New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus Pseudoanthidium (Apoidea: Megachilidae)

<!DOCTYPE html> <html lang="en-US" xml:lang="en-US"> <head> <meta charset="utf-8"> <meta name="viewport" content="width=device-width, initial-scale=1.0"> <title> New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus Pseudoanthidium (Apoidea: Megachilidae) | Zootaxa </title> <meta name="generator" content="Open Journal Systems 3.3.0.6"> <link rel="schema.DC" href="http://purl.org/dc/elements/1.1/" /> <meta name="DC.Creator.PersonalName" content="MAX KASPAREK"/> <meta name="DC.Date.created" scheme="ISO8601" content="2024-11-25"/> <meta name="DC.Date.dateSubmitted" scheme="ISO8601" content="2024-11-21"/> <meta name="DC.Date.issued" scheme="ISO8601" content="2024-11-25"/> <meta name="DC.Date.modified" scheme="ISO8601" content="2024-11-25"/> <meta name="DC.Description" xml:lang="en" content="Wool carder bees of the genus Pseudoanthidium comprise approximately 60–65 species, which are found in the Palaearctic, Indo-Malayan and Afrotropical realms. Their taxonomic relationships are little understood. Herein, I revised West and Central Palaearctic members of the genus. Four species are described as new, namely P. farsiense sp. nov. from Iran, P. microrubrum sp. nov. from Morocco, P. syriacum sp. nov. from Syria, and P. tajikistanicum sp. nov. from Tajikistan. The largely overlooked species Anthidium fulviventre Friese, 1917, described from Russia, and Anthidium ivanovi Mavromoustakis, 1954, described from Tajikistan, are recognized as members of the Pseudoanthidium genus, as P. fulviventre (Friese, 1917) comb. nov. and P. ivanovi (Mavromoustakis, 1954) stat. resurrect. &amp; comb. nov. Anthidium moricei Friese, 1911, from Jordan, and A. royoi Dusmet, 1915, from Morocco, are resurrected from synonymy with P. melanurum and suggested to be treated as P. moricei (Friese, 1911) stat. resurrect. and P. royoi (Dusmet, 1915) stat. resurrect. &amp; comb. nov. The hitherto unknown males of P. moricei (Friese, 1911) and P. rubellulum Pasteels, 1969 are described based on material from Jordan and Israel, respectively. Royanthidium bicoloripenne Pasteels, 1981 (syn. nov.) from Morocco, is revealed to be a junior synonym of P. octodentatum (Pérez, 1895). Morphological traits, along with DNA sequences of the mitochondrial COI gene (“barcoding gene”), allowed clustering the species in five polytypic and five monotypic species groups. As key character traits of the type species of nominate Pseudoanthidium largely fit the subgeneric characters of the subgenus Royanthidium Pasteels, 1969, Royanthidium is regarded as a junior synonym (syn. nov.) of nominate Pseudoanthidium. The species of the subgenus Exanthidium Pasteels, 1969 form a uniform clade both in terms of morphology and DNA marker. An examination of the non-Palaearctic Pseudoanthidium species is suggested to determine whether Exanthidium deserves subgenus status. "/> <meta name="DC.Format" scheme="IMT" content="application/pdf"/> <meta name="DC.Identifier" content="zootaxa.5541.1.1"/> <meta name="DC.Identifier.pageNumber" content="1-50 "/> <meta name="DC.Identifier.DOI" content="10.11646/zootaxa.5541.1.1"/> <meta name="DC.Identifier.URI" content="https://mapress.com/zt/article/view/zootaxa.5541.1.1"/> <meta name="DC.Language" scheme="ISO639-1" content="en"/> <meta name="DC.Rights" content="Copyright (c) 2024 "/> <meta name="DC.Rights" content=""/> <meta name="DC.Source" content="Zootaxa"/> <meta name="DC.Source.ISSN" content="1175-5334"/> <meta name="DC.Source.Issue" content="1"/> <meta name="DC.Source.Volume" content="5541"/> <meta name="DC.Source.URI" content="https://mapress.com/zt"/> <meta name="DC.Subject" xml:lang="en" content="identification key"/> <meta name="DC.Title" content="<strong>New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus <em>Pseudoanthidium</em> (Apoidea: Megachilidae)</strong>"/> <meta name="DC.Type" content="Text.Serial.Journal"/> <meta name="DC.Type.articleType" content="Article"/> <meta name="gs_meta_revision" content="1.1"/> <meta name="citation_journal_title" content="Zootaxa"/> <meta name="citation_journal_abbrev" content="Zootaxa"/> <meta name="citation_issn" content="1175-5334"/> <meta name="citation_author" content="MAX KASPAREK"/> <meta name="citation_author_institution" content="Mönchhofstr. 16; 69120 Heidelberg; Germany"/> <meta name="citation_title" content="<strong>New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus <em>Pseudoanthidium</em> (Apoidea: Megachilidae)</strong>"/> <meta name="citation_language" content="en"/> <meta name="citation_date" content="2024/11/25"/> <meta name="citation_volume" content="5541"/> <meta name="citation_issue" content="1"/> <meta name="citation_firstpage" content="1"/> <meta name="citation_lastpage" content="50"/> <meta name="citation_doi" content="10.11646/zootaxa.5541.1.1"/> <meta name="citation_abstract_html_url" content="https://mapress.com/zt/article/view/zootaxa.5541.1.1"/> <meta name="citation_keywords" xml:lang="en" content="Hymenoptera"/> <meta name="citation_keywords" xml:lang="en" content="taxonomy"/> <meta name="citation_keywords" xml:lang="en" content="new species"/> <meta name="citation_keywords" xml:lang="en" content="synonymy"/> <meta name="citation_keywords" xml:lang="en" content="Anthidium"/> <meta name="citation_keywords" xml:lang="en" content="wool carder bees"/> <meta name="citation_keywords" xml:lang="en" content="identification key"/> <meta name="citation_pdf_url" content="https://mapress.com/zt/article/download/zootaxa.5541.1.1/54884"/> <meta name="citation_reference" content="Ascher, J.S. & Pickering, J. (2024) Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila). Available from: http://www.discoverlife.org/mp/20q?guide=Apoidea_species (accessed 1 August 20024)"/> <meta name="citation_reference" content="Baker, D.B. (2004) Type material of Hymenoptera described by O. I. Radoszkowsky in the Natural History Museum, London, and the localities of A. P. Fedtschenko’s Reise in Turkestan. Mitteilungen aus dem Museum für Naturkunde Berlin, Deutsche Entomologische Zeitschrift, 51, 231–252. https://doi.org/10.1002/mmnd.4810510207"/> <meta name="citation_reference" content="Boustani, M., Dathe, H., Ghisbain, G., Kasparek, M., Michez, D., Nemer, N., Pauly, A., Risch, S., Terzo, M., Van Achter, X., Wood, T. & Rasmont, P. (2021) The bees of Lebanon (Hymenoptera: Apoidea: Anthophila). Zootaxa, 4976 (1), 1–146. https://doi.org/10.11646/zootaxa.4976.1.1"/> <meta name="citation_reference" content="Dusmet y Alonso, J.M. (1915) Ápidos de Marruecos de los gen. Anthidium, Nomada, Melecta, Crocisa, Coelioxys y Phiarus. Memorias de la Real Sociedad Española de Historia Natural, 8, 293–334."