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But the title continues, …or the preservation of favoured races in the struggle for life. This is more ambiguous, especially to the modern reader, for" /> <meta name="generator" content="Drupal 7 (https://www.drupal.org)" /> <link rel="canonical" href="https://www.darwinproject.ac.uk/commentary/life-sciences/species-and-varieties" /> <link rel="shortlink" href="https://www.darwinproject.ac.uk/node/1096" /> <meta property="og:site_name" content="Darwin Correspondence Project" /> <meta property="og:type" content="article" /> <meta property="og:url" content="https://www.darwinproject.ac.uk/commentary/life-sciences/species-and-varieties" /> <meta property="og:title" content="Species and varieties" /> <meta property="og:description" content="On the origin of species by means of natural selection …so begins the title of Darwin’s most famous book, and the reader would rightly assume that such a thing as ‘species’ must therefore exist and be subject to description. But the title continues, …or the preservation of favoured races in the struggle for life. This is more ambiguous, especially to the modern reader, for whom race carries a different and highly charged meaning. In the context of natural history, when Darwin used the term here, he simply meant ‘variety’, as in ‘a fast-growing race of wheat’." /> <meta property="og:updated_time" content="2022-11-23T10:20:17+00:00" /> <meta property="og:image" content="https://www.darwinproject.ac.uk/sites/default/files/Origin_1859_Title%20page_c.jpg" /> <meta name="twitter:card" content="summary" /> <meta name="twitter:url" content="https://www.darwinproject.ac.uk/commentary/life-sciences/species-and-varieties" /> <meta name="twitter:title" content="Species and varieties" /> <meta property="article:published_time" content="2022-11-18T17:08:59+00:00" /> <meta property="article:modified_time" content="2022-11-18T17:08:59+00:00" /> <title>Species and varieties | Darwin Correspondence Project</title> <meta name="viewport" content="width=device-width, initial-scale=1.0"/> <style type="text/css" media="all"> @import url("https://www.darwinproject.ac.uk/modules/system/system.base.css?sh7yfi"); @import 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easy-breadcrumb_segment-title" itemprop="title">Species and varieties</span></li> </ul> </div> </div> </div> </div> <h1 class="campl-page-title">Darwin Correspondence Project</h1> </div> </div> </div> </div> <div class="campl-row campl-page-header"> <div class="campl-wrap"> <div class="region region-horizontal-navigation"> <div id="block-menu-block-1" class="block block-menu-block"> <div> <div class="menu-block-wrapper menu-block-1 menu-name-main-menu parent-mlid-0 menu-level-1"> <div class="campl-wrap clearfix campl-local-navigation"><div class="campl-local-navigation-container"><ul class="campl-unstyled-list"><li class="first leaf menu-mlid-198"><a href="/">Home</a></li> <li class="expanded menu-mlid-2309"><a href="/about-darwin">About Darwin</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-2309"><a href="/about-darwin">About Darwin overview</a></li> <li class="expanded menu-mlid-869"><a href="/people/about-darwin/family-life">Family life</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-869"><a href="/people/about-darwin/family-life">Family life overview</a></li> <li class="leaf menu-mlid-1125"><a href="/people/about-darwin/family-life/darwin-childhood">Darwin on childhood</a></li> <li class="leaf menu-mlid-1038"><a href="/tags/about-darwin/family-life/darwin-marriage">Darwin on marriage</a></li> <li class="leaf menu-mlid-1258"><a href="/people/about-darwin/family-life/darwin-s-observations-his-children">Darwin’s observations on his children</a></li> <li class="leaf menu-mlid-2295"><a href="/people/about-darwin/family-life/darwin-and-fatherhood">Darwin and fatherhood</a></li> <li class="leaf menu-mlid-1039"><a href="/people/about-darwin/family-life/death-anne-elizabeth-darwin">The death of Annie Darwin</a></li> <li class="last leaf menu-mlid-1051"><a href="/people/about-darwin/family-life/visiting-darwins">Visiting the Darwins</a></li> </ul></li> <li class="leaf menu-mlid-3416"><a href="/commentary/voyage-hms-beagle" title="">Voyage of HMS Beagle</a></li> <li class="expanded menu-mlid-1035"><a href="/people/about-darwin/what-darwin-read">What Darwin read</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-1035"><a href="/people/about-darwin/what-darwin-read">What Darwin read overview</a></li> <li class="leaf menu-mlid-1130"><a href="/people/about-darwin/what-darwin-read/darwin-s-student-booklist">Darwin’s student booklist</a></li> <li class="leaf menu-mlid-933"><a href="/people/about-darwin/what-darwin-read/books-beagle">Books on the Beagle</a></li> <li class="last leaf menu-mlid-1036"><a href="/people/about-darwin/what-darwin-read/darwin-s-reading-notebooks">Darwin’s reading notebooks</a></li> </ul></li> <li class="expanded menu-mlid-1059"><a href="/about-darwin/origin-species">On the Origin of Species</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-1059"><a href="/about-darwin/origin-species">On the Origin of Species overview</a></li> <li class="leaf menu-mlid-1060"><a href="/people/about-darwin/origin-species/writing-origin">The writing of "Origin"</a></li> <li class="leaf menu-mlid-1084"><a href="/people/about-darwin/origin-species/abstract-darwin-s-theory">Abstract of Darwin’s theory</a></li> <li class="leaf menu-mlid-1093"><a href="/people/about-darwin/origin-species/alfred-russel-wallace-s-essay-varieties">Alfred Russel Wallace’s essay on varieties</a></li> <li class="leaf menu-mlid-1094"><a href="/charles-darwin-and-his-publisher">Charles Darwin and his publisher</a></li> <li class="last leaf menu-mlid-1147"><a href="/people/about-darwin/origin-species/review-origin-species">Review: The Origin of Species</a></li> </ul></li> <li class="leaf menu-mlid-1126"><a href="/people/about-darwin/darwins-health">Darwin’s health</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first last leaf menu-mlid-2917"><a href="/tags/darwin/darwin-on-his-health">Darwin's notes for his physician, 1865</a></li> </ul></li> <li class="leaf menu-mlid-1128"><a href="/about-darwin/darwin-s-photographic-portraits">Darwin’s photographic portraits</a></li> <li class="leaf menu-mlid-2896"><a href="/have-you-read-one-about">Have you read the one about....</a></li> <li class="expanded menu-mlid-2654"><a href="/people/about-darwin/six-things-darwin-never-said">Six things Darwin never said – and one he did</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-2654"><a href="/people/about-darwin/six-things-darwin-never-said">Six things Darwin never said – and one he did overview</a></li> <li class="last leaf menu-mlid-2892"><a href="/people/about-darwin/six-things-darwin-never-said/evolution-misquotation">The evolution of a misquotation</a></li> </ul></li> <li class="expanded menu-mlid-3673"><a href="/about-darwin/portraits-charles-darwin-catalogue">Portraits of Charles Darwin: a catalogue</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-3673"><a href="/about-darwin/portraits-charles-darwin-catalogue">Portraits of Charles Darwin: a catalogue overview</a></li> <li class="leaf menu-mlid-3676"><a href="/about-darwin/portraits-charles-darwin-catalogue/11-ellen-sharples-pastel">1.1 Ellen Sharples pastel</a></li> <li class="leaf menu-mlid-3709"><a href="/about-darwin/portraits-charles-darwin-catalogue/12-george-richmond-marriage-portrait">1.2 George Richmond, marriage portrait</a></li> <li class="leaf menu-mlid-3718"><a href="/about-darwin/portraits-charles-darwin-catalogue/13-thomas-herbert-maguire-lithograph">1.3 Thomas Herbert Maguire, lithograph</a></li> <li class="leaf menu-mlid-3721"><a href="/about-darwin/portraits-charles-darwin-catalogue/14-samuel-laurence-drawing-1">1.4 Samuel Laurence drawing 1</a></li> <li class="leaf menu-mlid-3724"><a href="/about-darwin/portraits-charles-darwin-catalogue/15-samuel-laurence-drawing-2">1.5 Samuel Laurence drawing 2</a></li> <li class="leaf menu-mlid-3727"><a href="/about-darwin/portraits-charles-darwin-catalogue/16-ouless-oil-portrait">1.6 Ouless oil portrait</a></li> <li class="leaf menu-mlid-3730"><a href="/about-darwin/portraits-charles-darwin-catalogue/17-ouless-replica">1.7 Ouless replica</a></li> <li class="leaf menu-mlid-3733"><a href="/about-darwin/portraits-charles-darwin-catalogue/18-anonymous-drawing-after-ouless">1.8 anonymous drawing, after Ouless</a></li> <li class="leaf menu-mlid-3736"><a href="/about-darwin/portraits-charles-darwin-catalogue/19-rajon-etching-after-ouless">1.9 Rajon, etching after Ouless</a></li> <li class="leaf menu-mlid-3679"><a href="/about-darwin/portraits-charles-darwin-catalogue/110-rajon-etching-variant-state">1.10 Rajon etching, variant state</a></li> <li class="leaf menu-mlid-3682"><a href="/about-darwin/portraits-charles-darwin-catalogue/111-laura-russell-oil">1.11 Laura Russell, oil</a></li> <li class="leaf menu-mlid-3685"><a href="/about-darwin/portraits-charles-darwin-catalogue/112-marian-huxley-drawing">1.12 Marian Huxley, drawing</a></li> <li class="leaf menu-mlid-3688"><a href="/about-darwin/portraits-charles-darwin-catalogue/113-louisa-nash-drawing">1.13 Louisa Nash, drawing</a></li> <li class="leaf menu-mlid-3691"><a href="/about-darwin/portraits-charles-darwin-catalogue/114-william-richmond-oil">1.14 William Richmond, oil</a></li> <li class="leaf menu-mlid-3694"><a href="/about-darwin/portraits-charles-darwin-catalogue/115-albert-goodwin-watercolour">1.15 Albert Goodwin, watercolour</a></li> <li class="leaf menu-mlid-3697"><a href="/about-darwin/portraits-charles-darwin-catalogue/116-alphonse-legros-drypoint">1.16 Alphonse Legros, drypoint</a></li> <li class="leaf menu-mlid-3700"><a href="/about-darwin/portraits-charles-darwin-catalogue/117-alphonse-legros-drawing">1.17 Alphonse Legros drawing</a></li> <li class="leaf menu-mlid-3703"><a href="/about-darwin/portraits-charles-darwin-catalogue/118-john-collier-oil-linnean">1.18 John Collier, oil in Linnean</a></li> <li class="leaf menu-mlid-3706"><a href="/about-darwin/portraits-charles-darwin-catalogue/119-john-collier-oil-npg">1.19 John Collier, oil in NPG</a></li> <li class="leaf menu-mlid-3712"><a href="/about-darwin/portraits-charles-darwin-catalogue/120-leopold-flameng-etching-after-collier">1.20 Leopold Flameng etching, after Collier</a></li> <li class="leaf menu-mlid-3715"><a href="/about-darwin/portraits-charles-darwin-catalogue/121-window-christs-college-cambridge">1.21 window at Christ's College Cambridge</a></li> <li class="leaf menu-mlid-3739"><a href="/about-darwin/portraits-charles-darwin-catalogue/21-thomas-woolner-bust">2.1 Thomas Woolner bust</a></li> <li class="leaf menu-mlid-3772"><a href="/about-darwin/portraits-charles-darwin-catalogue/22-thomas-woolner-metal-plaque">2.2 Thomas Woolner metal plaque</a></li> <li class="leaf menu-mlid-3802"><a href="/about-darwin/portraits-charles-darwin-catalogue/23-wedgwood-medallions">2.3 Wedgwood medallions</a></li> <li class="leaf menu-mlid-3805"><a href="/about-darwin/portraits-charles-darwin-catalogue/24-wedgwood-plaque">2.4 Wedgwood plaque</a></li> <li class="leaf