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class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Benzer_vs._Hirsch"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.2.1</span> <span>Benzer vs. Hirsch</span> </div> </a> <ul id="toc-Benzer_vs._Hirsch-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Research_accomplishments" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Research_accomplishments"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.2.2</span> <span>Research accomplishments</span> </div> </a> <ul id="toc-Research_accomplishments-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Cancer_research" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Cancer_research"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.3</span> <span>Cancer research</span> </div> </a> <ul id="toc-Cancer_research-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Honors_and_awards" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Honors_and_awards"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Honors and awards</span> </div> </a> <ul id="toc-Honors_and_awards-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Books" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Books"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Books</span> </div> </a> <ul id="toc-Books-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-See_also" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#See_also"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>See also</span> </div> </a> <ul id="toc-See_also-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" 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href="https://azb.wikipedia.org/wiki/%D8%B3%DB%8C%D9%85%D9%88%D8%B1_%D8%A8%D9%86%D8%B2%D8%B1" title="سیمور بنزر – South Azerbaijani" lang="azb" hreflang="azb" data-title="سیمور بنزر" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-zh-min-nan mw-list-item"><a href="https://zh-min-nan.wikipedia.org/wiki/Seymour_Benzer" title="Seymour Benzer – Minnan" lang="nan" hreflang="nan" data-title="Seymour Benzer" data-language-autonym="閩南語 / Bân-lâm-gú" data-language-local-name="Minnan" class="interlanguage-link-target"><span>閩南語 / Bân-lâm-gú</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%A1%D0%B5%D0%B9%D0%BC%D1%83%D1%80_%D0%91%D0%B5%D0%BD%D0%B7%D0%B5%D1%80" title="Сеймур Бензер – Belarusian" lang="be" hreflang="be" data-title="Сеймур Бензер" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Seymour_Benzer" title="Seymour Benzer – Catalan" lang="ca" hreflang="ca" data-title="Seymour Benzer" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Seymour_Benzer" title="Seymour Benzer – German" lang="de" hreflang="de" data-title="Seymour Benzer" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%B3%DB%8C%D9%85%D9%88%D8%B1_%D8%A8%D9%86%D8%B2%D8%B1" title="سیمور بنزر – Persian" lang="fa" hreflang="fa" data-title="سیمور بنزر" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Seymour_Benzer" title="Seymour Benzer – French" lang="fr" hreflang="fr" data-title="Seymour Benzer" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Seymour_Benzer" title="Seymour Benzer – Italian" lang="it" hreflang="it" data-title="Seymour Benzer" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D8%B3%D9%8A%D9%85%D9%88%D8%B1_%D8%A8%D9%86%D8%B2%D8%B1" title="سيمور بنزر – Egyptian Arabic" lang="arz" hreflang="arz" data-title="سيمور بنزر" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%B7%E3%83%BC%E3%83%A2%E3%82%A2%E3%83%BB%E3%83%99%E3%83%B3%E3%82%B6%E3%83%BC" title="シーモア・ベンザー – Japanese" lang="ja" hreflang="ja" data-title="シーモア・ベンザー" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Seymour_Benzer" title="Seymour Benzer – Polish" lang="pl" hreflang="pl" data-title="Seymour Benzer" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Seymour_Benzer" title="Seymour Benzer – Portuguese" lang="pt" hreflang="pt" data-title="Seymour Benzer" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%91%D0%B5%D0%BD%D0%B7%D0%B5%D1%80,_%D0%A1%D0%B5%D0%B9%D0%BC%D1%83%D1%80" title="Бензер, Сеймур – Russian" lang="ru" hreflang="ru" data-title="Бензер, Сеймур" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Seymour_Benzer" title="Seymour Benzer – Simple English" lang="en-simple" hreflang="en-simple" data-title="Seymour Benzer" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link 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searchaux" style="display:none">American geneticist</div> <style data-mw-deduplicate="TemplateStyles:r1257001546">.mw-parser-output .infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox-full-data:not(.notheme)>div:not(.notheme)[style]{background:#1f1f23!important;color:#f8f9fa}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox-full-data:not(.notheme) div:not(.notheme){background:#1f1f23!important;color:#f8f9fa}}@media(min-width:640px){body.skin--responsive .mw-parser-output .infobox-table{display:table!important}body.skin--responsive .mw-parser-output .infobox-table>caption{display:table-caption!important}body.skin--responsive .mw-parser-output .infobox-table>tbody{display:table-row-group}body.skin--responsive .mw-parser-output .infobox-table tr{display:table-row!important}body.skin--responsive .mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox biography vcard"><tbody><tr><th colspan="2" class="infobox-above" style="font-size:125%;"><div class="fn">Seymour Benzer</div></th></tr><tr><td colspan="2" class="infobox-image"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Seymour_Benzer.gif" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/39/Seymour_Benzer.gif/220px-Seymour_Benzer.gif" decoding="async" width="220" height="167" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/39/Seymour_Benzer.gif/330px-Seymour_Benzer.gif 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/39/Seymour_Benzer.gif/440px-Seymour_Benzer.gif 2x" data-file-width="1777" data-file-height="1351" /></a></span><div class="infobox-caption">Benzer with a <i><a href="/wiki/Drosophila" title="Drosophila">Drosophila</a></i> model, 1974</div></td></tr><tr><th scope="row" class="infobox-label">Born</th><td class="infobox-data"><span style="display:none">(<span class="bday">1921-10-15</span>)</span>October 15, 1921<br /><div style="display:inline" class="birthplace"><a href="/wiki/Bensonhurst,_Brooklyn" title="Bensonhurst, Brooklyn">Bensonhurst</a>, <a href="/wiki/New_York_City" title="New York City">New York City</a>, U.S.</div></td></tr><tr><th scope="row" class="infobox-label">Died</th><td class="infobox-data">November 30, 2007<span style="display:none">(2007-11-30)</span> (aged 86)<br /><div style="display:inline" class="deathplace"><a href="/wiki/Pasadena,_California" title="Pasadena, California">Pasadena, California</a>, U.S.</div></td></tr><tr><th scope="row" class="infobox-label">Alma mater</th><td class="infobox-data"><a href="/wiki/Brooklyn_College" title="Brooklyn College">Brooklyn College</a> (<a href="/wiki/B.S." class="mw-redirect" title="B.S.">BS</a>)<br /><a href="/wiki/Purdue_University" title="Purdue University">Purdue University</a> (<a href="/wiki/PhD" class="mw-redirect" title="PhD">PhD</a>)</td></tr><tr><th scope="row" class="infobox-label">Awards</th><td class="infobox-data"><a href="/wiki/Gairdner_Foundation_International_Award" class="mw-redirect" title="Gairdner Foundation International Award">Gairdner Foundation International Award</a> <small>(1964, 2004)</small><br /><a href="/wiki/Louisa_Gross_Horwitz_Prize" title="Louisa Gross Horwitz Prize">Louisa Gross Horwitz Prize</a> <small>(1976)</small><br /><a href="/wiki/Harvey_Prize" title="Harvey Prize">Harvey Prize</a> <span style="font-size:85%;">(1977)</span><br /><a href="/wiki/Thomas_Hunt_Morgan_Medal" title="Thomas Hunt Morgan Medal">Thomas Hunt Morgan Medal</a> <span style="font-size:85%;">(1986)</span><br /><a href="/wiki/Wolf_Prize_in_Medicine" title="Wolf Prize in Medicine">Wolf Prize in Medicine</a> <span style="font-size:85%;">(1991)</span><br /><a href="/wiki/Crafoord_Prize" title="Crafoord Prize">Crafoord Prize</a> <span style="font-size:85%;">(1993)</span><br /><a href="/wiki/Mendel_Medal_(genetics)" class="mw-redirect" title="Mendel Medal (genetics)">Mendel Medal</a> <span style="font-size:85%;">(1994)</span><br /><a href="/wiki/International_Prize_for_Biology" title="International Prize for Biology">International Prize for Biology</a> <span style="font-size:85%;">(2000)</span><br /><a href="/wiki/Royal_Society" title="Royal Society">Fellow of the Royal Society</a><sup id="cite_ref-frs_1-0" class="reference"><a href="#cite_note-frs-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><br /><a href="/wiki/Gruber_Prize_in_Neuroscience" title="Gruber Prize in Neuroscience">Gruber Prize in Neuroscience</a> <span style="font-size:85%;">(2004)</span></td></tr><tr><td colspan="2" class="infobox-full-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1257001546"><b>Scientific career</b></td></tr><tr><th scope="row" class="infobox-label">Fields</th><td class="infobox-data category"><a href="/wiki/Physics" title="Physics">Physics</a>, <a href="/wiki/Molecular_biology" title="Molecular biology">molecular biology</a>, <a href="/wiki/Behavioral_genetics" class="mw-redirect" title="Behavioral genetics">behavioral genetics</a>, <a href="/wiki/Chronobiology" title="Chronobiology">chronobiology</a>, <a href="/wiki/Neurogenetics" title="Neurogenetics">neurogenetics</a></td></tr><tr><th scope="row" class="infobox-label">Institutions</th><td class="infobox-data"><a href="/wiki/Purdue_University" title="Purdue University">Purdue University</a><br /><a href="/wiki/California_Institute_of_Technology" title="California Institute of Technology">California Institute of Technology</a></td></tr><tr><th scope="row" class="infobox-label"><a href="/wiki/Thesis" title="Thesis">Thesis</a></th><td class="infobox-data"><i><a rel="nofollow" class="external text" href="http://search.proquest.com/docview/301828715/">Photoelectric Effects in Germanium</a></i> <span style="font-size:97%;">(1947)</span></td></tr><tr style="display:none"><td colspan="2"> </td></tr></tbody></table> <p><b>Seymour Benzer</b> (October 15, 1921 – November 30, 2007) was an American <a href="/wiki/Physicist" title="Physicist">physicist</a>, <a href="/wiki/Molecular_biologist" class="mw-redirect" title="Molecular biologist">molecular biologist</a> and <a href="/wiki/Behavioral_geneticist" class="mw-redirect" title="Behavioral geneticist">behavioral geneticist</a>. His career began during the molecular biology revolution of the 1950s, and he eventually rose to prominence in the fields of molecular and behavioral genetics. He led a productive genetics research lab both at Purdue University and as the James G. Boswell Professor of Neuroscience, emeritus, at the <a href="/wiki/California_Institute_of_Technology" title="California Institute of Technology">California Institute of Technology</a>.<sup id="cite_ref-frs_1-1" class="reference"><a href="#cite_note-frs-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Greenspan_2008_R106–10_2-0" class="reference"><a href="#cite_note-Greenspan_2008_R106–10-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-nytzimmer_3-0" class="reference"><a href="#cite_note-nytzimmer-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Biography">Biography</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Seymour_Benzer&action=edit&section=1" title="Edit section: Biography"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Early_life_and_education">Early life and education</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Seymour_Benzer&action=edit&section=2" title="Edit section: Early life and education"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Benzer was born in the <a href="/wiki/South_Bronx" title="South Bronx">South Bronx</a> to Meir Benzer and Eva Naidorf, both <a href="/wiki/Ashkenazi_Jews" title="Ashkenazi Jews">Jews from Poland</a>.<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> He had two older sisters, and his parents favored him as the only boy.<sup id="cite_ref-Holmes_6-0" class="reference"><a href="#cite_note-Holmes-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> One of Benzer's earliest scientific experiences was dissecting frogs he had caught as a boy. In an interview at Caltech, Benzer also remembered receiving a microscope for his 13th birthday, “and that opened up the whole world.”<sup id="cite_ref-oralhistory_7-0" class="reference"><a href="#cite_note-oralhistory-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> The book <i><a href="/wiki/Arrowsmith_(novel)" title="Arrowsmith (novel)">Arrowsmith</a></i> by <a href="/wiki/Sinclair_Lewis" title="Sinclair Lewis">Sinclair Lewis</a> heavily influenced the young Benzer, and he even imitated the handwriting of Max Gottlieb, a scientist character in the novel. Benzer graduated from <a href="/wiki/New_Utrecht_High_School" title="New Utrecht High School">New Utrecht High School</a> at 15 years old.<sup id="cite_ref-Weiner_8-0" class="reference"><a href="#cite_note-Weiner-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p><p>In 1938 he enrolled at <a href="/wiki/Brooklyn_College" title="Brooklyn College">Brooklyn College</a> where he majored in physics.<sup id="cite_ref-nytzimmer_3-1" class="reference"><a href="#cite_note-nytzimmer-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Benzer then moved on to <a href="/wiki/Purdue_University" title="Purdue University">Purdue University</a> to earn his Ph.D. in solid state physics. While there he was recruited for a secret military project to develop improved radar. He performed research that led to the development of stable <a href="/wiki/Germanium" title="Germanium">germanium</a> rectifiers and discovered a germanium crystal able to be used at high voltages, among the scientific work that led to the first <a href="/wiki/Transistor" title="Transistor">transistor</a>.