CINXE.COM

<!DOCTYPE html><html lang="en-US"><head><title>Just a moment...</title><meta http-equiv="Content-Type" content="text/html; charset=UTF-8"><meta http-equiv="X-UA-Compatible" content="IE=Edge"><meta name="robots" content="noindex,nofollow"><meta name="viewport" content="width=device-width,initial-scale=1"><style>*{box-sizing:border-box;margin:0;padding:0}html{line-height:1.15;-webkit-text-size-adjust:100%;color:#313131;font-family:system-ui,-apple-system,BlinkMacSystemFont,Segoe UI,Roboto,Helvetica Neue,Arial,Noto Sans,sans-serif,Apple Color Emoji,Segoe UI Emoji,Segoe UI Symbol,Noto Color Emoji}body{display:flex;flex-direction:column;height:100vh;min-height:100vh}.main-content{margin:8rem auto;max-width:60rem;padding-left:1.5rem}@media (width <= 720px){.main-content{margin-top:4rem}}.h2{font-size:1.5rem;font-weight:500;line-height:2.25rem}@media (width <= 720px){.h2{font-size:1.25rem;line-height:1.5rem}}#challenge-error-text{background-image:url(data:image/svg+xml;base64,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);background-repeat:no-repeat;background-size:contain;padding-left:34px}@media (prefers-color-scheme:dark){body{background-color:#222;color:#d9d9d9}}</style><meta http-equiv="refresh" content="390"></head><body class="no-js"><div class="main-wrapper" role="main"><div class="main-content"><noscript><div class="h2"><span id="challenge-error-text">Enable JavaScript and cookies to continue</span></div></noscript></div></div><script>(function(){window._cf_chl_opt={cvId: '3',cZone: "onlinelibrary.wiley.com",cType: 'managed',cRay: '927a93ca7f17fdc8',cH: 'atpiM83lK0VKv2HTdcfo_OyEImgUBaaC.ezf55Insh4-1743201082-1.2.1.1-ALq5q3alvwChZqR_VsQTNaplJQ1nT3ZvbgLWlBEOTyNwljlGZfIgtsK0p7Za8uz4',cUPMDTk: "\/doi\/abs\/10.1096\/fj.201600698R?__cf_chl_tk=GdYCD7xCf2yK3M6tYulwbGt6mhOVNOjg6946xoMnljI-1743201082-1.0.1.1-iZcjBd8MM2NaOsCIP3XFDQaH23RLTod3q1pPV4itrfE",cFPWv: 'b',cITimeS: '1743201082',cTTimeMs: '1000',cMTimeMs: '390000',cTplC: 0,cTplV: 5,cTplB: 'cf',cK: "",fa: "\/doi\/abs\/10.1096\/fj.201600698R?