tropics, and has become an invasive species of concern in <a href="/wiki/Florida" title="Florida">Florida</a> and <a href="/wiki/Georgia_(U.S._state)" title="Georgia (U.S. state)">Georgia</a> in the US.</li> <li><i><a href="/wiki/Hemidactylus_mabouia" class="mw-redirect" title="Hemidactylus mabouia">Hemidactylus mabouia</a></i>, the tropical house gecko, Afro-American house gecko, or cosmopolitan house gecko, is a species of house gecko native to sub-Saharan Africa and also currently found in North, Central, and South America and the Caribbean.</li> <li><i><a href="/wiki/Mediterranean_house_gecko" title="Mediterranean house gecko">Hemidactylus turcicus</a></i>, the Mediterranean house gecko, is frequently found in and around buildings, and is an <a href="/wiki/Introduced_species" title="Introduced species">introduced species</a> in the US.</li></ul></li> <li><i><a href="/wiki/Lepidodactylus_lugubris" title="Lepidodactylus lugubris">Lepidodactylus lugubris</a></i>, the mourning gecko, is originally an East Asian and Pacific species; it is equally at home in the wild and residential neighborhoods.</li> <li><i><a href="/wiki/Bibron%27s_thick-toed_gecko" title="Bibron&#39;s thick-toed gecko">Pachydactylus bibroni</a></i>, Bibron's gecko, is native to southern Africa; this hardy <a href="/wiki/Arboreal_locomotion" title="Arboreal locomotion">arboreal</a> gecko is considered a household pest.</li> <li><i><a href="/wiki/Gold_dust_day_gecko" title="Gold dust day gecko">Phelsuma laticauda</a></i>, the gold dust day gecko, is diurnal; it lives in northern <a href="/wiki/Madagascar" title="Madagascar">Madagascar</a> and on the <a href="/wiki/Comoros" title="Comoros">Comoros</a>. It is also an introduced species in <a href="/wiki/Hawaiian_Islands" title="Hawaiian Islands">Hawaii</a>.</li> <li><i><a href="/wiki/Ptychozoon" title="Ptychozoon">Ptychozoon</a></i> is a genus of arboreal geckos from Southeast Asia also known as flying or parachute geckos; they have wing-like flaps from the neck to the upper leg to help them conceal themselves on trees and provide lift while jumping.</li> <li><i><a href="/wiki/Rhacodactylus" title="Rhacodactylus">Rhacodactylus</a></i> is genus of geckos native to <a href="/wiki/New_Caledonia" title="New Caledonia">New Caledonia</a>. <ul><li><i><a href="/wiki/Crested_gecko" title="Crested gecko">Rhacodactylus ciliatus</a></i> (now assigned to the genus <i>Correlophus</i>), the crested gecko, was believed extinct until rediscovered in 1994, and is gaining popularity as a pet.</li> <li><i><a href="/wiki/Rhacodactylus_leachianus" title="Rhacodactylus leachianus">Rhacodactylus leachianus</a></i>, the New Caledonian giant gecko, was first described by Cuvier in 1829; it is the largest living species of gecko.</li></ul></li> <li><i><a href="/wiki/Sphaerodactylus_ariasae" title="Sphaerodactylus ariasae">Sphaerodactylus ariasae</a></i>, the dwarf gecko, is native to the Caribbean Islands; it is the world's smallest lizard.</li> <li><i><a href="/wiki/Tarentola_mauritanica" title="Tarentola mauritanica">Tarentola mauritanica</a></i>, the crocodile or Moorish gecko, is commonly found in the <a href="/wiki/Mediterranean_Sea" title="Mediterranean Sea">Mediterranean</a> region from the <a href="/wiki/Iberian_Peninsula" title="Iberian Peninsula">Iberian Peninsula</a> and southern <a href="/wiki/France" title="France">France</a> to <a href="/wiki/Greece" title="Greece">Greece</a> and <a href="/wiki/North_Africa" title="North Africa">northern Africa</a>; their most distinguishing characteristics are their pointed heads, spiked skin, and tails resembling those of a <a href="/wiki/Crocodile" title="Crocodile">crocodile</a>.</li></ul> <div class="mw-heading mw-heading2"><h2 id="Reproduction">Reproduction</h2></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951" /><div role="note" class="hatnote navigation-not-searchable">See also: <a href="/wiki/Parthenogenesis_in_squamata#Lizards" class="mw-redirect" title="Parthenogenesis in squamata">Parthenogenesis in squamata §&#160;Lizards</a></div> <style data-mw-deduplicate="TemplateStyles:r1251242444">.mw-parser-output .ambox{border:1px solid #a2a9b1;border-left:10px solid #36c;background-color:#fbfbfb;box-sizing:border-box}.mw-parser-output .ambox+link+.ambox,.mw-parser-output .ambox+link+style+.ambox,.mw-parser-output .ambox+link+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+style+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+link+.ambox{margin-top:-1px}html body.mediawiki .mw-parser-output .ambox.mbox-small-left{margin:4px 1em 4px 0;overflow:hidden;width:238px;border-collapse:collapse;font-size:88%;line-height:1.25em}.mw-parser-output .ambox-speedy{border-left:10px solid #b32424;background-color:#fee7e6}.mw-parser-output .ambox-delete{border-left:10px solid #b32424}.mw-parser-output .ambox-content{border-left:10px solid #f28500}.mw-parser-output .ambox-style{border-left:10px solid #fc3}.mw-parser-output .ambox-move{border-left:10px solid #9932cc}.mw-parser-output .ambox-protection{border-left:10px solid #a2a9b1}.mw-parser-output .ambox .mbox-text{border:none;padding:0.25em 0.5em;width:100%}.mw-parser-output .ambox .mbox-image{border:none;padding:2px 0 2px 0.5em;text-align:center}.mw-parser-output .ambox .mbox-imageright{border:none;padding:2px 0.5em 2px 0;text-align:center}.mw-parser-output .ambox .mbox-empty-cell{border:none;padding:0;width:1px}.mw-parser-output .ambox .mbox-image-div{width:52px}@media(min-width:720px){.mw-parser-output .ambox{margin:0 10%}}@media print{body.ns-0 .mw-parser-output .ambox{display:none!important}}</style><table class="box-Expand_section plainlinks metadata ambox mbox-small-left ambox-content" role="presentation"><tbody><tr><td class="mbox-image"><span typeof="mw:File"><a href="/wiki/File:Wiki_letter_w_cropped.svg" class="mw-file-description"><img alt="[icon]" src="//upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Wiki_letter_w_cropped.svg/20px-Wiki_letter_w_cropped.svg.png" decoding="async" width="20" height="14" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/1c/Wiki_letter_w_cropped.svg/40px-Wiki_letter_w_cropped.svg.png 1.5x" data-file-width="44" data-file-height="31" /></a></span></td><td class="mbox-text"><div class="mbox-text-span">This section <b>needs expansion</b>. You can help by <span class="anonymous-show"><span class="plainlinks"><a class="external text" href="https://en.wikipedia.org/w/index.php?title=Talk%3AGecko&amp;preload=Template%3ASubmit+an+edit+request%2Fpreload&amp;action=edit&amp;section=new&amp;editintro=Template%3AEdit+protected%2Feditintro&amp;preloadtitle=Protected+edit+request+on+9+April+2025&amp;preloadparams%5B%5D=edit+fully-protected&amp;preloadparams%5B%5D=Gecko">making an edit request</a></span></span><span class="user-show"><a class="external text" href="https://en.wikipedia.org/w/index.php?title=Gecko&amp;action=edit&amp;section=">adding to it</a> </span>. <span class="date-container"><i>(<span class="date">February 2020</span>)</i></span></div></td></tr></tbody></table> <p>Most geckos lay a small clutch of eggs. Some are live-bearing, and a few can <a href="/wiki/Asexual_reproduction" title="Asexual reproduction">reproduce asexually</a> via <a href="/wiki/Parthenogenesis" title="Parthenogenesis">parthenogenesis</a>. Geckos also have a large diversity of sex-determining mechanisms, including <a href="/wiki/Temperature-dependent_sex_determination" title="Temperature-dependent sex determination">temperature-dependent sex determination</a> and both XX/XY and ZZ/ZW <a href="/wiki/Sex_chromosome" title="Sex chromosome">sex chromosomes</a> with multiple transitions among them over evolutionary time.<sup id="cite_ref-58" class="reference"><a href="#cite_note-58"><span class="cite-bracket">&#91;</span>58<span class="cite-bracket">&#93;</span></a></sup> Madagascar day geckos engage in a mating ritual in which sexually mature males produce a waxy substance from pores on the back of their legs. Males approach females with a head swaying motion along with rapid tongue flicking in the female.<sup id="cite_ref-59" class="reference"><a href="#cite_note-59"><span class="cite-bracket">&#91;</span>59<span class="cite-bracket">&#93;</span></a></sup> </p><p>Obligate <a href="/wiki/Parthenogenesis" title="Parthenogenesis">parthenogenesis</a> as a reproductive system has evolved multiple times in the family Gekkonidae.<sup id="cite_ref-Dedukh2022_60-0" class="reference"><a href="#cite_note-Dedukh2022-60"><span class="cite-bracket">&#91;</span>60<span class="cite-bracket">&#93;</span></a></sup> It has been shown that <a href="/wiki/Oocyte" title="Oocyte">oocytes</a> are able to undergo <a href="/wiki/Meiosis" title="Meiosis">meiosis</a> in three different obligate parthenogenetic complexes of geckos. An extra premeiotic endoreplication of <a href="/wiki/Chromosome" title="Chromosome">chromosomes</a> is essential for obligate parthenogenesis in these geckos.