menu-mlid-3808"><a href="/about-darwin/portraits-charles-darwin-catalogue/25-wedgwood-medallions-2nd-type">2.5 Wedgwood medallions, 2nd type</a></li> <li class="leaf menu-mlid-3811"><a href="/about-darwin/portraits-charles-darwin-catalogue/26-adolf-von-hildebrand-bust">2.6 Adolf von Hildebrand bust</a></li> <li class="leaf menu-mlid-3814"><a href="/about-darwin/portraits-charles-darwin-catalogue/27-joseph-moore-midland-union-medal">2.7 Joseph Moore, Midland Union medal</a></li> <li class="leaf menu-mlid-3817"><a href="/about-darwin/portraits-charles-darwin-catalogue/28-alphonse-legros-medallion">2.8 Alphonse Legros medallion</a></li> <li class="leaf menu-mlid-3820"><a href="/about-darwin/portraits-charles-darwin-catalogue/29-legros-medallion-plaster-model">2.9 Legros medallion, plaster model</a></li> <li class="leaf menu-mlid-3742"><a href="/about-darwin/portraits-charles-darwin-catalogue/210-moritz-klinkicht-print-legros">2.10 Moritz Klinkicht, print from Legros</a></li> <li class="leaf menu-mlid-3745"><a href="/about-darwin/portraits-charles-darwin-catalogue/211-christian-lehr-plaster-bust">2.11 Christian Lehr, plaster bust</a></li> <li class="leaf menu-mlid-3748"><a href="/about-darwin/portraits-charles-darwin-catalogue/212-allan-wyon-royal-society-medal">2.12 Allan Wyon, Royal Society medal</a></li> <li class="leaf menu-mlid-3751"><a href="/about-darwin/portraits-charles-darwin-catalogue/213-edgar-boehm-statue-nhm">2.13 Edgar Boehm, statue in the NHM</a></li> <li class="leaf menu-mlid-3754"><a href="/about-darwin/portraits-charles-darwin-catalogue/214-boehm-westminster-abbey-roundel">2.14 Boehm, Westminster Abbey roundel</a></li> <li class="leaf menu-mlid-3757"><a href="/about-darwin/portraits-charles-darwin-catalogue/215-boehm-terracotta-bust-npg">2.15 Boehm terracotta bust (NPG)</a></li> <li class="leaf menu-mlid-3760"><a href="/about-darwin/portraits-charles-darwin-catalogue/216-horace-montford-statue-shrewsbury">2.16 Horace Montford statue, Shrewsbury</a></li> <li class="leaf menu-mlid-3763"><a href="/about-darwin/portraits-charles-darwin-catalogue/217-montford-statuette">2.17 Montford, statuette</a></li> <li class="leaf menu-mlid-3766"><a href="/about-darwin/portraits-charles-darwin-catalogue/218-montford-carnegie-bust">2.18 Montford, Carnegie bust</a></li> <li class="leaf menu-mlid-3769"><a href="/about-darwin/portraits-charles-darwin-catalogue/219-montford-bust-royal-society">2.19 Montford, bust at the Royal Society</a></li> <li class="leaf menu-mlid-3775"><a href="/about-darwin/portraits-charles-darwin-catalogue/220-montford-terracotta-bust-npg">2.20 Montford, terracotta bust, NPG</a></li> <li class="leaf menu-mlid-3778"><a href="/about-darwin/portraits-charles-darwin-catalogue/221-montford-relief-christs-college">2.21 Montford, relief at Christ's College</a></li> <li class="leaf menu-mlid-3781"><a href="/about-darwin/portraits-charles-darwin-catalogue/222-l-j-chavalliaud-statue-liverpool">2.22 L.-J. Chavalliaud statue in Liverpool</a></li> <li class="leaf menu-mlid-3784"><a href="/about-darwin/portraits-charles-darwin-catalogue/223-hope-pinker-statue-oxford-museum">2.23 Hope Pinker statue, Oxford Museum</a></li> <li class="leaf menu-mlid-3787"><a href="/about-darwin/portraits-charles-darwin-catalogue/224-herbert-hampton-statue-lancaster">2.24 Herbert Hampton statue, Lancaster</a></li> <li class="leaf menu-mlid-3790"><a href="/about-darwin/portraits-charles-darwin-catalogue/225-henry-pegram-statue-birmingham">2.25 Henry Pegram statue, Birmingham</a></li> <li class="leaf menu-mlid-3793"><a href="/about-darwin/portraits-charles-darwin-catalogue/226-linnean-society-medal">2.26 Linnean Society medal</a></li> <li class="leaf menu-mlid-3796"><a href="/about-darwin/portraits-charles-darwin-catalogue/227-william-couper-bust-new-york">2.27 William Couper bust, New York</a></li> <li class="leaf menu-mlid-3799"><a href="/about-darwin/portraits-charles-darwin-catalogue/228-couper-bust-cambridge">2.28 Couper bust in Cambridge</a></li> <li class="leaf menu-mlid-3823"><a href="/about-darwin/portraits-charles-darwin-catalogue/31-antoine-claudet-daguerreotype">3.1 Antoine Claudet, daguerreotype</a></li> <li class="leaf menu-mlid-3856"><a href="/about-darwin/portraits-charles-darwin-catalogue/32-maull-and-polyblank-photo-1">3.2 Maull and Polyblank photo 1</a></li> <li class="leaf menu-mlid-3865"><a href="/about-darwin/portraits-charles-darwin-catalogue/33-maull-and-polyblank-photo-2">3.3 Maull and Polyblank photo 2</a></li> <li class="leaf menu-mlid-3868"><a href="/about-darwin/portraits-charles-darwin-catalogue/34-william-darwin-photo-1">3.4 William Darwin, photo 1</a></li> <li class="leaf menu-mlid-3871"><a href="/about-darwin/portraits-charles-darwin-catalogue/35-william-darwin-photo-2">3.5 William Darwin, photo 2</a></li> <li class="leaf menu-mlid-3874"><a href="/about-darwin/portraits-charles-darwin-catalogue/36-william-darwin-photo-3">3.6 William Darwin, photo 3</a></li> <li class="leaf menu-mlid-3877"><a href="/about-darwin/portraits-charles-darwin-catalogue/37-leonard-darwin-photo-verandah">3.7 Leonard Darwin, photo on verandah</a></li> <li class="leaf menu-mlid-3880"><a href="/about-darwin/portraits-charles-darwin-catalogue/38-leonard-darwin-interior-photo">3.8 Leonard Darwin, interior photo</a></li> <li class="leaf menu-mlid-3883"><a href="/about-darwin/portraits-charles-darwin-catalogue/39-leonard-darwin-photo-horseback">3.9 Leonard Darwin, photo on horseback</a></li> <li class="leaf menu-mlid-3826"><a href="/about-darwin/portraits-charles-darwin-catalogue/310-ernest-edwards-men-eminence">3.10 Ernest Edwards, 'Men of Eminence'</a></li> <li class="leaf menu-mlid-3829"><a href="/about-darwin/portraits-charles-darwin-catalogue/311-edwards-illustrated-london-news">3.11 Edwards, in Illustrated London News</a></li> <li class="leaf menu-mlid-3832"><a href="/about-darwin/portraits-charles-darwin-catalogue/312-edwards-second-group-photos">3.12 Edwards, second group of photos</a></li> <li class="leaf menu-mlid-3835"><a href="/about-darwin/portraits-charles-darwin-catalogue/313-edwards-representative-men">3.13 Edwards 'Representative Men'</a></li> <li class="leaf menu-mlid-3838"><a href="/about-darwin/portraits-charles-darwin-catalogue/314-julia-margaret-cameron-photos">3.14 Julia Margaret Cameron, photos</a></li> <li class="leaf menu-mlid-3841"><a href="/about-darwin/portraits-charles-darwin-catalogue/315-george-charles-wallich-photo">3.15 George Charles Wallich, photo</a></li> <li class="leaf menu-mlid-3844"><a href="/about-darwin/portraits-charles-darwin-catalogue/316-oscar-rejlander-photos">3.16 Oscar Rejlander, photos</a></li> <li class="leaf menu-mlid-3847"><a href="/about-darwin/portraits-charles-darwin-catalogue/317-lock-and-whitfield-men-mark">3.17 Lock and Whitfield, 'Men of Mark'</a></li> <li class="leaf menu-mlid-3850"><a href="/about-darwin/portraits-charles-darwin-catalogue/318-elliott-and-fry-photos-c1869-1871">3.18 Elliott and Fry photos, c.1869-1871</a></li> <li class="leaf menu-mlid-3853"><a href="/about-darwin/portraits-charles-darwin-catalogue/319-elliott-and-fry-photos-c1880-1">3.19 Elliott and Fry photos c.1880-1</a></li> <li class="leaf menu-mlid-3859"><a href="/about-darwin/portraits-charles-darwin-catalogue/320-elliott-and-fry-c1880-1-verandah">3.20 Elliott and Fry, c.1880-1, verandah</a></li> <li class="leaf menu-mlid-3862"><a href="/about-darwin/portraits-charles-darwin-catalogue/321-herbert-rose-barraud-photos">3.21 Herbert Rose Barraud, photos</a></li> <li class="leaf menu-mlid-3886"><a href="/about-darwin/portraits-charles-darwin-catalogue/41-albert-way-comic-drawings">4.1 Albert Way, comic drawings</a></li> <li class="leaf menu-mlid-3919"><a href="/about-darwin/portraits-charles-darwin-catalogue/42-augustus-earle-caricature-drawing">4.2 Augustus Earle, caricature drawing</a></li> <li class="leaf menu-mlid-3952"><a href="/about-darwin/portraits-charles-darwin-catalogue/43-alfred-crowquill-caricature">4.3 Alfred Crowquill, caricature</a></li> <li class="leaf menu-mlid-3985"><a href="/about-darwin/portraits-charles-darwin-catalogue/44-thomas-huxley-caricature-sketch">4.4 Thomas Huxley, caricature sketch</a></li> <li class="leaf menu-mlid-4018"><a href="/about-darwin/portraits-charles-darwin-catalogue/45-william-beard-comic-painting">4.5 William Beard, comic painting</a></li> <li class="leaf menu-mlid-4051"><a href="/about-darwin/portraits-charles-darwin-catalogue/46-thomas-nast-cartoon">4.6 Thomas Nast, cartoon</a></li> <li class="leaf menu-mlid-4054"><a href="/about-darwin/portraits-charles-darwin-catalogue/47-vanity-fair-caricature">4.7 'Vanity Fair', caricature</a></li> <li class="leaf menu-mlid-4057"><a href="/about-darwin/portraits-charles-darwin-catalogue/48-vanity-fair-preliminary-study">4.8 'Vanity Fair', preliminary study</a></li> <li class="leaf menu-mlid-4060"><a href="/about-darwin/portraits-charles-darwin-catalogue/49-graphic-cartoon">4.9 'Graphic', cartoon</a></li> <li class="leaf menu-mlid-3889"><a href="/about-darwin/portraits-charles-darwin-catalogue/410-hornet-caricature-darwin">4.10 'Hornet' caricature of Darwin</a></li> <li class="leaf menu-mlid-3892"><a href="/about-darwin/portraits-charles-darwin-catalogue/411-fun-cartoon-little-lecture">4.11 'Fun' cartoon, 'A little lecture'</a></li> <li class="leaf menu-mlid-3895"><a href="/about-darwin/portraits-charles-darwin-catalogue/412-fun-wedding-procession">4.12 'Fun', Wedding procession</a></li> <li class="leaf menu-mlid-3898"><a href="/about-darwin/portraits-charles-darwin-catalogue/413-fun-cartoon-griset-emotional">4.13 'Fun' cartoon by Griset, 'Emotional'</a></li> <li class="leaf menu-mlid-3901"><a href="/about-darwin/portraits-charles-darwin-catalogue/414-fun-cartoon-troubles">4.14 'Fun' cartoon, 'That troubles'</a></li> <li class="leaf menu-mlid-3904"><a href="/about-darwin/portraits-charles-darwin-catalogue/415-george-cruikshank-comic-drawing">4.15 George Cruikshank, comic drawing</a></li> <li class="leaf menu-mlid-3907"><a href="/about-darwin/portraits-charles-darwin-catalogue/416-joseph-simms-physiognomy">4.16 Joseph Simms, physiognomy</a></li> <li class="leaf menu-mlid-3910"><a href="/about-darwin/portraits-charles-darwin-catalogue/417-figaro-unidentifiable-1871">4.17 'Figaro', unidentifiable 1871</a></li> <li class="leaf menu-mlid-3913"><a href="/about-darwin/portraits-charles-darwin-catalogue/418-figaro-chromolithograph-1">4.18 'Figaro' chromolithograph 1</a></li> <li class="leaf menu-mlid-3916"><a href="/about-darwin/portraits-charles-darwin-catalogue/419-george-montbard-caricature">4.19 George Montbard, caricature</a></li> <li class="leaf menu-mlid-3922"><a href="/about-darwin/portraits-charles-darwin-catalogue/420-frederick-waddy-caricature">4.20 Frederick Waddy, caricature</a></li> <li class="leaf menu-mlid-3925"><a href="/about-darwin/portraits-charles-darwin-catalogue/421-gegeef-our-national-church-1">4.21 Gegeef, 'Our National Church', 1</a></li> <li class="leaf menu-mlid-3928"><a href="/about-darwin/portraits-charles-darwin-catalogue/422-gegeef-et-al-our-national-church-2">4.22 Gegeef et al., 'Our National Church', 2</a></li> <li class="leaf menu-mlid-3931"><a