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<!doctype html> <html class="no-js" xml:lang="en" lang="en" dir="ltr"> <head> <meta http-equiv="X-UA-Compatible" content="IE=Edge" /> <title>Neuroscience (PhD) | UC San Francisco</title> <meta http-equiv="Content-Type" content="text/html; charset=utf-8" /> <link rel="search" type="application/opensearchdescription+xml" href="/search/opensearch.xml" title="Catalog" /> <meta name="viewport" content="width=device-width, initial-scale=1.0, minimum-scale=1.0" /> <link href="/images/favicon.ico" rel="shortcut icon" /> <link rel="stylesheet" type="text/css" href="/css/reset.css" /> <link href="https://fonts.googleapis.com/css?family=Roboto:400,400i,500,500i,700,700i" rel="stylesheet"> <link rel="stylesheet" type="text/css" href="/fonts/font-awesome/font-awesome.min.css" /> <link rel="stylesheet" type="text/css" href="/css/courseleaf.css" /> <link rel="stylesheet" type="text/css" href="/css/screen.css?v=2024-2025" media="screen" /> <link rel="stylesheet" type="text/css" href="/css/print.css" media="print" /> <script type="text/javascript" src="/js/jquery.js"></script> <script type="text/javascript" src="/js/lfjs.js"></script> <script type="text/javascript" src="/js/lfjs_any.js"></script> <link rel="stylesheet" type="text/css" href="/js/lfjs.css" /> <script type="text/javascript" src="/js/courseleaf.js"></script> <script type="text/javascript" src="/js/custom.js?v=2024-2025"></script> </head> <body class=""> <nav aria-label="Skip content menu" class="accessible"> <div class="accessible-menu"> <ul> <li><a href="#contentarea" rel="section">Skip to Content</a></li> <li><a href="/azindex/">AZ Index</a></li> <li><a href="/">Catalog Home</a></li> </ul> </div> </nav> <header id="header"> <section class="header-universal"> <div class="wrap"> <p class="header-universal__title">University of California San Francisco</p> <a class="header-universal__give" href="https://giving.ucsf.edu/">Give to UCSF</a> </div> </section> <section class="header-main"> <div class="wrap"> <h1 id="site-title"><a href="/" rel="home">General Catalog</a></h1> <div class="mobile-toggles"> <button type="button" class="toggle-search" aria-controls="cat-search" aria-expanded="false" data-toggle="#cat-search"> <span class="sr-only">toggle search</span> <span class="toggle-search__icon" aria-hidden="true"></span> </button> <button id="hamburger" aria-controls="navigation" data-toggle="#navigation" aria-expanded="false"> <span class="sr-only">Toggle menu</span> <span class="burgertime" aria-hidden="true"> <span class="icon"></span> </span> </button> </div> <nav id="navigation" aria-label="Secondary" aria-hidden="false"> <ul id="main-nav" class="top clearfix"> <li class="isparent"> <button type="button">Courses</button> <ul> <li><a href="/course-catalog/">Course Listings</a></li> <li><a href="/course-search/">Search Courses</a></li> </ul> </li> <li><a href="/programs/">Academic Programs</a></li> <li class="isparent"> <button type="button">Schools and Programs</button> <ul> <li><a href="/schools/school-dentistry/">School of Dentistry</a></li> <li><a href="/schools/school-medicine/">School of Medicine</a></li> <li><a href="/schools/school-nursing/">School of Nursing</a></li> <li><a href="/schools/school-pharmacy/">School of Pharmacy</a></li> <li><a href="/schools/graduate-division/">Graduate Division</a></li> <li><a href="/schools/institute-global-health-sciences/">Institute for Global Health Sciences</a></li> </ul> </li> <li><a href="/university/">University Support Offices</a></li> <li><a href="/archive/">Archive</a></li> <li class="no-mobile"> <button type="button" class="toggle-search" aria-controls="cat-search" aria-expanded="false" data-toggle="#cat-search"> <span class="sr-only">open search</span> <span class="toggle-search__icon" aria-hidden="true"></span> </button> </li> </ul> <a class="header-main__give" href="https://giving.ucsf.edu/">Give to UCSF</a> </nav> <div class="search" id="cat-search"> <form action="/search/"> <label