The formation of human populations in South and Central Asia

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class="collections-action-panel-form action-panel-content action-form action-panel-smaller-selectors" data-existing-collections-url="/list-existing-collections/" data-add-to-existing-collection-url="/add-to-existing-collection/" data-create-and-add-to-new-collection-url="/create-and-add-to-new-collection/" data-get-article-ids-by-search-url="/get-article-ids-by-search/" data-myncbi-max-collection-name-length="100" data-add-to-collection-max-amount="1000" data-collections-root-url="https://www.ncbi.nlm.nih.gov/myncbi/collections/"> <input type="hidden" name="csrfmiddlewaretoken" value="Zjw1SLo8jwK5oLn7QHYMivcac3dQ1qvFsM7jr9G6jB9itd0gc105ZFeQhKtFPbwQ"> <div class="choice-group" role="radiogroup"> <ul class="radio-group-items"> <li> <input type="radio" id="collections-action-panel-new" class="collections-new" name="collections" value="new" data-ga-category="save_share" data-ga-action="collections" data-ga-label="collections_radio_new"> <label for="collections-action-panel-new">Create a new collection</label> </li> <li> <input type="radio" id="collections-action-panel-existing" class="collections-existing" name="collections" value="existing" checked="true" data-ga-category="save_share" data-ga-action="collections" data-ga-label="collections_radio_existing"> <label for="collections-action-panel-existing">Add to an existing collection</label> </li> </ul> </div> <div class="controls-wrapper"> <div class="action-panel-control-wrap new-collections-controls"> <label for="collections-action-panel-add-to-new" class="action-panel-label required-field-asterisk"> Name your collection: </label> <input type="text" name="add-to-new-collection" id="collections-action-panel-add-to-new" class="collections-action-add-to-new" pattern="[^"&=<>\/]*" title="The following characters are not allowed in the Name field: "&=<>/" maxlength="100" data-ga-category="save_share" data-ga-action="create_collection" data-ga-label="non_favorties_collection"> <div class="collections-new-name-too-long usa-input-error-message selection-validation-message"> Name must be less than 100 characters </div> </div> <div class="action-panel-control-wrap existing-collections-controls"> <label for="collections-action-panel-add-to-existing" class="action-panel-label"> Choose a collection: </label> <select id="collections-action-panel-add-to-existing" class="action-panel-selector collections-action-add-to-existing" data-ga-category="save_share" data-ga-action="select_collection" data-ga-label="($('#collections-action-add-to-existing').val() === 'Favorites') ? 'Favorites' : 'non_favorites_collection'"> </select> <div class="collections-retry-load-on-error usa-input-error-message selection-validation-message"> Unable to load your collection due to an error<br> <a href="#">Please try again</a> </div> </div> </div> <div class="action-panel-actions"> <button class="action-panel-submit" type="submit" data-loading-label="Adding..." data-pinger-ignore data-ga-category="save_share" data-ga-action="collections" data-ga-label="add"> Add </button> <button class="action-panel-cancel" aria-label="Close 'Add to Collections' panel" ref="linksrc=close_collections_panel" aria-controls="collections-action-panel" aria-expanded="false" data-ga-category="save_share" data-ga-action="collections" data-ga-label="cancel"> Cancel </button> </div> </form> </div> </div> <div id="bibliography-action-panel" class="bibliography-action-panel action-panel in-progress-dots-panel" aria-hidden="true" data-bibliography-open-panel-enabled="false" data-bibliography-open-panel-url-hash="#open-bibliography-panel"> <div class="inner-wrap"> <h3 class="action-panel-heading"> Add to My Bibliography </h3> <form id="bibliography-action-panel-form" class="bibliography-action-panel-form action-panel-content action-form action-panel-smaller-selectors" data-add-to-bibliography-max-amount="100" data-add-to-bibliography-batch-size="10" data-bibliography-delegates-url="/list-bibliography-delegates/" data-add-to-bibliography-url="/add-to-bibliography/" data-get-article-ids-by-search-url="/get-article-ids-by-search/" data-mybib-root-url="https://www.ncbi.nlm.nih.gov/myncbi/collections/mybibliography/"> <input type="hidden" name="csrfmiddlewaretoken" value="Zjw1SLo8jwK5oLn7QHYMivcac3dQ1qvFsM7jr9G6jB9itd0gc105ZFeQhKtFPbwQ"> <div class="action-panel-control-wrap bibliographies-controls"> <div class="choice-group"> <ul class="bibliographies-action-add radio-group-items"> <li> <input name="bibliography" id="my-bibliography" class="my-bibliography" type="radio" checked/> <label for="my-bibliography">My Bibliography</label> </li> </ul> </div> </div> <div class="bibliographies-retry-load-on-error usa-input-error-message selection-validation-message"> Unable to load your delegates due to an error<br> <a href="#">Please try again</a> </div> <div class="action-panel-actions"> <button class="action-panel-submit" type="submit" data-loading-label="Adding..." data-pinger-ignore> Add </button> <button class="action-panel-cancel" aria-label="Close 'Add to bibliography' panel" ref="linksrc=close_bibliography_panel" aria-controls="bibliography-action-panel" aria-expanded="false" data-ga-category="save_share" data-ga-action="mybib" data-ga-label="cancel"> Cancel </button> </div> </form> </div> </div> <div id="saved-search-action-panel" class="saved-search-action-panel action-panel " aria-hidden="true" data-saved-search-open-panel-enabled="false" data-saved-search-open-panel-url-hash="#open-saved-search-panel"> <div class="inner-wrap"> <h2 class="action-panel-heading"> Your saved search </h2> <form id="saved-search-action-panel-form" class="saved-search-action-panel-form action-panel-content action-form" data-create-saved-search-url="/create-saved-search/" data-try-search-terms-url="/try-search-term/" data-saved-search-root-url="https://www.ncbi.nlm.nih.gov/myncbi/searches/"> <input type="hidden" name="csrfmiddlewaretoken" value="Zjw1SLo8jwK5oLn7QHYMivcac3dQ1qvFsM7jr9G6jB9itd0gc105ZFeQhKtFPbwQ"> <div class="action-panel-control-wrap"> <label for="saved-search-name" class="action-panel-label saved-search-name-label required-field-asterisk"> Name of saved search: </label> <input maxlength="200" type="text" name="saved-search-name" id="saved-search-name" class="saved-search-name" value="" required pattern="[^"&=<>\/]*" title="The following characters are not allowed in the Name field: "&=<>/"> </div> <div class="action-panel-control-wrap"> <label for="saved-search-term" class="action-panel-label required-field-asterisk"> Search terms: </label> <textarea name="saved-search-term" id="saved-search-term" class="saved-search-term" required></textarea> </div> <div class="test-search-term-wrap"> <a href="#" class="try-search-term">Test search terms</a> </div> <div class="choice-group action-panel-extra-margin-top"> <span class="action-panel-label" id="fieldset-label"> Would you like email updates of new search results? </span> <fieldset id="saved-search-alert" aria-describedby="fieldset-label"> <legend class="usa-sr-only">Saved Search Alert Radio