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data-event-name="pinnable-header.vector-toc.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-toc.unpin">hide</button> </div> <ul class="vector-toc-contents" id="mw-panel-toc-list"> <li id="toc-mw-content-text" class="vector-toc-list-item vector-toc-level-1"> <a href="#" class="vector-toc-link"> <div class="vector-toc-text">(Top)</div> </a> </li> <li id="toc-Comparing_Respiratory_Pigments" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Comparing_Respiratory_Pigments"> <div class="vector-toc-text"> <span class="vector-toc-numb">1</span> <span>Comparing Respiratory Pigments</span> </div> </a> <ul id="toc-Comparing_Respiratory_Pigments-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Globins" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Globins"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Globins</span> </div> </a> <button aria-controls="toc-Globins-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Globins subsection</span> </button> <ul id="toc-Globins-sublist" class="vector-toc-list"> <li id="toc-Vertebrate_hemoglobin" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Vertebrate_hemoglobin"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.1</span> <span>Vertebrate hemoglobin</span> </div> </a> <ul id="toc-Vertebrate_hemoglobin-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Other_animal_hemoglobins" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Other_animal_hemoglobins"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.2</span> <span>Other animal hemoglobins</span> </div> </a> <ul id="toc-Other_animal_hemoglobins-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Leghemoglobin" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Leghemoglobin"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.3</span> <span>Leghemoglobin</span> </div> </a> <ul id="toc-Leghemoglobin-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Hemocyanin" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Hemocyanin"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Hemocyanin</span> </div> </a> <ul id="toc-Hemocyanin-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Notes" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Notes"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Notes</span> </div> </a> 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<div class="mw-indicators"> </div> <div id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Metalloprotein</div> <p>A <b>respiratory pigment</b> is a <a href="/wiki/Metalloprotein" title="Metalloprotein">metalloprotein</a> that serves a variety of important functions, its main being O<sub>2</sub> transport.<sup id="cite_ref-:0_1-0" class="reference"><a href="#cite_note-:0-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> Other functions performed include O<sub>2</sub> storage, CO<sub>2</sub> transport, and transportation of substances other than respiratory gases. There are four major classifications of respiratory pigment: <a href="/wiki/Hemoglobin" title="Hemoglobin">hemoglobin</a>, <a href="/wiki/Hemocyanin" title="Hemocyanin">hemocyanin</a>, erythrocruorin–<a href="/wiki/Chlorocruorin" class="mw-redirect" title="Chlorocruorin">chlorocruorin</a>, and <a href="/wiki/Hemerythrin" title="Hemerythrin">hemerythrin</a>. The heme-containing globin<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>a<span class="cite-bracket">]</span></a></sup> is the most commonly-occurring respiratory pigment, occurring in at least 9 different phyla of animals.<sup id="cite_ref-:1_3-0" class="reference"><a href="#cite_note-:1-3"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Comparing_Respiratory_Pigments">Comparing Respiratory Pigments</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Respiratory_pigment&action=edit&section=1" title="Edit section: Comparing Respiratory Pigments"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <table class="wikitable"> <caption> </caption> <tbody><tr> <th rowspan="2">Metalloprotein </th> <th colspan="2">Globins </th> <th rowspan="2">Hemocyanin </th> <th rowspan="2">Hemerythrin </th></tr> <tr> <th>Hemoglobin<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>b<span class="cite-bracket">]</span></a></sup> </th> <th>Erythrocruorin and chlorocruorin </th></tr> <tr> <th scope="row">O<sub>2</sub> Binding Material </th> <td>Iron<sup id="cite_ref-:2_5-0" class="reference"><a href="#cite_note-:2-5"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </td> <td>Iron<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </td> <td>Copper<sup id="cite_ref-:2_5-1" class="reference"><a href="#cite_note-:2-5"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </td> <td>Iron<sup id="cite_ref-:2_5-2" class="reference"><a href="#cite_note-:2-5"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </td></tr> <tr> <th scope="row">Location </th> <td>Intracellular<sup id="cite_ref-:1_3-1" class="reference"><a href="#cite_note-:1-3"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </td> <td>Extracellular<sup id="cite_ref-:3_7-0" class="reference"><a href="#cite_note-:3-7"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </td> <td>Extracellular<sup id="cite_ref-:1_3-2" class="reference"><a href="#cite_note-:1-3"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </td> <td>Intracellular<sup id="cite_ref-:1_3-3" class="reference"><a href="#cite_note-:1-3"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </td></tr> <tr> <th scope="row">Source Organism </th> <td>Almost all <a href="/wiki/Vertebrate" title="Vertebrate">vertebrates</a><sup id="cite_ref-:1_3-4" class="reference"><a href="#cite_note-:1-3"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </td> <td><style