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class="vector-toc-numb">2.2</span> <span>Galton's taxonomic revisions</span> </div> </a> <ul id="toc-Galton's_taxonomic_revisions-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Classification" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Classification"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Classification</span> </div> </a> <ul id="toc-Classification-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Paleobiology_and_paleoecology" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Paleobiology_and_paleoecology"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Paleobiology and paleoecology</span> </div> </a> <ul id="toc-Paleobiology_and_paleoecology-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 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title="Nanosaurus – German" lang="de" hreflang="de" data-title="Nanosaurus" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D9%86%D8%A7%D9%86%D9%88%D8%B3%D9%88%D8%B1" title="نانوسور – Persian" lang="fa" hreflang="fa" data-title="نانوسور" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Nanosaurus" title="Nanosaurus – French" lang="fr" hreflang="fr" data-title="Nanosaurus" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%A0%D7%A0%D7%95%D7%96%D7%90%D7%95%D7%A8%D7%95%D7%A1" title="ננוזאורוס – Hebrew" lang="he" hreflang="he" data-title="ננוזאורוס" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-mk mw-list-item"><a href="https://mk.wikipedia.org/wiki/%D0%9D%D0%B0%D0%BD%D0%BE%D1%81%D0%B0%D1%83%D1%80%D1%83%D1%81" title="Наносаурус – Macedonian" lang="mk" hreflang="mk" data-title="Наносаурус" data-language-autonym="Македонски" data-language-local-name="Macedonian" class="interlanguage-link-target"><span>Македонски</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Nanosaurus" title="Nanosaurus – Dutch" lang="nl" hreflang="nl" data-title="Nanosaurus" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Nanozaur" title="Nanozaur – Polish" lang="pl" hreflang="pl" data-title="Nanozaur" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Nanosaurus" title="Nanosaurus – Portuguese" lang="pt" hreflang="pt" data-title="Nanosaurus" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/Nanosaurus" title="Nanosaurus – Russian" lang="ru" hreflang="ru" data-title="Nanosaurus" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Nanosaurus" title="Nanosaurus – Simple English" lang="en-simple" hreflang="en-simple" data-title="Nanosaurus" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Nanosaurus" title="Nanosaurus – Finnish" lang="fi" hreflang="fi" data-title="Nanosaurus" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Nanosaurus" title="Nanosaurus – Vietnamese" lang="vi" hreflang="vi" data-title="Nanosaurus" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-zh 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</nav> <nav class="vector-appearance-landmark" aria-label="Appearance"> <div id="vector-appearance-pinned-container" class="vector-pinned-container"> <div id="vector-appearance" class="vector-appearance vector-pinnable-element"> <div class="vector-pinnable-header vector-appearance-pinnable-header vector-pinnable-header-pinned" data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">Appearance</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">move to sidebar</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">hide</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"><span class="mw-redirectedfrom">(Redirected from <a href="/w/index.php?title=Othnielosaurus&redirect=no" class="mw-redirect" title="Othnielosaurus">Othnielosaurus</a>)</span></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Extinct genus of dinosaurs</div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">Not to be confused with <a href="/wiki/Nanosaur" title="Nanosaur">Nanosaur</a>.</div> <p class="mw-empty-elt"> </p> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><i>Nanosaurus</i><br /><div style="font-size: 85%;">Temporal range: <a href="/wiki/Late_Jurassic" title="Late Jurassic">Late Jurassic</a>, <span class="noprint"><span style="display:inline-block;"></span><span style="display:inline-block;">155–148 <a href="/wiki/Megaannum" class="mw-redirect" title="Megaannum">Ma</a></span> <span style="display:inline-block;"></span><div id="Timeline-row" style="margin: 4px auto 0; clear:both; width:220px; padding:0px; height:18px; overflow:visible; white-space:nowrap; border:1px #666; border-style:solid none; position:relative; z-index:0; font-size:97%;"> <div style="position:absolute; height:100%; left:0px; width:207.23076923077px; padding-left:5px; text-align:left; background-color:rgb(254,217,106); background-image: linear-gradient(to right, rgba(255,255,255,1), rgba(254,217,106,1) 15%, rgba(254,217,106,1));"><a href="/wiki/Precambrian" title="Precambrian">PreꞒ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,160,86); left:37.636923076923px; width:18.073846153846px;"><a href="/wiki/Cambrian" title="Cambrian">Ꞓ</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(0,146,112); left:55.710769230769px; width:14.08px;"><a href="/wiki/Ordovician" title="Ordovician">O</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(179,225,182); left:69.790769230769px; width:8.3261538461539px;"><a href="/wiki/Silurian" title="Silurian">S</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(203,140,55); left:78.116923076923px; width:20.409230769231px;"><a href="/wiki/Devonian" title="Devonian">D</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(103,165,153); left:98.526153846154px; width:20.307692307692px;"><a href="/wiki/Carboniferous" title="Carboniferous">C</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(240,64,40); left:118.83384615385px; width:15.907015384615px;"><a href="/wiki/Permian" title="Permian">P</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(129,43,146); left:134.74086153846px; width:17.092984615385px;"><a href="/wiki/Triassic" title="Triassic">T</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(52,178,201); left:151.83384615385px; width:19.089230769231px;"><a href="/wiki/Jurassic" title="Jurassic">J</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(127,198,78); left:170.92307692308px; width:26.738461538462px;"><a href="/wiki/Cretaceous" title="Cretaceous">K</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(253,154,82); left:197.66153846154px; width:14.543692307692px;"><a href="/wiki/Paleogene" title="Paleogene">Pg</a></div> <div style="position:absolute; height:100%; text-align:center; background-color:rgb(255,230,25); left:212.20523076923px; width:6.9215384615385px;"><a href="/wiki/Neogene" title="Neogene">N</a></div> <div id="end-border" style="position:absolute; height:100%; background-color:#666; width:1px; left:219px"></div><div style="margin:0 auto; line-height:0; clear:both; width:220px; padding:0px; height:8px; overflow:visible; background-color:transparent; position:relative; top:-4px; z-index:100;"><div style="position:absolute; height:8px; left:167.53846153846px; width:2.3692307692308px; background-color:#360; opacity:0.42;"></div> <div style="position:absolute; height:8px; left:167.53846153846px; width:2.3692307692308px; background-color:#360; opacity:1;"></div> <div style="position:absolute; height:6px; top:1px; left:168.53846153846px; width:0.36923076923077px; background-color:#6c3;"></div> </div> </div></span></div> </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Othnielosaurus_fruita_1.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/6/64/Othnielosaurus_fruita_1.jpg/220px-Othnielosaurus_fruita_1.jpg" decoding="async" width="220" height="147" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/64/Othnielosaurus_fruita_1.jpg/330px-Othnielosaurus_fruita_1.