Buttons</legend> <ul class="radio-group-items"> <li> <input type="radio" id="saved-search-alert-yes" class="saved-search-alert-yes" name="saved-search-alert" value="yes" checked> <label for="saved-search-alert-yes" class="action-panel-label">Yes</label> </li> <li> <input aria-label="No radio input" type="radio" id="saved-search-alert-no" class="saved-search-alert-no" name="saved-search-alert" value="no"> <label for="saved-search-alert-no" class="action-panel-label">No</label> </li> </ul> </fieldset> </div> <div class="alert-schedule-wrap"> <div class="action-panel-control-wrap"> <label class="action-panel-label"> Email: </label> <span aria-label="Email address" id="saved-search-email" class="action-panel-label"><span class="action-panel-label-bold"></span> (<a class="myncbi-account-settings" href="https://www.ncbi.nlm.nih.gov/account/settings/">change</a>)</span> </div> <div class="action-panel-control-wrap action-panel-extra-margin-top"> <label for="saved-search-frequency" class="action-panel-label"> Frequency: </label> <select id="saved-search-frequency" class="no-border-panel-selector saved-search-frequency"> <option value="monthly">Monthly</option> <option value="weekly">Weekly</option> <option value="daily">Daily</option> </select> </div> <div class="action-panel-control-wrap saved-search-monthly-additional"> <label for="saved-search-monthly-on-day" class="action-panel-label"> Which day? </label> <select id="saved-search-monthly-on-day" class="no-border-panel-selector"> <option value="Sunday">The first Sunday</option> <option value="Monday">The first Monday</option> <option value="Tuesday">The first Tuesday</option> <option value="Wednesday">The first Wednesday</option> <option value="Thursday">The first Thursday</option> <option value="Friday">The first Friday</option> <option value="Saturday">The first Saturday</option> <option value="day">The first day</option> <option value="weekday">The first weekday</option> </select> </div> <div class="action-panel-control-wrap saved-search-weekly-additional"> <label for="saved-search-weekly-on-day" class="action-panel-label"> Which day? </label> <select id="saved-search-weekly-on-day" class="no-border-panel-selector saved-search-weekly-on-day"> <option value="Sunday">Sunday</option> <option value="Monday">Monday</option> <option value="Tuesday">Tuesday</option> <option value="Wednesday">Wednesday</option> <option value="Thursday">Thursday</option> <option value="Friday">Friday</option> <option value="Saturday">Saturday</option> </select> </div> <div class="action-panel-control-wrap"> <label for="saved-search-report" class="action-panel-label"> Report format: </label> <select id="saved-search-report" class="no-border-panel-selector saved-search-report"> <option value="DocSum">Summary</option> <option value="DocSumText">Summary (text)</option> <option value="Abstract">Abstract</option> <option value="AbstractText">Abstract (text)</option> <option value="MEDLINE">PubMed</option> </select> </div> <div class="action-panel-control-wrap"> <label for="saved-search-amount" class="action-panel-label"> Send at most: </label> <select id="saved-search-amount" class="no-border-panel-selector saved-search-amount"> <option value="1">1 item</option> <option value="5" selected>5 items</option> <option value="10">10 items</option> <option value="20">20 items</option> <option value="50">50 items</option> <option value="100">100 items</option> <option value="200">200 items</option> </select> </div> <div> <input type="checkbox" id="saved-search-send-if-no-result" class="saved-search-send-if-no-result" name="saved-search-send-if-no-result"> <label for="saved-search-send-if-no-result" class="action-panel-label smaller-checkbox"> Send even when there aren't any new results </label> </div> <div class="action-panel-control-wrap option-text-in-email-wrap"> <label for="saved-search-email-text" class="action-panel-label"> Optional text in email: </label> <textarea name="saved-search-email-text" id="saved-search-email-text" class="saved-search-email-text"></textarea> </div> </div> <div class="action-panel-actions"> <button class="action-panel-submit" type="submit" data-loading-label="Saving..." data-ga-category="save_share" data-ga-action="alert" data-ga-label="save"> Save </button> <button class="action-panel-cancel" aria-label="Close 'Your saved search' panel" ref="linksrc=close_saved_search_panel" aria-controls="saved-search-action-panel" aria-expanded="false" data-ga-category="save_share" data-ga-action="alert" data-ga-label="cancel"> Cancel </button> </div> </form> </div> </div> <div id="citation-manager-action-panel" class="citation-manager-action-panel action-panel" aria-hidden="true"> <div class="inner-wrap"> <h2 class="action-panel-heading"> Create a file for external citation management software </h2> <form id="citation-manager-action-panel-form" class="action-panel-content action-form" action="/results-export-ids/" data-by-search-action="/results-export-search-data/" data-by-ids-action="/results-export-ids/" method="post" data-by-search-method="post" data-by-ids-method="post"> <input type="hidden" name="csrfmiddlewaretoken" value="Zjw1SLo8jwK5oLn7QHYMivcac3dQ1qvFsM7jr9G6jB9itd0gc105ZFeQhKtFPbwQ"> <input name="results-format" type="hidden" value="pubmed"/> <div class="action-panel-actions"> <button class="action-panel-submit" type="submit" data-loading-label="Sending..." data-ga-category="save_share" data-ga-action="citation_manager" data-ga-label="save"> Create file </button> <button class="action-panel-cancel" aria-label="Close 'Send citations to citation manager' panel" ref="linksrc=close_citation_manager_panel" aria-controls="citation-manager-action-panel" aria-expanded="false" data-ga-category="save_share" data-ga-action="citation_manager" data-ga-label="cancel"> Cancel </button> </div> </form> </div> </div> <div id="rss-action-panel" class="rss-action-panel action-panel " aria-hidden="true"> <div class="inner-wrap"> <h2 class="action-panel-heading"> Your RSS Feed </h2> <form id="rss-action-panel-form" class="rss-action-panel-form action-panel-content action-form" data-create-rss-feed-url="/create-rss-feed-url/" data-search-form-term-value=""> <input type="hidden" name="csrfmiddlewaretoken" value="Zjw1SLo8jwK5oLn7QHYMivcac3dQ1qvFsM7jr9G6jB9itd0gc105ZFeQhKtFPbwQ"> <div class="action-panel-control-wrap"> <label for="rss-name" class="action-panel-label required-field-asterisk"> Name of RSS Feed: </label> <input maxlength="200" placeholder="Name" type="text" name="rss-name" id="rss-name" class="rss-name" value='' required pattern="[^"&=<>\/]*" title="The following characters are not allowed in the Name field: "&=<>/"> </div> <div class="rss-limit-wrap"> <div class="action-panel-control-wrap action-panel-extra-margin-top"> <label for="rss-limit" class="action-panel-label"> Number of items displayed: </label> <select id="rss-limit" class="no-border-panel-selector rss-limit"> <option value="5">5</option> <option value="10">10</option> <option value="15" selected="selected">15</option> <option value="20">20</option> <option value="50">50</option> <option value="100">100</option> </select> </div> </div> <div