data-mw-deduplicate="TemplateStyles:r1126788409">.mw-parser-output .plainlist ol,.mw-parser-output .plainlist ul{line-height:inherit;list-style:none;margin:0;padding:0}.mw-parser-output .plainlist ol li,.mw-parser-output .plainlist ul li{margin-bottom:0}</style><div class="plainlist"> <ul><li>Annelids and arthropods</li> <li>Chlorocruorin: 4 families of marine <a href="/wiki/Polychaete" title="Polychaete">polychaetes</a><sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup></li></ul> </div> </td> <td><a href="/wiki/Arthropod" title="Arthropod">Arthropoda</a> and <a href="/wiki/Mollusca" title="Mollusca">Mollusca</a><sup id="cite_ref-:1_3-5" class="reference"><a href="#cite_note-:1-3"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </td> <td> <p><a href="/wiki/Sipuncula" title="Sipuncula">Sipuncula</a>, <a href="/wiki/Priapulida" title="Priapulida">priapulida</a>, some <a href="/wiki/Brachiopod" title="Brachiopod">brachiopoda</a>, and a single <a href="/wiki/Annelid" title="Annelid">annelid</a> genus <sup id="cite_ref-:2_5-3" class="reference"><a href="#cite_note-:2-5"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p> </td></tr> <tr> <th scope="row">Oxygenated Color </th> <td>Bright red<sup id="cite_ref-:0_1-1" class="reference"><a href="#cite_note-:0-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </td> <td><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"><div class="plainlist"> <ul><li>Erythrocruorin: Bright red</li> <li>Chlorocruorin: Green when diluted, red when concentrated <sup id="cite_ref-:3_7-1" class="reference"><a href="#cite_note-:3-7"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup></li></ul> </div> </td> <td>Blue<sup id="cite_ref-:0_1-2" class="reference"><a href="#cite_note-:0-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </td> <td>Violet<sup id="cite_ref-:0_1-3" class="reference"><a href="#cite_note-:0-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </td></tr> <tr> <th scope="row">Deoxygenated Color </th> <td>Crimson<sup id="cite_ref-:0_1-4" class="reference"><a href="#cite_note-:0-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </td> <td><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1126788409"><div class="plainlist"> <ul><li>Erythrocruorin: Dark red</li> <li>Chlorocruorin: Green when diluted, brown-red when concentrated</li></ul> </div> </td> <td>Colorless<sup id="cite_ref-:0_1-5" class="reference"><a href="#cite_note-:0-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </td> <td>Colorless<sup id="cite_ref-:0_1-6" class="reference"><a href="#cite_note-:0-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </td></tr></tbody></table> <p>Hemoglobin, <a href="/wiki/Erythrocruorin" title="Erythrocruorin">erythrocruorin</a>, and chlorocruorin are all <a href="/wiki/Globin" title="Globin">globins</a>, iron-heme proteins with a common core. Their color comes from the absorption spectra of <a href="/wiki/Heme" title="Heme">heme</a> with Fe<sup>2+</sup>. Erythrocruorin and chlorocruorin are closely related giant globins found used by some invertebrates. Chlorocruorin has a special heme group, giving it different colors. </p><p><span class="citation-needed-content" style="padding-left:0.1em; padding-right:0.1em; color:var(--color-subtle, #54595d); border:1px solid var(--border-color-subtle, #c8ccd1);">Any of various coloured conjugated proteins, such as hemoglobin, occur in living organisms and function in oxygen transfer in cellular respiration.</span><sup class="noprint Inline-Template Template-Fact" style="margin-left:0.1em; white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (October 2019)">citation needed</span></a></i>]</sup> </p> <div class="mw-heading mw-heading2"><h2 id="Globins">Globins</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Respiratory_pigment&action=edit&section=2" title="Edit section: Globins"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The globin is thought to be a very ancient molecule, even acting as a molecular clock of sorts. It has even been used to date the separation of vertebrates and invertebrates more than 1 billion years ago. Globin enjoys a large biological distribution, not only occurring among more than 9 different phyla of animals but occurring in some fungi and bacteria as well, even being identified in nitrogen-fixing nodules on the roots of some leguminous plants. The isolation of the globin gene from plant root cells has suggested that the globin genes that were inherited from a common ancestor shared by plants and animals may be present in all plants.<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Vertebrate_hemoglobin">Vertebrate hemoglobin</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Respiratory_pigment&action=edit&section=3" title="Edit section: Vertebrate hemoglobin"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Vertebrates use a tetrameric hemoglobin, carried in red blood cells, to breathe. There are multiple types of hemoglobin that have been found in the human body alone. Hemoglobin A is the “normal” hemoglobin, the variant of hemoglobin that is most common after birth. Hemoglobin A2 is a minor component of hemoglobin found in red blood cells. Hemoglobin A2 makes up less than 3% of total red blood cell hemoglobin. Hemoglobin F typically is only found in the fetal stage of development. While Hemoglobin F falls dramatically after birth, it is possible for some people to produce some levels of Hemoglobin F throughout their full life.<sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Other_animal_hemoglobins">Other animal hemoglobins</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Respiratory_pigment&action=edit&section=4" title="Edit section: Other animal hemoglobins"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1251242444">.mw-parser-output .ambox{border:1px solid #a2a9b1;border-left:10px solid #36c;background-color:#fbfbfb;box-sizing:border-box}.mw-parser-output .ambox+link+.ambox,.mw-parser-output .ambox+link+style+.ambox,.mw-parser-output .ambox+link+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+style+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+link+.ambox{margin-top:-1px}html body.mediawiki .mw-parser-output .ambox.mbox-small-left{margin:4px 1em 4px 0;overflow:hidden;width:238px;border-collapse:collapse;font-size:88%;line-height:1.25em}.mw-parser-output .ambox-speedy{border-left:10px solid #b32424;background-color:#fee7e6}.mw-parser-output .ambox-delete{border-left:10px solid #b32424}.mw-parser-output .ambox-content{border-left:10px solid #f28500}.mw-parser-output .ambox-style{border-left:10px solid #fc3}.mw-parser-output .ambox-move{border-left:10px solid #9932cc}.mw-parser-output .ambox-protection{border-left:10px solid #a2a9b1}.mw-parser-output .ambox .mbox-text{border:none;padding:0.25em 0.5em;width:100%}.mw-parser-output .ambox .mbox-image{border:none;padding:2px 0 2px 0.5em;text-align:center}.mw-parser-output .ambox .mbox-imageright{border:none;padding:2px 0.5em 2px 0;text-align:center}.mw-parser-output .ambox .mbox-empty-cell{border:none;padding:0;width:1px}.mw-parser-output .ambox .mbox-image-div{width:52px}@media(min-width:720px){.mw-parser-output .ambox{margin:0 10%}}@media print{body.ns-0 .mw-parser-output .ambox{display:none!important}}</style><table class="box-Missing_information plainlinks metadata ambox ambox-content" role="presentation"><tbody><tr><td class="mbox-image"><div class="mbox-image-div"><span typeof="mw:File"><a href="/wiki/File:Wiki_letter_w.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/en/thumb/6/6c/Wiki_letter_w.svg/44px-Wiki_letter_w.svg.png" decoding="async" width="44" height="44" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/6/6c/Wiki_letter_w.svg/66px-Wiki_letter_w.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/6/6c/Wiki_letter_w.svg/88px-Wiki_letter_w.svg.png 2x" data-file-width="44" data-file-height="44" /></a></span></div></td><td class="mbox-text"><div class="mbox-text-span">This section <b>is missing information</b> about phylogeny link: is the "tetrameric intracellular" group monophyletic? What about the others?.<span class="hide-when-compact"> Please expand the section to include this information. Further details may exist on the <a href="/wiki/Talk:Respiratory_pigment" title="Talk:Respiratory pigment">talk page</a>.</span> <span class="date-container"><i>(<span class="date">November 2020</span>)</i></span></div></td></tr></tbody></table> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">Further information: <a href="/wiki/Hemoglobin#Analogues_in_non-vertebrate_organisms" title="Hemoglobin">Hemoglobin § Analogues in non-vertebrate organisms</a></div> <p>Animals use a great variety of globins for respiration. By structure, they can be classified as:<sup id="cite_ref-Weber2001_11-0" class="reference"><a href="#cite_note-Weber2001-11"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup><sup class="reference nowrap"><span title="Location: Fig. 1">: Fig. 1 </span></sup> </p> <ul><li>Intracellular Hbs. These globins reside inside a cell, much like the vertebrate Hb.</li> <li>Multi-subunit Hbs. These globins form complexes and work outside a cell.</li> <li>Multi-domain, multisubunit Hbs. These globins form complexes, work outside a cell, and have multiple globin domains per peptide chain.</li></ul> <p>Erythrocruorin and chlorocruorin belong to the multisubunit Hbs, specifically of the 12-dodecamer type. </p> <div class="mw-heading mw-heading3"><h3 id="Leghemoglobin">Leghemoglobin</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Respiratory_pigment&action=edit&section=5" title="Edit section: Leghemoglobin"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">See also: <a href="/wiki/Phytoglobin" title="Phytoglobin">phytoglobin</a></div> <p><a href="/wiki/Leghemoglobin" title="Leghemoglobin">Leghemoglobin</a> is a molecular similar in structure to myoglobin that is currently being used in artificial meat products, such as the <i>Impossible Burger</i>, to simulate both the color and taste of meat.<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> Similar in function to hemoglobin, leghemoglobin contains trace amounts of iron, but it is primarily found in plant roots.<sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Hemocyanin">Hemocyanin</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Respiratory_pigment&action=edit&section=6" title="Edit section: Hemocyanin"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Hemocyanin is a respiratory pigment that uses copper as its oxygen-binding molecule, as opposed to iron with hemoglobin. Hemocyanin is found in both <a href="/wiki/Arthropod" title="Arthropod">arthropods</a> and <a href="/wiki/Mollusca" title="Mollusca">Mollusca</a>, however it is thought that the molecule independently evolved in both phyla. There are several other molecules that exist in arthropods and Mollusca that are similar in structure to hemocyanin but serve entirely different purposes. For example, there are copper-containing tyrosinases that play significant roles in immune defense, wound healing, and the arthropod's cuticle. Molecules similar to hemocyanin in structure are grouped in under the hemocyanin superfamily.<sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Notes">Notes</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Respiratory_pigment&action=edit&section=7" title="Edit section: Notes"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-lower-alpha"> <div class="mw-references-wrap"><ol class="references"> <li id="cite_note-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-2">^</a></b></span> <span class="reference-text">also known as "hemoglobin", in a lax sense</span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text">in the strict sense, for the tetrameric form</span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Respiratory_pigment&action=edit&section=8" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1266661725">.mw-parser-output .portalbox{padding:0;margin:0.5em 0;display:table;box-sizing:border-box;max-width:175px;list-style:none}.mw-parser-output .portalborder{border:1px solid var(--border-color-base,#a2a9b1);padding:0.1em;background:var(--background-color-neutral-subtle,#f8f9fa)}.mw-parser-output .portalbox-entry{display:table-row;font-size:85%;line-height:110%;height:1.9em;font-style:italic;font-weight:bold}.mw-parser-output .portalbox-image{display:table-cell;padding:0.2em;vertical-align:middle;text-align:center}.mw-parser-output .portalbox-link{display:table-cell;padding:0.2em 0.2em 0.2em 0.3em;vertical-align:middle}@media(min-width:720px){.mw-parser-output .portalleft{margin:0.5em 1em 0.5em 0}.mw-parser-output .portalright{clear:right;float:right;margin:0.5em 0 0.5em 1em}}</style><ul role="navigation" aria-label="Portals" class="noprint portalbox portalborder portalright"> <li class="portalbox-entry"><span class="portalbox-image"><span class="noviewer" typeof="mw:File"><span><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/d6/WHO_Rod.svg/12px-WHO_Rod.svg.png" decoding="async" width="12" height="28" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/d6/WHO_Rod.svg/18px-WHO_Rod.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/d6/WHO_Rod.svg/24px-WHO_Rod.svg.png 2x" data-file-width="107" data-file-height="250" /></span></span></span><span class="portalbox-link"><a href="/wiki/Portal:Medicine" title="Portal:Medicine">Medicine portal</a></span></li></ul> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239543626"><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-:0-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-:0_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-:0_1-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-:0_1-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-:0_1-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-:0_1-4"><sup><i><b>e</b></i></sup></a> <a href="#cite_ref-:0_1-5"><sup><i><b>f</b></i></sup></a> <a href="#cite_ref-:0_1-6"><sup><i><b>g</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFUrich1994" class="citation cs2">Urich, Klaus (1994), Urich, Klaus (ed.), <a rel="nofollow" class="external text" href="https://doi.org/10.1007/978-3-662-06303-3_7">"Respiratory Pigments"</a>, <i>Comparative Animal Biochemistry</i>, Berlin, Heidelberg: Springer, pp. <span class="nowrap">249–</span>287, <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1007%2F978-3-662-06303-3_7">10.1007/978-3-662-06303-3_7</a>, <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-3-662-06303-3" title="Special:BookSources/978-3-662-06303-3"><bdi>978-3-662-06303-3</bdi></a><span class="reference-accessdate">, retrieved <span class="nowrap">2020-11-21</span></span></cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Comparative+Animal+Biochemistry&rft.atitle=Respiratory+Pigments&rft.pages=%3Cspan+class%3D%22nowrap%22%3E249-%3C%2Fspan%3E287&rft.date=1994&rft_id=info%3Adoi%2F10.1007%2F978-3-662-06303-3_7&rft.isbn=978-3-662-06303-3&rft.aulast=Urich&rft.aufirst=Klaus&rft_id=https%3A%2F%2Fdoi.org%2F10.1007%2F978-3-662-06303-3_7&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARespiratory+pigment" class="Z3988"></span></span> </li> <li id="cite_note-:1-3"><span class="mw-cite-backlink">^ <a href="#cite_ref-:1_3-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-:1_3-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-:1_3-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-:1_3-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-:1_3-4"><sup><i><b>e</b></i></sup></a> <a href="#cite_ref-:1_3-5"><sup><i><b>f</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHillWyseAnderson2017" class="citation book cs1">Hill, Richard W.; Wyse, Gordon A.; Anderson, Margaret (5 October 2017). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20201101112542/https://animalphys4e.sinauer.com/summary24.html"><i>Transport of Oxygen and Carbon Dioxide in Body Fluids (with an Introduction to Acid–Base Physiology)</i></a>. Sinauer Associates. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-1605357379" title="Special:BookSources/978-1605357379"><bdi>978-1605357379</bdi></a>. Archived from <a rel="nofollow" class="external text" href="https://animalphys4e.sinauer.com/summary24.html">the original</a> on 1 November 2020<span class="reference-accessdate">. Retrieved <span class="nowrap">10 November</span> 2020</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Transport+of+Oxygen+and+Carbon+Dioxide+in+Body+Fluids+%28with+an+Introduction+to+Acid%E2%80%93Base+Physiology%29&rft.pub=Sinauer+Associates&rft.date=2017-10-05&rft.isbn=978-1605357379&rft.aulast=Hill&rft.aufirst=Richard+W.&rft.au=Wyse%2C+Gordon+A.&rft.au=Anderson%2C+Margaret&rft_id=https%3A%2F%2Fanimalphys4e.sinauer.com%2Fsummary24.