href="/about-darwin/portraits-charles-darwin-catalogue/423-gegeef-battle-field-science">4.23 Gegeef, 'Battle Field of Science'</a></li> <li class="leaf menu-mlid-3934"><a href="/about-darwin/portraits-charles-darwin-catalogue/424-daily-graphic-nast-satire">4.24 'Daily Graphic', Nast satire</a></li> <li class="leaf menu-mlid-3937"><a href="/about-darwin/portraits-charles-darwin-catalogue/425-punch-1877-re-cambridge-doctorate">4.25 'Punch' 1877 re. Cambridge doctorate</a></li> <li class="leaf menu-mlid-3940"><a href="/about-darwin/portraits-charles-darwin-catalogue/426-christmas-card-caricature-monkeys">4.26 Christmas card caricature, monkeys</a></li> <li class="leaf menu-mlid-3943"><a href="/about-darwin/portraits-charles-darwin-catalogue/427-four-founders-darwinismus">4.27 'Four founders of Darwinismus'</a></li> <li class="leaf menu-mlid-3946"><a href="/about-darwin/portraits-charles-darwin-catalogue/428-english-celebrities-montage">4.28 'English celebrities' montage</a></li> <li class="leaf menu-mlid-3949"><a href="/about-darwin/portraits-charles-darwin-catalogue/429-richard-grant-white-fall-man">4.29 Richard Grant White, 'Fall of man'</a></li> <li class="leaf menu-mlid-3955"><a href="/about-darwin/portraits-charles-darwin-catalogue/430-la-petite-lune-gill-cartoon">4.30 'La Petite Lune', Gill cartoon</a></li> <li class="leaf menu-mlid-3958"><a href="/about-darwin/portraits-charles-darwin-catalogue/431-la-lune-rousse-gill-cartoon">4.31 'La Lune Rousse', Gill cartoon</a></li> <li class="leaf menu-mlid-3961"><a href="/about-darwin/portraits-charles-darwin-catalogue/432-anis-liqueur-label">4.32 Anis liqueur label</a></li> <li class="leaf menu-mlid-3964"><a href="/about-darwin/portraits-charles-darwin-catalogue/433-harpers-weekly-bellew-caricature">4.33 'Harper's Weekly', Bellew caricature</a></li> <li class="leaf menu-mlid-3967"><a href="/about-darwin/portraits-charles-darwin-catalogue/434-punch-sambourne-cartoon-1">4.34 'Punch', Sambourne cartoon 1</a></li> <li class="leaf menu-mlid-3970"><a href="/about-darwin/portraits-charles-darwin-catalogue/435-frederick-sem-caricature">4.35 Frederick Sem, caricature</a></li> <li class="leaf menu-mlid-3973"><a href="/about-darwin/portraits-charles-darwin-catalogue/436-sem-chistmas-card">4.36 Sem, Chistmas card</a></li> <li class="leaf menu-mlid-3976"><a href="/about-darwin/portraits-charles-darwin-catalogue/437-mosquito-satire">4.37 'Mosquito' satire</a></li> <li class="leaf menu-mlid-3979"><a href="/about-darwin/portraits-charles-darwin-catalogue/438-franz-goedecker-caricature">4.38 Franz Goedecker, caricature</a></li> <li class="leaf menu-mlid-3982"><a href="/about-darwin/portraits-charles-darwin-catalogue/439-moonshine-magazine-cartoon">4.39 'Moonshine' magazine cartoon</a></li> <li class="leaf menu-mlid-3988"><a href="/about-darwin/portraits-charles-darwin-catalogue/440-phrenological-magazine">4.40 'Phrenological Magazine'</a></li> <li class="leaf menu-mlid-3991"><a href="/about-darwin/portraits-charles-darwin-catalogue/441-punch-sambourne-cartoon-2">4.41 'Punch', Sambourne cartoon 2</a></li> <li class="leaf menu-mlid-3994"><a href="/about-darwin/portraits-charles-darwin-catalogue/442-punch-sambourne-cartoon-3">4.42 'Punch' Sambourne cartoon 3</a></li> <li class="leaf menu-mlid-3997"><a href="/about-darwin/portraits-charles-darwin-catalogue/443-illustrated-london-news-article">4.43 'Illustrated London News' article</a></li> <li class="leaf menu-mlid-4000"><a href="/about-darwin/portraits-charles-darwin-catalogue/444-puck-cartoon-1">4.44 'Puck' cartoon 1</a></li> <li class="leaf menu-mlid-4003"><a href="/about-darwin/portraits-charles-darwin-catalogue/445-puck-cartoon-2">4.45 'Puck' cartoon 2</a></li> <li class="leaf menu-mlid-4006"><a href="/about-darwin/portraits-charles-darwin-catalogue/446-puck-cartoon-3">4.46 'Puck' cartoon 3</a></li> <li class="leaf menu-mlid-4009"><a href="/about-darwin/portraits-charles-darwin-catalogue/447-puck-cartoon-4">4.47 'Puck' cartoon 4</a></li> <li class="leaf menu-mlid-4012"><a href="/about-darwin/portraits-charles-darwin-catalogue/448-puck-cartoon-5">4.48 'Puck', cartoon 5</a></li> <li class="leaf menu-mlid-4015"><a href="/about-darwin/portraits-charles-darwin-catalogue/449-alfred-bryan-caricature">4.49 Alfred Bryan, caricature</a></li> <li class="leaf menu-mlid-4021"><a href="/about-darwin/portraits-charles-darwin-catalogue/450-cigar-box-lid-design">4.50 Cigar box lid design</a></li> <li class="leaf menu-mlid-4024"><a href="/about-darwin/portraits-charles-darwin-catalogue/451-frederick-holder-life-and-work">4.51 Frederick Holder 'Life and Work'</a></li> <li class="leaf menu-mlid-4027"><a href="/about-darwin/portraits-charles-darwin-catalogue/452-wasp-caricature">4.52 'Wasp' caricature</a></li> <li class="leaf menu-mlid-4030"><a href="/about-darwin/portraits-charles-darwin-catalogue/453-claud-warren-outlines-hands">4.53 Claud Warren, 'Outlines of Hands'</a></li> <li class="leaf menu-mlid-4033"><a href="/about-darwin/portraits-charles-darwin-catalogue/454-jubilees-queen-victoria">4.54 jubilees of Queen Victoria</a></li> <li class="leaf menu-mlid-4036"><a href="/about-darwin/portraits-charles-darwin-catalogue/455-harry-furniss-caricature">4.55 Harry Furniss caricature</a></li> <li class="leaf menu-mlid-4039"><a href="/about-darwin/portraits-charles-darwin-catalogue/456-larks-cartoon">4.56 'Larks' cartoon</a></li> <li class="leaf menu-mlid-4042"><a href="/about-darwin/portraits-charles-darwin-catalogue/457-silhouette-cartoon">4.57 silhouette cartoon</a></li> <li class="leaf menu-mlid-4045"><a href="/about-darwin/portraits-charles-darwin-catalogue/458-simian-savage-drawings">4.58 'Simian, savage' . . . drawings</a></li> <li class="last leaf menu-mlid-4048"><a href="/about-darwin/portraits-charles-darwin-catalogue/459-simplicissimus-cartoon">4.59 'Simplicissimus' cartoon</a></li> </ul></li> <li class="expanded menu-mlid-4066"><a href="/about-darwin/darwin-and-experimental-life">Darwin and the experimental life</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-4066"><a href="/about-darwin/darwin-and-experimental-life">Darwin and the experimental life overview</a></li> <li class="leaf menu-mlid-4069"><a href="/about-darwin/darwin-and-experimental-life/what-experiment">What is an experiment?</a></li> <li class="leaf menu-mlid-4081"><a href="/about-darwin/darwin-and-experimental-life/morphology-movement-observation-and-experiment">From morphology to movement: observation and experiment</a></li> <li class="leaf menu-mlid-4078"><a href="/about-darwin/darwin-and-experimental-life/fools-experiments">Fool's experiments</a></li> <li class="leaf menu-mlid-4075"><a href="/about-darwin/darwin-and-experimental-life/experimenting-emotions">Experimenting with emotions</a></li> <li class="last leaf menu-mlid-4072"><a href="/about-darwin/darwin-and-experimental-life/animals-ethics-and-progress-science">Animals, ethics, and the progress of science</a></li> </ul></li> <li class="leaf menu-mlid-3366"><a href="/fake-darwin">Fake Darwin: myths and misconceptions</a></li> <li class="leaf menu-mlid-3402"><a href="/darwins-bad-days">Darwin's bad days</a></li> <li class="last leaf menu-mlid-3644"><a href="/people/about-darwin/darwin-s-first-love">Darwin’s first love</a></li> </ul></li> <li class="expanded menu-mlid-800"><a href="/letters" title="">The letters</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-800"><a href="/letters" title="">The letters overview</a></li> <li class="expanded menu-mlid-2080"><a href="/letters/darwins-life-letters">Darwin's life in letters</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-2080"><a href="/letters/darwins-life-letters">Darwin's life in letters overview</a></li> <li class="leaf menu-mlid-1097"><a href="/letters/darwins-life-letters/darwin-letters-1821-1836-childhood-beagle-voyage">1821-1836: Childhood to the Beagle voyage</a></li> <li class="leaf menu-mlid-1106"><a href="/letters/darwins-life-letters/darwin-letters-1837-1843-london-years-natural-selection" title="Charles Darwin's life seen through his letters, 1837-43">1837-43: The London years to 'natural selection'</a></li> <li class="leaf menu-mlid-1107"><a href="/letters/darwins-life-letters/darwin-letters-1844-1846-building-scientific-network">1844-1846: Building a scientific network</a></li> <li class="leaf menu-mlid-1108"><a href="/letters/darwins-life-letters/darwin-letters-1847-1850-microscopes-and-barnacles">1847-1850: Microscopes and barnacles</a></li> <li class="leaf menu-mlid-1109"><a href="/letters/darwins-life-letters/darwin-letters-1851-1855-death-daughter">1851-1855: Death of a daughter</a></li> <li class="leaf menu-mlid-1110"><a href="/letters/darwins-life-letters/darwin-letters-1856-1857-big-book">1856-1857: The 'Big Book'</a></li> <li class="leaf menu-mlid-936"><a href="/letters/darwins-life-letters/darwin-letters-1858-1859-origin">1858-1859: Origin</a></li> <li class="leaf menu-mlid-937"><a href="/letters/darwins-life-letters/darwin-letters-1860-answering-critics">1860: Answering critics</a></li> <li class="leaf menu-mlid-1111"><a href="/letters/darwins-life-letters/darwin-letters-1861-gaining-allies">1861: Gaining allies</a></li> <li class="leaf menu-mlid-1098"><a href="/letters/darwins-life-letters/darwin-letters-1862-multiplicity-experiments">1862: A multiplicity of experiments</a></li> <li class="leaf menu-mlid-1099"><a href="/letters/darwins-life-letters/darwin-letters-1863-quarrels-home-honours-abroad">1863: Quarrels at home, honours abroad</a></li> <li class="leaf menu-mlid-1100"><a href="/letters/darwins-life-letters/darwin-letters-1864-failing-health">1864: Failing health</a></li> <li class="leaf menu-mlid-935"><a href="/letters/darwins-life-letters/darwin-letters-1865-delays-and-disappointments">1865: Delays and disappointments</a></li> <li class="leaf menu-mlid-1101"><a href="/letters/darwins-life-letters/darwin-letters1866-survival-fittest">1866: Survival of the fittest</a></li> <li class="leaf menu-mlid-1102"><a href="/letters/darwins-life-letters/darwin-letters-1867-civilised-dispute">1867: A civilised dispute</a></li> <li class="leaf menu-mlid-1103"><a href="/letters/darwins-life-letters/darwin-letters-1868-studying-sex">1868: Studying sex</a></li> <li class="leaf menu-mlid-1104"><a href="/letters/darwins-life-letters/darwin-letters-1869-forward-all-fronts">1869: Forward on all fronts</a></li> <li class="leaf menu-mlid-1105"><a href="/letters/darwins-life-letters/darwin-letters1870-human-evolution">1870: Human evolution</a></li> <li class="leaf menu-mlid-1151"><a href="/letters/darwins-life-letters/darwin-letters-1871-emptying-nest">1871: An emptying nest</a></li> <li class="leaf menu-mlid-1152"><a href="/letters/darwins-life-letters/darwin-letters-1872-job-done">1872: Job done?</a></li> <li class="leaf menu-mlid-1049"><a href="/letters/darwins-life-letters/darwins-letters-1873-animal-or-vegetable">1873: Animal or vegetable?