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糖ヌクレオチド - Wikipedia
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id="mw-navigation"> <nav id="mw-panel" class="vector-main-menu-landmark" aria-label="サイト"> <div id="vector-main-menu-pinned-container" class="vector-pinned-container"> </div> </nav> </div> </div> <div class="vector-sticky-pinned-container"> <nav id="mw-panel-toc" aria-label="目次" data-event-name="ui.sidebar-toc" class="mw-table-of-contents-container vector-toc-landmark"> <div id="vector-toc-pinned-container" class="vector-pinned-container"> <div id="vector-toc" class="vector-toc vector-pinnable-element"> <div class="vector-pinnable-header vector-toc-pinnable-header vector-pinnable-header-pinned" data-feature-name="toc-pinned" data-pinnable-element-id="vector-toc" > <h2 class="vector-pinnable-header-label">目次</h2> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-toc.pin">サイドバーに移動</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-toc.unpin">非表示</button> </div> <ul class="vector-toc-contents" id="mw-panel-toc-list"> <li id="toc-mw-content-text" class="vector-toc-list-item vector-toc-level-1"> <a href="#" class="vector-toc-link"> <div class="vector-toc-text">ページ先頭</div> </a> </li> <li id="toc-歴史" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#歴史"> <div class="vector-toc-text"> <span class="vector-toc-numb">1</span> <span>歴史</span> </div> </a> <ul id="toc-歴史-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-生化学的重要性" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#生化学的重要性"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>生化学的重要性</span> </div> </a> <ul id="toc-生化学的重要性-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-種類" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#種類"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>種類</span> </div> </a> <ul id="toc-種類-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-脚注" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#脚注"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>脚注</span> </div> </a> <ul id="toc-脚注-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="目次" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="目次の表示・非表示を切り替え" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">目次の表示・非表示を切り替え</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">糖ヌクレオチド</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="特定の記事の別の言語版に移動します。 利用可能な言語8件" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-8" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">8の言語版</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Nucleotide_sugar" title="英語: Nucleotide sugar" lang="en" hreflang="en" data-title="Nucleotide sugar" data-language-autonym="English" data-language-local-name="英語" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Nucl%C3%A9otide-ose" title="フランス語: Nucléotide-ose" lang="fr" hreflang="fr" data-title="Nucléotide-ose" data-language-autonym="Français" data-language-local-name="フランス語" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Nucle%C3%B3tido_azucre" title="ガリシア語: Nucleótido azucre" lang="gl" hreflang="gl" data-title="Nucleótido azucre" data-language-autonym="Galego" data-language-local-name="ガリシア語" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EB%8B%B9%EB%89%B4%ED%81%B4%EB%A0%88%EC%98%A4%ED%83%80%EC%9D%B4%EB%93%9C" title="韓国語: 