for="cat-search-term" class="sr-only">Search catalog</label> <input class="search-field" type="text" name="search" id="cat-search-term" placeholder="Search catalog..." /> <button class="search-button" type="submit"> <span class="sr-only">Submit search</span> </button>  </form> </div> </div> </section> </header> <section id="content-container"> <div class="wrap"> <div id="col-content"> <nav id="breadcrumb" aria-label="Breadcrumbs"> <ul><li><a href="/">Home</a><span class="crumbsep">></span></li><li><a href="/programs/">Academic Programs</a><span class="crumbsep">></span></li><li><span class="active">Neuroscience (PhD)</span></li></ul> </nav> <main id="contentarea"> <header> <h1 class="page-title">Neuroscience (PhD)</h1> </header> <script type="text/javascript"> //<![CDATA[ var defshow = "text"; var bodycontainer = "#body"; var validhashes = ",text,requirementstext,"; if(window.location.hash.length) { var hash = cleanHash(window.location.hash); if(validhashes.indexOf("," + hash + ",") != -1) defshow = hash; } var ss = document.styleSheets; var workss = null; for(var i=0; i < ss.length; i++) { var currmedia = ""; if (typeof ss[i].media == "string") currmedia = ss[i].media else if (typeof ss[i].media == "object" && ss[i].media.mediaText) currmedia = ss[i].media.mediaText; if((currmedia == "screen" || currmedia == "all") && (ss[i].href.indexOf("/") == 0 || ss[i].href.indexOf(document.location.host) >= 0)) { workss = ss[i]; break; } } if(workss && workss.insertRule) { workss.insertRule(".tab_content { display: none; }", workss.cssRules.length) workss.insertRule(".sectionheader { display: none; }", workss.cssRules.length) workss.insertRule("#" + defshow + "container { display: block; }", workss.cssRules.length) } else if(workss && workss.addRule) { workss.addRule(".tab_content", "display: none"); workss.addRule(".sectionheader", "display: none"); workss.addRule("#" + defshow + "container", "display: block"); } else { if (window.console) window.console.log('screen.css <link> needs attribute media="screen"'); } //]]> </script> <nav id="tabs" aria-label="Page content tabs"> <ul class="clearfix" role="tablist"> <li id="texttab" role="presentation"> <a href="#textcontainer" role="tab" aria-controls="textcontainer" onclick="return showSection('text');">Overview</a></li> <li id="requirementstexttab" role="presentation"> <a href="#requirementstextcontainer" role="tab" aria-controls="requirementstextcontainer" onclick="return showSection('requirementstext');" >Details</a></li> </ul> </nav> <div id="textcontainer" class="page_content tab_content" role="tabpanel" aria-labelledby="texttab"> <p><a href="http://neurograd.ucsf.edu" target="_blank" rel="noopener">Visit program website.</a></p><p><a href="https://gradapplication.ucsf.edu/" target="_blank" rel="noopener" class="applyBtn">Apply now</a></p><p><strong>Degree Offered:</strong> PhD<br/> <strong>Program Leadership:</strong><br/> Samuel Pleasure, MD, PhD, Program Director<br/> <strong>Admissions Inquiries:</strong><br/> Neuroscience Graduate Program, <a href="mailto:neurosciadmin@listsrv.ucsf.edu">neurosciadmin@listsrv.ucsf.edu</a></p> <h2>Program Description</h2> <p>The Neuroscience PhD program prepares students for independent research and teaching in neuroscience. It seeks to train students who will be expert in one particular approach to neuroscientific research, but who will also have a strong general background in other areas of neuroscience and related disciplines. To achieve this objective, students take interdisciplinary core and advanced courses in neuroscience, as well as related courses sponsored by other graduate programs. In addition, students carry out research under the supervision of faculty members in the program.