</a></li> <li class="leaf menu-mlid-1050"><a href="/letters/darwins-life-letters/darwin-letters-1874-turbulent-year">1874: A turbulent year</a></li> <li class="leaf menu-mlid-2275"><a href="/letters/darwins-life-letters/darwin-letters-1875-pulling-strings">1875: Pulling strings</a></li> <li class="leaf menu-mlid-2894"><a href="/letters/darwins-life-letters/darwin-letters-1876-midst-life">1876: In the midst of life</a></li> <li class="leaf menu-mlid-3363"><a href="/letters/darwins-life-letters/darwin-letters-1877-flowers-and-honours">1877: Flowers and honours</a></li> <li class="leaf menu-mlid-3383"><a href="/letters/darwins-life-letters/darwin-letters-1878-movement-and-sleep">1878: Movement and sleep</a></li> <li class="leaf menu-mlid-3394"><a href="/letters/darwins-life-letters/darwin-letters-1879-tracing-roots">1879: Tracing roots</a></li> <li class="leaf menu-mlid-3661"><a href="/letters/darwins-life-letters/darwin-letters-1880-sensitivity-and-worms">1880: Sensitivity and worms</a></li> <li class="leaf menu-mlid-4063"><a href="/letters/darwins-life-letters/darwin-letters-1881-old-friends-and-new-admirers">1881: Old friends and new admirers</a></li> <li class="last leaf menu-mlid-4102"><a href="/letters/darwins-life-letters/darwin-letters-1882-nothing-too-great-or-too-small">1882: Nothing too great or too small</a></li> </ul></li> <li class="expanded menu-mlid-3384"><a href="/letters/darwins-works-letters">Darwin's works in letters</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-3384"><a href="/letters/darwins-works-letters">Darwin's works in letters overview</a></li> <li class="leaf menu-mlid-3385"><a href="/letters/darwins-works-letters/journal-researches">Journal of researches</a></li> <li class="leaf menu-mlid-3386"><a href="/letters/darwins-works-letters/living-and-fossil-cirripedia">Living and fossil cirripedia</a></li> <li class="expanded menu-mlid-3400"><a href="/letters/darwins-works-letters/origin-big-book">Before Origin: the ‘big book’</a><ul class="menu"><li class="first collapsed menu-mlid-3400"><a href="/letters/darwins-works-letters/origin-big-book">Before Origin: the ‘big book’ overview</a></li> <li class="last leaf menu-mlid-3401"><a href="/letters/darwins-works-letters/origin-big-book/dates-composition-darwins-manuscript-species">Dates of composition of Darwin's manuscript on species</a></li> </ul></li> <li class="leaf menu-mlid-3406"><a href="/letters/darwins-works-letters/origin">Origin</a></li> <li class="expanded menu-mlid-3407"><a href="/letters/darwins-works-letters/rewriting-origin-later-editions">Rewriting Origin - the later editions</a><ul class="menu"><li class="first collapsed menu-mlid-3407"><a href="/letters/darwins-works-letters/rewriting-origin-later-editions">Rewriting Origin - the later editions overview</a></li> <li class="leaf menu-mlid-3412"><a href="/letters/darwins-works-letters/rewriting-origin-later-editions/how-old-earth">How old is the earth?</a></li> <li class="leaf menu-mlid-3411"><a href="/letters/darwins-works-letters/rewriting-origin-later-editions/whale-bear">The whale-bear</a></li> <li class="last leaf menu-mlid-3413"><a href="/letters/darwins-works-letters/rewriting-origin-later-editions/origin-lost-changes-second-german">Origin: the lost changes for the second German edition</a></li> </ul></li> <li class="leaf menu-mlid-3404"><a href="/letters/darwins-works-letters/orchids">Orchids</a></li> <li class="leaf menu-mlid-3662"><a href="/letters/darwins-works-letters/climbing-plants">Climbing plants</a></li> <li class="leaf menu-mlid-3387"><a href="/letters/darwins-works-letters/descent">Descent</a></li> <li class="leaf menu-mlid-3388"><a href="/letters/darwins-works-letters/expression">Expression</a></li> <li class="leaf menu-mlid-3389"><a href="/letters/darwins-works-letters/insectivorous-plants">Insectivorous plants</a></li> <li class="leaf menu-mlid-3663"><a href="/letters/darwins-works-letters/forms-flowers">Forms of flowers</a></li> <li class="leaf menu-mlid-4084"><a href="/letters/darwins-works-letters/cross-and-self-fertilisation">Cross and self fertilisation</a></li> <li class="leaf menu-mlid-3405"><a href="/letters/darwins-works-letters/life-erasmus-darwin">Life of Erasmus Darwin</a></li> <li class="last leaf menu-mlid-3665"><a href="/letters/darwins-works-letters/movement-plants">Movement in Plants</a></li> </ul></li> <li class="leaf menu-mlid-2264"><a href="/letters/about-letters">About the letters</a></li> <li class="expanded menu-mlid-2408"><a href="/letters/lifecycle-letter-film">Lifecycle of a letter film</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-2408"><a href="/letters/lifecycle-letter-film">Lifecycle of a letter film overview</a></li> <li class="leaf menu-mlid-2411"><a href="/letters/lifecycle-letter-film/editing-letter">Editing a Letter</a></li> <li class="last leaf menu-mlid-2409"><a href="/letters/lifecycle-letter-film/working-darwin-archive">Working in the Darwin archive</a></li> </ul></li> <li class="leaf menu-mlid-2296"><a href="/letters/capturing-darwin-s-voice-audio-selected-letters">Capturing Darwin’s voice: audio of selected letters</a></li> <li class="leaf menu-mlid-2266"><a href="/letters/correspondence-women">Correspondence with women</a></li> <li class="leaf menu-mlid-878"><a href="/hunt-new-letters">The hunt for new letters</a></li> <li class="expanded menu-mlid-1136"><a href="/letters/editorial-policy-and-practice">Editorial policy and practice</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-1136"><a href="/letters/editorial-policy-and-practice">Editorial policy and practice overview</a></li> <li class="last leaf menu-mlid-3646"><a href="/letters/editorial-policy-and-practice/full-notes-editorial-policy">Full notes on editorial policy</a></li> </ul></li> <li class="leaf menu-mlid-2263"><a href="/letters/symbols-and-abbreviations">Symbols and abbreviations</a></li> <li class="leaf menu-mlid-2905"><a href="/letters/darwins-letters-timeline">Darwin's letters: a timeline</a></li> <li class="leaf menu-mlid-2909"><a href="/world-map">Darwin's letters: World Map</a></li> <li class="leaf menu-mlid-3364"><a href="https://www.darwinproject.ac.uk/have-you-read-one-about" title="">Have you read the one about...</a></li> <li class="leaf menu-mlid-4183"><a href="/letters/charles-darwin-life-letters">Charles Darwin: A Life in Letters</a></li> <li class="leaf menu-mlid-4158"><a href="/darwin-conversation-exhibition">Darwin in Conversation exhibition</a></li> <li class="leaf menu-mlid-4099"><a href="/letters/diagrams-and-drawings-letters">Diagrams and drawings in letters</a></li> <li class="expanded menu-mlid-4105"><a href="/letters/favourite-letters">Favourite Letters</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-4105"><a href="/letters/favourite-letters">Favourite Letters overview</a></li> <li class="leaf menu-mlid-4162"><a href="/letters/favourite-letters/be-envious-ripe-oranges-w-d-fox-may-1832">Be envious of ripe oranges: To W. D. Fox, May 1832</a></li> <li class="leaf menu-mlid-4111"><a href="/letters/favourite-letters/monstrous-stain-j-m-herbert-2-june-1833">That monstrous stain: To J. M. Herbert, 2 June 1833</a></li> <li class="leaf menu-mlid-4166"><a href="/letters/favourite-letters/my-most-solemn-request-emma-darwin-5-july-1844">My most solemn request: To Emma Darwin, 5 July 1844</a></li> <li class="leaf menu-mlid-4178"><a href="/letters/favourite-letters/our-poor-dear-dear-child-emma-darwin-23-april-1851">Our poor dear dear child: To Emma Darwin, [23 April 1851]</a></li> <li class="leaf menu-mlid-4174"><a href="/letters/favourite-letters/i-beg-million-pardons-john-lubbock-3-september-1862">I beg a million pardons: To John Lubbock, [3 September 1862]</a></li> <li class="leaf menu-mlid-4129"><a href="/letters/favourite-letters/prize-possessions-henry-denny-17-january-1865">Prize possessions: To Henry Denny, 17 January [1865]</a></li> <li class="leaf menu-mlid-4114"><a href="/letters/favourite-letters/how-manage-it-j-d-hooker-17-june-1865">How to manage it: To J. D. Hooker, [17 June 1865]</a></li> <li class="leaf menu-mlid-4117"><a href="/letters/favourite-letters/fly-flower-hermann-m-ller-23-october-1867">A fly on the flower: From Hermann Müller, 23 October 1867</a></li> <li class="leaf menu-mlid-4138"><a href="/letters/favourite-letters/reading-my-roommate-s-illustrious-ancestor-t-h-huxley-10-june-1868">Reading my roommate’s illustrious ancestor: To T. H. Huxley, 10 June 1868</a></li> <li class="leaf menu-mlid-4169"><a href="/letters/favourite-letters/beginning-something-j-d-hooker-22-january-1869">A beginning, & that is something: To J. D. Hooker, [22 January 1869]</a></li> <li class="leaf menu-mlid-4135"><a href="/letters/favourite-letters/perfect-copper-plate-hand-adolf-reuter-30-may-1869">Perfect copper-plate hand: From Adolf Reuter, 30 May 1869</a></li> <li class="leaf menu-mlid-4132"><a href="/letters/favourite-letters/darwin-s-favourite-photographer-o-g-rejlander-30-april-1871">Darwin’s favourite photographer: From O. G. Rejlander, 30 April 1871</a></li> <li class="leaf menu-mlid-4172"><a href="/letters/favourite-letters/your-letter-eternalized-us-n-d-doedes-27-march-1873">Your letter eternalized before us: From N. D. Doedes, 27 March 1873</a></li> <li class="leaf menu-mlid-4120"><a href="/letters/favourite-letters/lost-translation-auguste-forel-12-november-1874">Lost in translation: From Auguste Forel, 12 November 1874</a></li> <li class="leaf menu-mlid-4182"><a href="/letters/favourite-letters/i-never-trusted-drosera-e-f-lubbock-after-2-july-1875">I never trusted Drosera: From E. F. Lubbock, [after 2 July] 1875</a></li> <li class="leaf menu-mlid-4126"><a href="/letters/favourite-letters/argus-pheasant-mivart-r-wallace-17-june-1876">From Argus pheasant to Mivart: To A. R. Wallace, 17 June 1876</a></li> <li class="leaf menu-mlid-4141"><a href="/letters/favourite-letters/wearing-his-knowledge-lightly-fritz-m-ller-5-april-1878">Wearing his knowledge lightly: From Fritz Müller, 5 April 1878</a></li> <li class="leaf menu-mlid-4123"><a href="/letters/favourite-letters/terms-engagement-julius-wiesner-25-october-1881">Terms of engagement: To Julius Wiesner, 25 October 1881</a></li> <li class="last leaf menu-mlid-4108"><a href="/letters/favourite-letters/intellectual-capacities-caroline-kennard-26-december-1881">Intellectual capacities: From Caroline Kennard, 26 December 1881</a></li> </ul></li> <li class="expanded menu-mlid-4144"><a href="/letters/darwin-plays">Darwin plays</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-4144"><a href="/letters/darwin-plays">Darwin plays overview</a></li> <li class="leaf menu-mlid-4150"><a href="/letters/darwin-plays/emma-audio-play">'Emma' audio play</a></li> <li class="leaf menu-mlid-4147"><a href="/letters/darwin-plays/frank-audio-play">'Frank' audio play</a></li> <li class="leaf menu-mlid-4153"><a href="/letters/darwin-plays/confessing-murder-audio-play">'Like confessing a murder' audio play</a></li> <li class="last expanded menu-mlid-2294"><a href="/letters/darwin-plays/re-design-dramatisation">'Re: Design' dramatisation</a><ul class="menu"><li class="first collapsed menu-mlid-2294"><a href="/letters/darwin-plays/re-design-dramatisation">'Re: Design' dramatisation overview</a></li> <li class="last leaf menu-mlid-1134"><a href="/commentary/religion/re-design-dramatisation/dramatisation-script">Dramatisation script</a></li> </ul></li> </ul></li> <li class="leaf menu-mlid-3377"><a href="https://www.darwinproject.ac.uk/search?sort=date&keyword=darwin&f1-document-type=letter" title="">Browse