<sup id="cite_ref-Greenspan_2008_R106–10_2-1" class="reference"><a href="#cite_note-Greenspan_2008_R106–10-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-nytzimmer_3-2" class="reference"><a href="#cite_note-nytzimmer-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-caltechobit_9-0" class="reference"><a href="#cite_note-caltechobit-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Personal_life">Personal life</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Seymour_Benzer&action=edit&section=3" title="Edit section: Personal life"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>At <a href="/wiki/Brooklyn_College" title="Brooklyn College">Brooklyn College</a>, as a sixteen-year-old freshman, Benzer met Dorothy Vlosky (nicknamed Dotty), a twenty-one-year-old nurse. He later married her in <a href="/wiki/New_York_City" title="New York City">New York City</a> in 1942.<sup id="cite_ref-Weiner_8-1" class="reference"><a href="#cite_note-Weiner-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> They had two daughters, Barbie (Barbara) and Martha Jane. </p><p>Benzer died of a stroke at the <a href="/wiki/Huntington_Hospital" title="Huntington Hospital">Huntington Hospital</a> in <a href="/wiki/Pasadena,_California" title="Pasadena, California">Pasadena, California</a>.<sup id="cite_ref-caltechobit_9-1" class="reference"><a href="#cite_note-caltechobit-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Scientific_career">Scientific career</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Seymour_Benzer&action=edit&section=4" title="Edit section: Scientific career"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Molecular_biology">Molecular biology</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Seymour_Benzer&action=edit&section=5" title="Edit section: Molecular biology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Upon receiving his Ph.D. in 1947, he was immediately hired as an assistant professor in physics at Purdue. However, Benzer was inspired by <a href="/wiki/Erwin_Schr%C3%B6dinger" title="Erwin Schrödinger">Erwin Schrödinger's</a> book <i><a href="/wiki/What_Is_Life%3F" title="What Is Life?">What Is Life?</a></i>, in which the physicist pondered the physical nature of the gene and a “code” of life. This catalyzed Benzer's shift in interest to biology, and he moved into the area of <a href="/wiki/Bacteriophage" title="Bacteriophage">bacteriophage</a> genetics.,<sup id="cite_ref-Retros_10-0" class="reference"><a href="#cite_note-Retros-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> spending two years as a postdoctoral fellow in <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Max Delbrück</a>'s laboratory at California Institute of Technology, and then returning to Purdue. At Purdue University, Benzer developed the <a href="/wiki/T4_rII_system" title="T4 rII system">T4 <i>rII</i> system</a>, a new genetic technique involving <a href="/wiki/Genetic_recombination" title="Genetic recombination">recombination</a> in T4 bacteriophage <i>rII</i> mutants.<sup id="cite_ref-topography_11-0" class="reference"><a href="#cite_note-topography-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> After observing that a particular <i>rII</i> mutant, a mutation that caused the bacteriophage to eliminate bacteria more rapidly than usual, was not exhibiting the expected phenotype, it occurred to Benzer that this strain might have come from a cross between two different <i>rII</i> mutants (each having part of the <i>rII</i> gene intact) wherein a <a href="/wiki/Genetic_recombination" title="Genetic recombination">recombination</a> event resulted in a normal <i>rII</i> sequence. Benzer realized that by generating many <i>r</i> mutants and recording the <a href="/wiki/Recombination_frequency" class="mw-redirect" title="Recombination frequency">recombination frequency</a> between different <i>r</i> strains, one could create a detailed map of the gene, much as <a href="/wiki/Alfred_Sturtevant" title="Alfred Sturtevant">Alfred Sturtevant</a> had done for <a href="/wiki/Chromosomes" class="mw-redirect" title="Chromosomes">chromosomes</a>.<sup id="cite_ref-Weiner_8-2" class="reference"><a href="#cite_note-Weiner-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> Taking advantage of the enormous number of recombinants that could be analyzed in the <i>rII</i> mutant system, Benzer was eventually able to map over 2400 <i>rII</i> mutations. The data he collected provided the first evidence that the gene is not an indivisible entity, as previously believed, and that genes were linear.<sup id="cite_ref-topology_12-0" class="reference"><a href="#cite_note-topology-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-topography_11-1" class="reference"><a href="#cite_note-topography-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> Benzer also proved that <a href="/wiki/Mutation" title="Mutation">mutations</a> were distributed in many different parts of a single gene, and the resolving power of his system allowed him to discern mutants that differ at the level of a single <a href="/wiki/Nucleotide" title="Nucleotide">nucleotide</a>. Based on his <i>rII</i> data, Benzer also proposed distinct classes of mutations including <a href="/wiki/Deletion_(genetics)" title="Deletion (genetics)">deletions</a>, <a href="/wiki/Point_mutations" class="mw-redirect" title="Point mutations">point mutations</a>, <a href="/wiki/Missense_mutations" class="mw-redirect" title="Missense mutations">missense mutations</a>, and <a href="/wiki/Nonsense_mutations" class="mw-redirect" title="Nonsense mutations">nonsense mutations</a>.<sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p><p>Benzer's work influenced many other scientists of his time (see <a href="/wiki/Phage_group" title="Phage group">Phage group</a>). In his molecular biology period, Benzer dissected the fine structure of a single gene, laying down the ground work for decades of mutation analysis and genetic engineering, and setting up a paradigm using the <i>rII</i> phage that would later be used by <a href="/wiki/Francis_Crick" title="Francis Crick">Francis Crick</a> and <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Sydney Brenner</a> to establish the <a href="/wiki/Triplet_code" class="mw-redirect" title="Triplet code">triplet code</a> of <a href="/wiki/DNA" title="DNA">DNA</a>. In addition, Benzer's mapping technique was taken up by <a href="/wiki/Richard_Feynman" title="Richard Feynman">Richard Feynman</a>.<sup id="cite_ref-Weiner_8-3" class="reference"><a href="#cite_note-Weiner-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p><p>In 1967, Benzer left the field of phage genetics and returned to the California Institute of Technology to work in <a href="/wiki/Behavioral_genetics" class="mw-redirect" title="Behavioral genetics">behavioral genetics</a>. </p> <div class="mw-heading mw-heading3"><h3 id="Behavioral_genetics">Behavioral genetics</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Seymour_Benzer&action=edit&section=6" title="Edit section: Behavioral genetics"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading4"><h4 id="Benzer_vs._Hirsch">Benzer vs. Hirsch</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Seymour_Benzer&action=edit&section=7" title="Edit section: Benzer vs. Hirsch"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Benzer was one of the first scientists to rise to prominence in the field of <a href="/wiki/Behavioral_genetics" class="mw-redirect" title="Behavioral genetics">behavioral genetics</a>. As the field began to emerge in the 1960s and 70s, Benzer found himself in scientific opposition to another of the field's leading researchers, <a href="/wiki/Jerry_Hirsch" title="Jerry Hirsch">Jerry Hirsch</a>. While Hirsch believed that behaviors were complex phenomena irreducible to the level of single genes, Benzer advocated that animal behaviors were not too complex to be directed by a single gene. This translated to methodological differences in the two researchers' experiments with <a href="/wiki/Drosophila" title="Drosophila">Drosophila</a> that profoundly influenced the field of behavioral genetics. Hirsch artificially selected for behaviors of interest over many generations, while Benzer primarily used <a href="/wiki/Forward_genetics" title="Forward genetics">forward genetic</a> mutagenesis screens to isolate mutants for a particular behavior.<sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> Benzer and Hirsch's competing philosophies served to provide necessary scientific tension in order to accelerate and enhance developments in behavioral genetics, helping it gain traction as a legitimate area of study in the scientific community. </p> <div class="mw-heading mw-heading4"><h4 id="Research_accomplishments">Research accomplishments</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Seymour_Benzer&action=edit&section=8" title="Edit section: Research accomplishments"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Benzer used forward genetics to investigate the genetic basis of various behaviors such as <a href="/wiki/Phototaxis" title="Phototaxis">phototaxis</a>, <a href="/wiki/Circadian_rhythms" class="mw-redirect" title="Circadian rhythms">circadian rhythms</a>, and learning by inducing mutations in a Drosophila population and then screening individuals for altered phenotypes of interest.<sup id="cite_ref-Retros_10-1" class="reference"><a href="#cite_note-Retros-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> To better identify mutants, Benzer developed novel apparatuses such as the countercurrent device, which was designed to separate flies according to the magnitude and direction of their phototactic response.<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> Benzer identified mutants for a wide variety of characteristics: vision (<i>nonphototactic</i>, <i>negative phototactic</i>, and <i>eyes absent</i><sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup>), locomotion (<i>sluggish</i>, <i>uncoordinated</i>), stress sensitivity (<i>freaked-out</i>), sexual function (<i>savoir-faire</i>, <i><a href="/wiki/Fruitless_(gene)" title="Fruitless (gene)">fruitless</a></i>), nerve and muscle function (<i>photoreceptor degeneration</i>, <i>drop-dead</i>), and learning and memory (<i><a href="/wiki/Rutabaga_(gene)" title="Rutabaga (gene)">rutabaga</a></i>, <i>dunce</i><sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup>).<sup id="cite_ref-genetobehavior_18-0" class="reference"><a href="#cite_note-genetobehavior-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> </p><p>Benzer and student <a href="/wiki/Ron_Konopka" class="mw-redirect" title="Ron Konopka">Ron Konopka</a> discovered the first circadian rhythm mutants. Three distinct mutant types—arrhythmic, shortened period, and lengthened period—were identified. These mutations all involved the same functional gene on the X chromosome and influenced the eclosion rhythm of the population as well as rhythms in individual flies' locomotor activity.<sup id="cite_ref-perMutants_19-0" class="reference"><a href="#cite_note-perMutants-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> To monitor <i>Drosophila</i> locomotor activity, Benzer and postdoctoral researcher, Yoshiki Hotta, designed a system using infrared light and solar cells.<sup id="cite_ref-perMutants_19-1" class="reference"><a href="#cite_note-perMutants-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> All three mutations were mapped to the X chromosome, zero centimorgans away from each other, indicating that the mutant phenotypes corresponded to alleles of the same gene, which Konopka named <i><a href="/wiki/Period_(gene)" title="Period (gene)">period</a></i>.<sup id="cite_ref-perMutants_19-2" class="reference"><a href="#cite_note-perMutants-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> This was the first of several seminal studies of single genes affecting behavior, studies that have been replicated in other animal models and are now the basis for the growing field of molecular biology of behavior. In 1992 Benzer, working with Michael Rosbash, furthered this work by showing that the PER protein, which <i><a href="/wiki/Period_(gene)" title="Period (gene)">period</a></i> codes for, is predominantly located in the nucleus.<sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> The work with Period mutants was catalytic in the study of circadian rhythms and served to propel the field forward. </p><p>On 2 October 2017, Dr. Rosbash, along with Drs. Michael W. Young and Jeffrey C. Hall, were awarded the Nobel Prize in Physiology or Medicine in recognition of their cloning of circadian rhythm genes, and the elucidation of the biochemical mechanisms by which the circadian rhythm protein products regulated behavior. </p><p>Benzer was at the forefront of the study of neurodegeneration in fruit flies, modeling human diseases and attempting to suppress them. He also contributed to the field of aging biology, looking for mutants with altered longevity and trying to dissect the mechanisms by which an organism can escape the inevitable functional downfall and its associated diseases.<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> In 1998, Benzer and his colleagues Yi-Jyun Lin and Laurent Seroude published findings of a long-life mutant in <i>Drosophila</i>, then named Methuselah. The mutant gene coded for a previously unknown member of the <a href="/wiki/GPCR" class="mw-redirect" title="GPCR">GPCR</a> family. By testing against temperature stress, it is thought that these mutants have an increased ability to respond to stress and thus to live longer.<sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup> One of Benzer's final research projects was on dietary restriction and longevity research. A paper was published, in '<a href="/wiki/Cell_magazine" class="mw-redirect" title="Cell magazine">'Cell'</a>', on the longevity effect of 4E-BP, a translational repressor, following dietary restriction. Although the research was done before his death, the paper was published afterwards and dedicated to his memory.