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/64/Othnielosaurus_fruita_1.jpg/440px-Othnielosaurus_fruita_1.jpg 2x" data-file-width="5472" data-file-height="3648" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%">Reconstructed skeleton cast, <a href="/wiki/Dinosaur_Journey_Museum" title="Dinosaur Journey Museum">Dinosaur Journey Museum</a> </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Taxonomy_(biology)" title="Taxonomy (biology)">Scientific classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Nanosaurus" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>Domain: </td> <td><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Animal" title="Animal">Animalia</a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Chordate" title="Chordate">Chordata</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><a href="/wiki/Dinosaur" title="Dinosaur">Dinosauria</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><span style="font-style:normal;font-weight:normal;">†</span><a href="/wiki/Ornithischia" title="Ornithischia">Ornithischia</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><span style="font-style:normal;font-weight:normal;">†</span><a href="/wiki/Genasauria" class="mw-redirect" title="Genasauria">Genasauria</a> </td></tr> <tr> <td><i>Clade</i>: </td> <td><span style="font-style:normal;font-weight:normal;">†</span><a href="/wiki/Neornithischia" title="Neornithischia">Neornithischia</a> </td></tr> <tr> <td>Family: </td> <td><span style="font-style:normal;font-weight:normal;">†</span><a class="mw-selflink selflink">Nanosauridae</a><br /><small><a href="/wiki/Othniel_Charles_Marsh" title="Othniel Charles Marsh">Marsh</a>, 1877</small> </td></tr> <tr> <td>Genus: </td> <td><span style="font-style:normal;font-weight:normal;">†</span><a class="mw-selflink selflink"><i>Nanosaurus</i></a><br /><small>Marsh, <a href="/wiki/1877_in_paleontology" title="1877 in paleontology">1877</a></small> </td></tr> <tr> <td>Species: </td> <td><div style="display:inline" class="species">†<i><b>N. agilis</b></i></div> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Binomial_nomenclature" title="Binomial nomenclature">Binomial name</a> </th></tr> <tr> <td colspan="2" style="text-align: center"><b><span class="binomial"><span style="font-weight:normal;">†</span><i>Nanosaurus agilis</i></span></b><br /><div style="font-size: 85%;">Marsh, 1877</div> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(235,235,210)"><a href="/wiki/Synonym_(taxonomy)" title="Synonym (taxonomy)">Synonyms</a> </th></tr> <tr> <td colspan="2" style="text-align: left"> <div class="collapsible-list mw-collapsible mw-collapsed" style="text-align: left;"> <div style="line-height: 1.6em; font-weight: bold;"><div><small>Genus synonymy</small></div></div> <ul class="mw-collapsible-content" style="margin-top: 0; margin-bottom: 0; line-height: inherit;"><li style="line-height: inherit; margin: 0"><i>Drinker</i> <small>Bakker et al., 1990</small> </li><li style="line-height: inherit; margin: 0"><i>Othnielia</i> <small>Galton, 1977</small> </li><li style="line-height: inherit; margin: 0"><i>Othnielosaurus</i> <small>Galton, 2007</small> </li></ul> </div> <div class="collapsible-list mw-collapsible mw-collapsed" style="text-align: left;"> <div style="line-height: 1.6em; font-weight: bold;"><div><small>Species synonymy</small></div></div> <ul class="mw-collapsible-content" style="margin-top: 0; margin-bottom: 0; line-height: inherit;"><li style="line-height: inherit; margin: 0"><i>Drinker nisti</i> <small>Bakker et al., 1990</small> </li><li style="line-height: inherit; margin: 0"><i>Laosaurus consors</i> <small>Marsh, 1894</small> </li><li style="line-height: inherit; margin: 0"><i>Nanosaurus rex</i> <small>Marsh, 1877</small> </li><li style="line-height: inherit; margin: 0"><i>Othnielia rex</i> <small>(Marsh, 1877)</small> </li><li style="line-height: inherit; margin: 0"><i>Othnielosaurus consors</i> <small>(Marsh, 1894)</small> </li></ul> </div> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p><i><b>Nanosaurus</b></i> ("small or dwarf lizard") is an extinct <a href="/wiki/Genus" title="Genus">genus</a> of <a href="/wiki/Neornithischian" class="mw-redirect" title="Neornithischian">neornithischian</a> <a href="/wiki/Dinosaur" title="Dinosaur">dinosaur</a> that lived about 155 to 148 million years ago, during the <a href="/wiki/Late_Jurassic" title="Late Jurassic">Late Jurassic</a> in North America. Its <a href="/wiki/Fossil" title="Fossil">fossils</a> are known from the <a href="/wiki/Morrison_Formation" title="Morrison Formation">Morrison Formation</a> of the south-western <a href="/wiki/United_States" title="United States">United States</a>. The <a href="/wiki/Type_species" title="Type species">type</a> and only <a href="/wiki/Species" title="Species">species</a>, <i><b>Nanosaurus agilis</b></i>, was <a href="/wiki/Species_description" title="Species description">described</a> and named by <a href="/wiki/Othniel_Charles_Marsh" title="Othniel Charles Marsh">Othniel Charles Marsh</a> in 1877. The <a href="/wiki/Taxon" title="Taxon">taxon</a> has a complicated taxonomic history, largely the work of Marsh and <a href="/wiki/Peter_M._Galton" class="mw-redirect" title="Peter M. Galton">Peter M. Galton</a>, involving the genera <i><a href="/wiki/Laosaurus" title="Laosaurus">Laosaurus</a></i>, <i><a href="/wiki/Hallopus" title="Hallopus">Hallopus</a></i>, <i>Drinker</i>, <i>Othnielia</i>, and <i>Othnielosaurus</i>, the latter three now being considered to be synonyms of <i>Nanosaurus</i>. It had historically been classified as a <a href="/wiki/Hypsilophodont" class="mw-redirect" title="Hypsilophodont">hypsilophodont</a> or <a href="/wiki/Fabrosauridae" class="mw-redirect" title="Fabrosauridae">fabrosaur</a>, types of generalized small bipedal herbivore, but more recent research has abandoned these groupings as <a href="/wiki/Paraphyletic" class="mw-redirect" title="Paraphyletic">paraphyletic</a> and <i>Nanosaurus</i> is today considered a <a href="/wiki/Basal_(phylogenetics)" title="Basal (phylogenetics)">basal</a> member of Neornithischia. </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Description">Description</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nanosaurus&action=edit&section=1" title="Edit section: Description"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Othnielosaurus_Size_Comparison_by_PaleoGeek.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/77/Othnielosaurus_Size_Comparison_by_PaleoGeek.svg/220px-Othnielosaurus_Size_Comparison_by_PaleoGeek.svg.png" decoding="async" width="220" height="176" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/77/Othnielosaurus_Size_Comparison_by_PaleoGeek.svg/330px-Othnielosaurus_Size_Comparison_by_PaleoGeek.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/77/Othnielosaurus_Size_Comparison_by_PaleoGeek.svg/440px-Othnielosaurus_Size_Comparison_by_PaleoGeek.svg.png 2x" data-file-width="886" data-file-height="709" /></a><figcaption>Size compared to a human</figcaption></figure> <p><i>Nanosaurus</i> is known from material from all parts of the body, including two good skeletons, although the skull is still poorly known.<sup id="cite_ref-NSWC04_1-0" class="reference"><a href="#cite_note-NSWC04-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> It was a small animal, with specimens previously assigned to <i>Drinker</i> and <i>Othnielosaurus</i> measuring 2–2.2 metres (6.6–7.2 ft) long and weighing 20–30 kilograms (44–66 lb).<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> </p><p>It was a bipedal dinosaur with short forelimbs and long hindlimbs with large <a href="/wiki/Process_(anatomy)" title="Process (anatomy)">processes</a> for <a href="/wiki/Muscle" title="Muscle">muscle</a> attachments.<sup id="cite_ref-SH04_3-0" class="reference"><a href="#cite_note-SH04-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> The hands were short and broad with short fingers. The head was small. It had small leaf-shaped cheek teeth (triangular and with small ridges and denticles lining the front and back edges), and <a href="/wiki/Premaxilla" title="Premaxilla">premaxillary</a> teeth with less ornamentation.<sup id="cite_ref-PMG07_4-0" class="reference"><a href="#cite_note-PMG07-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p><p>Like several other neornithischian dinosaurs, such as <i><a href="/wiki/Hypsilophodon" title="Hypsilophodon">Hypsilophodon</a></i>, <i><a href="/wiki/Thescelosaurus" title="Thescelosaurus">Thescelosaurus</a></i>, and <i><a href="/wiki/Talenkauen" title="Talenkauen">Talenkauen</a></i>, <i>Nanosaurus</i> had thin plates lying along the ribs. Called intercostal plates, these structures were <a href="/wiki/Cartilage" title="Cartilage">cartilaginous</a> in origin.