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Available in 17 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-17" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">17 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D9%83%D9%8A%D8%AB_%D9%83%D8%A7%D9%85%D8%A8%D9%84" title="كيث كامبل – Arabic" lang="ar" hreflang="ar" data-title="كيث كامبل" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Keith_Campbell_(biolog)" title="Keith Campbell (biolog) – Danish" lang="da" hreflang="da" data-title="Keith Campbell (biolog)" data-language-autonym="Dansk" data-language-local-name="Danish" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Keith_Campbell_(Biologe)" title="Keith Campbell (Biologe) – German" lang="de" hreflang="de" data-title="Keith Campbell (Biologe)" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Keith_Campbell_(bi%C3%B3logo)" title="Keith Campbell (biólogo) – Spanish" lang="es" hreflang="es" data-title="Keith Campbell (biólogo)" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Keith_Campbell" title="Keith Campbell – French" lang="fr" hreflang="fr" data-title="Keith Campbell" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%ED%82%A4%EC%8A%A4_%EC%BA%A0%EB%B2%A8_(%EC%83%9D%EB%AC%BC%ED%95%99%EC%9E%90)" title="키스 캠벨 (생물학자) – Korean" lang="ko" hreflang="ko" data-title="키스 캠벨 (생물학자)" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Keith_Campbell_(ahli_biologi)" title="Keith Campbell (ahli biologi) – Indonesian" lang="id" hreflang="id" data-title="Keith Campbell (ahli biologi)" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-mg mw-list-item"><a href="https://mg.wikipedia.org/wiki/Keith_Campbell" title="Keith Campbell – Malagasy" lang="mg" hreflang="mg" data-title="Keith Campbell" data-language-autonym="Malagasy" data-language-local-name="Malagasy" class="interlanguage-link-target"><span>Malagasy</span></a></li><li class="interlanguage-link interwiki-xmf mw-list-item"><a href="https://xmf.wikipedia.org/wiki/%E1%83%99%E1%83%98%E1%83%A2_%E1%83%99%E1%83%94%E1%83%9B%E1%83%9E%E1%83%91%E1%83%94%E1%83%9A%E1%83%98" title="კიტ კემპბელი – Mingrelian" lang="xmf" hreflang="xmf" data-title="კიტ კემპბელი" data-language-autonym="მარგალური" data-language-local-name="Mingrelian" class="interlanguage-link-target"><span>მარგალური</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D9%83%D9%8A%D8%AB_%D9%83%D8%A7%D9%85%D8%A8%D9%84_(%D8%A7%D8%B3%D8%AA%D8%A7%D8%B0_%D8%AC%D8%A7%D9%85%D8%B9%D8%A9_%D9%85%D9%86_%D8%A7%D9%84%D9%85%D9%85%D9%84%D9%83%D9%87_%D8%A7%D9%84%D9%85%D8%AA%D8%AD%D8%AF%D9%87)" title="كيث كامبل (استاذ جامعة من المملكه المتحده) – Egyptian Arabic" lang="arz" hreflang="arz" data-title="كيث كامبل (استاذ جامعة من المملكه المتحده)" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Keith_Campbell_(bioloog)" title="Keith Campbell (bioloog) – Dutch" lang="nl" hreflang="nl" data-title="Keith Campbell (bioloog)" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%82%AD%E3%83%BC%E3%82%B9%E3%83%BB%E3%82%AD%E3%83%A3%E3%83%B3%E3%83%99%E3%83%AB_(%E7%94%9F%E7%89%A9%E5%AD%A6%E8%80%85)" title="キース・キャンベル (生物学者) – Japanese" lang="ja" hreflang="ja" data-title="キース・キャンベル (生物学者)" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Keith_Campbell" title="Keith Campbell – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Keith Campbell" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Keith_Campbell" title="Keith Campbell – Polish" lang="pl" hreflang="pl" data-title="Keith Campbell" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Keith_Campbell" title="Keith Campbell – Portuguese" lang="pt" hreflang="pt" data-title="Keith Campbell" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%9A%D1%8D%D0%BC%D0%BF%D0%B1%D0%B5%D0%BB%D0%BB,_%D0%9A%D0%B8%D1%82" title="Кэмпбелл, Кит – Russian" lang="ru" hreflang="ru" data-title="Кэмпбелл, Кит" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E5%9F%BA%E6%80%9D%C2%B7%E5%9D%8E%E8%B4%9D%E5%B0%94" title="基思·坎贝尔 – Chinese" lang="zh" hreflang="zh" data-title="基思·坎贝尔" 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<div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">British biologist and university professor (1954–2012)</div> <p class="mw-empty-elt"> </p> <style data-mw-deduplicate="TemplateStyles:r1257001546">.mw-parser-output .infobox-subbox{padding:0;border:none;margin:-3px;width:auto;min-width:100%;font-size:100%;clear:none;float:none;background-color:transparent}.mw-parser-output .infobox-3cols-child{margin:auto}.mw-parser-output .infobox .navbar{font-size:100%}@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox-full-data:not(.notheme)>div:not(.notheme)[style]{background:#1f1f23!important;color:#f8f9fa}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox-full-data:not(.notheme) div:not(.notheme){background:#1f1f23!important;color:#f8f9fa}}@media(min-width:640px){body.skin--responsive .mw-parser-output .infobox-table{display:table!important}body.skin--responsive .mw-parser-output .infobox-table>caption{display:table-caption!important}body.skin--responsive .mw-parser-output .infobox-table>tbody{display:table-row-group}body.skin--responsive .mw-parser-output .infobox-table tr{display:table-row!important}body.skin--responsive .mw-parser-output .infobox-table th,body.skin--responsive .mw-parser-output .infobox-table td{padding-left:inherit;padding-right:inherit}}</style><table class="infobox biography vcard"><tbody><tr><th colspan="2" class="infobox-above" style="font-size:125%;"><div class="fn">Keith Campbell</div></th></tr><tr><th scope="row" class="infobox-label">Born</th><td class="infobox-data"><div style="display:inline" class="nickname">Keith Henry Stockman Campbell</div><br /><span style="display:none">(<span class="bday">1954-05-23</span>)</span>23 May 1954<br /><div style="display:inline" class="birthplace"><a href="/wiki/Birmingham" title="Birmingham">Birmingham</a>, England</div></td></tr><tr><th scope="row" class="infobox-label">Died</th><td class="infobox-data">5 October 2012<span style="display:none">(2012-10-05)</span> (aged 58)<br /><div style="display:inline" class="deathplace"><a href="/wiki/Ingleby,_Derbyshire" title="Ingleby, Derbyshire">Ingleby, Derbyshire</a>, England</div></td></tr><tr><th scope="row" class="infobox-label">Alma