__cf_chl_f_tk=GdYCD7xCf2yK3M6tYulwbGt6mhOVNOjg6946xoMnljI-1743201082-1.0.1.1-iZcjBd8MM2NaOsCIP3XFDQaH23RLTod3q1pPV4itrfE",md: "KFIV0LIUSTLtM.VE1hUuZtc7._.zO2T5gM.YLdzYppg-1743201082-1.2.1.1-tgVuFVUeuQG.9TFrEZQbV8xJJLieyf74vPg58kZ.ECIzf8MGo8v1a0fJ._zrkVaSrbEGvD07tkqV.37FkC1iOsHj3QFi5925JFRiKDyw7N3ZKFFT5cs32rHlEShCuRbdlvMcetwFChFBs1hVFw2EwIUIzXmYlrreBksvUTC4BkCXkxP5leMqvBF584sT3M59nbX1XLOzHSiG6.5mp3ECGvPtWqcZVYeibaFxMnTvzDhi1JhUHZAAsYLHRIGcJrOelKqXwiNPO95Xm29R.eaJFFS8uHe7LhB_3YeKOpJfacGnGExdbtZN_mcfdeXYnHMrHZ4RBpNm.sjOvPydPFrLQ_cXVCIP2Nz4ajPU_l90CNy7lyUG65yhO5epNoZ94PVD4RatMePEaANVeQTRDUhdyAlnt5q77YKjepn.48NHyTXfhie6Lhaxf_lmwiOmQZ2uH4972btrqfYGixCWEVABWhZ4bhgJf.OkGGzShXao4d6BKGBtTlSuLdmb6Z5CO0jXkcyQujogOOTnxv2b_2DG2TNnqxf9n6NTTAsI_uEHuaqBx8ykGSlWZRsrCKGq8HGGZrRora2pgGJ_fjUwompqWn7QofJLl5HhC73BpZWbuA5.xcU2Kd29gu56p03VR5fNrN9xO6.ShBUIhlVA2_P6IwlAazsofc6wERxPtwu_ZrQnexUSedWCjLgjBDm12bEaEPjkhnou11bzJ44FNHHNGVknQcwgbPxHFplmypk4lE6OVtQs1Ue_._FIXru9Hq4DOjeDY60JjWAU3lRCSj.acigBQI2f4Dt_MYytUyQKH7PxNSto3bXBRzUS6E.sYRjSzx1XGKiiK9oDxJ.x.1ISvDccrEN6IfQP9fXKrw5cd8Bx9wbXNjeul7qZKgfaxZil0Mt_Wvk.kiLbhrDgSQGz1tjmL6dfLqgsaVN.7GCP0A9TO_DhMVO_zKOVBqdizoMqdCCXdmufwYc6YTF7dyXlH0Emgck27xXj_hvtrcyt7jRLRWVokIL4FVM30LlFkUGXlSM.mHSH8hoZTXXI0d6zMCfSP73qmo6pOVQXzFKeBIRnWiMlZw5v16o9TiM8xZLTxso_S0YfK5f_S1Ev9CRMH9xFRIvCjPDhuvFmrx9tZHoKxiCp8XmkFpqWfD7KQU0q9Qp1SN8X4bp32NOhCDLjRxOCxwIbafUVeACLmFEHGEj8gtrHuthfBTJ8PsYYv9Qxs.Y2f1CsVhujVgwfqc6SciEHr5MFmo_8cwv3PFAiXG48N2BzfpOE_NOymQfx3gVTRMKL1RMKl7X0wsBHq9pXyeb.VbEX37hnfGdSuzXQMFI",mdrd: "QPqjwbTlaWDqzbqTPCGEAbaYL475.5JKpss4v.aC.Tc-1743201082-1.2.1.1-KBErWHI9EKVLXRkQZVhNiQXNsDWGqGCvf3s6jtoO.TO2ZG3MLZw_P7sZ1UbLw3qASERxEZHQLcxvSEE.g6lGNxg2qcg1JSIJL2M8rpbQFaUjcre8T11YGrje9RiS2FTuXRtmM5nF6rud1qqyqwDMh2aOv2OmD6ZXlAmaoKCTftqiXRLBy2A_l2od4goqXwdAzLRWrpfzOwSFWrv89fnGL2Uhrgn_HF_jTbM_fezyFjXjIm_6l3iGGYx6KDjrKJ9gUawCviK7XKHNudtG8s9zsXRywBC66en3pt3Hk6hStmzij94ceenGQcVdxHNTKeMZpKjTT_mvz5nyk9OcKkis7Bzc.tAW3NswNCWCbiAmmb3TBm7Z2h1h.tnM3G18ZegLjy4rRVbRjSiKNK8rEE2V9M2lGcpMKiGzkK5vIZgkdv8NI5XrciOJ06KQ2FoCek6r7QerMP66t97Mq3aJ7moNmid58SBMsDV16NtD9iYWe47WmPphhrne6XmsK.qsXHzqh2PQiOtLxRzU3TWVMH321r5cREq8.Ilv5YOy5EDL5Ky4KIfq_OVg3fxe82dq2zK5cu6SYgdSPUMJeprSg0HLGP17Kc1nUsfuh7De_zUuXE3GW9btlnSH6dKe80D34NbyYATyY8hwRjrUHYp7_PQE0UPbCKbqMqGgeh17lTztNRYKOL4heJACOKRj9CUji419.xq1lM3tXVbYi1qMtVjNMrKW4_pYAc0FoDjw5OFU_v4URvkU1EmcYrTufNez7bBTq_Hd9cUjGWPdyc7NxzYUcEEZeC8zR9q_2tweLGMC0AvjUMYDP.EmGVS5hAG96