/> <meta name="citation_reference" content="Friese, H. (1898) Die Bienen Europa‘s (Apidae europaeae) nach ihren Gattungen, Arten und Varietäten auf vergleichend morphologisch-biologischer Grundlage. Theil IV: Solitäre Apiden: Genus Eriades. Genus Trachusa. Genus Anthidium. Akademische Druck- und Verlagsanstalt, Innsbruck, 303 pp."/> <meta name="citation_reference" content="Friese, H. (1917) Neue Arten der Bienengattung Anthidium (Hym.) (Paläarktische Region und von Formosa). Deutsche Entomologische Zeitschrift, 1917, 49–60."/> <meta name="citation_reference" content="Kasparek, M. (2017) Resin bees of the anthidiine genus Trachusa. Identification, taxonomy, distribution and biology of the Old World species. Entomofauna Supplement, 21, 1–152."/> <meta name="citation_reference" content="Kasparek, M. (2021) The bee genus Pseudoanthidium: Revision of the subgenus Exanthidium with the description of a new species (Apoidea: Megachilidae). Fragmenta entomologica, 53, 333–346."/> <meta name="citation_reference" content="Kasparek, M. (2022) The Resin and Wool Carder Bees (Anthidiini) of Europe and Western Turkey. Identification. Distribution. Biology. Chimaira, Frankfurt, 292 pp. [unchanged reprint 2023]"/> <meta name="citation_reference" content="Kasparek, M. & Ebmer, A.W. (2023) The wool carder bee Pseudoanthidium alpinum (Morawitz, 1873): identity of the enigmatic type species of the genus Pseudoanthidium (Hymenoptera: Megachilidae: Anthidiini). Osmia, 11, 39–50. https://doi.org/10.47446/OSMIA11.7"/> <meta name="citation_reference" content="Kasparek, M., Hauser, M. & Schmid-Egger, Ch. (2023) Taxonomic revision of the Afro-Arabian anthidiine bee Pseudoanthidium ochrognathum sensu lato (Apoidea: Anthidiini) based on morphological and genetic data. Integrative Systematics, 6 (2), 55–69. https://doi.org/10.18476/2023.526995"/> <meta name="citation_reference" content="Kasparek, M. & Kuhlmann, M. (2024) Do the members of the rare African bee genus Anthidioma deserve distinct genus status? Linzer biologische Beiträge, 56 (1), 111–120."/> <meta name="citation_reference" content="Kasparek, M. & J. Monks (2024) Not all male bees posses thirteen antennal segments: New megachilid species from the Afro-Arabian and Afrotropical regions (Apoidea: Anthidiini: Pseudoanthidium) with deviant numbers of segments. Zoology in the Middle East, 70, 1–18."/> <meta name="citation_reference" content="Kumar, V., Griswold, T. & Belavadi, V.V. (2017) The resin and carder bees of south India (Hymenoptera: Megachilidae: Anthidiini). Zootaxa, 4317 (3), 436–468. https://doi.org/10.11646/zootaxa.4317.3.2"/> <meta name="citation_reference" content="Lhomme, P., Michez, D., Christmann, S., Scheuchl, E., El Abdouni, I., Hamroud, L., Ihsane, O., Sentil, A., Smaili, M.C., Schwarz, M., Dathe, H., Straka, J., Pauly, A., Schmid-Egger, Ch., Patiny, S., Müller, A., Praz, Ch., Risch, S., Kasparek, M., Kuhlmann, M., Wood, Th. J., Bogusch, P., Ascher, J. & Rasmont, R. (2020) The wild bees (Hymenoptera: Apoidea) of Morocco. Zootaxa, 4892 (1), 1–159. https://doi.org/10.11646/zootaxa.4892.1.1"/> <meta name="citation_reference" content="Litman, J. R., Griswold, T. & Danforth, B.N. (2016) Phylogenetic systematics and a revised generic classification of anthidiine bees (Hymenoptera: Megachilidae). Molecular Phylogenetics and Evolution, 100, 183–198. https://doi.org/10.1016/j.ympev.2016.03.018"/> <meta name="citation_reference" content="Litman, J.R., Fateryga, A.V., Griswold, T.L., Aubert, M., Proshchalykin, M.Yu., Le Divelec, R., Burrows, S. & Praz, Ch.J. (2021) Paraphyly and low levels of genetic divergence in morphologically distinct taxa: revision of the Pseudoanthidium scapulare complex of carder bees (Apoidea: Megachilidae: Anthidiini). Zoological Journal of the Linnean Society, 195, 1287–1337. [2022] https://doi.org/10.1093/zoolinnean/zlab062"/> <meta name="citation_reference" content="Mavromoustakis, G.A. (1948) On some anthidiine bees (Apoidea) from Palestine. Part III. Annals and Magazine of Natural History, Series 11, 14, 428–434. https://doi.org/10.1080/00222934708654652"/> <meta name="citation_reference" content="Mavromoustakis, G.A. (1954) New and little-known bees of the subfamily Anthidiinae (Apoidea). Part VII. Annals and Magazine for Natural History, Series 12, 7, 249–252. https://doi.org/10.1080/00222935408651726"/> <meta name="citation_reference" content="Michener, Ch.D. (2007) The Bees of the World. Second edition. The Johns Hopkins University Press, Baltimore, Maryland, 992 pp."/> <meta name="citation_reference" content="Michener, Ch.D. & Griswold, T.L. (1994) The classification of Old World Anthidiini (Hymenoptera, Megachilidae). The University of Kansas Science Bulletin, 55, 299–327."/> <meta name="citation_reference" content="MoA [Ministry of Agriculture and Natural Resources, Department of Agriculture] (1989) Hymenoptera. The systematic list of the Mavromoustakis collection. Nicosia."/> <meta name="citation_reference" content="Mocsáry, A. (1884) Species generis Anthidium Fabr. regionis Palaearcticae. Természetrajzi Füzetek Kiadija a Magyar Nemzeti Müzeum [Naturhistorische Hefte herausgegeben vom Ungarischen National-Museum], 8, 241–278."/> <meta name="citation_reference" content="Morawitz, F.F. (1873 [1874]) Die Bienen Daghestans. Horae Societatis Entomologicae Rossicae (Trudy Russkago éntomologicheskago obshchestva), 10, 129–189."/> <meta name="citation_reference" content="Morawitz, F.F. (1875) A travel to Turkestan by the member-founder of the society A. P. Fedtschenko accomplished from the Imperial Society of Naturalists, Anthropologists, and Etnographists on a Commission from the General-Governor of Turkestan K. P. von Kaufmann (Issue 9). Vol. 11. Zoogeographical Investigations. Pt V. (Division 7). Bees (Mellifera). Izvestiya Imperatorskogo Obshchestva Lyubiteley Estestvoznaniya, Anthropologii i Ethnografii, 19, 1–160."