<sup id="cite_ref-23" class="reference"><a href="#cite_note-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Cancer_research">Cancer research</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Seymour_Benzer&action=edit&section=9" title="Edit section: Cancer research"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>In 1978, Dotty was in the hospital with breast cancer, and Seymour's friend, colleague, and mentor <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Max Delbrück</a> was diagnosed with cancer. Consequently, Seymour Benzer took interest in cancer biology and attended several conferences on breast cancer.<sup id="cite_ref-Weiner_8-4" class="reference"><a href="#cite_note-Weiner-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> Benzer later remarried with Carol Miller, a <a href="/wiki/Neuropathology" title="Neuropathology">neuropathologist</a>. Together, in the early 1980s, they used <a href="/wiki/Immunostaining" title="Immunostaining">antibody staining</a> techniques to find nearly identical <a href="/wiki/Genes" class="mw-redirect" title="Genes">genes</a> between flies and humans.<sup id="cite_ref-Weiner_8-5" class="reference"><a href="#cite_note-Weiner-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Honors_and_awards">Honors and awards</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Seymour_Benzer&action=edit&section=10" title="Edit section: Honors and awards"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Seymour_Benzer_1986.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Seymour_Benzer_1986.png/220px-Seymour_Benzer_1986.png" decoding="async" width="220" height="187" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Seymour_Benzer_1986.png/330px-Seymour_Benzer_1986.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/ad/Seymour_Benzer_1986.png/440px-Seymour_Benzer_1986.png 2x" data-file-width="942" data-file-height="800" /></a><figcaption>Benzer with <a href="/wiki/Ronald_Reagan" title="Ronald Reagan">Ronald Reagan</a> in 1986</figcaption></figure> <ul><li>Fellow of the <a href="/wiki/American_Academy_of_Arts_and_Sciences" title="American Academy of Arts and Sciences">American Academy of Arts and Sciences</a> (1959)<sup id="cite_ref-AAAS_24-0" class="reference"><a href="#cite_note-AAAS-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Gairdner_Foundation_International_Award" class="mw-redirect" title="Gairdner Foundation International Award">Gairdner Foundation International Award</a> (1964)</li> <li><a href="/wiki/Albert_Lasker_Award_for_Basic_Medical_Research" title="Albert Lasker Award for Basic Medical Research">Albert Lasker Award for Basic Medical Research</a> (1971)</li> <li><a href="/wiki/Louisa_Gross_Horwitz_Prize" title="Louisa Gross Horwitz Prize">Louisa Gross Horwitz Prize</a> from <a href="/wiki/Columbia_University" title="Columbia University">Columbia University</a> (1976),<sup id="cite_ref-25" class="reference"><a href="#cite_note-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Harvey_Prize" title="Harvey Prize">Harvey Prize</a> (1977)</li> <li><a href="/wiki/Dickson_Prize_in_Science" class="mw-redirect" title="Dickson Prize in Science">Dickson Prize in Science</a> (1978)</li> <li><a href="/wiki/National_Medal_of_Science" title="National Medal of Science">National Medal of Science</a> (1982)</li> <li><a href="/wiki/Rosenstiel_Award" title="Rosenstiel Award">Rosenstiel Award</a> (1985)</li> <li><a href="/wiki/Thomas_Hunt_Morgan_Medal" title="Thomas Hunt Morgan Medal">Thomas Hunt Morgan Medal</a> (1986)</li> <li><a href="/wiki/Ralph_W._Gerard_Prize_in_Neuroscience" title="Ralph W. Gerard Prize in Neuroscience">Ralph W. Gerard Prize in Neuroscience</a> (1989)</li> <li><a href="/wiki/Wolf_Prize_in_Medicine" title="Wolf Prize in Medicine">Wolf Prize in Medicine</a> (1991)</li> <li><a href="/wiki/Crafoord_Prize" title="Crafoord Prize">Crafoord Prize</a> (1993)<sup id="cite_ref-Crafoord_Prize_Benzer_1993_26-0" class="reference"><a href="#cite_note-Crafoord_Prize_Benzer_1993-26"><span class="cite-bracket">[</span>26<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Feltrinelli_Prize" title="Feltrinelli Prize">Feltrinelli Prize</a> (1994)</li> <li><a href="/wiki/International_Prize_for_Biology" title="International Prize for Biology">International Prize for Biology</a> (2000)</li> <li><a href="/wiki/NAS_Award_in_the_Neurosciences" title="NAS Award in the Neurosciences">NAS Award in the Neurosciences</a> from the <a href="/wiki/United_States_National_Academy_of_Sciences" class="mw-redirect" title="United States National Academy of Sciences">National Academy of Sciences</a> (2001)<sup id="cite_ref-Elliot_27-0" class="reference"><a href="#cite_note-Elliot-27"><span class="cite-bracket">[</span>27<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/March_of_Dimes_Prize_in_Developmental_Biology" title="March of Dimes Prize in Developmental Biology">March of Dimes Prize in Developmental Biology</a> (2002)</li> <li><a href="/wiki/Gairdner_Foundation_International_Award" class="mw-redirect" title="Gairdner Foundation International Award">Gairdner Foundation International Award</a> (2004) (second award)</li> <li><a href="/wiki/The_Franklin_Institute_Awards" class="mw-redirect" title="The Franklin Institute Awards">Bower Award and Prize for Achievement in Science (2004)</a></li> <li><a href="/wiki/Albany_Medical_Center_Prize" title="Albany Medical Center Prize">Albany Medical Center Prize</a> (2006)</li></ul> <p>He was a member of the <a href="/wiki/French_Academy_of_Sciences" title="French Academy of Sciences">French Academy of Sciences</a>, the U.S. National Academy of Sciences, the <a href="/wiki/American_Philosophical_Society" title="American Philosophical Society">American Philosophical Society</a> and the <a href="/wiki/Royal_Society" title="Royal Society">Royal Society</a>. </p> <div class="mw-heading mw-heading2"><h2 id="Books">Books</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Seymour_Benzer&action=edit&section=11" title="Edit section: Books"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Benzer is the subject of the 1999 book <i><a href="/wiki/Time,_Love,_Memory" title="Time, Love, Memory">Time, Love, Memory</a>: A Great Biologist and His Quest for the Origins of Behavior</i> by <a href="/wiki/Pulitzer_Prize" title="Pulitzer Prize">Pulitzer laureate</a> <a href="/wiki/Jonathan_Weiner" title="Jonathan Weiner">Jonathan Weiner</a>,<sup id="cite_ref-Weiner_8-6" class="reference"><a href="#cite_note-Weiner-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> and <i>Reconceiving the Gene: Seymour Benzer's Adventures in Phage Genetics</i> by Lawrence Holmes.<sup id="cite_ref-Holmes_6-1" class="reference"><a href="#cite_note-Holmes-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="See_also">See also</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Seymour_Benzer&action=edit&section=12" title="Edit section: See also"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li><a href="/wiki/Phage_group" title="Phage group">Phage group</a></li> <li><a href="/wiki/T4_rII_system" title="T4 rII system">T4 rII system</a></li></ul> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Seymour_Benzer&action=edit&section=13" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width" style="column-width: 30em;"> <ol class="references"> <li id="cite_note-frs-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-frs_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-frs_1-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFGreenspan2012" class="citation journal cs1">Greenspan, R. 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(2012). <a rel="nofollow" class="external text" href="https://doi.org/10.1098%2Frsbm.2012.0031">"Seymour Benzer. 15 October 1921 -- 30 November 2007"</a>. <i><a href="/wiki/Biographical_Memoirs_of_Fellows_of_the_Royal_Society" title="Biographical Memoirs of Fellows of the Royal Society">Biographical Memoirs of Fellows of the Royal Society</a></i>. <b>58</b>: 23–32. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1098%2Frsbm.2012.0031">10.1098/rsbm.2012.0031</a></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Biographical+Memoirs+of+Fellows+of+the+Royal+Society&rft.atitle=Seymour+Benzer.+15+October+1921+--+30+November+2007&rft.volume=58&rft.pages=23-32&rft.date=2012&rft_id=info%3Adoi%2F10.1098%2Frsbm.2012.0031&rft.aulast=Greenspan&rft.aufirst=R.+J.&rft_id=https%3A%2F%2Fdoi.org%2F10.1098%252Frsbm.2012.0031&rfr_id=info%3Asid%2Fen.wikipedia.org%3ASeymour+Benzer" class="Z3988"></span></span> </li> <li id="cite_note-Greenspan_2008_R106–10-2"><span class="mw-cite-backlink">^ <a href="#cite_ref-Greenspan_2008_R106–10_2-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Greenspan_2008_R106–10_2-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFGreenspan2008" class="citation journal cs1">Greenspan, R. J. (2008). <a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.cub.2007.12.039">"Seymour Benzer (1921–2007)"</a>. <i>Current Biology</i>. <b>18</b> (3): R106–R110. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2008CBio...18.R106G">2008CBio...18.R106G</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.cub.2007.12.039">10.1016/j.cub.2007.12.039</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/18345547">18345547</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:2094146">2094146</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Current+Biology&rft.atitle=Seymour+Benzer+%281921%E2%80%932007%29&rft.volume=18&rft.issue=3&rft.pages=R106-R110&rft.date=2008&rft_id=info%3Adoi%2F10.1016%2Fj.cub.2007.12.039&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A2094146%23id-name%3DS2CID&rft_id=info%3Apmid%2F18345547&rft_id=info%3Abibcode%2F2008CBio...18.R106G&rft.aulast=Greenspan&rft.aufirst=R.+J.&rft_id=https%3A%2F%2Fdoi.org%2F10.1016%252Fj.cub.2007.12.039&rfr_id=info%3Asid%2Fen.wikipedia.org%3ASeymour+Benzer" class="Z3988"></span></span> </li> <li id="cite_note-nytzimmer-3"><span class="mw-cite-backlink">^ <a href="#cite_ref-nytzimmer_3-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-nytzimmer_3-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-nytzimmer_3-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCarl_Zimmer2007" class="citation news cs1"><a href="/wiki/Carl_Zimmer" title="Carl Zimmer">Carl Zimmer</a> (8 December 2007). <a rel="nofollow" class="external text" href="https://www.nytimes.com/2007/12/08/science/08benzer.html?ref=science">"Seymour Benzer, geneticist, is dead at 86"</a>. <i>The New York Times</i><span class="reference-accessdate">. Retrieved <span class="nowrap">2008-02-18</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+New+York+Times&rft.atitle=Seymour+Benzer%2C+geneticist%2C+is+dead+at+86&rft.date=2007-12-08&rft.au=Carl+Zimmer&rft_id=https%3A%2F%2Fwww.nytimes.com%2F2007%2F12%2F08%2Fscience%2F08benzer.html%3Fref%3Dscience&rfr_id=info%3Asid%2Fen.wikipedia.org%3ASeymour+Benzer" class="Z3988"></span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.geni.com/people/Seymor-Benzer/6000000040552313938">"Seymor Benzer"</a>. 15 October 1921.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Seymor+Benzer&rft.date=1921-10-15&rft_id=https%3A%2F%2Fwww.geni.com%2Fpeople%2FSeymor-Benzer%2F6000000040552313938&rfr_id=info%3Asid%2Fen.wikipedia.org%3ASeymour+Benzer" class="Z3988"></span></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-5">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://resolver.caltech.edu/CaltechOH:OH_Benzer_S">"Interview with Seymour Benzer"</a>. <i>CaltechOralHistories</i>. January 2002<span class="reference-accessdate">. Retrieved <span class="nowrap">28 June</span> 2024</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=CaltechOralHistories&rft.atitle=Interview+with+Seymour+Benzer&rft.date=2002-01&rft_id=https%3A%2F%2Fresolver.caltech.edu%2FCaltechOH%3AOH_Benzer_S&rfr_id=info%3Asid%2Fen.wikipedia.org%3ASeymour+Benzer" class="Z3988"></span></span> </li> <li id="cite_note-Holmes-6"><span class="mw-cite-backlink">^ <a href="#cite_ref-Holmes_6-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Holmes_6-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHolmesSummers,_William_C.2006" class="citation book cs1">Holmes, Frederic Lawrence; Summers, William C. (2006). <i>Reconceiving the Gene: Seymour Benzer's Adventures in Phage Genetics</i>. New Haven: Yale University Press. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-300-11078-4" title="Special:BookSources/978-0-300-11078-4"><bdi>978-0-300-11078-4</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Reconceiving+the+Gene%3A+Seymour+Benzer%27s+Adventures+in+Phage+Genetics&rft.place=New+Haven&rft.pub=Yale+University+Press&rft.date=2006&rft.isbn=978-0-300-11078-4&rft.aulast=Holmes&rft.aufirst=Frederic+Lawrence&rft.au=Summers%2C+William+C.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ASeymour+Benzer" class="Z3988"></span></span> </li> <li id="cite_note-oralhistory-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-oralhistory_7-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBenzer1991" class="citation web cs1">Benzer, Seymour (1991). <a rel="nofollow" class="external text" href="http://resolver.caltech.edu/CaltechOH:OH_Benzer_S">"Seymour Benzer (1921-2007) Interviewed by Heidi Asputrian"</a>. <i>Oral History Project</i>. California Institute of Technology Archives<span class="reference-accessdate">. Retrieved <span class="nowrap">19 April</span> 2011</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Oral+History+Project&rft.atitle=Seymour+Benzer+%281921-2007%29+Interviewed+by+Heidi+Asputrian&rft.date=1991&rft.aulast=Benzer&rft.aufirst=Seymour&rft_id=http%3A%2F%2Fresolver.caltech.edu%2FCaltechOH%3AOH_Benzer_S&rfr_id=info%3Asid%2Fen.wikipedia.org%3ASeymour+Benzer" class="Z3988"></span></span> </li> <li id="cite_note-Weiner-8"><span class="mw-cite-backlink">^ <a href="#cite_ref-Weiner_8-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Weiner_8-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-Weiner_8-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-Weiner_8-3"><sup><i><b>d</b></i></sup></a> <a href="#cite_ref-Weiner_8-4"><sup><i><b>e</b></i></sup></a> <a href="#cite_ref-Weiner_8-5"><sup><i><b>f</b></i></sup></a> <a href="#cite_ref-Weiner_8-6"><sup><i><b>g</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWeiner1999" class="citation book cs1">Weiner, Jonathan (1999). <span class="id-lock-registration" title="Free registration required"><a rel="nofollow" class="external text" href="https://archive.org/details/timelovememorygr00wein"><i>Time, Love, Memory: A Great Biologist and His Quest for the Origins of Behavior</i></a></span>. New York: Knopf. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-679-44435-0" title="Special:BookSources/978-0-679-44435-0"><bdi>978-0-679-44435-0</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Time%2C+Love%2C+Memory%3A+A+Great+Biologist+and+His+Quest+for+the+Origins+of+Behavior&rft.place=New+York&rft.pub=Knopf&rft.date=1999&rft.isbn=978-0-679-44435-0&rft.aulast=Weiner&rft.aufirst=Jonathan&rft_id=https%3A%2F%2Farchive.org%2Fdetails%2Ftimelovememorygr00wein&rfr_id=info%3Asid%2Fen.wikipedia.org%3ASeymour+Benzer" class="Z3988"></span></span> </li> <li id="cite_note-caltechobit-9"><span class="mw-cite-backlink">^ <a href="#cite_ref-caltechobit_9-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-caltechobit_9-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation pressrelease cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20080208080709/http://mr.caltech.edu/media/Press_Releases/PR13076.html">"Neurogenetics Pioneer Seymour Benzer Dies"</a> (Press release). California Institute of Technology. 30 November 2007. Archived from <a rel="nofollow" class="external text" href="http://mr.caltech.edu/media/Press_Releases/PR13076.html">the original</a> on 8 February 2008<span class="reference-accessdate">. Retrieved <span class="nowrap">2008-02-18</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Neurogenetics+Pioneer+Seymour+Benzer+Dies&rft.pub=California+Institute+of+Technology&rft.date=2007-11-30&rft_id=http%3A%2F%2Fmr.caltech.edu%2Fmedia%2FPress_Releases%2FPR13076.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3ASeymour+Benzer" class="Z3988"></span></span> </li> <li id="cite_note-Retros-10"><span class="mw-cite-backlink">^ <a href="#cite_ref-Retros_10-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Retros_10-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFJanJan2008" class="citation journal cs1">Jan, Y. -N.; Jan, L. (2008). "Retrospective: Seymour Benzer (1921-2007)". <i>Science</i>. <b>319</b> (5859): 45. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.1154050">10.1126/science.1154050</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/18174427">18174427</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:206511019">206511019</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Science&rft.atitle=Retrospective%3A+Seymour+Benzer+%281921-2007%29&rft.volume=319&rft.issue=5859&rft.pages=45&rft.date=2008&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A206511019%23id-name%3DS2CID&rft_id=info%3Apmid%2F18174427&rft_id=info%3Adoi%2F10.1126%2Fscience.1154050&rft.aulast=Jan&rft.aufirst=Y.+-N.&rft.au=Jan%2C+L.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ASeymour+Benzer" class="Z3988"></span></span> </li> <li id="cite_note-topography-11"><span class="mw-cite-backlink">^ <a href="#cite_ref-topography_11-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-topography_11-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBenzer1961" class="citation journal cs1"><a class="mw-selflink selflink">Benzer, S.</a> (1961). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC221592">"On the Topography of the Genetic Fine Structure"</a>. <i>Proceedings of the National Academy of Sciences</i>. <b>47</b> (3): 403–415. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1961PNAS...47..403B">1961PNAS...47..403B</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1073%2Fpnas.47.3.403">10.1073/pnas.47.3.403</a></span>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC221592">221592</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/16590840">16590840</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Proceedings+of+the+National+Academy+of+Sciences&rft.atitle=On+the+Topography+of+the+Genetic+Fine+Structure&rft.volume=47&rft.issue=3&rft.pages=403-415&rft.date=1961&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC221592%23id-name%3DPMC&rft_id=info%3Apmid%2F16590840&rft_id=info%3Adoi%2F10.1073%2Fpnas.47.3.403&rft_id=info%3Abibcode%2F1961PNAS...47..403B&rft.aulast=Benzer&rft.aufirst=S.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC221592&rfr_id=info%3Asid%2Fen.wikipedia.org%3ASeymour+Benzer" class="Z3988"></span></span> </li> <li id="cite_note-topology-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-topology_12-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBenzer1959" class="citation journal cs1"><a class="mw-selflink selflink">Benzer, S.</a> (1959). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC222769">"On the Topology of the Genetic Fine Structure"</a>. <i>Proceedings of the National Academy of Sciences</i>. <b>45</b> (11): 1607–20. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1959PNAS...45.1607B">1959PNAS...45.1607B</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1073%2Fpnas.45.11.1607">10.1073/pnas.45.11.1607</a></span>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC222769">222769</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/16590553">16590553</a>.</cite><span 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Columbia University Medical Center<span class="reference-accessdate">. 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abbr{font-variant:small-caps;border-bottom:none;text-decoration:none;cursor:inherit}.mw-parser-output .navbar-ct-full{font-size:114%;margin:0 7em}.mw-parser-output .navbar-ct-mini{font-size:114%;margin:0 4em}html.skin-theme-clientpref-night .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}@media(prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navbar li a abbr{color:var(--color-base)!important}}@media print{.mw-parser-output .navbar{display:none!important}}</style><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Wolf_Prize_in_Medicine" title="Template:Wolf Prize in Medicine"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Wolf_Prize_in_Medicine" title="Template talk:Wolf Prize in Medicine"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Wolf_Prize_in_Medicine" title="Special:EditPage/Template:Wolf Prize in Medicine"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Laureates_of_the_Wolf_Prize_in_Medicine" style="font-size:114%;margin:0 4em">Laureates of the <a href="/wiki/Wolf_Prize_in_Medicine" title="Wolf Prize in Medicine">Wolf Prize in Medicine</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1970s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/George_Davis_Snell" title="George Davis Snell">George Snell</a> / <a href="/wiki/Jean_Dausset" title="Jean Dausset">Jean Dausset</a> / <a href="/wiki/Jon_van_Rood" title="Jon van Rood">Jon J. van Rood</a> (1978)</li> <li><a href="/wiki/Roger_Wolcott_Sperry" title="Roger Wolcott Sperry">Roger Sperry</a> / <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Arvid Carlsson</a> / <a href="/wiki/Oleh_Hornykiewicz" title="Oleh Hornykiewicz">Oleh Hornykiewicz</a> (1979)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1980s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">César Milstein</a> / <a href="/wiki/Leo_Sachs" title="Leo Sachs">Leo Sachs</a> / <a href="/wiki/James_Learmonth_Gowans" title="James Learmonth Gowans">James L. Gowans</a> (1980)</li> <li><a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a> / <a href="/wiki/Stanley_Norman_Cohen" title="Stanley Norman Cohen">Stanley Norman Cohen</a> (1981)</li> <li><a href="/wiki/Jean-Pierre_Changeux" title="Jean-Pierre Changeux">Jean-Pierre Changeux</a> / <a href="/wiki/Solomon_H._Snyder" title="Solomon H. Snyder">Solomon H. Snyder</a> / <a href="/wiki/James_Black_(pharmacologist)" title="James Black (pharmacologist)">James W. Black</a> (1982)</li> <li><a href="/wiki/Donald_F._Steiner" title="Donald F. Steiner">Donald F. Steiner</a> (1984/5)</li> <li><a href="/wiki/Osamu_Hayaishi" title="Osamu Hayaishi">Osamu Hayaishi</a> (1986)</li> <li><a href="/wiki/Pedro_Cuatrecasas" title="Pedro Cuatrecasas">Pedro Cuatrecasas</a> / <a href="/wiki/Meir_Wilchek" title="Meir Wilchek">Meir Wilchek</a> (1987)</li> <li><a href="/wiki/Henri_G._Hers" title="Henri G. Hers">Henri G. Hers</a> / <a href="/wiki/Elizabeth_F._Neufeld" title="Elizabeth F. Neufeld">Elizabeth F. Neufeld</a> (1988)</li> <li><a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a> / <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Edward B. Lewis</a> (1989)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">1990s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Maclyn_McCarty" title="Maclyn McCarty">Maclyn McCarty</a> (1990)</li> <li><a class="mw-selflink selflink">Seymour Benzer</a> (1991)</li> <li><a href="/wiki/Judah_Folkman" title="Judah Folkman">Judah Folkman</a> (1992)</li> <li><a href="/wiki/Michael_Berridge" title="Michael Berridge">Michael Berridge</a> / <a href="/wiki/Yasutomi_Nishizuka" title="Yasutomi Nishizuka">Yasutomi Nishizuka</a> (1994/5)</li> <li><a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Stanley B. Prusiner</a> (1995/6)</li> <li><a href="/wiki/Mary_F._Lyon" title="Mary F. Lyon">Mary F. Lyon</a> (1996/7)</li> <li><a href="/wiki/Michael_Sela" title="Michael Sela">Michael Sela</a> / <a href="/wiki/Ruth_Arnon" title="Ruth Arnon">Ruth Arnon</a> (1998)</li> <li><a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric Kandel</a> (1999)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">2000s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Avram_Hershko" title="Avram Hershko">Avram Hershko</a> / <a href="/wiki/Alexander_Varshavsky" title="Alexander Varshavsky">Alexander Varshavsky</a> (2001)</li> <li><a href="/wiki/Ralph_L._Brinster" title="Ralph L. Brinster">Ralph L. Brinster</a> / <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Mario Capecchi</a> / <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a> (2002/3)</li> <li><a href="/wiki/Robert_Weinberg_(biologist)" title="Robert Weinberg (biologist)">Robert Weinberg</a> / <a href="/wiki/Roger_Y._Tsien" title="Roger Y. Tsien">Roger Y. Tsien</a> (2004)</li> <li><a href="/wiki/Alexander_Levitzki" title="Alexander Levitzki">Alexander Levitzki</a> / <a href="/wiki/Anthony_R._Hunter" title="Anthony R. Hunter">Anthony R. Hunter</a> / <a href="/wiki/Tony_Pawson_(biochemist)" title="Tony Pawson (biochemist)">Tony Pawson</a> (2005)</li> <li><a href="/wiki/Howard_Cedar" title="Howard Cedar">Howard Cedar</a> / <a href="/wiki/Aharon_Razin" title="Aharon Razin">Aharon Razin</a> (2008)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">2010s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Axel_Ullrich" title="Axel Ullrich">Axel Ullrich</a> (2010)</li> <li><a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Shinya Yamanaka</a> / <a href="/wiki/Rudolf_Jaenisch" title="Rudolf Jaenisch">Rudolf Jaenisch</a> (2011)</li> <li><a href="/wiki/Ronald_M._Evans" title="Ronald M. Evans">Ronald M. Evans</a> (2012)</li> <li><a href="/wiki/Nahum_Sonenberg" title="Nahum Sonenberg">Nahum Sonenberg</a> / <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Gary Ruvkun</a> / <a href="/wiki/Victor_Ambros" title="Victor Ambros">Victor Ambros</a> (2014)</li> <li><a href="/wiki/John_Kappler" title="John Kappler">John Kappler</a> / <a href="/wiki/Philippa_Marrack" title="Philippa Marrack">Philippa Marrack</a> / <a href="/wiki/Jeffrey_V._Ravetch" title="Jeffrey V. Ravetch">Jeffrey V. Ravetch</a> (2015)</li> <li><a href="/wiki/Lewis_C._Cantley" title="Lewis C. Cantley">Lewis C. Cantley</a> / <a href="/wiki/C._Ronald_Kahn" title="C. Ronald Kahn">C. Ronald Kahn</a> (2016)</li> <li><a href="/wiki/James_P._Allison" title="James P. Allison">James P. Allison</a> (2017)</li> <li><a href="/wiki/Jeffrey_M._Friedman" title="Jeffrey M. Friedman">Jeffrey M. Friedman</a> (2019)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">2020s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Emmanuelle_Charpentier" title="Emmanuelle Charpentier">Emmanuelle Charpentier</a> / <a href="/wiki/Jennifer_Doudna" title="Jennifer Doudna">Jennifer Doudna</a> (2020)</li> <li><a href="/wiki/Joan_Steitz" class="mw-redirect" title="Joan Steitz">Joan Steitz</a> / <a href="/wiki/Lynne_Maquat" class="mw-redirect" title="Lynne Maquat">Lynne Elizabeth Maquat</a> / <a href="/wiki/Adrian_R._Krainer" class="mw-redirect" title="Adrian R. Krainer">Adrian Krainer</a> (2021)</li> <li><a href="/wiki/Daniel_J._Drucker" title="Daniel J. Drucker">Daniel J. Drucker</a> (2023)</li> <li><a href="/wiki/Botond_Roska" title="Botond Roska">Botond Roska</a> / <a href="/wiki/Jos%C3%A9-Alain_Sahel" title="José-Alain Sahel">José-Alain Sahel</a> (2024)</li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="United_States_National_Medal_of_Science_laureates" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:National_Medal_of_Science" title="Template:National Medal of Science"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:National_Medal_of_Science" title="Template talk:National Medal of Science"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:National_Medal_of_Science" title="Special:EditPage/Template:National Medal of Science"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="United_States_National_Medal_of_Science_laureates" style="font-size:114%;margin:0 4em"><a href="/wiki/National_Medal_of_Science" title="National Medal of Science">United States National Medal of Science</a> <a href="/wiki/Laureate" title="Laureate">laureates</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Behavioral_and_social_science" style="font-size:114%;margin:0 4em">Behavioral and social science</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">1960s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1964</dt> <dd><a href="/wiki/Neal_E._Miller" title="Neal E. Miller">Neal Elgar Miller</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1980s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1986</dt> <dd><a href="/wiki/Herbert_A._Simon" title="Herbert