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Hopi Hoekstra - Wikipedia

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dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Evolutionary biologist</div> <style data-mw-deduplicate="TemplateStyles:r1257001546">.mw-parser-output .infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox-full-data:not(.notheme)>div:not(.notheme)[style]{background:#1f1f23!important;color:#f8f9fa}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox-full-data:not(.notheme) div:not(.notheme){background:#1f1f23!important;color:#f8f9fa}}@media(min-width:640px){body.skin--responsive .mw-parser-output .infobox-table{display:table!important}body.skin--responsive .mw-parser-output .infobox-table>caption{display:table-caption!important}body.skin--responsive .mw-parser-output .infobox-table>tbody{display:table-row-group}body.skin--responsive .mw-parser-output .infobox-table tr{display:table-row!important}body.skin--responsive .mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox vcard"><tbody><tr><th colspan="2" class="infobox-above" style="font-size: 100%;"><div class="fn" style="font-size:125%;">Hopi Hoekstra</div></th></tr><tr><td colspan="2" class="infobox-full-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1257001546"></td></tr><tr><th colspan="2" class="infobox-header" style="color: #202122; background:lavender;line-height:normal;padding:0.2em;">Dean of the <a href="/wiki/Harvard_Faculty_of_Arts_and_Sciences" title="Harvard Faculty of Arts and Sciences">Harvard Faculty of Arts and Sciences</a></th></tr><tr><td colspan="2" class="infobox-full-data" style="border-bottom:none"><div class="skin-nightmode-reset-color" style="width:100%; margin:0; color: black; background-color: lavender"><b><a href="/wiki/Incumbent" title="Incumbent">Incumbent</a></b></div></td></tr><tr><td colspan="2" class="infobox-full-data" style="border-bottom:none"><span class="nowrap"><b>Assumed office</b></span>&#32;<br />August 1, 2023</td><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1257001546"></tr><tr><th scope="row" class="infobox-label" style="text-align:left"><span class="nowrap">Preceded by</span></th><td class="infobox-data"><a href="/wiki/Claudine_Gay" title="Claudine Gay">Claudine Gay</a></td></tr><tr style="display:none"><td colspan="2"> </td></tr><tr><th colspan="2" class="infobox-header" style="color: #202122; background:lavender">Personal details</th></tr><tr><th scope="row" class="infobox-label">Born</th><td class="infobox-data">July 1972</td></tr><tr><th scope="row" class="infobox-label"><a href="/wiki/Alma_mater" title="Alma mater">Alma mater</a></th><td class="infobox-data"><a href="/wiki/University_of_California,_Berkeley" title="University of California, Berkeley">University of California, Berkeley</a> <span style="font-size:85%;">(B.A.)</span><br /><a href="/wiki/University_of_Washington" title="University of Washington">University of Washington</a> <span style="font-size:85%;">(Ph.D)</span></td></tr><tr><th scope="row" class="infobox-label">Awards</th><td class="infobox-data">National Academy of Sciences (2016)<br />Richard Lounsbery Award (2015)<br /><a href="/wiki/C._Hart_Merriam_Award" title="C. Hart Merriam Award">C. Hart Merriam Award</a> (2019)</td></tr><tr><td colspan="2" class="infobox-below" style="border-top: 1px solid right;"><div></div></td></tr></tbody></table> <p><b>Danielle</b> "<b>Hopi</b>" <b>Elisabeth Hoekstra</b> (born 1972) is an <a href="/wiki/Evolutionary_biology" title="Evolutionary biology">evolutionary biologist</a> working at <a href="/wiki/Harvard_University" title="Harvard University">Harvard University</a> in <a href="/wiki/Cambridge,_Massachusetts" title="Cambridge, Massachusetts">Cambridge, Massachusetts</a>, where she is Dean of the <a href="/wiki/Harvard_Faculty_of_Arts_and_Sciences" title="Harvard Faculty of Arts and Sciences">Harvard Faculty of Arts and Sciences</a>. Her lab uses natural populations of rodents to study the genetic basis of adaptation.<sup id="cite_ref-:0_1-0" class="reference"><a href="#cite_note-:0-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-NYT_2-0" class="reference"><a href="#cite_note-NYT-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-sciencedaily_3-0" class="reference"><a href="#cite_note-sciencedaily-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-PNAS_2015_4-0" class="reference"><a href="#cite_note-PNAS_2015-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> She is the C.Y. Chan Professor of Arts and Sciences and the Xiaomeng Tong and Yu Chen Professor of Life Sciences in the Department of Organismic and Evolutionary Biology and the Department of Molecular and Cellular Biology at Harvard University.<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">&#91;</span>5<span class="cite-bracket">&#93;</span></a></sup> She is also the Curator of Mammals at the Museum of Comparative Zoology and a Harvard College Professor. In 2014, Hoekstra became a <a href="/wiki/Howard_Hughes_Medical_Institute" title="Howard Hughes Medical Institute">Howard Hughes Medical Institute</a> Investigator.<sup id="cite_ref-:0_1-1" class="reference"><a href="#cite_note-:0-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> In 2016, she was elected to the <a href="/wiki/National_Academy_of_Sciences" title="National Academy of Sciences">National Academy of Sciences</a>,<sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">&#91;</span>6<span class="cite-bracket">&#93;</span></a></sup> and in 2017, she was elected to the <a href="/wiki/American_Academy_of_Arts_and_Sciences" title="American Academy of Arts and Sciences">American Academy of Arts and Sciences</a>.<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">&#91;</span>7<span class="cite-bracket">&#93;</span></a></sup> Hoekstra became the Edgerley Family Dean of Harvard's Faculty of Arts and Sciences in August 2023.<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">&#91;</span>8<span class="cite-bracket">&#93;</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Early_life">Early life</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hopi_Hoekstra&amp;action=edit&amp;section=1" title="Edit section: Early life"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Hoekstra was born to a family of Dutch ancestry. Hoekstra's first name "Hopi" is derived from a <a href="/wiki/Dutch_language" title="Dutch language">Dutch</a> term of endearment.