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Its members, the '''malacostracans''', display a great diversity of body forms and include [[crab]]s, [[lobster]]s, [[crayfish]], [[Caridea|shrimp]], [[krill]], [[Dendrobranchiata|prawns]], [[woodlouse|woodlice]], [[Amphipoda|amphipods]], [[mantis shrimp]], [[Cymothoa_exigua|tongue-eating lice]] and many other less familiar animals. They are abundant in all marine environments and have colonised freshwater and terrestrial habitats. They are [[Segmentation (biology)|segmented]] animals, united by a common [[body plan]] comprising 20 body segments (rarely 21), and divided into a head, thorax, and abdomen. ==Etymology== The name Malacostraca is {{etymology|grc|''{{wikt-lang|grc|μαλακός}}'' ({{grc-transl|μαλακός}})|soft||''{{wikt-lang|grc|όστρακον}}'' ({{grc-transl|όστρακον}})|shell}}. The word was used by [[Aristotle's biology|Aristotle]], who contrasted them with [[oyster]]s, in comparison with which their shells are pliable.&lt;ref>{{OED|malacostracan}}&lt;/ref> It was applied to this taxon by French zoologist [[Pierre André Latreille]] in 1802. He was curator of the arthropod collection at the [[National Museum of Natural History (France)|National Museum of Natural History]] in Paris.&lt;ref>{{cite journal |author= Dupuis, Claude |year=1974 |title=Pierre André Latreille (1762–1833): the foremost entomologist of his time |journal=Annual Review of Entomology |volume=19 |pages=1–14 |doi=10.1146/annurev.en.19.010174.000245}}&lt;/ref> This scientific name is misleading, since the shell is soft only immediately after [[ecdysis|moulting]], and is usually hard.&lt;ref name="Rich">{{cite book |author=Rich, Patricia Vickers |author2=Fenton, Mildred Adams |author2-link=Mildred Adams Fenton |author3=Fenton, Caroll Lane |author3-link=Carroll Lane Fenton |author4=Rich, Thomas Hewitt |year=1996 |title=The Fossil Book: a Record of Prehistoric Life |edition=2nd |publisher=[[Courier Dover Publications]] |isbn=978-0-486-29371-4 |chapter=Crustaceans |pages=[https://archive.org/details/fossilbookrecor00rich/page/213 213–221] |chapter-url=https://books.google.com/books?id=_ntSspji0LYC&amp;pg=PA217 |url-access=registration |url=https://archive.org/details/fossilbookrecor00rich/page/213 }}&lt;/ref> Malacostracans are sometimes contrasted with entomostracans, a name applied to all crustaceans outside the Malacostraca, and named after the obsolete taxon [[Entomostraca]].&lt;ref name="Alberta">{{cite book |author= Clifford, Hugh F. |year=1991 |title=Aquatic Invertebrates of Alberta: an Illustrated Guide |publisher=[[University of Alberta]] |isbn=978-0-88864-234-9 |chapter=Introduction to the Malacostraca |pages=173–175 |chapter-url=https://books.google.com/books?id=8UQ4jHev6voC&amp;pg=PA173}}&lt;/ref> ==Description== [[File:Nebalia bipes.jpg|thumb|right|[[Leptostraca]] such as ''[[Nebalia bipes]]'' retain the primitive condition of having seven abdominal segments.]] The class Malacostraca includes about 40,000 species,&lt;ref>{{cite book |year=2002 |series=Zoological catalogue of Australia |volume=19.2A |title=Crustacea: Malacostraca |publisher=[[CSIRO Publishing]] |isbn=978-0-643-06901-5 |chapter=Introduction |author=Poore, Hugh F. |pages=1–7 |chapter-url=https://books.google.com/books?id=ww6RzBz42-4C&amp;pg=PA1}}&lt;/ref> and "arguably ... contains a greater diversity of body forms than any other class in the animal kingdom".&lt;ref name="Barnes">{{cite book |author1=Barnes, R. S. K. |author2=Calow, P. |author3=Olive, P. J. W. |author4=Golding, D. W. |author5=Spicer, J. I. |year=2001 |title=The Invertebrates: a Synthesis |edition=3rd |publisher=[[Wiley-Blackwell]] |isbn=978-0-632-04761-1 |chapter=Invertebrates with legs: the arthropods and similar groups |pages=168–206 |chapter-url=https://books.google.com/books?id=TBMsbe9efPgC&amp;pg=PA202}}&lt;/ref> Its members are characterised by the presence of three [[Tagma (arthropod anatomy)|tagmata]] (specialized groupings of multiple segments) – a five-segmented head, an eight-segmented [[thorax]] and an abdomen with six segments and a [[telson]], except in the [[Leptostraca]], which retain the [[plesiomorphy|ancestral condition]] of seven abdominal segments.&lt;ref name="Barnes"/> Malacostracans have abdominal appendages, a fact that differentiates them from all other major crustacean taxa except [[Remipedia]].