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARespiratory+pigment" class="Z3988"></span> <span class="cs1-visible-error citation-comment"><code class="cs1-code">{{<a href="/wiki/Template:Cite_book" title="Template:Cite book">cite book</a>}}</code>: </span><span class="cs1-visible-error citation-comment"><code class="cs1-code">|website=</code> ignored (<a href="/wiki/Help:CS1_errors#periodical_ignored" title="Help:CS1 errors">help</a>)</span></span> </li> <li id="cite_note-:2-5"><span class="mw-cite-backlink">^ <a href="#cite_ref-:2_5-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-:2_5-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-:2_5-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-:2_5-3"><sup><i><b>d</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFLamyTruchotGillesInternational_Union_of_Biological_Sciences1985" class="citation book cs1">Lamy, Jean; Truchot, J.-P; Gilles, R; International Union of Biological Sciences; Section of Comparative Physiology and Biochemistry; International Congress of Comparative Physiology and Biochemistry, eds. (1985). <a rel="nofollow" class="external text" href="https://www.worldcat.org/oclc/12558726"><i>Respiratory pigments in animals: relation, structure-function</i></a>. Berlin; New York: Springer-Verlag. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-387-15629-3" title="Special:BookSources/978-0-387-15629-3"><bdi>978-0-387-15629-3</bdi></a>. <a href="/wiki/OCLC_(identifier)" class="mw-redirect" title="OCLC (identifier)">OCLC</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/oclc/12558726">12558726</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Respiratory+pigments+in+animals%3A+relation%2C+structure-function&rft.place=Berlin%3B+New+York&rft.pub=Springer-Verlag&rft.date=1985&rft_id=info%3Aoclcnum%2F12558726&rft.isbn=978-0-387-15629-3&rft_id=https%3A%2F%2Fwww.worldcat.org%2Foclc%2F12558726&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARespiratory+pigment" class="Z3988"></span></span> </li> <li id="cite_note-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-6">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFFox1949" class="citation journal cs1">Fox, H. Munro (1949). <a rel="nofollow" class="external text" href="https://www.jstor.org/stable/82565">"On Chlorocruorin and Haemoglobin"</a>. <i>Proceedings of the Royal Society of London. Series B, Biological Sciences</i>. <b>136</b> (884): <span class="nowrap">378–</span>388. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1949RSPSB.136..378F">1949RSPSB.136..378F</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1098%2Frspb.1949.0031">10.1098/rspb.1949.0031</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0080-4649">0080-4649</a>. <a href="/wiki/JSTOR_(identifier)" class="mw-redirect" title="JSTOR (identifier)">JSTOR</a> <a rel="nofollow" class="external text" href="https://www.jstor.org/stable/82565">82565</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/18143368">18143368</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:6133526">6133526</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Proceedings+of+the+Royal+Society+of+London.+Series+B%2C+Biological+Sciences&rft.atitle=On+Chlorocruorin+and+Haemoglobin&rft.volume=136&rft.issue=884&rft.pages=%3Cspan+class%3D%22nowrap%22%3E378-%3C%2Fspan%3E388&rft.date=1949&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A6133526%23id-name%3DS2CID&rft_id=info%3Abibcode%2F1949RSPSB.136..378F&rft_id=info%3Adoi%2F10.1098%2Frspb.1949.0031&rft.issn=0080-4649&rft_id=https%3A%2F%2Fwww.jstor.org%2Fstable%2F82565%23id-name%3DJSTOR&rft_id=info%3Apmid%2F18143368&rft.aulast=Fox&rft.aufirst=H.+Munro&rft_id=https%3A%2F%2Fwww.jstor.org%2Fstable%2F82565&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARespiratory+pigment" class="Z3988"></span></span> </li> <li id="cite_note-:3-7"><span class="mw-cite-backlink">^ <a href="#cite_ref-:3_7-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-:3_7-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFFoxGardiner1932" class="citation journal cs1">Fox, Harold Munro; Gardiner, John Stanley (1932-09-01). <a rel="nofollow" class="external text" href="https://doi.org/10.1098%2Frspb.1932.0060">"The oxygen affinity of chlorocruorin"</a>. <i>Proceedings of the Royal Society of London. Series B, Containing Papers of a Biological Character</i>. <b>111</b> (772): <span class="nowrap">356–</span>363. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1098%2Frspb.1932.0060">10.1098/rspb.1932.0060</a></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Proceedings+of+the+Royal+Society+of+London.+Series+B%2C+Containing+Papers+of+a+Biological+Character&rft.atitle=The+oxygen+affinity+of+chlorocruorin&rft.volume=111&rft.issue=772&rft.pages=%3Cspan+class%3D%22nowrap%22%3E356-%3C%2Fspan%3E363&rft.date=1932-09-01&rft_id=info%3Adoi%2F10.1098%2Frspb.1932.0060&rft.aulast=Fox&rft.aufirst=Harold+Munro&rft.au=Gardiner%2C+John+Stanley&rft_id=https%3A%2F%2Fdoi.org%2F10.1098%252Frspb.1932.0060&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARespiratory+pigment" class="Z3988"></span></span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-8">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFImaiYoshikawa1985" class="citation journal cs1">Imai, Kiyohiro; Yoshikawa, Shinya (1985). "Oxygen-binding characteristics of Potamilla chlorocruorin". <i>European Journal of Biochemistry</i>. <b>147</b> (3): <span class="nowrap">453–</span>463. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1111%2Fj.0014-2956.1985.00453.x">10.1111/j.0014-2956.1985.00453.x</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/1432-1033">1432-1033</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/3979380">3979380</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=European+Journal+of+Biochemistry&rft.atitle=Oxygen-binding+characteristics+of+Potamilla+chlorocruorin&rft.volume=147&rft.issue=3&rft.pages=%3Cspan+class%3D%22nowrap%22%3E453-%3C%2Fspan%3E463&rft.date=1985&rft.issn=1432-1033&rft_id=info%3Apmid%2F3979380&rft_id=info%3Adoi%2F10.1111%2Fj.0014-2956.1985.00453.x&rft.aulast=Imai&rft.aufirst=Kiyohiro&rft.au=Yoshikawa%2C+Shinya&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARespiratory+pigment" class="Z3988"></span></span> </li> <li id="cite_note-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-9">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFGlomskiTamburlin1989" class="citation journal cs1">Glomski, Chester; Tamburlin, Judith (1989). <a rel="nofollow" class="external text" href="https://digitum.um.es/digitum/bitstream/10201/18688/1/The%20phylogenetic%20odyssey%20of%20the%20erythrocyte.