class="action-panel-actions"> <button class="action-panel-submit" type="submit" data-loading-label="Creating..." data-ga-category="save_share" data-ga-action="alert" data-ga-label="save"> Create RSS </button> <button class="action-panel-cancel" aria-label="Close 'Your RSS' panel" ref="linksrc=close_rss_panel" aria-controls="rss-action-panel" aria-expanded="false" data-ga-category="save_share" data-ga-action="alert" data-ga-label="cancel"> Cancel </button> </div> <div class="action-panel-control-wrap rss-link-copy-wrap"> <label for="rss-link" class="usa-sr-only">RSS Link</label> <input placeholder="Your RSS Feed Link" type="text" name="rss-link" id="rss-link" class="rss-link" title="RSS Link"> <button type="button" disabled class="rss-link-copy-button disabled" data-ga-category="save_share" data-ga-action="rss" data-ga-label="copy"> Copy </button> </div> </form> </div> </div> </div> </div> <div class="article-page" id="article-page" data-article-pmid="31488661"> <aside class="page-sidebar"> <div class="inner-wrap"> <div class="full-text-links"> <div class="full-view"> <h3 class="title"> Full text links </h3> <div class="full-text-links-list"> <a class="link-item dialog-focus" 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data-collections-root-url="https://www.ncbi.nlm.nih.gov/myncbi/collections/"><input type="hidden" name="csrfmiddlewaretoken" value="Zjw1SLo8jwK5oLn7QHYMivcac3dQ1qvFsM7jr9G6jB9itd0gc105ZFeQhKtFPbwQ"><div class="choice-group" role="radiogroup"><ul class="radio-group-items"><li><input type="radio" id="collections-action-dialog-new" class="collections-new" name="collections" value="new" data-ga-category="collections_button" data-ga-action="click" data-ga-label="collections_radio_new"><label for="collections-action-dialog-new">Create a new collection</label></li><li><input type="radio" id="collections-action-dialog-existing" class="collections-existing" name="collections" value="existing" checked="true" data-ga-category="collections_button" data-ga-action="click" data-ga-label="collections_radio_existing"><label for="collections-action-dialog-existing">Add to an existing collection</label></li></ul></div><div class="controls-wrapper"><div class="action-panel-control-wrap new-collections-controls"><label for="collections-action-dialog-add-to-new" class="action-panel-label required-field-asterisk"> Name your collection: </label><input type="text" name="add-to-new-collection" id="collections-action-dialog-add-to-new" class="collections-action-add-to-new" pattern="[^"&=<>\/]*" title="The following characters are not allowed in the Name field: "&=<>/" maxlength="100" data-ga-category="collections_button" data-ga-action="create_collection" data-ga-label="non_favorties_collection"><div class="collections-new-name-too-long usa-input-error-message selection-validation-message"> Name must be less than 100 characters </div></div><div class="action-panel-control-wrap existing-collections-controls"><label for="collections-action-dialog-add-to-existing" class="action-panel-label"> Choose a collection: </label><select id="collections-action-dialog-add-to-existing" class="action-panel-selector collections-action-add-to-existing" data-ga-category="collections_button" data-ga-action="select_collection" data-ga-label="($('.collections-action-add-to-existing').val() === 'Favorites') ? 'Favorites' : 'non_favorites_collection'"></select><div class="collections-retry-load-on-error usa-input-error-message selection-validation-message"> Unable to load your collection due to an error<br><a href="#">Please try again</a></div></div></div><div class="action-panel-actions"><button class="action-panel-submit" type="submit" data-loading-label="Adding..." data-pinger-ignore data-ga-category="collections_button" data-ga-action="click" data-ga-label="add"> Add </button><button class="action-panel-cancel" aria-label="Close 'Add to Collections' panel" ref="linksrc=close_collections_panel" aria-controls="collections-action-panel" aria-expanded="false" data-ga-category="collections_button" data-ga-action="click" data-ga-label="cancel"> Cancel </button></div></form></div></div></div></div><button class="more-actions-button more-actions-dialog-trigger" title="Open dialog with more actions to take" ref="linksrc=more_actions_btn"></button><div class="display-options" 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id="full-view-journal-trigger" class="journal-actions-trigger trigger" ref="linksrc=journal_actions_btn" title="Science (New York, N.Y.)" tabindex="0" aria-controls="full-view-journal" aria-expanded="false" aria-label="Toggle dropdown menu for journal Science (New York, N.Y.)" data-pinger-ignore> Science </button><div id="full-view-journal" class="journal-actions-dropdown dropdown dropdown-container" hidden aria-label="Dropdown menu for journal Science (New York, N.Y.)"><div class="title">Actions</div><div class="content"><ul class="journal-actions-links"><li><a class="search-in-pubmed-link dropdown-block-link" href="/?term=%22Science%22%5Bjour%5D&sort=date&sort_order=desc" data-href="/?term=%22Science%22%5Bjour%5D&sort=date&sort_order=desc" ref="linksrc=search_in_pubmed_journal_name_link&journal_abbrev=Science" data-ga-category="search" data-ga-action="journal_link" data-ga-label="Science"> Search in PubMed </a></li><li><a class="search-in-nlm-catalog-link dropdown-block-link" ref ="linksrc=search_in_nlm_catalog_link" href="https://www.ncbi.nlm.nih.gov/nlmcatalog?term=%22Science%22%5BTitle+Abbreviation%5D" data-ga-category="search_catalog" data-ga-action="journal_link" data-ga-label="Science"> Search in NLM Catalog </a></li><li><a class="add-to-search-link dropdown-block-link" ref ="linksrc=add_to_search_link" role="button" data-search-term='"Science"[jour]' data-ga-category="search" data-ga-action="add_to_search" data-ga-label='"Science"[jour]' href="#"> Add to Search </a></li></ul></div></div></div><span class="period">. </span><span class="cit">2019 Sep 6;365(6457):eaat7487.</span> </div> <span class="citation-doi"> doi: 10.1126/science.aat7487. </span> </div> <h1 class="heading-title"> The formation of human populations in South and Central Asia </h1> <div class="inline-authors"> <div class="authors"> <div class="authors-list"> <span class="authors-list-item "><a class="full-name" href="/?term=Narasimhan+VM&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Vagheesh M Narasimhan">Vagheesh M Narasimhan</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu." href="#full-view-affiliation-1" ref="linksrc=author_aff"> 1 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Patterson+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Nick Patterson">Nick Patterson</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Broad Institute of Harvard and MIT, Cambridge, MA 02142, USA. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu." href="#full-view-affiliation-2" ref="linksrc=author_aff"> 2 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Radcliffe Institute for Advanced Study, Harvard University, Cambridge, MA 02138, USA." href="#full-view-affiliation-3" ref="linksrc=author_aff"> 3 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Moorjani+P&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Priya Moorjani">Priya Moorjani</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Molecular and Cell Biology, University of California, Berkeley, CA 94720, USA." href="#full-view-affiliation-4" ref="linksrc=author_aff"> 4 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Center for Computational Biology, University of California, Berkeley, CA 94720, USA." href="#full-view-affiliation-5" ref="linksrc=author_aff"> 5 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Rohland+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Nadin Rohland">Nadin Rohland</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Broad Institute of Harvard and MIT, Cambridge, MA 02142, USA." href="#full-view-affiliation-7" ref="linksrc=author_aff"> 7 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Bernardos+R&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Rebecca Bernardos">Rebecca Bernardos</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Mallick+S&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Swapan Mallick">Swapan Mallick</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Broad Institute of Harvard and MIT, Cambridge, MA 02142, USA." href="#full-view-affiliation-7" ref="linksrc=author_aff"> 7 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Lazaridis+I&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Iosif Lazaridis">Iosif Lazaridis</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Nakatsuka+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Nathan Nakatsuka">Nathan Nakatsuka</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Harvard-MIT Division of Health Sciences and Technology, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-9" ref="linksrc=author_aff"> 9 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Olalde+I&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Iñigo Olalde">Iñigo Olalde</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Lipson+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Mark Lipson">Mark Lipson</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Kim+AM&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Alexander M Kim">Alexander M Kim</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, Harvard University, Cambridge, MA 02138, USA." href="#full-view-affiliation-10" ref="linksrc=author_aff"> 10 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Olivieri+LM&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Luca M Olivieri">Luca M Olivieri</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="ISMEO - International Association of Mediterranean and Oriental Studies, Italian Archaeological Mission in Pakistan, 19200 Saidu Sharif (Swat), Pakistan." href="#full-view-affiliation-11" ref="linksrc=author_aff"> 11 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Coppa+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Alfredo Coppa">Alfredo Coppa</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Environmental Biology, Sapienza University, Rome 00185, Italy." href="#full-view-affiliation-12" ref="linksrc=author_aff"> 12 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Vidale+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Massimo Vidale">Massimo Vidale</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="ISMEO - International Association of Mediterranean and Oriental Studies, Italian Archaeological Mission in Pakistan, 19200 Saidu Sharif (Swat), Pakistan." href="#full-view-affiliation-11" ref="linksrc=author_aff"> 11 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Cultural Heritage: Archaeology and History of Art, Cinema and Music, University of Padua, Padua 35139, Italy." href="#full-view-affiliation-13" ref="linksrc=author_aff"> 13 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Mallory+J&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="James Mallory">James Mallory</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="School of Natural and Built Environment, Queen's University Belfast, Belfast BT7 1NN, Northern Ireland, UK." href="#full-view-affiliation-14" ref="linksrc=author_aff"> 14 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Moiseyev+V&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Vyacheslav Moiseyev">Vyacheslav Moiseyev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Peter the Great Museum of Anthropology and Ethnography (Kunstkamera), Russian Academy of Science, St. Petersburg 199034, Russia." href="#full-view-affiliation-15" ref="linksrc=author_aff"> 15 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Kitov+E&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Egor Kitov">Egor Kitov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Center of Physical Anthropology, Institute of Ethnology and Anthropology, Russian Academy of Sciences, Moscow 119991, Russia." href="#full-view-affiliation-16" ref="linksrc=author_aff"> 16 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan." href="#full-view-affiliation-17" ref="linksrc=author_aff"> 17 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Al-Farabi Kazakh National University, Almaty 050040, Kazakhstan." href="#full-view-affiliation-18" ref="linksrc=author_aff"> 18 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Monge+J&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Janet Monge">Janet Monge</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="University of Pennsylvania Museum of Archaeology and Anthropology, Philadelphia, PA 19104, USA." href="#full-view-affiliation-19" ref="linksrc=author_aff"> 19 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Adamski+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Nicole Adamski">Nicole Adamski</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Alex+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Neel Alex">Neel Alex</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Electrical Engineering and Computer Science, University of California, Berkeley, CA 94720, USA." href="#full-view-affiliation-20" ref="linksrc=author_aff"> 20 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Broomandkhoshbacht+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Nasreen Broomandkhoshbacht">Nasreen Broomandkhoshbacht</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Candilio+F&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Francesca Candilio">Francesca Candilio</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland." href="#full-view-affiliation-21" ref="linksrc=author_aff"> 21 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Soprintendenza Archeologia, Belle Arti e Paesaggio per la Città Metropolitana di Cagliari e le Province di Oristano e Sud Sardegna, Cagliari 09124, Italy." href="#full-view-affiliation-22" ref="linksrc=author_aff"> 22 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Callan+K&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Kimberly Callan">Kimberly Callan</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Cheronet+O&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Olivia Cheronet">Olivia