<sup id="cite_ref-BG08_5-0" class="reference"><a href="#cite_note-BG08-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="History_and_taxonomy">History and taxonomy</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nanosaurus&action=edit&section=2" title="Edit section: History and taxonomy"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Marsh's_original_groundwork"><span id="Marsh.27s_original_groundwork"></span>Marsh's original groundwork</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nanosaurus&action=edit&section=3" title="Edit section: Marsh's original groundwork"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Nanosaurus.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c2/Nanosaurus.jpg/220px-Nanosaurus.jpg" decoding="async" width="220" height="60" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c2/Nanosaurus.jpg/330px-Nanosaurus.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c2/Nanosaurus.jpg/440px-Nanosaurus.jpg 2x" data-file-width="605" data-file-height="164" /></a><figcaption>Holotype dentary and ilium</figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Nano_type.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/9/90/Nano_type.jpg/220px-Nano_type.jpg" decoding="async" width="220" height="132" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/90/Nano_type.jpg/330px-Nano_type.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/90/Nano_type.jpg/440px-Nano_type.jpg 2x" data-file-width="3235" data-file-height="1939" /></a><figcaption>Type specimen of <i>Nanosaurus agilis</i> Marsh 1877 (YPM VP 1913) as illustrated in 1908 (left) and cast of the bone impressions after the fragmentary bone was removed (right) </figcaption></figure> <p><i>Nanosaurus</i> has had a long and complicated taxonomic history. In 1877, Marsh named two species of <i>Nanosaurus</i> in separate publications, based on partial remains from the Morrison Formation of <a href="/wiki/Garden_Park,_Colorado" title="Garden Park, Colorado">Garden Park</a>, <a href="/wiki/Colorado" title="Colorado">Colorado</a>. One paper described <i>N. agilis</i>, based on <a href="/wiki/Peabody_Museum_of_Natural_History" title="Peabody Museum of Natural History">YPM</a> 1913, with remains including impressions of a <a href="/wiki/Dentary" class="mw-redirect" title="Dentary">dentary</a>, and <a href="/wiki/Postcrania" class="mw-redirect" title="Postcrania">postcranial</a> bits including an <a href="/wiki/Ilium_(bone)" title="Ilium (bone)">ilium</a>, <a href="/wiki/Femur" title="Femur">thigh bones</a>, <a href="/wiki/Tibia" title="Tibia">shin bones</a>, and a <a href="/wiki/Fibula" title="Fibula">fibula</a>.<sup id="cite_ref-OCM77a_6-0" class="reference"><a href="#cite_note-OCM77a-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> The other paper named <i>N. rex</i>, a second species which Marsh based on YPM 1915 (also called 1925 in Galton, 2007), a complete thigh bone.<sup id="cite_ref-PMG07_4-1" class="reference"><a href="#cite_note-PMG07-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-OCM77b_7-0" class="reference"><a href="#cite_note-OCM77b-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> He regarded both species as small ("fox-sized") animals.<sup id="cite_ref-OCM77b_7-1" class="reference"><a href="#cite_note-OCM77b-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> A third species, <i><a href="/wiki/Hallopus" title="Hallopus">N. victor</a></i>, was named, which he soon recognized to be something completely different, and is now known as the small, bipedal <a href="/wiki/Crocodylomorph" class="mw-redirect" title="Crocodylomorph">crocodylomorph</a> <i><a href="/wiki/Hallopus" title="Hallopus">Hallopus</a></i>.<sup id="cite_ref-OCM77a_6-1" class="reference"><a href="#cite_note-OCM77a-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-OCM81_8-0" class="reference"><a href="#cite_note-OCM81-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> </p> <figure class="mw-default-size mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Marsh_laosaurus.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/6/66/Marsh_laosaurus.jpg/220px-Marsh_laosaurus.jpg" decoding="async" width="220" height="156" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/6/66/Marsh_laosaurus.jpg/330px-Marsh_laosaurus.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/6/66/Marsh_laosaurus.jpg/440px-Marsh_laosaurus.jpg 2x" data-file-width="2063" data-file-height="1464" /></a><figcaption><a href="/wiki/Othniel_Charles_Marsh" title="Othniel Charles Marsh">Othniel Charles Marsh</a>'s 1896 skeletal restoration of "<a href="/wiki/Laosaurus" title="Laosaurus">Laosaurus</a>" <i>consors</i> (now <i>Nanosaurus</i>).</figcaption></figure> <p>The next year, he named the new genus <i><a href="/wiki/Laosaurus" title="Laosaurus">Laosaurus</a></i> on material collected by <a href="/wiki/Samuel_Wendell_Williston" title="Samuel Wendell Williston">Samuel Wendell Williston</a> from <a href="/wiki/Como_Bluff" title="Como Bluff">Como Bluff</a>, <a href="/wiki/Wyoming" title="Wyoming">Wyoming</a>. Two species were named: the <a href="/wiki/Type_species" title="Type species">type species</a> <i>L. celer</i>, based on parts of eleven <a href="/wiki/Vertebra" title="Vertebra">vertebrae</a> (YPM 1875);<sup id="cite_ref-OCM78_9-0" class="reference"><a href="#cite_note-OCM78-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> and the "smaller" <i>L. gracilis</i>, originally based on a back vertebra's <a href="/wiki/Vertebra" title="Vertebra">centrum</a>, a <a href="/wiki/Tail" title="Tail">caudal</a> centrum, and part of an <a href="/wiki/Ulna" title="Ulna">ulna</a> (review by <a href="/wiki/Peter_Galton" title="Peter Galton">Peter Galton</a> in 1983 finds the specimen to now consist of thirteen back and eight caudal centra, and portions of both hindlimbs).<sup id="cite_ref-OCM78_9-1" class="reference"><a href="#cite_note-OCM78-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-PMG83_10-0" class="reference"><a href="#cite_note-PMG83-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> A third species, <i>L. consors</i>, was established by Marsh in 1894 for YPM 1882, which consists of most of one articulated skeleton and part of at least one other individual.<sup id="cite_ref-OCM94_11-0" class="reference"><a href="#cite_note-OCM94-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> The skull was only partially preserved, and the fact that the vertebrae were represented only by centra suggests a partially grown individual. Galton (1983) notes that much of the current mounted skeleton was restored in plaster, or had paint applied.<sup id="cite_ref-PMG83_10-1" class="reference"><a href="#cite_note-PMG83-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Galton's_taxonomic_revisions"><span id="Galton.27s_taxonomic_revisions"></span>Galton's taxonomic revisions</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nanosaurus&action=edit&section=4" title="Edit section: Galton's taxonomic revisions"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size mw-halign-left" typeof="mw:File/Thumb"><a href="/wiki/File:Othnielia.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/24/Othnielia.png/220px-Othnielia.png" decoding="async" width="220" height="92" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/24/Othnielia.png/330px-Othnielia.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/24/Othnielia.png/440px-Othnielia.png 2x" data-file-width="1800" data-file-height="749" /></a><figcaption>Cast mounted as if a herd running, Denver Museum of Nature and Science.</figcaption></figure> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Nanosaurus_BYU_163.tif" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/2a/Nanosaurus_BYU_163.tif/lossy-page1-220px-Nanosaurus_BYU_163.tif.jpg" decoding="async" width="220" height="304" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/2a/Nanosaurus_BYU_163.tif/lossy-page1-330px-Nanosaurus_BYU_163.tif.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/2a/Nanosaurus_BYU_163.tif/lossy-page1-440px-Nanosaurus_BYU_163.tif.jpg 2x" data-file-width="2386" data-file-height="3300" /></a><figcaption>Skull-less skeleton of a juvenile <i>Nanosaurus</i> skeleton (BYU 163)</figcaption></figure> <p>These animals attracted little professional attention until the 1970s and 1980s, when Peter Galton reviewed many of the "hypsilophodonts" in a series of papers. In 1973, he and <a href="/wiki/James_A._Jensen" title="James A. Jensen">Jim Jensen</a> described a partial skeleton (<a href="/wiki/Brigham_Young_University" title="Brigham Young University">BYU ESM 163</a> as of Galton, 2007) missing the head, hands, and tail as <i>Nanosaurus</i>? <i>rex</i>, which had been damaged by other collectors prior to description.