A. Simon">Herbert A. Simon</a></dd> <dt>1987</dt> <dd><a href="/wiki/Anne_Anastasi" title="Anne Anastasi">Anne Anastasi</a></dd> <dd><a href="/wiki/George_Stigler" title="George Stigler">George J. Stigler</a></dd> <dt>1988</dt> <dd><a href="/wiki/Milton_Friedman" title="Milton Friedman">Milton Friedman</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1990s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1990</dt> <dd><a href="/wiki/Leonid_Hurwicz" title="Leonid Hurwicz">Leonid Hurwicz</a></dd> <dd><a href="/wiki/Patrick_Suppes" title="Patrick Suppes">Patrick Suppes</a></dd> <dt>1991</dt> <dd><a href="/wiki/George_Armitage_Miller" title="George Armitage Miller">George A. Miller</a></dd> <dt>1992</dt> <dd><a href="/wiki/Eleanor_J._Gibson" title="Eleanor J. Gibson">Eleanor J. Gibson</a></dd> <dt>1994</dt> <dd><a href="/wiki/Robert_K._Merton" title="Robert K. Merton">Robert K. Merton</a></dd> <dt>1995</dt> <dd><a href="/wiki/Roger_Shepard" title="Roger Shepard">Roger N. Shepard</a></dd> <dt>1996</dt> <dd><a href="/wiki/Paul_Samuelson" title="Paul Samuelson">Paul Samuelson</a></dd> <dt>1997</dt> <dd><a href="/wiki/William_Kaye_Estes" title="William Kaye Estes">William K. Estes</a></dd> <dt>1998</dt> <dd><a href="/wiki/William_Julius_Wilson" title="William Julius Wilson">William Julius Wilson</a></dd> <dt>1999</dt> <dd><a href="/wiki/Robert_Solow" title="Robert Solow">Robert M. Solow</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2000s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>2000</dt> <dd><a href="/wiki/Gary_Becker" title="Gary Becker">Gary Becker</a></dd> <dt>2003</dt> <dd><a href="/wiki/R._Duncan_Luce" title="R. Duncan Luce">R. Duncan Luce</a></dd> <dt>2004</dt> <dd><a href="/wiki/Kenneth_Arrow" title="Kenneth Arrow">Kenneth Arrow</a></dd> <dt>2005</dt> <dd><a href="/wiki/Gordon_H._Bower" title="Gordon H. Bower">Gordon H. Bower</a></dd> <dt>2008</dt> <dd><a href="/wiki/Michael_Posner_(psychologist)" title="Michael Posner (psychologist)">Michael I. Posner</a></dd> <dt>2009</dt> <dd><a href="/wiki/Mortimer_Mishkin" title="Mortimer Mishkin">Mortimer Mishkin</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2010s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>2011</dt> <dd><a href="/wiki/Anne_Treisman" title="Anne Treisman">Anne Treisman</a></dd> <dt>2014</dt> <dd><a href="/wiki/Robert_Axelrod_(political_scientist)" title="Robert Axelrod (political scientist)">Robert Axelrod</a></dd> <dt>2015</dt> <dd><a href="/wiki/Albert_Bandura" title="Albert Bandura">Albert Bandura</a></dd></dl> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible uncollapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Biological_sciences" style="font-size:114%;margin:0 4em">Biological sciences</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">1960s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1963</dt> <dd><a href="/wiki/C._B._van_Niel" title="C. B. van Niel">C. B. van Niel</a></dd> <dt>1964</dt> <dd><a href="/wiki/Theodosius_Dobzhansky" title="Theodosius Dobzhansky">Theodosius Dobzhansky</a></dd> <dd><a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Marshall W. Nirenberg</a></dd> <dt>1965</dt> <dd><a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Francis P. Rous</a></dd> <dd><a href="/wiki/George_Gaylord_Simpson" title="George Gaylord Simpson">George G. Simpson</a></dd> <dd><a href="/wiki/Donald_Van_Slyke" title="Donald Van Slyke">Donald D. Van Slyke</a></dd> <dt>1966</dt> <dd><a href="/wiki/Edward_F._Knipling" title="Edward F. Knipling">Edward F. Knipling</a></dd> <dd><a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Fritz Albert Lipmann</a></dd> <dd><a href="/wiki/William_Cumming_Rose" title="William Cumming Rose">William C. Rose</a></dd> <dd><a href="/wiki/Sewall_Wright" title="Sewall Wright">Sewall Wright</a></dd> <dt>1967</dt> <dd><a href="/wiki/Kenneth_Stewart_Cole" title="Kenneth Stewart Cole">Kenneth S. Cole</a></dd> <dd><a href="/wiki/Harry_Harlow" title="Harry Harlow">Harry F. Harlow</a></dd> <dd><a href="/wiki/Michael_Heidelberger" title="Michael Heidelberger">Michael Heidelberger</a></dd> <dd><a href="/wiki/Alfred_Sturtevant" title="Alfred Sturtevant">Alfred H. Sturtevant</a></dd> <dt>1968</dt> <dd><a href="/wiki/Horace_Barker" title="Horace Barker">Horace Barker</a></dd> <dd><a href="/wiki/Bernard_Brodie_(biochemist)" class="mw-redirect" title="Bernard Brodie (biochemist)">Bernard B. Brodie</a></dd> <dd><a href="/wiki/Detlev_Bronk" title="Detlev Bronk">Detlev W. Bronk</a></dd> <dd><a href="/wiki/Jay_Laurence_Lush" title="Jay Laurence Lush">Jay Lush</a></dd> <dd><a href="/wiki/B._F._Skinner" title="B. F. Skinner">Burrhus Frederic Skinner</a></dd> <dt>1969</dt> <dd><a href="/wiki/Robert_Huebner" title="Robert Huebner">Robert Huebner</a></dd> <dd><a href="/wiki/Ernst_Mayr" title="Ernst Mayr">Ernst Mayr</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1970s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1970</dt> <dd><a href="/wiki/Barbara_McClintock" title="Barbara McClintock">Barbara McClintock</a></dd> <dd><a href="/wiki/Albert_Sabin" title="Albert Sabin">Albert B. Sabin</a></dd> <dt>1973</dt> <dd><a href="/wiki/Daniel_I._Arnon" title="Daniel I. Arnon">Daniel I. Arnon</a></dd> <dd><a href="/wiki/Earl_W._Sutherland_Jr." class="mw-redirect" title="Earl W. Sutherland Jr.">Earl W. Sutherland Jr.</a></dd> <dt>1974</dt> <dd><a href="/wiki/Britton_Chance" title="Britton Chance">Britton Chance</a></dd> <dd><a href="/wiki/Erwin_Chargaff" title="Erwin Chargaff">Erwin Chargaff</a></dd> <dd><a href="/wiki/James_V._Neel" title="James V. Neel">James V. Neel</a></dd> <dd><a href="/wiki/James_Augustine_Shannon" class="mw-redirect" title="James Augustine Shannon">James Augustine Shannon</a></dd> <dt>1975</dt> <dd><a href="/wiki/Hallowell_Davis" title="Hallowell Davis">Hallowell Davis</a></dd> <dd><a href="/wiki/Paul_Gyorgy" title="Paul Gyorgy">Paul Gyorgy</a></dd> <dd><a href="/wiki/Sterling_B._Hendricks" title="Sterling B. Hendricks">Sterling B. Hendricks</a></dd> <dd><a href="/wiki/Orville_Vogel" title="Orville Vogel">Orville Alvin Vogel</a></dd> <dt>1976</dt> <dd><a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Roger Guillemin</a></dd> <dd><a href="/wiki/Keith_Roberts_Porter" class="mw-redirect" title="Keith Roberts Porter">Keith Roberts Porter</a></dd> <dd><a href="/wiki/Efraim_Racker" title="Efraim Racker">Efraim Racker</a></dd> <dd><a href="/wiki/E._O._Wilson" title="E. O. Wilson">E. O. Wilson</a></dd> <dt>1979</dt> <dd><a href="/wiki/Robert_H._Burris" title="Robert H. Burris">Robert H. Burris</a></dd> <dd><a href="/wiki/Elizabeth_C._Crosby" title="Elizabeth C. Crosby">Elizabeth C. Crosby</a></dd> <dd><a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Arthur Kornberg</a></dd> <dd><a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Severo Ochoa</a></dd> <dd><a href="/wiki/Earl_Reece_Stadtman" title="Earl Reece Stadtman">Earl Reece Stadtman</a></dd> <dd><a href="/wiki/G._Ledyard_Stebbins" title="G. Ledyard Stebbins">George Ledyard Stebbins</a></dd> <dd><a href="/wiki/Paul_Alfred_Weiss" title="Paul Alfred Weiss">Paul Alfred Weiss</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1980s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1981</dt> <dd><a href="/wiki/Philip_Handler" title="Philip Handler">Philip Handler</a></dd> <dt>1982</dt> <dd><a class="mw-selflink selflink">Seymour Benzer</a></dd> <dd><a href="/wiki/Glenn_W._Burton" title="Glenn W. Burton">Glenn W. Burton</a></dd> <dd><a href="/wiki/Mildred_Cohn" title="Mildred Cohn">Mildred Cohn</a></dd> <dt>1983</dt> <dd><a href="/wiki/Howard_Bachrach" title="Howard Bachrach">Howard L. Bachrach</a></dd> <dd><a href="/wiki/Paul_Berg" title="Paul Berg">Paul Berg</a></dd> <dd><a href="/wiki/Wendell_L._Roelofs" title="Wendell L. Roelofs">Wendell L. Roelofs</a></dd> <dd><a href="/wiki/Berta_Scharrer" title="Berta Scharrer">Berta Scharrer</a></dd> <dt>1986</dt> <dd><a href="/wiki/Stanley_Cohen_(biochemist)" title="Stanley Cohen (biochemist)">Stanley Cohen</a></dd> <dd><a href="/wiki/Donald_Henderson" title="Donald Henderson">Donald A. Henderson</a></dd> <dd><a href="/wiki/Vernon_Mountcastle" class="mw-redirect" title="Vernon Mountcastle">Vernon B. Mountcastle</a></dd> <dd><a href="/wiki/George_Emil_Palade" title="George Emil Palade">George Emil Palade</a></dd> <dd><a href="/wiki/Joan_A._Steitz" title="Joan A. Steitz">Joan A. Steitz</a></dd> <dt>1987</dt> <dd><a href="/wiki/Michael_E._DeBakey" class="mw-redirect" title="Michael E. DeBakey">Michael E. DeBakey</a></dd> <dd><a href="/wiki/Theodor_O._Diener" class="mw-redirect" title="Theodor O. Diener">Theodor O. Diener</a></dd> <dd><a href="/wiki/Harry_Eagle" title="Harry Eagle">Harry Eagle</a></dd> <dd><a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Har Gobind Khorana</a></dd> <dd><a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Rita Levi-Montalcini</a></dd> <dt>1988</dt> <dd><a href="/wiki/Michael_S._Brown" class="mw-redirect" title="Michael S. Brown">Michael S. Brown</a></dd> <dd><a href="/wiki/Stanley_Norman_Cohen" title="Stanley Norman Cohen">Stanley Norman Cohen</a></dd> <dd><a href="/wiki/Joseph_L._Goldstein" title="Joseph L. Goldstein">Joseph L. Goldstein</a></dd> <dd><a href="/wiki/Maurice_Hilleman" title="Maurice Hilleman">Maurice R. Hilleman</a></dd> <dd><a href="/wiki/Eric_Kandel" title="Eric Kandel">Eric R. Kandel</a></dd> <dd><a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Rosalyn Sussman Yalow</a></dd> <dt>1989</dt> <dd><a href="/wiki/Katherine_Esau" title="Katherine Esau">Katherine Esau</a></dd> <dd><a href="/wiki/Viktor_Hamburger" title="Viktor Hamburger">Viktor Hamburger</a></dd> <dd><a href="/wiki/Philip_Leder" title="Philip Leder">Philip Leder</a></dd> <dd><a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Joshua Lederberg</a></dd> <dd><a href="/wiki/Roger_W._Sperry" class="mw-redirect" title="Roger W. Sperry">Roger W. Sperry</a></dd> <dd><a href="/wiki/Harland_G._Wood" title="Harland G. Wood">Harland G. Wood</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1990s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1990</dt> <dd><a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Baruj Benacerraf</a></dd> <dd><a href="/wiki/Herbert_W._Boyer" class="mw-redirect" title="Herbert W. Boyer">Herbert W. Boyer</a></dd> <dd><a href="/wiki/Daniel_E._Koshland_Jr." title="Daniel E. Koshland Jr.">Daniel E. Koshland Jr.</a></dd> <dd><a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Edward B. Lewis</a></dd> <dd><a href="/wiki/David_G._Nathan" title="David G. Nathan">David G. Nathan</a></dd> <dd><a href="/wiki/E._Donnall_Thomas" title="E. Donnall Thomas">E. Donnall Thomas</a></dd> <dt>1991</dt> <dd><a href="/wiki/Mary_Ellen_Avery" title="Mary Ellen Avery">Mary Ellen Avery</a></dd> <dd><a href="/wiki/G._Evelyn_Hutchinson" title="G. Evelyn Hutchinson">G. Evelyn Hutchinson</a></dd> <dd><a href="/wiki/Elvin_A._Kabat" title="Elvin A. Kabat">Elvin A. Kabat</a></dd> <dd><a href="/wiki/Robert_Kates" title="Robert Kates">Robert W. Kates</a></dd> <dd><a href="/wiki/Salvador_Luria" title="Salvador Luria">Salvador Luria</a></dd> <dd><a href="/wiki/Paul_Marks_(scientist)" title="Paul Marks (scientist)">Paul A. Marks</a></dd> <dd><a href="/wiki/Folke_K._Skoog" title="Folke K. Skoog">Folke K. Skoog</a></dd> <dd><a href="/wiki/Paul_C._Zamecnik" class="mw-redirect" title="Paul C. Zamecnik">Paul C. Zamecnik</a></dd> <dt>1992</dt> <dd><a href="/wiki/Maxine_Singer" title="Maxine Singer">Maxine Singer</a></dd> <dd><a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Howard Martin Temin</a></dd> <dt>1993</dt> <dd><a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Daniel Nathans</a></dd> <dd><a href="/wiki/Salome_G._Waelsch" class="mw-redirect" title="Salome G. Waelsch">Salome G. Waelsch</a></dd> <dt>1994</dt> <dd><a href="/wiki/Thomas_Eisner" title="Thomas Eisner">Thomas Eisner</a></dd> <dd><a href="/wiki/Elizabeth_F._Neufeld" title="Elizabeth F. Neufeld">Elizabeth F. Neufeld</a></dd> <dt>1995</dt> <dd><a href="/wiki/Alexander_Rich" title="Alexander Rich">Alexander Rich</a></dd> <dt>1996</dt> <dd><a href="/wiki/Ruth_Patrick" title="Ruth Patrick">Ruth Patrick</a></dd> <dt>1997</dt> <dd><a href="/wiki/James_Watson" title="James Watson">James Watson</a></dd> <dd><a href="/wiki/Robert_A._Weinberg" class="mw-redirect" title="Robert A. Weinberg">Robert A. Weinberg</a></dd> <dt>1998</dt> <dd><a href="/wiki/Bruce_Ames" title="Bruce Ames">Bruce Ames</a></dd> <dd><a href="/wiki/Janet_Rowley" title="Janet Rowley">Janet Rowley</a></dd> <dt>1999</dt> <dd><a href="/wiki/David_Baltimore" title="David Baltimore">David Baltimore</a></dd> <dd><a href="/wiki/Jared_Diamond" title="Jared Diamond">Jared Diamond</a></dd> <dd><a href="/wiki/Lynn_Margulis" title="Lynn Margulis">Lynn Margulis</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2000s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>2000</dt> <dd><a href="/wiki/Nancy_C._Andreasen" class="mw-redirect" title="Nancy C. Andreasen">Nancy C. Andreasen</a></dd> <dd><a href="/wiki/Peter_H._Raven" title="Peter H. Raven">Peter H. Raven</a></dd> <dd><a href="/wiki/Carl_Woese" title="Carl Woese">Carl Woese</a></dd> <dt>2001</dt> <dd><a href="/wiki/Francisco_J._Ayala" title="Francisco J. Ayala">Francisco J. Ayala</a></dd> <dd><a href="/wiki/George_Bass_(archaeologist)" title="George Bass (archaeologist)">George F. Bass</a></dd> <dd><a href="/wiki/Mario_R._Capecchi" class="mw-redirect" title="Mario R. Capecchi">Mario R. Capecchi</a></dd> <dd><a href="/wiki/Ann_Graybiel" title="Ann Graybiel">Ann Graybiel</a></dd> <dd><a href="/wiki/Gene_Likens" title="Gene Likens">Gene E. Likens</a></dd> <dd><a href="/wiki/Victor_A._McKusick" title="Victor A. McKusick">Victor A. McKusick</a></dd> <dd><a href="/wiki/Harold_Varmus" class="mw-redirect" title="Harold Varmus">Harold Varmus</a></dd> <dt>2002</dt> <dd><a href="/wiki/James_E._Darnell" title="James E. Darnell">James E. Darnell</a></dd> <dd><a href="/wiki/Evelyn_M._Witkin" title="Evelyn M. Witkin">Evelyn M. Witkin</a></dd> <dt>2003</dt> <dd><a href="/wiki/J._Michael_Bishop" title="J. Michael Bishop">J. Michael Bishop</a></dd> <dd><a