iOCW1CSWn.Ib_ZT3uOj8s3UgRNqJcLhSsBmU21dZVAbF8x4k9CfQVipx2WEfROmJBkEBa.TdMKSftHmuo_SOJ2Q7kaByQD8wLoPcOylKrgvwe6Ey_k1JTsz1g0w1AiwLYMBqiBL05jpcheybN__FJOX6Zw21jErLpewXWvFP5q7mnukiE9JldoGV0w4DlaR64CQoyzBLwveWhlbH9GiTe1XPMbjprJ16Vo7p9_JbU5r0HldHrdl6WK7QzyC1ELoWt8DmU7tF7ipYtN5a5EtJPsmH3H88oCx2k8ssKHcHm1R0ZuBWT7LF6r9qOaufpGH_2ACQ6RYOqz77K1cOp_nFVgVDkmHxUJMCsunwFJj1gn3nTJlLkAC4opQRik3xZAFeX5XQI8x7YQYVhRVwe8EJFyGX6aTQ8Xa3mVaAXEOpTI5Z8qZ658ozqV_JkjVXX9kD_NBv_kmk1gRVjThL.dm0_gNtUrE3QdOlqzTWdMiCrDf3Kc79PG5jVruL29rm_NnSroAEh0w.LDUsLhMmNG3BJAGfJOhiTQUDwtojUT8uYcjFSr5vXaBvQUJzYw.GCi5b_FHWjskp4ZeD0Ka4_j6uX0ClEqxF3dJvOqKSxG3WKC2qPT7egp4NNj1qw1NYtoEw.j5bMfPInjnsyCdwtqVAvc0Wryx6ZWjCwsPIrmROSifNnpsjVNac9vcTqnYTzPRYxnk0e5FnKrR.bLkN6aWknleuPrDBc5.dD96EWsGdtSzcjSrThhQOSDP.s0aM1AKAU8hQ8T92dj5B2cmDvXR88QK10gN_1b_jDFircy57SMPGPWnxYzE4oFkEjBb_7Jq_dULPrsezO9DibqDLlnAjGAih_0tgWbjt4T7rDb5yS5Bux02_qr5NKgskUtXNkVpgqbuDsaUGg3HSZU8I9gNkpmjnbPuJ3QaCj_YLdlpPirWHaX6XHNMOCY7mGNnZe2CftjakS1cR7KJfs0gbuBs.fu8MjVvWPqrBtH.hrnOC7DIjYOogwfjn06nt3RRD2pRVeg2PMV7XkKkFk2AaVxXO0MjQszeCm.3ReD.SXy_C2i44HpZF_UEseq8AdRga4KT2xxDOsRixm7Lit9mn6CdnGZh7clj1f9nUnaF23zTUKEg91oxnXwN1W8C2KI8DaIRB.cMPThBdqboTVBB4FfABHgCdzKCrNq9HwTXXF5SxRD1X3ExjUN_G7Q6cdjjO9zEE5vibeqIbKKzgHebt_xqxiHsWOwhrT_LfIU6QKc6H_6haMJxQ_hM_jyEZftl8NIyD19M"};var cpo = document.createElement('script');cpo.src = '/cdn-cgi/challenge-platform/h/b/orchestrate/chl_page/v1?ray=927a93ca7f17fdc8';window._cf_chl_opt.cOgUHash = location.hash === '' && location.href.indexOf('#') !== -1 ? '#' : location.hash;window._cf_chl_opt.cOgUQuery = location.search === '' && location.href.slice(0, location.href.length - window._cf_chl_opt.cOgUHash.length).indexOf('?') !== -1 ? '?' : location.search;if (window.history && window.history.replaceState) {var ogU = location.pathname + window._cf_chl_opt.cOgUQuery + window._cf_chl_opt.cOgUHash;history.replaceState(null, null, "\/doi\/abs\/10.1096\/fj.201600698R?__cf_chl_rt_tk=GdYCD7xCf2yK3M6tYulwbGt6mhOVNOjg6946xoMnljI-1743201082-1.0.1.1-iZcjBd8MM2NaOsCIP3XFDQaH23RLTod3q1pPV4itrfE" + window._cf_chl_opt.cOgUHash);cpo.onload = function() {history.replaceState(null, null, ogU);}}document.getElementsByTagName('head')[0].appendChild(cpo);}());</script></body></html>