%20I%20Hemoglobin%20the%20universal%20respiratory%20pigment.pdf">"The phylogenetic odyssey of the erythrocyte. I Hemoglobin: the universal respiratory pigment"</a> <span class="cs1-format">(PDF)</span>. <i>Histol Histopath</i>. <b>4</b> (4): <span class="nowrap">509–</span>514. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/2520483">2520483</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Histol+Histopath&rft.atitle=The+phylogenetic+odyssey+of+the+erythrocyte.+I+Hemoglobin%3A+the+universal+respiratory+pigment&rft.volume=4&rft.issue=4&rft.pages=%3Cspan+class%3D%22nowrap%22%3E509-%3C%2Fspan%3E514&rft.date=1989&rft_id=info%3Apmid%2F2520483&rft.aulast=Glomski&rft.aufirst=Chester&rft.au=Tamburlin%2C+Judith&rft_id=https%3A%2F%2Fdigitum.um.es%2Fdigitum%2Fbitstream%2F10201%2F18688%2F1%2FThe%2520phylogenetic%2520odyssey%2520of%2520the%2520erythrocyte.%2520I%2520Hemoglobin%2520the%2520universal%2520respiratory%2520pigment.pdf&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARespiratory+pigment" class="Z3988"></span></span> </li> <li id="cite_note-10"><span class="mw-cite-backlink"><b><a href="#cite_ref-10">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://sickle.bwh.harvard.edu/hemoglobinopathy.html">"Hemoglobinopathies"</a>. <i>sickle.bwh.harvard.edu</i><span class="reference-accessdate">. Retrieved <span class="nowrap">2020-11-21</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=sickle.bwh.harvard.edu&rft.atitle=Hemoglobinopathies&rft_id=https%3A%2F%2Fsickle.bwh.harvard.edu%2Fhemoglobinopathy.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARespiratory+pigment" class="Z3988"></span></span> </li> <li id="cite_note-Weber2001-11"><span class="mw-cite-backlink"><b><a href="#cite_ref-Weber2001_11-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWeberVinogradov2001" class="citation journal cs1">Weber RE, Vinogradov SN (April 2001). "Nonvertebrate Hemoglobins: Functions and Molecular Adaptations". <i>Physiological Reviews</i>. <b>81</b> (2): <span class="nowrap">569–</span>628. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1152%2Fphysrev.2001.81.2.569">10.1152/physrev.2001.81.2.569</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/11274340">11274340</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:10863037">10863037</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Physiological+Reviews&rft.atitle=Nonvertebrate+Hemoglobins%3A+Functions+and+Molecular+Adaptations&rft.volume=81&rft.issue=2&rft.pages=%3Cspan+class%3D%22nowrap%22%3E569-%3C%2Fspan%3E628&rft.date=2001-04&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A10863037%23id-name%3DS2CID&rft_id=info%3Apmid%2F11274340&rft_id=info%3Adoi%2F10.1152%2Fphysrev.2001.81.2.569&rft.aulast=Weber&rft.aufirst=RE&rft.au=Vinogradov%2C+SN&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARespiratory+pigment" class="Z3988"></span></span> </li> <li id="cite_note-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-12">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFLeeYongKimChoi2020" class="citation journal cs1">Lee, Hyun Jung; Yong, Hae In; Kim, Minsu; Choi, Yun-Sang; Jo, Cheorun (October 2020). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7463075">"Status of meat alternatives and their potential role in the future meat market — A review"</a>. <i>Asian-Australasian Journal of Animal Sciences</i>. <b>33</b> (10): <span class="nowrap">1533–</span>1543. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.5713%2Fajas.20.0419">10.5713/ajas.20.0419</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/1011-2367">1011-2367</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7463075">7463075</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/32819080">32819080</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Asian-Australasian+Journal+of+Animal+Sciences&rft.atitle=Status+of+meat+alternatives+and+their+potential+role+in+the+future+meat+market+%E2%80%94+A+review&rft.volume=33&rft.issue=10&rft.pages=%3Cspan+class%3D%22nowrap%22%3E1533-%3C%2Fspan%3E1543&rft.date=2020-10&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC7463075%23id-name%3DPMC&rft.issn=1011-2367&rft_id=info%3Apmid%2F32819080&rft_id=info%3Adoi%2F10.5713%2Fajas.20.0419&rft.aulast=Lee&rft.aufirst=Hyun+Jung&rft.au=Yong%2C+Hae+In&rft.au=Kim%2C+Minsu&rft.au=Choi%2C+Yun-Sang&rft.au=Jo%2C+Cheorun&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC7463075&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARespiratory+pigment" class="Z3988"></span></span> </li> <li id="cite_note-13"><span class="mw-cite-backlink"><b><a href="#cite_ref-13">^</a></b></span> <span class="reference-text">Seehafer, A., & Bartels, M. (2019). Meat 2.0 the regulatory environment of plant-based and cultured meat. <i>European Food and Feed Law Review (EFFL), 14(4),</i>323-331.</span> </li> <li id="cite_note-14"><span class="mw-cite-backlink"><b><a href="#cite_ref-14">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBurmester2001" class="citation journal cs1">Burmester, Thorsten (2001-02-01). <a rel="nofollow" class="external text" href="https://academic.oup.com/mbe/article/18/2/184/1079272">"Molecular Evolution of the Arthropod Hemocyanin Superfamily"</a>. <i>Molecular Biology and Evolution</i>. <b>18</b> (2): <span class="nowrap">184–</span>195. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Foxfordjournals.molbev.a003792">10.1093/oxfordjournals.molbev.a003792</a></span>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/0737-4038">0737-4038</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/11158377">11158377</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Molecular+Biology+and+Evolution&rft.atitle=Molecular+Evolution+of+the+Arthropod+Hemocyanin+Superfamily&rft.volume=18&rft.issue=2&rft.pages=%3Cspan+class%3D%22nowrap%22%3E184-%3C%2Fspan%3E195&rft.date=2001-02-01&rft.issn=0737-4038&rft_id=info%3Apmid%2F11158377&rft_id=info%3Adoi%2F10.1093%2Foxfordjournals.molbev.a003792&rft.aulast=Burmester&rft.aufirst=Thorsten&rft_id=https%3A%2F%2Facademic.oup.com%2Fmbe%2Farticle%2F18%2F2%2F184%2F1079272&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARespiratory+pigment" class="Z3988"></span></span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="Further_reading">Further reading</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Respiratory_pigment&action=edit&section=9" title="Edit section: Further reading"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCoatesDecker2017" class="citation journal cs1">Coates, CJ; Decker, H (January 2017). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5219038">"Immunological properties of oxygen-transport proteins: hemoglobin, hemocyanin and hemerythrin"</a>. <i>Cellular and Molecular Life Sciences</i>. <b>74</b> (2): <span class="nowrap">293–</span>317. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1007%2Fs00018-016-2326-7">10.1007/s00018-016-2326-7</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5219038">5219038</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/27518203">27518203</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Cellular+and+Molecular+Life+Sciences&rft.atitle=Immunological+properties+of+oxygen-transport+proteins%3A+hemoglobin%2C+hemocyanin+and+hemerythrin.&rft.volume=74&rft.issue=2&rft.pages=%3Cspan+class%3D%22nowrap%22%3E293-%3C%2Fspan%3E317&rft.date=2017-01&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5219038%23id-name%3DPMC&rft_id=info%3Apmid%2F27518203&rft_id=info%3Adoi%2F10.1007%2Fs00018-016-2326-7&rft.aulast=Coates&rft.aufirst=CJ&rft.au=Decker%2C+H&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5219038&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARespiratory+pigment" class="Z3988"></span></li></ul> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Respiratory_pigment&action=edit&section=10" title="Edit section: External links"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li>Gene Ontology – <a rel="nofollow" class="external text" href="http://amigo.geneontology.org/amigo/term/GO:0005344">GO:0005344: oxygen carrier activity</a></li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output 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.navbar{float:left;text-align:left;margin-right:0.5em}body.skin--responsive .mw-parser-output .navbox-image img{max-width:none!important}@media print{body.ns-0 .mw-parser-output .navbox{display:none!important}}</style></div><div role="navigation" class="navbox" aria-labelledby="Proteins_that_contain_heme_(hemoproteins)178" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><style data-mw-deduplicate="TemplateStyles:r1239400231">.mw-parser-output .navbar{display:inline;font-size:88%;font-weight:normal}.mw-parser-output .navbar-collapse{float:left;text-align:left}.mw-parser-output .navbar-boxtext{word-spacing:0}.mw-parser-output .navbar ul{display:inline-block;white-space:nowrap;line-height:inherit}.mw-parser-output .navbar-brackets::before{margin-right:-0.125em;content:"[ "}.mw-parser-output .navbar-brackets::after{margin-left:-0.125em;content:" ]"}.mw-parser-output .navbar li{word-spacing:-0.125em}.mw-parser-output .navbar a>span,.mw-parser-output .navbar a>abbr{text-decoration:inherit}.mw-parser-output .navbar-mini abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Hemeproteins" title="Template:Hemeproteins"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Hemeproteins" title="Template talk:Hemeproteins"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Hemeproteins" title="Special:EditPage/Template:Hemeproteins"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Proteins_that_contain_heme_(hemoproteins)178" style="font-size:114%;margin:0 4em"><a href="/wiki/Protein" title="Protein">Proteins</a> that contain <a href="/wiki/Heme" title="Heme">heme</a> (<a href="/wiki/Hemoprotein" title="Hemoprotein">hemoproteins</a>)</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Globin" title="Globin">Globins</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Hemoglobin" title="Hemoglobin">Hemoglobin</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Subunits</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i>Alpha locus on <a href="/wiki/Chromosome_16" title="Chromosome 16">16</a>:</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>α</i> <ul><li><a href="/wiki/Hemoglobin_subunit_alpha" title="Hemoglobin subunit alpha">HBA1</a></li> <li><a href="/wiki/Hemoglobin,_alpha_2" title="Hemoglobin, alpha 2">HBA2</a></li> <li><a href="/wiki/HBAP1" title="HBAP1">pseudo</a></li></ul></li> <li><i>ζ</i> <ul><li><a href="/wiki/Hemoglobin_subunit_zeta" title="Hemoglobin subunit zeta">HBZ</a></li></ul></li> <li><i>θ</i> <ul><li><a href="/wiki/HBQ1" title="HBQ1">HBQ1</a></li></ul></li> <li><i>μ</i> <ul><li><a href="/wiki/Mu_hemoglobin" title="Mu hemoglobin">HBM</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Human_%CE%B2-globin_locus" title="Human β-globin locus">Beta locus</a> on <a href="/wiki/Chromosome_11" title="Chromosome 