href="/wiki/Solomon_H._Snyder" title="Solomon H. Snyder">Solomon H. Snyder</a></dd> <dd><a href="/wiki/Charles_Yanofsky" title="Charles Yanofsky">Charles Yanofsky</a></dd> <dt>2004</dt> <dd><a href="/wiki/Norman_Borlaug" title="Norman Borlaug">Norman E. Borlaug</a></dd> <dd><a href="/wiki/Phillip_A._Sharp" class="mw-redirect" title="Phillip A. Sharp">Phillip A. Sharp</a></dd> <dd><a href="/wiki/Thomas_Starzl" title="Thomas Starzl">Thomas E. Starzl</a></dd> <dt>2005</dt> <dd><a href="/wiki/Anthony_Fauci" title="Anthony Fauci">Anthony Fauci</a></dd> <dd><a href="/wiki/Torsten_N._Wiesel" class="mw-redirect" title="Torsten N. Wiesel">Torsten N. Wiesel</a></dd> <dt>2006</dt> <dd><a href="/wiki/Rita_R._Colwell" title="Rita R. Colwell">Rita R. Colwell</a></dd> <dd><a href="/wiki/Nina_Fedoroff" title="Nina Fedoroff">Nina Fedoroff</a></dd> <dd><a href="/wiki/Lubert_Stryer" title="Lubert Stryer">Lubert Stryer</a></dd> <dt>2007</dt> <dd><a href="/wiki/Robert_Lefkowitz" title="Robert Lefkowitz">Robert J. Lefkowitz</a></dd> <dd><a href="/wiki/Bert_W._O%27Malley" title="Bert W. O'Malley">Bert W. O'Malley</a></dd> <dt>2008</dt> <dd><a href="/wiki/Francis_Collins" title="Francis Collins">Francis S. Collins</a></dd> <dd><a href="/wiki/Elaine_Fuchs" title="Elaine Fuchs">Elaine Fuchs</a></dd> <dd><a href="/wiki/Craig_Venter" title="Craig Venter">J. Craig Venter</a></dd> <dt>2009</dt> <dd><a href="/wiki/Susan_L._Lindquist" class="mw-redirect" title="Susan L. Lindquist">Susan L. Lindquist</a></dd> <dd><a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Stanley B. Prusiner</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2010s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>2010</dt> <dd><a href="/wiki/Ralph_L._Brinster" title="Ralph L. Brinster">Ralph L. Brinster</a></dd> <dd><a href="/wiki/Rudolf_Jaenisch" title="Rudolf Jaenisch">Rudolf Jaenisch</a></dd> <dt>2011</dt> <dd><a href="/wiki/Lucy_Shapiro" title="Lucy Shapiro">Lucy Shapiro</a></dd> <dd><a href="/wiki/Leroy_Hood" title="Leroy Hood">Leroy Hood</a></dd> <dd><a href="/wiki/Sallie_Chisholm" class="mw-redirect" title="Sallie Chisholm">Sallie Chisholm</a></dd> <dt>2012</dt> <dd><a href="/wiki/May_Berenbaum" title="May Berenbaum">May Berenbaum</a></dd> <dd><a href="/wiki/Bruce_Alberts" title="Bruce Alberts">Bruce Alberts</a></dd> <dt>2013</dt> <dd><a href="/wiki/Rakesh_K._Jain" class="mw-redirect" title="Rakesh K. Jain">Rakesh K. Jain</a></dd> <dt>2014</dt> <dd><a href="/wiki/Stanley_Falkow" title="Stanley Falkow">Stanley Falkow</a></dd> <dd><a href="/wiki/Mary-Claire_King" title="Mary-Claire King">Mary-Claire King</a></dd> <dd><a href="/wiki/Simon_Levin" class="mw-redirect" title="Simon Levin">Simon Levin</a></dd></dl> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Chemistry" style="font-size:114%;margin:0 4em">Chemistry</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">1960s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1964</dt> <dd><a href="/wiki/Roger_Adams" title="Roger Adams">Roger Adams</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1980s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1982</dt> <dd><a href="/wiki/F._Albert_Cotton" title="F. Albert Cotton">F. Albert Cotton</a></dd> <dd><a href="/wiki/Gilbert_Stork" title="Gilbert Stork">Gilbert Stork</a></dd> <dt>1983</dt> <dd><a href="/wiki/Roald_Hoffmann" title="Roald Hoffmann">Roald Hoffmann</a></dd> <dd><a href="/wiki/George_C._Pimentel" title="George C. Pimentel">George C. Pimentel</a></dd> <dd><a href="/wiki/Richard_N._Zare" class="mw-redirect" title="Richard N. Zare">Richard N. Zare</a></dd> <dt>1986</dt> <dd><a href="/wiki/Harry_B._Gray" title="Harry B. Gray">Harry B. Gray</a></dd> <dd><a href="/wiki/Yuan_Tseh_Lee" class="mw-redirect" title="Yuan Tseh Lee">Yuan Tseh Lee</a></dd> <dd><a href="/wiki/Carl_S._Marvel" class="mw-redirect" title="Carl S. Marvel">Carl S. Marvel</a></dd> <dd><a href="/wiki/Frank_H._Westheimer" class="mw-redirect" title="Frank H. Westheimer">Frank H. Westheimer</a></dd> <dt>1987</dt> <dd><a href="/wiki/William_Summer_Johnson" title="William Summer Johnson">William S. Johnson</a></dd> <dd><a href="/wiki/Walter_H._Stockmayer" title="Walter H. Stockmayer">Walter H. Stockmayer</a></dd> <dd><a href="/wiki/Max_Tishler" title="Max Tishler">Max Tishler</a></dd> <dt>1988</dt> <dd><a href="/wiki/William_O._Baker" title="William O. Baker">William O. Baker</a></dd> <dd><a href="/wiki/Konrad_E._Bloch" class="mw-redirect" title="Konrad E. Bloch">Konrad E. Bloch</a></dd> <dd><a href="/wiki/Elias_J._Corey" class="mw-redirect" title="Elias J. Corey">Elias J. Corey</a></dd> <dt>1989</dt> <dd><a href="/wiki/Richard_Barry_Bernstein" title="Richard Barry Bernstein">Richard B. Bernstein</a></dd> <dd><a href="/wiki/Melvin_Calvin" title="Melvin Calvin">Melvin Calvin</a></dd> <dd><a href="/wiki/Rudolph_A._Marcus" title="Rudolph A. Marcus">Rudolph A. Marcus</a></dd> <dd><a href="/wiki/Harden_M._McConnell" title="Harden M. McConnell">Harden M. McConnell</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1990s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1990</dt> <dd><a href="/wiki/Elkan_Blout" title="Elkan Blout">Elkan Blout</a></dd> <dd><a href="/wiki/Karl_August_Folkers" title="Karl August Folkers">Karl Folkers</a></dd> <dd><a href="/wiki/John_D._Roberts" title="John D. Roberts">John D. Roberts</a></dd> <dt>1991</dt> <dd><a href="/wiki/Ronald_Breslow" title="Ronald Breslow">Ronald Breslow</a></dd> <dd><a href="/wiki/Gertrude_B._Elion" title="Gertrude B. Elion">Gertrude B. Elion</a></dd> <dd><a href="/wiki/Dudley_R._Herschbach" title="Dudley R. Herschbach">Dudley R. Herschbach</a></dd> <dd><a href="/wiki/Glenn_T._Seaborg" title="Glenn T. Seaborg">Glenn T. Seaborg</a></dd> <dt>1992</dt> <dd><a href="/wiki/Howard_Ensign_Simmons_Jr." title="Howard Ensign Simmons Jr.">Howard E. Simmons Jr.</a></dd> <dt>1993</dt> <dd><a href="/wiki/Donald_J._Cram" title="Donald J. Cram">Donald J. Cram</a></dd> <dd><a href="/wiki/Norman_Hackerman" title="Norman Hackerman">Norman Hackerman</a></dd> <dt>1994</dt> <dd><a href="/wiki/George_S._Hammond" title="George S. Hammond">George S. Hammond</a></dd> <dt>1995</dt> <dd><a href="/wiki/Thomas_Cech" title="Thomas Cech">Thomas Cech</a></dd> <dd><a href="/wiki/Isabella_L._Karle" class="mw-redirect" title="Isabella L. Karle">Isabella L. Karle</a></dd> <dt>1996</dt> <dd><a href="/wiki/Norman_Davidson_(biologist)" title="Norman Davidson (biologist)">Norman Davidson</a></dd> <dt>1997</dt> <dd><a href="/wiki/Darleane_C._Hoffman" title="Darleane C. Hoffman">Darleane C. Hoffman</a></dd> <dd><a href="/wiki/Harold_S._Johnston" title="Harold S. Johnston">Harold S. Johnston</a></dd> <dt>1998</dt> <dd><a href="/wiki/John_W._Cahn" title="John W. Cahn">John W. Cahn</a></dd> <dd><a href="/wiki/George_M._Whitesides" title="George M. Whitesides">George M. Whitesides</a></dd> <dt>1999</dt> <dd><a href="/wiki/Stuart_A._Rice" title="Stuart A. Rice">Stuart A. Rice</a></dd> <dd><a href="/wiki/John_Ross_(chemist)" title="John Ross (chemist)">John Ross</a></dd> <dd><a href="/wiki/Susan_Solomon" title="Susan Solomon">Susan Solomon</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2000s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>2000</dt> <dd><a href="/wiki/John_D._Baldeschwieler" title="John D. Baldeschwieler">John D. Baldeschwieler</a></dd> <dd><a href="/wiki/Ralph_F._Hirschmann" title="Ralph F. Hirschmann">Ralph F. Hirschmann</a></dd> <dt>2001</dt> <dd><a href="/wiki/Ernest_R._Davidson" title="Ernest R. Davidson">Ernest R. Davidson</a></dd> <dd><a href="/wiki/G%C3%A1bor_A._Somorjai" title="Gábor A. Somorjai">Gábor A. Somorjai</a></dd> <dt>2002</dt> <dd><a href="/wiki/John_I._Brauman" class="mw-redirect" title="John I. Brauman">John I. Brauman</a></dd> <dt>2004</dt> <dd><a href="/wiki/Stephen_J._Lippard" title="Stephen J. Lippard">Stephen J. Lippard</a></dd> <dt>2005</dt> <dd><a href="/wiki/Tobin_J._Marks" title="Tobin J. Marks">Tobin J. Marks</a></dd> <dt>2006</dt> <dd><a href="/wiki/Marvin_H._Caruthers" title="Marvin H. Caruthers">Marvin H. Caruthers</a></dd> <dd><a href="/wiki/Peter_Dervan" title="Peter Dervan">Peter B. Dervan</a></dd> <dt>2007</dt> <dd><a href="/wiki/Mostafa_El-Sayed" title="Mostafa El-Sayed">Mostafa A. El-Sayed</a></dd> <dt>2008</dt> <dd><a href="/wiki/Joanna_Fowler" title="Joanna Fowler">Joanna Fowler</a></dd> <dd><a href="/wiki/JoAnne_Stubbe" title="JoAnne Stubbe">JoAnne Stubbe</a></dd> <dt>2009</dt> <dd><a href="/wiki/Stephen_J._Benkovic" title="Stephen J. Benkovic">Stephen J. Benkovic</a></dd> <dd><a href="/wiki/Marye_Anne_Fox" title="Marye Anne Fox">Marye Anne Fox</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2010s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>2010</dt> <dd><a href="/wiki/Jacqueline_K._Barton" class="mw-redirect" title="Jacqueline K. Barton">Jacqueline K. Barton</a></dd> <dd><a href="/wiki/Peter_J._Stang" title="Peter J. Stang">Peter J. Stang</a></dd> <dt>2011</dt> <dd><a href="/wiki/Allen_J._Bard" title="Allen J. Bard">Allen J. Bard</a></dd> <dd><a href="/wiki/M._Frederick_Hawthorne" title="M. Frederick Hawthorne">M. Frederick Hawthorne</a></dd> <dt>2012</dt> <dd><a href="/wiki/Judith_P._Klinman" class="mw-redirect" title="Judith P. Klinman">Judith P. Klinman</a></dd> <dd><a href="/wiki/Jerrold_Meinwald" title="Jerrold Meinwald">Jerrold Meinwald</a></dd> <dt>2013</dt> <dd><a href="/wiki/Geraldine_L._Richmond" title="Geraldine L. Richmond">Geraldine L. Richmond</a></dd> <dt>2014</dt> <dd><a href="/wiki/A._Paul_Alivisatos" class="mw-redirect" title="A. Paul Alivisatos">A. Paul Alivisatos</a></dd></dl> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Engineering_sciences" style="font-size:114%;margin:0 4em">Engineering sciences</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">1960s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1962</dt> <dd><a href="/wiki/Theodore_von_K%C3%A1rm%C3%A1n" title="Theodore von Kármán">Theodore von Kármán</a></dd> <dt>1963</dt> <dd><a href="/wiki/Vannevar_Bush" title="Vannevar Bush">Vannevar Bush</a></dd> <dd><a href="/wiki/John_Robinson_Pierce" class="mw-redirect" title="John Robinson Pierce">John Robinson Pierce</a></dd> <dt>1964</dt> <dd><a href="/wiki/Charles_S._Draper" class="mw-redirect" title="Charles S. Draper">Charles S. Draper</a></dd> <dd><a href="/wiki/Othmar_Ammann" title="Othmar Ammann">Othmar H. Ammann</a></dd> <dt>1965</dt> <dd><a href="/wiki/Hugh_L._Dryden" class="mw-redirect" title="Hugh L. Dryden">Hugh L. Dryden</a></dd> <dd><a href="/wiki/Clarence_L._Johnson" class="mw-redirect" title="Clarence L. Johnson">Clarence L. Johnson</a></dd> <dd><a href="/wiki/Warren_K._Lewis" title="Warren K. Lewis">Warren K. Lewis</a></dd> <dt>1966</dt> <dd><a href="/wiki/Claude_E._Shannon" class="mw-redirect" title="Claude E. Shannon">Claude E. Shannon</a></dd> <dt>1967</dt> <dd><a href="/wiki/Edwin_H._Land" title="Edwin H. Land">Edwin H. Land</a></dd> <dd><a href="/wiki/Igor_I._Sikorsky" class="mw-redirect" title="Igor I. Sikorsky">Igor I. Sikorsky</a></dd> <dt>1968</dt> <dd><a href="/wiki/J._Presper_Eckert" title="J. Presper Eckert">J. Presper Eckert</a></dd> <dd><a href="/wiki/Nathan_M._Newmark" title="Nathan M. Newmark">Nathan M. Newmark</a></dd> <dt>1969</dt> <dd><a href="/wiki/Jack_St._Clair_Kilby" class="mw-redirect" title="Jack St. Clair Kilby">Jack St. Clair Kilby</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1970s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1970</dt> <dd><a href="/wiki/George_E._Mueller" class="mw-redirect" title="George E. Mueller">George E. Mueller</a></dd> <dt>1973</dt> <dd><a href="/wiki/Harold_E._Edgerton" class="mw-redirect" title="Harold E. Edgerton">Harold E. Edgerton</a></dd> <dd><a href="/wiki/Richard_T._Whitcomb" title="Richard T. Whitcomb">Richard T. Whitcomb</a></dd> <dt>1974</dt> <dd><a href="/wiki/Rudolf_Kompfner" title="Rudolf Kompfner">Rudolf Kompfner</a></dd> <dd><a href="/wiki/Ralph_Brazelton_Peck" title="Ralph Brazelton Peck">Ralph Brazelton Peck</a></dd> <dd><a href="/wiki/Abel_Wolman" title="Abel Wolman">Abel Wolman</a></dd> <dt>1975</dt> <dd><a href="/wiki/Manson_Benedict" title="Manson Benedict">Manson Benedict</a></dd> <dd><a href="/wiki/William_Hayward_Pickering" class="mw-redirect" title="William Hayward Pickering">William Hayward Pickering</a></dd> <dd><a href="/wiki/Frederick_Terman" title="Frederick Terman">Frederick E. Terman</a></dd> <dd><a href="/wiki/Wernher_von_Braun" title="Wernher von Braun">Wernher von Braun</a></dd> <dt>1976</dt> <dd><a href="/wiki/Morris_Cohen_(scientist)" title="Morris Cohen (scientist)">Morris Cohen</a></dd> <dd><a href="/wiki/Peter_C._Goldmark" class="mw-redirect" title="Peter C. Goldmark">Peter C. Goldmark</a></dd> <dd><a href="/wiki/Erwin_Wilhelm_M%C3%BCller" title="Erwin Wilhelm Müller">Erwin Wilhelm Müller</a></dd> <dt>1979</dt> <dd><a href="/wiki/Emmett_N._Leith" class="mw-redirect" title="Emmett N. Leith">Emmett N. Leith</a></dd> <dd><a href="/wiki/Raymond_D._Mindlin" title="Raymond D. Mindlin">Raymond D. Mindlin</a></dd> <dd><a href="/wiki/Robert_N._Noyce" class="mw-redirect" title="Robert N. Noyce">Robert N. Noyce</a></dd> <dd><a href="/wiki/Earl_R._Parker" title="Earl R. Parker">Earl R. Parker</a></dd> <dd><a href="/wiki/Simon_Ramo" title="Simon Ramo">Simon Ramo</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1980s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1982</dt> <dd><a href="/wiki/Edward_H._Heinemann" class="mw-redirect" title="Edward H. Heinemann">Edward H. Heinemann</a></dd> <dd><a href="/wiki/Donald_L._Katz" title="Donald L. Katz">Donald L. Katz</a></dd> <dt>1983</dt> <dd><a href="/wiki/Bill_Hewlett" title="Bill Hewlett">Bill Hewlett</a></dd> <dd><a href="/wiki/George_Low" title="George Low">George Low</a></dd> <dd><a href="/wiki/John_G._Trump" title="John G. Trump">John G. Trump</a></dd> <dt>1986</dt> <dd><a href="/wiki/Hans_Wolfgang_Liepmann" class="mw-redirect" title="Hans Wolfgang Liepmann">Hans Wolfgang Liepmann</a></dd> <dd><a href="/wiki/Tung-Yen_Lin" title="Tung-Yen Lin">Tung-Yen Lin</a></dd> <dd><a href="/wiki/Bernard_M._Oliver" title="Bernard M. Oliver">Bernard M. Oliver</a></dd> <dt>1987</dt> <dd><a href="/wiki/Robert_Byron_Bird" title="Robert Byron Bird">Robert Byron Bird</a></dd> <dd><a href="/wiki/H._Bolton_Seed" class="mw-redirect" title="H. Bolton Seed">H. Bolton Seed</a></dd> <dd><a href="/wiki/Ernst_Weber_(engineer)" title="Ernst Weber (engineer)">Ernst Weber</a></dd> <dt>1988</dt> <dd><a href="/wiki/Daniel_C._Drucker" title="Daniel C. Drucker">Daniel C. Drucker</a></dd> <dd><a href="/wiki/Willis_M._Hawkins" class="mw-redirect" title="Willis M. Hawkins">Willis M. Hawkins</a></dd> <dd><a href="/wiki/George_W._Housner" title="George W. Housner">George W. Housner</a></dd> <dt>1989</dt> <dd><a href="/wiki/Harry_George_Drickamer" title="Harry George Drickamer">Harry George Drickamer</a></dd> <dd><a href="/wiki/Herbert_E._Grier" title="Herbert E. Grier">Herbert E. Grier</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1990s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1990</dt> <dd><a href="/wiki/Mildred_Dresselhaus" title="Mildred Dresselhaus">Mildred Dresselhaus</a></dd> <dd><a href="/wiki/Nick_Holonyak_Jr." class="mw-redirect" title="Nick Holonyak Jr.">Nick Holonyak Jr.