<sup id="cite_ref-GJ73_12-0" class="reference"><a href="#cite_note-GJ73-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> By 1977, he had concluded that <i>Nanosaurus agilis</i> was quite different from <i>N. rex</i> and the new skeleton, and coined <i><a href="/wiki/Othnielia" class="mw-redirect" title="Othnielia">Othnielia</a></i> for the latter species. The paper (primarily concerning the transcontinental nature of <i><a href="/wiki/Dryosaurus" title="Dryosaurus">Dryosaurus</a></i>) considered <i>Laosaurus consors</i> and <i>L. gracilis</i> synonyms of <i>O. rex</i> without elaboration, and considered <i>L. celer</i> an invalid <i><a href="/wiki/Nomen_nudum" title="Nomen nudum">nomen nudum</a></i>.<sup id="cite_ref-PMG77_13-0" class="reference"><a href="#cite_note-PMG77-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> </p><p>In 1990, <a href="/wiki/Robert_T._Bakker" title="Robert T. Bakker">Robert Bakker</a>, Peter Galton, James Siegwarth, and James Filla described remains of a dinosaur they named <i>Drinker nisti</i>. The name is somewhat ironic; <i>Drinker</i>, named for renowned <a href="/wiki/Palaeontologist" class="mw-redirect" title="Palaeontologist">palaeontologist</a> <a href="/wiki/Edward_Drinker_Cope" title="Edward Drinker Cope">Edward Drinker Cope</a> whose infamous "<a href="/wiki/Bone_wars" class="mw-redirect" title="Bone wars">bone wars</a>" with rival <a href="/wiki/Othniel_Charles_Marsh" title="Othniel Charles Marsh">Othniel Charles Marsh</a> produced many dinosaur fossils which are world-famous today, was described as a probable close relative of <i>Othnielia</i>, named for Marsh. The species name refers to the <b>N</b>ational <b>I</b>nstitute of <b>S</b>tandards and <b>T</b>echnology (<a href="/wiki/NIST" class="mw-redirect" title="NIST">NIST</a>). Discovered by Siegwarth and Filla in upper <a href="/wiki/Morrison_Formation" title="Morrison Formation">Morrison Formation</a> beds at <a href="/wiki/Como_Bluff" title="Como Bluff">Como Bluff</a>, <a href="/wiki/Wyoming" title="Wyoming">Wyoming</a>, it was <a href="/wiki/Holotype" title="Holotype">based on</a> a partial <a href="/wiki/Juvenile_(organism)" title="Juvenile (organism)">subadult</a> skeleton (listed as CPS 106 originally, then as Tate 4001 by Bakker 1996<sup id="cite_ref-galton2018_14-0" class="reference"><a href="#cite_note-galton2018-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup>) including partial jaws, vertebrae, and partial limbs. Several other specimens found in the same area were assigned to it, mostly consisting of vertebral and hindlimb remains, and teeth.<sup id="cite_ref-BGSF90_15-0" class="reference"><a href="#cite_note-BGSF90-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> The holotype specimen's current location is unknown; according to Carpenter and Galton (2018), the previous two institutions reported to have had it did not ever curate the specimen, and the collection it was originally said to be in never existed at all.<sup id="cite_ref-galton2018_14-1" class="reference"><a href="#cite_note-galton2018-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> </p> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Othnielosaurus.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/b/b9/Othnielosaurus.jpg/220px-Othnielosaurus.jpg" decoding="async" width="220" height="77" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/b/b9/Othnielosaurus.jpg/330px-Othnielosaurus.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/b/b9/Othnielosaurus.jpg/440px-Othnielosaurus.jpg 2x" data-file-width="3484" data-file-height="1227" /></a><figcaption><a href="/wiki/Life_restoration" class="mw-redirect" title="Life restoration">Life restoration</a></figcaption></figure> <p>Several decades later, in his 2007 study of the teeth of Morrison ornithischians, Galton concluded that the holotype femur of <i>Othnielia rex</i> is not diagnostic, and reassigned the BYU skeleton to <i>Laosaurus consors</i>, which is based on better material. As the genus <i>Laosaurus</i> is also based on nondiagnostic material, he gave the species <i>L. consors</i> its own genus, <i>Othnielosaurus</i>. As a result, in practical terms, what had been thought of as <i>Othnielia</i> is now known as <i>Othnielosaurus consors</i>. Regarding <i>Nanosaurus agilis</i>, Galton considered it a potentially valid basal <a href="/wiki/Ornithopoda" title="Ornithopoda">ornithopod</a>, and noted similarities to <a href="/wiki/Heterodontosauridae" title="Heterodontosauridae">heterodontosaurids</a> in the thigh bone. He tentatively assigned to it some teeth that had been referred to <i><a href="/wiki/Drinker_(dinosaur)" class="mw-redirect" title="Drinker (dinosaur)">Drinker</a></i>.<sup id="cite_ref-PMG07_4-2" class="reference"><a href="#cite_note-PMG07-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> </p><p>Another decade later, in 2018, Galton, alongside <a href="/wiki/Kenneth_Carpenter" title="Kenneth Carpenter">Kenneth Carpenter</a>, described a new ornithischian specimen. They found it very similar to the fragmentary holotype of <i>Nanosaurus</i>, but more clear in its anatomical features. Their new specimen was also found to display extreme similarity with the specimens of <i>Othnielosaurus</i> and <i>Drinker</i>. Due to the new data, they concluded that all three species, alongside <i>Othnielia</i>, represented the same animal, united under the name <i>Nanosaurus agilis</i>. This painted a new picture of a singular, very common small dinosaur known from a large amount of material.<sup id="cite_ref-galton2018_14-2" class="reference"><a href="#cite_note-galton2018-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> This conclusion has been recognized by papers since, some of which incorporating the new, all-encompassing taxon into their <a href="/wiki/Phylogenetic_analyses" class="mw-redirect" title="Phylogenetic analyses">phylogenetic analyses</a>.<sup id="cite_ref-herne2019_16-0" class="reference"><a href="#cite_note-herne2019-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-li2019_17-0" class="reference"><a href="#cite_note-li2019-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-wilson2019_18-0" class="reference"><a href="#cite_note-wilson2019-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Classification">Classification</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nanosaurus&action=edit&section=5" title="Edit section: Classification"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The <a href="/wiki/Cladogram" title="Cladogram">cladogram</a> below results from analysis by Herne et al., 2019.<sup id="cite_ref-herne2019_16-1" class="reference"><a href="#cite_note-herne2019-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> </p> <div class="clade"><style data-mw-deduplicate="TemplateStyles:r1258728058">body.skin-vector-2022 .mw-parser-output div.clade,body.skin-minerva .mw-parser-output div.clade{overflow-x:auto;overflow-y:hidden}body.skin-minerva .mw-parser-output div.clade p{font-size:inherit}.mw-parser-output table.clade{border-spacing:0;margin:0;font-size:100%;line-height:100%;border-collapse:separate;width:auto}.mw-parser-output table.clade table.clade{width:100%;line-height:inherit}.mw-parser-output table.clade td.clade-label{min-width:0.2em;width:0.2em;padding:0.1em 0.25em;vertical-align:bottom;text-align:center;border-left:1px solid;border-bottom:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-label::before,.mw-parser-output table.clade td.clade-slabel::before{content:"\2060 "}.mw-parser-output table.clade td.clade-fixed-width{overflow:hidden;text-overflow:ellipsis}.mw-parser-output table.clade td.clade-fixed-width:hover{overflow:visible}.mw-parser-output table.clade td.clade-label.first{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-label.