&lt;ref name=Ruppert>{{cite book |title=Invertebrate Zoology |edition=7th |last1=Ruppert |first1=Edward E. |last2=Fox |first2=Richard, S. |last3=Barnes |first3=Robert D. |year=2004 |publisher=Cengage Learning |isbn=978-81-315-0104-7 |pages=625–626 }}&lt;/ref> Each body segment bears a pair of jointed [[appendage]]s, although these may be lost secondarily.&lt;ref name="ADW">{{cite web |url=http://animaldiversity.ummz.umich.edu/site/accounts/information/Malacostraca.html |author1=Atwater, Dan |author2=Fautin, Daphne G. |year=2001 |title=Class Malacostraca: crabs, krill, pill bugs, shrimp, and relatives |work=[[Animal Diversity Web]] |publisher=[[University of Michigan]] |access-date=November 23, 2010}}&lt;/ref> ===Tagmata=== The head bears two pairs of [[antenna (biology)|antennae]], the first of which is often [[Arthropod leg#Biramous and uniramous|biramous]] (branching into two parts) and the second pair bear [[exopod]]s (outer branches) which are often flattened into antennal scales known as [[scaphocerite]]s.&lt;ref name=Ruppert/> The mouthparts consist of pairs each of [[Mandible (arthropod mouthpart)|mandibles]], maxillules (second pair of mouthparts) and [[Maxilla (arthropod mouthpart)|maxillae]]. Except for [[Anostraca|fairy shrimps]], malacostracans are the only extant arthropods with [[compound eye]]s placed on moveable stalks,&lt;ref>{{cite book | url=https://books.google.com/books?id=x7vtCAAAQBAJ&amp;q=Anostraca+and+the+Malacostraca+&amp;pg=PA148 | title=Multicellular Animals: Volume II: The Phylogenetic System of the Metazoa | isbn=9783662103968 | last1=Ax | first1=Peter | date=9 March 2013 | publisher=Springer }}&lt;/ref>&lt;ref>{{cite journal | url=https://www.jstor.org/stable/1547926 | jstor=1547926 | title=Optical Design and Evolutionary Adaptation in Crustacean Compound Eyes | last1=Cronin | first1=Thomas W. | journal=Journal of Crustacean Biology | date=1986 | volume=6 | issue=1 | pages=1–23 | doi=10.2307/1547926 }}&lt;/ref> although in some taxa the eyes are unstalked, reduced or lost.&lt;ref name="Davie"/>&lt;ref name="Hayward">{{cite book |editor1=Hayward, P. J. |editor2=Ryland, John Stanley |year=1995 |title=Handbook of the Marine Fauna of North-West Europe |publisher=[[Oxford University Press]] |isbn=978-0-19-854055-7 |chapter=Crustaceans |pages=290–461 |author1=Hayward, P. J. |author2=Isaac, M. J. |author3=Makings, P. |author4=Moyse, J. |author5=Naylor, E. |author6=Smaldon, G. }}&lt;/ref> Up to three thoracic segments may be fused with the head to form a [[cephalothorax]]; the associated appendages turn forward and are modified as [[maxilliped]]s (accessory [[arthropod mouthparts|mouthparts]]).&lt;ref name=Ruppert/> A [[carapace]] may be absent, present or [[secondary loss|secondarily lost]], and may cover the head, part or all of the thorax and some of the abdomen.&lt;ref name="Barnes"/> It is variable in form and may be fused dorsally with some of the thoracic segments or occasionally be in two parts, hinged dorsally.&lt;ref name="Davie"/> Typically, each of the thoracic appendages is biramous and the endopods are the better developed of the branches, being used for crawling or grasping. Each endopod consist of seven articulating segments; the coxa, basis, ischium, merus, carpus, propodus and dactylus. In [[Decapoda|decapods]], the claw is formed by the articulation of the dactylus against an outgrowth of the propodus. In some taxa, the exopods are lost and the appendages are uniramous.&lt;ref name=Ruppert/> There is a clear demarcation between the thorax and the six or seven-segmented abdomen. In most taxa, each abdominal segment except the last carries a pair of biramous [[pleopod]]s used for swimming, burrowing, gas exchange, creating a current or brooding eggs. The first and second abdominal pleopods may be modified in the male to form [[gonopod]]s (accessory copulatory appendages).&lt;ref name=Ruppert/> The appendages of the last segment are typically flattened into [[uropod]]s, which together with the terminal telson, make up the "tail fan".&lt;ref name="Hayward"/> It is the sudden flexion of this tail fan that provides the thrust for the rapid escape response of these crustaceans and the tail fan is also used in steering.&lt;ref name=Ruppert/> In Leptostraca, the appendages on the telson instead form [[Caudal ramus|caudal rami]] (spine-like protrusions).