Cheronet</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland." href="#full-view-affiliation-21" ref="linksrc=author_aff"> 21 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="School of Archaeology, University College Dublin, Dublin 4, Ireland." href="#full-view-affiliation-23" ref="linksrc=author_aff"> 23 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Evolutionary Anthropology, University of Vienna, 1090 Vienna, Austria." href="#full-view-affiliation-24" ref="linksrc=author_aff"> 24 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Culleton+BJ&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Brendan J Culleton">Brendan J Culleton</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institutes of Energy and the Environment, Pennsylvania State University, University Park, PA 16802, USA." href="#full-view-affiliation-25" ref="linksrc=author_aff"> 25 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Ferry+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Matthew Ferry">Matthew Ferry</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Fernandes+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Daniel Fernandes">Daniel Fernandes</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland." href="#full-view-affiliation-21" ref="linksrc=author_aff"> 21 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="School of Archaeology, University College Dublin, Dublin 4, Ireland." href="#full-view-affiliation-23" ref="linksrc=author_aff"> 23 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Evolutionary Anthropology, University of Vienna, 1090 Vienna, Austria." href="#full-view-affiliation-24" ref="linksrc=author_aff"> 24 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="CIAS, Department of Life Sciences, University of Coimbra, Coimbra 3000-456, Portugal." href="#full-view-affiliation-26" ref="linksrc=author_aff"> 26 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Freilich+S&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Suzanne Freilich">Suzanne Freilich</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Evolutionary Anthropology, University of Vienna, 1090 Vienna, Austria." href="#full-view-affiliation-24" ref="linksrc=author_aff"> 24 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Gamarra+B&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Beatriz Gamarra">Beatriz Gamarra</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland." href="#full-view-affiliation-21" ref="linksrc=author_aff"> 21 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="School of Archaeology, University College Dublin, Dublin 4, Ireland." href="#full-view-affiliation-23" ref="linksrc=author_aff"> 23 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Catalan Institute of Human Paleoecology and Social Evolution (IPHES), Tarragona 43007, Spain." href="#full-view-affiliation-27" ref="linksrc=author_aff"> 27 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Gaudio+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Daniel Gaudio">Daniel Gaudio</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland." href="#full-view-affiliation-21" ref="linksrc=author_aff"> 21 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="School of Archaeology, University College Dublin, Dublin 4, Ireland." href="#full-view-affiliation-23" ref="linksrc=author_aff"> 23 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Hajdinjak+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Mateja Hajdinjak">Mateja Hajdinjak</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Max Planck Institute for Evolutionary Anthropology, Leipzig 04103, Germany." href="#full-view-affiliation-28" ref="linksrc=author_aff"> 28 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Harney+%C3%89&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Éadaoin Harney">Éadaoin Harney</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-8" ref="linksrc=author_aff"> 8 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA 02138, USA." href="#full-view-affiliation-29" ref="linksrc=author_aff"> 29 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Harper+TK&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Thomas K Harper">Thomas K Harper</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, Pennsylvania State University, University Park, PA 16802, USA." href="#full-view-affiliation-30" ref="linksrc=author_aff"> 30 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Keating+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Denise Keating">Denise Keating</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland." href="#full-view-affiliation-21" ref="linksrc=author_aff"> 21 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Lawson+AM&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Ann Marie Lawson">Ann Marie Lawson</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Mah+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Matthew Mah">Matthew Mah</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Broad Institute of Harvard and MIT, Cambridge, MA 02142, USA." href="#full-view-affiliation-7" ref="linksrc=author_aff"> 7 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Mandl+K&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Kirsten Mandl">Kirsten Mandl</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Evolutionary Anthropology, University of Vienna, 1090 Vienna, Austria." href="#full-view-affiliation-24" ref="linksrc=author_aff"> 24 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Michel+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Megan Michel">Megan Michel</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Novak+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Mario Novak">Mario Novak</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland." href="#full-view-affiliation-21" ref="linksrc=author_aff"> 21 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute for Anthropological Research, Zagreb 10000, Croatia." href="#full-view-affiliation-31" ref="linksrc=author_aff"> 31 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Oppenheimer+J&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Jonas Oppenheimer">Jonas Oppenheimer</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Rai+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Niraj Rai">Niraj Rai</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="CSIR-Centre for Cellular and Molecular Biology, Hyderabad 500 007, India." href="#full-view-affiliation-32" ref="linksrc=author_aff"> 32 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Birbal Sahni Institute of Palaeosciences, Lucknow 226007, India." href="#full-view-affiliation-33" ref="linksrc=author_aff"> 33 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Sirak+K&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Kendra Sirak">Kendra Sirak</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland." href="#full-view-affiliation-21" ref="linksrc=author_aff"> 21 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, Emory University, Atlanta, GA 30322, USA." href="#full-view-affiliation-34" ref="linksrc=author_aff"> 34 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Slon+V&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Viviane