11">11</a>:</i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i>β</i> <ul><li><a href="/wiki/Hemoglobin_subunit_beta" title="Hemoglobin subunit beta">HBB</a></li></ul></li> <li><i>δ</i> <ul><li><a href="/wiki/HBD" title="HBD">HBD</a></li></ul></li> <li><i>γ</i> <ul><li><a href="/wiki/HBG1" title="HBG1">HBG1</a></li> <li><a href="/wiki/HBG2" title="HBG2">HBG2</a></li></ul></li> <li><i>ε</i> <ul><li><a href="/wiki/HBE1" title="HBE1">HBE1</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Tetrameric_protein" title="Tetrameric protein">Tetramers</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i>stages of <br />development:</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">Embryonic</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Embryonic_hemoglobin" title="Embryonic hemoglobin">HbE Gower 1</a> (ζ<sub>2</sub>ε<sub>2</sub>)</li> <li><a href="/wiki/Embryonic_hemoglobin" title="Embryonic hemoglobin">HbE Gower 2</a> (α<sub>2</sub>ε<sub>2</sub>)</li> <li><a href="/wiki/Embryonic_hemoglobin" title="Embryonic hemoglobin">HbE Portland I</a> (ζ<sub>2</sub>γ<sub>2</sub>)</li> <li><a href="/wiki/Embryonic_hemoglobin" title="Embryonic hemoglobin">HbE Portland II</a> (ζ<sub>2</sub>β<sub>2</sub>)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Fetal</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Fetal_hemoglobin" title="Fetal hemoglobin">HbF/Fetal</a> (α<sub>2</sub>γ<sub>2</sub>)</li> <li><a href="/wiki/Hemoglobin_A" title="Hemoglobin A">HbA</a> (α<sub>2</sub>β<sub>2</sub>)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Adult</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Hemoglobin_A" title="Hemoglobin A">HbA</a> (α<sub>2</sub>β<sub>2</sub>)</li> <li><a href="/wiki/Hemoglobin_A2" title="Hemoglobin A2">HbA<sub>2</sub></a> (α<sub>2</sub>δ<sub>2</sub>)</li> <li><a href="/wiki/Fetal_hemoglobin" title="Fetal hemoglobin">HbF/Fetal</a> (α<sub>2</sub>γ<sub>2</sub>)</li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i>pathology:</i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Alpha-thalassemia" title="Alpha-thalassemia">HbH</a> (β<sub>4</sub>)</li> <li><a href="/wiki/Hemoglobin_Barts" title="Hemoglobin Barts">Barts</a> (γ<sub>4</sub>)</li> <li><a href="/wiki/HbD" class="mw-redirect" title="HbD">HbD</a> (α<sub>2</sub>β<sup>D</sup><sub>2</sub>)</li> <li><a href="/wiki/Sickle_cell_disease" title="Sickle cell disease">HbS</a> (α<sub>2</sub>β<sup>S</sup><sub>2</sub>)</li> <li><a href="/wiki/Hemoglobin_C" title="Hemoglobin C">HbC</a> (α<sub>2</sub>β<sup>C</sup><sub>2</sub>)</li> <li><a href="/wiki/Hemoglobin_E" title="Hemoglobin E">HbE</a> (α<sub>2</sub>β<sup>E</sup><sub>2</sub>)</li> <li><a href="/wiki/Hemoglobin_O" title="Hemoglobin O">HbO</a> (α<sub>2</sub>β<sup>O</sup><sub>2</sub>)</li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Compounds</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Carboxyhemoglobin" title="Carboxyhemoglobin">Carboxyhemoglobin</a></li> <li><a href="/wiki/Carbaminohemoglobin" title="Carbaminohemoglobin">Carbaminohemoglobin</a></li> <li><a href="/wiki/Hemoglobin" title="Hemoglobin">Oxyhemoglobin</a>/<a href="/wiki/Deoxyhemoglobin" class="mw-redirect" title="Deoxyhemoglobin">Deoxyhemoglobin</a></li> <li><a href="/wiki/Sulfhemoglobinemia" title="Sulfhemoglobinemia">Sulfhemoglobin</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other human</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Glycated_hemoglobin" title="Glycated hemoglobin">Glycated hemoglobin</a></li> <li><a href="/wiki/Methemoglobin" title="Methemoglobin">Methemoglobin</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Nonhuman</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Erythrocruorin" title="Erythrocruorin">Chlorocruorin</a></li> <li><a href="/wiki/Erythrocruorin" title="Erythrocruorin">Erythrocruorin</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i>human:</i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Myoglobin" title="Myoglobin">Myoglobin</a> <ul><li><a href="/wiki/Metmyoglobin" title="Metmyoglobin">Metmyoglobin</a></li></ul></li> <li><a href="/wiki/Neuroglobin" title="Neuroglobin">Neuroglobin</a></li> <li><a href="/wiki/Cytoglobin" title="Cytoglobin">Cytoglobin</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i>plant:</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Leghemoglobin" title="Leghemoglobin">Leghemoglobin</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cytochrome" title="Cytochrome">Cytochrome</a> <ul><li><a href="/wiki/Cytochrome_b" title="Cytochrome b">Cytochrome b</a></li> <li><a href="/wiki/Cytochrome_P450" title="Cytochrome P450">Cytochrome P450</a></li></ul></li> <li><a href="/wiki/Methemalbumin" title="Methemalbumin">Methemalbumin</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="2" style="background: transparent; padding: 0px;"><div><i>see also <a href="/wiki/Template:Disorders_of_globin_and_globulin_proteins" title="Template:Disorders of globin and globulin proteins">disorders of globin and globulin proteins</a></i></div></td></tr></tbody></table></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?useformat=desktop&type=1x1&usesul3=0" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Retrieved from "<a dir="ltr" href="https://en.wikipedia.org/w/index.php?title=Respiratory_pigment&oldid=1240205085">https://en.wikipedia.org/w/index.php?title=Respiratory_pigment&oldid=1240205085</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Help:Category" title="Help:Category">Category</a>: <ul><li><a href="/wiki/Category:Respiratory_pigments" title="Category:Respiratory pigments">Respiratory pigments</a></li></ul></div><div id="mw-hidden-catlinks" class="mw-hidden-catlinks 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