CINXE.COM

<!DOCTYPE html><html lang="en-US"><head><title>Just a moment...</title><meta http-equiv="Content-Type" content="text/html; charset=UTF-8"><meta http-equiv="X-UA-Compatible" content="IE=Edge"><meta name="robots" content="noindex,nofollow"><meta name="viewport" content="width=device-width,initial-scale=1"><style>*{box-sizing:border-box;margin:0;padding:0}html{line-height:1.15;-webkit-text-size-adjust:100%;color:#313131;font-family:system-ui,-apple-system,BlinkMacSystemFont,Segoe UI,Roboto,Helvetica Neue,Arial,Noto Sans,sans-serif,Apple Color Emoji,Segoe UI Emoji,Segoe UI Symbol,Noto Color Emoji}body{display:flex;flex-direction:column;height:100vh;min-height:100vh}.main-content{margin:8rem auto;max-width:60rem;padding-left:1.5rem}@media (width <= 720px){.main-content{margin-top:4rem}}.h2{font-size:1.5rem;font-weight:500;line-height:2.25rem}@media (width <= 720px){.h2{font-size:1.25rem;line-height:1.5rem}}#challenge-error-text{background-image:url(data:image/svg+xml;base64,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);background-repeat:no-repeat;background-size:contain;padding-left:34px}@media (prefers-color-scheme:dark){body{background-color:#222;color:#d9d9d9}}</style><meta http-equiv="refresh" content="120"></head><body class="no-js"><div class="main-wrapper" role="main"><div class="main-content"><noscript><div class="h2"><span id="challenge-error-text">Enable JavaScript and cookies to continue</span></div></noscript></div></div><script>(function(){window._cf_chl_opt={cvId: '3',cZone: "publications.waset.org",cType: 'non-interactive',cRay: '9134ffb44945f8d0',cH: 'WazVIyCNpnwKKrJ1EF4LM.uog_vrcvXnvCBunlifyGY-1739787144-1.2.1.1-q9Z5v.MqLhHer2aC4Fak4sTz0HVWhmzj9T52C_hqQBpODQHUVoeIXxwh.oAKgrT1',cUPMDTk: "\/abstracts\/search?q=fish%20species&page=2&__cf_chl_tk=tupcEJIRwF1dGC2h8J5a0JaifvKmcyPlvaoieWroK7s-1739787144-1.0.1.1-3eZjedB_tPRJUIJaLZhkUYMx4tstc0MASRRm4OL0jeM",cFPWv: 'b',cITimeS: '1739787144',cTTimeMs: '1000',cMTimeMs: '120000',cTplC: 0,cTplV: 5,cTplB: 'cf',cK: "",fa: "\/abstracts\/search?q=fish%20species&page=2&__cf_chl_f_tk=tupcEJIRwF1dGC2h8J5a0JaifvKmcyPlvaoieWroK7s-1739787144-1.0.1.1-3eZjedB_tPRJUIJaLZhkUYMx4tstc0MASRRm4OL0jeM",md: "tQbE04lzBd9QvcLes.cPpIceMOYQB.IKyfD4Y18DoN8-1739787144-1.2.1.1-E.IIoI8o3lr.BfmxtSisZZeEu67J_yqhhU2KsDIhXH9mNeqap2iPFL8Jj8d7VFbKRt5NkQQAzHSDMpEHgxcgKu6Mqp7h9e4UpzFTvn2UKAwRlKRlMh61obxTT5zhlvrZlXJbz.bsRrmBrBN8UCF8M6AkcX3xNKLH9lcQRtSK9FS1t2VVQW5lf8qw1e5NmyafTlTtTV2x_12vxc65Gd_aH5AQhQ2u1c.4kz.HKjx1fRkA1Tnxxdxr7KPc9Qe5Q4EG7abrRIOvI4s9iFuMHzSCN70uurayxBY6K3vyCbJIKnz1Q2ae.RNcfmLmD5kH7ib.2mH6vVpV1AlxNM3Lb5ANSD0r0TCm8OGRp9FbKqDGbC9ByJnPi57s0fPWifXtz4Za4Mg_mpQ6Adkh0p5fHmESl87y53THwjIkGx3mIeLme.UjYNNcjzKWdNkhBh6.wvtUvxaTaFYwwZP9zxluyF5rvkkYYrVTElUZBZriyaGZlf0c1qMfEfEQqUfRglrj9eNVg3rlDDfKLvKk5PVsF1hEyC6iZzB1T4gG.TjlYI2.dipN_ijVOyclxKYQxsj7r4l2bDk1HAB1yR1Qz0LQA84KqvJrFVNkScmb__c7g7yyJoYAPkVj0Y_gLUuhWuUCTH4wkfRtkFWXPGZc2KXWjGSygp0N7DHeIDqdQXQf.7WhHNAD.URuN1VC98avZpUiGTGMHMLQD83o9UbI55sy6Ii7SOYAI5FWIbi3sHdZLDl4kQnqsON934v.DnVQKulWgqlGUnkxb765DHNrnXM2mg0KIfqyj2em3u4afyWhLCnju0PRvwDQUoH_f_RXLBTFnpzWiVLk1z0q4qTjZC6NyTacZGjczbWD_AQ155Bok20eB_xbDDT.jtmimAQgd.D8wY_1wSCJReANZ.IGNhRp0e4leg8pZ70wUz5HTg4dyspygJJRAVz1PzjslU3Ue94Bhnr_SQhF3hlPVOWlqAx2WF6wlUrwy2TcTPdoUFTO07qP6ab2N7ViClEbn4rVen62B_tuSftXm3Z15yZpFqRrXHvWDuUpj6Hvy10l3LuffN47Jl0drVmSYq_oMVV7RbCH1eVn3SKWzO5v280A3V4or7xSE19EwpbTr5pvPLolo.D9tL9oO0t5VuzCUlrzO4uUp_E.Ndmof5dicVcV2XBvOEgLDXwyR3nxGHkVNxshkexrH_Wlc.hdBJFYlqPogXb1zYfbXM0PXjydTmK08GZkvQmAzODgwnBiE2JNy4l9sextTyJ00H7vVFzdSmuzMWhLqcjukFFh0qU5b20UDMcrytX_S6CUw30OwUo2rJDVCK0SNZFhlcXJ0lc5T4mujptd6BL4IpJv9QWPr32gZsPliigJ3Vzz4FyQOJ8CRdn6Befuxe1KH1uylxl1sclvq9dpS5ps0xmoK5ivFTOKdVr165Q3vVQG_ZPZtvf6nVSKtQErmBK_5J.rV4YwFj7zX.3XUY0Xipve7fhQ_MtLMrnpYc2CezgPtQnRqdeiNgueWKou.bhXrhmDK0QA7NFcplht_tT1l243X9x5l2MdHRIrMyuT_ZeJnzHGd8AOp29NtBsG23UY8Ffsr7tSIOEmUMFsz67h5RsKhEYH7BJt2g8PQi2yP3QH9TpF8H0FGmeDbuMuMivVsLVtxtPZ8EP5oySZ0UyBxOrh7Wwi3trOf4yiHtoaZmxuB9TV3B0.1u1JQjCopKNlLWQcqi61dBbobBMpwEDsY8_n5X7Ks52sIrAFs.MKW4w0e_51EaffBv9gr3_mIAHjrKciXY90owt73EQlsLlc7v11vgmVJWXspcSSk9KqxJLVmdcH8RN7PuFrhrIy2Pu9..yigVQwljz.pNLoMthZIIXAs3nTfsYlLc4JzDrOwymEdaGcROJus.6tqx7zfDnbuTVZtlr.RMyPDVg4r7Ti1ZBbEep_IndzdkzQoe389XMngK2hhGKY7_anwdm5bS171X7yZQh3FPCTOqUfMcKC",mdrd: "vZ7bJvgC3GnCTCy0QxyKrwxnYz3UnMvAgu5UC.jczBs-1739787144-1.2.1.1-39xl6WX0nVrZ6f9X6hDLuaOiRGe5C91WeQROU0pNml3jOg2a_8UqeGgezBURVUK_K0lumfAwQPkj_rcJquplfrT_vX5y1np9dCWopzuGoE5.iOTN6DEM_9YMPk465Uao0rtz1J9R2nmY7HkL9aXreQIYisl..wytTUcnLUnlZGz_fPqBZUDMUOklSRV62sgOC2kjrsitgG4FFGmoBE_FXc5hH3EDqzGX9.yikhKOBSu6CCv3gZrJ3CbXd_m4b4.zXOo9wj.F58TTjB1mWiFMkrNpWngPvgEsXIsXSlKllFyTbNnDgerdxVNCSC2Fsexd2LCJuCgkPa2ZGi7NVaDMhLsmo1.8p9wHDwgdusHpJikXuPYJ477w2JPorXI2UK2MkRwJEII2EaPmmm3MvioMFr8lv8rs6QbwlJYXsQXYv1EK_AwvHy2OPXK8XpoeFLMgTh5UJ8_4njIjYqgGzJw9L3bikR8X44wJQ3FAsTNT9plzEzBXvbPcVyNOW6u0VE8nUDE1lUdvPMq7hmsV_sX9.0