&lt;ref>{{cite book |author1=Ricketts, Edward F. |author1-link=Ed Ricketts |author2=Calvin, Jack |author3=Phillips, David W. |author4=Hedgpeth, Joel W. |year=1992 |title=Between Pacific Tides |edition=5th |publisher=[[Stanford University Press]] |isbn=978-0-8047-2068-7 |chapter=Rocky shores of bays and estuaries |pages=[https://archive.org/details/betweenpacificti0000rick/page/269 269–316] |chapter-url=https://books.google.com/books?id=tUl5ESavtRIC&amp;pg=PA279 |title-link=Between Pacific Tides }}&lt;/ref> ===Internal anatomy=== The digestive tract is straight and the foregut consists of a short oesophagus and a two-chambered stomach, the first part of which contains a gizzard-like "gastric mill" for grinding food. The walls of this have [[chitin]]ous ridges, teeth and calcareous ossicles. The fine particles and soluble material are then moved into the midgut where chemical processing and absorption takes place in one or more pairs of large digestive caeca. The hindgut is concerned with water reclamation and the formation of faeces and the anus is situated at the base of the telson.&lt;ref name=IZ>{{cite book |title=Invertebrate Zoology |edition=7th |last1=Ruppert |first1=Edward E. |last2=Fox |first2=Richard, S. |last3=Barnes |first3=Robert D. |year=2004 |publisher=Cengage Learning |isbn=978-81-315-0104-7 |pages=610–613 }}&lt;/ref> Like other crustaceans, malacostracans have an [[open circulatory system]] in which the heart pumps blood into the [[Circulatory system#Open circulatory system|hemocoel]] (body cavity) where it supplies the needs of the organs for oxygen and nutrients before diffusing back to the heart.&lt;ref>{{cite web |url=http://www2.gsu.edu/~bioasx/closeopen.html |title=Closed and open circulatory system |author=Sakurai, Akira |publisher=Georgia State University |access-date=2014-05-21 |archive-date=2016-09-17 |archive-url=https://web.archive.org/web/20160917173108/http://www2.gsu.edu/~bioasx/closeopen.html |url-status=dead }}&lt;/ref> The typical [[respiratory pigment]] in malacostracans is [[haemocyanin]].&lt;ref>{{cite book |author=Urich, Klaus |year=1994 |title=Comparative Animal Biochemistry |publisher=[[Springer Science+Business Media|Springer]] |isbn=978-3-540-57420-0 |chapter=Respiratory pigments |pages=249–287 |chapter-url=https://books.google.com/books?id=GLbcWyeaCGQC&amp;pg=PA272}}&lt;/ref> Structures that function as kidneys are located near the base of the antennae. A brain exists in the form of ganglia close to the antennae, there are ganglia in each segment and a collection of major ganglia below the oesophagus.&lt;ref name="Encarta">{{cite book |title=Encarta |year=2005 |chapter=Crustacean |author= Ghiselin, Michael T. |author-link=Michael Ghiselin |publisher=[[Microsoft]]|title-link=Encarta }}&lt;/ref> Sensory organs include compound eyes (often stalked), [[Simple eye in invertebrates|ocelli]] (simple eyes), [[statocyst]]s and sensory [[seta|bristles]]. The naupliar eye is a characteristic of the [[Crustacean larvae#Nauplius|nauplius]] larva and consists of four cup-shaped ocelli facing in different directions and able to distinguish between light and darkness.&lt;ref name=IZ/> ==Ecology== [[File:Sally lightfoot crab.jpg|thumb|left|''[[Grapsus grapsus]]'', a terrestrial crab]] Malacostracans live in a wide range of [[marine (ocean)|marine]] and [[freshwater]] habitats, and three orders have [[terrestrial animal|terrestrial]] members: Amphipoda ([[Talitridae]]), Isopoda (Oniscidea, the [[woodlouse|woodlice]]) and Decapoda ([[Coenobitidae|terrestrial hermit crabs]], crabs of the families [[Ocypodidae]], [[Gecarcinidae]], and [[Grapsidae]], and terrestrial [[crayfish]]).&lt;ref>{{cite book |author=Little, Colin |year=1983 |title=The Colonisation of Land: Origins and Adaptations of Terrestrial Animals |publisher=[[Cambridge University Press]] |isbn=978-0-521-25218-8 |chapter=Crustaceans and the evolution of the arthropods |pages=63–106 |chapter-url=https://books.google.com/books?id=tfs8AAAAIAAJ&amp;pg=PA67}}&lt;/ref> They are abundant in all marine ecosystems, and most species are [[scavenger]]s, although some, such as the [[porcelain crab]]s, are [[filter feeder]]s, and some, such as mantis shrimps, are [[carnivore]]s.&lt;ref name="Hayward"/> ==Life cycle== Most species of malacostracans have distinct sexes (a phenomenon known as [[gonochorism]]), although a few species exhibit [[hermaphrodite|hermaphroditism]].&lt;ref name="Hayward"/> The female genital openings or [[gonopore]]s are located on the sixth thoracic segment or its appendages, while the male gonopores are on the eighth segment or its appendages, or in a small number of species, on the seventh.