<sup id="cite_ref-NYT_2-1" class="reference"><a href="#cite_note-NYT-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-:1_9-0" class="reference"><a href="#cite_note-:1-9"><span class="cite-bracket">&#91;</span>9<span class="cite-bracket">&#93;</span></a></sup> Hoekstra attended Los Altos High School in California. She attended college at the <a href="/wiki/University_of_California,_Berkeley" title="University of California, Berkeley">University of California, Berkeley</a>, where she initially intended to study <a href="/wiki/Political_science" title="Political science">political science</a>. She has stated that at one point she wanted to become the U.S. <a href="/wiki/Ambassador" title="Ambassador">ambassador</a> to the Netherlands, but she was drawn into biology by a class on <a href="/wiki/Biomechanics" title="Biomechanics">biomechanics</a> taught by Robert J. Full. She went on to work in Full's lab, studying the biomechanics of animal locomotion.<sup id="cite_ref-NYT_2-2" class="reference"><a href="#cite_note-NYT-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-:1_9-1" class="reference"><a href="#cite_note-:1-9"><span class="cite-bracket">&#91;</span>9<span class="cite-bracket">&#93;</span></a></sup> One factor for choosing UC Berkeley was that she wanted to play Pac10 volleyball, which she did for two years. </p> <div class="mw-heading mw-heading2"><h2 id="Career">Career</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hopi_Hoekstra&amp;action=edit&amp;section=2" title="Edit section: Career"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Hoekstra received her B.A. in Integrative Biology from the University of California, Berkeley. Before her graduate studies, she researched <a href="/wiki/Grizzly_bear" title="Grizzly bear">grizzly bears</a> for a year in Yellowstone National Park. She obtained her Ph.D. in <a href="/wiki/Zoology" title="Zoology">Zoology</a> as a Howard Hughes Predoctoral Fellow at the <a href="/wiki/University_of_Washington" title="University of Washington">University of Washington</a>.<sup id="cite_ref-:0_1-2" class="reference"><a href="#cite_note-:0-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> For her postdoctoral work, she studied the genetic basis of adaptive melanism in pocket mice at the <a href="/wiki/University_of_Arizona" title="University of Arizona">University of Arizona</a>. In 2003, she became an assistant professor at the <a href="/wiki/University_of_California,_San_Diego" title="University of California, San Diego">University of California, San Diego</a>. In 2007, she moved to <a href="/wiki/Harvard_University" title="Harvard University">Harvard University</a>, where she received tenure in 2010.<sup id="cite_ref-:0_1-3" class="reference"><a href="#cite_note-:0-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-NYT_2-3" class="reference"><a href="#cite_note-NYT-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> She has served on the advisory board of several foundations (e.g. Searle Scholars Program, Max Plank Society), magazines (<i>Scientific American</i>, <i>Quanta</i>) and journals (<i>PNAS</i>, <i>Current Biology</i>, <i>PLoS Genetics</i>, <i>Development</i>, <i>bioRxiv</i>). In June 2023, she was named as the Dean of Harvard's Faculty of Arts and Sciences, succeeding <a href="/wiki/Claudine_Gay" title="Claudine Gay">Claudine Gay</a>, who had assumed the University's presidency a month prior.<sup id="cite_ref-10" class="reference"><a href="#cite_note-10"><span class="cite-bracket">&#91;</span>10<span class="cite-bracket">&#93;</span></a></sup> Hoekstra assumed office on August 1, 2023. </p> <div class="mw-heading mw-heading2"><h2 id="Research">Research</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hopi_Hoekstra&amp;action=edit&amp;section=3" title="Edit section: Research"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Hoekstra spent her scientific career working to understand the fundamental processes by which organisms, including humans, differ in the natural world. She capitalizes on natural variation in non-traditional model organisms, most notably the deer mouse,<sup id="cite_ref-:0_1-4" class="reference"><a href="#cite_note-:0-1"><span class="cite-bracket">&#91;</span>1<span class="cite-bracket">&#93;</span></a></sup> a system she pioneered.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">&#91;</span>11<span class="cite-bracket">&#93;</span></a></sup> Her work is characterized by its an interdisciplinary approach, utilizing both field and lab experiments. Her laboratory’s overarching research strategy is to use tools from genetics, development and neuroscience to discover novel mechanisms by which evolution shapes biodiversity and, conversely, to use biodiversity as a tool to reveal general principles in biology. </p><p>Based on this work, she has been featured in <i>National Geographic</i><sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">&#91;</span>12<span class="cite-bracket">&#93;</span></a></sup> and profiled in the <i>New York Times</i>.<sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">&#91;</span>13<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Behavioral_Genetics">Behavioral Genetics</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hopi_Hoekstra&amp;action=edit&amp;section=4" title="Edit section: Behavioral Genetics"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Hoekstra is best known for studying the genetic mechanisms that influence the evolution of highly complex natural behaviors.<sup id="cite_ref-nytimes2013-01-16_14-0" class="reference"><a href="#cite_note-nytimes2013-01-16-14"><span class="cite-bracket">&#91;</span>14<span class="cite-bracket">&#93;</span></a></sup> In 2013, Hoekstra published an article in the journal <i><a href="/wiki/Nature_(journal)" title="Nature (journal)">Nature</a></i> on the genetics of burrowing behavior in two sister species of <i>Peromyscus</i> mice; the <a href="/wiki/Oldfield_mouse" title="Oldfield mouse">oldfield mouse</a> (<i>P. polionotus</i>), which builds elaborate burrows complete with an escape tunnel, and the deer mouse (<i>P. maniculatis</i>), which builds a simple and shallow nest.<sup id="cite_ref-nytimes2013-01-16_14-1" class="reference"><a href="#cite_note-nytimes2013-01-16-14"><span class="cite-bracket">&#91;</span>14<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-NYT_2-4" class="reference"><a href="#cite_note-NYT-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> Using a combination of behavioral assays and classical genetic strategies, Hoekstra and her students identified four regions of <a href="/wiki/DNA" title="DNA">DNA</a> which control the length of the tunnels dug by the mice.