CINXE.COM

<!DOCTYPE html><html lang="en-US"><head><title>Just a moment...</title><meta http-equiv="Content-Type" content="text/html; charset=UTF-8"><meta http-equiv="X-UA-Compatible" content="IE=Edge"><meta name="robots" content="noindex,nofollow"><meta name="viewport" content="width=device-width,initial-scale=1"><style>*{box-sizing:border-box;margin:0;padding:0}html{line-height:1.15;-webkit-text-size-adjust:100%;color:#313131;font-family:system-ui,-apple-system,BlinkMacSystemFont,Segoe UI,Roboto,Helvetica Neue,Arial,Noto Sans,sans-serif,Apple Color Emoji,Segoe UI Emoji,Segoe UI Symbol,Noto Color Emoji}body{display:flex;flex-direction:column;height:100vh;min-height:100vh}.main-content{margin:8rem auto;max-width:60rem;padding-left:1.5rem}@media (width <= 720px){.main-content{margin-top:4rem}}.h2{font-size:1.5rem;font-weight:500;line-height:2.25rem}@media (width <= 720px){.h2{font-size:1.25rem;line-height:1.5rem}}#challenge-error-text{background-image:url(data:image/svg+xml;base64,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);background-repeat:no-repeat;background-size:contain;padding-left:34px}@media (prefers-color-scheme:dark){body{background-color:#222;color:#d9d9d9}}</style><meta http-equiv="refresh" content="390"></head><body class="no-js"><div class="main-wrapper" role="main"><div class="main-content"><noscript><div class="h2"><span id="challenge-error-text">Enable JavaScript and cookies to continue</span></div></noscript></div></div><script>(function(){window._cf_chl_opt={cvId: '3',cZone: "onlinelibrary.wiley.com",cType: 'managed',cRay: '8e96dda0bf94ab47',cH: 'K0U0jfXQmsI.6tJ3Nj0_6YJNCHTZVDPLqE3Z_fJwUr4-1732760289-1.2.1.1-1Nwyd_LXrB9UUzEQ.TPEGYW_ycq9oz807ClPIFMagj0NFRWljGfQMjpEzVKVO6TF',cUPMDTk: "\/resolve\/doi?DOI=10.1890\/0012-9658(2006)87%5b1627:MSITAM%5d2.0.CO;2&__cf_chl_tk=olhK73zQqbKbEklhrIDzRR5fH0AZsN8jHCm3B76ZJhM-1732760289-1.0.1.1-ztn4q0pV.VJLDU373GyhmNNZ08uDung0KaVD7RnlCEw",cFPWv: 'b',cITimeS: '1732760289',cTTimeMs: '1000',cMTimeMs: '390000',cTplC: 0,cTplV: 5,cTplB: 'cf',cK: "",fa: "\/resolve\/doi?DOI=10.1890\/0012-9658(2006)87%5b1627:MSITAM%5d2.0.CO;2&__cf_chl_f_tk=olhK73zQqbKbEklhrIDzRR5fH0AZsN8jHCm3B76ZJhM-1732760289-1.0.1.1-ztn4q0pV.VJLDU373GyhmNNZ08uDung0KaVD7RnlCEw",md: "JLNJEXuNxBt_OFY8v_obXupxKq13EY8KMJzS4Ah1fgs-1732760289-1.2.1.1-2O1SUmVlOsWDGfdaP8WWEc9oLfGINiN8HJHKH5J3KGdPfP74wpUnKiALOaWlzRvyYe9O1TsfYxCVpIWP1dMQ6d3.RuKdNf.IzTbvD7nBKI.eY8rP44EwTIWET5D1SZJ9NU3TBhyzvJb8x6gVJq5FP1FNjORwXfZirT.0Ac8Xkrgnxq8GOWSWTO1OyV6DLuSi_tAaCJRz9dx0q5UkFnmh8E1uGsgpHW9TiGGGQaSzcJri8jx3im0D8tKvTT8.BeZeVXYT4n24AKtgYM_ts.MhoN4X_Cro.sShW70L9tIH1Tsi22j.6GEZCOKu_ovbsZW_pQpUK.UqxXjXDjhyraVT5.CrgYqGa4lOo9OvvbAxcJDM_gSWmB6yVuDWuefPb6EOdlKVeXjdCTbytOvhIhl9e33hvLiqL5RSSHvfKSnjxVWRaPGp5UyYqTsN9WqaEvKpLva_9C352V4Ix2.jzvlURaRVste9wE1UI07vTz92m90BpzT.tIFWD442y.TnbDlfpWQRjh3wWS0qMplvhX4ICOI.U6K.vwfSBv04fhSYqlECTykduMD8a76JQnuDn0c9acKah4y91a93kEHRPiKetL7UH.PvI.4gJu8pZ8nV5xC0TDroPcXWlLU6C9LL9wSJw8zMsm6egyR0gZ33rNE9iKYZvUdGM0zrNycCy78SU8GLzcrUM8tpXpwXrpWbSoyaNWOnrjWcSkzIsR.RVg1HwCmSsNfbyZyQ8G5G_HJQHJpskwe7ZJ2bLnE6qkwz94D7fBIQujKuiT64siQdDslLHy2dU0QO3aQNSF3wEDHvz_bSXoOb05KDI0dyxwaZaKKM5wwC4EcNwov6w1HUVgRgZchy00OhSrMLS1gWpKyHIO68MivbokQiNuY5c6glpW1Ug2eo_ASnNIi9J_NOn6j1mxFPM0ujo7eT9.vqMrT0FRY1CA5oDwytoyfysWnO_XL8f0oxkyiL8K7s5MYOy_E6hTLhM1Ix5RV9CpMoIlhjP1zUQpCIfDKFYsjqNljYSIxDc.hjNWtF1JzZP.1EJxkATGz5g3lq5ovcRnjhV.rdQDs8wb3X3YY8QWELGq.A9NtzzWLbG8HI7O3b7p4p3l.I4H.4pwSPwtDhpEfGak5g9TlYnLlKi3VUtsd0aE0GntZWES.WHY_Jw6zRN68yUBxss2Hq_vJ8Nio4ph.NYVh7xpvdUz0v19oE0fLCbj2MkqZM4EuwEZ8hBqHV6phkmGnflWRu6yqlkZDj_HGLr3bNZNFp5n1gdc0guWQU15z.Rudq5MsIMf9.oUER_PwwBxOVYz8iyM6y2gHAC.JdfRazzHsp9Zp1flPHoz6OWd7Y0JbzklEUGznGX11qaBSYxHSBiMAejeT_fs5KqpwFepvwnTfTypddKpvEAo9IXpfYRV6gr351X6FvuQ86CzXMPdvPjO1GuksDxEjJO976fc1pLodIUn8h1iC0faZzvS4Yw28QhGmXcThfk9Ka7i5_FPLVk5R7h.9iCB2r_p4.gn0M4R29igwC6OJdGJasPY7SaP8qIKOQPcUY1WBbyfYYWfv4GWKlRe0ToH7ZKKM9ky22J4SLn9g4JEV57UjGMXT2epKWzzpOzE.MECHPN7iq3x9etknJPnadd_.hH5YFpzCu.53TN.xaAk0Tm2VCEuftgISQN4h4UnQhjqG1OoyUSfsM4tLRA3E53OtvsLDjeI6PX.dBtSg4kQkb.PnasSVeB6FVzIAvF1c.YqevQra_MTrotDzMvD_Ef6QHMpwPYWWFq04DaFsE7JsZWYsHOStnqNxgYKmmdxeDAHndQEfsWQGns5OhM0Js5fuvH5dRbd0CG3ryXma3FIBQ3x3Yn.svEf.vfkKXSMgQN_n4UwTyouKQebAQZHoCrXUofzPMVEm68jLImHEXT6IXGwUTpAOQd.1KW02gLqrVIbZ8zzg4fnRXgiNtpKU2EAdBVNQ3wbOBUcql4CnQj6xem80fdPBV_SMlE9SyguocytOzv02OuUD3PaMRBWYwd5Rp5OOd3MDlSlk.Wxg8oUBYhilEuQizALGReQ2X77itZyT5VKq.GH0SwK0o_gSMyGhBveEFzkTpqwmYaaf8gLhQj6weHU6Vx4x9UWPIoEcjs6djkTP31Ty.WaH6uUFq9emfjKnoYYJbuQQOXcjnG2A9vqFpYOg5kg8LeBSm0C8LnfLWDVM.Y4N9ADueQGCASyRo3VAfMhNYHbWZEzgwdDMu3AMpm0KGY7asMS1LiTh6evHBLhbjhkoWbiyUPrIvnQHzqDT0yF3LK7o.LQr5JWb8Xa7VJiQiOrHZPT66uD2gFtx4J6SiuJWmAJBVt7Ax6l0z62yxjyGgVg6VJK6TXckw8vqR5xNTzFKNo41pzd5Tz4knLLfWewjdeEPlRizLMpcG3vxHtXPTyueX5AcKSahmwQ6Rh1VDRmMbbkPAJ3dxkUrk4RbLUCdBZ35LEWq7SUBxRajkrVsJCTlH7w_vjd3TVWZq5nxVAhhjjpaKRx2xTZol_fxwXiBJ7_MFfm_h7dK8wV7in8JGtI9DH_AdEMdW.X.rMgN2CmvewFqChjhG99LAgFZkvALDXBo4N30DaMLmUb0t7CKCE8i6VS1yfuXSGzQDnq3QmaPf5.L5DDXGRQp4FPepn8YPa1kpZGfCPXF5fpepGgev_jscR.6NRbwaR.tlPR4En3rm8K2ng5DfcdKpfBOJvO2P8dc8y.mAbX0o4zGW5TSGA_YJutZHB3gHsdQqMxOkzc1q4lSIdzuldPdSDl9PZm.Ck4Kbe8BkJpK8ogWAgPf5t1XYA6QowMhI_b0VMW0EbY3nbAy5qJ7xSuG1QvilF6YJvLLO92WA6vwRkeyfbXbvxW54JSa5jtEL3LyIxHRizr3wlzrVK9XipuEHPxRAGxAHwA6VuB2aFTjQQXNSTxlE8aXcQPIvkel8kUjFbYzsU21PJqFLiWXphKDz08AbuHHkdNhLvLJd8JCQcx7A9XvUVBmRPd26dfwoUMxTXFsHvYYVKOvkGQXXwX1LmALuo9APM8SZuo.Kba9OSsQDqItQ_U8CsXEx1XOpj0umXeAf0g425ZfMWfcL2Gc7hGWICP26dQ",mdrd: "wFDgPF13rzrIqbYDhFfZumKoeZQBfBsWAhhyMA0upNM-1732760289-1.2.1.1-0POk2k28TXauSUQadvwBqMC0gU8KUODdQaJ.d0KvP938SSgTbuxKHgkhltVshBldFiylI3e1FIBfe74dXOQu9IUkDV.EDyh_T48kEi4uxjKz0Kf1d13cZLYhj8d1y2k3OWhEDHrbrteK77pRNinfWAn6iAy1JcQ0tvph8hq8BUfk.FbAJS5S7BPNo._jYJK4WWaRE6h3KvxqeoKpECdjg5VCgoB9yWRT_0sdsgiKFXs5bSUQSWup085SwKA42lX7lAKzkfsbv6.SG9xEddPjbPNgeVm8yssIH_ABMuVr_g3PpNilSbUG6Z5yM84pkAI1oMrxyxmHvkTs3MMV2sPhAPNrkz8aHRUu553xwMu7PPRq1sBARNwD1y.gmRnFPEToslVPmzfji_hS8K75j37pQ1ZN5ygSFHWp3wbUdzHMueiC3dOZL_I4RdZGBUxWzkfKZ8oQelwAZCLvHto5amS72tkZxuQx4o8EkLe3anwVFhEuF8b24yIc7Ql5Hdzq6HKIUMgyq0N5z2PSH.TYQo2sh_z.dJwHhEeVFmvAuE_hsjSFaLoELRP_noww.aLtsaiw5yhgZGpD9e5rOGSDoZUj.fKCtIqALf3XqmT4us17dRQDzWz28pFz2L.4g8CMchVzBZ3TnJYnl7hG67nNA6cDCapLAc4o1HN4GRPc3nQXbF1uCp6gz4YJTYhB8Ksx1fc8T5mRcthSftFCLT9i4lQViQ9GpulExHHI6Gl4zu.XYlZbfE4ujdX2tNotlMCn7TNiaKgQkIMk5gT9IRqzl8FKZEydsFnDqjI759fnelObzWNgFWNKVNGhZ88zdC0PiAok3mRIDYNK2qjlpSa49LXSHW6udbJ0eUEmEqjjKF83