/> <meta name="citation_reference" content="Morawitz, F.F. (1888) Hymenoptera aculeata nova. Horae Societatis Entomologicae Rossicae (Trudy Russkago éntomologicheskago obshchestva), 22, 224–302."/> <meta name="citation_reference" content="Morawitz, F.F. (1894 [1893]) Supplement zur Bienenfauna Turkestans. Horae Societatis Entomologicae Rossicae (Trudy Russkago éntomologicheskago obshchestva), 28, 3–87."/> <meta name="citation_reference" content="Nalinrachatakan, P., Ascher, J.S., Kasparek, M., Traiyasut, P., Thanoosing, Ch. & Warrit, N. (2023) A review of the anthidiine bees (Apoidea, Megachilidae) in Thailand. Zookeys, 1186, 235–284. https://doi.org/10.3897/zookeys.1186.95203"/> <meta name="citation_reference" content="Niu, Z.-Q., Luo, A.-R., Griswold, T. & Zhu, C.-D. (2021) Review of the bee genus Pseudoanthidium Friese, 1898 (Hymenoptera: Apoidea: Megachilidae: Anthidiini) of China with descriptions of three new species. Zootaxa, 4996 (1), 133–152. https://doi.org/10.11646/zootaxa.4996.1.5"/> <meta name="citation_reference" content="Ornosa, C., Ortiz-Sánchez, F.J. & Torres, F. (2008) Catálogo de los Megachilidae del Mediterráneo occidental (Hymenoptera, Apoidea). III. Anthidiini y Dioxyini. Graellsia, 64, 61–86. https://doi.org/10.3989/graellsia.2008.v64.i1.55"/> <meta name="citation_reference" content="Pagliano, G. (1994) Catalogo degli Imenotteri Italiani. IV. (Apoidea: Colletidae, Andrenidae, Megachilidae, Anthophoridae, Apidae). Memorie della Societa Entomologica Italiana, 72, 331–467."/> <meta name="citation_reference" content="Pasteels, J.J. (1969) La systématique generique et subgenerique des Anthidiinae (Hymenoptera, Apoidea, Megachilidae) de l‘Ancien Monde. Memoires de la Société Royale d‘Entomologie de Belgique, 31, 3–148."/> <meta name="citation_reference" content="Pasteels, J.J. (1984) Révision des Anthidiinae (Hymenoptera, Apoidea, Megachilidae) de l’Afrique subsaharienne. Académie Royale de Belgique, Mémoires de la Classe des Sciences, Collection in-4°-2e Série, 19 (1), 1–165."/> <meta name="citation_reference" content="Pérez, J. (1895) Espèces nouvelles de mellifères de Barbarie (diagnoses préliminaires). G. Gounouihou, Bordeaux, 64 pp"/> <meta name="citation_reference" content="Popov, V.B. (1950) Generic groupings of the mid-Asian bees of the subgenus Anthiniinae (Hymenoptera, Megachilidae). Doklady Akademii Nauk SSSR, 70, 315–318."/> <meta name="citation_reference" content="Popov, V.B. (1952) The fauna of bees (Hymenoptera, Apoidea) of south-western Turkmenia and their landscape distribution. Trudy Zoologicheskogo Instituta, Akademiya Nauk SSSR, Leningrad, 10, 61–117."/> <meta name="citation_reference" content="Proshkalykin, M.Yu. & Astafurova, Y.V. (2017) The history of study of the Russian bees (Hymenoptera, Anthophila). A.I. Kurentsov’s Annual Memorial Meetings, 28, 26–34."/> <meta name="citation_reference" content="Proshchalykin, M.Yu. & Fateryga, A.V. (2017) 77. Family Megachilidae. In: Belokobylskij, S.A. & Lelej, A.S. (Eds.), Annotated Catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute of the Russian Academy of Sciences Supplement, 6, pp. 295–308."/> <meta name="citation_reference" content="Rasmussen, C. & Ascher, J.S. (2008) Heinrich Friese (1860–1948): Names proposed and notes on a pioneer melittologist (Hymenoptera, Anthophila). Zootaxa, 1833 (1), 1–118. https://doi.org/10.11646/zootaxa.1833.1.1"/> <meta name="citation_reference" content="Sandhouse, G.A. (1943) The type species of the genera and subgenera of bees. Proceedings of the United States National Museum, 92, 519–619. https://doi.org/10.5479/si.00963801.3156.519"/> <meta name="citation_reference" content="Tamura, K.. Stecher, G. & Kumar, S. (2021) MEGA11: Molecular Evolutionary Genetics Analysis version 11. Molecular Biology and Evolution, 38, 3022–3027. https://doi.org/10.1093/molbev/msab120)"/> <meta name="citation_reference" content="Vereecken, N.J., Ruiz, C., Marshall, L., Pérez-Gil, M., Molenberg, J.-M., Jacobi, B., La Roche, F. & Litman, J.R. (2023) A new small carder bee species from the eastern Canary Islands (Hymenoptera, Megachilidae, Anthidiini). Journal of Hymenoptera Research, 96, 983–1015. https://doi.org/10.3897/jhr.96.111550"/> <meta name="citation_reference" content="Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpaläarktis und im turkestanischen Becken. Entomofauna, 1, 119–209."/> <meta name="citation_reference" content="Warncke, K. (1982) Beitrag zur Bienenfauna des Iran. 15. Die Gattung Anthidium F. Bollettino del Museo Civico di Storia Naturale di Venezia, 32 (1981), 171–196."/> <meta name="citation_reference" content="Zwickl, D.J. & Hillis, D.M. (2002) Increased taxon sampling greatly reduces phylogenetic error. Systematic Biology, 51 (4), 588–598. https://doi.org/10.1080/10635150290102339"/> <link rel="alternate" type="application/atom+xml" href="https://mapress.com/zt/gateway/plugin/WebFeedGatewayPlugin/atom"> <link rel="alternate" type="application/rdf+xml" href="https://mapress.com/zt/gateway/plugin/WebFeedGatewayPlugin/rss"> <link rel="alternate" type="application/rss+xml" href="https://mapress.com/zt/gateway/plugin/WebFeedGatewayPlugin/rss2"> <link rel="stylesheet" href="https://mapress.com/plugins/themes/oldGregg/resources/bootstrap/css/bootstrap.min.css?v=3.3.0.6" type="text/css" /><link rel="stylesheet" href="https://mapress.com/zt/$$$call$$$/page/page/css?name=less" type="text/css" /><link rel="stylesheet" href="https://fonts.googleapis.com/css?family=Lora:400,400i,700,700i&display=swap&subset=cyrillic,vietnamese" rel="stylesheet//fonts.googleapis.com/css?family=Lora" type="text/css" /><link rel="stylesheet" href="//fonts.googleapis.com/css?family=PT+Serif:400,400i,700,700i&display=swap&subset=cyrillic" type="text/css" /><link rel="stylesheet" href="//fonts.googleapis.com/css?family=Source+Sans+Pro:400,600,700,900&display=swap&subset=cyrillic" type="text/css" /><link rel="stylesheet" href="https://mapress.com/plugins/generic/citations/css/citations.css?v=3.3.0.6" type="text/css" /><link