당뉴클레오타이드" lang="ko" hreflang="ko" data-title="당뉴클레오타이드" data-language-autonym="한국어" data-language-local-name="韓国語" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/A%C3%A7%C3%BAcar_nucleot%C3%ADdeo" title="ポルトガル語: Açúcar nucleotídeo" lang="pt" hreflang="pt" data-title="Açúcar nucleotídeo" data-language-autonym="Português" data-language-local-name="ポルトガル語" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/Nukleotidni_%C5%A1e%C4%87er" title="セルボ・クロアチア語: Nukleotidni šećer" lang="sh" hreflang="sh" data-title="Nukleotidni šećer" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="セルボ・クロアチア語" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Nukleotidsacharid" title="スロバキア語: Nukleotidsacharid" lang="sk" hreflang="sk" data-title="Nukleotidsacharid" data-language-autonym="Slovenčina" data-language-local-name="スロバキア語" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/Nukleotidni_%C5%A1e%C4%87er" title="セルビア語: Nukleotidni šećer" lang="sr" hreflang="sr" data-title="Nukleotidni šećer" data-language-autonym="Српски / srpski" data-language-local-name="セルビア語" class="interlanguage-link-target"><span>Српски / srpski</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q117973#sitelinks-wikipedia" title="言語間リンクを編集" class="wbc-editpage">リンクを編集</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="名前空間"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/%E7%B3%96%E3%83%8C%E3%82%AF%E3%83%AC%E3%82%AA%E3%83%81%E3%83%89" title="本文を閲覧 [c]" accesskey="c"><span>ページ</span></a></li><li id="ca-talk" class="new vector-tab-noicon mw-list-item"><a href="/w/index.php?title=%E3%83%8E%E3%83%BC%E3%83%88:%E7%B3%96%E3%83%8C%E3%82%AF%E3%83%AC%E3%82%AA%E3%83%81%E3%83%89&action=edit&redlink=1" rel="discussion" class="new" title="「本文ページについての議論」 (存在しないページ) [t]" accesskey="t"><span>ノート</span></a></li> </ul> </div> </div> <div id="vector-variants-dropdown" class="vector-dropdown emptyPortlet" > <input type="checkbox" id="vector-variants-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-variants-dropdown" class="vector-dropdown-checkbox " aria-label="別の言語に切り替える" > <label id="vector-variants-dropdown-label" for="vector-variants-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">日本語</span> </label> <div class="vector-dropdown-content"> <div id="p-variants" class="vector-menu mw-portlet mw-portlet-variants emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> </div> </div> </nav> </div> <div id="right-navigation" class="vector-collapsible"> <nav aria-label="表示"> <div id="p-views" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-views" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-view" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/%E7%B3%96%E3%83%8C%E3%82%AF%E3%83%AC%E3%82%AA%E3%83%81%E3%83%89"><span>閲覧</span></a></li><li id="ca-edit" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=%E7%B3%96%E3%83%8C%E3%82%AF%E3%83%AC%E3%82%AA%E3%83%81%E3%83%89&action=edit" title="このページのソースコードを編集する [e]" accesskey="e"><span>編集</span></a></li><li id="ca-history" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=%E7%B3%96%E3%83%8C%E3%82%AF%E3%83%AC%E3%82%AA%E3%83%81%E3%83%89&action=history" title="このページの過去の版 [h]" accesskey="h"><span>履歴表示</span></a></li> </ul> </div> </div> </nav> <nav class="vector-page-tools-landmark" aria-label="ページツール"> <div id="vector-page-tools-dropdown" class="vector-dropdown vector-page-tools-dropdown" > <input type="checkbox" id="vector-page-tools-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-tools-dropdown" class="vector-dropdown-checkbox " aria-label="ツール" > <label id="vector-page-tools-dropdown-label" for="vector-page-tools-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">ツール</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-tools-unpinned-container" class="vector-unpinned-container"> <div id="vector-page-tools" class="vector-page-tools vector-pinnable-element"> <div class="vector-pinnable-header vector-page-tools-pinnable-header vector-pinnable-header-unpinned" data-feature-name="page-tools-pinned" data-pinnable-element-id="vector-page-tools" data-pinned-container-id="vector-page-tools-pinned-container" data-unpinned-container-id="vector-page-tools-unpinned-container" > <div class="vector-pinnable-header-label">ツール</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-page-tools.pin">サイドバーに移動</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-page-tools.unpin">非表示</button> </div> <div id="p-cactions" class="vector-menu mw-portlet mw-portlet-cactions emptyPortlet vector-has-collapsible-items" title="その他の操作" > <div class="vector-menu-heading"> 操作 </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-more-view" class="selected vector-more-collapsible-item mw-list-item"><a href="/wiki/%E7%B3%96%E3%83%8C%E3%82%AF%E3%83%AC%E3%82%AA%E3%83%81%E3%83%89"><span>閲覧</span></a></li><li id="ca-more-edit" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=%E7%B3%96%E3%83%8C%E3%82%AF%E3%83%AC%E3%82%AA%E3%83%81%E3%83%89&action=edit" title="このページのソースコードを編集する [e]" accesskey="e"><span>編集</span></a></li><li id="ca-more-history" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=%E7%B3%96%E3%83%8C%E3%82%AF%E3%83%AC%E3%82%AA%E3%83%81%E3%83%89&action=history"><span>履歴表示</span></a></li> </ul> </div> </div> <div id="p-tb" class="vector-menu mw-portlet mw-portlet-tb" > <div class="vector-menu-heading"> 全般 </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="t-whatlinkshere" class="mw-list-item"><a href="/wiki/%E7%89%B9%E5%88%A5:%E3%83%AA%E3%83%B3%E3%82%AF%E5%85%83/%E7%B3%96%E3%83%8C%E3%82%AF%E3%83%AC%E3%82%AA%E3%83%81%E3%83%89" title="ここにリンクしている全ウィキページの一覧 [j]" accesskey="j"><span>リンク元</span></a></li><li id="t-recentchangeslinked" class="mw-list-item"><a href="/wiki/%E7%89%B9%E5%88%A5:%E9%96%A2%E9%80%A3%E3%83%9A%E3%83%BC%E3%82%B8%E3%81%AE%E6%9B%B4%E6%96%B0%E7%8A%B6%E6%B3%81/%E7%B3%96%E3%83%8C%E3%82%AF%E3%83%AC%E3%82%AA%E3%83%81%E3%83%89" rel="nofollow" title="このページからリンクしているページの最近の更新 [k]" accesskey="k"><span>関連ページの更新状況</span></a></li><li id="t-upload" class="mw-list-item"><a href="/wiki/Wikipedia:%E3%83%95%E3%82%A1%E3%82%A4%E3%83%AB%E3%81%AE%E3%82%A2%E3%83%83%E3%83%97%E3%83%AD%E3%83%BC%E3%83%89" title="ファイルをアップロードする [u]" accesskey="u"><span>ファイルをアップロード</span></a></li><li id="t-specialpages" class="mw-list-item"><a href="/wiki/%E7%89%B9%E5%88%A5:%E7%89%B9%E5%88%A5%E3%83%9A%E3%83%BC%E3%82%B8%E4%B8%80%E8%A6%A7" title="特別ページの一覧 [q]" accesskey="q"><span>特別ページ</span></a></li><li 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href="https://www.wikidata.org/wiki/Special:EntityPage/Q117973" title="関連付けられたデータリポジトリ項目へのリンク [g]" accesskey="g"><span>ウィキデータ項目</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> </div> </div> </div> <div class="vector-column-end"> <div class="vector-sticky-pinned-container"> <nav class="vector-page-tools-landmark" aria-label="ページツール"> <div