lLecE.KGa7P90wFG4uXHIFbSD1Uht_1E2e0u65tzPOUT_2XF5xbGh1KFbOMslcsZ3aCYXjYr0YCjQzBEInqHYBuz2s4n2hkNjfeY3ARsxpMALsstREUEyHaB8S6NI7OM.GREKTwfmUAiXqO9zyOUWfh_PbqT3f8AXoztxysP9l9ZVVoD95T5760QPufP5RDB_laZykCx2tDhP7G2QEBMZVxA9VAR4Mz8jFRqu.lE01Em44B3_.jHG5odLO4bL3J7fqA_Ck_uaJCkT74S3R0JmMefL_BOlrTf1mfoC_fVXHEujqsd5OQgsGrnQjJn_z6LeMT9DfYdNj1UwfUcR3rgT2hUkuFSlrPlxaiMocv8iObUfjumKaX88O3PO0stFZTVMn7ZBKvRM0YbDUQvQmAd4gkIR2mI4XmbxaI16CvQLb1JMbPiZMLh1P4q0HmLyrhx.pFBaHFRSjiC_z6POAHCRRcIX6EIaQZr5HYliNgv1agRRNJe8vatVX2iP6VOsE0MVb4SZVzW7PlSSlrILsauiVou9llVK2yS2vYhgknx_AfZNbDwaTJJu.ODjdTdWBZWVld1o_YQc2hZU8BvM8Ykcc1AZM5hmT2IcUfujJ6Pa6XI7YmCX8HizCbvmGIru8wReZAYD2rdpyKQlHZQYeCHP7AWGAKmHImi90E.etYldEK1rk_lYcv3eyUfry07u15WYvddVQvrBEonbHLI0Rp5VXpNqG.v70N2NKAcUJd5h1bjEnLX.jYRMHL_MHUtST9sXzdYbnh5gWGtx2_5yxpsV9frZ5eh1Nx4Jt1Xgkcjj05qjL0ne.FsFTqTgw2gYd3BkfjPAknldRHS9xhBfSxRhzVLl0axDDzrMlTn4p89CCU_CEfL4qISGquj.bq1I0GiUXEmb_3N6y41IbQOfu9mjGvQ6804Y71Dy04kTuby1VnESzrVIZDMDWNYXBc0Hs2fSR.F.L5aC8txWAxtRNnFE_2S1QQ4hzSe9W_eqX1u2hzRNszSJNhLk4tZPd.I4eWt.m7ykiepYz2YKsgvKtb3OnUFCP9Z2qBOPgcAq7Z5hhnG1Z_TXvoCGR7tTH1mWNMjwham6sg_GKQ2UvGg9taF6b98YkezzlnaHz_YG8b3sqdVKg"};var cpo = document.createElement('script');cpo.src = '/cdn-cgi/challenge-platform/h/b/orchestrate/chl_page/v1?ray=8e96dda0bf94ab47';window._cf_chl_opt.cOgUHash = location.hash === '' && location.href.indexOf('#') !== -1 ? '#' : location.hash;window._cf_chl_opt.cOgUQuery = location.search === '' && location.href.slice(0, location.href.length - window._cf_chl_opt.cOgUHash.length).indexOf('?') !== -1 ? '?' : location.search;if (window.history && window.history.replaceState) {var ogU = location.pathname + window._cf_chl_opt.cOgUQuery + window._cf_chl_opt.cOgUHash;history.replaceState(null, null, "\/resolve\/doi?DOI=10.1890\/0012-9658(2006)87%5b1627:MSITAM%5d2.0.CO;2&__cf_chl_rt_tk=olhK73zQqbKbEklhrIDzRR5fH0AZsN8jHCm3B76ZJhM-1732760289-1.0.1.1-ztn4q0pV.VJLDU373GyhmNNZ08uDung0KaVD7RnlCEw" + window._cf_chl_opt.cOgUHash);cpo.onload = function() {history.replaceState(null, null, ogU);}}document.getElementsByTagName('head')[0].appendChild(cpo);}());</script></body></html>