<sup id="cite_ref-Dedukh2022_60-1" class="reference"><a href="#cite_note-Dedukh2022-60"><span class="cite-bracket">&#91;</span>60<span class="cite-bracket">&#93;</span></a></sup> Appropriate segregation during meiosis to form viable progeny is facilitated by the formation of <a href="/wiki/Bivalent_(genetics)" title="Bivalent (genetics)">bivalents</a> made from copies of identical chromosomes. </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFVillaWingsRabi2022" class="citation journal cs1">Villa, Andrea; Wings, Oliver; Rabi, Márton (2022). Angielczyk, Kenneth (ed.). <a rel="nofollow" class="external text" href="https://ddd.uab.cat/pub/artpub/2022/259562/pappal_a2022v8n3ae1434.pdf">"A new gecko (Squamata, Gekkota) from the Eocene of Geiseltal (Germany) implies long-term persistence of European Sphaerodactylidae"</a> <span class="cs1-format">(PDF)</span>. <i>Papers in Palaeontology</i>. <b>8</b> (3). <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2022PPal....8E1434V">2022PPal....8E1434V</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1002%2Fspp2.1434">10.1002/spp2.1434</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a>&#160;<a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/2056-2799">2056-2799</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a>&#160;<a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:249358350">249358350</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Papers+in+Palaeontology&amp;rft.atitle=A+new+gecko+%28Squamata%2C+Gekkota%29+from+the+Eocene+of+Geiseltal+%28Germany%29+implies+long-term+persistence+of+European+Sphaerodactylidae&amp;rft.volume=8&amp;rft.issue=3&amp;rft.date=2022&amp;rft_id=info%3Adoi%2F10.1002%2Fspp2.1434&amp;rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A249358350%23id-name%3DS2CID&amp;rft.issn=2056-2799&amp;rft_id=info%3Abibcode%2F2022PPal....8E1434V&amp;rft.aulast=Villa&amp;rft.aufirst=Andrea&amp;rft.au=Wings%2C+Oliver&amp;rft.au=Rabi%2C+M%C3%A1rton&amp;rft_id=https%3A%2F%2Fddd.uab.cat%2Fpub%2Fartpub%2F2022%2F259562%2Fpappal_a2022v8n3ae1434.pdf&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AGecko" class="Z3988"></span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-2">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://reptile-database.reptarium.cz/advanced_search?taxon=gecko&amp;submit=Search">"Search results – gecko"</a>. <i>Reptile-Database.Reptarium.cz</i>. 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Archived from <a rel="nofollow" class="external text" href="http://www.GeckoCare.net/shedding.php">the original</a> on 29 May 2013<span class="reference-accessdate">. Retrieved <span class="nowrap">19 April</span> 2013</span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=unknown&amp;rft.jtitle=www.GeckoCare.net&amp;rft.atitle=GeckoCare+-+shedding&amp;rft_id=http%3A%2F%2Fwww.GeckoCare.net%2Fshedding.php&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AGecko" class="Z3988"></span></span> </li> <li id="cite_note-17"><span class="mw-cite-backlink"><b><a href="#cite_ref-17">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://buddygenius.com/crested-geckos-shed/">"Crested geckos shedding"</a>. <i>BuddyGenius.com</i>. Buddy Genius. 5 July 2020. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20180116140204/https://buddygenius.com/crested-geckos-shed/">Archived</a> from the original on 16 January 2018<span class="reference-accessdate">. Retrieved <span class="nowrap">1 February</span> 2022</span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=unknown&amp;rft.jtitle=BuddyGenius.com&amp;rft.atitle=Crested+geckos+shedding&amp;rft.date=2020-07-05&amp;rft_id=https%3A%2F%2Fbuddygenius.com%2Fcrested-geckos-shed%2F&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AGecko" class="Z3988"></span></span> </li> <li id="cite_note-plosone.org-18"><span class="mw-cite-backlink">^ <a href="#cite_ref-plosone.org_18-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-plosone.org_18-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-plosone.org_18-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFGambleGreenbaumJackmanRussell2012" class="citation journal cs1">Gamble, Tony; Greenbaum, Eli; Jackman, Todd R.; Russell, Anthony P.; Bauer, Aaron M. (27 June 2012). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3384654">"Repeated Origin and Loss of Adhesive Toepads in Geckos"</a>. <i>PLOS ONE</i>. <b>7</b> (6): e39429. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2012PLoSO...739429G">2012PLoSO...739429G</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1371%2Fjournal.pone.0039429">10.1371/journal.pone.0039429</a></span>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a>&#160;<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3384654">3384654</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/22761794">22761794</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=PLOS+ONE&amp;rft.atitle=Repeated+Origin+and+Loss+of+Adhesive+Toepads+in+Geckos&amp;rft.volume=7&amp;rft.issue=6&amp;rft.pages=e39429&amp;rft.date=2012-06-27&amp;rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3384654%23id-name%3DPMC&amp;rft_id=info%3Apmid%2F22761794&amp;rft_id=info%3Adoi%2F10.1371%2Fjournal.pone.0039429&amp;rft_id=info%3Abibcode%2F2012PLoSO...739429G&amp;rft.aulast=Gamble&amp;rft.aufirst=Tony&amp;rft.au=Greenbaum%2C+Eli&amp;rft.au=Jackman%2C+Todd+R.