Slon">Viviane Slon</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Max Planck Institute for Evolutionary Anthropology, Leipzig 04103, Germany." href="#full-view-affiliation-28" ref="linksrc=author_aff"> 28 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Stewardson+K&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Kristin Stewardson">Kristin Stewardson</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Zalzala+F&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Fatma Zalzala">Fatma Zalzala</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Zhang+Z&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Zhao Zhang">Zhao Zhang</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-6" ref="linksrc=author_aff"> 6 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Akhatov+G&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Gaziz Akhatov">Gaziz Akhatov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan." href="#full-view-affiliation-17" ref="linksrc=author_aff"> 17 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Bagashev+AN&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Anatoly N Bagashev">Anatoly N Bagashev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Tyumen Scientific Centre SB RAS, Institute of the Problems of Northern Development, Tyumen 625003, Russia." href="#full-view-affiliation-35" ref="linksrc=author_aff"> 35 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Bagnera+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Alessandra Bagnera">Alessandra Bagnera</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="ISMEO - International Association of Mediterranean and Oriental Studies, Italian Archaeological Mission in Pakistan, 19200 Saidu Sharif (Swat), Pakistan." href="#full-view-affiliation-11" ref="linksrc=author_aff"> 11 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Baitanayev+B&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Bauryzhan Baitanayev">Bauryzhan Baitanayev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan." href="#full-view-affiliation-17" ref="linksrc=author_aff"> 17 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Bendezu-Sarmiento+J&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Julio Bendezu-Sarmiento">Julio Bendezu-Sarmiento</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="CNRS-EXT500, Directeur de la Delegation Archaologique Francaise en Afghanistan (DAFA), Embassy of France in Kabul, Afghanistan." href="#full-view-affiliation-36" ref="linksrc=author_aff"> 36 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Bissembaev+AA&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Arman A Bissembaev">Arman A Bissembaev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan." href="#full-view-affiliation-17" ref="linksrc=author_aff"> 17 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Aktobe Regional Historical Museum, Aktobe 030006, Kazakhstan." href="#full-view-affiliation-37" ref="linksrc=author_aff"> 37 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Bonora+GL&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Gian Luca Bonora">Gian Luca Bonora</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Archaeology of Asia Department, ISMEO - International Association of Mediterranean and Oriental Studies, Rome RM00186, Italy." href="#full-view-affiliation-38" ref="linksrc=author_aff"> 38 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Chargynov+TT&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Temirlan T Chargynov">Temirlan T Chargynov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Kyrgyz National University, 720033 Bishkek, Kyrgyzstan." href="#full-view-affiliation-39" ref="linksrc=author_aff"> 39 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Chikisheva+T&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Tatiana Chikisheva">Tatiana Chikisheva</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute of Archaeology and Ethnography, Siberian Branch, Russian Academy of Sciences, Novosibirsk 630090, Russia." href="#full-view-affiliation-40" ref="linksrc=author_aff"> 40 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Dashkovskiy+PK&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Petr K Dashkovskiy">Petr K Dashkovskiy</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Political History, National and State-Confessional Relations, Altai State University, Barnaul 656049, Russia." href="#full-view-affiliation-41" ref="linksrc=author_aff"> 41 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Derevianko+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Anatoly Derevianko">Anatoly Derevianko</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute of Archaeology and Ethnography, Siberian Branch, Russian Academy of Sciences, Novosibirsk 630090, Russia." href="#full-view-affiliation-40" ref="linksrc=author_aff"> 40 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Dobe%C5%A1+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Miroslav Dobeš">Miroslav Dobeš</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute of Archaeology, Czech Academy of Sciences, Prague 118 01, Czech Republic." href="#full-view-affiliation-42" ref="linksrc=author_aff"> 42 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Douka+K&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Katerina Douka">Katerina Douka</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Max Planck Institute for the Science of Human History, Jena 07745, Germany." href="#full-view-affiliation-43" ref="linksrc=author_aff"> 43 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Oxford Radiocarbon Accelerator Unit, Research Laboratory for Archaeology and the History of Art, University of Oxford, Oxford OX1 3QY, UK." href="#full-view-affiliation-44" ref="linksrc=author_aff"> 44 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Dubova+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Nadezhda Dubova">Nadezhda Dubova</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Center of Physical Anthropology, Institute of Ethnology and Anthropology, Russian Academy of Sciences, Moscow 119991, Russia." href="#full-view-affiliation-16" ref="linksrc=author_aff"> 16 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Duisengali+MN&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Meiram N Duisengali">Meiram N Duisengali</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Aktobe Regional Historical Museum, Aktobe 030006, Kazakhstan." href="#full-view-affiliation-37" ref="linksrc=author_aff"> 37 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Enshin+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Dmitry Enshin">Dmitry Enshin</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Tyumen Scientific Centre SB RAS, Institute of the Problems of Northern Development, Tyumen 625003, Russia." href="#full-view-affiliation-35" ref="linksrc=author_aff"> 35 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Epimakhov+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Andrey Epimakhov">Andrey Epimakhov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute of History and Archaeology, Ural Branch RAS, Yekaterinburg 