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-slabel{padding:0.1em 0.25em;vertical-align:top;text-align:center;border-left:1px solid;white-space:nowrap}.mw-parser-output table.clade td.clade-slabel:hover{overflow:visible}.mw-parser-output table.clade td.clade-slabel.last{border-left:none;border-right:none}.mw-parser-output table.clade td.clade-slabel.reverse{border-left:none;border-right:1px solid}.mw-parser-output table.clade td.clade-bar{vertical-align:middle;text-align:left;padding:0 0.5em;position:relative}.mw-parser-output table.clade td.clade-bar.reverse{text-align:right;position:relative}.mw-parser-output table.clade td.clade-leaf{border:0;padding:0;text-align:left}.mw-parser-output table.clade td.clade-leafR{border:0;padding:0;text-align:right}.mw-parser-output table.clade td.clade-leaf.reverse{text-align:right}.mw-parser-output table.clade:hover span.linkA{background-color:yellow}.mw-parser-output table.clade:hover span.linkB{background-color:green}</style> <table class="clade" style="font-size:85%; line-height:85%"> <tbody><tr> <td class="clade-label first"><b><a href="/wiki/Ornithischia" title="Ornithischia">Ornithischia</a></b> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><a href="/wiki/Heterodontosauridae" title="Heterodontosauridae">Heterodontosauridae</a> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Eocursor" title="Eocursor">Eocursor</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><b><a href="/wiki/Thyreophora" title="Thyreophora">Thyreophora</a></b> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><b><a href="/wiki/Neornithischia" title="Neornithischia">Neornithischia</a></b> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Lesothosaurus" title="Lesothosaurus">Lesothosaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Agilisaurus" title="Agilisaurus">Agilisaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Hexinlusaurus" title="Hexinlusaurus">Hexinlusaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Yandusaurus" title="Yandusaurus">Yandusaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><b>Nanosaurus</b></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"><a href="/wiki/Jeholosauridae" title="Jeholosauridae">Jeholosauridae</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Haya_(dinosaur)" class="mw-redirect" title="Haya (dinosaur)">Haya</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Jeholosaurus" title="Jeholosaurus">Jeholosaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Changchunsaurus" title="Changchunsaurus">Changchunsaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><a href="/wiki/Thescelosauridae" title="Thescelosauridae">Thescelosauridae</a> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Orodromeus" title="Orodromeus">Orodromeus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Koreanosaurus" title="Koreanosaurus">Koreanosaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Zephyrosaurus" title="Zephyrosaurus">Zephyrosaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Yueosaurus" title="Yueosaurus">Yueosaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><i><a href="/wiki/Thescelosaurus" title="Thescelosaurus">Thescelosaurus</a></i> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"><b><a href="/wiki/Cerapoda" class="mw-redirect" title="Cerapoda">Cerapoda</a></b> </td> <td rowspan="2" class="clade-leaf"> <div><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1258728058"> <table class="clade"> <tbody><tr> <td class="clade-label first"> </td> <td rowspan="2" class="clade-leaf"> <p><b><a href="/wiki/Marginocephalia" title="Marginocephalia">Marginocephalia</a></b> </p> </td></tr> <tr> <td class="clade-slabel"> </td></tr> <tr> <td class="clade-label"> </td> <td rowspan="2" class="clade-leaf"> <p><b><a href="/wiki/Ornithopoda" title="Ornithopoda">Ornithopoda</a></b> </p> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> </td></tr> <tr> <td class="clade-slabel last"> </td></tr></tbody></table></div> <div class="mw-heading mw-heading2"><h2 id="Paleobiology_and_paleoecology">Paleobiology and paleoecology</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nanosaurus&action=edit&section=6" title="Edit section: Paleobiology and paleoecology"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Fighting_Othnielia.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8f/Fighting_Othnielia.jpg/220px-Fighting_Othnielia.jpg" decoding="async" width="220" height="128" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8f/Fighting_Othnielia.jpg/330px-Fighting_Othnielia.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8f/Fighting_Othnielia.jpg/440px-Fighting_Othnielia.jpg 2x" data-file-width="3231" data-file-height="1880" /></a><figcaption>Casts mounted as if fighting, Wyoming Dinosaur Center</figcaption></figure> <p><i>Nanosaurus</i> was one of the smaller members of the diverse <a href="/wiki/Morrison_Formation" title="Morrison Formation">Morrison Formation</a> dinosaur <a href="/wiki/Fauna" title="Fauna">fauna</a>, diminutive in comparison to the giant <a href="/wiki/Sauropod" class="mw-redirect" title="Sauropod">sauropods</a>.<sup id="cite_ref-JRF03a2_19-0" class="reference"><a href="#cite_note-JRF03a2-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> The Morrison Formation is interpreted as a <a href="/wiki/Semiarid" class="mw-redirect" title="Semiarid">semiarid</a> environment with distinct <a href="/wiki/Wet_season" title="Wet season">wet</a> and <a href="/wiki/Dry_season" title="Dry season">dry seasons</a>, and flat <a href="/wiki/Floodplain" title="Floodplain">floodplains</a>.<sup id="cite_ref-DAR89_20-0" class="reference"><a href="#cite_note-DAR89-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> Vegetation varied from <a href="/wiki/River" title="River">river</a>-lining <a href="/wiki/Gallery_forest" title="Gallery forest">gallery forests</a> of <a href="/wiki/Conifer" title="Conifer">conifers</a>, <a href="/wiki/Tree_fern" title="Tree fern">tree ferns</a>, and <a href="/wiki/Fern" title="Fern">ferns</a>, to fern <a href="/wiki/Savanna" title="Savanna">savannas</a> with rare trees.<sup id="cite_ref-KC06_21-0" class="reference"><a href="#cite_note-KC06-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> It has been a rich fossil hunting ground, holding fossils of <a href="/wiki/Chlorophyta" title="Chlorophyta">green algae</a>, <a href="/wiki/Fungi" class="mw-redirect" title="Fungi">fungi</a>, <a href="/wiki/Moss" title="Moss">mosses</a>, <a href="/wiki/Equisetum" title="Equisetum">horsetails</a>, ferns, <a href="/wiki/Cycad" title="Cycad">cycads</a>, <a href="/wiki/Ginkgo" title="Ginkgo">ginkgoes</a>, and several families of <a href="/wiki/Conifer" title="Conifer">conifers</a>. Other fossils discovered include <a href="/wiki/Bivalve" class="mw-redirect" title="Bivalve">bivalves</a>, <a href="/wiki/Snail" title="Snail">snails</a>, <a href="/wiki/Actinopterygii" title="Actinopterygii">ray-finned fishes</a>, <a href="/wiki/Frog" title="Frog">frogs</a>, <a href="/wiki/Salamander" title="Salamander">salamanders</a>, <a href="/wiki/Turtle" title="Turtle">turtles</a>, <a href="/wiki/Sphenodontia" class="mw-redirect" title="Sphenodontia">sphenodonts</a>, <a href="/wiki/Lizard" title="Lizard">lizards</a>, terrestrial and aquatic <a href="/wiki/Crocodylomorpha" title="Crocodylomorpha">crocodylomorphs</a>, several species of <a href="/wiki/Pterosaur" title="Pterosaur">pterosaur</a>, numerous dinosaur species, and early <a href="/wiki/Mammal" title="Mammal">mammals</a> such as <a href="/wiki/Docodonta" title="Docodonta">docodonts</a>, <a href="/wiki/Multituberculata" title="Multituberculata">multituberculates</a>, <a href="/wiki/Symmetrodonta" title="Symmetrodonta">symmetrodonts</a>, and <a href="/wiki/Triconodonta" class="mw-redirect" title="Triconodonta">triconodonts</a>. Such dinosaurs as the theropods <i><a href="/wiki/Ceratosaurus" title="Ceratosaurus">Ceratosaurus</a></i>, <i><a href="/wiki/Allosaurus" title="Allosaurus">Allosaurus</a></i>, <i><a href="/wiki/Ornitholestes" title="Ornitholestes">Ornitholestes</a></i>, and <i><a href="/wiki/Torvosaurus" title="Torvosaurus">Torvosaurus</a></i>, the sauropods <i><a href="/wiki/Apatosaurus" title="Apatosaurus">Apatosaurus</a></i>, <i><a href="/wiki/Brachiosaurus" title="Brachiosaurus">Brachiosaurus</a></i>, <i><a href="/wiki/Camarasaurus" title="Camarasaurus">Camarasaurus</a></i>, and <i><a href="/wiki/Diplodocus" title="Diplodocus">Diplodocus</a></i>, and the <a href="/wiki/Ornithischia" title="Ornithischia">ornithischians</a> <i><a href="/wiki/Camptosaurus" title="Camptosaurus">Camptosaurus</a></i>, <i><a href="/wiki/Dryosaurus" title="Dryosaurus">Dryosaurus</a></i>, and <i><a href="/wiki/Stegosaurus" title="Stegosaurus">Stegosaurus</a></i> are known from the Morrison.<sup id="cite_ref-DJCetal06_22-0" class="reference"><a href="#cite_note-DJCetal06-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup> <i>Nanosaurus</i> is present in stratigraphic zones 2-5.