mater</th><td class="infobox-data"><a href="/wiki/Queen_Elizabeth_College" title="Queen Elizabeth College">Queen Elizabeth College</a>,BSc (now part of <a href="/wiki/King%27s_College_London" title="King's College London">King's College London</a>)<br /><a href="/wiki/University_of_Sussex" title="University of Sussex">University of Sussex</a>, PhD</td></tr><tr><th scope="row" class="infobox-label">Known for</th><td class="infobox-data"><a href="/wiki/Dolly_(sheep)" title="Dolly (sheep)">Dolly (sheep)</a> (1996)</td></tr><tr><th scope="row" class="infobox-label">Awards</th><td class="infobox-data"><a href="/wiki/Shaw_Prize" title="Shaw Prize">Shaw Prize</a> (2008) <br /> IETS Pioneer Award (2015)</td></tr><tr><td colspan="2" class="infobox-full-data"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1257001546"><b>Scientific career</b></td></tr><tr><th scope="row" class="infobox-label">Institutions</th><td class="infobox-data"><a href="/wiki/University_of_London" title="University of London">University of London</a><br /><a href="/wiki/University_of_Sussex" title="University of Sussex">University of Sussex</a><br /><a href="/wiki/Roslin_Institute" title="Roslin Institute">Roslin Institute</a><br /><a href="/wiki/University_of_Nottingham" title="University of Nottingham">University of Nottingham</a></td></tr><tr><th scope="row" class="infobox-label"><a href="/wiki/Thesis" title="Thesis">Thesis</a></th><td class="infobox-data"><i><a rel="nofollow" class="external text" href="https://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.512104">Aspects of cell cycle regulation in yeast and Xenopus</a></i> <span style="font-size:97%;">(1988)</span></td></tr><tr style="display:none"><td colspan="2"> </td></tr></tbody></table> <p><b>Keith Henry Stockman Campbell</b> (23 May 1954 – 5 October 2012)<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> was a British <a href="/wiki/Biologist" title="Biologist">biologist</a> who was a member of the team at <a href="/wiki/Roslin_Institute" title="Roslin Institute">Roslin Institute</a> that in 1996 first cloned a mammal, a <a href="/wiki/Finnish_Dorset_(sheep)" class="mw-redirect" title="Finnish Dorset (sheep)">Finnish Dorset</a> lamb named <a href="/wiki/Dolly_(sheep)" title="Dolly (sheep)">Dolly</a>, from fully differentiated adult mammary cells. He was Professor of Animal Development at the <a href="/wiki/University_of_Nottingham" title="University of Nottingham">University of Nottingham</a>.<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-natureobit_5-0" class="reference"><a href="#cite_note-natureobit-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-6" class="reference"><a href="#cite_note-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> In 2008, he received the <a href="/wiki/Shaw_Prize" title="Shaw Prize">Shaw Prize</a> for Medicine and Life Sciences jointly with <a href="/wiki/Ian_Wilmut" title="Ian Wilmut">Ian Wilmut</a> and <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Shinya Yamanaka</a> for "their works on the cell differentiation in mammals". </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Education">Education</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Keith_Campbell_(biologist)&action=edit&section=1" title="Edit section: Education"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Campbell was born in <a href="/wiki/Birmingham" title="Birmingham">Birmingham</a>, England, to an English mother and Scottish father. He started his education in <a href="/wiki/Perth,_Scotland" title="Perth, Scotland">Perth</a>, Scotland, but, when he was eight years old, his family returned to Birmingham, where he attended <a href="/wiki/King_Edward_VI_Camp_Hill_School_for_Boys" title="King Edward VI Camp Hill School for Boys">King Edward VI Camp Hill School for Boys</a>.<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> He obtained his Bachelor of Science degree in <a href="/wiki/Microbiology" title="Microbiology">microbiology</a> from the <a href="/wiki/Queen_Elizabeth_College" title="Queen Elizabeth College">Queen Elizabeth College</a>, <a href="/wiki/University_of_London" title="University of London">University of London</a> (now part of <a href="/wiki/King%27s_College_London" title="King's College London">King's College London</a>).<sup id="cite_ref-9" class="reference"><a href="#cite_note-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> In 1983 Campbell was awarded the <a href="/wiki/Marie_Curie" title="Marie Curie">Marie Curie</a> Research Scholarship, which led to postgraduate studies and later his PhD from the <a href="/wiki/University_of_Sussex" title="University of Sussex">University of Sussex</a><sup id="cite_ref-campbellphd_10-0" class="reference"><a href="#cite_note-campbellphd-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> (Brighton, England, UK).<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-13" class="reference"><a href="#cite_note-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Research_and_career">Research and career</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Keith_Campbell_(biologist)&action=edit&section=2" title="Edit section: Research and career"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Campbell's interest in cloning mammals was inspired by work done by Karl Illmensee and <a href="/wiki/John_Gurdon" title="John Gurdon">John Gurdon</a>.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (August 2016)">citation needed</span></a></i>]</sup> Working at the <a href="/wiki/Roslin_Institute" title="Roslin Institute">Roslin Institute</a> since 1991, Campbell became involved with the cloning efforts led by <a href="/wiki/Ian_Wilmut" title="Ian Wilmut">Ian Wilmut</a>. In July 1995 Keith Campbell and Bill Ritchie succeeded in producing a pair of lambs, Megan and Morag from embryonic cells, which had differentiated in culture. </p><p>In 1996, a team led by <a href="/wiki/Ian_Wilmut" title="Ian Wilmut">Ian Wilmut</a> with Keith Campbell as the main contributor, used the same technique and shocked the world by successfully cloning a sheep from adult mammary cells. Dolly, a Finn Dorset sheep named after the singer <a href="/wiki/Dolly_Parton" title="Dolly Parton">Dolly Parton</a>, was born in 1996 and lived to be six years old (dying from a viral infection and not old age, as has been suggested). Campbell had a key role in the creation of Dolly, as he had the crucial idea of co-ordinating the stages of the "cell cycle" of the donor somatic cells and the recipient eggs and using diploid quiescent or "G0" arrested somatic cells as nuclear donors. In 2006, <a href="/wiki/Ian_Wilmut" title="Ian Wilmut">Ian Wilmut</a> admitted that Campbell deserved "66 per cent" of the credit.<sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> </p><p>In 1997, Ritchie and Campbell in collaboration with PPL (Pharmaceutical Proteins Limited) created another sheep named "Polly", created from genetically altered skin cells containing a human gene.<sup id="cite_ref-15" class="reference"><a href="#cite_note-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> In 2000, after joining PPL Ltd, Campbell and his PPL team (based in North America) were successful in producing the world's first piglets by <a href="/wiki/Somatic-cell_nuclear_transfer" class="mw-redirect" title="Somatic-cell nuclear transfer">Somatic-cell nuclear transfer</a> (SCNT), the so-called cloning technique. Furthermore, the PPL teams based in Roslin, Scotland and Blacksburg (USA) used the technique to produce the first gene targeted domestic animals as well as a range of animals producing human therapeutic proteins in their milk.<sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> </p><p>From November 1999, Campbell held the post of Professor of Animal Development, Division of Animal Physiology, School of Biosciences at the <a href="/wiki/University_of_Nottingham" title="University of Nottingham">University of Nottingham</a> where he continued to study embryo growth and differentiation. He supported the use of SCNT for the production of personalised stem cell therapies and for the study of human diseases and the use of cybrid embryo production to overcome the lack of human eggs available for research. Stem cells can be isolated from embryonic, fetal and adult derived material and more recently by overexpression of certain genes for the production of "induced pluripotent cells". Campbell believed all potential stem cell populations should be used for both basic and applied research which may provide basic scientific knowledge and lead to the development of cell therapies.<sup id="cite_ref-18" class="reference"><a href="#cite_note-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Awards_and_honours">Awards and honours</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Keith_Campbell_(biologist)&action=edit&section=3" title="Edit section: Awards and honours"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>In 2008, he received the <a href="/wiki/Shaw_Prize" title="Shaw Prize">Shaw Prize</a> for Medicine and Life Sciences jointly with Ian Wilmut and Shinya Yamanaka.<sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup> He was awarded the Pioneer Award from the International Embryo Transfer Society posthumously in 2015.<sup id="cite_ref-23" class="reference"><a href="#cite_note-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Personal_life">Personal life</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Keith_Campbell_(biologist)&action=edit&section=4" title="Edit section: Personal life"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Campbell died on 5 October 2012, aged 58, after accidentally hanging himself in his bedroom at his <a href="/wiki/Ingleby,_Derbyshire" title="Ingleby, Derbyshire">Ingleby, Derbyshire</a> home, whilst heavily intoxicated. It was determined at the inquest that he had been behaving erratically at the time and had no actual intention to kill himself; the verdict was a <a href="/wiki/Death_by_misadventure" title="Death by misadventure">death by misadventure</a>.<sup id="cite_ref-24" class="reference"><a href="#cite_note-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-25" class="reference"><a href="#cite_note-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup> He was buried at Bretby Crematorium, <a href="/wiki/Derbyshire" title="Derbyshire">Derbyshire</a>.<sup id="cite_ref-26" class="reference"><a href="#cite_note-26"><span class="cite-bracket">[</span>26<span class="cite-bracket">]</span></a></sup> He is survived by his wife, Kathy, and two daughters, Claire and Lauren. </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Keith_Campbell_(biologist)&action=edit&section=5" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist reflist-columns references-column-width" style="column-width: 30em;"> <ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFSylvia_Hui2012" class="citation news cs1">Sylvia Hui (11 October 2012). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20121015000131/http://www.startribune.com/lifestyle/173675601.html">"Keith Campbell, UK biologist who helped create cloned sheep Dolly, dies"</a>. <i><a href="/wiki/Star_Tribune" class="mw-redirect" title="Star Tribune">Star Tribune</a></i>. <a href="/wiki/Associated_Press" title="Associated Press">Associated Press</a>. Archived from <a rel="nofollow" class="external text" href="http://www.startribune.com/lifestyle/173675601.html">the original</a> on 15 October 2012<span class="reference-accessdate">. Retrieved <span class="nowrap">11 October</span> 2012</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Star+Tribune&rft.atitle=Keith+Campbell%2C+UK+biologist+who+helped+create+cloned+sheep+Dolly%2C+dies&rft.date=2012-10-11&rft.au=Sylvia+Hui&rft_id=http%3A%2F%2Fwww.startribune.com%2Flifestyle%2F173675601.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-2">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation news cs1"><a rel="nofollow" class="external text" href="https://www.telegraph.co.uk/news/obituaries/science-obituaries/9602836/Professor-Keith-Campbell.html">"Professor Keith Campbell"</a>. <i>The Daily Telegraph</i>. London. 11 October 2012.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+Daily+Telegraph&rft.atitle=Professor+Keith+Campbell&rft.date=2012-10-11&rft_id=https%3A%2F%2Fwww.telegraph.co.uk%2Fnews%2Fobituaries%2Fscience-obituaries%2F9602836%2FProfessor-Keith-Campbell.