<sup id="cite_ref-nytimes2013-01-16_14-2" class="reference"><a href="#cite_note-nytimes2013-01-16-14"><span class="cite-bracket">&#91;</span>14<span class="cite-bracket">&#93;</span></a></sup> Trainees in her lab have also identified a specific gene that affects parental behavior<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">&#91;</span>15<span class="cite-bracket">&#93;</span></a></sup> and also are genetically dissecting variation in other behaviors such as vocalization and skilled motor behavior. </p> <div class="mw-heading mw-heading3"><h3 id="Color_Adaptation">Color Adaptation</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hopi_Hoekstra&amp;action=edit&amp;section=5" title="Edit section: Color Adaptation"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Hoekstra started her career studying the evolution of mouse fur color and its significance for <a href="/wiki/Adaptation" title="Adaptation">adaptation</a>.<sup id="cite_ref-NYT_2-5" class="reference"><a href="#cite_note-NYT-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> She was among the first to identify a specific DNA mutation and directly link it to fitness in the wild, a result found in many modern textbooks. In 2013, her team published an article in the journal <i><a href="/wiki/Science_(journal)" title="Science (journal)">Science</a></i>, describing how coat color in mice was controlled by nine separate mutations within a single gene, named "agouti."<sup id="cite_ref-sciencedaily_3-1" class="reference"><a href="#cite_note-sciencedaily-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> Speaking about this discovery, Hoekstra said, "The question has always been whether evolution is dominated by these big leaps or smaller steps. When we first implicated the agouti gene, we could have stopped there and concluded that evolution takes these big steps as only one major gene was involved, but that would have been wrong. When we looked more closely, within this gene, we found that even within this single locus, there are, in fact, many small steps."<sup id="cite_ref-sciencedaily_3-2" class="reference"><a href="#cite_note-sciencedaily-3"><span class="cite-bracket">&#91;</span>3<span class="cite-bracket">&#93;</span></a></sup> Her work supports the hypothesis that <a href="/wiki/Evolution" title="Evolution">evolution</a> can occur through incremental changes.<sup id="cite_ref-PNAS_2015_4-1" class="reference"><a href="#cite_note-PNAS_2015-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup> Recently, Hoekstra has found evidence linking the mutation the Agouti gene to survival in mice.<sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">&#91;</span>16<span class="cite-bracket">&#93;</span></a></sup> More specifically, the study showed how a sequence variant in the Agouti gene changes the phenotype and then linked those changes to changes in population allele frequency, demonstrating evolution of trait by natural selection.<sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">&#91;</span>17<span class="cite-bracket">&#93;</span></a></sup> More recently, her lab has discovered the developmental origins of complex color patterns.<sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">&#91;</span>18<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Honors_and_awards">Honors and awards</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hopi_Hoekstra&amp;action=edit&amp;section=6" title="Edit section: Honors and awards"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li>2022 Awarded the <a href="/wiki/Lowell_Thomas_Award" class="mw-redirect" title="Lowell Thomas Award">Lowell Thomas Award</a><sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">&#91;</span>19<span class="cite-bracket">&#93;</span></a></sup></li> <li>2021 Elected as a Fellow to the American Association for the Advancement of Science<sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">&#91;</span>20<span class="cite-bracket">&#93;</span></a></sup></li> <li>2019 Awarded the <a href="/wiki/Hart_Merriam_Award" class="mw-redirect" title="Hart Merriam Award">Hart Merriam Award</a><sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">&#91;</span>21<span class="cite-bracket">&#93;</span></a></sup></li> <li>2018 Elected to the <a href="/wiki/American_Philosophical_Society" title="American Philosophical Society">American Philosophical Society</a><sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">&#91;</span>22<span class="cite-bracket">&#93;</span></a></sup></li> <li>2017 Elected to the <a href="/wiki/American_Academy_of_Arts_and_Sciences" title="American Academy of Arts and Sciences">American Academy of Arts and Sciences</a><sup id="cite_ref-23" class="reference"><a href="#cite_note-23"><span class="cite-bracket">&#91;</span>23<span class="cite-bracket">&#93;</span></a></sup></li> <li>2016 Elected to the <a href="/wiki/National_Academy_of_Sciences" title="National Academy of Sciences">National Academy of Sciences</a><sup id="cite_ref-24" class="reference"><a href="#cite_note-24"><span class="cite-bracket">&#91;</span>24<span class="cite-bracket">&#93;</span></a></sup></li> <li>2015 <a href="/wiki/Richard_Lounsbery_Award" title="Richard Lounsbery Award">Richard Lounsbery Award</a>, <a href="/wiki/National_Academy_of_Sciences" title="National Academy of Sciences">National Academy of Sciences</a><sup id="cite_ref-PNAS_2015_4-2" class="reference"><a href="#cite_note-PNAS_2015-4"><span class="cite-bracket">&#91;</span>4<span class="cite-bracket">&#93;</span></a></sup><sup id="cite_ref-25" class="reference"><a href="#cite_note-25"><span class="cite-bracket">&#91;</span>25<span class="cite-bracket">&#93;</span></a></sup></li> <li>2006 Beckman Young Investigator Award, Arnold and Mabel Beckman Foundation<sup id="cite_ref-26" class="reference"><a href="#cite_note-26"><span class="cite-bracket">&#91;</span>26<span class="cite-bracket">&#93;</span></a></sup></li> <li>2003 Jasper J. Loftus-Hills Young Investigator Prize, <a href="/wiki/American_Society_of_Naturalists" title="American Society of Naturalists">American Society of Naturalists</a><sup id="cite_ref-27" class="reference"><a href="#cite_note-27"><span class="cite-bracket">&#91;</span>27<span class="cite-bracket">&#93;</span></a></sup></li> <li>1998 Ernst Mayr Award, <a href="/wiki/Society_of_Systematic_Biologists" title="Society of Systematic Biologists">Society of Systematic Biologists</a><sup id="cite_ref-Ernst_Mayr_Award_28-0" class="reference"><a href="#cite_note-Ernst_Mayr_Award-28"><span class="cite-bracket">&#91;</span>28<span class="cite-bracket">&#93;</span></a></sup></li></ul> <div class="mw-heading mw-heading2"><h2 id="Family">Family</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hopi_Hoekstra&amp;action=edit&amp;section=7" title="Edit section: Family"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Hoekstra lives in <a href="/wiki/Cambridge,_Massachusetts" title="Cambridge, Massachusetts">Cambridge, Massachusetts</a>, with her son and her husband, <a href="/wiki/James_Mallet" title="James Mallet">James Mallet</a>. Mallet is also an evolutionary biologist at Harvard.