&amp;rft.au=Russell%2C+Anthony+P.&amp;rft.au=Bauer%2C+Aaron+M.&amp;rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3384654&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AGecko" class="Z3988"></span></span> </li> <li id="cite_note-19"><span class="mw-cite-backlink"><b><a href="#cite_ref-19">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.nisenet.org/scientific-images/gecko_toe">"Scientific image – gecko toe"</a>. <i>www.NISEnet.org</i>. NISE Network. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20130509025409/http://www.nisenet.org/scientific-images/gecko_toe">Archived</a> from the original on 2013-05-09<span class="reference-accessdate">. Retrieved <span class="nowrap">2022-02-01</span></span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=unknown&amp;rft.jtitle=www.NISEnet.org&amp;rft.atitle=Scientific+image+%E2%80%93+gecko+toe&amp;rft_id=http%3A%2F%2Fwww.nisenet.org%2Fscientific-images%2Fgecko_toe&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AGecko" class="Z3988"></span></span> </li> <li id="cite_note-Santos2007-20"><span class="mw-cite-backlink"><b><a href="#cite_ref-Santos2007_20-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFSantosSpenkoParnessSangbae2007" class="citation journal cs1">Santos, Daniel; Spenko, Matthew; Parness, Aaron; Sangbae, Kim; Cutkosky, Mark (2007). <a rel="nofollow" class="external text" href="http://www.brill.nl/journal-adhesion-science-and-technology">"Directional adhesion for climbing: Theoretical and practical considerations"</a>. <i><a href="/w/index.php?title=Journal_of_Adhesion_Science_and_Technology&amp;action=edit&amp;redlink=1" class="new" title="Journal of Adhesion Science and Technology (page does not exist)">Journal of Adhesion Science and Technology</a></i>. <b>21</b> (<span class="nowrap">12–</span>13): <span class="nowrap">1317–</span>1341. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1163%2F156856107782328399">10.1163/156856107782328399</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a>&#160;<a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:53470787">53470787</a>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20120115143737/http://www.brill.nl/journal-adhesion-science-and-technology">Archived</a> from the original on 2012-01-15<span class="reference-accessdate">. Retrieved <span class="nowrap">2012-02-04</span></span>. <q><i>Gecko "feet and toes are a hierarchical system of complex structures consisting of lamellae, setae, and spatulae. The distinguishing characteristics of the gecko adhesion system have been described [as] (1) anisotropic attachment, (2) high pulloff force to preload ratio, (3) low detachment force, (4) material independence, (5) self-cleaning, (6) antiself sticking and (7) nonsticky default state. ... The gecko's adhesive structures are made from ß-keratin (modulus of elasticity [about] 2 GPa). Such a stiff material is not inherently sticky; however, because of the gecko adhesive's hierarchical nature and extremely small distal features (spatulae are [about] 200 nm in size), the gecko's foot is able to intimately conform to the surface and generate significant attraction using <a href="/wiki/Van_der_Waals_force" title="Van der Waals force">van der Waals forces</a>.</i></q></cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Journal+of+Adhesion+Science+and+Technology&amp;rft.atitle=Directional+adhesion+for+climbing%3A+Theoretical+and+practical+considerations&amp;rft.volume=21&amp;rft.issue=%3Cspan+class%3D%22nowrap%22%3E12%E2%80%93%3C%2Fspan%3E13&amp;rft.pages=%3Cspan+class%3D%22nowrap%22%3E1317-%3C%2Fspan%3E1341&amp;rft.date=2007&amp;rft_id=info%3Adoi%2F10.1163%2F156856107782328399&amp;rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A53470787%23id-name%3DS2CID&amp;rft.aulast=Santos&amp;rft.aufirst=Daniel&amp;rft.au=Spenko%2C+Matthew&amp;rft.au=Parness%2C+Aaron&amp;rft.au=Sangbae%2C+Kim&amp;rft.au=Cutkosky%2C+Mark&amp;rft_id=http%3A%2F%2Fwww.brill.nl%2Fjournal-adhesion-science-and-technology&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AGecko" class="Z3988"></span></span> </li> <li id="cite_note-Puthoff-21"><span class="mw-cite-backlink">^ <a href="#cite_ref-Puthoff_21-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Puthoff_21-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Puthoff_21-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-Puthoff_21-3"><sup><i><b>d</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFPuthoffProwseWilkinsonAutumn2010" class="citation journal cs1">Puthoff, J.B.; Prowse, M.; Wilkinson, M.; Autumn, K. 