620990, Russia." href="#full-view-affiliation-45" ref="linksrc=author_aff"> 45 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="South Ural State University, Chelyabinsk 454080, Russia." href="#full-view-affiliation-46" ref="linksrc=author_aff"> 46 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Fribus+AV&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Alexey V Fribus">Alexey V Fribus</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Kemerovo State University, Kemerovo 650043, Russia." href="#full-view-affiliation-47" ref="linksrc=author_aff"> 47 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Fuller+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Dorian Fuller">Dorian Fuller</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute of Archaeology, University College London, London WC1H 0PY, UK." href="#full-view-affiliation-48" ref="linksrc=author_aff"> 48 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="School of Cultural Heritage, Northwest University, Shanxi, 710069, China." href="#full-view-affiliation-49" ref="linksrc=author_aff"> 49 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Goryachev+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Alexander Goryachev">Alexander Goryachev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Tyumen Scientific Centre SB RAS, Institute of the Problems of Northern Development, Tyumen 625003, Russia." href="#full-view-affiliation-35" ref="linksrc=author_aff"> 35 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Gromov+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Andrey Gromov">Andrey Gromov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Peter the Great Museum of Anthropology and Ethnography (Kunstkamera), Russian Academy of Science, St. Petersburg 199034, Russia." href="#full-view-affiliation-15" ref="linksrc=author_aff"> 15 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Grushin+SP&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Sergey P Grushin">Sergey P Grushin</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Ethnography and Museology, Altai State University, Barnaul 656049, Russia." href="#full-view-affiliation-50" ref="linksrc=author_aff"> 50 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Hanks+B&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Bryan Hanks">Bryan Hanks</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, University of Pittsburgh, Pittsburgh, PA 15260, USA." href="#full-view-affiliation-51" ref="linksrc=author_aff"> 51 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Judd+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Margaret Judd">Margaret Judd</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, University of Pittsburgh, Pittsburgh, PA 15260, USA." href="#full-view-affiliation-51" ref="linksrc=author_aff"> 51 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Kazizov+E&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Erlan Kazizov">Erlan Kazizov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan." href="#full-view-affiliation-17" ref="linksrc=author_aff"> 17 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Khokhlov+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Aleksander Khokhlov">Aleksander Khokhlov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Samara State University of Social Sciences and Education, Samara 443099, Russia." href="#full-view-affiliation-52" ref="linksrc=author_aff"> 52 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Krygin+AP&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Aleksander P Krygin">Aleksander P Krygin</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="West Kazakhstan Regional Center for History and Archaeology, Uralsk 090000, Kazakhstan." href="#full-view-affiliation-53" ref="linksrc=author_aff"> 53 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Kupriyanova+E&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Elena Kupriyanova">Elena Kupriyanova</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Scientific and Educational Center of Study on the Problem of Nature and Man, Chelyabinsk State University, Chelyabinsk 454021, Russia." href="#full-view-affiliation-54" ref="linksrc=author_aff"> 54 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Kuznetsov+P&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Pavel Kuznetsov">Pavel Kuznetsov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Samara State University of Social Sciences and Education, Samara 443099, Russia." href="#full-view-affiliation-52" ref="linksrc=author_aff"> 52 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Luiselli+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Donata Luiselli">Donata Luiselli</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Cultural Heritage, University of Bologna, 48121 Ravenna, Italy." href="#full-view-affiliation-55" ref="linksrc=author_aff"> 55 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Maksudov+F&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Farhod Maksudov">Farhod Maksudov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute for Archaeological Research, Uzbekistan Academy of Sciences, Samarkand 140151, Uzbekistan." href="#full-view-affiliation-56" ref="linksrc=author_aff"> 56 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Mamedov+AM&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Aslan M Mamedov">Aslan M Mamedov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Center for Research, Restoration and Protection of Historical and Cultural Heritage of Aktobe Region, Aktobe 030007, Kazakhstan." href="#full-view-affiliation-57" ref="linksrc=author_aff"> 57 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Mamirov+TB&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Talgat B Mamirov">Talgat B Mamirov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan." href="#full-view-affiliation-17" ref="linksrc=author_aff"> 17 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Meiklejohn+C&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Christopher Meiklejohn">Christopher Meiklejohn</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, University of Winnipeg, Winnipeg, MB R3B 2E9, Canada." href="#full-view-affiliation-58" ref="linksrc=author_aff"> 58 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Merrett+DC&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Deborah C Merrett">Deborah C Merrett</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Simon Fraser University, Burnaby, BC V5A 1S6, Canada." href="#full-view-affiliation-59" ref="linksrc=author_aff"> 59 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Micheli+R&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Roberto Micheli">Roberto Micheli</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="ISMEO - International Association of Mediterranean and Oriental Studies, Italian Archaeological Mission in Pakistan, 19200 Saidu Sharif (Swat), Pakistan." href="#full-view-affiliation-11" ref="linksrc=author_aff"> 11 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="MiBAC - Ministero per i Beni e le Attività Culturali - Soprintendenza Archeologia, belle arti e paesaggio del