rel="stylesheet" href="https://mapress.com/public/journals/1/styleSheet.css?d=2024-01-25+09%3A25%3A41" type="text/css" /><link rel="stylesheet" href="https://mapress.com/public/site/styleSheet.css?v=3.3.0.6" type="text/css" /> <meta property="og:image" content="https://mapress.com/public/journals/1/article_53498_cover_en_US.jpg" />  <script> var _paq = window._paq = window._paq || []; /* tracker methods like "setCustomDimension" should be called before "trackPageView" */ _paq.push(['trackPageView']); _paq.push(['enableLinkTracking']); (function() { var u="//mapress.com/publisher/wp-admin/matomo/"; _paq.push(['setTrackerUrl', u+'matomo.php']); _paq.push(['setSiteId', '1']); var d=document, g=d.createElement('script'), s=d.getElementsByTagName('script')[0]; g.async=true; g.src=u+'matomo.js'; s.parentNode.insertBefore(g,s); })(); </script>  </head> <body class="pkp_page_article pkp_op_view has_site_logo" dir="ltr"> <div class="cmp_skip_to_content"> <a class="sr-only" href="#header">Skip to main content</a> <a class="sr-only" href="#content">Skip to main navigation menu</a> <a class="sr-only" href="#footer">Skip to site footer</a> </div> <header class="container-fluid"> <div class="column"> <div class="journal_branding col-md-4"> <a href=" https://mapress.com/zt/index " class="is_img"> <img src="https://mapress.com/public/journals/1/pageHeaderLogoImage_en_US.png" width="715" height="205" class="img-fluid" alt="Responsive image" /> </a> </div> <div class="navigation_wrapper col-md-8"> <nav class="usernav navbar navbar-expand navbar-light" aria-label="User Navigation"> <ul id="navigationUser" class=" navbar-nav ml-md-auto"> <li class="profile nav-item"> <a class="nav-link" href="https://mapress.com/zt/user/register">Register</a> </li> <li class="profile nav-item"> <a class="nav-link" href="https://mapress.com/zt/login">Login</a> </li> </ul> </nav> <nav class="primaryNav navbar navbar-expand-lg navbar-light" aria-label="Site Navigation"> <button class="navbar-toggler" type="button" data-toggle="collapse" data-target="#primaryNavbarContent" aria-controls="primaryNavbarContent" aria-expanded="false" aria-label="Toggle navigation"> <span class="navbar-toggler-icon"></span> </button> <div class="collapse navbar-collapse" id="primaryNavbarContent"> <ul id="navigationPrimary" class=" navbar-nav mr-auto"> <li class="nav-item"> <a class="nav-link" href="https://mapress.com/zt/index"> Home </a> </li> <li class=" nav-item"> <a class="nav-link" href="https://store.mapress.com">Store</a> </li> <li class=" nav-item"> <a class="nav-link" href="https://mapress.com/zt/issue/current">Current</a> </li> <li class=" nav-item"> <a class="nav-link" href="https://mapress.com/zt/publishedarticles">Articles</a> </li> <li class=" nav-item"> <a class="nav-link" href="https://mapress.com/zt/issue/archive">Archives</a> </li> <li class=" nav-item"> <a class="nav-link" href="https://mapress.com/zt/sv">Special volume</a> </li> <li class=" nav-item dropdown"> <a class="nav-link dropdown-toggle" href="https://mapress.com/zt/about" id="navbarDropdown" role="button" data-toggle="dropdown" aria-haspopup="true" aria-expanded="false"> About </a> <div class="dropdown-menu" aria-labelledby="navbarDropdown"> <a class=" dropdown-item" href="https://mapress.com/zt/about"> About the Journal </a> <a class=" dropdown-item" href="https://mapress.com/zt/about/submissions"> Submissions </a> <a class=" dropdown-item" href="https://mapress.com/zt/announcement"> Announcements </a> <a class=" dropdown-item" href="https://mapress.com/zt/about/privacy"> Privacy Statement </a> <a class=" dropdown-item" href="https://mapress.com/zt/qna"> Q & A </a> <a class=" dropdown-item" href="https://mapress.com/zt/Advantage"> Advantages </a> <a class=" dropdown-item" href="https://mapress.com/zt/about/contact"> Contact </a> </div> </li> <li class=" nav-item"> <a class="nav-link" href="https://mapress.com/zt/subscriptioninfo">Subscription</a> </li> <li class=" nav-item"> <a class="nav-link" href="https://mapress.com/zt/search/search">Search</a> </li> <li class=" nav-item"> <a class="nav-link" href="https://mapress.com/zt/Editors">Full editor list:</a> </li> <li class=" nav-item"> <a class="nav-link" href="https://mapress.com/zt/general">Theory & Methodology</a> </li> <li class=" nav-item"> <a class="nav-link" href="https://mapress.com/zt/Protist">Protist</a> </li> <li class=" nav-item dropdown"> <a class="nav-link dropdown-toggle" href="https://mapress.com" id="navbarDropdown" role="button" data-toggle="dropdown" aria-haspopup="true" aria-expanded="false"> VERTEBRATES </a> <div class="dropdown-menu" aria-labelledby="navbarDropdown"> <a class=" dropdown-item" href="https://mapress.com/zt/Pisces"> Pisces-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Amphibia"> Amphibia-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Reptilia"> Reptilia-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Aves"> Aves-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Mammalia"> Mammalia-Editors </a> </div> </li> <li class=" nav-item dropdown"> <a class="nav-link dropdown-toggle" href="https://mapress.com" id="navbarDropdown" role="button" data-toggle="dropdown" aria-haspopup="true" aria-expanded="false"> ARTHROPODS </a> <div class="dropdown-menu" aria-labelledby="navbarDropdown"> <a class=" dropdown-item" href="https://mapress.com/zt/Trilobita"> Trilobita-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Pycnogonida"> Pycnogonida-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Opiliones"> Opiliones-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Scorpiones"> Scorpiones-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Solifugae"> Solifugae-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Pseudoscorpiones"> Pseudoscorpiones-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Palpigradi"> Palpigradi-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Ricinulei"> Ricinulei-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Acari"> Acari-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Araneae"> Araneae-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Amblypygi"> Amblypygi-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Uropygida"> Thelyphonida-(Previously-Uropygida)-