&lt;ref name="Davie">{{cite book |title=Crustacea: Malacostraca. Phyllocarida, Hoplocarida, Eucarida (Part 1) |series=Volume 19.3A of Zoological Catalogue of Australia |author=Davie, P. J. F. |year=2002 |isbn=978-0-643-06791-2 |publisher=[[CSIRO Publishing]] |chapter=Class Malacostraca. Introduction |page=23 |chapter-url=https://books.google.com/books?id=Sc7i6AL-GewC&amp;pg=PA23}}&lt;/ref> The naupliar larval stages are often reduced and take place before hatching, but where they occur, a [[metamorphosis]] usually occurs between the larval and the adult forms. Primitive malacostracans have a free-swimming naupliar larval stage.&lt;ref name="Davie"/> Research suggests the common ancestor of Malacostraca had lost the free-living nauplius larval stage, but re-evolved it again through [[heterochrony]] in Dendrobranchiata and Euphausiacea, which both has a lecithotrophic (non-feeding) nauplius stage.&lt;ref>{{cite journal | pmc=4429915 | year=2015 | last1=Jirikowski | first1=G. J. | last2=Wolff | first2=C. | last3=Richter | first3=S. | title=Evolution of eumalacostracan development—new insights into loss and reacquisition of larval stages revealed by heterochrony analysis | journal=Evodevo | volume=6 | page=4 | doi=10.1186/2041-9139-6-4 | pmid=25973168 | doi-access=free }}&lt;/ref>&lt;ref>{{cite journal | pmc=4684318 | year=2015 | last1=Akther | first1=H. | last2=Agersted | first2=M. D. | last3=Olesen | first3=J. | title=Naupliar and Metanaupliar Development of Thysanoessa raschii (Malacostraca, Euphausiacea) from Godthåbsfjord, Greenland, with a Reinstatement of the Ancestral Status of the Free-Living Nauplius in Malacostracan Evolution | journal=PLOS ONE | volume=10 | issue=12 | pages=e0141955 | doi=10.1371/journal.pone.0141955 | pmid=26682744 | bibcode=2015PLoSO..1041955A | doi-access=free }}&lt;/ref> ==Mating== Mating behavior has been studied in the freshwater shrimp ''Caridina ensifera''.&lt;ref name=Yue>{{cite journal |vauthors=Yue GH, Chang A |title=Molecular evidence for high frequency of multiple paternity in a freshwater shrimp species Caridina ensifera |journal=PLOS ONE |volume=5 |issue=9 |pages=e12721 |year=2010 |pmid=20856862 |pmc=2939052 |doi=10.1371/journal.pone.0012721 |bibcode=2010PLoSO...512721Y |doi-access=free }}&lt;/ref> Multiple paternity, common in the ''Malacostrica'', also occurs in ''C. ensifera''. Reproductive success of sires was found to correlate inversely with their genetic relatedness to the mother.&lt;ref name=Yue /> This finding suggests that sperm competition and/or pre- and post-copulatory female choice occurs. Female choice may increase the fitness of progeny by avoiding [[inbreeding]] that can lead to expression of [[Zygosity#Homozygous|homozygous]] deleterious [[Dominance (genetics)|recessive]] mutations.&lt;ref name="pmid19834483">{{cite journal |vauthors=Charlesworth D, Willis JH |title=The genetics of inbreeding depression |journal=Nat. Rev. Genet. |volume=10 |issue=11 |pages=783–96 |year=2009 |pmid=19834483 |doi=10.1038/nrg2664 |s2cid=771357 }}&lt;/ref> ==Phylogenetics== {{Main|Phylogeny of Malacostraca}} The [[monophyly]] of Malacostraca is widely accepted. This is supported by several common morphological traits which are present throughout the group and is confirmed by molecular studies.&lt;ref>{{cite journal |author=Hassanin, Alexandre |title=Phylogeny of Arthropoda inferred from mitochondrial sequences: Strategies for limiting the misleading effects of multiple changes in pattern and rates of substitution |year=2006 |journal=[[Molecular Phylogenetics and Evolution]] |volume=38 |issue=1 |pages=100–116 |doi=10.1016/j.ympev.2005.09.012|url=http://www.csulb.edu/~dlunderw/entomology/Arthropodphylogeny2006.pdf |pmid=16290034|bibcode=2006MolPE..38..100H }}&lt;/ref> However, a number of problems make it difficult to determine the relationships between the orders of Malacostraca. These include differences in [[mutation rate]]s in different [[Lineage (evolution)|lineages]], different patterns of evolution being apparent in different sources of data, including [[convergent evolution]], and [[long branch attraction]].