<sup id="cite_ref-foster-appendix_23-0" class="reference"><a href="#cite_note-foster-appendix-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup> </p><p>Typically, <i>Nanosaurus</i> has been interpreted like other hypsilophodonts as a small, <a href="/wiki/Cursorial" title="Cursorial">swift</a> <a href="/wiki/Herbivore" title="Herbivore">herbivore</a>,<sup id="cite_ref-NSWC042_24-0" class="reference"><a href="#cite_note-NSWC042-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup> although Bakker (1986) interpreted <i>Nanosaurus</i> as an <a href="/wiki/Omnivore" title="Omnivore">omnivore</a>.<sup id="cite_ref-RTB86_25-0" class="reference"><a href="#cite_note-RTB86-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Nanosaurus&action=edit&section=7" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-NSWC04-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-NSWC04_1-0">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite id="CITEREFNormanSues,_Hans-DieterWitmer,_Larry_M.Coria,_Rodolfo_A.2004" class="citation book cs1"><a href="/wiki/David_B._Norman" title="David B. Norman">Norman, David B.</a>; Sues, Hans-Dieter; Witmer, Larry M.; Coria, Rodolfo A. (2004). "Basal Ornithopoda". In Weishampel, David B.; Dodson, Peter; Osmólska, Halszka (eds.). <span class="id-lock-limited" title="Free access subject to limited trial, subscription normally required"><a rel="nofollow" class="external text" href="https://archive.org/details/dinosauriandedit00weis"><i>The Dinosauria</i></a></span> (2nd ed.). Berkeley: University of California Press. pp. <a rel="nofollow" class="external text" href="https://archive.org/details/dinosauriandedit00weis/page/n411">393</a>–412. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/0-520-24209-2" title="Special:BookSources/0-520-24209-2"><bdi>0-520-24209-2</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=Basal+Ornithopoda&rft.btitle=The+Dinosauria&rft.place=Berkeley&rft.pages=393-412&rft.edition=2nd&rft.pub=University+of+California+Press&rft.date=2004&rft.isbn=0-520-24209-2&rft.aulast=Norman&rft.aufirst=David+B.&rft.au=Sues%2C+Hans-Dieter&rft.au=Witmer%2C+Larry+M.&rft.au=Coria%2C+Rodolfo+A.&rft_id=https%3A%2F%2Farchive.org%2Fdetails%2Fdinosauriandedit00weis&rfr_id=info%3Asid%2Fen.wikipedia.org%3ANanosaurus" class="Z3988"></span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-2">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPaul2016" class="citation book cs1">Paul, Gregory S. (2016). <a rel="nofollow" class="external text" href="http://worldcat.org/oclc/985402380"><i>The Princeton Field Guide to Dinosaurs</i></a>. Princeton University Press. p. 306. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-1-78684-190-2" title="Special:BookSources/978-1-78684-190-2"><bdi>978-1-78684-190-2</bdi></a>. <a href="/wiki/OCLC_(identifier)" class="mw-redirect" title="OCLC (identifier)">OCLC</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/oclc/985402380">985402380</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=The+Princeton+Field+Guide+to+Dinosaurs&rft.pages=306&rft.pub=Princeton+University+Press&rft.date=2016&rft_id=info%3Aoclcnum%2F985402380&rft.isbn=978-1-78684-190-2&rft.aulast=Paul&rft.aufirst=Gregory+S.&rft_id=http%3A%2F%2Fworldcat.org%2Foclc%2F985402380&rfr_id=info%3Asid%2Fen.wikipedia.org%3ANanosaurus" class="Z3988"></span></span> </li> <li id="cite_note-SH04-3"><span class="mw-cite-backlink"><b><a href="#cite_ref-SH04_3-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFScott_Hartman" class="citation web cs1">Scott Hartman. <a rel="nofollow" class="external text" href="http://skeletaldrawing.com/psgallery/pages/othnielia.html">"othnielia"</a><span class="reference-accessdate">. Retrieved <span class="nowrap">2007-01-25</span></span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=unknown&rft.btitle=othnielia&rft.au=Scott+Hartman&rft_id=http%3A%2F%2Fskeletaldrawing.com%2Fpsgallery%2Fpages%2Fothnielia.html&rfr_id=info%3Asid%2Fen.wikipedia.org%3ANanosaurus" class="Z3988"></span></span> </li> <li id="cite_note-PMG07-4"><span class="mw-cite-backlink">^ <a href="#cite_ref-PMG07_4-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-PMG07_4-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-PMG07_4-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFGalton2007" class="citation book cs1">Galton, Peter M. (2007). "Teeth of ornithischian dinosaurs (mostly Ornithopoda) from the Morrison Formation (Upper Jurassic) of the western United States". In Carpenter, Kenneth (ed.). <i>Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs</i>. Bloomington and Indianapolis: Indiana University Press. pp. 17–47. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-253-34817-3" title="Special:BookSources/978-0-253-34817-3"><bdi>978-0-253-34817-3</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=Teeth+of+ornithischian+dinosaurs+%28mostly+Ornithopoda%29+from+the+Morrison+Formation+%28Upper+Jurassic%29+of+the+western+United+States&rft.btitle=Horns+and+Beaks%3A+Ceratopsian+and+Ornithopod+Dinosaurs&rft.place=Bloomington+and+Indianapolis&rft.pages=17-47&rft.pub=Indiana+University+Press&rft.date=2007&rft.isbn=978-0-253-34817-3&rft.aulast=Galton&rft.aufirst=Peter+M.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ANanosaurus" class="Z3988"></span></span> </li> <li id="cite_note-BG08-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-BG08_5-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFButlerGalton,_Peter_M.2008" class="citation journal cs1">Butler, Richard J.; Galton, Peter M. (2008). "The 'dermal armour' of the ornithopod dinosaur <i>Hypsilophodon</i> from the Wealden (Early Cretaceous: Barremian) of the Isle of Wight: a reappraisal". <i>Cretaceous Research</i>. <b>29</b> (4): 636–642. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2008CrRes..29..636B">2008CrRes..29..636B</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.cretres.2008.02.002">10.1016/j.cretres.2008.02.002</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Cretaceous+Research&rft.atitle=The+%27dermal+armour%27+of+the+ornithopod+dinosaur+Hypsilophodon+from+the+Wealden+%28Early+Cretaceous%3A+Barremian%29+of+the+Isle+of+Wight%3A+a+reappraisal&rft.volume=29&rft.issue=4&rft.pages=636-642&rft.date=2008&rft_id=info%3Adoi%2F10.1016%2Fj.cretres.2008.02.002&rft_id=info%3Abibcode%2F2008CrRes..29..636B&rft.aulast=Butler&rft.aufirst=Richard+J.&rft.au=Galton%2C+Peter+M.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ANanosaurus" class="Z3988"></span></span> </li> <li id="cite_note-OCM77a-6"><span class="mw-cite-backlink">^ <a href="#cite_ref-OCM77a_6-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-OCM77a_6-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMarsh1877" class="citation journal cs1"><a href="/wiki/Othniel_Charles_Marsh" title="Othniel Charles Marsh">Marsh, Othniel Charles</a> (1877). <a rel="nofollow" class="external text" href="https://zenodo.org/record/2429857">"Notice of some new vertebrate fossils"</a>. <i>American Journal of Science and Arts</i>. <b>14</b> (81): 249–256. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1877AmJS...14..249M">1877AmJS...14..249M</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.2475%2Fajs.s3-14.81.249">10.2475/ajs.s3-14.81.249</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:131230859">131230859</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=American+Journal+of+Science+and+Arts&rft.atitle=Notice+of+some+new+vertebrate+fossils&rft.volume=14&rft.issue=81&rft.pages=249-256&rft.date=1877&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A131230859%23id-name%3DS2CID&rft_id=info%3Adoi%2F10.2475%2Fajs.s3-14.81.249&rft_id=info%3Abibcode%2F1877AmJS...14..249M&rft.aulast=Marsh&rft.aufirst=Othniel+Charles&rft_id=https%3A%2F%2Fzenodo.org%2Frecord%2F2429857&rfr_id=info%3Asid%2Fen.wikipedia.org%3ANanosaurus" class="Z3988"></span></span> </li> <li id="cite_note-OCM77b-7"><span class="mw-cite-backlink">^ <a href="#cite_ref-OCM77b_7-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-OCM77b_7-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMarsh1877" class="citation journal cs1"><a href="/wiki/Othniel_Charles_Marsh" title="Othniel Charles Marsh">Marsh, Othniel Charles</a> (1877). <a rel="nofollow" class="external text" href="https://zenodo.org/record/1450040">"Notice of new dinosaurian reptiles from the Jurassic formations"</a>. <i>American Journal of Science and Arts</i>. <b>14</b> (84): 514–516. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/1877AmJS...14..514M">1877AmJS...14..514M</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.2475%2Fajs.s3-14.84.514">10.2475/ajs.s3-14.84.514</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:130488291">130488291</a>.</cite><span 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"Photogrammetry of the Oryctodromeus cubicularis type locality burrow and the utility of preexisting, standard field photographs for three dimensional digital reconstruction". <i>Historical Biology</i>. <b>32</b> (8): 1054–1061. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1080%2F08912963.2018.1563783">10.1080/08912963.2018.1563783</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:91500384">91500384</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Historical+Biology&rft.atitle=Photogrammetry+of+the+Oryctodromeus+cubicularis+type+locality+burrow+and+the+utility+of+preexisting%2C+standard+field+photographs+for+three+dimensional+digital+reconstruction&rft.volume=32&rft.issue=8&rft.pages=1054-1061&rft.date=2019&rft_id=info%3Adoi%2F10.1080%2F08912963.2018.1563783&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A91500384%23id-name%3DS2CID&rft.aulast=Wilson&rft.aufirst=John+P.&rft.au=Varricchio%2C+David+J.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ANanosaurus" class="Z3988"></span></span> </li> <li id="cite_note-JRF03a2-19"><span class="mw-cite-backlink"><b><a href="#cite_ref-JRF03a2_19-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFFoster2003" class="citation journal cs1">Foster, John R. 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this template">e</abbr></a></li></ul></div><div id="Ornithischia" style="font-size:114%;margin:0 4em"><a href="/wiki/Ornithischia" title="Ornithischia">Ornithischia</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="*_Kingdom:_Animalia_*_Phylum:_Chordata_*_Class:_Sauropsida_*_Clade:_Avemetatarsalia_*_Clade:_Dinosauria" style="font-size:114%;margin:0 4em"><span style="font-weight:normal; font-style:normal"><span style="font-size:85%;"> <ul><li>Kingdom: <a href="/wiki/Animal" title="Animal">Animalia</a></li> <li>Phylum: <a href="/wiki/Chordate" title="Chordate">Chordata</a></li> <li>Class: <a href="/wiki/Reptile" title="Reptile">Sauropsida</a></li> <li><i>Clade</i>: <a href="/wiki/Avemetatarsalia" title="Avemetatarsalia">Avemetatarsalia</a></li> <li><i>Clade</i>: <a href="/wiki/Dinosaur" title="Dinosaur">Dinosauria</a></li></ul> </span></span></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Avemetatarsalia" scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Avemetatarsalia" title="Avemetatarsalia">Avemetatarsalia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Avemetatarsalia" title="Template:Avemetatarsalia">Avemetatarsalia</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ornithischia" title="Ornithischia">Ornithischia</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see below↓</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible uncollapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="3"><div id="Ornithischia" style="font-size:114%;margin:0 4em"><a href="/wiki/Ornithischia" title="Ornithischia">Ornithischia</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Ornithischia" scope="row" class="navbox-group" style="width:1%"><div style="position: absolute; transform: translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Ornithischia" title="Ornithischia">Ornithischia</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Amanasaurus" title="Amanasaurus">Amanasaurus</a></i>?</li> <li><i><a href="/wiki/Asilisaurus" title="Asilisaurus">Asilisaurus</a></i>?</li> <li><i><a href="/wiki/Diodorus_scytobrachion" title="Diodorus scytobrachion">Diodorus</a></i>?</li> <li><i><a href="/wiki/Eocursor" title="Eocursor">Eocursor</a></i></li> <li><i><a href="/wiki/Eucoelophysis" title="Eucoelophysis">Eucoelophysis</a></i>?</li> <li><i><a href="/wiki/Gamatavus" title="Gamatavus">Gamatavus</a></i>?</li> <li><i><a href="/wiki/Ignotosaurus" title="Ignotosaurus">Ignotosaurus</a></i>?</li> <li><i><a href="/wiki/Kwanasaurus" title="Kwanasaurus">Kwanasaurus</a></i>?</li> <li><i><a href="/wiki/Laquintasaura" title="Laquintasaura">Laquintasaura</a></i></li> <li><i><a href="/wiki/Lesothosaurus" title="Lesothosaurus">Lesothosaurus</a></i></li> <li><i><a href="/wiki/Lewisuchus" title="Lewisuchus">Lewisuchus</a></i>?</li> <li><i><a href="/wiki/Lutungutali" title="Lutungutali">Lutungutali</a></i>?</li> <li><i><a href="/wiki/Pisanosaurus" title="Pisanosaurus">Pisanosaurus</a></i>?</li> <li><i><a href="/wiki/Sacisaurus" title="Sacisaurus">Sacisaurus</a></i>?</li> <li><i><a href="/wiki/Silesaurus" title="Silesaurus">Silesaurus</a></i>?</li> <li><i><a href="/wiki/Soumyasaurus" title="Soumyasaurus">Soumyasaurus</a></i>?</li> <li><i><a href="/wiki/Technosaurus" title="Technosaurus">Technosaurus</a></i>?</li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Heterodontosauridae" title="Heterodontosauridae">Heterodontosauridae</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Abrictosaurus" title="Abrictosaurus">Abrictosaurus</a></i></li> <li><i><a href="/wiki/Echinodon" title="Echinodon">Echinodon</a></i></li> <li><i><a href="/wiki/Fruitadens" title="Fruitadens">Fruitadens</a></i></li> <li><i><a href="/wiki/Heterodontosaurus" title="Heterodontosaurus">Heterodontosaurus</a></i></li> <li><i><a href="/wiki/Lycorhinus" title="Lycorhinus">Lycorhinus</a></i></li> <li><i><a href="/wiki/Manidens" title="Manidens">Manidens</a></i></li> <li><i><a href="/wiki/Pegomastax" title="Pegomastax">Pegomastax</a></i></li> <li><i><a href="/wiki/Tianyulong" title="Tianyulong">Tianyulong</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Thyreophora" title="Thyreophora">Thyreophora</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Emausaurus" title="Emausaurus">Emausaurus</a></i></li> <li><i><a href="/wiki/Scelidosaurus" title="Scelidosaurus">Scelidosaurus</a></i></li> <li><i><a href="/wiki/Scutellosaurus" title="Scutellosaurus">Scutellosaurus</a></i></li> <li><i><a href="/wiki/Yuxisaurus" title="Yuxisaurus">Yuxisaurus</a></i></li> <li><i><a href="/wiki/Jakapil" title="Jakapil">Jakapil</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ankylosauria" title="Ankylosauria">Ankylosauria</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Ankylosauria" title="Template:Ankylosauria">Ankylosauria</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Stegosauria" title="Stegosauria">Stegosauria</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Stegosauria" title="Template:Stegosauria">Stegosauria</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><div style="position: absolute; transform: translate(-50%,-50%) rotate(270deg);"><a href="/wiki/Neornithischia" title="Neornithischia">Neornithischia</a></div></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Agilisaurus" title="Agilisaurus">Agilisaurus</a></i></li> <li><i><a href="/wiki/Hexinlusaurus" title="Hexinlusaurus">Hexinlusaurus</a></i></li> <li><i><a href="/wiki/Hypsilophodon" title="Hypsilophodon">Hypsilophodon</a></i></li> <li><i><a href="/wiki/Kulindadromeus" title="Kulindadromeus">Kulindadromeus</a></i></li> <li><i><a href="/wiki/Minimocursor" title="Minimocursor">Minimocursor</a></i></li> <li><i><a class="mw-selflink selflink">Nanosaurus</a></i></li> <li><i><a href="/wiki/Phyllodon" title="Phyllodon">Phyllodon</a></i></li> <li><i><a href="/wiki/Sanxiasaurus" title="Sanxiasaurus">Sanxiasaurus</a></i></li> <li><i><a href="/wiki/Xiaosaurus" title="Xiaosaurus">Xiaosaurus</a></i></li> <li><i><a href="/wiki/Yandusaurus" title="Yandusaurus">Yandusaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Thescelosauridae" title="Thescelosauridae">Thescelosauridae</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Albertadromeus" title="Albertadromeus">Albertadromeus</a></i></li> <li><i><a href="/wiki/Changchunsaurus" title="Changchunsaurus">Changchunsaurus</a></i>?</li> <li><i><a href="/wiki/Changmiania" title="Changmiania">Changmiania</a></i>?</li> <li><i><a href="/wiki/Fona" title="Fona">Fona</a></i></li> <li><i><a href="/wiki/Haya_griva" title="Haya griva">Haya</a></i>?</li> <li><i><a href="/wiki/Jeholosaurus" title="Jeholosaurus">Jeholosaurus</a></i>?</li> <li><i><a href="/wiki/Koreanosaurus" title="Koreanosaurus">Koreanosaurus</a></i>?</li> <li><i><a class="mw-selflink selflink">Nanosaurus</a></i>?