all Darwin letters in date order</a></li> <li class="last leaf menu-mlid-3391"><a href="/letters/list-correspondents">List of correspondents</a></li> </ul></li> <li class="expanded active-trail menu-mlid-824"><a href="/commentary" class="active-trail campl-selected">Commentary</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed active-trail menu-mlid-824"><a href="/commentary" class="active-trail">Commentary overview</a></li> <li class="expanded menu-mlid-2269"><a href="/commentary/evolution" title="">Evolution</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-2269"><a href="/commentary/evolution" title="">Evolution overview</a></li> <li class="leaf menu-mlid-2262"><a href="/commentary/evolution/natural-selection">Natural selection</a></li> <li class="leaf menu-mlid-2293"><a href="/commentary/evolution/sexual-selection">Sexual selection</a></li> <li class="leaf menu-mlid-2299"><a href="/commentary/evolution/inheritance">Inheritance</a></li> <li class="leaf menu-mlid-2298"><a href="/commentary/evolution/divergence">Divergence</a></li> <li class="leaf menu-mlid-3410"><a href="/commentary/evolution/correlation-growth-deaf-blue-eyed-cats-pigs-and-poison">Correlation of growth: deaf blue-eyed cats, pigs, and poison</a></li> <li class="leaf menu-mlid-3414"><a href="/commentary/evolution/natural-selection-trouble-terminology-part-i">Natural Selection: the trouble with terminology Part I</a></li> <li class="leaf menu-mlid-3415"><a href="/commentary/evolution/survival-fittest-trouble-terminology-part-ii">Survival of the fittest: the trouble with terminology Part II</a></li> <li class="last leaf menu-mlid-3671"><a href="/commentary/evolution/darwin-s-species-notebooks-i-think">Darwin’s species notebooks: ‘I think . . .’</a></li> </ul></li> <li class="expanded menu-mlid-871"><a href="/commentary/geology" title="">Geology</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-871"><a href="/commentary/geology" title="">Geology overview</a></li> <li class="leaf menu-mlid-2259"><a href="/commentary/geology/darwin-geology">Darwin and geology</a></li> <li class="leaf menu-mlid-1127"><a href="/topics/geology/darwin-s-introduction-geology">Darwin’s introduction to geology</a></li> <li class="leaf menu-mlid-1270"><a href="/commentary/geology/geology-beagle-voyage">The geology of the Beagle voyage</a></li> <li class="leaf menu-mlid-1122"><a href="/commentary/geology/darwin-coral-reefs">Darwin and coral reefs</a></li> <li class="leaf menu-mlid-2258"><a href="/commentary/geology/darwin-s-earthquakes">Darwin’s earthquakes</a></li> <li class="leaf menu-mlid-2257"><a href="/topics/geology/darwin-geological-society" title="">Darwin and the Geological Society</a></li> <li class="leaf menu-mlid-1123"><a href="/commentary/geology/darwin-glen-roy">Darwin and Glen Roy</a></li> <li class="last leaf menu-mlid-1087"><a href="/topics/geology/bibliography-darwin-s-geological-publications">Bibliography of Darwin’s geological publications</a></li> </ul></li> <li class="expanded active-trail menu-mlid-2247"><a href="/commentary/life-sciences" title="" class="active-trail">Life sciences</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed active-trail menu-mlid-2247"><a href="/commentary/life-sciences" title="" class="active-trail">Life sciences overview</a></li> <li class="expanded menu-mlid-1117"><a href="/commentary/life-sciences/darwin-and-down">Darwin and Down</a><ul class="menu"><li class="first collapsed menu-mlid-1117"><a href="/commentary/life-sciences/darwin-and-down">Darwin and Down overview</a></li> <li class="last leaf menu-mlid-1149"><a href="/commentary/life-sciences/darwin-and-down/darwin-s-hothouse-and-lists-hothouse-plants">Darwin’s hothouse and lists of hothouse plants</a></li> </ul></li> <li class="leaf active-trail active menu-mlid-4087 campl-current-page"><a href="/commentary/life-sciences/species-and-varieties" class="active-trail active">Species and varieties</a></li> <li class="leaf menu-mlid-1058"><a href="/commentary/life-sciences/evolution-honeycomb">The evolution of honeycomb</a></li> <li class="leaf menu-mlid-1083"><a href="/commentary/life-sciences/tale-two-bees">A tale of two bees</a></li> <li class="expanded menu-mlid-1088"><a href="/commentary/life-sciences/beauty-and-seed">Beauty and the seed</a><ul class="menu"><li class="first collapsed menu-mlid-1088"><a href="/commentary/life-sciences/beauty-and-seed">Beauty and the seed overview</a></li> <li class="last leaf menu-mlid-1055"><a href="/commentary/life-sciences/beauty-and-seed/mauro-galetti-profile-ecologist">Mauro Galetti: profile of an ecologist</a></li> </ul></li> <li class="leaf menu-mlid-2261"><a href="/commentary/life-sciences/casting-about-darwin-worms">Casting about: Darwin on worms</a></li> <li class="leaf menu-mlid-1081"><a href="/commentary/life-sciences/was-darwin-ecologist">Was Darwin an ecologist?</a></li> <li class="leaf menu-mlid-3641"><a href="/commentary/life-sciences/dipsacus-and-drosera-frank-s-favourite-carnivores">Dipsacus and Drosera</a></li> <li class="expanded menu-mlid-2318"><a href="/commentary/life-sciences/darwin-and-barnacles">Darwin and barnacles</a><ul class="menu"><li class="first collapsed menu-mlid-2318"><a href="/commentary/life-sciences/darwin-and-barnacles">Darwin and barnacles overview</a></li> <li class="last leaf menu-mlid-2317"><a href="/commentary/life-sciences/darwin-and-barnacles/darwin-s-study-cirripedia">Darwin’s study of the Cirripedia</a></li> </ul></li> <li class="expanded menu-mlid-2280"><a href="/commentary/life-sciences/darwin-and-vivisection">Darwin and vivisection</a><ul class="menu"><li class="first collapsed menu-mlid-2280"><a href="/commentary/life-sciences/darwin-and-vivisection">Darwin and vivisection overview</a></li> <li class="leaf menu-mlid-2284"><a href="/topics/life-sciences/darwin-and-vivisection/vivisection-draft-petition">Vivisection: draft petition</a></li> <li class="leaf menu-mlid-2285"><a href="/topics/life-sciences/darwin-and-vivisection/vivisection-baas-committee-report">Vivisection: BAAS committee report</a></li> <li class="leaf menu-mlid-2286"><a href="/topics/life-sciences/darwin-and-vivisection/vivisection-first-sketch-bill">Vivisection: first sketch of the bill</a></li> <li class="leaf menu-mlid-2287"><a href="/topics/life-sciences/darwin-and-vivisection/vivisection-darwins-testimony-royal-commission">Vivisection: Darwin's testimony</a></li> <li class="last leaf menu-mlid-2288"><a href="/topics/life-sciences/darwin-and-vivisection/appeal-against-animal-cruelty">'An Appeal' against animal cruelty</a></li> </ul></li> <li class="last leaf menu-mlid-2916"><a href="/commentary/life-sciences/biodiversity-and-its-histories">Biodiversity and its histories</a></li> </ul></li> <li class="expanded menu-mlid-873"><a href="/commentary/human-nature" title="">Human nature</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-873"><a href="/commentary/human-nature" title="">Human nature overview</a></li> <li class="leaf menu-mlid-2604"><a href="/commentary/human-nature/darwin-human-evolution">Darwin on human evolution</a></li> <li class="expanded menu-mlid-1269"><a href="/commentary/human-nature/expression-emotions">The expression of emotions</a><ul class="menu"><li class="first collapsed menu-mlid-1269"><a href="/commentary/human-nature/expression-emotions">The expression of emotions overview</a></li> <li class="expanded menu-mlid-1262"><a href="/commentary/human-nature/expression-emotions/emotion-experiment">Emotion experiment</a><ul class="menu"><li class="first collapsed menu-mlid-1262"><a href="/commentary/human-nature/expression-emotions/emotion-experiment">Emotion experiment overview</a></li> <li class="last leaf menu-mlid-1263"><a href="/commentary/human-nature/expression-emotions/emotion-experiment/results-darwin-online-emotions">Results of the Darwin Online Emotions Experiment</a></li> </ul></li> <li class="leaf menu-mlid-2256"><a href="/commentary/human-nature/expression-emotions/face-emotion">Face of emotion</a></li> <li class="last leaf menu-mlid-2292"><a href="/commentary/human-nature/expression-emotions/darwin-s-queries-expression">Darwin’s queries on expression</a></li> </ul></li> <li class="expanded menu-mlid-1052"><a href="/commentary/human-nature/origin-language">The origin of language</a><ul class="menu"><li class="first collapsed menu-mlid-1052"><a href="/commentary/human-nature/origin-language">The origin of language overview</a></li> <li class="leaf menu-mlid-1053"><a href="/commentary/human-nature/origin-language/language-key-letters">Language: key letters</a></li> <li class="last leaf menu-mlid-1153"><a href="/commentary/human-nature/origin-language/language-interview-gregory-radick">Language: Interview with Gregory Radick</a></li> </ul></li> <li class="last leaf menu-mlid-1140"><a href="/commentary/human-nature/darwin-and-human-nature-film-series">Film series podcasts</a></li> </ul></li> <li class="expanded menu-mlid-874"><a href="/commentary/religion" title="">Religion</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-874"><a href="/commentary/religion" title="">Religion overview</a></li> <li class="leaf menu-mlid-1033"><a href="/commentary/religion/darwin-and-design">Darwin and design</a></li> <li class="leaf menu-mlid-1288"><a href="/commentary/religion/what-did-darwin-believe">What did Darwin believe?</a></li> <li class="leaf menu-mlid-1034"><a href="/commentary/religion/darwin-and-church">Darwin and the Church</a></li> <li class="leaf menu-mlid-1089"><a href="/commentary/religion/british-association-meeting-1860">British Association meeting 1860</a></li> <li class="leaf menu-mlid-1121"><a href="/commentary/religion/darwin-and-religion-america">Darwin and religion in America</a></li> <li class="expanded menu-mlid-1045"><a href="/commentary/religion/essays-reviews-asa-gray">Essays and reviews by Asa Gray</a><ul class="menu"><li class="first collapsed menu-mlid-1045"><a href="/commentary/religion/essays-reviews-asa-gray">Essays and reviews by Asa Gray overview</a></li> <li class="leaf menu-mlid-2271"><a href="/commentary/religion/essays-reviews-asa-gray/darwiniana-preface">Darwiniana – Preface</a></li> <li class="leaf menu-mlid-2272"><a href="/commentary/religion/essays-reviews-asa-gray/essay-design-versus-necessity">Essay: Design versus necessity</a></li> <li class="leaf menu-mlid-1046"><a href="/commentary/religion/essays-reviews-asa-gray/essay-natural-selection-natural-theology">Essay: Natural selection & natural theology</a></li> <li class="leaf menu-mlid-1146"><a href="/commentary/religion/essays-reviews-asa-gray/essay-evolution-theology">Essay: Evolution and theology</a></li> <li class="leaf menu-mlid-2273"><a href="/commentary/religion/essays-reviews-asa-gray/essay-what-darwinism">Essay: What is Darwinism?