</p> <p>The UCSF Neuroscience program faculty, who are among the world leaders in their respective areas of neuroscience, utilize innovative cellular, computational, electrophysiological, genetic, imaging, and molecular strategies to address outstanding problems in neuroscience. These approaches are employed in an integrative manner to engage in research in all areas of neuroscience, including behavior, biophysics, cell biology, development, neural systems, and disorders of the nervous system. The collaborative nature of the UCSF environment offers a unique opportunity for students to take advantage of the interdisciplinary nature of research at the frontier of modern neuroscience.</p> <p>The UCSF Neuroscience Program is committed to a diverse and inclusive graduate student population, which enhances the educational experience, the workplace, and the nature of scientific research.</p> <h3>Faculty</h3> <p>The Neuroscience program currently has 100 faculty members from the following departments and areas: Anatomy, Biochemistry and Biophysics, Bioengineering, the Brain and Spinal Cord Injury Center, Cell and Tissue Biology, CMP, CVRI, the Diabetes Center, IND, Gladstone Institutes, Neurological Surgery, Neurology, the Neuroscience Imaging Center, Ophthalmology, Oral and Maxillofacial Surgery, Otolaryngology, Pathology, Pediatrics-Medical Genetics, Pediatrics-Neonatology, Pharmaceutical Chemistry, Physical Therapy, Physiology, and Psychiatry.</p> <p>The Neuroscience program is a member of the <a href="https://pibs.ucsf.edu/" target="_blank">Program in Biological Sciences</a> (PIBS).</p> <h3>Sub-Disciplines</h3> <ul> <li>Addiction Research</li> <li><span>Behavioral Neuroscience</span></li> <li><span>Cell and Molecular Neuroscience</span></li> <li><span>Developmental Neuroscience</span></li> <li><span>Nervous System Disease</span></li> <li><span>Synaptic Physiology</span></li> <li><span>Systems Neuroscience</span></li> <li><span>Theoretical/Computational Neuroscience</span></li> </ul> <p>The Neuroscience program is based primarily at Mission Bay, but also has faculty at Parnassus, the UCSF affiliated Veterans Administration Medical Center, and San Francisco General Hospital. Visit the <a href="https://neurograd.ucsf.edu/" target="_blank">program website</a> for more information.</p> <p>The Neuroscience program is offered by the UCSF Graduate Division, administered by the UCSF School of Medicine, and delivered by faculty members in the UCSF schools of dentistry, medicine, and pharmacy.</p> <h2>Admission Requirements</h2> <ul> <li>Undergraduate courses in biology, physics, inorganic and organic chemistry, and calculus.</li> <li>Prior research experience (undergraduate thesis projects, internships, technician positions, summer research programs, etc.)</li> </ul> <h2>Learning Outcomes</h2> <ol> <li>General knowledge and understanding of modern neuroscience.</li> <li>Theoretical understanding of a broad range of modern neuroscientific techniques.</li> <li>Ability to read, critically assess, and review contemporary scientific literature.</li> <li>Ability to identify outstanding questions in the field, and to design experiments that are ethical and viable to advance the field.</li> <li>Practical expertise in techniques relevant to a subfield of neuroscientific research.</li> <li>Capacity to produce high-quality grant proposals and scientific papers.</li> <li>Clear communication of scientific research to both scientific and non-scientific audiences.</li> <li>Conceptualization of the importance of mentorship, the variety of needs that accompany students of different backgrounds, and demonstration of mentorship ability.</li> </ol> <h2>Additional Information</h2> <h3>Program Core Faculty</h3> <ul> <li><em>Find a <a href="https://neurograd.ucsf.edu/faculty" target="_blank">program faculty list</a> on the program website.</em></li> </ul> <h3>Career Outcomes</h3> <ul> <li><em>Find <a href="https://graduate.ucsf.edu/program-statistics#career" target="_blank">career outcomes and other data on PhD programs</a> on the Graduate Division website.