&lt;ref>{{cite journal |author1=Jenner, Ronald A. |author2=Ní Dhubhghaill, Ciara |author3=Ferla, Matteo P. |author4=Wills, Matthew A. |year=2009 |title=Eumalacostracan phylogeny and total evidence: limitations of the usual suspects |journal=[[BMC Evolutionary Biology]] |volume=9 |issue=1 |page=21 |doi=10.1186/1471-2148-9-21 |url= |pmc=2640363 |pmid=19173741 |doi-access=free |bibcode=2009BMCEE...9...21J }}&lt;/ref> There is less agreement on the status of the subclass [[Phyllocarida]] with its single extant order, Leptostraca, depending on whether foliaceous (leaf-like) limbs have a single or multiple origin. Some authors advocate placing Phyllocarida in Phyllopoda, a group used in former classification systems, which would then include branchiopods, cephalocarids and leptostracans. A molecular study by American biologists Trisha Spears and Lawrence Abele concluded that phylogenetic evidence did not support the monophyly of this grouping, and that Phyllocarida should be regarded as a subclass of Malacostraca that had diverged from the main lineage at an early date.&lt;ref name="Davie"/>&lt;ref>{{cite journal |author1=Spears, Trisha |author2=Abele, Lawrence G. |year=1999 |title=Phylogenetic Relationships of Crustaceans with Foliaceous Limbs: An 18S rDNA Study of Branchiopoda, Cephalocarida, and Phyllocarida |journal=Journal of Crustacean Biology |volume=19 |issue=4 |pages=825–843 |jstor=1549304 |doi=10.1163/193724099x00538|doi-access=free }}&lt;/ref> The following cladogram is based on the 2001 [[phylogenetics|phylogenetic analysis]] of Richter &amp; Scholtz.&lt;ref>{{cite journal|vauthors=Richter, Scholtz|date=January 2002|title=Phylogenetic analysis of the Malacostraca (Crustacea)|journal=Journal of Zoological Systematics and Evolutionary Research|volume=39|issue=3|pages=113–136|doi=10.1046/j.1439-0469.2001.00164.x|doi-access=free}}&lt;/ref> {{clade |label1=Malacostraca |1={{clade |1=[[Phyllocarida]] |2={{clade |1=[[Hoplocarida]] |label2=[[Eumalacostraca]] |2={{clade |1={{clade |1=[[Decapoda]] |2=[[Amphionidacea]] }} |2={{clade |1={{clade |1=[[Euphausiacea]] |label2=[[Peracarida]] |2={{clade |1=[[Thermosbaenacea]] |2={{clade |1={{clade |1=[[Mysida]] |2=[[Lophogastrida]] }} |2={{clade |1={{clade |1=[[Spelaeogriphacea]] |2=[[Mictacea]] }} |2={{clade |1=[[Cumacea]] |2={{clade |1=[[Tanaidacea]] |2=[[Isopoda]] }} }} }} }} }} }} |label2=[[Syncarida]] |2={{clade |1=[[Anaspidacea]] |2=[[Bathynellacea]] }} }} }} }} }} }} ===Subclass Phyllocarida=== Leptostraca is the only extant order of Phyllocarida, the other two orders, [[Archaeostraca]] and [[Hoplostraca]] being extinct. Leptostracans are thought to be the most primitive of the malacostracans and date back to the Cambrian period. They range in length from {{convert|1|to|4|cm|1|abbr=on}}, most being suspension feeders though some are carnivores or scavengers. They have a two part carapace which encloses the head, the whole thorax and part of the abdomen and are the only malacostracans with seven abdominal segments. Three families are known with several genera and about twenty species. They are found worldwide from the intertidal zone to the deep ocean, all but one species being [[Benthos|benthic]] (living on the seabed).&lt;ref name=Ruppert/>&lt;ref name="Davie"/> ===Subclass Hoplocarida=== [[File:Squilla empusa.jpg|left|thumb|upright|''Squilla empusa'',&lt;br/> a mantis shrimp]] [[Stomatopoda]] is the only extant order of [[Hoplocarida]], the other two orders, Aeschronectida and Archaeostomatopoda being extinct. Stomatopodans, commonly known as mantis shrimps, range in length from {{convert|5|to|36|cm|0|abbr=on}} and are predators. They have a [[Anatomical terms of location#Dorsal and ventral|dorso-ventrally]] flattened body and a shield-like carapace and are armed with powerful, raptorial claws normally carried in a folded position. There are about 300 species, most living in tropical and subtropical seas although some live in temperate areas. They are benthic, mostly hiding in cracks and crevices or living in burrows, some emerging to forage while others are ambush predators.&lt;ref name=Ruppert/>&lt;ref name="Davie"/> ===Subclass Eumalacostraca=== The Eumalocostraca contains the vast majority of the approximately 40,000 living species of malacostracans and consists of three superorders, [[Syncarida]], [[Peracarida]] and [[Eucarida]]. Syncaridans are mostly small and found in freshwater and subterranean habitats. Peracaridans are characterised by having a [[Brood pouch (Peracarida)|marsupium]] in which they brood their young. They are found in marine, freshwater and terrestrial habitats and include [[Amphipoda]], [[Cumacea]], [[Isopoda]] and [[Mysida]]. Eucarida includes lobsters, crabs, shrimps, [[prawn]]s and krill.