<sup id="cite_ref-NYT_2-6" class="reference"><a href="#cite_note-NYT-2"><span class="cite-bracket">&#91;</span>2<span class="cite-bracket">&#93;</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Selected_publications">Selected publications</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hopi_Hoekstra&amp;action=edit&amp;section=8" title="Edit section: Selected publications"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li>Hager, Emily R.; Harringmeyer, Olivia S.; Wooldridge, T. Brock; Theingi, Shunn; Gable, Jacob T.; McFadden, Sade; Neugeboren, Beverly; Turner, Kyle M.; Jensen, Jeffrey D.; Hoekstra, Hopi E. (2022-07-22). "A chromosomal inversion contributes to divergence in multiple traits between deer mouse ecotypes". <i>Science</i>. <b>377</b> (6604): 399–405. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:10.1126/science.abg0718. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a>&#160;1095-9203. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a>&#160;9571565. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;35862520.</li> <li>Barrett, Rowan D. H.; Laurent, Stefan; Mallarino, Ricardo; Pfeifer, Susanne P.; Xu, Charles C. Y.; Foll, Matthieu; Wakamatsu, Kazumasa; Duke-Cohan, Jonathan S.; Jensen, Jeffrey D.; Hoekstra, Hopi E. (2019-02-01). "Linking a mutation to survival in wild mice". <i>Science</i>. <b>363</b> (6426): 499–504. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:10.1126/science.aav3824. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a>&#160;1095-9203. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;30705186.</li> <li>Bendesky, Andres; Kwon, Young-Mi; Lassance, Jean-Marc; Lewarch, Caitlin L.; Yao, Shenqin; Peterson, Brant K.; He, Meng Xiao; Dulac, Catherine; Hoekstra, Hopi E. (2017-04-27). "The genetic basis of parental care evolution in monogamous mice". <i>Nature</i>. <b>544</b> (7651): 434–439. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:10.1038/nature22074. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a>&#160;1476-4687. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a>&#160;5600873. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;28424518.</li> <li>Mallarino, Ricardo; Henegar, Corneliu; Mirasierra, Mercedes; Manceau, Marie; Schradin, Carsten; Vallejo, Mario; Beronja, Slobodan; Barsh, Gregory S.; Hoekstra, Hopi E. (2016-11-24). "Developmental mechanisms of stripe patterns in rodents". <i>Nature</i>. <b>539</b> (7630): 518–523. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:10.1038/nature20109. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a>&#160;1476-4687. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a>&#160;5292240. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;27806375.</li> <li><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFLinnenPohPetersonBarrett2013" class="citation journal cs1">Linnen, C.R.; Poh, Y.-P.; Peterson, B.K.; Barrett, R.D.H.; Larson, J.G; Jensen, J.; Hoekstra, H.E. (2013). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3836219">"Adaptive evolution of multiple traits through multiple mutations at a single gene"</a>. <i>Science</i>. <b>339</b> (6125): 1312–1316. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2013Sci...339.1312L">2013Sci...339.1312L</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.1233213">10.1126/science.1233213</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a>&#160;<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3836219">3836219</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/23493712">23493712</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Science&amp;rft.atitle=Adaptive+evolution+of+multiple+traits+through+multiple+mutations+at+a+single+gene&amp;rft.volume=339&amp;rft.issue=6125&amp;rft.pages=1312-1316&amp;rft.date=2013&amp;rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3836219%23id-name%3DPMC&amp;rft_id=info%3Apmid%2F23493712&amp;rft_id=info%3Adoi%2F10.1126%2Fscience.1233213&amp;rft_id=info%3Abibcode%2F2013Sci...339.1312L&amp;rft.aulast=Linnen&amp;rft.aufirst=C.R.&amp;rft.au=Poh%2C+Y.-P.&amp;rft.au=Peterson%2C+B.K.&amp;rft.au=Barrett%2C+R.D.H.&amp;rft.au=Larson%2C+J.G&amp;rft.au=Jensen%2C+J.&amp;rft.au=Hoekstra%2C+H.E.&amp;rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3836219&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AHopi+Hoekstra" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWeberPetersonHoekstra2013" class="citation journal cs1">Weber, J.N.; Peterson, B.K.; Hoekstra, H.E. (2013). "Discrete genetic modules are responsible for the evolution of complex burrowing behaviour in deer mice". <i>Nature</i>. <b>493</b> (7432): 4202–405. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2Fnature11816">10.1038/nature11816</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/23325221">23325221</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a>&#160;<a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:4361153">4361153</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Nature&amp;rft.atitle=Discrete+genetic+modules+are+responsible+for+the+evolution+of+complex+burrowing+behaviour+in+deer+mice&amp;rft.volume=493&amp;rft.issue=7432&amp;rft.pages=4202-405&amp;rft.date=2013&amp;rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A4361153%23id-name%3DS2CID&amp;rft_id=info%3Apmid%2F23325221&amp;rft_id=info%3Adoi%2F10.1038%2Fnature11816&amp;rft.aulast=Weber&amp;rft.aufirst=J.N.&amp;rft.au=Peterson%2C+B.K.&amp;rft.au=Hoekstra%2C+H.E.&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AHopi+Hoekstra" class="Z3988"></span></li> <li>Manceau, Marie; Domingues, Vera S.; Mallarino, Ricardo; Hoekstra, Hopi E. 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On the origin and spread of an adaptive allele in deer mice. Science. 2009 Aug 28;325(5944):1095-8. <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2736094/">doi: 10.1126/science.1175826</a>. PMID 19713521; PMCID: PMC2736094.</li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFHoekstraHirschmannBundeyInsel2006" class="citation journal cs1">Hoekstra, Hopi E.; Hirschmann, Rachel J.; Bundey, Richard A.; Insel, Paul A.; Crossland, Janet P. (2006). "A single amino acid mutation contributes to adaptive beach mouse color pattern". <i>Science</i>. <b>313</b> (5783): 101–104. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2006Sci...313..101H">2006Sci...313..101H</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.1126121">10.1126/science.1126121</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a>&#160;<a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/16825572">16825572</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a>&#160;<a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:33376626">33376626</a>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=article&amp;rft.jtitle=Science&amp;rft.atitle=A+single+amino+acid+mutation+contributes+to+adaptive+beach+mouse+color+pattern&amp;rft.volume=313&amp;rft.issue=5783&amp;rft.pages=101-104&amp;rft.date=2006&amp;rft_id=info%3Adoi%2F10.1126%2Fscience.1126121&amp;rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A33376626%23id-name%3DS2CID&amp;rft_id=info%3Apmid%2F16825572&amp;rft_id=info%3Abibcode%2F2006Sci...313..101H&amp;rft.aulast=Hoekstra&amp;rft.aufirst=Hopi+E.&amp;rft.au=Hirschmann%2C+Rachel+J.&amp;rft.au=Bundey%2C+Richard+A.&amp;rft.au=Insel%2C+Paul+A.&amp;rft.au=Crossland%2C+Janet+P.&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AHopi+Hoekstra" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFNachmanHoekstraD&#39;Agostino2003" class="citation journal cs1">Nachman, M.W.; Hoekstra, H.E.; D'Agostino, S. L. 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Retrieved <span class="nowrap">2023-11-13</span></span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=unknown&amp;rft.jtitle=www.nasonline.org&amp;rft.atitle=Hopi+E.+Hoekstra&amp;rft_id=https%3A%2F%2Fwww.nasonline.org%2Fmember-directory%2Fmembers%2F20038955.html&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AHopi+Hoekstra" class="Z3988"></span></span> </li> <li id="cite_note-25"><span class="mw-cite-backlink"><b><a href="#cite_ref-25">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.nasonline.org/programs/awards/richard-lounsbery-award.html">"Richard Lounsbery Award"</a>. <i>www.nasonline.org</i><span class="reference-accessdate">. Retrieved <span class="nowrap">2023-11-13</span></span>.</cite><span title="ctx_ver=Z39.88-2004&amp;rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&amp;rft.genre=unknown&amp;rft.jtitle=www.nasonline.org&amp;rft.atitle=Richard+Lounsbery+Award&amp;rft_id=https%3A%2F%2Fwww.nasonline.org%2Fprograms%2Fawards%2Frichard-lounsbery-award.html&amp;rfr_id=info%3Asid%2Fen.wikipedia.org%3AHopi+Hoekstra" class="Z3988"></span></span> </li> <li id="cite_note-26"><span class="mw-cite-backlink"><b><a href="#cite_ref-26">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20180801221136/http://www.beckman-foundation.org/beckman-young-investigators/hopi-hoekstra">"Hopi Hoekstra"</a>. <i>Arnold and Mabel Beckman Foundation</i>. Archived from <a rel="nofollow" class="external text" href="http://www.beckman-foundation.org/beckman-young-investigators/hopi-hoekstra">the original</a> on 1 August 2018<span class="reference-accessdate">. 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style="color:white">Alan Garber</span></a></li> <li><a href="/wiki/Harvard_Board_of_Overseers" title="Harvard Board of Overseers"><span class="tmp-color" style="color:white">Board of Overseers</span></a></li> <li><a href="/wiki/President_and_Fellows_of_Harvard_College" title="President and Fellows of Harvard College"><span class="tmp-color" style="color:white">President and Fellows</span></a></li> <li><span class="tmp-color" style="color:white">Provost</span> <a href="/wiki/John_F._Manning" title="John F. Manning"><span class="tmp-color" style="color:white">John Manning</span></a></li> <li><a href="/wiki/Harvard_Library" title="Harvard Library"><span class="tmp-color" style="color:white">Library</span></a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%"><a href="/wiki/Harvard_Faculty_of_Arts_and_Sciences" title="Harvard Faculty of Arts and Sciences"><span style="color:white">Arts and<br /> Sciences</span></a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Dean <a class="mw-selflink selflink">Hopi Hoekstra</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%"><a href="/wiki/Harvard_College" title="Harvard College"><span style="color:white">College</span></a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Dean <a href="/wiki/Rakesh_Khurana" title="Rakesh Khurana">Rakesh Khurana</a></li> <li><a href="/wiki/Radcliffe_College" title="Radcliffe College">Radcliffe College</a></li></ul> <ul><li><a href="/wiki/List_of_Harvard_College_freshman_dormitories" title="List of Harvard College freshman dormitories">Freshman dormitories</a></li> <li>Upperclass houses <ul><li><a href="/wiki/Adams_House_(Harvard_College)" title="Adams House (Harvard College)">Adams</a></li> <li><a href="/wiki/Cabot_House" title="Cabot House">Cabot</a></li> <li><a href="/wiki/Currier_House_(Harvard_College)" title="Currier House (Harvard College)">Currier</a></li> <li><a href="/wiki/Dudley_Community" title="Dudley Community">Dudley</a></li> <li><a href="/wiki/Dunster_House" title="Dunster House">Dunster</a></li> <li><a href="/wiki/Eliot_House_(Harvard_College)" class="mw-redirect" title="Eliot House (Harvard College)">Eliot</a></li> <li><a href="/wiki/Kirkland_House" title="Kirkland House">Kirkland</a></li> <li><a href="/wiki/Leverett_House" title="Leverett House">Leverett</a></li> <li><a href="/wiki/Lowell_House" title="Lowell House">Lowell</a></li> <li><a href="/wiki/Mather_House_(Harvard_College)" title="Mather House (Harvard College)">Mather</a></li> <li><a href="/wiki/Pforzheimer_House" title="Pforzheimer House">Pforzheimer</a></li> <li><a href="/wiki/Quincy_House_(Harvard_College)" title="Quincy House (Harvard College)">Quincy</a></li> <li><a href="/wiki/Winthrop_House" title="Winthrop House">Winthrop</a></li></ul></li></ul> <ul><li><a href="/wiki/List_of_Harvard_College_undergraduate_organizations" title="List of Harvard College undergraduate organizations">Undergraduate organizations</a></li> <li><a href="/wiki/Hasty_Pudding_Theatricals" title="Hasty Pudding Theatricals">Hasty Pudding Theatricals</a></li> <li><i><a href="/wiki/The_Harvard_Advocate" title="The Harvard Advocate">The Harvard Advocate</a></i></li> <li><i><a href="/wiki/The_Harvard_Crimson" title="The Harvard Crimson">The Harvard Crimson</a></i></li> <li><i><a href="/wiki/The_Harvard_Independent" title="The Harvard Independent">The Harvard Independent</a></i></li> <li><i><a href="/wiki/The_Harvard_Lampoon" title="The Harvard Lampoon">The Harvard Lampoon</a></i></li></ul> <ul><li><a href="/wiki/Harvard_Crimson" title="Harvard Crimson">Athletics</a> <ul><li><a href="/wiki/Harvard_Crimson_baseball" title="Harvard Crimson baseball">Baseball</a></li> <li><a href="/wiki/Harvard_Crimson_men%27s_basketball" title="Harvard Crimson men&#39;s basketball">Men's basketball</a></li> <li><a href="/wiki/Harvard_Crimson_women%27s_basketball" title="Harvard Crimson women&#39;s