</em></li> </ul></div> <div id="requirementstextcontainer" class="page_content tab_content" role="tabpanel" aria-labelledby="requirementstexttab" aria-hidden="true"> <a name="requirementstext"></a><h2>Degree Requirements</h2> <p>In order to assess the successful completion of the goals set out for students of the Neuroscience program, the program requires:</p> <ol> <li>An agreement of the student鈥檚 thesis committee that the student has demonstrated sufficient aptitude toward the above goals, as well as</li> <li>The production of a graduate thesis in the form of a dissertation that demonstrates: <ol start="1" type="1"> <li>Minimum GPA of 3.0</li> <li>All core courses and required activities taken and passed</li> <li>Six quarters in residence including a minimum of three registered quarters after advancement to candidacy</li> <li>Pass qualifying examination</li> <li>Completion and submission of the dissertation</li> <li>For additional details, please see <a href="https://graduate.ucsf.edu/phd-degree" target="_blank">graduate.ucsf.edu/phd-degree</a><br/> <br/> In addition to the dissertation, students are required to:</li> <li>Hold a thesis seminar, in which a curated portion of the dissertation is presented to an audience consisting of both neuroscientists and non-neuroscientists.</li> <li>Have published, or have begun the process of publishing, a first-author scientific paper in a reputable peer-reviewed journal (under exceptional circumstances the thesis committee may assess the adequacy for graduation based on meeting all the other requirements for graduation without a first-author paper).</li> </ol> </li> </ol> <h2>Core Courses</h2> <table class="sc_courselist"> <caption class="hidden noscript">Course List</caption> <colgroup> <col class="codecol"/> <col class="titlecol"/> <col align="char" char="." class="hourscol"/> </colgroup> <thead> <tr class="hidden noscript"> <th scope="col">Code</th> <th scope="col">Title</th> <th scope="col" class="hourscol">Units</th> </tr> </thead> <tbody> <tr class="even firstrow"><td class="codecol"><a href="/search/?P=NEUROSCI%20200" title="NEUROSCI聽200" class="bubblelink code" onclick="return showCourse(this, 'NEUROSCI 200');">NEUROSCI聽200</a></td><td>Introduction to Neuroscience. Essential Concepts & Methods</td><td class="hourscol">2.5</td></tr> <tr class="odd"><td class="codecol"><a href="/search/?P=NEUROSCI%20201A" title="NEUROSCI聽201A" class="bubblelink code" onclick="return showCourse(this, 'NEUROSCI 201A');">NEUROSCI聽201A</a></td><td>Basic Concepts in Cellular and Molecular Neuroscience</td><td class="hourscol">5</td></tr> <tr class="even"><td class="codecol"><a href="/search/?P=NEUROSCI%20201B" title="NEUROSCI聽201B" class="bubblelink code" onclick="return showCourse(this, 'NEUROSCI 201B');">NEUROSCI聽201B</a></td><td>Basic Concepts for Cellular and Developmental Neuroscience</td><td class="hourscol">4</td></tr> <tr class="odd"><td class="codecol"><a href="/search/?P=NEUROSCI%20201C" title="NEUROSCI聽201C" class="bubblelink code" onclick="return showCourse(this, 'NEUROSCI 201C');">NEUROSCI聽201C</a></td><td>Introduction to Systems and Behavioral Neuroscience</td><td class="hourscol">4</td></tr> <tr class="even"><td class="codecol"><a href="/search/?P=NEUROSCI%20215" title="NEUROSCI聽215" class="bubblelink code" onclick="return showCourse(this, 'NEUROSCI 215');">NEUROSCI聽215</a></td><td>Laboratory Rotation</td><td class="hourscol">8-12</td></tr> <tr class="odd"><td class="codecol"><a href="/search/?P=NEUROSCI%20221" title="NEUROSCI聽221" class="bubblelink code" onclick="return showCourse(this, 'NEUROSCI 221');">NEUROSCI聽221</a></td><td>Current Topics in Neuroscience</td><td class="hourscol">1</td></tr> <tr class="even"><td class="codecol"><a href="/search/?P=NEUROSCI%20250" title="NEUROSCI聽250" class="bubblelink code" onclick="return showCourse(this, 'NEUROSCI 250');">NEUROSCI聽250</a></td><td>Research</td><td class="hourscol">1-8</td></tr> <tr class="odd"><td class="codecol"><a href="/search/?P=GRAD%20214" title="GRAD聽214" class="bubblelink code" onclick="return showCourse(this, 'GRAD 214');">GRAD聽214</a></td><td>Responsible Conduct of Research and Rigor & Reproducibility</td><td class="hourscol">1.5</td></tr> <tr class="even