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-3">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFKindMcCreathHowcroftCampbell2000" class="citation journal cs1">Kind, A. J.; McCreath, K. J.; Howcroft, J.; <a class="mw-selflink selflink">Campbell, K. H. S.</a>; Colman, A.; Schnieke, A. E. (2000). "Production of gene-targeted sheep by nuclear transfer from cultured somatic cells". <i>Nature</i>. <b>405</b> (6790): 1066–1069. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2000Natur.405.1066M">2000Natur.405.1066M</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2F35016604">10.1038/35016604</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/10890449">10890449</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:4419356">4419356</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nature&rft.atitle=Production+of+gene-targeted+sheep+by+nuclear+transfer+from+cultured+somatic+cells&rft.volume=405&rft.issue=6790&rft.pages=1066-1069&rft.date=2000&rft_id=info%3Adoi%2F10.1038%2F35016604&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A4419356%23id-name%3DS2CID&rft_id=info%3Apmid%2F10890449&rft_id=info%3Abibcode%2F2000Natur.405.1066M&rft.aulast=Kind&rft.aufirst=A.+J.&rft.au=McCreath%2C+K.+J.&rft.au=Howcroft%2C+J.&rft.au=Campbell%2C+K.+H.+S.&rft.au=Colman%2C+A.&rft.au=Schnieke%2C+A.+E.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPolejaevaChenVaughtPage2000" class="citation journal cs1">Polejaeva, I. A.; Chen, S. H.; Vaught, T. D.; Page, R. L.; Mullins, J.; Ball, S.; Dai, Y.; Boone, J.; Walker, S.; Ayares, D. L.; Colman, A.; Campbell, K. H. S. (2000). "Cloned pigs produced by nuclear transfer from adult somatic cells". <i>Nature</i>. <b>407</b> (6800): 86–90. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2000Natur.407...86P">2000Natur.407...86P</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2F35024082">10.1038/35024082</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/10993078">10993078</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:4373081">4373081</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nature&rft.atitle=Cloned+pigs+produced+by+nuclear+transfer+from+adult+somatic+cells&rft.volume=407&rft.issue=6800&rft.pages=86-90&rft.date=2000&rft_id=info%3Adoi%2F10.1038%2F35024082&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A4373081%23id-name%3DS2CID&rft_id=info%3Apmid%2F10993078&rft_id=info%3Abibcode%2F2000Natur.407...86P&rft.aulast=Polejaeva&rft.aufirst=I.+A.&rft.au=Chen%2C+S.+H.&rft.au=Vaught%2C+T.+D.&rft.au=Page%2C+R.+L.&rft.au=Mullins%2C+J.&rft.au=Ball%2C+S.&rft.au=Dai%2C+Y.&rft.au=Boone%2C+J.&rft.au=Walker%2C+S.&rft.au=Ayares%2C+D.+L.&rft.au=Colman%2C+A.&rft.au=Campbell%2C+K.+H.+S.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-natureobit-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-natureobit_5-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFTrounson2012" class="citation journal cs1"><a href="/wiki/Alan_O._Trounson" class="mw-redirect" title="Alan O. Trounson">Trounson, A.</a> (2012). <a rel="nofollow" class="external text" href="https://doi.org/10.1038%2F491193a">"Keith H. Campbell (1954–2012)"</a>. <i>Nature</i>. <b>491</b> (7423): 193. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2012Natur.491..193T">2012Natur.491..193T</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1038%2F491193a">10.1038/491193a</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/23135461">23135461</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nature&rft.atitle=Keith+H.+Campbell+%281954%E2%80%932012%29&rft.volume=491&rft.issue=7423&rft.pages=193&rft.date=2012&rft_id=info%3Apmid%2F23135461&rft_id=info%3Adoi%2F10.1038%2F491193a&rft_id=info%3Abibcode%2F2012Natur.491..193T&rft.aulast=Trounson&rft.aufirst=A.&rft_id=https%3A%2F%2Fdoi.org%2F10.1038%252F491193a&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-6"><span class="mw-cite-backlink"><b><a href="#cite_ref-6">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWilmut2012" class="citation journal cs1"><a href="/wiki/Ian_Wilmut" title="Ian Wilmut">Wilmut, I.</a> (2012). <a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.1233495">"Keith Campbell (1954-2012)"</a>. <i>Science</i>. <b>338</b> (6114): 1553. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2012Sci...338.1553W">2012Sci...338.1553W</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.1233495">10.1126/science.1233495</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/23258883">23258883</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:206547014">206547014</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Science&rft.atitle=Keith+Campbell+%281954-2012%29&rft.volume=338&rft.issue=6114&rft.pages=1553&rft.date=2012&rft_id=info%3Adoi%2F10.1126%2Fscience.1233495&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A206547014%23id-name%3DS2CID&rft_id=info%3Apmid%2F23258883&rft_id=info%3Abibcode%2F2012Sci...338.1553W&rft.aulast=Wilmut&rft.aufirst=I.&rft_id=https%3A%2F%2Fdoi.org%2F10.1126%252Fscience.1233495&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-7">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20060303054113/http://www.nottingham.ac.uk/biosciences/anphy/academic/campbell.html">"Keith Campbell Profile with selected references"</a>. Archived from the original on 3 March 2006<span class="reference-accessdate">. Retrieved <span class="nowrap">6 June</span> 2008</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Keith+Campbell+Profile+with+selected+references&rft_id=http%3A%2F%2Fwww.nottingham.ac.uk%2Fbiosciences%2Fanphy%2Facademic%2Fcampbell.