basketball">Women's basketball</a></li> <li><a href="/wiki/Harvard_Crimson_fencing" title="Harvard Crimson fencing">Fencing</a></li> <li><a href="/wiki/Harvard_Crimson_football" title="Harvard Crimson football">Football</a></li> <li><a href="/wiki/Harvard_Crimson_men%27s_ice_hockey" title="Harvard Crimson men&#39;s ice hockey">Men's ice hockey</a></li> <li><a href="/wiki/Harvard_Crimson_women%27s_ice_hockey" title="Harvard Crimson women&#39;s ice hockey">Women's ice hockey</a></li> <li><a href="/wiki/Harvard_Crimson_men%27s_lacrosse" title="Harvard Crimson men&#39;s lacrosse">Men's lacrosse‎</a></li> <li><a href="/wiki/Harvard_rugby" title="Harvard rugby">Men's rugby</a></li> <li><a href="/wiki/Harvard_women%27s_rugby" title="Harvard women&#39;s rugby">Women's rugby</a></li> <li><a href="/wiki/Harvard_Crimson_men%27s_soccer" title="Harvard Crimson men&#39;s soccer">Men's soccer</a></li> <li><a href="/wiki/Harvard_Crimson_women%27s_soccer" title="Harvard Crimson women&#39;s soccer">Women's soccer</a></li> <li><a href="/wiki/Harvard_Crimson_men%27s_squash" title="Harvard Crimson men&#39;s squash">Men's squash</a></li> <li><a href="/wiki/Harvard_Crimson_men%27s_volleyball" title="Harvard Crimson men&#39;s volleyball">Men's volleyball</a></li> <li><a href="/wiki/Harvard_Crimson_women%27s_volleyball" title="Harvard Crimson women&#39;s volleyball">Women's volleyball</a></li></ul></li> <li><a href="/wiki/Ivy_League" title="Ivy League">Ivy League</a></li> <li><a href="/wiki/Cornell%E2%80%93Harvard_hockey_rivalry" title="Cornell–Harvard hockey rivalry">Cornell hockey rivalry</a></li> <li><a href="/wiki/Harvard%E2%80%93Yale_football_rivalry" title="Harvard–Yale football rivalry">Yale football rivalry</a></li> <li>Stadiums and arenas <ul><li><a href="/wiki/Beanpot_(ice_hockey)" title="Beanpot (ice hockey)">Beanpot</a></li> <li><a href="/wiki/Bright-Landry_Hockey_Center" title="Bright-Landry Hockey Center">Bright-Landry Hockey Center</a></li> <li><a href="/wiki/Harvard_Stadium" title="Harvard Stadium">Harvard Stadium</a></li> <li><a href="/wiki/Jordan_Field" title="Jordan Field">Jordan Field</a></li> <li><a href="/wiki/Lavietes_Pavilion" title="Lavietes Pavilion">Lavietes Pavilion</a></li> <li><a href="/wiki/Ohiri_Field" title="Ohiri Field">Ohiri Field</a></li> <li><a href="/wiki/Malkin_Athletic_Center" title="Malkin Athletic Center">Malkin Athletic Center</a></li></ul></li> <li>Boathouses <ul><li><a href="/wiki/Newell_Boathouse" title="Newell Boathouse">Newell Boathouse</a></li> <li><a href="/wiki/Weld_Boathouse" title="Weld Boathouse">Weld Boathouse</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%"><a href="/wiki/Harvard_Division_of_Continuing_Education" title="Harvard Division of Continuing Education"><span style="color:white">Continuing Education</span></a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Dean Nancy Coleman</li> <li><a href="/wiki/Harvard_Extension_School" title="Harvard Extension School">Extension School</a> <ul><li><a href="/wiki/History_of_Harvard_Extension_School" title="History of Harvard Extension School">Extension School history</a></li></ul></li> <li><a href="/wiki/Harvard_Summer_School" title="Harvard Summer School">Summer School</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%"><a href="/wiki/Harvard_John_A._Paulson_School_of_Engineering_and_Applied_Sciences" title="Harvard John A. Paulson School of Engineering and Applied Sciences"><span style="color:white">Engineering and<br />Applied Sciences</span></a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Dean <a href="/wiki/David_C._Parkes" title="David C. Parkes">David C. Parkes</a></li> <li><a href="/wiki/Lyman_Laboratory_of_Physics" title="Lyman Laboratory of Physics">Lyman Laboratory of Physics</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%"><a href="/wiki/Harvard_Graduate_School_of_Arts_and_Sciences" class="mw-redirect" title="Harvard Graduate School of Arts and Sciences"><span style="color:white">Graduate School</span></a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Dean <a href="/wiki/Emma_Dench" title="Emma Dench">Emma Dench</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%">Libraries</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cabot_Science_Library" title="Cabot Science Library">Cabot</a></li> <li><a href="/wiki/Harvard%E2%80%93Yenching_Library" title="Harvard–Yenching Library">Harvard-Yenching</a></li> <li><a href="/wiki/Houghton_Library" title="Houghton Library">Houghton</a> <ul><li><i><a href="/wiki/Harvard_Review" title="Harvard Review">Harvard Review</a></i></li></ul></li> <li><a href="/wiki/Lamont_Library" title="Lamont Library">Lamont</a></li> <li><a href="/wiki/Pusey_Library" title="Pusey Library">Pusey</a></li> <li><a href="/wiki/Widener_Library" title="Widener Library">Widener</a> <ul><li><a href="/wiki/Harry_Elkins_Widener" title="Harry Elkins Widener">Harry Elkins Widener</a></li> <li><a href="/wiki/Eleanor_Elkins_Widener" title="Eleanor Elkins Widener">Eleanor Elkins Widener</a></li></ul></li> <li><a href="/wiki/Grossman_Library" title="Grossman Library">Grossman</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%">Centers, institutes,<br />and societies</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Harvard_University_Asia_Center" title="Harvard University Asia Center">Asia Center</a></li> <li><a href="/wiki/Hutchins_Center_for_African_and_African_American_Research" title="Hutchins Center for African and African American Research">Center for African and African American Research</a> <ul><li><a href="/wiki/W._E._B._Du_Bois_Research_Institute" title="W. E. B. Du Bois Research Institute">Du Bois Institute</a></li></ul></li> <li><a href="/wiki/Harvard%E2%80%93Smithsonian_Center_for_Astrophysics" title="Harvard–Smithsonian Center for Astrophysics">Center for Astrophysics</a></li> <li><a href="/wiki/Fairbank_Center_for_Chinese_Studies" title="Fairbank Center for Chinese Studies">Center for Chinese Studies</a></li> <li><a href="/wiki/Minda_de_Gunzburg_Center_for_European_Studies" title="Minda de Gunzburg Center for European Studies">Center for European Studies</a></li> <li><a href="/wiki/Carpenter_Center_for_the_Visual_Arts" title="Carpenter Center for the Visual Arts">Center for Visual Arts</a></li> <li><a href="/wiki/Center_for_Hellenic_Studies" title="Center for Hellenic Studies">Center for Hellenic Studies</a></li> <li><a href="/wiki/Dumbarton_Oaks" title="Dumbarton Oaks">Dumbarton Oaks</a></li> <li><a href="/wiki/Harvard_Forest" title="Harvard Forest">Harvard Forest</a></li> <li><a href="/wiki/The_Rowland_Institute_at_Harvard" title="The Rowland Institute at Harvard">Rowland Institute</a></li> <li><a