lastrow"><td class="codecol"><a href="/search/?P=GRAD%20215" title="GRAD聽215" class="bubblelink code" onclick="return showCourse(this, 'GRAD 215');">GRAD聽215</a></td><td>Managing your Research for Reproducibility</td><td class="hourscol">1</td></tr> <tr class="listsum"><td colspan="2">Total Units</td><td class="hourscol">28-39</td></tr></tbody> </table> <h3>Approved Electives</h3> <table class="sc_courselist"> <caption class="hidden noscript">Course List</caption> <colgroup> <col class="codecol"/> <col class="titlecol"/> <col align="char" char="." class="hourscol"/> </colgroup> <thead> <tr class="hidden noscript"> <th scope="col">Code</th> <th scope="col">Title</th> <th scope="col" class="hourscol">Units</th> </tr> </thead> <tbody> <tr class="even firstrow"><td class="codecol"><a href="/search/?P=NEUROSCI%20219" title="NEUROSCI聽219" class="bubblelink code" onclick="return showCourse(this, 'NEUROSCI 219');">NEUROSCI聽219</a></td><td>Special Topics in Basic and Translational Neuroscience</td><td class="hourscol">3</td></tr> <tr class="odd"><td class="codecol"><a href="/search/?P=NEUROSCI%20220" title="NEUROSCI聽220" class="bubblelink code" onclick="return showCourse(this, 'NEUROSCI 220');">NEUROSCI聽220</a></td><td>Neuroscience Journal Club</td><td class="hourscol">1</td></tr> <tr class="even"><td class="codecol"><a href="/search/?P=BIOMED%20SCI%20270" title="BIOMED聽SCI聽270" class="bubblelink code" onclick="return showCourse(this, 'BIOMED SCI 270');">BIOMED聽SCI聽270</a></td><td>Special Topics in Biomedical Sciences</td><td class="hourscol">3</td></tr> <tr class="odd"><td class="codecol"><a href="/search/?P=BIOPHYSICS%20219" title="BIOPHYSICS聽219" class="bubblelink code" onclick="return showCourse(this, 'BIOPHYSICS 219');">BIOPHYSICS聽219</a></td><td>Special Topics in Biophysics</td><td class="hourscol">3</td></tr> <tr class="even"><td class="codecol"><a href="/search/?P=CHEMISTRY%20219" title="CHEMISTRY聽219" class="bubblelink code" onclick="return showCourse(this, 'CHEMISTRY 219');">CHEMISTRY聽219</a></td><td>Special Topics in Basic and Translational Chemical Biology</td><td class="hourscol">3</td></tr> <tr class="odd"><td class="codecol"><a href="/search/?P=DEV%20STMCEL%20270" title="DEV聽STMCEL聽270" class="bubblelink code" onclick="return showCourse(this, 'DEV STMCEL 270');">DEV聽STMCEL聽270</a></td><td>Special Topics in Developmental & Stem Cell Biology</td><td class="hourscol">3</td></tr> <tr class="even"><td class="codecol"><a href="/search/?P=GRAD%20202" title="GRAD聽202" class="bubblelink code" onclick="return showCourse(this, 'GRAD 202');">GRAD聽202</a></td><td>Racism in Science</td><td class="hourscol">3</td></tr> <tr class="odd"><td class="codecol"><a href="/search/?P=GRAD%20210" title="GRAD聽210" class="bubblelink code" onclick="return showCourse(this, 'GRAD 210');">GRAD聽210</a></td><td>Justice, Equity, Diversity and Inclusion Academic Leadership</td><td class="hourscol">4</td></tr> <tr class="even"><td class="codecol"><a href="/search/?P=GRAD%20213" title="GRAD聽213" class="bubblelink code" onclick="return showCourse(this, 'GRAD 213');">GRAD聽213</a></td><td>Motivating INformed Decisions (MIND) Catalytic Course</td><td class="hourscol">2</td></tr> <tr class="odd"><td class="codecol"><a href="/search/?P=GRAD%20286" title="GRAD聽286" class="bubblelink code" onclick="return showCourse(this, 'GRAD 286');">GRAD聽286</a></td><td>GSICE Curricular Practicum</td><td class="hourscol">1</td></tr> <tr class="even"><td class="codecol"><a href="/search/?P=PHARMGENOM%20219" title="PHARMGENOM聽219" class="bubblelink code" onclick="return showCourse(this, 'PHARMGENOM 219');">PHARMGENOM聽219</a></td><td>Special Topics in Pharm Sci and Pharmacogenomics</td><td class="hourscol">3</td></tr> <tr class="odd lastrow"><td colspan="2"><span class="courselistcomment">Other elective courses may be approved by the program as appropriate.</span></td><td class="hourscol"></td></tr> </tbody> </table> <h2>Non-course Core Requirements</h2> 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