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span><span class="cs1-maint citation-comment"><code class="cs1-code">{{<a href="/wiki/Template:Cite_web" title="Template:Cite web">cite web</a>}}</code>: CS1 maint: bot: original URL status unknown (<a href="/wiki/Category:CS1_maint:_bot:_original_URL_status_unknown" title="Category:CS1 maint: bot: original URL status unknown">link</a>)</span></span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-8">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFIan_Wilmut2012" class="citation news cs1">Ian Wilmut (16 October 2012). <a rel="nofollow" class="external text" href="https://www.theguardian.com/science/2012/oct/16/keith-campbell">"Keith Campbell obituary"</a>. The Guardian, London<span class="reference-accessdate">. Retrieved <span class="nowrap">16 May</span> 2013</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=Keith+Campbell+obituary&rft.date=2012-10-16&rft.au=Ian+Wilmut&rft_id=https%3A%2F%2Fwww.theguardian.com%2Fscience%2F2012%2Foct%2F16%2Fkeith-campbell&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-9">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20151222084607/http://www.shawprize.org/en/shaw.php?tmp=3&twoid=49&threeid=56&fourid=72&fiveid=15">"Autobiography of Keith H S Campbell"</a>. The Shaw Prize Foundation. Archived from <a rel="nofollow" class="external text" href="http://www.shawprize.org/en/shaw.php?tmp=3&twoid=49&threeid=56&fourid=72&fiveid=15">the original</a> on 22 December 2015<span class="reference-accessdate">. Retrieved <span class="nowrap">13 December</span> 2015</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Autobiography+of+Keith+H+S+Campbell&rft.pub=The+Shaw+Prize+Foundation&rft_id=http%3A%2F%2Fwww.shawprize.org%2Fen%2Fshaw.php%3Ftmp%3D3%26twoid%3D49%26threeid%3D56%26fourid%3D72%26fiveid%3D15&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-campbellphd-10"><span class="mw-cite-backlink"><b><a href="#cite_ref-campbellphd_10-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCambpell1988" class="citation thesis cs1">Cambpell, Keith Henry Stockman (1988). <a rel="nofollow" class="external text" href="https://ethos.bl.uk/OrderDetails.do?uin=uk.bl.ethos.512104"><i>Aspects of cell cycle regulation in yeast and Xenopus</i></a> (PhD thesis). University of Sussex. <a href="/wiki/OCLC_(identifier)" class="mw-redirect" title="OCLC (identifier)">OCLC</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/oclc/757094117">757094117</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Adissertation&rft.title=Aspects+of+cell+cycle+regulation+in+yeast+and+Xenopus&rft.degree=PhD&rft.inst=University+of+Sussex&rft.date=1988&rft_id=info%3Aoclcnum%2F757094117&rft.aulast=Cambpell&rft.aufirst=Keith+Henry+Stockman&rft_id=http%3A%2F%2Fethos.bl.uk%2FOrderDetails.do%3Fuin%3Duk.bl.ethos.512104&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-11"><span class="mw-cite-backlink"><b><a href="#cite_ref-11">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.ars.usda.gov/News/docs.htm?docid=11639">"Lecture editorial for Keith Campbell, PhD, Professor of Animal Development at the University of Nottingham"</a>. ars.usda.gov. 13 November 2003<span class="reference-accessdate">. Retrieved <span class="nowrap">11 October</span> 2012</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Lecture+editorial+for+Keith+Campbell%2C+PhD%2C+Professor+of+Animal+Development+at+the+University+of+Nottingham&rft.pub=ars.usda.gov&rft.date=2003-11-13&rft_id=https%3A%2F%2Fwww.ars.usda.gov%2FNews%2Fdocs.htm%3Fdocid%3D11639&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-12">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://www.cirs-tm.org/researchers/researchers.php?id=674">Keith Campbell Profile at CIRS</a></span> </li> <li id="cite_note-13"><span class="mw-cite-backlink"><b><a href="#cite_ref-13">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="http://library.thinkquest.org/24355/data/light/details/profiles/campbell.html">Kieth Campbell Profile at thinkquest</a> <a rel="nofollow" class="external text" href="https://web.archive.org/web/20080328152836/http://library.thinkquest.org/24355/data/light/details/profiles/campbell.html">Archived</a> 28 March 2008 at the <a href="/wiki/Wayback_Machine" title="Wayback Machine">Wayback Machine</a></span> </li> <li id="cite_note-14"><span class="mw-cite-backlink"><b><a href="#cite_ref-14">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation news cs1"><a rel="nofollow" class="external text" href="https://www.telegraph.co.uk/news/uknews/1512377/I-didnt-clone-Dolly-the-sheep-says-prof.html">"I didn't clone Dolly the sheep, says prof"</a>. The Daily Telegraph. 8 March 2006<span class="reference-accessdate">. Retrieved <span class="nowrap">24 March</span> 2016</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=I+didn%27t+clone+Dolly+the+sheep%2C+says+prof&rft.date=2006-03-08&rft_id=https%3A%2F%2Fwww.telegraph.co.uk%2Fnews%2Fuknews%2F1512377%2FI-didnt-clone-Dolly-the-sheep-says-prof.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-15"><span class="mw-cite-backlink"><b><a href="#cite_ref-15">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFSchnieke1997" class="citation journal cs1">Schnieke; et al. (1997). "Human factor IX transgenic sheep produced by transfer of nuclei from transfected fetal fibroblasts". <i>Science</i>. <b>278</b> (5346): 2130–3. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1997Sci...278.2130S">1997Sci...278.2130S</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.278.5346.2130">10.1126/science.278.5346.2130</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/9405350">9405350</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:36257561">36257561</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Science&rft.atitle=Human+factor+IX+transgenic+sheep+produced+by+transfer+of+nuclei+from+transfected+fetal+fibroblasts.&rft.volume=278&rft.issue=5346&rft.pages=2130-3&rft.date=1997&rft_id=info%3Adoi%2F10.1126%2Fscience.278.5346.2130&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A36257561%23id-name%3DS2CID&rft_id=info%3Apmid%2F9405350&rft_id=info%3Abibcode%2F1997Sci...278.2130S&rft.au=Schnieke&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-16"><span class="mw-cite-backlink"><b><a href="#cite_ref-16">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCampbell1999" class="citation journal cs1"><a class="mw-selflink selflink">Campbell, K. H. S.</a> (1999). "Nuclear Equivalence, Nuclear Transfer, and the Cell Cycle". <i>Cloning</i>. <b>1</b> (1): 3–15. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1089%2F15204559950020058">10.1089/15204559950020058</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/16218826">16218826</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Cloning&rft.atitle=Nuclear+Equivalence%2C+Nuclear+Transfer%2C+and+the+Cell+Cycle&rft.volume=1&rft.issue=1&rft.pages=3-15&rft.date=1999&rft_id=info%3Adoi%2F10.1089%2F15204559950020058&rft_id=info%3Apmid%2F16218826&rft.aulast=Campbell&rft.aufirst=K.