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rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/30938766">30938766</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Integrative+and+Comparative+Biology&amp;rft.atitle=Gecko+Adhesion+in+Space+and+Time%3A+A+Phylogenetic+Perspective+on+the+Scansorial+Success+Story&amp;rft.volume=59&amp;rft.issue=1&amp;rft.pages=%3Cspan+class%3D%22nowrap%22%3E117-%3C%2Fspan%3E130&amp;rft.date=2019-07-01&amp;rft.issn=1540-7063&amp;rft_id=info%3Apmid%2F30938766&amp;rft_id=info%3Adoi%2F10.1093%2Ficb%2Ficz020&amp;rft.aulast=Bauer&amp;rft.aufirst=A+M&amp;rft_id=https%3A%2F%2Fdoi.org%2F10.1093%252Ficb%252Ficz020&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AGecko" class="Z3988"></span></span> </li> <li id="cite_note-57"><span class="mw-cite-backlink"><b><a href="#cite_ref-57">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.reptile-database.org/">"THE REPTILE DATABASE"</a>. <i>www.reptile-database.org</i>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20211029024411/http://www.reptile-database.org/">Archived</a> from the original on 2021-10-29<span class="reference-accessdate">. Retrieved <span class="nowrap">2016-09-20</span></span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=unknown&amp;rft.jtitle=www.reptile-database.org&amp;rft.atitle=THE+REPTILE+DATABASE&amp;rft_id=http%3A%2F%2Fwww.reptile-database.org%2F&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AGecko" class="Z3988"></span></span> </li> <li id="cite_note-58"><span class="mw-cite-backlink"><b><a href="#cite_ref-58">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFGambleCoryell,_J.Ezaz,_T.Lynch,_J.2015" class="citation journal cs1">Gamble, Tony; Coryell, J.; Ezaz, T.; Lynch, J.; Scantlebury, D.; Zarkower, D. (2015). <a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fmolbev%2Fmsv023">"Restriction site-associated DNA sequencing (RAD-seq) reveals an extraordinary number of transitions among gecko sex-determining systems"</a>. <i>Molecular Biology and Evolution</i>. <b>32</b> (5): <span class="nowrap">1296–</span>1309. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fmolbev%2Fmsv023">10.1093/molbev/msv023</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/25657328">25657328</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Molecular+Biology+and+Evolution&amp;rft.atitle=Restriction+site-associated+DNA+sequencing+%28RAD-seq%29+reveals+an+extraordinary+number+of+transitions+among+gecko+sex-determining+systems&amp;rft.volume=32&amp;rft.issue=5&amp;rft.pages=%3Cspan+class%3D%22nowrap%22%3E1296-%3C%2Fspan%3E1309&amp;rft.date=2015&amp;rft_id=info%3Adoi%2F10.1093%2Fmolbev%2Fmsv023&amp;rft_id=info%3Apmid%2F25657328&amp;rft.aulast=Gamble&amp;rft.aufirst=Tony&amp;rft.au=Coryell%2C+J.&amp;rft.au=Ezaz%2C+T.&amp;rft.au=Lynch%2C+J.&amp;rft.au=Scantlebury%2C+D.&amp;rft.au=Zarkower%2C+D.&amp;rft_id=https%3A%2F%2Fdoi.org%2F10.1093%252Fmolbev%252Fmsv023&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AGecko" class="Z3988"></span></span> </li> <li id="cite_note-59"><span class="mw-cite-backlink"><b><a href="#cite_ref-59">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFFryRoycroft" class="citation web cs1">Fry, Courtney; Roycroft, Carl. <a rel="nofollow" class="external text" href="https://animaldiversity.org/accounts/Phelsuma_madagascariensis/">"Phelsuma madagascariensis (Madagascar Day Gecko)"</a>. <i>Animal Diversity Web</i>. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20231001011236/https://animaldiversity.org/accounts/Phelsuma_madagascariensis/">Archived</a> from the original on 2023-10-01<span class="reference-accessdate">. Retrieved <span class="nowrap">2024-02-07</span></span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=unknown&amp;rft.jtitle=Animal+Diversity+Web&amp;rft.atitle=Phelsuma+madagascariensis+%28Madagascar+Day+Gecko%29&amp;rft.aulast=Fry&amp;rft.aufirst=Courtney&amp;rft.au=Roycroft%2C+Carl&amp;rft_id=https%3A%2F%2Fanimaldiversity.org%2Faccounts%2FPhelsuma_madagascariensis%2F&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AGecko" class="Z3988"></span></span> </li> <li id="cite_note-Dedukh2022-60"><span class="mw-cite-backlink">^ <a href="#cite_ref-Dedukh2022_60-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Dedukh2022_60-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFDedukhAltmanováKlímaKratochvíl2022" class="citation journal cs1">Dedukh, Dmitrij; Altmanová, Marie; Klíma, Jiří; Kratochvíl, Lukáš (2022-04-01). <a rel="nofollow" class="external text" href="https://journals.biologists.com/dev/article/149/7/dev200345/275038/Premeiotic-endoreplication-is-essential-for">"Premeiotic endoreplication is essential for obligate parthenogenesis in geckos"</a>. <i>Development</i>. <b>149</b> (7): dev200345. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1242%2Fdev.200345">10.1242/dev.200345</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a>&#160;<a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/1477-9129">1477-9129</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/35388415">35388415</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a>&#160;<a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:248001402">248001402</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Development&amp;rft.atitle=Premeiotic+endoreplication+is+essential+for+obligate+parthenogenesis+in+geckos&amp;rft.volume=149&amp;rft.issue=7&amp;rft.pages=dev200345&amp;rft.date=2022-04-01&amp;rft.issn=1477-9129&amp;rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A248001402%23id-name%3DS2CID&amp;rft_id=info%3Apmid%2F35388415&amp;rft_id=info%3Adoi%2F10.1242%2Fdev.200345&amp;rft.aulast=Dedukh&amp;rft.aufirst=Dmitrij&amp;rft.au=Altmanov%C3%A1%2C+Marie&amp;rft.au=Kl%C3%ADma%2C+Ji%C5%99%C3%AD&amp;rft.au=Kratochv%C3%ADl%2C+Luk%C3%A1%C5%A1&amp;rft_id=https%3A%2F%2Fjournals.biologists.com%2Fdev%2Farticle%2F149%2F7%2Fdev200345%2F275038%2FPremeiotic-endoreplication-is-essential-for&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AGecko" class="Z3988"></span></span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="Further_reading">Further reading</h2></div> <ul><li>Forbes, Peter (4th Estate, London 2005) <i>The Gecko's Foot—Bio Inspiration: Engineered from Nature</i> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a>&#160;<a href="/wiki/Special:BookSources/0-00-717990-1" title="Special:BookSources/0-00-717990-1">0-00-717990-1</a> in H/B</li> <li>Zug, George. <a rel="nofollow" class="external text" href="http://www.sil.si.edu/smithsoniancontributions/Zoology/sc_RecordSingle.cfm?filename=SCtZ-0631"><i>Speciation and Dispersal in a Low Diversity Taxon: The Slender Geckos</i> Hemiphyllodactylus <i>(Reptilia, Gekkonidae)</i></a>. Smithsonian Contributions to Zoology, no. 631. Washington, D.C.: Smithsonian Institution Scholarly Press, 2010.</li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222" /><cite id="CITEREFGambleGreenbaumJackmanRussell2012" class="citation journal cs1">Gamble, T.; Greenbaum, E.; Jackman, T.R.; Russell, A.P.; Bauer, A.M. (2012). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3384654">"Repeated origin and loss of adhesive toepads in geckos"</a>. <i>PLOS ONE</i>. <b>7</b> (6): e39429. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2012PLoSO...739429G">2012PLoSO...739429G</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1371%2Fjournal.pone.0039429">10.1371/journal.pone.0039429</a></span>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a>&#160;<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3384654">3384654</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/22761794">22761794</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=PLOS+ONE&amp;rft.atitle=Repeated+origin+and+loss+of+adhesive+toepads+in+geckos&amp;rft.volume=7&amp;rft.issue=6&amp;rft.pages=e39429&amp;rft.date=2012&amp;rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3384654%23id-name%3DPMC&amp;rft_id=info%3Apmid%2F22761794&amp;rft_id=info%3Adoi%2F10.1371%2Fjournal.pone.0039429&amp;rft_id=info%3Abibcode%2F2012PLoSO...739429G&amp;rft.aulast=Gamble&amp;rft.aufirst=T.&amp;rft.au=Greenbaum%2C+E.&amp;rft.au=Jackman%2C+T.R.&amp;rft.au=Russell%2C+A.P.&amp;rft.au=Bauer%2C+A.M.&amp;rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3384654&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AGecko" 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title="wiktionary:gecko">gecko</a></b></i> in Wiktionary, the free dictionary.