&lt;ref>{{cite book |title=Crustacea: Malacostraca. Phyllocarida, Hoplocarida, Eucarida (Part 1) |series=Volume 19.3A of Zoological Catalogue of Australia |author=Davie, P. J. F. |year=2002 |isbn=978-0-643-06791-2 |publisher=[[CSIRO Publishing]] |chapter=Class Malacostraca. Introduction |pages=91 |chapter-url=https://books.google.com/books?id=Sc7i6AL-GewC&amp;pg=PA23}}&lt;/ref> ===Fossil record=== The first malacostracans appeared sometime in the [[Cambrian]], when animals belonging to the Phyllocarida appeared.&lt;ref>{{cite journal |author1=Collette, Joseph H. |author2=Hagadorn, James W. |year=2010 |title=Three-dimensionally preserved arthropods from Cambrian Lagerstätten of Quebec and Wisconsin |journal=[[Journal of Paleontology]] |volume=84 |issue=4 |pages=646–667 |doi=10.1666/09-075.1|s2cid=130064618 }}&lt;/ref>&lt;ref>{{cite journal |author1=Collette, Joseph H. |author2=Hagadorn, James W. |year=2010 |title=Early evolution of phyllocarid arthropods: phylogeny and systematics of Cambrian–Devonian archaeostracans |journal=[[Journal of Paleontology]] |volume=84 |issue=5 |pages=795–820 |doi=10.1666/09-092.1|bibcode=2010JPal...84..795C |s2cid=85074218 }}&lt;/ref> ==Classification== {{See also|List of prehistoric malacostracans}} The following classification of living malacostracans is based on ''An Updated Classification of the Recent Crustacea'' (2001) by the American marine biologists [[Joel W. Martin]], curator of crustaceans at the [[Natural History Museum of Los Angeles County]], and George E. Davies.&lt;ref name="M&amp;D">{{cite book |url=http://atiniui.nhm.org/pdfs/3839/3839.pdf |title=An Updated Classification of the Recent Crustacea |author1=Martin, Joel W. |author2=Davis, George E. |year=2001 |pages=132 |publisher=[[Natural History Museum of Los Angeles County]] |access-date=2009-12-14 |archive-date=2011-07-11 |archive-url=https://web.archive.org/web/20110711181351/http://atiniui.nhm.org/pdfs/3839/3839.pdf |url-status=dead }}&lt;/ref> Extinct orders have been added to this&lt;ref name="Jenner">{{cite journal |journal=[[Contributions to Zoology]] |volume=67 |issue=3 |pages=155–186 |year=1998 |title=Palaeo- and archaeostomatopods (Hoplocarida: Crustacea) from the Bear Gulch Limestone, Mississippian (Namurian), of central Montana |author1=Jenner, Ronald A. |author2=Hof, Cees H. J. |author3=Schram, Frederick R. |doi=10.1163/18759866-06703001 |doi-access=free }}&lt;/ref>&lt;ref>{{cite book |author1=Camacho, A. I. |author2=Valdecasas, A. G. |year=2008 |title=Freshwater Animal Diversity Assessment |series=Developments in Hydrobiology |volume=198 |pages=257–266 |chapter=Global diversity of syncarids (Syncarida; Crustacea) in freshwater |doi=10.1007/978-1-4020-8259-7_28 |editor1=Balian, E. V. |editor2=Lévêque, C. |editor3=Segers, H. |editor4=Martens, K. |publisher=[[Springer Science+Business Media|Springer]]|isbn=978-1-4020-8258-0 }}&lt;/ref>&lt;ref>{{cite book |author=Davie, P. J. F. |year=2001 |title=Crustacea: Malacostraca: Phyllocarida, Hoplocarida, Eucarida (Part 1) |series=Zoological catalogue of Australia |volume=19.3A |publisher=[[CSIRO Publishing]] |isbn=978-0-643-06791-2 |chapter=Subclass: Phyllocarida, Introduction |page=24 |chapter-url=https://books.google.com/books?id=Sc7i6AL-GewC&amp;pg=PA24}}&lt;/ref> and are indicated by an [[Dagger (typography)|obelisk]] (†). [[File:Odontodactylus scyllarus3.jpg|thumb|''[[Peacock mantis shrimp|Odontodactylus scyllarus]]'' (Hoplocarida: [[mantis shrimp|Stomatopoda]])]] [[File:Porcellio scaber and Oniscus asellus - Zalné20070205.jpg|thumb|''[[Porcellio scaber]]'' and ''[[Oniscus asellus]]'' (Peracarida: [[Isopoda]])]] [[File:Cancer pagurus.jpg|thumb|''[[Cancer pagurus]]'' (Eucarida: [[Decapoda]])]] Class '''Malacostraca''' &lt;small>[[Pierre André Latreille|Latreille]], 1802&lt;/small> * Subclass [[Phyllocarida]] &lt;small>Packard, 1879&lt;/small> :* † [[Archaeostraca]] &lt;small>Claus 1888&lt;/small> :* † [[Hoplostraca]] &lt;small>Schram, 1973&lt;/small> :* [[Leptostraca]] &lt;small>Claus, 1880&lt;/small> * Subclass [[Hoplocarida]] &lt;small>Calman, 1904&lt;/small> :* † [[Aeschronectida]] &lt;small>Schram, 1969&lt;/small> :* † [[Archaeostomatopoda]] &lt;small>Schram, 1969&lt;/small> :* [[Stomatopoda]] &lt;small>Latreille, 1817&lt;/small> * Subclass [[Eumalacostraca]] &lt;small>Grobben, 1892&lt;/small> ** Superorder [[Syncarida]] &lt;small>Packard, 1885&lt;/small> *** † [[Palaeocaridacea]] &lt;small>Brooks, 