+H.+S.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-17"><span class="mw-cite-backlink"><b><a href="#cite_ref-17">^</a></b></span> <span class="reference-text">Campbell, K. & Wilmut, I. (1998) "Nuclear Transfer". <i>Animal Breeding: Technology for the 21st Century</i>. (ed: A.J. Clark; Harwood Academic Publishers). pp. 47–62</span> </li> <li id="cite_note-18"><span class="mw-cite-backlink"><b><a href="#cite_ref-18">^</a></b></span> <span class="reference-text">Campbell, K. (1998). "Cloning Dolly: Implications for Human Medicine", <i>Fertility and Reproductive Medicine</i> (eds: R.D. Kempers, J. Cohen, A.F. Haney, and J.B. Younger), pp. 3–11. (excerpted from Medica: International Congress, series 1183)</span> </li> <li id="cite_note-19"><span class="mw-cite-backlink"><b><a href="#cite_ref-19">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCampbell1997" class="citation journal cs1"><a class="mw-selflink selflink">Campbell, K.</a> (1997). "Totipotency or multipotentiality of cultured cells: Applications and progress". <i>Theriogenology</i>. <b>47</b>: 63–72. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2FS0093-691X%2896%2900340-8">10.1016/S0093-691X(96)00340-8</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Theriogenology&rft.atitle=Totipotency+or+multipotentiality+of+cultured+cells%3A+Applications+and+progress&rft.volume=47&rft.pages=63-72&rft.date=1997&rft_id=info%3Adoi%2F10.1016%2FS0093-691X%2896%2900340-8&rft.aulast=Campbell&rft.aufirst=K.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-20"><span class="mw-cite-backlink"><b><a href="#cite_ref-20">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFWilmutSchniekeMcWhirKind1997" class="citation journal cs1"><a href="/wiki/Ian_Wilmut" title="Ian Wilmut">Wilmut, I.</a>; Schnieke, A. E.; McWhir, J.; Kind, A. J.; <a class="mw-selflink selflink">Campbell, K. H. S.</a> (1997). "Viable offspring derived from fetal and adult mammalian cells". <i>Nature</i>. <b>385</b> (6619): 810–813. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1997Natur.385..810W">1997Natur.385..810W</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2F385810a0">10.1038/385810a0</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/9039911">9039911</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:4260518">4260518</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nature&rft.atitle=Viable+offspring+derived+from+fetal+and+adult+mammalian+cells&rft.volume=385&rft.issue=6619&rft.pages=810-813&rft.date=1997&rft_id=info%3Adoi%2F10.1038%2F385810a0&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A4260518%23id-name%3DS2CID&rft_id=info%3Apmid%2F9039911&rft_id=info%3Abibcode%2F1997Natur.385..810W&rft.aulast=Wilmut&rft.aufirst=I.&rft.au=Schnieke%2C+A.+E.&rft.au=McWhir%2C+J.&rft.au=Kind%2C+A.+J.&rft.au=Campbell%2C+K.+H.+S.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-21"><span class="mw-cite-backlink"><b><a href="#cite_ref-21">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFSchniekeKindRitchieMycock1997" class="citation journal cs1">Schnieke, A. E.; Kind, A. J.; Ritchie, W. A.; Mycock, K.; Scott, A. R.; Ritchie, M.; <a href="/wiki/Ian_Wilmut" title="Ian Wilmut">Wilmut, I.</a>; Colman, A.; <a class="mw-selflink selflink">Campbell, K. H.</a> (1997). "Human Factor IX Transgenic Sheep Produced by Transfer of Nuclei from Transfected Fetal Fibroblasts". <i>Science</i>. <b>278</b> (5346): 2130–2133. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1997Sci...278.2130S">1997Sci...278.2130S</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1126%2Fscience.278.5346.2130">10.1126/science.278.5346.2130</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/9405350">9405350</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:36257561">36257561</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Science&rft.atitle=Human+Factor+IX+Transgenic+Sheep+Produced+by+Transfer+of+Nuclei+from+Transfected+Fetal+Fibroblasts&rft.volume=278&rft.issue=5346&rft.pages=2130-2133&rft.date=1997&rft_id=info%3Adoi%2F10.1126%2Fscience.278.5346.2130&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A36257561%23id-name%3DS2CID&rft_id=info%3Apmid%2F9405350&rft_id=info%3Abibcode%2F1997Sci...278.2130S&rft.aulast=Schnieke&rft.aufirst=A.+E.&rft.au=Kind%2C+A.+J.&rft.au=Ritchie%2C+W.+A.&rft.au=Mycock%2C+K.&rft.au=Scott%2C+A.+R.&rft.au=Ritchie%2C+M.&rft.au=Wilmut%2C+I.&rft.au=Colman%2C+A.&rft.au=Campbell%2C+K.+H.&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-22"><span class="mw-cite-backlink"><b><a href="#cite_ref-22">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20121013004459/http://www.nottingham.ac.uk/biosciences/people/keith.campbell">"Biography: Keith H. S. Campbell, BSc D.Phil"</a>. Faculty of Science, Nottingham University. Archived from <a rel="nofollow" class="external text" href="https://www.nottingham.ac.uk/biosciences/people/keith.campbell">the original</a> on 13 October 2012<span class="reference-accessdate">. Retrieved <span class="nowrap">11 October</span> 2012</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Biography%3A+Keith+H.+S.+Campbell%2C+BSc+D.Phil&rft.pub=Faculty+of+Science%2C+Nottingham+University&rft_id=https%3A%2F%2Fwww.nottingham.ac.uk%2Fbiosciences%2Fpeople%2Fkeith.campbell&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-23"><span class="mw-cite-backlink"><b><a href="#cite_ref-23">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.http://www.iets.org/pdf/awards/2015PioneerCampbellCitation.pdf">"Recipient of the 2015 IETS Pioneer Award: Keith Henry Stockman Campbell, PhD"</a> <span class="cs1-format">(PDF)</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Recipient+of+the+2015+IETS+Pioneer+Award%3A+Keith+Henry+Stockman+Campbell%2C+PhD&rft_id=https%3A%2F%2Fwww.http%3A%2F%2Fwww.iets.org%2Fpdf%2Fawards%2F2015PioneerCampbellCitation.pdf&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span><sup class="noprint Inline-Template"><span style="white-space: nowrap;">[<i><a href="/wiki/Wikipedia:Link_rot" title="Wikipedia:Link rot"><span title=" Dead link tagged May 2017">permanent dead link</span></a></i><span style="visibility:hidden; color:transparent; padding-left:2px">‍</span>]</span></sup></span> </li> <li id="cite_note-24"><span class="mw-cite-backlink"><b><a href="#cite_ref-24">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBritten2013" class="citation news cs1">Britten, Nick (18 May 2013). <a rel="nofollow" class="external text" href="https://www.telegraph.co.uk/news/uknews/10065584/Scientist-behind-Dolly-the-sheep-killed-himself-by-mistake-in-drunken-fury.html">"Keith Campbell hanged himself by mistake"</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.atitle=Keith+Campbell+hanged+himself+by+mistake&rft.date=2013-05-18&rft.aulast=Britten&rft.aufirst=Nick&rft_id=https%3A%2F%2Fwww.telegraph.co.uk%2Fnews%2Fuknews%2F10065584%2FScientist-behind-Dolly-the-sheep-killed-himself-by-mistake-in-drunken-fury.