</div></div> </div> <ul><li><a rel="nofollow" class="external text" href="http://www.theinformationarchives.com/Gecko/">Gecko gallery and information</a></li> <li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20090214232218/http://lclark.edu/~autumn/dept/geckostory.html"><i>How Geckos Stick to Walls</i></a></li> <li><a rel="nofollow" class="external text" href="http://www.geckocare.net/">Comprehensive gecko care information</a></li> <li><a rel="nofollow" class="external text" href="http://www.gekkota.com">Global gecko association site with pictures, caresheets, species list</a></li> <li><a rel="nofollow" class="external text" href="https://www.flickr.com/photos/thirtypounces/54014101/">Gecko anatomy picture</a></li> <li><a rel="nofollow" class="external text" href="http://www.null-hypothesis.co.uk/science/reviews/books/geckos_foot_peter_forbes_reviewed"><i>The Gecko's Foot</i></a></li> <li><a rel="nofollow" class="external text" href="http://news.bbc.co.uk/1/hi/sci/tech/6967474.stm">Artificial gecko feet for a Spiderman suit (BBC 2007-08-28)</a></li> <li><a rel="nofollow" class="external text" href="http://www.geckotime.com/">Gecko Time Online <i>Gecko Magazine</i></a></li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl ol,.mw-parser-output .hlist dl ul,.mw-parser-output .hlist ol dl,.mw-parser-output .hlist ol ol,.mw-parser-output .hlist ol ul,.mw-parser-output .hlist ul dl,.mw-parser-output .hlist ul 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navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Gekkonidae" title="Gekkonidae">Gekkonidae</a></li> <li><a href="/wiki/Carphodactylidae" title="Carphodactylidae">Carphodactylidae</a></li> <li><a href="/wiki/Diplodactylidae" title="Diplodactylidae">Diplodactylidae</a></li> <li><a href="/wiki/Eublepharidae" title="Eublepharidae">Eublepharidae</a></li> <li><a href="/wiki/Phyllodactylidae" title="Phyllodactylidae">Phyllodactylidae</a></li> <li><a href="/wiki/Pygopodidae" title="Pygopodidae">Pygopodidae</a></li> <li><a href="/wiki/Sphaerodactylidae" title="Sphaerodactylidae">Sphaerodactylidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Scincoidea" class="mw-redirect" title="Scincoidea">Scincoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cordylidae" title="Cordylidae">Cordylidae</a></li> <li><a href="/wiki/Gerrhosauridae" title="Gerrhosauridae">Gerrhosauridae</a></li> <li><a href="/wiki/Skink" title="Skink">Scincidae</a></li> <li><a href="/wiki/Xantusiidae" class="mw-redirect" title="Xantusiidae">Xantusiidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Laterata" class="mw-redirect" title="Laterata">Laterata</a> / <a href="/wiki/Lacertoidea" title="Lacertoidea">Lacertoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Lacertidae" title="Lacertidae">Lacertidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/w/index.php?title=Gymnophthalmoidea&amp;action=edit&amp;redlink=1" class="new" title="Gymnophthalmoidea (page does not exist)">Gymnophthalmoidea</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Alopoglossidae" class="mw-redirect" title="Alopoglossidae">Alopoglossidae</a></li> <li><a href="/wiki/Gymnophthalmidae" title="Gymnophthalmidae">Gymnophthalmidae</a></li> <li><a href="/wiki/Teiidae" title="Teiidae">Teiidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Amphisbaenia" title="Amphisbaenia">Amphisbaenia</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Amphisbaenidae" title="Amphisbaenidae">Amphisbaenidae</a></li> <li><a href="/wiki/Bipedidae" title="Bipedidae">Bipedidae</a></li> <li><a href="/wiki/Blanidae" class="mw-redirect" title="Blanidae">Blanidae</a></li> <li><a href="/wiki/Cadeidae" class="mw-redirect" title="Cadeidae">Cadeidae</a></li> <li><a href="/wiki/Rhineuridae" title="Rhineuridae">Rhineuridae</a></li> <li><a href="/wiki/Trogonophidae" title="Trogonophidae">Trogonophidae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Anguimorpha" title="Anguimorpha">Anguimorpha</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Shinisauridae" class="mw-redirect" title="Shinisauridae">Shinisauridae</a></li> <li><a href="/wiki/Lanthanotidae" class="mw-redirect" title="Lanthanotidae">Lanthanotidae</a></li> <li><a href="/wiki/Varanidae" title="Varanidae">Varanidae</a></li> <li><a href="/wiki/Helodermatidae" title="Helodermatidae">Helodermatidae</a></li> <li><a href="/wiki/Xenosauridae" title="Xenosauridae">Xenosauridae</a></li> <li><a href="/wiki/Anguidae" title="Anguidae">Anguidae</a></li> <li><a href="/wiki/Anniellidae" class="mw-redirect" title="Anniellidae">Anniellidae</a></li> <li><a href="/wiki/Diploglossidae" title="Diploglossidae">Diploglossidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Iguania" class="mw-redirect" title="Iguania">Iguania</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Agamidae" title="Agamidae">Agamidae</a></li> <li><a href="/wiki/Chamaeleonidae" class="mw-redirect" title="Chamaeleonidae">Chamaeleonidae</a></li> <li><a href="/wiki/Corytophanidae" title="Corytophanidae">Corytophanidae</a></li> <li><a href="/wiki/Crotaphytidae" title="Crotaphytidae">Crotaphytidae</a></li> <li><a