</a></dd> <dt>1991</dt> <dd><a href="/wiki/George_H._Heilmeier" title="George H. Heilmeier">George H. Heilmeier</a></dd> <dd><a href="/wiki/Luna_B._Leopold" class="mw-redirect" title="Luna B. Leopold">Luna B. Leopold</a></dd> <dd><a href="/wiki/H._Guyford_Stever" class="mw-redirect" title="H. Guyford Stever">H. Guyford Stever</a></dd> <dt>1992</dt> <dd><a href="/wiki/Calvin_F._Quate" class="mw-redirect" title="Calvin F. Quate">Calvin F. Quate</a></dd> <dd><a href="/wiki/John_Roy_Whinnery" title="John Roy Whinnery">John Roy Whinnery</a></dd> <dt>1993</dt> <dd><a href="/wiki/Alfred_Y._Cho" title="Alfred Y. Cho">Alfred Y. Cho</a></dd> <dt>1994</dt> <dd><a href="/wiki/Ray_W._Clough" class="mw-redirect" title="Ray W. Clough">Ray W. Clough</a></dd> <dt>1995</dt> <dd><a href="/wiki/Hermann_A._Haus" title="Hermann A. Haus">Hermann A. Haus</a></dd> <dt>1996</dt> <dd><a href="/wiki/James_L._Flanagan" title="James L. Flanagan">James L. Flanagan</a></dd> <dd><a href="/wiki/C._Kumar_N._Patel" title="C. Kumar N. Patel">C. Kumar N. Patel</a></dd> <dt>1998</dt> <dd><a href="/wiki/Eli_Ruckenstein" title="Eli Ruckenstein">Eli Ruckenstein</a></dd> <dt>1999</dt> <dd><a href="/wiki/Kenneth_N._Stevens" title="Kenneth N. Stevens">Kenneth N. Stevens</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2000s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>2000</dt> <dd><a href="/wiki/Yuan-Cheng_Fung" title="Yuan-Cheng Fung">Yuan-Cheng B. Fung</a></dd> <dt>2001</dt> <dd><a href="/wiki/Andreas_Acrivos" title="Andreas Acrivos">Andreas Acrivos</a></dd> <dt>2002</dt> <dd><a href="/wiki/Leo_Beranek" title="Leo Beranek">Leo Beranek</a></dd> <dt>2003</dt> <dd><a href="/wiki/John_Prausnitz" title="John Prausnitz">John M. Prausnitz</a></dd> <dt>2004</dt> <dd><a href="/wiki/Edwin_N._Lightfoot" title="Edwin N. Lightfoot">Edwin N. Lightfoot</a></dd> <dt>2005</dt> <dd><a href="/wiki/Jan_D._Achenbach" title="Jan D. Achenbach">Jan D. Achenbach</a></dd> <dt>2006</dt> <dd><a href="/wiki/Robert_S._Langer" title="Robert S. Langer">Robert S. Langer</a></dd> <dt>2007</dt> <dd><a href="/wiki/David_J._Wineland" title="David J. Wineland">David J. Wineland</a></dd> <dt>2008</dt> <dd><a href="/wiki/Rudolf_E._K%C3%A1lm%C3%A1n" title="Rudolf E. Kálmán">Rudolf E. Kálmán</a></dd> <dt>2009</dt> <dd><a href="/wiki/Amnon_Yariv" title="Amnon Yariv">Amnon Yariv</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2010s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>2010</dt> <dd><a href="/wiki/Shu_Chien" title="Shu Chien">Shu Chien</a></dd> <dt>2011</dt> <dd><a href="/wiki/John_B._Goodenough" title="John B. Goodenough">John B. Goodenough</a></dd> <dt>2012</dt> <dd><a href="/wiki/Thomas_Kailath" title="Thomas Kailath">Thomas Kailath</a></dd></dl> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Mathematical,_statistical,_and_computer_sciences" style="font-size:114%;margin:0 4em">Mathematical, statistical, and computer sciences</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">1960s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1963</dt> <dd><a href="/wiki/Norbert_Wiener" title="Norbert Wiener">Norbert Wiener</a></dd> <dt>1964</dt> <dd><a href="/wiki/Solomon_Lefschetz" title="Solomon Lefschetz">Solomon Lefschetz</a></dd> <dd><a href="/wiki/Marston_Morse" title="Marston Morse">H. Marston Morse</a></dd> <dt>1965</dt> <dd><a href="/wiki/Oscar_Zariski" title="Oscar Zariski">Oscar Zariski</a></dd> <dt>1966</dt> <dd><a href="/wiki/John_Milnor" title="John Milnor">John Milnor</a></dd> <dt>1967</dt> <dd><a href="/wiki/Paul_Cohen" title="Paul Cohen">Paul Cohen</a></dd> <dt>1968</dt> <dd><a href="/wiki/Jerzy_Neyman" title="Jerzy Neyman">Jerzy Neyman</a></dd> <dt>1969</dt> <dd><a href="/wiki/William_Feller" title="William Feller">William Feller</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1970s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1970</dt> <dd><a href="/wiki/Richard_Brauer" title="Richard Brauer">Richard Brauer</a></dd> <dt>1973</dt> <dd><a href="/wiki/John_Tukey" title="John Tukey">John Tukey</a></dd> <dt>1974</dt> <dd><a href="/wiki/Kurt_G%C3%B6del" title="Kurt Gödel">Kurt Gödel</a></dd> <dt>1975</dt> <dd><a href="/wiki/John_W._Backus" class="mw-redirect" title="John W. Backus">John W. Backus</a></dd> <dd><a href="/wiki/Shiing-Shen_Chern" title="Shiing-Shen Chern">Shiing-Shen Chern</a></dd> <dd><a href="/wiki/George_Dantzig" title="George Dantzig">George Dantzig</a></dd> <dt>1976</dt> <dd><a href="/wiki/Kurt_O._Friedrichs" class="mw-redirect" title="Kurt O. Friedrichs">Kurt Otto Friedrichs</a></dd> <dd><a href="/wiki/Hassler_Whitney" title="Hassler Whitney">Hassler Whitney</a></dd> <dt>1979</dt> <dd><a href="/wiki/Joseph_L._Doob" title="Joseph L. Doob">Joseph L. Doob</a></dd> <dd><a href="/wiki/Donald_Knuth" title="Donald Knuth">Donald E. Knuth</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1980s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1982</dt> <dd><a href="/wiki/Marshall_H._Stone" title="Marshall H. Stone">Marshall H. Stone</a></dd> <dt>1983</dt> <dd><a href="/wiki/Herman_Goldstine" title="Herman Goldstine">Herman Goldstine</a></dd> <dd><a href="/wiki/Isadore_Singer" title="Isadore Singer">Isadore Singer</a></dd> <dt>1986</dt> <dd><a href="/wiki/Peter_Lax" title="Peter Lax">Peter Lax</a></dd> <dd><a href="/wiki/Antoni_Zygmund" title="Antoni Zygmund">Antoni Zygmund</a></dd> <dt>1987</dt> <dd><a href="/wiki/Raoul_Bott" title="Raoul Bott">Raoul Bott</a></dd> <dd><a href="/wiki/Michael_Freedman" title="Michael Freedman">Michael Freedman</a></dd> <dt>1988</dt> <dd><a href="/wiki/Ralph_E._Gomory" title="Ralph E. Gomory">Ralph E. Gomory</a></dd> <dd><a href="/wiki/Joseph_B._Keller" class="mw-redirect" title="Joseph B. Keller">Joseph B. Keller</a></dd> <dt>1989</dt> <dd><a href="/wiki/Samuel_Karlin" title="Samuel Karlin">Samuel Karlin</a></dd> <dd><a href="/wiki/Saunders_Mac_Lane" title="Saunders Mac Lane">Saunders Mac Lane</a></dd> <dd><a href="/wiki/Donald_C._Spencer" title="Donald C. Spencer">Donald C. Spencer</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1990s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1990</dt> <dd><a href="/wiki/George_F._Carrier" title="George F. Carrier">George F. Carrier</a></dd> <dd><a href="/wiki/Stephen_Cole_Kleene" title="Stephen Cole Kleene">Stephen Cole Kleene</a></dd> <dd><a href="/wiki/John_McCarthy_(computer_scientist)" title="John McCarthy (computer scientist)">John McCarthy</a></dd> <dt>1991</dt> <dd><a href="/wiki/Alberto_Calder%C3%B3n" title="Alberto Calderón">Alberto Calderón</a></dd> <dt>1992</dt> <dd><a href="/wiki/Allen_Newell" title="Allen Newell">Allen Newell</a></dd> <dt>1993</dt> <dd><a href="/wiki/Martin_David_Kruskal" title="Martin David Kruskal">Martin David Kruskal</a></dd> <dt>1994</dt> <dd><a href="/wiki/John_Cocke_(computer_scientist)" title="John Cocke (computer scientist)">John Cocke</a></dd> <dt>1995</dt> <dd><a href="/wiki/Louis_Nirenberg" title="Louis Nirenberg">Louis Nirenberg</a></dd> <dt>1996</dt> <dd><a href="/wiki/Richard_Karp" class="mw-redirect" title="Richard Karp">Richard Karp</a></dd> <dd><a href="/wiki/Stephen_Smale" title="Stephen Smale">Stephen Smale</a></dd> <dt>1997</dt> <dd><a href="/wiki/Shing-Tung_Yau" title="Shing-Tung Yau">Shing-Tung Yau</a></dd> <dt>1998</dt> <dd><a href="/wiki/Cathleen_Synge_Morawetz" title="Cathleen Synge Morawetz">Cathleen Synge Morawetz</a></dd> <dt>1999</dt> <dd><a href="/wiki/Felix_Browder" title="Felix Browder">Felix Browder</a></dd> <dd><a href="/wiki/Ronald_Coifman" title="Ronald Coifman">Ronald R. Coifman</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2000s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>2000</dt> <dd><a href="/wiki/John_Griggs_Thompson" class="mw-redirect" title="John Griggs Thompson">John Griggs Thompson</a></dd> <dd><a href="/wiki/Karen_Uhlenbeck" title="Karen Uhlenbeck">Karen Uhlenbeck</a></dd> <dt>2001</dt> <dd><a href="/wiki/Calyampudi_Radhakrishna_Rao" class="mw-redirect" title="Calyampudi Radhakrishna Rao">Calyampudi R. Rao</a></dd> <dd><a href="/wiki/Elias_M._Stein" title="Elias M. Stein">Elias M. Stein</a></dd> <dt>2002</dt> <dd><a href="/wiki/James_Glimm" title="James Glimm">James G. Glimm</a></dd> <dt>2003</dt> <dd><a href="/wiki/Carl_R._de_Boor" title="Carl R. de Boor">Carl R. de Boor</a></dd> <dt>2004</dt> <dd><a href="/wiki/Dennis_Sullivan" title="Dennis Sullivan">Dennis P. Sullivan</a></dd> <dt>2005</dt> <dd><a href="/wiki/Bradley_Efron" title="Bradley Efron">Bradley Efron</a></dd> <dt>2006</dt> <dd><a href="/wiki/Hyman_Bass" title="Hyman Bass">Hyman Bass</a></dd> <dt>2007</dt> <dd><a href="/wiki/Leonard_Kleinrock" title="Leonard Kleinrock">Leonard Kleinrock</a></dd> <dd><a href="/wiki/Andrew_Viterbi" title="Andrew Viterbi">Andrew J. Viterbi</a></dd> <dt>2009</dt> <dd><a href="/wiki/David_B._Mumford" class="mw-redirect" title="David B. Mumford">David B. Mumford</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2010s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>2010</dt> <dd><a href="/wiki/Richard_A._Tapia" title="Richard A. Tapia">Richard A. Tapia</a></dd> <dd><a href="/wiki/S._R._Srinivasa_Varadhan" title="S. R. Srinivasa Varadhan">S. R. Srinivasa Varadhan</a></dd> <dt>2011</dt> <dd><a href="/wiki/Solomon_W._Golomb" title="Solomon W. Golomb">Solomon W. Golomb</a></dd> <dd><a href="/wiki/Barry_Mazur" title="Barry Mazur">Barry Mazur</a></dd> <dt>2012</dt> <dd><a href="/wiki/Alexandre_Chorin" title="Alexandre Chorin">Alexandre Chorin</a></dd> <dd><a href="/wiki/David_Blackwell" title="David Blackwell">David Blackwell</a></dd> <dt>2013</dt> <dd><a href="/wiki/Michael_Artin" title="Michael Artin">Michael Artin</a></dd></dl> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Physical_sciences" style="font-size:114%;margin:0 4em">Physical sciences</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">1960s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1963</dt> <dd><a href="/wiki/Luis_Walter_Alvarez" title="Luis Walter Alvarez">Luis W. Alvarez</a></dd> <dt>1964</dt> <dd><a href="/wiki/Julian_Schwinger" title="Julian Schwinger">Julian Schwinger</a></dd> <dd><a href="/wiki/Harold_Urey" title="Harold Urey">Harold Urey</a></dd> <dd><a href="/wiki/Robert_Burns_Woodward" title="Robert Burns Woodward">Robert Burns Woodward</a></dd> <dt>1965</dt> <dd><a href="/wiki/John_Bardeen" title="John Bardeen">John Bardeen</a></dd> <dd><a href="/wiki/Peter_Debye" title="Peter Debye">Peter Debye</a></dd> <dd><a href="/wiki/Leon_M._Lederman" title="Leon M. Lederman">Leon M. Lederman</a></dd> <dd><a href="/wiki/William_Rubey" class="mw-redirect" title="William Rubey">William Rubey</a></dd> <dt>1966</dt> <dd><a href="/wiki/Jacob_Bjerknes" title="Jacob Bjerknes">Jacob Bjerknes</a></dd> <dd><a href="/wiki/Subrahmanyan_Chandrasekhar" title="Subrahmanyan Chandrasekhar">Subrahmanyan Chandrasekhar</a></dd> <dd><a href="/wiki/Henry_Eyring_(chemist)" title="Henry Eyring (chemist)">Henry Eyring</a></dd> <dd><a href="/wiki/John_H._Van_Vleck" class="mw-redirect" title="John H. Van Vleck">John H. Van Vleck</a></dd> <dd><a href="/wiki/Vladimir_K._Zworykin" title="Vladimir K. Zworykin">Vladimir K. Zworykin</a></dd> <dt>1967</dt> <dd><a href="/wiki/Jesse_Beams" title="Jesse Beams">Jesse Beams</a></dd> <dd><a href="/wiki/Francis_Birch_(geophysicist)" title="Francis Birch (geophysicist)">Francis Birch</a></dd> <dd><a href="/wiki/Gregory_Breit" title="Gregory Breit">Gregory Breit</a></dd> <dd><a href="/wiki/Louis_Hammett" class="mw-redirect" title="Louis Hammett">Louis Hammett</a></dd> <dd><a href="/wiki/George_Kistiakowsky" title="George Kistiakowsky">George Kistiakowsky</a></dd> <dt>1968</dt> <dd><a href="/wiki/Paul_Doughty_Bartlett" title="Paul Doughty Bartlett">Paul Bartlett</a></dd> <dd><a href="/wiki/Herbert_Friedman" title="Herbert Friedman">Herbert Friedman</a></dd> <dd><a href="/wiki/Lars_Onsager" title="Lars Onsager">Lars Onsager</a></dd> <dd><a href="/wiki/Eugene_Wigner" title="Eugene Wigner">Eugene Wigner</a></dd> <dt>1969</dt> <dd><a href="/wiki/Herbert_C._Brown" title="Herbert C. Brown">Herbert C. Brown</a></dd> <dd><a href="/wiki/Wolfgang_Panofsky" class="mw-redirect" title="Wolfgang Panofsky">Wolfgang Panofsky</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1970s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1970</dt> <dd><a href="/wiki/Robert_H._Dicke" title="Robert H. Dicke">Robert H. Dicke</a></dd> <dd><a href="/wiki/Allan_R._Sandage" class="mw-redirect" title="Allan R. Sandage">Allan R. Sandage</a></dd> <dd><a href="/wiki/John_C._Slater" title="John C. Slater">John C. Slater</a></dd> <dd><a href="/wiki/John_A._Wheeler" class="mw-redirect" title="John A. Wheeler">John A. Wheeler</a></dd> <dd><a href="/wiki/Saul_Winstein" title="Saul Winstein">Saul Winstein</a></dd> <dt>1973</dt> <dd><a href="/wiki/Carl_Djerassi" title="Carl Djerassi">Carl Djerassi</a></dd> <dd><a href="/wiki/Maurice_Ewing" title="Maurice Ewing">Maurice Ewing</a></dd> <dd><a href="/wiki/Arie_Jan_Haagen-Smit" title="Arie