</li> <li><i><a href="/wiki/Nevadadromeus" title="Nevadadromeus">Nevadadromeus</a></i></li> <li><i><a href="/wiki/Orodromeus" title="Orodromeus">Orodromeus</a></i></li> <li><i><a href="/wiki/Oryctodromeus" title="Oryctodromeus">Oryctodromeus</a></i></li> <li><i><a href="/wiki/Parksosaurus" title="Parksosaurus">Parksosaurus</a></i></li> <li><i><a href="/wiki/Thescelosaurus" title="Thescelosaurus">Thescelosaurus</a></i></li> <li><i><a href="/wiki/Yueosaurus" title="Yueosaurus">Yueosaurus</a></i>?</li> <li><i><a href="/wiki/Zephyrosaurus" title="Zephyrosaurus">Zephyrosaurus</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ornithopoda" title="Ornithopoda">Ornithopoda</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Ornithopoda" title="Template:Ornithopoda">Ornithopoda</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Marginocephalia" title="Marginocephalia">Marginocephalia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Ceratopsia" title="Ceratopsia">Ceratopsia</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Ceratopsia" title="Template:Ceratopsia">Ceratopsia</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Pachycephalosauria" title="Pachycephalosauria">Pachycephalosauria</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><ul><li>see <a href="/wiki/Template:Pachycephalosauria" title="Template:Pachycephalosauria">Pachycephalosauria</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td><td class="noviewer navbox-image" rowspan="1" style="width:1px;padding:0 0 0 2px"><div><span style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Laquintasaura_fixed_by_Tom_Parker.png" class="mw-file-description" title="Laquintasaura venezuelae"><img alt="Laquintasaura venezuelae" src="//upload.wikimedia.org/wikipedia/commons/thumb/a/ae/Laquintasaura_fixed_by_Tom_Parker.png/100px-Laquintasaura_fixed_by_Tom_Parker.png" decoding="async" width="100" height="37" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/a/ae/Laquintasaura_fixed_by_Tom_Parker.png/150px-Laquintasaura_fixed_by_Tom_Parker.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/a/ae/Laquintasaura_fixed_by_Tom_Parker.png/200px-Laquintasaura_fixed_by_Tom_Parker.png 2x" data-file-width="1694" data-file-height="622" /></a></span></span> <p><span typeof="mw:File"><a href="/wiki/File:Heterodontosaurus_restoration.jpg" class="mw-file-description" title="Heterodontosaurus tucki"><img alt="Heterodontosaurus tucki" src="//upload.wikimedia.org/wikipedia/commons/thumb/1/14/Heterodontosaurus_restoration.jpg/100px-Heterodontosaurus_restoration.jpg" decoding="async" width="100" height="46" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/1/14/Heterodontosaurus_restoration.jpg/150px-Heterodontosaurus_restoration.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/1/14/Heterodontosaurus_restoration.jpg/200px-Heterodontosaurus_restoration.jpg 2x" data-file-width="1812" data-file-height="825" /></a></span> <span style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Scelidosaurus_harrisonii.png" class="mw-file-description" title="Scutellosaurus lawleri"><img alt="Scutellosaurus lawleri" src="//upload.wikimedia.org/wikipedia/commons/thumb/0/02/Scelidosaurus_harrisonii.png/100px-Scelidosaurus_harrisonii.png" decoding="async" width="100" height="37" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/0/02/Scelidosaurus_harrisonii.png/150px-Scelidosaurus_harrisonii.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/0/02/Scelidosaurus_harrisonii.png/200px-Scelidosaurus_harrisonii.png 2x" data-file-width="3497" data-file-height="1291" /></a></span></span> <span style="display: inline-block; transform: scaleX(-1);"><span typeof="mw:File"><a href="/wiki/File:Kulindadromeus_by_Tom_Parker.png" class="mw-file-description" title="Kulindadromeus zabaikalicus"><img alt="Kulindadromeus zabaikalicus" src="//upload.wikimedia.org/wikipedia/commons/thumb/d/da/Kulindadromeus_by_Tom_Parker.png/100px-Kulindadromeus_by_Tom_Parker.png" decoding="async" width="100" height="42" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/d/da/Kulindadromeus_by_Tom_Parker.png/150px-Kulindadromeus_by_Tom_Parker.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/d/da/Kulindadromeus_by_Tom_Parker.png/200px-Kulindadromeus_by_Tom_Parker.png 2x" data-file-width="2125" data-file-height="885" /></a></span></span> </p> <span typeof="mw:File"><a href="/wiki/File:Thescelosaurus_filamented.jpg" class="mw-file-description" title="Thescelosaurus neglectus"><img alt="Thescelosaurus neglectus" src="//upload.wikimedia.org/wikipedia/commons/thumb/b/b4/Thescelosaurus_filamented.jpg/100px-Thescelosaurus_filamented.jpg" decoding="async" width="100" height="33" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/b/b4/Thescelosaurus_filamented.jpg/150px-Thescelosaurus_filamented.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/b/b4/Thescelosaurus_filamented.jpg/200px-Thescelosaurus_filamented.jpg 2x" data-file-width="1140" data-file-height="379" /></a></span></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="See_also" style="font-size:114%;margin:0 4em">See also</div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Nomen_dubium" title="Nomen dubium">Nomina dubia</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alocodon" title="Alocodon">Alocodon</a></i></li> <li><i><a href="/wiki/Bienosaurus" title="Bienosaurus">Bienosaurus</a></i></li> <li><i><a href="/wiki/Craspedodon" title="Craspedodon">Craspedodon</a></i></li> <li><i><a href="/wiki/Fabrosaurus" title="Fabrosaurus">Fabrosaurus</a></i></li> <li><i><a href="/wiki/Ferganocephale" title="Ferganocephale">Ferganocephale</a></i></li> <li><i><a href="/wiki/Geranosaurus" title="Geranosaurus">Geranosaurus</a></i></li> <li><i><a href="/wiki/Gongbusaurus" title="Gongbusaurus">Gongbusaurus</a></i></li> <li><i><a href="/wiki/Laosaurus" title="Laosaurus">Laosaurus</a></i></li> <li><i><a href="/wiki/Notoceratops" title="Notoceratops">Notoceratops</a></i></li> <li><i><a href="/wiki/Siluosaurus" title="Siluosaurus">Siluosaurus</a></i></li> <li><i><a href="/wiki/Syngonosaurus" title="Syngonosaurus">Syngonosaurus</a></i></li> <li><i><a href="/wiki/Tatisaurus" title="Tatisaurus">Tatisaurus</a></i></li> <li><i><a href="/wiki/Tichosteus" title="Tichosteus">Tichosteus</a></i></li> <li><i><a href="/wiki/Trimucrodon" title="Trimucrodon">Trimucrodon</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Incertae_sedis" title="Incertae sedis">Incertae sedis</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Ajkaceratops" title="Ajkaceratops">Ajkaceratops</a></i></li> <li><i><a href="/wiki/Dravidosaurus" title="Dravidosaurus">Dravidosaurus</a></i>?</li> <li><i><a href="/wiki/Jakapil" title="Jakapil">Jakapil</a></i></li> <li><i><a href="/wiki/Serendipaceratops" title="Serendipaceratops">Serendipaceratops</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other taxa</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Ornithoscelida" title="Ornithoscelida">Ornithoscelida</a></li> <li><a href="/wiki/Phytodinosauria" title="Phytodinosauria">Phytodinosauria</a></li> <li><a href="/wiki/Silesauridae" title="Silesauridae">Silesauridae</a></li> <li><a href="/wiki/Hypsilophodontidae" title="Hypsilophodontidae">Hypsilophodontidae</a></li> <li><a href="/wiki/Jeholosauridae" title="Jeholosauridae">Jeholosauridae</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other articles</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/List_of_stratigraphic_units_with_ornithischian_tracks" title="List of stratigraphic units with ornithischian tracks">List of stratigraphic units with ornithischian tracks</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Nanosaurus</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q1827215" class="extiw" title="wikidata:Q1827215">Q1827215</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/47460279">47460279</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/4823089">4823089</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&id=1307280">1307280</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Open_Tree_of_Life" title="Open Tree of Life">Open Tree of Life</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://tree.opentreeoflife.org/taxonomy/browse?id=4946578">4946578</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Paleobiology_Database" title="Paleobiology Database">Paleobiology Database</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://paleobiodb.org/classic/basicTaxonInfo?taxon_no=38738">38738</a></span></span></li></ul> </div></td></tr></tbody></table></div>    </div><noscript><img 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