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-25"><span class="mw-cite-backlink"><b><a href="#cite_ref-25">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://www.independent.co.uk/news/obituaries/professor-keith-campbell-biologist-who-played-a-leading-role-in-cloning-dolly-the-sheep-8209724.htmll">"Obituary: Professor Keith Campbell: Biologist who plarole in cloning Dolly the sheep"</a>. <i><a href="/wiki/Independent.co.uk" class="mw-redirect" title="Independent.co.uk">Independent.co.uk</a></i>. 13 October 2012.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=Independent.co.uk&rft.atitle=Obituary%3A+Professor+Keith+Campbell%3A+Biologist+who+plarole+in+cloning+Dolly+the+sheep&rft.date=2012-10-13&rft_id=https%3A%2F%2Fwww.independent.co.uk%2Fnews%2Fobituaries%2Fprofessor-keith-campbell-biologist-who-played-a-leading-role-in-cloning-dolly-the-sheep-8209724.htmll&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> <li id="cite_note-26"><span class="mw-cite-backlink"><b><a href="#cite_ref-26">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation web cs1"><a rel="nofollow" class="external text" href="http://www.nottingham.ac.uk/news/keith-campbell/keith-campbell-book-of-condolence.aspx">"Keith Campbell – Book of Condolence"</a>. University of Nottingham. 11 October 2012<span class="reference-accessdate">. Retrieved <span class="nowrap">16 May</span> 2013</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=Keith+Campbell+%E2%80%93+Book+of+Condolence&rft.pub=University+of+Nottingham&rft.date=2012-10-11&rft_id=http%3A%2F%2Fwww.nottingham.ac.uk%2Fnews%2Fkeith-campbell%2Fkeith-campbell-book-of-condolence.aspx&rfr_id=info%3Asid%2Fen.wikipedia.org%3AKeith+Campbell+%28biologist%29" class="Z3988"></span></span> </li> </ol></div> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output .hlist.inline dl,.mw-parser-output .hlist.inline ol,.mw-parser-output .hlist.inline ul,.mw-parser-output .hlist dl dl,.mw-parser-output .hlist dl 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title="Saul Perlmutter">Saul Perlmutter</a>, <a href="/wiki/Adam_Riess" title="Adam Riess">Adam Riess</a> and <a href="/wiki/Brian_Schmidt" title="Brian Schmidt">Brian Schmidt</a> (2006)</li> <li><a href="/wiki/Peter_Goldreich" title="Peter Goldreich">Peter Goldreich</a> (2007)</li> <li><a href="/wiki/Reinhard_Genzel" title="Reinhard Genzel">Reinhard Genzel</a> (2008)</li> <li><a href="/wiki/Frank_Shu" title="Frank Shu">Frank Shu</a> (2009)</li> <li><a href="/wiki/Charles_L._Bennett" title="Charles L. Bennett">Charles Bennett</a>, <a href="/wiki/Lyman_Page" title="Lyman Page">Lyman Page</a> and <a href="/wiki/David_Spergel" title="David Spergel">David Spergel</a> (2010)</li> <li><a href="/wiki/Enrico_Costa_(physicist)" title="Enrico Costa (physicist)">Enrico Costa</a> and <a href="/wiki/Gerald_J._Fishman" title="Gerald J. Fishman">Gerald Fishman</a> (2011)</li> <li><a href="/wiki/David_C._Jewitt" title="David C. Jewitt">David C. Jewitt</a> and <a href="/wiki/Jane_Luu" title="Jane Luu">Jane Luu</a> (2012)</li> <li><a href="/wiki/Steven_Balbus" title="Steven Balbus">Steven Balbus</a> and <a href="/wiki/John_F._Hawley" title="John F. Hawley">John F. Hawley</a> (2013)</li> <li><a href="/wiki/Daniel_Eisenstein" title="Daniel Eisenstein">Daniel Eisenstein</a>, <a href="/wiki/Shaun_Cole" title="Shaun Cole">Shaun Cole</a> and <a href="/wiki/John_A._Peacock" title="John A. Peacock">John A. Peacock</a> (2014)</li> <li><a href="/wiki/William_J._Borucki" title="William J. Borucki">William J. Borucki</a> (2015)</li> <li><a href="/wiki/Ronald_Drever" title="Ronald Drever">Ronald Drever</a>, <a href="/wiki/Kip_Thorne" title="Kip Thorne">Kip Thorne</a> and <a href="/wiki/Rainer_Weiss" title="Rainer Weiss">Rainer Weiss</a> (2016)</li> <li><a href="/wiki/Simon_White" title="Simon White">Simon White</a> (2017)</li> <li><a href="/wiki/Jean-Loup_Puget" title="Jean-Loup Puget">Jean-Loup Puget</a> (2018)</li> <li><a href="/wiki/Edward_C._Stone" title="Edward C. Stone">Edward C. Stone</a> (2019)</li> <li><a href="/wiki/Roger_Blandford" title="Roger Blandford">Roger Blandford</a> (2020)</li> <li><a href="/wiki/Victoria_Kaspi" title="Victoria Kaspi">Victoria Kaspi</a> and <a href="/wiki/Chryssa_Kouveliotou" title="Chryssa Kouveliotou">Chryssa Kouveliotou</a> (2021)</li> <li><a href="/wiki/Lennart_Lindegren" title="Lennart Lindegren">Lennart Lindegren</a> and <a href="/wiki/Michael_Perryman" title="Michael Perryman">Michael Perryman</a> (2022)</li> <li><a href="/wiki/Matthew_Bailes" title="Matthew Bailes">Matthew Bailes</a>, <a href="/wiki/Duncan_Lorimer" title="Duncan Lorimer">Duncan Lorimer</a> and <a href="/wiki/Maura_McLaughlin" title="Maura McLaughlin">Maura McLaughlin</a> (2023)</li> <li><a href="/wiki/Shrinivas_R._Kulkarni" class="mw-redirect" title="Shrinivas R. Kulkarni">Shrinivas R. Kulkarni</a> (2024)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">Life science<br />and medicine</th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Stanley_Norman_Cohen" title="Stanley Norman Cohen">Stanley Norman Cohen</a>, <a href="/wiki/Herbert_Boyer" title="Herbert Boyer">Herbert Boyer</a>, <a href="/wiki/Yuet_Wai_Kan" title="Yuet Wai Kan">Yuet-Wai Kan</a> and <a href="/wiki/Richard_Doll" title="Richard Doll">Richard Doll</a> (2004)</li> <li><a