id="vector-page-tools-pinned-container" class="vector-pinned-container"> </div> </nav> <nav class="vector-appearance-landmark" aria-label="表示"> <div id="vector-appearance-pinned-container" class="vector-pinned-container"> <div id="vector-appearance" class="vector-appearance vector-pinnable-element"> <div class="vector-pinnable-header vector-appearance-pinnable-header vector-pinnable-header-pinned" data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">表示</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">サイドバーに移動</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">非表示</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">出典: フリー百科事典『ウィキペディア(Wikipedia)』</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="ja" dir="ltr"><p><b>糖ヌクレオチド</b>は<a href="/wiki/%E4%BB%A3%E8%AC%9D" title="代謝">代謝</a>における<a href="/wiki/%E5%8D%98%E7%B3%96" title="単糖">単糖</a>の活性型である。糖ヌクレオチドはグリコシル化では糖の供給源となり、この反応は<a href="/wiki/%E3%82%B0%E3%83%AA%E3%82%B3%E3%82%B7%E3%83%AB%E3%83%88%E3%83%A9%E3%83%B3%E3%82%B9%E3%83%95%E3%82%A7%E3%83%A9%E3%83%BC%E3%82%BC" title="グリコシルトランスフェラーゼ">グリコシルトランスフェラーゼ</a>によって触媒される。 </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="歴史"><span id=".E6.AD.B4.E5.8F.B2"></span>歴史</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%E7%B3%96%E3%83%8C%E3%82%AF%E3%83%AC%E3%82%AA%E3%83%81%E3%83%89&action=edit&section=1" title="節を編集: 歴史"><span>編集</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>オリゴ糖の同化作用と糖ヌクレオチドの役割は、1950年代にルロアールらがグリコシルトランスフェラーゼを発見したことにより明かになった。グリコシルトランスフェラーゼはグリコシル基を糖ヌクレオチドから受容体に転移させる酵素である<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>。 </p> <div class="mw-heading mw-heading2"><h2 id="生化学的重要性"><span id=".E7.94.9F.E5.8C.96.E5.AD.A6.E7.9A.84.E9.87.8D.E8.A6.81.E6.80.A7"></span>生化学的重要性</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%E7%B3%96%E3%83%8C%E3%82%AF%E3%83%AC%E3%82%AA%E3%83%81%E3%83%89&action=edit&section=2" title="節を編集: 生化学的重要性"><span>編集</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>グリコシル基供与体として作用するためにはその単糖が高いエネルギーを持たなければならないため、糖ヌクレオチドとなる必要がある。この糖ヌクレオチドはヌクレオチド三リン酸とグリコシル一リン酸との反応で合成される。 </p> <figure class="mw-halign-center" typeof="mw:File"><a href="/wiki/%E3%83%95%E3%82%A1%E3%82%A4%E3%83%AB:Activatedmonosaccahride.png" class="mw-file-description" title="Activation of Monosaccharides"><img alt="Activation of Monosaccharides" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/90/Activatedmonosaccahride.png/600px-Activatedmonosaccahride.png" decoding="async" width="600" height="79" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/90/Activatedmonosaccahride.png/900px-Activatedmonosaccahride.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/90/Activatedmonosaccahride.png/1200px-Activatedmonosaccahride.png 2x" data-file-width="2219" data-file-height="293" /></a><figcaption>Activation of Monosaccharides</figcaption></figure> <div class="mw-heading mw-heading2"><h2 id="種類"><span id=".E7.A8.AE.E9.A1.9E"></span>種類</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%E7%B3%96%E3%83%8C%E3%82%AF%E3%83%AC%E3%82%AA%E3%83%81%E3%83%89&action=edit&section=3" title="節を編集: 種類"><span>編集</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>動物にはグリコシル基供与体として9種の糖ヌクレオチドが存在し、ヌクレオシドの違いによって分類することができる<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup>。 </p> <ul><li><a href="/wiki/%E3%82%A6%E3%83%AA%E3%82%B8%E3%83%B3%E4%BA%8C%E3%83%AA%E3%83%B3%E9%85%B8" title="ウリジン二リン酸">ウリジン二リン酸</a> - <a href="/wiki/%E3%82%A6%E3%83%AA%E3%82%B8%E3%83%B3%E4%BA%8C%E3%83%AA%E3%83%B3%E9%85%B8%E3%82%B0%E3%83%AB%E3%82%B3%E3%83%BC%E3%82%B9" title="ウリジン二リン酸グルコース">ウリジン二リン酸グルコース</a>、<a href="/wiki/%E3%82%A6%E3%83%AA%E3%82%B8%E3%83%B3%E4%BA%8C%E3%83%AA%E3%83%B3%E9%85%B8%E3%82%AC%E3%83%A9%E3%82%AF%E3%83%88%E3%83%BC%E3%82%B9" title="ウリジン二リン酸ガラクトース">ウリジン二リン酸ガラクトース</a>、<a href="/wiki/%E3%82%A6%E3%83%AA%E3%82%B8%E3%83%B3%E4%BA%8C%E3%83%AA%E3%83%B3%E9%85%B8-N-%E3%82%A2%E3%82%BB%E3%83%81%E3%83%AB%E3%82%B0%E3%83%AB%E3%82%B3%E3%82%B5%E3%83%9F%E3%83%B3" title="ウリジン二リン酸-N-アセチルグルコサミン">ウリジン二リン酸-N-アセチルグルコサミン</a>、<a href="/wiki/%E3%82%A6%E3%83%AA%E3%82%B8%E3%83%B3%E4%BA%8C%E3%83%AA%E3%83%B3%E9%85%B8%E3%82%B0%E3%83%AB%E3%82%AF%E3%83%AD%E3%83%B3%E9%85%B8" title="ウリジン二リン酸グルクロン酸">ウリジン二リン酸グルクロン酸</a>、<a href="/w/index.php?title=%E3%82%A6%E3%83%AA%E3%82%B8%E3%83%B3%E4%BA%8C%E3%83%AA%E3%83%B3%E9%85%B8%E3%82%AD%E3%82%B7%E3%83%AD%E3%83%BC%E3%82%B9&action=edit&redlink=1" class="new" title="「ウリジン二リン酸キシロース」 (存在しないページ)">ウリジン二リン酸キシロース</a></li> <li><a href="/wiki/%E3%82%B0%E3%82%A2%E3%83%8E%E3%82%B7%E3%83%B3%E4%BA%8C%E3%83%AA%E3%83%B3%E9%85%B8" title="グアノシン二リン酸">グアノシン二リン酸</a> - <a href="/w/index.php?title=%E3%82%B0%E3%82%A2%E3%83%8E%E3%82%B7%E3%83%B3%E4%BA%8C%E3%83%AA%E3%83%B3%E9%85%B8%E3%83%9E%E3%83%B3%E3%83%8E%E3%83%BC%E3%82%B9&action=edit&redlink=1" class="new" title="「グアノシン二リン酸マンノース」 (存在しないページ)">グアノシン二リン酸マンノース</a>、<a href="/w/index.php?title=%E3%82%B0%E3%82%A2%E3%83%8E%E3%82%B7%E3%83%B3%E4%BA%8C%E3%83%AA%E3%83%B3%E9%85%B8%E3%83%95%E3%82%B3%E3%83%BC%E3%82%B9&action=edit&redlink=1" class="new" title="「グアノシン二リン酸フコース」 (存在しないページ)">グアノシン二リン酸フコース</a></li> <li><a href="/wiki/%E3%82%B7%E3%83%81%E3%82%B8%E3%83%B3%E4%B8%80%E3%83%AA%E3%83%B3%E9%85%B8" class="mw-redirect" title="シチジン一リン酸">シチジン一リン酸</a> - <a href="/w/index.php?title=%E3%82%B7%E3%83%81%E3%82%B8%E3%83%B3%E4%B8%80%E3%83%AA%E3%83%B3%E9%85%B8-N-%E3%82%A2%E3%82%BB%E3%83%81%E3%83%AB%E3%83%8E%E3%82%A4%E3%83%A9%E3%83%9F%E3%83%B3%E9%85%B8&action=edit&redlink=1" class="new" title="「シチジン一リン酸-N-アセチルノイラミン酸」 (存在しないページ)">シチジン一リン酸-N-アセチルノイラミン酸</a></li></ul> <p>植物およびバクテリアでは他にも多くの糖が用いられる。特に<a href="/w/index.php?title=%E3%82%B7%E3%83%81%E3%82%B8%E3%83%B3%E4%BA%8C%E3%83%AA%E3%83%B3%E9%85%B8%E3%82%B0%E3%83%AB%E3%82%B3%E3%83%BC%E3%82%B9&action=edit&redlink=1" class="new" title="「シチジン二リン酸グルコース」 (存在しないページ)">シチジン二リン酸グルコース</a>と<a href="/w/index.php?title=%E3%83%81%E3%83%9F%E3%82%B8%E3%83%B3%E4%BA%8C%E3%83%AA%E3%83%B3%E9%85%B8%E3%82%B0%E3%83%AB%E3%82%B3%E3%83%BC%E3%82%B9&action=edit&redlink=1" class="new" title="「チミジン二リン酸グルコース」 (存在しないページ)">チミジン二リン酸グルコース</a>が自然界に多く見られ、他にも多くのCDP, TDP-糖が供与体ヌクレオチドとして存在する<sup id="cite_ref-Reeves_3-0" class="reference"><a href="#cite_note-Reeves-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Liu_4-0" class="reference"><a href="#cite_note-Liu-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> 。 </p> <div class="mw-heading mw-heading2"><h2 id="脚注"><span id=".E8.84.9A.E6.B3.A8"></span>脚注</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=%E7%B3%96%E3%83%8C%E3%82%AF%E3%83%AC%E3%82%AA%E3%83%81%E3%83%89&action=edit&section=4" title="節を編集: 脚注"><span>編集</span></a><span class="mw-editsection-bracket">]</span></span></div> <ol class="references"> <li id="cite_note-1"><b><a href="#cite_ref-1">^</a></b> <span class="reference-text"><cite style="font-style:normal" class="citation journal">Derek Horton (2008). “The Development of Carbohydrate Chemistry and Biology”. <i>Carbohydrate Chemistry, Biology and Medical Applications</i>: 1–28. <a href="/wiki/Doi_(%E8%AD%98%E5%88%A5%E5%AD%90)" class="mw-redirect" title="Doi (識別子)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2FB978-0-08-054816-6.00001-X">10.1016/B978-0-08-054816-6.00001-X</a>.