</a></li> <li class="last leaf menu-mlid-2274"><a href="/commentary/religion/essays-reviews-asa-gray/essay-evolutionary-teleology">Essay: Evolutionary teleology</a></li> </ul></li> <li class="last expanded menu-mlid-2249"><a href="/commentary/religion/science-and-religion-interviews">Science and religion Interviews</a><ul class="menu"><li class="first collapsed menu-mlid-2249"><a href="/commentary/religion/science-and-religion-interviews">Science and religion Interviews overview</a></li> <li class="leaf menu-mlid-2250"><a href="/commentary/religion/science-and-religion-interviews/interview-emily-ballou">Interview with Emily Ballou</a></li> <li class="leaf menu-mlid-2251"><a href="/commentary/religion/science-and-religion-interviews/interview-simon-conway-morris">Interview with Simon Conway Morris</a></li> <li class="leaf menu-mlid-2252"><a href="/commentary/religion/science-and-religion-interviews/interview-john-hedley-brooke">Interview with John Hedley Brooke</a></li> <li class="leaf menu-mlid-2253"><a href="/commentary/religion/science-and-religion-interviews/interview-randal-keynes">Interview with Randal Keynes</a></li> <li class="leaf menu-mlid-2254"><a href="/commentary/religion/science-and-religion-interviews/interview-tim-lewens">Interview with Tim Lewens</a></li> <li class="last leaf menu-mlid-2255"><a href="/commentary/religion/science-and-religion-interviews/interview-pietro-corsi">Interview with Pietro Corsi</a></li> </ul></li> </ul></li> <li class="last expanded menu-mlid-3395"><a href="/commentary/curious" title="">For the curious...</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-3395"><a href="/commentary/curious" title="">For the curious... overview</a></li> <li class="leaf menu-mlid-4155"><a href="/commentary/curious/cordillera-beagle-expedition">Cordillera Beagle expedition</a></li> <li class="leaf menu-mlid-4096"><a href="/commentary/curious/darwin-family">The Darwin family</a></li> <li class="leaf menu-mlid-4093"><a href="/commentary/curious/darwin-s-plant-experiments">Darwin’s plant experiments</a></li> <li class="leaf menu-mlid-4090"><a href="/commentary/curious/behind-scenes">Behind the scenes</a></li> <li class="leaf menu-mlid-3664"><a href="/commentary/curious/darwin-s-networks">Darwin’s Networks</a></li> <li class="leaf menu-mlid-3643"><a href="/commentary/curious/darwin-and-beagle-voyage">Darwin and the Beagle voyage</a></li> <li class="leaf menu-mlid-3403"><a href="/commentary/curious/darwin-and-working-home">Darwin and working from home</a></li> <li class="leaf menu-mlid-3409"><a href="/commentary/curious/darwin-cats-and-cat-shows">Darwin, cats and cat shows</a></li> <li class="leaf menu-mlid-3396"><a href="/commentary/curious/darwin-and-dogs">Darwin and dogs</a></li> <li class="leaf menu-mlid-3397"><a href="/commentary/curious/darwins-illness">Darwin's illness</a></li> <li class="leaf menu-mlid-3398"><a href="/commentary/curious/plant-or-animal-or-don-t-try-home">Plant or animal? (Or: Don’t try this at home!)</a></li> <li class="last leaf menu-mlid-3399"><a href="/commentary/curious/strange-things-sent-darwin-post">Strange things sent to Darwin in the post</a></li> </ul></li> </ul></li> <li class="expanded menu-mlid-823"><a href="/people">People</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-823"><a href="/people">People overview</a></li> <li class="leaf menu-mlid-2150"><a href="/commentary/key-correspondents" title="">Key correspondents</a></li> <li class="leaf menu-mlid-2153"><a href="/commentary/beagle-voyage-networks" title="">Beagle voyage networks</a></li> <li class="leaf menu-mlid-2152"><a href="/commentary/family-and-friends" title="">Family and friends</a></li> <li class="leaf menu-mlid-2159"><a href="/commentary/darwins-scientific-network" title="">Darwin's scientific network</a></li> <li class="leaf menu-mlid-2155"><a href="/tags/readers-and-critics" title="">Readers and critics</a></li> <li class="leaf menu-mlid-2162"><a href="/tags/publishers-artists-and-illustrators" title="">Publishers, artists and illustrators</a></li> <li class="leaf menu-mlid-3381"><a href="/correspondents-alphabetical" title="">People pages in alphabetical order</a></li> <li class="expanded menu-mlid-3367"><a href="/people/german-and-dutch-photograph-albums">German and Dutch photograph albums</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-3367"><a href="/people/german-and-dutch-photograph-albums">German and Dutch photograph albums overview</a></li> <li class="leaf menu-mlid-3372"><a href="/people/german-and-dutch-photograph-albums/photograph-album-german-and-austrian-scientists">Photograph album of German and Austrian scientists</a></li> <li class="leaf menu-mlid-3371"><a href="/people/german-and-dutch-photograph-albums/photograph-album-dutch-admirers">Photograph album of Dutch admirers</a></li> <li class="last leaf menu-mlid-3370"><a href="/german-poems-presented-darwin">German poems presented to Darwin</a></li> </ul></li> <li class="last leaf menu-mlid-3380"><a href="https://www.darwinproject.ac.uk/search?text=%22%22;f1-document-type=people;sort=name" title="">List of all people mentioned in letters</a></li> </ul></li> <li class="expanded menu-mlid-825"><a href="/learning-resources" title="">Learning</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-825"><a href="/learning-resources" title="">Learning overview</a></li> <li class="expanded menu-mlid-1739"><a href="/learning/7-11">Ages 7-11</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-1739"><a href="/learning/7-11">Ages 7-11 overview</a></li> <li class="leaf menu-mlid-2601"><a href="/learning/7-11/darwin-the-collector">Darwin The Collector</a></li> <li class="leaf menu-mlid-2602"><a href="/learning/7-11/detecting-darwin">Detecting Darwin</a></li> <li class="leaf menu-mlid-2603"><a href="/learning/7-11/darwin-and-evolution">Darwin And Evolution</a></li> <li class="leaf menu-mlid-2600"><a href="/learning/7-11/darwins-fantastical-voyage">Darwin's Fantastical Voyage</a></li> <li class="last leaf menu-mlid-3408"><a href="/learning/7-11/home-learning-7-11-years">Home learning: 7-11 years</a></li> </ul></li> <li class="expanded menu-mlid-1744"><a href="/learning/11-14">Ages 11-14</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-1744"><a href="/learning/11-14">Ages 11-14 overview</a></li> <li class="leaf menu-mlid-2899"><a href="/learning/11-14/darwin-and-religion">Darwin and Religion</a></li> <li class="leaf menu-mlid-2898"><a href="/learning/11-14/doing-darwins-experiments">Doing Darwin’s Experiments</a></li> <li class="leaf menu-mlid-2900"><a href="/learning/11-14/how-dangerous-was-darwin">How dangerous was Darwin?</a></li> <li class="leaf menu-mlid-2901"><a href="/learning/11-14/offer-of-a-lifetime">Offer of a lifetime</a></li> <li class="leaf menu-mlid-2903"><a href="/learning/11-14/darwin-and-slavery">Darwin and slavery</a></li> <li class="leaf menu-mlid-2902"><a href="/learning/11-14/beagle-voyage">Beagle Voyage</a></li> <li class="leaf menu-mlid-2904"><a href="/learning/11-14/darwins-scientific-women">Darwin’s scientific women</a></li> <li class="last leaf menu-mlid-1154"><a href="/case-studies-using-darwin%E2%80%99s-letters-classroom">Schools Gallery: Using Darwin’s letters in the classroom</a></li> </ul></li> <li class="expanded menu-mlid-833"><a href="/learning/universities">Universities</a><ul class="campl-unstyled-list local-dropdown-menu"><li class="first collapsed menu-mlid-833"><a href="/learning/universities">Universities overview</a></li> <li class="expanded menu-mlid-1063"><a href="/learning/universities/letters-primary-source" title="">Letters as a primary source</a><ul class="menu"><li class="first collapsed menu-mlid-1063"><a href="/learning/universities/letters-primary-source" title="">Letters as a primary source overview</a></li> <li class="leaf menu-mlid-2276"><a href="/learning/universities/letters-primary-source/scientific-networks">Scientific networks</a></li> <li class="leaf menu-mlid-2277"><a href="/learning/universities/letters-primary-source/scientific-practice">Scientific practice</a></li> <li class="leaf menu-mlid-2278"><a href="/learning/universities/letters-primary-source/controversy">Controversy</a></li> <li class="leaf menu-mlid-2279"><a href="/learning/universities/letters-primary-source/religion">Religion</a></li> <li class="leaf menu-mlid-1066"><a href="/learning/universities/letters-primary-source/discussion-questions-and-essay-questions">Discussion questions and essay questions</a></li> <li class="last leaf menu-mlid-1286"><a href="/learning/universities/letters-primary-source/suggested-reading">Suggested reading</a></li> </ul></li> <li class="expanded menu-mlid-1550"><a href="/learning/universities/getting-know-darwins-science">Getting to know Darwin's science</a><ul class="menu"><li class="first collapsed menu-mlid-1550"><a href="/learning/universities/getting-know-darwins-science">Getting to know Darwin's science overview</a></li> <li class="leaf menu-mlid-2144"><a href="/learning/universities/getting-know-darwins-science/early-days">Early days</a></li> <li class="leaf menu-mlid-1086"><a href="/learning/universities/getting-know-darwins-science/barnacles">Barnacles</a></li> <li class="leaf menu-mlid-932"><a href="/learning/universities/getting-know-darwins-science/biogeography">Biogeography</a></li> <li class="leaf menu-mlid-1287"><a href="/learning/universities/getting-know-darwins-science/variation-under-domestication">Variation under domestication</a></li> <li class="leaf menu-mlid-1259"><a href="/learning/universities/getting-know-darwins-science/orchids">Orchids</a></li> <li class="leaf menu-mlid-1150"><a href="/learning/universities/getting-know-darwins-science/instinct-and-evolution-mind">Instinct and the evolution of mind</a></li> <li class="leaf menu-mlid-1048"><a href="/learning/universities/getting-know-darwins-science/insectivorous-plants">Insectivorous plants</a></li> <li class="leaf menu-mlid-1095"><a href="/learning/universities/getting-know-darwins-science/climbing-plants">Climbing plants</a></li> <li class="leaf menu-mlid-1141"><a href="/learning/universities/getting-know-darwins-science/floral-dimorphism">Floral dimorphism</a></li> <li class="leaf menu-mlid-1260"><a href="/learning/universities/getting-know-darwins-science/power-movement-plants">Power of movement in plants</a></li> <li class="leaf menu-mlid-1135"><a href="/learning/universities/getting-know-darwins-science/earthworms">Earthworms</a></li> <li class="last leaf menu-mlid-1132"><a href="/learning/universities/getting-know-darwins-science/dining-down-house">Dining at Down House</a></li> </ul></li> <li class="expanded menu-mlid-2314"><a href="/learning/universities/darwin-and-human-nature">Darwin and human nature</a><ul class="menu"><li class="first collapsed menu-mlid-2314"><a href="/learning/universities/darwin-and-human-nature">Darwin and human nature overview</a></li> <li class="leaf menu-mlid-1155"><a href="/learning/universities/darwin-and-human-nature/moral-nature">Moral nature</a></li> <li class="last leaf menu-mlid-1261"><a href="/learning/universities/darwin-and-human-nature/race-civilization-and-progress">Race, civilization, and progress</a></li> </ul></li> <li class="last expanded menu-mlid-1289"><a href="/learning/universities/women-and-science">Women and science</a><ul class="menu"><li class="first collapsed menu-mlid-1289"><a href="/learning/universities/women-and-science">Women and science overview</a></li> <li class="leaf menu-mlid-2281"><a href="/learning/universities/women-and-science/women-s-scientific-participation">Women’s scientific participation</a></li> <li 