href="/wiki/Harvard_Ukrainian_Research_Institute" title="Harvard Ukrainian Research Institute">Ukrainian Research Institute</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%"><a href="/wiki/Harvard_Business_School" title="Harvard Business School"><span style="color:white">Business</span></a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Dean <a href="/wiki/Srikant_Datar" title="Srikant Datar">Srikant Datar</a></li> <li><a href="/wiki/Baker_Library/Bloomberg_Center" title="Baker Library/Bloomberg Center">Baker Library/Bloomberg Center</a></li> <li><a href="/wiki/Harvard_Business_Publishing" title="Harvard Business Publishing">Harvard Business Publishing</a> <ul><li><i><a href="/wiki/Harvard_Business_Review" title="Harvard Business Review">Harvard Business Review</a></i></li></ul></li> <li><a href="/wiki/Harvard_Business_School_RFC" title="Harvard Business School RFC">Harvard Business School RFC</a></li> <li><a href="/wiki/Spangler_Center" title="Spangler Center">Spangler Center</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%"><a href="/wiki/Harvard_Graduate_School_of_Design" title="Harvard Graduate School of Design"><span style="color:white">Design</span></a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Dean <a href="/wiki/Sarah_Whiting" title="Sarah Whiting">Sarah Whiting</a></li> <li><i><a href="/wiki/Harvard_Design_Magazine" title="Harvard Design Magazine">Harvard Design Magazine</a></i></li> <li><a href="/wiki/Joint_Center_for_Housing_Studies" title="Joint Center for Housing Studies">Center for Housing Studies</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%"><a href="/wiki/Harvard_Divinity_School" title="Harvard Divinity School"><span style="color:white">Divinity</span></a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Dean <a href="/wiki/David_N._Hempton" title="David N. Hempton">David N. Hempton</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%"><a href="/wiki/Harvard_Graduate_School_of_Education" title="Harvard Graduate School of Education"><span style="color:white">Education</span></a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Dean <a href="/wiki/Bridget_Terry_Long" title="Bridget Terry Long">Bridget Terry Long</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%"><a href="/wiki/Harvard_Kennedy_School" title="Harvard Kennedy School"><span style="color:white">Government</span></a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Dean Jeremy M. Weinstein</li> <li><a href="/wiki/Ash_Center_for_Democratic_Governance_and_Innovation" title="Ash Center for Democratic Governance and Innovation">Ash Center</a></li> <li><a href="/wiki/Belfer_Center_for_Science_and_International_Affairs" title="Belfer Center for Science and International Affairs">Belfer Center</a></li> <li><a href="/wiki/Carr_Center_for_Human_Rights_Policy" title="Carr Center for Human Rights Policy">Carr Center</a></li> <li><a href="/wiki/Edmond_%26_Lily_Safra_Center_for_Ethics" title="Edmond &amp; Lily Safra Center for Ethics">Center for Ethics</a></li> <li><a href="/wiki/Joint_Center_for_Housing_Studies" title="Joint Center for Housing Studies">Center for Housing Studies</a></li> <li><a href="/wiki/Hauser_Center_for_Nonprofit_Organizations" title="Hauser Center for Nonprofit Organizations">Center for Nonprofit Organizations</a></li> <li><a href="/wiki/Center_for_Public_Leadership" title="Center for Public Leadership">Center for Public Leadership</a></li> <li><a href="/wiki/Harvard_Institute_of_Politics" title="Harvard Institute of Politics">Institute of Politics</a></li> <li><a href="/wiki/Rappaport_Institute_for_Greater_Boston" title="Rappaport Institute for Greater Boston">Rappaport Institute</a></li> <li><a href="/wiki/Shorenstein_Center_on_Media,_Politics_and_Public_Policy" title="Shorenstein Center on Media, Politics and Public Policy">Shorenstein Center</a> <ul><li><a href="/wiki/The_Journalist%27s_Resource" title="The Journalist&#39;s Resource">The Journalist's Resource</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%"><a href="/wiki/Harvard_Law_School" title="Harvard Law School"><span style="color:white">Law</span></a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Dean <a href="/wiki/John_F._Manning" title="John F. Manning">John F. Manning</a></li> <li><a href="/wiki/Berkman_Klein_Center_for_Internet_%26_Society" title="Berkman Klein Center for Internet &amp; Society">Berkman Klein Center</a></li> <li><i><a href="/wiki/Harvard_Civil_Rights%E2%80%93Civil_Liberties_Law_Review" title="Harvard Civil Rights–Civil Liberties Law Review">Civil Rights-Civil Liberties Law Review</a></i></li> <li><i><a href="/wiki/Harvard_International_Law_Journal" title="Harvard International Law Journal">International Law Journal</a></i></li> <li><i><a href="/wiki/Harvard_Journal_of_Law_%26_Technology" title="Harvard Journal of Law &amp; Technology">Journal of Law &amp; Technology</a></i></li> <li><i><a href="/wiki/Harvard_Journal_on_Legislation" title="Harvard Journal on Legislation">Journal on Legislation</a></i></li> <li><i><a href="/wiki/Harvard_Law_Record" title="Harvard Law Record">Law Record</a></i></li> <li><i><a href="/wiki/Harvard_Law_Review" title="Harvard Law Review">Law Review</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%"><a href="/wiki/Harvard_Medical_School" title="Harvard Medical School"><span style="color:white">Medicine</span></a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Dean <a href="/wiki/George_Q._Daley" title="George Q. Daley">George Q. Daley</a></li> <li><a href="/wiki/Broad_Institute" title="Broad Institute">Broad Institute</a></li> <li><a href="/wiki/Boston_Medical_Library" title="Boston Medical Library">Medical Library</a> <ul><li><a href="/wiki/Warren_Anatomical_Museum" title="Warren Anatomical Museum">Anatomical Museum</a></li></ul></li> <li><a href="/wiki/Harvard%E2%80%93MIT_Program_in_Health_Sciences_and_Technology" title="Harvard–MIT Program in Health Sciences and Technology">Program in Health Sciences</a></li> <li><a href="/wiki/Harvard_School_of_Dental_Medicine" title="Harvard School of Dental Medicine">School of Dental Medicine</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%"><a href="/wiki/Harvard_T.H._Chan_School_of_Public_Health" title="Harvard T.H. Chan School of Public Health"><span style="color:white">Public Health</span></a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>Dean <a href="/wiki/Michelle_Ann_Williams" title="Michelle Ann Williams">Michelle Ann Williams</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%">Museums and<br /> arboretum</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Arnold_Arboretum" title="Arnold Arboretum">Arnold Arboretum</a></li> <li><a href="/wiki/Harvard_Art_Museums" title="Harvard