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Schizomida"> Schizomida-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Myriapoda"> Myriapoda-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Crustacea"> Crustacea-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Collembola"> Collembola-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Protura"> Protura-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Diplura"> Diplura-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Microcoryphia"> Microcoryphia-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Zygentoma"> Zygentoma-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Ephemeroptera"> Ephemeroptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Odonata"> Odonata-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Titanoptera"> Titanoptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Orthoptera"> Orthoptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Phasmatodea"> Phasmatodea-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Embioptera"> Embioptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Grylloblattodea"> Grylloblattodea-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Plecoptera"> Plecoptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Dermaptera"> Dermaptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Zoraptera"> Zoraptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Mantodea"> Mantodea-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Blattodea"> Blattodea-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Isoptera"> Isoptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Psocoptera"> Psocoptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Phthiraptera"> Phthiraptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Thysanoptera"> Thysanoptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Hemiptera"> Hemiptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Hymenoptera"> Hymenoptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Strepsiptera"> Strepsiptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Coleoptera"> Coleoptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Megaloptera-Neuroptera-Raphidioptera"> Megaloptera-Neuroptera-Raphidioptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Trichoptera"> Trichoptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Lepidoptera"> Lepidoptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Diptera"> Diptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Siphonaptera"> Siphonaptera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Mecoptera"> Mecoptera-Editors </a> </div> </li> <li class=" nav-item dropdown"> <a class="nav-link dropdown-toggle" href="https://mapress.com" id="navbarDropdown" role="button" data-toggle="dropdown" aria-haspopup="true" aria-expanded="false"> OTHER INVERTEBRATES </a> <div class="dropdown-menu" aria-labelledby="navbarDropdown"> <a class=" dropdown-item" href="https://mapress.com/zt/Ctenophora"> Ctenophora-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Porifera"> Porifera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Coelenterata"> Coelenterata-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Xenoturbellida"> Xenoturbellida-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Acoelomorpha"> Acoelomorpha-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Tunicata"> Tunicata-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Ascidiacea"> Ascidiacea-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Echinodermata"> Echinodermata-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Hemichordata"> Hemichordata-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Nematoda"> Nematoda-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Nematomorpha"> Nematomorpha-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Tardigrada"> Tardigrada-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Onychophora"> Onychophora-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Loricifera"> Loricifera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Kinorhyncha"> Kinorhyncha-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Bryozoa"> Bryozoa-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Entoprocta"> Entoprocta-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Annelida"> Annelida-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Sipuncula"> Sipuncula-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Mollusca"> Mollusca-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Nemertea"> Nemertea-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Echiura"> Echiura-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Brachiopoda"> Brachiopoda-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Phoronida"> Phoronida-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Gastrotricha"> Gastrotricha-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Platyhelminthes"> Platyhelminthes-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Gnathostomulida"> Gnathostomulida-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Rotifera"> Rotifera-Editors </a> <a class=" dropdown-item" href="https://mapress.com/zt/Acanthocephala"> Acanthocephala-Editors </a> </div> </li> </ul> </div> </nav> </div> </div> </header> <main> <div class="jatsParser__container"> <div class="jatsParser__meta"> <div class="jatsParser__cover-wrapper"> <img class="jatsParser__cover" src="https://mapress.com/public/journals/1/article_53498_cover_en_US.jpg"> </div> <div class="jatsParser__meta-row"> <div class="jatsParser__meta-section-title"> <span>Issue: <a href="https://mapress.com/zt/issue/view/zootaxa.5541.1"> Vol. 5541 No. 1: 25 Nov. 2024 </a> </span> </div> <div class="jatsParser__meta-section-title"> <span>Type: Article</span> </div> <div class="jatsParser__meta-date-published"> <span>Published:<span> <span>2024-11-25</span> </div> <div class="jatsParser__meta-doi"> <span class="jatsParser__doi-label"> DOI: </span> <span class="jatsParser__meta-doi-value"> <a href="https://doi.org/10.11646/zootaxa.5541.1.1"> 10.11646/zootaxa.5541.1.1 </a> </span> </div> <div class="pages"> <span>Page range:</span> 1-50 </div> <div class="views"> <span>Abstract views:</span> 139 </div> <div class="views"> <span>PDF downloaded:</span> 13 </div> </div> <h1 class="jatsParser__meta-title"><strong>New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus <em>Pseudoanthidium</em> (Apoidea: Megachilidae)</strong></h1> <ul class="jatsParser__meta-authors"> <li><a class="jatsParser__meta-author-string-href" href="#author-1"><span class="jatsParser__meta-author">MAX KASPAREK</span><sup class="jatsParser__meta-symbol jatsParser__symbol-plus">+</sup><sup class="jatsParser__meta-symbol jatsParser__symbol-minus jatsParser__hide">−</sup></a><a class="jatsParser__meta-orcidImage" href="https://orcid.org/0000-0002-5604-6791"><img src="https://mapress.com/plugins/themes/oldGregg/templates/images/orcid.png"></a></li> </ul> <div class="jatsParser__details-authors"> <div class="jatsParser__details-author jatsParser__hideAuthor" id="jatsParser__author-1"> <div class="jatsParser__details-author-affiliation">Mönchhofstr. 