Jan Haagen-Smit">Arie Jan Haagen-Smit</a></dd> <dd><a href="/wiki/Vladimir_Haensel" title="Vladimir Haensel">Vladimir Haensel</a></dd> <dd><a href="/wiki/Frederick_Seitz" title="Frederick Seitz">Frederick Seitz</a></dd> <dd><a href="/wiki/Robert_Rathbun_Wilson" class="mw-redirect" title="Robert Rathbun Wilson">Robert Rathbun Wilson</a></dd> <dt>1974</dt> <dd><a href="/wiki/Nicolaas_Bloembergen" title="Nicolaas Bloembergen">Nicolaas Bloembergen</a></dd> <dd><a href="/wiki/Paul_Flory" title="Paul Flory">Paul Flory</a></dd> <dd><a href="/wiki/William_Alfred_Fowler" title="William Alfred Fowler">William Alfred Fowler</a></dd> <dd><a href="/wiki/Linus_Carl_Pauling" class="mw-redirect" title="Linus Carl Pauling">Linus Carl Pauling</a></dd> <dd><a href="/wiki/Kenneth_Sanborn_Pitzer" class="mw-redirect" title="Kenneth Sanborn Pitzer">Kenneth Sanborn Pitzer</a></dd> <dt>1975</dt> <dd><a href="/wiki/Hans_A._Bethe" class="mw-redirect" title="Hans A. Bethe">Hans A. Bethe</a></dd> <dd><a href="/wiki/Joseph_O._Hirschfelder" title="Joseph O. Hirschfelder">Joseph O. Hirschfelder</a></dd> <dd><a href="/wiki/Lewis_Sarett" class="mw-redirect" title="Lewis Sarett">Lewis Sarett</a></dd> <dd><a href="/wiki/Edgar_Bright_Wilson" title="Edgar Bright Wilson">Edgar Bright Wilson</a></dd> <dd><a href="/wiki/Chien-Shiung_Wu" title="Chien-Shiung Wu">Chien-Shiung Wu</a></dd> <dt>1976</dt> <dd><a href="/wiki/Samuel_Goudsmit" title="Samuel Goudsmit">Samuel Goudsmit</a></dd> <dd><a href="/wiki/Herbert_S._Gutowsky" title="Herbert S. Gutowsky">Herbert S. Gutowsky</a></dd> <dd><a href="/wiki/Frederick_Rossini" title="Frederick Rossini">Frederick Rossini</a></dd> <dd><a href="/wiki/Verner_Suomi" class="mw-redirect" title="Verner Suomi">Verner Suomi</a></dd> <dd><a href="/wiki/Henry_Taube" title="Henry Taube">Henry Taube</a></dd> <dd><a href="/wiki/George_Uhlenbeck" title="George Uhlenbeck">George Uhlenbeck</a></dd> <dt>1979</dt> <dd><a href="/wiki/Richard_P._Feynman" class="mw-redirect" title="Richard P. Feynman">Richard P. Feynman</a></dd> <dd><a href="/wiki/Herman_Mark" class="mw-redirect" title="Herman Mark">Herman Mark</a></dd> <dd><a href="/wiki/Edward_M._Purcell" class="mw-redirect" title="Edward M. Purcell">Edward M. Purcell</a></dd> <dd><a href="/wiki/John_Sinfelt" class="mw-redirect" title="John Sinfelt">John Sinfelt</a></dd> <dd><a href="/wiki/Lyman_Spitzer" title="Lyman Spitzer">Lyman Spitzer</a></dd> <dd><a href="/wiki/Victor_F._Weisskopf" class="mw-redirect" title="Victor F. Weisskopf">Victor F. Weisskopf</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1980s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1982</dt> <dd><a href="/wiki/Philip_Warren_Anderson" class="mw-redirect" title="Philip Warren Anderson">Philip W. Anderson</a></dd> <dd><a href="/wiki/Yoichiro_Nambu" title="Yoichiro Nambu">Yoichiro Nambu</a></dd> <dd><a href="/wiki/Edward_Teller" title="Edward Teller">Edward Teller</a></dd> <dd><a href="/wiki/Charles_H._Townes" title="Charles H. Townes">Charles H. Townes</a></dd> <dt>1983</dt> <dd><a href="/wiki/E._Margaret_Burbidge" class="mw-redirect" title="E. Margaret Burbidge">E. Margaret Burbidge</a></dd> <dd><a href="/wiki/Maurice_Goldhaber" title="Maurice Goldhaber">Maurice Goldhaber</a></dd> <dd><a href="/wiki/Helmut_Landsberg" title="Helmut Landsberg">Helmut Landsberg</a></dd> <dd><a href="/wiki/Walter_Munk" title="Walter Munk">Walter Munk</a></dd> <dd><a href="/wiki/Frederick_Reines" title="Frederick Reines">Frederick Reines</a></dd> <dd><a href="/wiki/Bruno_B._Rossi" class="mw-redirect" title="Bruno B. Rossi">Bruno B. Rossi</a></dd> <dd><a href="/wiki/J._Robert_Schrieffer" class="mw-redirect" title="J. Robert Schrieffer">J. Robert Schrieffer</a></dd> <dt>1986</dt> <dd><a href="/wiki/Solomon_J._Buchsbaum" title="Solomon J. Buchsbaum">Solomon J. Buchsbaum</a></dd> <dd><a href="/wiki/H._Richard_Crane" title="H. Richard Crane">H. Richard Crane</a></dd> <dd><a href="/wiki/Herman_Feshbach" title="Herman Feshbach">Herman Feshbach</a></dd> <dd><a href="/wiki/Robert_Hofstadter" title="Robert Hofstadter">Robert Hofstadter</a></dd> <dd><a href="/wiki/Yang_Chen-Ning" title="Yang Chen-Ning">Chen-Ning Yang</a></dd> <dt>1987</dt> <dd><a href="/wiki/Philip_Abelson" title="Philip Abelson">Philip Abelson</a></dd> <dd><a href="/wiki/Walter_Elsasser" class="mw-redirect" title="Walter Elsasser">Walter Elsasser</a></dd> <dd><a href="/wiki/Paul_C._Lauterbur" class="mw-redirect" title="Paul C. Lauterbur">Paul C. Lauterbur</a></dd> <dd><a href="/wiki/George_Pake" title="George Pake">George Pake</a></dd> <dd><a href="/wiki/James_A._Van_Allen" class="mw-redirect" title="James A. Van Allen">James A. Van Allen</a></dd> <dt>1988</dt> <dd><a href="/wiki/D._Allan_Bromley" title="D. Allan Bromley">D. Allan Bromley</a></dd> <dd><a href="/wiki/Chu_Ching-wu" class="mw-redirect" title="Chu Ching-wu">Paul Ching-Wu Chu</a></dd> <dd><a href="/wiki/Walter_Kohn" title="Walter Kohn">Walter Kohn</a></dd> <dd><a href="/wiki/Norman_Foster_Ramsey_Jr." class="mw-redirect" title="Norman Foster Ramsey Jr.">Norman Foster Ramsey Jr.</a></dd> <dd><a href="/wiki/Jack_Steinberger" title="Jack Steinberger">Jack Steinberger</a></dd> <dt>1989</dt> <dd><a href="/wiki/Arnold_O._Beckman" class="mw-redirect" title="Arnold O. Beckman">Arnold O. Beckman</a></dd> <dd><a href="/wiki/Eugene_Parker" title="Eugene Parker">Eugene Parker</a></dd> <dd><a href="/wiki/Robert_P._Sharp" title="Robert P. Sharp">Robert Sharp</a></dd> <dd><a href="/wiki/Henry_Stommel" title="Henry Stommel">Henry Stommel</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">1990s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>1990</dt> <dd><a href="/wiki/Allan_McLeod_Cormack" class="mw-redirect" title="Allan McLeod Cormack">Allan M. Cormack</a></dd> <dd><a href="/wiki/Edwin_M._McMillan" class="mw-redirect" title="Edwin M. McMillan">Edwin M. McMillan</a></dd> <dd><a href="/wiki/Robert_Pound" title="Robert Pound">Robert Pound</a></dd> <dd><a href="/wiki/Roger_Revelle" title="Roger Revelle">Roger Revelle</a></dd> <dt>1991</dt> <dd><a href="/wiki/Arthur_L._Schawlow" class="mw-redirect" title="Arthur L. Schawlow">Arthur L. Schawlow</a></dd> <dd><a href="/wiki/Edward_C._Stone" title="Edward C. Stone">Ed Stone</a></dd> <dd><a href="/wiki/Steven_Weinberg" title="Steven Weinberg">Steven Weinberg</a></dd> <dt>1992</dt> <dd><a href="/wiki/Eugene_M._Shoemaker" class="mw-redirect" title="Eugene M. Shoemaker">Eugene M. Shoemaker</a></dd> <dt>1993</dt> <dd><a href="/wiki/Val_Logsdon_Fitch" title="Val Logsdon Fitch">Val Fitch</a></dd> <dd><a href="/wiki/Vera_Rubin" title="Vera Rubin">Vera Rubin</a></dd> <dt>1994</dt> <dd><a href="/wiki/Albert_Overhauser" title="Albert Overhauser">Albert Overhauser</a></dd> <dd><a href="/wiki/Frank_Press" title="Frank Press">Frank Press</a></dd> <dt>1995</dt> <dd><a href="/wiki/Hans_Dehmelt" class="mw-redirect" title="Hans Dehmelt">Hans Dehmelt</a></dd> <dd><a href="/wiki/Peter_Goldreich" title="Peter Goldreich">Peter Goldreich</a></dd> <dt>1996</dt> <dd><a href="/wiki/Wallace_S._Broecker" class="mw-redirect" title="Wallace S. Broecker">Wallace S. Broecker</a></dd> <dt>1997</dt> <dd><a href="/wiki/Marshall_Rosenbluth" title="Marshall Rosenbluth">Marshall Rosenbluth</a></dd> <dd><a href="/wiki/Martin_Schwarzschild" title="Martin Schwarzschild">Martin Schwarzschild</a></dd> <dd><a href="/wiki/George_Wetherill" title="George Wetherill">George Wetherill</a></dd> <dt>1998</dt> <dd><a href="/wiki/Don_L._Anderson" title="Don L. Anderson">Don L. Anderson</a></dd> <dd><a href="/wiki/John_N._Bahcall" title="John N. Bahcall">John N. Bahcall</a></dd> <dt>1999</dt> <dd><a href="/wiki/James_Cronin" title="James Cronin">James Cronin</a></dd> <dd><a href="/wiki/Leo_Kadanoff" title="Leo Kadanoff">Leo Kadanoff</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2000s</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>2000</dt> <dd><a href="/wiki/Willis_E._Lamb" class="mw-redirect" title="Willis E. Lamb">Willis E. Lamb</a></dd> <dd><a href="/wiki/Jeremiah_P._Ostriker" title="Jeremiah P. Ostriker">Jeremiah P. Ostriker</a></dd> <dd><a href="/wiki/Gilbert_F._White" title="Gilbert F. White">Gilbert F. White</a></dd> <dt>2001</dt> <dd><a href="/wiki/Marvin_L._Cohen" title="Marvin L. Cohen">Marvin L. Cohen</a></dd> <dd><a href="/wiki/Raymond_Davis_Jr." title="Raymond Davis Jr.">Raymond Davis Jr.</a></dd> <dd><a href="/wiki/Charles_Keeling" class="mw-redirect" title="Charles Keeling">Charles Keeling</a></dd> <dt>2002</dt> <dd><a href="/wiki/Richard_Garwin" title="Richard Garwin">Richard Garwin</a></dd> <dd><a href="/wiki/W._Jason_Morgan" title="W. Jason Morgan">W. Jason Morgan</a></dd> <dd><a href="/wiki/Edward_Witten" title="Edward Witten">Edward Witten</a></dd> <dt>2003</dt> <dd><a href="/wiki/G._Brent_Dalrymple" class="mw-redirect" title="G. Brent Dalrymple">G. Brent Dalrymple</a></dd> <dd><a href="/wiki/Riccardo_Giacconi" title="Riccardo Giacconi">Riccardo Giacconi</a></dd> <dt>2004</dt> <dd><a href="/wiki/Robert_N._Clayton" title="Robert N. Clayton">Robert N. Clayton</a></dd> <dt>2005</dt> <dd><a href="/wiki/Ralph_A._Alpher" class="mw-redirect" title="Ralph A. Alpher">Ralph A. Alpher</a></dd> <dd><a href="/wiki/Lonnie_Thompson" title="Lonnie Thompson">Lonnie Thompson</a></dd> <dt>2006</dt> <dd><a href="/wiki/Daniel_Kleppner" title="Daniel Kleppner">Daniel Kleppner</a></dd> <dt>2007</dt> <dd><a href="/wiki/Fay_Ajzenberg-Selove" title="Fay Ajzenberg-Selove">Fay Ajzenberg-Selove</a></dd> <dd><a href="/wiki/Charles_Pence_Slichter" title="Charles Pence Slichter">Charles P. Slichter</a></dd> <dt>2008</dt> <dd><a href="/wiki/Berni_Alder" title="Berni Alder">Berni Alder</a></dd> <dd><a href="/wiki/James_E._Gunn_(astronomer)" class="mw-redirect" title="James E. Gunn (astronomer)">James E. Gunn</a></dd> <dt>2009</dt> <dd><a href="/wiki/Yakir_Aharonov" title="Yakir Aharonov">Yakir Aharonov</a></dd> <dd><a href="/wiki/Esther_M._Conwell" title="Esther M. Conwell">Esther M. Conwell</a></dd> <dd><a href="/wiki/Warren_M._Washington" title="Warren M. Washington">Warren M. Washington</a></dd></dl> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">2010s</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <dl><dt>2011</dt> <dd><a href="/wiki/Sidney_Drell" title="Sidney Drell">Sidney Drell</a></dd> <dd><a href="/wiki/Sandra_Faber" title="Sandra Faber">Sandra Faber</a></dd> <dd><a href="/wiki/Sylvester_James_Gates" title="Sylvester James Gates">Sylvester James Gates</a></dd> <dt>2012</dt> <dd><a href="/wiki/Burton_Richter" title="Burton Richter">Burton Richter</a></dd> <dd><a href="/wiki/Sean_Solomon" title="Sean Solomon">Sean C. Solomon</a></dd> <dt>2014</dt> <dd><a href="/wiki/Shirley_Ann_Jackson" title="Shirley Ann Jackson">Shirley Ann Jackson</a></dd></dl> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox authority-control" aria-labelledby="Authority_control_databases_frameless&#124;text-top&#124;10px&#124;alt=Edit_this_at_Wikidata&#124;link=https&#58;//www.wikidata.org/wiki/Q1975294#identifiers&#124;class=noprint&#124;Edit_this_at_Wikidata" style="padding:3px"><table class="nowraplinks hlist mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Authority_control_databases_frameless&#124;text-top&#124;10px&#124;alt=Edit_this_at_Wikidata&#124;link=https&#58;//www.wikidata.org/wiki/Q1975294#identifiers&#124;class=noprint&#124;Edit_this_at_Wikidata" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Authority_control" title="Help:Authority control">Authority control databases</a> <span class="mw-valign-text-top noprint" typeof="mw:File/Frameless"><a href="https://www.wikidata.org/wiki/Q1975294#identifiers" title="Edit this at Wikidata"><img alt="Edit this at Wikidata" src="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/10px-OOjs_UI_icon_edit-ltr-progressive.svg.png" decoding="async" width="10" height="10" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/8/8a/OOjs_UI_icon_edit-ltr-progressive.svg/15px-OOjs_UI_icon_edit-ltr-progressive.svg.png 1.5x, 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style="width:1%">National</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://d-nb.info/gnd/121508498">Germany</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/n98085907">United States</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://catalogue.bnf.fr/ark:/12148/cb15570644f">France</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://data.bnf.fr/ark:/12148/cb15570644f">BnF data</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://id.ndl.go.jp/auth/ndlna/00867221">Japan</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://aleph.nkp.cz/F/?func=find-c&local_base=aut&ccl_term=ica=jn20040505004&CON_LNG=ENG">Czech Republic</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="http://data.bibliotheken.nl/id/thes/p099674645">Netherlands</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="http://olduli.nli.org.il/F/?func=find-b&local_base=NLX10&find_code=UID&request=987007450315305171">Israel</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Academics</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.scopus.com/authid/detail.uri?authorId=7004332698">Scopus</a></span></li></ul></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"><ul><li><span class="uid"><a rel="nofollow" class="external text" href="https://www.idref.fr/094820791">IdRef</a></span></li><li><span class="uid"><a rel="nofollow" class="external text" href="https://snaccooperative.org/ark:/99166/w6pv75mf">SNAC</a></span></li></ul></div></td></tr></tbody></table></div>    </div><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?type=1x1" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Retrieved from "<a dir="ltr" href="https://en.wikipedia.org/w/index.php?title=Seymour_Benzer&oldid=1245492695">https://en.wikipedia.org/w/index.php?title=Seymour_Benzer&oldid=1245492695</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Help:Category" title="Help:Category">Categories</a>: <ul><li><a href="/wiki/Category:1921_births" title="Category:1921 births">1921 births</a></li><li><a href="/wiki/Category:2007_deaths" 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