</cite><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=The+Development+of+Carbohydrate+Chemistry+and+Biology&rft.jtitle=Carbohydrate+Chemistry%2C+Biology+and+Medical+Applications&rft.aulast=Derek+Horton&rft.au=Derek+Horton&rft.date=2008&rft.pages=1%E2%80%9328&rft_id=info:doi/10.1016%2FB978-0-08-054816-6.00001-X&rfr_id=info:sid/ja.wikipedia.org:%E7%B3%96%E3%83%8C%E3%82%AF%E3%83%AC%E3%82%AA%E3%83%81%E3%83%89"><span style="display: none;"> </span></span></span> </li> <li id="cite_note-2"><b><a href="#cite_ref-2">^</a></b> <span class="reference-text"> <a rel="nofollow" class="external text" href="http://www.cshlpress.com/default.tpl?action=full&--eqskudatarq=666">Cold Spring Harbor Laboratory Press</a> Essentials of Glycobiology, Second Edition</span> </li> <li id="cite_note-Reeves-3"><b><a href="#cite_ref-Reeves_3-0">^</a></b> <span class="reference-text"><cite style="font-style:normal" class="citation journal">Samuel G, Reeves P (2003). “Biosynthesis of O-antigens: genes and pathways involved in nucleotide sugar precursor synthesis and O-antigen assembly”. <i>Carbohydr. Res.</i> <b>338</b> (23): 2503–19. <a href="/wiki/Doi_(%E8%AD%98%E5%88%A5%E5%AD%90)" class="mw-redirect" title="Doi (識別子)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.carres.2003.07.009">10.1016/j.carres.2003.07.009</a>. <style data-mw-deduplicate="TemplateStyles:r101121245">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation.cs-ja1 q,.mw-parser-output .citation.cs-ja2 q{quotes:"「""」""『""』"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free a,.mw-parser-output .citation .cs1-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited a,.mw-parser-output .id-lock-registration a,.mw-parser-output .citation .cs1-lock-limited a,.mw-parser-output .citation .cs1-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription a,.mw-parser-output .citation .cs1-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:var(--color-success,#3a3);margin-left:0.3em}.mw-parser-output .cs1-format{font-size:95%}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}</style><a href="/wiki/PMID_(%E8%AD%98%E5%88%A5%E5%AD%90)" class="mw-redirect" title="PMID (識別子)">PMID</a> <a rel="nofollow" class="external text" href="//pubmed.ncbi.nlm.nih.gov/14670712/">14670712</a>.</cite><span class="Z3988" title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=Biosynthesis+of+O-antigens%3A+genes+and+pathways+involved+in+nucleotide+sugar+precursor+synthesis+and+O-antigen+assembly&rft.jtitle=Carbohydr.+Res.&rft.aulast=Samuel+G%2C+Reeves+P&rft.au=Samuel+G%2C+Reeves+P&rft.date=2003&rft.volume=338&rft.issue=23&rft.pages=2503%E2%80%9319&rft_id=info:doi/10.1016%2Fj.carres.2003.07.009&rft_id=info:pmid/14670712&rfr_id=info:sid/ja.wikipedia.org:%E7%B3%96%E3%83%8C%E3%82%AF%E3%83%AC%E3%82%AA%E3%83%81%E3%83%89"><span style="display: none;"> </span></span></span> </li> <li id="cite_note-Liu-4"><b><a href="#cite_ref-Liu_4-0">^</a></b> <span class="reference-text"><cite style="font-style:normal" class="citation journal">Xue M. 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