16; 69120 Heidelberg; Germany</div> </div> </div> <div class="jatsParser__keywords-wrapper"> <div class="jatsParser__keywords-row"> <span class="jatsParser__keyword">Hymenoptera</span> <span class="jatsParser__keyword">taxonomy</span> <span class="jatsParser__keyword">new species</span> <span class="jatsParser__keyword">synonymy</span> <span class="jatsParser__keyword">Anthidium</span> <span class="jatsParser__keyword">wool carder bees</span> <span class="jatsParser__keyword">identification key</span> </div> </div> </div> <div class="jatsParser__articleView"> <div class="jatsParser__left-article-block"> <div class="article-page__galleys"> <ul class="list-galleys primary-galleys"> <li> <a class="obj_galley_link pdf restricted" href="https://mapress.com/zt/article/view/zootaxa.5541.1.1/54884"> <span class="pkp_screen_reader"> Requires Subscription or Fee </span> PDF(38.15MB) </a> </li> </ul> </div> </div> <div class="jatsParser__center-article-block"> <div class="jatsParser__article-fulltext" id="jatsParserFullText"> <h2 class="article-section-title jatsParser__abstract">Abstract</h2> <p>Wool carder bees of the genus <em>Pseudoanthidium</em> comprise approximately 60–65 species, which are found in the Palaearctic, Indo-Malayan and Afrotropical realms. Their taxonomic relationships are little understood. Herein, I revised West and Central Palaearctic members of the genus. Four species are described as new, namely <em>P. farsiense</em> <strong>sp. nov. </strong>from Iran, <em>P. microrubrum</em> <strong>sp. nov. </strong>from Morocco, <em>P. syriacum</em> <strong>sp. nov.</strong> from Syria, and <em>P. tajikistanicum</em> <strong>sp. nov.</strong> from Tajikistan. The largely overlooked species <em>Anthidium fulviventre</em> Friese, 1917, described from Russia, and <em>Anthidium ivanovi</em> Mavromoustakis, 1954, described from Tajikistan, are recognized as members of the <em>Pseudoanthidium </em>genus, as <em>P. fulviventre</em> (Friese, 1917) <strong>comb. nov.</strong> and <em>P. ivanovi</em> (Mavromoustakis, 1954) <strong>stat. resurrect. & comb. nov.</strong> <em>Anthidium moricei</em> Friese, 1911, from Jordan, and <em>A. royoi </em>Dusmet, 1915, from Morocco, are resurrected from synonymy with <em>P. melanurum</em> and suggested to be treated as <em>P. moricei</em> (Friese, 1911) <strong>stat. resurrect.</strong> and <em>P. royoi </em>(Dusmet, 1915) <strong>stat. resurrect. & comb. nov. </strong>The hitherto unknown males of <em>P. moricei</em> (Friese, 1911) and <em>P. rubellulum</em> Pasteels, 1969 are described based on material from Jordan and Israel, respectively. <em>Royanthidium bicoloripenne </em>Pasteels, 1981 <strong>(syn. nov.)</strong> from Morocco, is revealed to be a junior synonym of <em>P. octodentatum</em> (Pérez, 1895). Morphological traits, along with DNA sequences of the mitochondrial COI gene (“barcoding gene”), allowed clustering the species in five polytypic and five monotypic species groups. As key character traits of the type species of nominate <em>Pseudoanthidium</em> largely fit the subgeneric characters of the subgenus <em>Royanthidium </em>Pasteels, 1969, <em>Royanthidium</em> is regarded as a junior synonym <strong>(syn. nov.)</strong> of nominate <em>Pseudoanthidium</em>. The species of the subgenus <em>Exanthidium</em> Pasteels, 1969 form a uniform clade both in terms of morphology and DNA marker. An examination of the non-Palaearctic <em>Pseudoanthidium</em> species is suggested to determine whether <em>Exanthidium</em> deserves subgenus status.</p> <p> </p> <h2 class="article-section-title jatsParser__references">References</h2> <ol class="jatsParser__references-list"> <li>Ascher, J.S. & Pickering, J. (2024) Discover Life bee species guide and world checklist (Hymenoptera: Apoidea: Anthophila). Available from: <a href="http://www.discoverlife.org/mp/20q?guide=Apoidea_species">http://www.discoverlife.org/mp/20q?guide=Apoidea_species</a> (accessed 1 August 20024) </li> <li>Baker, D.B. (2004) Type material of Hymenoptera described by O. I. Radoszkowsky in the Natural History Museum, London, and the localities of A. P. Fedtschenko’s Reise in Turkestan. Mitteilungen aus dem Museum für Naturkunde Berlin, Deutsche Entomologische Zeitschrift, 51, 231–252. <a href="https://doi.org/10.1002/mmnd.4810510207">https://doi.org/10.1002/mmnd.4810510207</a> </li> <li>Boustani, M., Dathe, H., Ghisbain, G., Kasparek, M., Michez, D., Nemer, N., Pauly, A., Risch, S., Terzo, M., Van Achter, X., Wood, T. & Rasmont, P. (2021) The bees of Lebanon (Hymenoptera: Apoidea: Anthophila). Zootaxa, 4976 (1), 1–146. <a href="https://doi.org/10.11646/zootaxa.4976.1.1">https://doi.org/10.11646/zootaxa.4976.1.1</a> </li> <li>Dusmet y Alonso, J.M. (1915) Ápidos de Marruecos de los gen. Anthidium, Nomada, Melecta, Crocisa, Coelioxys y Phiarus. Memorias de la Real Sociedad Española de Historia Natural, 8, 293–334. </li> <li>Friese, H. (1898) Die Bienen Europa‘s (Apidae europaeae) nach ihren Gattungen, Arten und Varietäten auf vergleichend morphologisch-biologischer Grundlage. Theil IV: Solitäre Apiden: Genus Eriades. Genus Trachusa. Genus Anthidium. Akademische Druck- und Verlagsanstalt, Innsbruck, 303 pp. </li> <li>Friese, H. (1917) Neue Arten der Bienengattung Anthidium (Hym.) (Paläarktische Region und von Formosa). Deutsche Entomologische Zeitschrift, 1917, 