1979&lt;/small> *** [[Bathynellacea]] &lt;small>Chappuis, 1915&lt;/small> *** [[Anaspidacea]] &lt;small>Calman, 1904&lt;/small> ** Superorder [[Peracarida]] &lt;small>Calman, 1904&lt;/small> *** [[Spelaeogriphacea]] &lt;small>Gordon, 1957&lt;/small> *** [[Thermosbaenacea]] &lt;small>Monod, 1927&lt;/small> *** [[Lophogastrida]] &lt;small>Sars, 1870&lt;/small> *** [[Mysida]] &lt;small>Haworth, 1825&lt;/small> *** [[Stygiomysida]] &lt;small>Tchindonova, 1981&lt;/small> *** [[Mictacea]] &lt;small>Bowman ''et al.'', 1985&lt;/small> *** [[Amphipoda]] &lt;small>Latreille, 1816&lt;/small> *** [[Isopoda]] &lt;small>Latreille, 1817&lt;/small> *** [[Tanaidacea]] &lt;small>Dana, 1849&lt;/small> *** [[Cumacea]] &lt;small>Krøyer, 1846&lt;/small> ** Superorder [[Eucarida]] &lt;small>Calman, 1904&lt;/small> *** † [[Angustidontida]] &lt;small>Gueriau, Charbonnier &amp; Clément, 2014&lt;/small>&lt;ref>{{Cite journal|last1=Gueriau|first1=Pierre|last2=Charbonnier|first2=Sylvain|last3=Clément|first3=Gaël|date=2014-09-01|title=Angustidontid crustaceans from the Late Devonian of Strud (Namur Province, Belgium): Insights into the origin of Decapoda|url=https://www.researchgate.net/publication/265211942|journal=Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen|volume=273|issue=3 |pages=327–337 |doi=10.1127/0077-7749/2014/0434}}&lt;/ref> *** [[Krill|Euphausiacea]] &lt;small>Dana, 1852&lt;/small> *** [[Decapoda]] &lt;small>Latreille, 1802&lt;/small> ==References== {{Reflist|32em}} ==External links== {{Portal|Crustaceans}} *{{Commons category-inline|Malacostraca}} *{{cite EB1911|first=Thomas Roscoe Rede |last=Stebbing |author-link=Thomas Roscoe Rede Stebbing |wstitle=Malacostraca |volume=17 |pages=457–459 |short=x}} *[http://tolweb.org/Malacostraca/6253 Malacostraca], [[Tree of Life Web Project]] *[http://www.ucmp.berkeley.edu/arthropoda/crustacea/malacostraca.html Introduction to the Malacostraca], [[University of California, Berkeley]] *[https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=22215 Malacostraca] , [[Paleobiology Database|The Paleobiology Database]] *[https://web.archive.org/web/20110706130418/http://www.tafi.org.au/zooplankton/imagekey/malacostraca/ Malacostraca image key - Guide to the marine zooplankton of south eastern Australia], Tasmanian Aquaculture and Fisheries Institute {{Arthropods}} {{Malacostraca}} {{Taxonbar|from=Q182978}} {{Authority control}} [[Category:Malacostraca| ]] [[Category:Arthropod classes]] [[Category:Extant Cambrian first appearances]] </textarea><div class="templatesUsed"><div class="mw-templatesUsedExplanation"><p><span id="templatesused">Pages transcluded onto the current version of this page<span class="posteditwindowhelplinks"> (<a href="/wiki/Help:Transclusion" title="Help:Transclusion">help</a>)</span>:</span> </p></div><ul> 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title="Template:Wikt-lang">Template:Wikt-lang</a> (<a href="/w/index.php?title=Template:Wikt-lang&amp;action=edit" title="Template:Wikt-lang">view source</a>) (extended confirmed protected)</li><li><a href="/wiki/Template:Yesno" title="Template:Yesno">Template:Yesno</a> (<a href="/w/index.php?title=Template:Yesno&amp;action=edit" title="Template:Yesno">view source</a>) (protected)</li><li><a href="/wiki/Template:Yesno-no" title="Template:Yesno-no">Template:Yesno-no</a> (<a href="/w/index.php?title=Template:Yesno-no&amp;action=edit" title="Template:Yesno-no">view source</a>) (template editor protected)</li><li><a href="/wiki/Template:Yesno-yes" title="Template:Yesno-yes">Template:Yesno-yes</a> (<a href="/w/index.php?title=Template:Yesno-yes&amp;action=edit" title="Template:Yesno-yes">view source</a>) (template editor protected)</li><li><a href="/wiki/Module:Ancient_Greek" title="Module:Ancient Greek">Module:Ancient Greek</a> (<a 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href="/wiki/Module:Category_handler/blacklist" title="Module:Category handler/blacklist">Module:Category handler/blacklist</a> (<a href="/w/index.php?title=Module:Category_handler/blacklist&amp;action=edit" title="Module:Category handler/blacklist">view source</a>) (protected)</li><li><a href="/wiki/Module:Category_handler/config" title="Module:Category handler/config">Module:Category handler/config</a> (<a href="/w/index.php?title=Module:Category_handler/config&amp;action=edit" title="Module:Category handler/config">view source</a>) (protected)</li><li><a