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class="rdf-meta element-hidden"></span><span property="sioc:num_replies" content="0" datatype="xsd:integer" class="rdf-meta element-hidden"></span> <div class="content campl-content-container"> <div class="field field-name-field-image field-type-file field-label-hidden"><div class="field-items"><div class="field-item even"><div id="file-4150" class="file file-image file-image-jpeg"> <h2 class="element-invisible"><a href="/file/origin1859title-pagecjpg">Origin_1859_Title page_c.jpg</a></h2> <div class="content"> <a href="https://www.darwinproject.ac.uk/sites/default/files/Origin_1859_Title%20page_c.jpg"><img typeof="foaf:Image" class="campl-scale-with-grid" src="https://www.darwinproject.ac.uk/sites/default/files/Origin_1859_Title%20page_c.jpg" width="786" height="1232" alt="Origin of species (1859) title page" title="Origin of species (1859) title page" /></a><div class="field field-name-field-file-caption field-type-text field-label-hidden"><div class="field-items"><div class="field-item even">Origin of species (1859) title page</div></div></div> </div> </div> </div></div></div><div class="field field-name-body field-type-text-with-summary field-label-hidden"><div class="field-items"><div class="field-item even" property="content:encoded"><p><em>On the origin of species by means of natural selection</em> …so begins the title of Darwin’s most famous book, and the reader would rightly assume that such a thing as ‘species’ must therefore exist and be subject to description. But the title continues, <em>…or the preservation of favoured races in the struggle for life</em>. This is more ambiguous, especially to the modern reader, for whom race carries a different and highly charged meaning. In the context of natural history, when Darwin used the term here, he simply meant ‘variety’, as in ‘a fast-growing race of wheat’. The question, then, is whether Darwin thought that species were ‘real’ or whether boundaries among races or varieties were so fluid that clearly defined species didn’t exist. The answer to the question is … both!</p> <p> </p> <p> <strong>‘Undiscoverable essence’</strong></p> <p> In one of his reading notebooks, under the lists of books he wanted to read (DAR 119: 2v), Darwin scribbled a reminder to himself in 1838 to ‘read Aristotle to see if any of my views are ancient’. He never got around to reading Aristotle beyond a few extracts, until shortly before his death, when William Ogle sent his translation, <em>Aristotle on the parts of animals</em> (Ogle trans. 1882). Darwin would have found that Aristotle seemed to deny the existence of fixed beings when he wrote, ‘For nature passes from lifeless objects to animals in such unbroken sequence, interposing between them beings which live and yet are not animals, that scarcely any difference seems to exist between two neighbouring groups owing to their close proximity’ (<em>ibid.</em>, p. 104). Contrast this with Aristotelian essentialism, the idea that certain attributes are essential to the characterisation of things, and you have, in a nutshell, the two sides of a debate about the nature of living things that continued from antiquity.</p> <p>The notion of an essence or essential characters that define or delimit organisms was central to Linnaeus’s great taxonomic project, <em>Systema naturæ</em>, and to most subsequent attempts at systematising organisms until Darwin published his own taxonomic works between 1851 and 1854. Linnaeus ordered the world according to an artificial system; that is, he chose a specific group of structural features by which he ordered organisms, without regard for environmental or other criteria. He was challenged by others who searched for a more ‘natural’ taxonomy that would exemplify the actual relationships of species, genera, orders, and classes. The notion of a defining characteristic seems straightforward only until it is put into practice when faced with a group of organisms. Choosing characters on which to base inclusion in a group is fraught with difficulty. In taxonomy, the characteristics that formed the basis for these choices were known as diagnostic features, but the trick was choosing which features were essential; that is, which features could be used to diagnose the place of an individual organism.</p> <p>Darwin himself did not set out to be a taxonomist, but in trying to understand some of the anatomical anomalies of a tiny parasitic barnacle, he was eventually led to revise the taxonomy of the whole group, including extinct species. His approach to taxonomy was similar to those who sought to determine actual relationships; that is, he focused on how one species or variety related to other similar ones. He went further, however, not just trying to determine similarities in structure and habits but trying to discover relationships based on descent from earlier forms. He always tried to make descriptions after having seen several individuals, having noted after years of observation just how much variability often existed within a species. The features he focused on were not simply morphological, but were those clearly related to each animal’s place in the environment: adaptive features. This taxonomy was underpinned by the idea that the relations in features reflected a real genealogical relationship over time. In the four volumes on the sub-class Cirripedia, Darwin maintained the distinction between species and varieties and even complained while working on his descriptions, ‘Literally not one species is properly defined: not one naturalist has ever taken the trouble to open the shell of any species to describe it scientifically, & yet all the genera have 1/2 a dozen synonyms’ (<a href="/letter/?docId=letters/DCP-LETT-1221.xml">letter to H. E. Strickland, [4 February 1849]</a>).</p> <p> In the conclusion to <em>Origin</em>, Darwin stated his belief about systematics informed by an evolutionary point of view, noting, ‘Systematists will be able to pursue their labours as at present; but they will not be incessantly haunted by the shadowy doubt whether this or that form be in essence a species.’ He continued, regarding species and varieties, ‘Hereafter we shall be compelled to acknowledge that the only distinction between species and well-marked varieties is, that the latter are known, or believed, to be connected at the present day by intermediate gradations, whereas species were formerly thus connected. Hence, without quite rejecting the consideration of the present existence of intermediate gradations between any two forms, we shall be led to weigh more carefully and to value higher the actual amount of difference between them. …we shall at least be freed from the vain search for the undiscovered and undiscoverable essence of the term species’ (<em>Origin</em>, p. 485).<br />  </p> <p><strong>‘To define the undefinable’</strong></p> <p> Darwin developed his theory of evolution by natural selection over many years and gave a lot of thought to definitions of species both implicit and explicit in the work of contemporary naturalists. In a letter to his friend Joseph Hooker, he wrote, ‘It is really laughable to see what different ideas are prominent in various naturalists minds, when they speak of “species” in some resemblance is everything & descent of little weight—in some resemblance seems to go for nothing & Creation the reigning idea—in some descent the key—in some sterility an unfailing test, with others not worth a farthing. It all comes, I believe, from trying to define the undefinable’ (<a href="/letter/?docId=letters/DCP-LETT-2022.xml">letter to  J. D. Hooker, 24 December [1856]</a>). The idea that sterility was a test of species was firmly held by Thomas Huxley, who argued that until Darwin could demonstrate the possibility of developing from a particular stock, by selective breeding, two forms, which should either be unable to cross one with another, or whose cross-bred offspring should be infertile with one another, the theory of evolution by natural selection would not be ‘beyond the reach of all possible assault’ (T. H. Huxley 1863a, p. 147).</p> <p>In <em>Origin </em>p. 272, Darwin had argued that the sterility of interspecific hybrids was not a special endowment but was gradually acquired through divergent modifications in the reproductive systems of the forms that were crossed; in other words, as incipient new species diverged from each other while undergoing various structural changes, some changes to reproductive organs would eventually result in two related species no longer being able either to breed, or to produce fertile offspring. Huxley’s challenge to create two forms infertile with one another would have been hard to achieve in practice, given that Darwin’s theory required that small variations over very long periods of time were required to effect change. Darwin began to look at sterility from a different perspective. In May 1860, he noticed differences in the flowers of cowslips, some plants having long stamens (male organs) and short pistils (female organs), others with short stamens and long pistils; he also noted differences in the size of the pollen in the two forms. Over the next decades, Darwin carried out numerous experiments with many plant species whose flowers had two or three different forms and published five articles and eventually a book, <em>The different forms of flowers on plants of the same species </em>(1877) What Darwin discovered was that many, but by no means all, different sexual forms were often sterile if crossed with the same form. Sometimes all different forms of a species were self-sterile, sometimes one form was self-sterile, while another was fertile with its own pollen. There seemed to be no universal law about sterility, even in members of the same species.</p> <p>Throughout the 1860s, Darwin vacillated about whether sterility could be ‘selected’. In 1862, he told Hooker, ‘I am now strongly inclined to believe that sterility is at first a selected quality to keep incipient species distinct’ (<a href="/letter/?docId=letters/DCP-LETT-4765A.xml">letter to J. D. Hooker, 12 [December 1862]</a>). In 1866, Darwin compared the sterility of hybrids, or the offspring of hybrids, with the sterility of same-form unions among dimorphic and trimorphic plants; he evidently still thought the latter might be caused by natural selection (<em>Origin</em> 4th ed., pp. 323–6). However, by 1868, in <em>The variation of animals and plants under domestication</em>, Darwin argued that degrees of lessened fertility could not have been slowly accumulated by means of natural selection because no advantage would accrue from fewer offspring resulting from crosses with other varieties, and therefore lessened fertility would not be 'selected'. (<em>Variation</em> 2: 186–7). One of the reasons that Darwin’s views on the subject crystallised at this time was his discussion of the issue with Alfred Wallace in the spring of 1868. Wallace had sent a concise elaboration of how the sterility of hybrids might be produced by natural selection (<a href="/letter/?docId=letters/DCP-LETT-5966.xml">letter from A. R. Wallace, 1 March 1868</a>). Darwin turned to his son George, a mathematician, who stated the problem as a mathematical equation, finding that natural selection could not increase sterility. Wallace was not convinced by the objections and the exchange continued briefly until he decided to ‘say no more but leave the problem as insoluble’ (<a href="/letter/?docId=letters/DCP-LETT-6104.xml">letter from A. R. Wallace, 8 [April] 1868</a>).