49–60. </li> <li>Kasparek, M. (2017) Resin bees of the anthidiine genus Trachusa. Identification, taxonomy, distribution and biology of the Old World species. Entomofauna Supplement, 21, 1–152. </li> <li>Kasparek, M. (2021) The bee genus Pseudoanthidium: Revision of the subgenus Exanthidium with the description of a new species (Apoidea: Megachilidae). Fragmenta entomologica, 53, 333–346. </li> <li>Kasparek, M. (2022) The Resin and Wool Carder Bees (Anthidiini) of Europe and Western Turkey. Identification. Distribution. Biology. Chimaira, Frankfurt, 292 pp. [unchanged reprint 2023] </li> <li>Kasparek, M. & Ebmer, A.W. (2023) The wool carder bee Pseudoanthidium alpinum (Morawitz, 1873): identity of the enigmatic type species of the genus Pseudoanthidium (Hymenoptera: Megachilidae: Anthidiini). Osmia, 11, 39–50. <a href="https://doi.org/10.47446/OSMIA11.7">https://doi.org/10.47446/OSMIA11.7</a> </li> <li>Kasparek, M., Hauser, M. & Schmid-Egger, Ch. (2023) Taxonomic revision of the Afro-Arabian anthidiine bee Pseudoanthidium ochrognathum sensu lato (Apoidea: Anthidiini) based on morphological and genetic data. Integrative Systematics, 6 (2), 55–69. <a href="https://doi.org/10.18476/2023.526995">https://doi.org/10.18476/2023.526995</a> </li> <li>Kasparek, M. & Kuhlmann, M. (2024) Do the members of the rare African bee genus Anthidioma deserve distinct genus status? Linzer biologische Beiträge, 56 (1), 111–120. </li> <li>Kasparek, M. & J. Monks (2024) Not all male bees posses thirteen antennal segments: New megachilid species from the Afro-Arabian and Afrotropical regions (Apoidea: Anthidiini: Pseudoanthidium) with deviant numbers of segments. Zoology in the Middle East, 70, 1–18. </li> <li>Kumar, V., Griswold, T. & Belavadi, V.V. (2017) The resin and carder bees of south India (Hymenoptera: Megachilidae: Anthidiini). Zootaxa, 4317 (3), 436–468. <a href="https://doi.org/10.11646/zootaxa.4317.3.2">https://doi.org/10.11646/zootaxa.4317.3.2</a> </li> <li>Lhomme, P., Michez, D., Christmann, S., Scheuchl, E., El Abdouni, I., Hamroud, L., Ihsane, O., Sentil, A., Smaili, M.C., Schwarz, M., Dathe, H., Straka, J., Pauly, A., Schmid-Egger, Ch., Patiny, S., Müller, A., Praz, Ch., Risch, S., Kasparek, M., Kuhlmann, M., Wood, Th. J., Bogusch, P., Ascher, J. & Rasmont, R. (2020) The wild bees (Hymenoptera: Apoidea) of Morocco. Zootaxa, 4892 (1), 1–159. <a href="https://doi.org/10.11646/zootaxa.4892.1.1">https://doi.org/10.11646/zootaxa.4892.1.1</a> </li> <li>Litman, J. R., Griswold, T. & Danforth, B.N. (2016) Phylogenetic systematics and a revised generic classification of anthidiine bees (Hymenoptera: Megachilidae). Molecular Phylogenetics and Evolution, 100, 183–198. <a href="https://doi.org/10.1016/j.ympev.2016.03.018">https://doi.org/10.1016/j.ympev.2016.03.018</a> </li> <li>Litman, J.R., Fateryga, A.V., Griswold, T.L., Aubert, M., Proshchalykin, M.Yu., Le Divelec, R., Burrows, S. & Praz, Ch.J. (2021) Paraphyly and low levels of genetic divergence in morphologically distinct taxa: revision of the Pseudoanthidium scapulare complex of carder bees (Apoidea: Megachilidae: Anthidiini). Zoological Journal of the Linnean Society, 195, 1287–1337. [2022] <a href="https://doi.org/10.1093/zoolinnean/zlab062">https://doi.org/10.1093/zoolinnean/zlab062</a> </li> <li>Mavromoustakis, G.A. (1948) On some anthidiine bees (Apoidea) from Palestine. Part III. Annals and Magazine of Natural History, Series 11, 14, 428–434. <a href="https://doi.org/10.1080/00222934708654652">https://doi.org/10.1080/00222934708654652</a> </li> <li>Mavromoustakis, G.A. (1954) New and little-known bees of the subfamily Anthidiinae (Apoidea). Part VII. Annals and Magazine for Natural History, Series 12, 7, 249–252. <a href="https://doi.org/10.1080/00222935408651726">https://doi.org/10.1080/00222935408651726</a> </li> <li>Michener, Ch.D. (2007) The Bees of the World. Second edition. The Johns Hopkins University Press, Baltimore, Maryland, 992 pp. </li> <li>Michener, Ch.D. & Griswold, T.L. (1994) The classification of Old World Anthidiini (Hymenoptera, Megachilidae). The University of Kansas Science Bulletin, 55, 299–327. </li> <li>MoA [Ministry of Agriculture and Natural Resources, Department of Agriculture] (1989) Hymenoptera. The systematic list of the Mavromoustakis collection. Nicosia. </li> <li>Mocsáry, A. (1884) Species generis Anthidium Fabr. regionis Palaearcticae. Természetrajzi Füzetek Kiadija a Magyar Nemzeti Müzeum [Naturhistorische Hefte herausgegeben vom Ungarischen National-Museum], 8, 241–278. </li> <li>Morawitz, F.F. (1873 [1874]) Die Bienen Daghestans. Horae Societatis Entomologicae Rossicae (Trudy Russkago éntomologicheskago obshchestva), 10, 129–189. </li> <li>Morawitz, F.F. (1875) A travel to Turkestan by the member-founder of the society A. P. Fedtschenko accomplished from the Imperial Society of Naturalists, Anthropologists, and Etnographists on a Commission from the General-Governor of Turkestan K. P. von Kaufmann (Issue 9). Vol. 11. Zoogeographical Investigations. Pt V. (Division 7). Bees (Mellifera). Izvestiya Imperatorskogo Obshchestva Lyubiteley Estestvoznaniya, Anthropologii i Ethnografii, 19, 1–160. </li> <li>Morawitz, F.F. (1888) Hymenoptera aculeata nova. Horae Societatis Entomologicae Rossicae (Trudy Russkago éntomologicheskago obshchestva), 22, 224–302. </li> <li>Morawitz, F.F. (1894 [1893]) Supplement zur Bienenfauna Turkestans. Horae Societatis Entomologicae Rossicae (Trudy Russkago éntomologicheskago obshchestva), 28, 3–87. </li> <li>Nalinrachatakan, P., Ascher, J.S., Kasparek, M., Traiyasut, P., Thanoosing, Ch. & Warrit, N. (2023) A review of the anthidiine bees (Apoidea, Megachilidae) in Thailand. Zookeys, 1186, 235–284. <a href="https://doi.org/10.3897/zookeys.1186.95203">https://doi.org/10.3897/zookeys.1186.95203</a> </li> <li>Niu, Z.-Q., Luo, A.-R., Griswold, T. & Zhu, C.