href="/wiki/Dactyloidae" title="Dactyloidae">Dactyloidae</a></li> <li><a href="/wiki/Hoplocercidae" title="Hoplocercidae">Hoplocercidae</a></li> <li><a href="/wiki/Iguanidae" title="Iguanidae">Iguanidae</a></li> <li><a href="/wiki/Leiocephalidae" class="mw-redirect" title="Leiocephalidae">Leiocephalidae</a></li> <li><a href="/wiki/Leiosauridae" title="Leiosauridae">Leiosauridae</a></li> <li><a href="/wiki/Liolaemidae" title="Liolaemidae">Liolaemidae</a></li> <li><a href="/wiki/Opluridae" title="Opluridae">Opluridae</a></li> <li><a href="/wiki/Phrynosomatidae" title="Phrynosomatidae">Phrynosomatidae</a></li> <li><a href="/wiki/Polychrotidae" title="Polychrotidae">Polychrotidae</a></li> <li><a href="/wiki/Tropiduridae" title="Tropiduridae">Tropiduridae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Serpentes" class="mw-redirect" title="Serpentes">Serpentes</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Scolecophidia" title="Scolecophidia">Scolecophidia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Anomalepididae" title="Anomalepididae">Anomalepididae</a></li> <li><a href="/wiki/Gerrhopilidae" title="Gerrhopilidae">Gerrhopilidae</a></li> <li><a href="/wiki/Leptotyphlopidae" title="Leptotyphlopidae">Leptotyphlopidae</a></li> <li><a href="/wiki/Typhlopidae" title="Typhlopidae">Typhlopidae</a></li> <li><a href="/wiki/Xenotyphlopidae" class="mw-redirect" title="Xenotyphlopidae">Xenotyphlopidae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Alethinophidia" title="Alethinophidia">Alethinophidia</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Acrochordidae" class="mw-redirect" title="Acrochordidae">Acrochordidae</a></li> <li><a href="/wiki/Aniliidae" class="mw-redirect" title="Aniliidae">Aniliidae</a></li> <li><a href="/wiki/Anomochilidae" class="mw-redirect" title="Anomochilidae">Anomochilidae</a></li> <li><a href="/wiki/Atractaspididae" title="Atractaspididae">Atractaspididae</a></li> <li><a href="/wiki/Boidae" title="Boidae">Boidae</a></li> <li><a href="/wiki/Bolyeriidae" title="Bolyeriidae">Bolyeriidae</a></li> <li><a href="/wiki/Colubridae" title="Colubridae">Colubridae</a></li> <li><a href="/wiki/Cylindrophis" title="Cylindrophis">Cylindrophiidae</a></li> <li><a href="/wiki/Elapidae" title="Elapidae">Elapidae</a></li> <li><a href="/wiki/Loxocemidae" class="mw-redirect" title="Loxocemidae">Loxocemidae</a></li> <li><a href="/wiki/Pythonidae" title="Pythonidae">Pythonidae</a></li> <li><a href="/wiki/Tropidophiidae" title="Tropidophiidae">Tropidophiidae</a></li> <li><a href="/wiki/Uropeltidae" title="Uropeltidae">Uropeltidae</a></li> <li><a href="/wiki/Viper" title="Viper">Viperidae</a></li> <li><a href="/wiki/Xenopeltidae" class="mw-redirect" title="Xenopeltidae">Xenopeltidae</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374" /><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235" /></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers1714" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers1714" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Gekkota</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q1008888" class="extiw" title="wikidata:Q1008888">Q1008888</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Gekkota" class="extiw" title="wikispecies:Gekkota">Gekkota</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Australian_Faunal_Directory" title="Australian Faunal Directory">AFD</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://biodiversity.org.au/afd/taxa/Gekkota">Gekkota</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Catalogue_of_Life" title="Catalogue of Life">CoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.catalogueoflife.org/data/taxon/87BW6">87BW6</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/2819161">2819161</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/INaturalist" title="INaturalist">iNaturalist</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://inaturalist.org/taxa/371057">371057</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Integrated_Taxonomic_Information_System" title="Integrated Taxonomic Information System">ITIS</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&amp;search_value=564528">564528</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&amp;id=8560">8560</a></span></span></li> <li><span style="white-space:nowrap;">NZOR: <span class="uid"><a rel="nofollow" class="external text" href="https://www.nzor.org.nz/names/ce4b96cb-960f-4f16-8c48-36916207d53d">ce4b96cb-960f-4f16-8c48-36916207d53d</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=92743">92743</a></span></span></li></ul> 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