</p> <p>Ultimately, Darwin’s view was practical and flexible. Species could be defined in relation to each other, and, over time, these relations could change in response to the continually changing environment. Varieties might be potential new species, and present species could once have been varieties. The relations among species were not derived from some idealistic notion of essences, but from actual genealogical relationships over time.</p> <p> </p> </div></div></div><div class="field field-name-field-terms field-type-taxonomy-term-reference field-label-above"><div class="field-label">Terms: </div><div class="field-items"><div class="field-item even"><a href="/commentary/life-sciences" typeof="skos:Concept" property="rdfs:label skos:prefLabel" datatype="">Life sciences</a></div></div></div><div class="field field-name-field-term-related-letters field-type-text field-label-above"><div class="field-label">Related letters: </div><div class="field-items"><div class="field-item even">1221</div><div class="field-item odd">2022</div><div class="field-item even">3855</div><div class="field-item odd">5966</div><div class="field-item even">6104</div></div></div><div class="field field-name-field-related-bibliographic-refe field-type-text field-label-above"><div class="field-label">Related bibliographic reference: </div><div class="field-items"><div class="field-item even"><div class="bibliography even"> Ogle, William, trans. 1882. <i>Aristotle on the parts of animals</i>. London: Kegan Paul, Trench & Co.</div></div><div class="field-item odd"><div class="bibliography even"> Linnaeus, Carolus (Carl von Linné). 1758–9. <i>Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis.</i> 10th edition. 2 vols. Stockholm: Laurentius Salvius.</div></div><div class="field-item even"><div class="bibliography even"> <i>Origin</i>: <i>On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life.</i> By Charles Darwin. London: John Murray. 1859.</div></div><div class="field-item odd"><div class="bibliography even"> Huxley, Thomas Henry. 1863a. <i>Evidence as to man’s place in nature.</i> London: Williams & Norgate.</div></div><div class="field-item even"><div class="bibliography even"> <i>Origin</i> 4th ed.: <i>On the origin of species by means of natural selection, or the preservation of favoured races in the struggle for life.</i> 4th edition, with additions and corrections. By Charles Darwin. London: John Murray. 1866.</div></div><div class="field-item odd"><div class="bibliography even"> <i>Variation</i>: <i>The variation of animals and plants under domestication.</i> By Charles Darwin. 2 vols. London: John Murray. 1868.</div></div><div class="field-item even"><div class="bibliography even"> <i>Forms of flowers</i>: <i>The different forms of flowers on plants of the same species.</i> By Charles Darwin. London: John Murray. 1877.</div></div></div></div> </div> </div> </div> </div> </div> </div> </div> <div class="campl-column3 campl-secondary-content "> <div class="region region-sidebar"> <div id="block-menu-block-3" class="block block-menu-block"> <div class="campl-content-container campl-no-bottom-padding"> <div class="campl-heading-container"> <h2>In this section:</h2> </div> </div> <div class="campl-content-container"> <div class="menu-block-wrapper menu-block-3 menu-name-main-menu parent-mlid-0 menu-level-3"> <ul class="menu"><li class="first collapsed menu-mlid-1117 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/Down-house1_4.jpg?itok=4_rtOp2x)"><a href="/commentary/life-sciences/darwin-and-down">Darwin and Down</a></li> <li class="leaf active-trail active menu-mlid-4087 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/Origin_1859_Title%20page_c.jpg?itok=7naCGxR8)"><a href="/commentary/life-sciences/species-and-varieties" class="active-trail active">Species and varieties</a></li> <li class="leaf menu-mlid-1058 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/Bees-cells-image-1-300x225.jpg?itok=Ha7HVE-3)"><a href="/commentary/life-sciences/evolution-honeycomb">The evolution of honeycomb</a></li> <li class="leaf menu-mlid-1083 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/Bombus-hypnorum.jpg?itok=K-XkShjF)"><a href="/commentary/life-sciences/tale-two-bees">A tale of two bees</a></li> <li class="collapsed menu-mlid-1088 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/CAMPBELL-G-D-01-00802.jpg?itok=37yjHVqZ)"><a href="/commentary/life-sciences/beauty-and-seed">Beauty and the seed</a></li> <li class="leaf menu-mlid-2261 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/PR-T-00992-B-00001-00045_worm.jpg?itok=QpmPFG6c)"><a href="/commentary/life-sciences/casting-about-darwin-worms">Casting about: Darwin on worms</a></li> <li class="leaf menu-mlid-1081 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/PR-Q-00340-00001-C-00007-00004-000-p471.jpg?itok=amNzEpLQ)"><a href="/commentary/life-sciences/was-darwin-ecologist">Was Darwin an ecologist?</a></li> <li class="leaf menu-mlid-3641 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/%28Dipsacus%20sylvestris%29-plate-1.jpg?itok=nYJvk7OO)"><a href="/commentary/life-sciences/dipsacus-and-drosera-frank-s-favourite-carnivores">Dipsacus and Drosera</a></li> <li class="collapsed menu-mlid-2318 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/MS-DAR-00029-00003-000-00272-crop.jpg?itok=REk6Bi3J)"><a href="/commentary/life-sciences/darwin-and-barnacles">Darwin and barnacles</a></li> <li class="collapsed menu-mlid-2280 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/COBBE-F-P-01-00984.jpg?itok=Mw540KZZ)"><a href="/commentary/life-sciences/darwin-and-vivisection">Darwin and vivisection</a></li> <li class="last leaf menu-mlid-2916 with-rh-icon" style="background-image:url(https://www.darwinproject.ac.uk/sites/default/files/styles/rh-icon/public/Hall%20of%20Biodiversity.jpg?itok=Hd9QNgWM)"><a href="/commentary/life-sciences/biodiversity-and-its-histories">Biodiversity and its histories</a></li> </ul></div> </div> </div> <div id="block-cudl-related-letters" class="block block-cudl"> <div class="campl-content-container campl-no-bottom-padding"> <div class="campl-heading-container"> <h2>Related letters</h2> </div> </div> <div class="campl-content-container"> <div class="letter even"> <h3><a href="/letter/?docId=letters/DCP-LETT-1221.xml">To H. E. Strickland [4 February 1849]</a></h3> <div class="summary"><p>HES’s arguments are of great weight, but CD cannot yet bring himself to reject well-known names for obscure ones. Sends four cases that he thinks will stagger HES. Cites his problems in classifying cirripedes. CD cannot bear to give new names, yet may do wrong to attach old ones. Not one species is correctly defined. The harm done by "species mongers".</p></div> </div><div class="letter odd"> <h3><a href="/letter/?docId=letters/DCP-LETT-2022.xml">To J. D. Hooker 24 December [1856]</a></h3> <div class="summary"><p>On the variety of species definitions prevalent among naturalists.</p></div> </div><div class="letter even"> <h3><a href="/letter/?docId=letters/DCP-LETT-3855.xml">To J. D. Hooker 12 [December 1862]</a></h3> <div class="summary"><p>Maintains his view on crossing. Thinks practical breeders would agree with him; doubts that variability and domestication are at all necessarily correlative.</p><p>Identical plants in different conditions a heavy argument against "direct action" [of physical conditions].</p><p>His 1000-pigeon case is altered if long-beaked are in least degree sterile with short-beaked.</p><p>His work on dimorphism inclines him to believe that sterility is at first a selected quality to keep incipient species distinct.</p><p>Case of easy modification of <i>Lythrum</i> pollen to favour or prevent crossing.</p><p>Monsters.</p><p>Has just finished chapter on variations of cultivated plants.</p><p>Edinburgh doctors have sent him Diploma of Medical Society.</p></div> </div><div class="letter odd"> <h3><a href="/letter/?docId=letters/DCP-LETT-5966.xml">From A. R. Wallace 1 March 1868</a></h3> <div class="summary"><p>Offers enclosure demonstrating that natural selection could produce sterility of hybrids.</p><p>More on Pangenesis and the inadequacy of H. Spencer’s approach.</p></div> </div><div class="letter even"> <h3><a href="/letter/?docId=letters/DCP-LETT-6104.xml">From A. R. Wallace 8 [April] 1868</a></h3> <div class="summary"><p>If CD is not convinced by his notes on sterility, ARW has little doubt that he is wrong. In fact he was only <i>half</i>-convinced by his own arguments.</p><p>Modifies his first proposition [a species varies occasionally in two directions, but owing to free inter-crossing the variations never increase] and further discusses the subject.</p><p>Encloses Berthold Seemann’s notes on flora of the Hawaiian Islands. Presence of European alpine species in Hawaiian volcanoes is a "hard nut" for geographical distribution [but see ARW’s <i>Island life</i> (1880), p. 323].</p></div> </div> </div> </div> <div id="block-views-my-sidebar-block" class="block block-views"> <div> <div class="view view-my-sidebar view-id-my_sidebar view-display-id-block view-dom-id-762dbbd0f61e052829691ade6c06389f"> <div class="view-content"> <div class="views-row views-row-1 views-row-odd views-row-first views-row-last"> <div class="views-field views-field-field-side-text"> <div class="field-content"></div> </div> </div> </div> </div> </div> </div> <div id="block-block-57" class="block block-block"> <div class="campl-content-container campl-no-bottom-padding"> <div class="campl-heading-container"> <h2>Darwin's letters: a timeline</h2> </div> </div> <div class="campl-content-container"> <p><a href="/letters/darwins-letters-timeline"><img alt="Timeline of letters to and from represented as a chart" src="/sites/all/modules/darwin_letter_timeline/timeline-promo.png" /></a></p> <p><a href="/letters/darwins-letters-timeline">Explore the letters to and from Charles Darwin over time</a></p> </div> </div> <div id="block-darwin-sharing-darwin-sharing-follow" class="block block-darwin-sharing"> <div class="campl-content-container">  <div class="social-media-follow"><a class="icon-sm darwin-twitter" href="https://twitter.com/mydeardarwin?lang=en" target="_blank"><i class="fab fa-twitter"></i> Follow on Twitter</a> <a class="icon-sm darwin-facebook" href="https://www.facebook.com/MyDearDarwin/" target="_blank"><i class="fab fa-facebook-f"></i> Follow on Facebook</a> <a class="icon-sm darwin-instagram" href="https://www.instagram.com/mydeardarwin/"><i class="fab fa-instagram"></i> Follow on Instagram</a></div>  </div> </div> </div> </div> </div> </div> <div class="campl-row campl-local-footer"> <div class="campl-wrap clearfix"> <div class="campl-column3 campl-footer-navigation"> <div 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