-D. (2021) Review of the bee genus Pseudoanthidium Friese, 1898 (Hymenoptera: Apoidea: Megachilidae: Anthidiini) of China with descriptions of three new species. Zootaxa, 4996 (1), 133–152. <a href="https://doi.org/10.11646/zootaxa.4996.1.5">https://doi.org/10.11646/zootaxa.4996.1.5</a> </li> <li>Ornosa, C., Ortiz-Sánchez, F.J. & Torres, F. (2008) Catálogo de los Megachilidae del Mediterráneo occidental (Hymenoptera, Apoidea). III. Anthidiini y Dioxyini. Graellsia, 64, 61–86. <a href="https://doi.org/10.3989/graellsia.2008.v64.i1.55">https://doi.org/10.3989/graellsia.2008.v64.i1.55</a> </li> <li>Pagliano, G. (1994) Catalogo degli Imenotteri Italiani. IV. (Apoidea: Colletidae, Andrenidae, Megachilidae, Anthophoridae, Apidae). Memorie della Societa Entomologica Italiana, 72, 331–467. </li> <li>Pasteels, J.J. (1969) La systématique generique et subgenerique des Anthidiinae (Hymenoptera, Apoidea, Megachilidae) de l‘Ancien Monde. Memoires de la Société Royale d‘Entomologie de Belgique, 31, 3–148. </li> <li>Pasteels, J.J. (1984) Révision des Anthidiinae (Hymenoptera, Apoidea, Megachilidae) de l’Afrique subsaharienne. Académie Royale de Belgique, Mémoires de la Classe des Sciences, Collection in-4°-2e Série, 19 (1), 1–165. </li> <li>Pérez, J. (1895) Espèces nouvelles de mellifères de Barbarie (diagnoses préliminaires). G. Gounouihou, Bordeaux, 64 pp </li> <li>Popov, V.B. (1950) Generic groupings of the mid-Asian bees of the subgenus Anthiniinae (Hymenoptera, Megachilidae). Doklady Akademii Nauk SSSR, 70, 315–318. </li> <li>Popov, V.B. (1952) The fauna of bees (Hymenoptera, Apoidea) of south-western Turkmenia and their landscape distribution. Trudy Zoologicheskogo Instituta, Akademiya Nauk SSSR, Leningrad, 10, 61–117. </li> <li>Proshkalykin, M.Yu. & Astafurova, Y.V. (2017) The history of study of the Russian bees (Hymenoptera, Anthophila). A.I. Kurentsov’s Annual Memorial Meetings, 28, 26–34. </li> <li>Proshchalykin, M.Yu. & Fateryga, A.V. (2017) 77. Family Megachilidae. In: Belokobylskij, S.A. & Lelej, A.S. (Eds.), Annotated Catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute of the Russian Academy of Sciences Supplement, 6, pp. 295–308. </li> <li>Rasmussen, C. & Ascher, J.S. (2008) Heinrich Friese (1860–1948): Names proposed and notes on a pioneer melittologist (Hymenoptera, Anthophila). Zootaxa, 1833 (1), 1–118. <a href="https://doi.org/10.11646/zootaxa.1833.1.1">https://doi.org/10.11646/zootaxa.1833.1.1</a> </li> <li>Sandhouse, G.A. (1943) The type species of the genera and subgenera of bees. Proceedings of the United States National Museum, 92, 519–619. <a href="https://doi.org/10.5479/si.00963801.3156.519">https://doi.org/10.5479/si.00963801.3156.519</a> </li> <li>Tamura, K.. Stecher, G. & Kumar, S. (2021) MEGA11: Molecular Evolutionary Genetics Analysis version 11. Molecular Biology and Evolution, 38, 3022–3027. <a href="https://doi.org/10.1093/molbev/msab120)">https://doi.org/10.1093/molbev/msab120)</a> </li> <li>Vereecken, N.J., Ruiz, C., Marshall, L., Pérez-Gil, M., Molenberg, J.-M., Jacobi, B., La Roche, F. & Litman, J.R. (2023) A new small carder bee species from the eastern Canary Islands (Hymenoptera, Megachilidae, Anthidiini). Journal of Hymenoptera Research, 96, 983–1015. <a href="https://doi.org/10.3897/jhr.96.111550">https://doi.org/10.3897/jhr.96.111550</a> </li> <li>Warncke, K. (1980) Die Bienengattung Anthidium Fabricius, 1804 in der Westpaläarktis und im turkestanischen Becken. Entomofauna, 1, 119–209. </li> <li>Warncke, K. (1982) Beitrag zur Bienenfauna des Iran. 15. Die Gattung Anthidium F. Bollettino del Museo Civico di Storia Naturale di Venezia, 32 (1981), 171–196. </li> <li>Zwickl, D.J. & Hillis, D.M. (2002) Increased taxon sampling greatly reduces phylogenetic error. Systematic Biology, 51 (4), 588–598. <a href="https://doi.org/10.1080/10635150290102339">https://doi.org/10.1080/10635150290102339</a> </li> </ol> </div> </div> <div class="jatsParser__right-article-block"> <div class="jatsParser__intraarticle-menu"> <div id="jatsParser__navbar-article" class="jatsParser__navbar"> <nav class="jatsParser__navbar-items" id="jatsParser__navbarItems"> </nav> </div> </div> </div> </div> </div> </main> <footer> <div class="container"> <div class="sidebar row hasUserFooter"> <div class="user_footer col-lg-6 col-md-12"> <div><img src="https://mapress.com/j/public/site/images/mapress/mpicon1.gif" alt="" /> </div> <div>ISSN 1175-5326 (Print Edition) & ISSN 1175-5334 (Online Edition)</div> <div>Published by <a href="https://mapress.com/">Magnolia Press</a>, Auckland, New Zealand</div> </div> <div class="pkp_block block_subscription"> <h2 class="title">Subscription</h2> <div class="content"> <p>Login to access subscriber-only resources.</p> </div> </div> <div class="pkp_block block_custom" id="customblock-scimagojr"> <h2 class="title pkp_screen_reader">scimagojr</h2> <div class="content"> <p><a title="SCImago Journal & Country Rank" href="https://www.scimagojr.com/journalsearch.php?q=4700151916&tip=sid&exact=no"><img src="https://www.scimagojr.com/journal_img.php?id=4700151916" alt="SCImago Journal & Country Rank" border="0"></a></p> </div> </div> </div> <div class="row"> </div> </div> </footer> <script src="https://mapress.com/plugins/themes/oldGregg/resources/jquery/jquery.min.js?v=3.3.0.6" type="text/javascript"></script><script src="https://mapress.com/plugins/themes/oldGregg/resources/bootstrap/js/popper.min.js?v=3.3.0.6" type="text/javascript"></script><script src="https://mapress.com/plugins/themes/oldGregg/resources/bootstrap/js/bootstrap.min.js?v=3.3.0.6" type="text/javascript"></script><script src="https://mapress.com/plugins/themes/oldGregg/resources/js/fontawesome-all.min.js?v=3.3.0.6" type="text/javascript"></script><script src="https://mapress.com/plugins/themes/oldGregg/resources/js/main.js?v=3.3.0.6" type="text/javascript"></script><script src="https://mapress.com/plugins/themes/oldGregg/resources/js/article.js?v=3.3.0.6" type="text/javascript"></script><script type="text/javascript"> (function(i,s,o,g,r,a,m){i['GoogleAnalyticsObject']=r;i[r]=i[r]||function(){ (i[r].q=i[r].q||[]).push(arguments)},i[r].l=1*new Date();a=s.createElement(o), m=s.getElementsByTagName(o)[0];a.async=1;a.src=g;m.parentNode.insertBefore(a,m) })(window,document,'script','https://www.google-analytics.com/analytics.js','ga'); ga('create', 'UA-71881734-1', 'auto'); ga('send', 'pageview'); </script> </body>

CINXE.COM

New species, new synonyms, and resurrected taxa: A review of West and Central Palaearctic members of the genus Pseudoanthidium (Apoidea: Megachilidae) | Zootaxa