href="/wiki/Michael_Berridge" title="Michael Berridge">Michael Berridge</a> (2005)</li> <li><a href="/wiki/Xiaodong_Wang_(biochemist)" title="Xiaodong Wang (biochemist)">Xiaodong Wang</a> (2006)</li> <li><a href="/wiki/Robert_Lefkowitz" title="Robert Lefkowitz">Robert Lefkowitz</a> (2007)</li> <li><a href="/wiki/Ian_Wilmut" title="Ian Wilmut">Ian Wilmut</a>, <a class="mw-selflink selflink">Keith H. S. Campbell</a> and <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Shinya Yamanaka</a> (2008)</li> <li><a href="/wiki/Douglas_L._Coleman" title="Douglas L. Coleman">Douglas Coleman</a> and <a href="/wiki/Jeffrey_M._Friedman" title="Jeffrey M. Friedman">Jeffrey Friedman</a> (2009)</li> <li><a href="/wiki/David_Julius" title="David Julius">David Julius</a> (2010)</li> <li><a href="/wiki/Jules_A._Hoffmann" title="Jules A. Hoffmann">Jules Hoffmann</a>, <a href="/wiki/Ruslan_Medzhitov" title="Ruslan Medzhitov">Ruslan Medzhitov</a> and <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Bruce Beutler</a> (2011)</li> <li><a href="/wiki/Franz-Ulrich_Hartl" title="Franz-Ulrich Hartl">Franz-Ulrich Hartl</a> and <a href="/wiki/Arthur_L._Horwich" title="Arthur L. Horwich">Arthur L. Horwich</a> (2012)</li> <li><a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Jeffrey C. Hall</a>, <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Michael Rosbash</a> and <a href="/wiki/Michael_W._Young" title="Michael W. Young">Michael W. Young</a> (2013)</li> <li><a href="/wiki/Kazutoshi_Mori" title="Kazutoshi Mori">Kazutoshi Mori</a> and <a href="/wiki/Peter_Walter" title="Peter Walter">Peter Walter</a> (2014)</li> <li><a href="/wiki/Bonnie_Bassler" title="Bonnie Bassler">Bonnie Bassler</a> and <a href="/wiki/Everett_Peter_Greenberg" title="Everett Peter Greenberg">Everett Peter Greenberg</a> (2015)</li> <li><a href="/wiki/Adrian_Bird" title="Adrian Bird">Adrian Bird</a> and <a href="/wiki/Huda_Zoghbi" title="Huda Zoghbi">Huda Zoghbi</a> (2016)</li> <li><a href="/wiki/Ian_R._Gibbons" title="Ian R. Gibbons">Ian R. Gibbons</a> and <a href="/wiki/Ronald_Vale" title="Ronald Vale">Ronald Vale</a> (2017)</li> <li><a href="/wiki/Mary-Claire_King" title="Mary-Claire King">Mary-Claire King</a> (2018)</li> <li><a href="/wiki/Maria_Jasin" title="Maria Jasin">Maria Jasin</a> (2019)</li> <li><a href="/wiki/Gero_Miesenb%C3%B6ck" title="Gero Miesenböck">Gero Miesenböck</a>, <a href="/wiki/Peter_Hegemann" title="Peter Hegemann">Peter Hegemann</a> and <a href="/wiki/Georg_Nagel" title="Georg Nagel">Georg Nagel</a> (2020)</li> <li><a href="/wiki/Scott_D._Emr" title="Scott D. Emr">Scott D. Emr</a> (2021)</li> <li><a href="/wiki/Paul_Negulescu" title="Paul Negulescu">Paul A. Negulescu</a> and <a href="/wiki/Michael_J._Welsh_(biologist)" title="Michael J. Welsh (biologist)">Michael J. Welsh</a> (2022)</li> <li><a href="/wiki/Patrick_Cramer" title="Patrick Cramer">Patrick Cramer</a> and <a href="/wiki/Eva_Nogales" title="Eva Nogales">Eva Nogales</a> (2023)</li> <li><a href="/wiki/Stuart_H._Orkin" class="mw-redirect" title="Stuart H. Orkin">Stuart H. Orkin</a> and <a href="/wiki/Swee_Lay_Thein" title="Swee Lay Thein">Swee Lay Thein</a> (2024)</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;background:#d3d3d3;">Mathematical<br />science</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Shiing-Shen_Chern" title="Shiing-Shen Chern">Shiing-Shen Chern</a> (2004)</li> <li><a href="/wiki/Andrew_Wiles" title="Andrew Wiles">Andrew Wiles</a> (2005)</li> <li><a href="/wiki/David_Mumford" title="David Mumford">David Mumford</a> and <a href="/wiki/Wu_Wenjun" title="Wu Wenjun">Wentsun Wu</a> (2006)</li> <li><a href="/wiki/Robert_Langlands" title="Robert Langlands">Robert Langlands</a> and <a href="/wiki/Richard_Taylor_(mathematician)" title="Richard Taylor (mathematician)">Richard Taylor</a> (2007)</li> <li><a href="/wiki/Vladimir_Arnold" title="Vladimir Arnold">Vladimir Arnold</a> and <a href="/wiki/Ludvig_Faddeev" title="Ludvig Faddeev">Ludwig Faddeev</a> (2008)</li> <li><a href="/wiki/Simon_Donaldson" title="Simon Donaldson">Simon Donaldson</a> and <a href="/wiki/Clifford_Taubes" title="Clifford Taubes">Clifford Taubes</a> (2009)</li> <li><a href="/wiki/Jean_Bourgain" title="Jean Bourgain">Jean Bourgain</a> (2010)</li> <li><a href="/wiki/Demetrios_Christodoulou" title="Demetrios Christodoulou">Demetrios Christodoulou</a> and <a href="/wiki/Richard_S._Hamilton" title="Richard S. Hamilton">Richard S. Hamilton</a> (2011)</li> <li><a href="/wiki/Maxim_Kontsevich" title="Maxim Kontsevich">Maxim Kontsevich</a> (2012)</li> <li><a href="/wiki/David_Donoho" title="David Donoho">David Donoho</a> (2013)</li> <li><a href="/wiki/George_Lusztig" title="George Lusztig">George Lusztig</a> (2014)</li> <li><a href="/wiki/Gerd_Faltings" title="Gerd Faltings">Gerd Faltings</a> and <a href="/wiki/Henryk_Iwaniec" title="Henryk Iwaniec">Henryk Iwaniec</a> (2015)</li> <li><a href="/wiki/Nigel_Hitchin" title="Nigel Hitchin">Nigel Hitchin</a> (2016)</li> <li><a href="/wiki/J%C3%A1nos_Koll%C3%A1r" title="János Kollár">János Kollár</a> and <a href="/wiki/Claire_Voisin" title="Claire Voisin">Claire Voisin</a> (2017)</li> <li><a href="/wiki/Luis_Caffarelli" title="Luis Caffarelli">Luis Caffarelli</a> (2018)</li> <li><a href="/wiki/Michel_Talagrand" title="Michel Talagrand">Michel Talagrand</a> (2019)</li> <li><a href="/wiki/Alexander_Beilinson" title="Alexander Beilinson">Alexander Beilinson</a> and <a href="/wiki/David_Kazhdan" title="David Kazhdan">David Kazhdan</a> (2020)</li> <li><a href="/wiki/Jean-Michel_Bismut" title="Jean-Michel Bismut">Jean-Michel Bismut</a> and <a href="/wiki/Jeff_Cheeger" title="Jeff Cheeger">Jeff Cheeger</a> (2021)</li> <li><a href="/wiki/Noga_Alon" title="Noga Alon">Noga Alon</a> and <a href="/wiki/Ehud_Hrushovski" title="Ehud Hrushovski">Ehud Hrushovski</a> (2022)</li> <li><a href="/wiki/Vladimir_Drinfeld" title="Vladimir Drinfeld">Vladimir Drinfeld</a> and <a href="/wiki/Shing-Tung_Yau" title="Shing-Tung Yau">Shing-Tung Yau</a> (2023)</li> <li><a href="/wiki/Peter_Sarnak" title="Peter Sarnak">Peter Sarnak</a> (2024)</li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox authority-control" 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