Art Museums">Harvard Art Museums</a></li> <li><a href="/wiki/Peabody_Museum_of_Archaeology_and_Ethnology" title="Peabody Museum of Archaeology and Ethnology">Archaeology and Ethnology Museum</a> <ul><li><a href="/wiki/Busch%E2%80%93Reisinger_Museum" class="mw-redirect" title="Busch–Reisinger Museum">Busch–Reisinger Museum</a></li> <li><a href="/wiki/Fogg_Museum" class="mw-redirect" title="Fogg Museum">Fogg Museum</a></li> <li><a href="/wiki/Arthur_M._Sackler_Museum" class="mw-redirect" title="Arthur M. Sackler Museum">Sackler Museum</a></li></ul></li> <li><a href="/wiki/Harvard_Collection_of_Historical_Scientific_Instruments" title="Harvard Collection of Historical Scientific Instruments">Collection of Scientific Instruments</a> <ul><li><a href="/wiki/Glass_Flowers" title="Glass Flowers">Glass Flowers</a></li> <li><a href="/wiki/Harvard_Mineralogical_Museum" title="Harvard Mineralogical Museum">Mineralogical Museum</a></li></ul></li> <li><a href="/wiki/General_Artemas_Ward_House" title="General Artemas Ward House">General Artemas Ward House</a></li> <li><a href="/wiki/Harvard_Museum_of_Natural_History" title="Harvard Museum of Natural History">Natural History Museum</a> <ul><li><a href="/wiki/Museum_of_Comparative_Zoology" title="Museum of Comparative Zoology">Comparative Zoology Museum</a></li> <li><a href="/wiki/Harvard_University_Herbaria" title="Harvard University Herbaria">Herbaria</a></li></ul></li> <li><a href="/wiki/Harvard_Museum_of_the_Ancient_Near_East" title="Harvard Museum of the Ancient Near East">Museum of the Ancient Near East</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%">Cambridge<br /> campus</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Boylston_Hall_(Harvard_University)" title="Boylston Hall (Harvard University)">Boylston Hall</a></li> <li><a href="/wiki/Robert_Stow_Bradley_Jr._Memorial" title="Robert Stow Bradley Jr. Memorial">Bradley Memorial</a></li> <li><i><a href="/wiki/Discobolus_(Harvard_University)" title="Discobolus (Harvard University)">Discobolus</a></i></li> <li><a href="/wiki/Gates_of_Harvard_Yard" title="Gates of Harvard Yard">Gates of Harvard Yard</a></li> <li><a href="/wiki/Harvard_Graduate_Center" title="Harvard Graduate Center">Graduate Center</a></li> <li><a href="/wiki/Harvard_Hall" title="Harvard Hall">Harvard Hall</a></li> <li><a href="/wiki/Memorial_Church_of_Harvard_University" title="Memorial Church of Harvard University">Memorial Church</a> <ul><li><a href="/wiki/Harvard_University_Choir" title="Harvard University Choir">Choir</a></li></ul></li> <li><a href="/wiki/Harvard_Yard" title="Harvard Yard">Harvard Yard</a></li> <li><a href="/wiki/Massachusetts_Hall_(Harvard_University)" title="Massachusetts Hall (Harvard University)">Massachusetts Hall</a></li> <li><a href="/wiki/Memorial_Hall_(Harvard_University)" title="Memorial Hall (Harvard University)">Memorial Hall</a></li> <li><i><a href="/wiki/Night_Wall_I" title="Night Wall I">Night Wall I</a></i></li> <li><a href="/wiki/Peabody_Terrace" title="Peabody Terrace">Peabody Terrace</a></li> <li><a href="/wiki/President%27s_House_(Harvard)" title="President&#39;s House (Harvard)">President's House</a></li> <li><a href="/wiki/Harvard_Science_Center" title="Harvard Science Center">Science Center</a></li> <li><a href="/wiki/Smith_Campus_Center" title="Smith Campus Center">Smith Campus Center</a></li> <li><a href="/wiki/University_Hall_(Harvard_University)" title="University Hall (Harvard University)">University Hall</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%">Centers and<br /> institutes</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Harvard_Innovation_Labs" title="Harvard Innovation Labs">Harvard Innovation Labs</a></li> <li><a href="/wiki/Massachusetts_Green_High_Performance_Computing_Center" title="Massachusetts Green High Performance Computing Center">Computing Center</a></li> <li><a href="/wiki/Harvard_Radcliffe_Institute" title="Harvard Radcliffe Institute">Harvard Radcliffe Institute</a> <ul><li><a href="/wiki/Schlesinger_Library" title="Schlesinger Library">Schlesinger Library</a></li></ul></li> <li><a href="/wiki/Institute_for_Advanced_Theater_Training" title="Institute for Advanced Theater Training">Institute for Advanced Theater Training</a></li> <li><a href="/wiki/Nieman_Foundation_for_Journalism" title="Nieman Foundation for Journalism">Nieman Foundation for Journalism</a></li> <li><a href="/wiki/Real_Colegio_Complutense" title="Real Colegio Complutense">Real Colegio Complutense</a></li> <li><a href="/wiki/Villa_I_Tatti" title="Villa I Tatti">Villa I Tatti</a></li> <li><a href="/wiki/Wyss_Institute_for_Biologically_Inspired_Engineering" title="Wyss Institute for Biologically Inspired Engineering">Wyss Institute</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%">People</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/List_of_Harvard_University_people" title="List of Harvard University people">List of Harvard University people</a> <ul><li><a href="/wiki/List_of_Harvard_University_non-graduate_alumni" title="List of Harvard University non-graduate alumni">Non-graduate alumni</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="background-color:#A31F36;color:white;box-shadow: inset 2px 2px 0 #2C2A29, inset -2px -2px 0 #2C2A29;;width:1%">Miscellaneous</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Fair_Harvard" title="Fair Harvard">Alma mater</a></li> <li><a href="/wiki/History_and_traditions_of_Harvard_commencements" title="History and traditions of Harvard commencements">Commencement traditions</a> <ul><li><a href="/wiki/Academic_regalia_of_Harvard_University" title="Academic regalia of Harvard University">Academic regalia</a></li></ul></li> <li><a href="/wiki/Harvard_Graduate_Students_Union" title="Harvard Graduate Students Union">Graduate Students Union</a></li> <li><a href="/wiki/Heraldry_of_Harvard_University" title="Heraldry of Harvard University">Heraldry</a></li> <li><a href="/wiki/Harvard_Tercentenary_celebration" title="Harvard Tercentenary celebration">Tercentenary celebration</a></li> <li><i><a href="/wiki/Harvard_Magazine" title="Harvard Magazine">Harvard Magazine</a></i></li> <li><a href="/wiki/Harvard_University_Press" title="Harvard University Press">Harvard University Press</a></li> <li><a href="/wiki/Lionel_de_Jersey_Harvard" title="Lionel de Jersey Harvard">Lionel de Jersey Harvard</a></li> <li><a href="/wiki/Math_55" title="Math 55">Math 55</a></li> <li><a href="/wiki/Harvard_Society_of_Fellows" title="Harvard Society of Fellows">Society of Fellows</a></li> <li><a href="/wiki/List_of_University_Professors_at_Harvard_University" 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