xBz9F4Es3Qa9f4pjX5k8e0ScDioMZCYLupNrSWQnAfxk19Pr7pCTnphQZpUnVbPRf3ICPyY1GLoHYdaNjKuvDRKyMSQLekHDabSqAvc0H5Gc6gR4yTng9koyOWkvQS7Mfht8vUdoMC.45o8lu_v02aQZa24yASjHAc34SOZoUbkgRAz6Ulj878l40kD26D.ZoE0UbpG901OmdpHBoKqAoErCN_HJR2BJhALSkS0KZT4Ib5Q9zCaailYxgX8R3eWZmCVHPZU47So0.1x9mRI1uRRdrsFnTjc9nh6BltNuKEuXms8NJGZcdhI5L55qtIKpIuRhO_KtyOl6Btl.oftsuMJ_0sBh.78ZrfQbf11u5LGZWi3_V29uXGqnn_HEa6Af9iQamBOyr1YmpEHKdLgJhL53Yo_tPNk2q7CXaZOOYdZ7i7YViZq1vSc8a_EKPD0RYhxVIffqlqQAa652CowymJwe6m22Gp5EIUIly_8ymdpSzX1YLpuWPXJPJAMhcvLcXrSviUt83KN9BAYvzNG0f7_TmqNr.qLImlNo4uBiwDnKnlE3XZiQFsxJPdRBDWM_PepmqAKJ4qKzBf6n8S5Ekt79XKxZ3oggfViTNz_cO0mP4ChUSEfZZ8wKtcxUmWMVe9z3iio4koyZV7MVEeaAqn4ePfff_b192SPs2bY7rRxT1X5BZEL2WCjI1gZOYXTUYmtCWKCWL0hnG82WPQlm82dQj7jaYd5.nAOP9SjY3c3w5YqlRsi8GrpHGQ_Samx5HY0q2f.7RWpXf3MmO_L_ipV.rmPvjuv9rbR8iAD0oBAREg3wpHuZhtxyamK_psl2EmcjXu1NkiYbGPTIzTUvl2eK_nFxQVvBHbC1A27Emq0aMuT94W26_x5Zsyrx9P0Ne9s.3Vt5WrDjN9NAE_8Zu0KqKIJQ7Zu6oDw3_EQPkhewMApRtThsnePxfeKOac4FBY5wgTMmKIGcUcfCzsM5LAiyNuNPqD6cUTz2pJWexRhOmIhbPWzG.QmrqkCyx7UzqOTjBPGk0Hbw9AyPrI.ytqOc8OvTyQe.d11w17fwbuwnquRhD7W9YA_Vf6vikkMm6U74kKhqqA5kvH1idAMAA.1lvEkuMp7fSQKoFHHz2QKgCZIbcl0sKVdXU3LRZB1FkV7yx3eNGYZzXTl4oyOqoTI9OaC1H2cfovc5LItj6aLCBcBlezC3jj8qcjsm8KswzVZi.ZADERKck1T2cNQ7E7I5tzYmrvOHLx2RIiYp3sXbgNt5mZ4bSYutP4GIeTH.DD6U.9kYUAv6Me7IRoLMXaF73d.NdlJVIJe3eWaWfEr7J7nSXv0m3VDh9GhxrP8USCJ.R4BHPZyYMziVG9F8_5pXsBFeqKy5iGXBMd2IzLOj5asfLTt3NBh97cxF6Rc5wZAmiCfq7M3iK1KH9SrO_7V5jXWdpOoP.tPLyP.arS0T1fO7EmYoEmGpmBwxsF7A"};var cpo = document.createElement('script');cpo.src = '/cdn-cgi/challenge-platform/h/b/orchestrate/chl_page/v1?ray=9134ffb44945f8d0';window._cf_chl_opt.cOgUHash = location.hash === '' && location.href.indexOf('#') !== -1 ? '#' : location.hash;window._cf_chl_opt.cOgUQuery = location.search === '' && location.href.slice(0, location.href.length - window._cf_chl_opt.cOgUHash.length).indexOf('?') !== -1 ? '?' : location.search;if (window.history && window.history.replaceState) {var ogU = location.pathname + window._cf_chl_opt.cOgUQuery + window._cf_chl_opt.cOgUHash;history.replaceState(null, null, "\/abstracts\/search?q=fish%20species&page=2&__cf_chl_rt_tk=tupcEJIRwF1dGC2h8J5a0JaifvKmcyPlvaoieWroK7s-1739787144-1.0.1.1-3eZjedB_tPRJUIJaLZhkUYMx4tstc0MASRRm4OL0jeM" + window._cf_chl_opt.cOgUHash);cpo.onload = function() {history.replaceState(null, null, ogU);}}document.getElementsByTagName('head')[0].appendChild(cpo);}());</script></body></html>