href="/wiki/Module:Category_handler/data" title="Module:Category handler/data">Module:Category handler/data</a> (<a href="/w/index.php?title=Module:Category_handler/data&amp;action=edit" title="Module:Category handler/data">view source</a>) (protected)</li><li><a href="/wiki/Module:Category_handler/shared" title="Module:Category handler/shared">Module:Category handler/shared</a> (<a 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title="Module:Citation/CS1/Configuration">Module:Citation/CS1/Configuration</a> (<a href="/w/index.php?title=Module:Citation/CS1/Configuration&amp;action=edit" title="Module:Citation/CS1/Configuration">view source</a>) (protected)</li><li><a href="/wiki/Module:Citation/CS1/Date_validation" title="Module:Citation/CS1/Date validation">Module:Citation/CS1/Date validation</a> (<a href="/w/index.php?title=Module:Citation/CS1/Date_validation&amp;action=edit" title="Module:Citation/CS1/Date validation">view source</a>) (protected)</li><li><a href="/wiki/Module:Citation/CS1/Identifiers" title="Module:Citation/CS1/Identifiers">Module:Citation/CS1/Identifiers</a> (<a href="/w/index.php?title=Module:Citation/CS1/Identifiers&amp;action=edit" title="Module:Citation/CS1/Identifiers">view source</a>) (protected)</li><li><a href="/wiki/Module:Citation/CS1/Utilities" title="Module:Citation/CS1/Utilities">Module:Citation/CS1/Utilities</a> (<a href="/w/index.php?title=Module:Citation/CS1/Utilities&amp;action=edit" title="Module:Citation/CS1/Utilities">view source</a>) (protected)</li><li><a href="/wiki/Module:Citation/CS1/Whitelist" title="Module:Citation/CS1/Whitelist">Module:Citation/CS1/Whitelist</a> (<a href="/w/index.php?title=Module:Citation/CS1/Whitelist&amp;action=edit" title="Module:Citation/CS1/Whitelist">view source</a>) (protected)</li><li><a href="/wiki/Module:Citation/CS1/styles.css" title="Module:Citation/CS1/styles.css">Module:Citation/CS1/styles.css</a> (<a href="/w/index.php?title=Module:Citation/CS1/styles.css&amp;action=edit" title="Module:Citation/CS1/styles.css">view source</a>) (protected)</li><li><a href="/wiki/Module:Clade" title="Module:Clade">Module:Clade</a> (<a href="/w/index.php?title=Module:Clade&amp;action=edit" title="Module:Clade">view source</a>) (template editor protected)</li><li><a href="/wiki/Module:Commons_link" title="Module:Commons link">Module:Commons link</a> (<a 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(protected)</li><li><a href="/wiki/Module:Disambiguation/templates" title="Module:Disambiguation/templates">Module:Disambiguation/templates</a> (<a href="/w/index.php?title=Module:Disambiguation/templates&amp;action=edit" title="Module:Disambiguation/templates">view source</a>) (protected)</li><li><a href="/wiki/Module:EditAtWikidata" title="Module:EditAtWikidata">Module:EditAtWikidata</a> (<a href="/w/index.php?title=Module:EditAtWikidata&amp;action=edit" title="Module:EditAtWikidata">view source</a>) (protected)</li><li><a href="/wiki/Module:Exponential_search" title="Module:Exponential search">Module:Exponential search</a> (<a href="/w/index.php?title=Module:Exponential_search&amp;action=edit" title="Module:Exponential search">view source</a>) (template editor protected)</li><li><a href="/wiki/Module:Footnotes" title="Module:Footnotes">Module:Footnotes</a> (<a href="/w/index.php?title=Module:Footnotes&amp;action=edit" title="Module:Footnotes">view source</a>) (template editor protected)</li><li><a href="/wiki/Module:Format_link" title="Module:Format link">Module:Format link</a> (<a href="/w/index.php?title=Module:Format_link&amp;action=edit" title="Module:Format link">view source</a>) (template editor protected)</li><li><a href="/wiki/Module:Hatnote" title="Module:Hatnote">Module:Hatnote</a> (<a href="/w/index.php?title=Module:Hatnote&amp;action=edit" title="Module:Hatnote">view source</a>) (template editor protected)</li><li><a href="/wiki/Module:Hatnote/styles.css" title="Module:Hatnote/styles.css">Module:Hatnote/styles.css</a> (<a href="/w/index.php?title=Module:Hatnote/styles.css&amp;action=edit" title="Module:Hatnote/styles.css">view source</a>) (template editor protected)</li><li><a href="/wiki/Module:Hatnote_list" title="Module:Hatnote list">Module:Hatnote list</a> (<a href="/w/index.php?title=Module:Hatnote_list&amp;action=edit" title="Module:Hatnote list">view source</a>) (template editor protected)</li><li><a href="/wiki/Module:ISO_639_name" 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