Friuli Venezia Giulia, 34135 Trieste, Italy." href="#full-view-affiliation-60" ref="linksrc=author_aff"> 60 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Mochalov+O&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Oleg Mochalov">Oleg Mochalov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Samara State University of Social Sciences and Education, Samara 443099, Russia." href="#full-view-affiliation-52" ref="linksrc=author_aff"> 52 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Mustafokulov+S&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Samariddin Mustafokulov">Samariddin Mustafokulov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute for Archaeological Research, Uzbekistan Academy of Sciences, Samarkand 140151, Uzbekistan." href="#full-view-affiliation-56" ref="linksrc=author_aff"> 56 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Afrosiab Museum, Samarkand 140151, Uzbekistan." href="#full-view-affiliation-61" ref="linksrc=author_aff"> 61 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Nayak+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Ayushi Nayak">Ayushi Nayak</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Max Planck Institute for the Science of Human History, Jena 07745, Germany." href="#full-view-affiliation-43" ref="linksrc=author_aff"> 43 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Pettener+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Davide Pettener">Davide Pettener</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Biological, Geological and Environmental Sciences, Alma Mater Studiorum - University of Bologna, Bologna 40126, Italy." href="#full-view-affiliation-62" ref="linksrc=author_aff"> 62 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Potts+R&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Richard Potts">Richard Potts</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Human Origins Program, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013, USA." href="#full-view-affiliation-63" ref="linksrc=author_aff"> 63 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Razhev+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Dmitry Razhev">Dmitry Razhev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Tyumen Scientific Centre SB RAS, Institute of the Problems of Northern Development, Tyumen 625003, Russia." href="#full-view-affiliation-35" ref="linksrc=author_aff"> 35 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Rykun+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Marina Rykun">Marina Rykun</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="National Research Tomsk State University, Tomsk 634050, Russia." href="#full-view-affiliation-64" ref="linksrc=author_aff"> 64 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Sarno+S&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Stefania Sarno">Stefania Sarno</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Biological, Geological and Environmental Sciences, Alma Mater Studiorum - University of Bologna, Bologna 40126, Italy." href="#full-view-affiliation-62" ref="linksrc=author_aff"> 62 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Savenkova+TM&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Tatyana M Savenkova">Tatyana M Savenkova</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="F. Voino-Yasenetsky Krasnoyarsk State Medical University, Krasnoyarsk 660022, Russia." href="#full-view-affiliation-65" ref="linksrc=author_aff"> 65 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Sikhymbaeva+K&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Kulyan Sikhymbaeva">Kulyan Sikhymbaeva</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Central State Museum Republic of Kazakhstan, Samal-1 Microdistrict, Almaty 050010, Kazakhstan." href="#full-view-affiliation-66" ref="linksrc=author_aff"> 66 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Slepchenko+SM&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Sergey M Slepchenko">Sergey M Slepchenko</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Tyumen Scientific Centre SB RAS, Institute of the Problems of Northern Development, Tyumen 625003, Russia." href="#full-view-affiliation-35" ref="linksrc=author_aff"> 35 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Soltobaev+OA&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Oroz A Soltobaev">Oroz A Soltobaev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Kyrgyz National University, 720033 Bishkek, Kyrgyzstan." href="#full-view-affiliation-39" ref="linksrc=author_aff"> 39 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Stepanova+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Nadezhda Stepanova">Nadezhda Stepanova</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute of Archaeology and Ethnography, Siberian Branch, Russian Academy of Sciences, Novosibirsk 630090, Russia." href="#full-view-affiliation-40" ref="linksrc=author_aff"> 40 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Svyatko+S&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Svetlana Svyatko">Svetlana Svyatko</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Peter the Great Museum of Anthropology and Ethnography (Kunstkamera), Russian Academy of Science, St. Petersburg 199034, Russia." href="#full-view-affiliation-15" ref="linksrc=author_aff"> 15 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="CHRONO Centre for Climate, the Environment, and Chronology, Queen's University of Belfast, Belfast BT7 1NN, Northern Ireland, UK." href="#full-view-affiliation-67" ref="linksrc=author_aff"> 67 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Tabaldiev+K&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Kubatbek Tabaldiev">Kubatbek Tabaldiev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Kyrgyz-Turkish Manas University, Bishkek, Kyrgyzstan." href="#full-view-affiliation-68" ref="linksrc=author_aff"> 68 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Teschler-Nicola+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Maria Teschler-Nicola">Maria Teschler-Nicola</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Evolutionary Anthropology, University of Vienna, 1090 Vienna, Austria." href="#full-view-affiliation-24" ref="linksrc=author_aff"> 24 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, Natural History Museum Vienna, 1010 Vienna, Austria." href="#full-view-affiliation-69" ref="linksrc=author_aff"> 69 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Tishkin+AA&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Alexey A Tishkin">Alexey A Tishkin</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Ethnography and Museology, The Laboratory of Interdisciplinary Studies in Archaeology of Western Siberia and Altai, Altai State University, Barnaul 656049, Russia." href="#full-view-affiliation-70" ref="linksrc=author_aff"> 70 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Tkachev+VV&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Vitaly V Tkachev">Vitaly V Tkachev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute of Steppe, Ural Branch RAS, Orenburg 460000, Russia." href="#full-view-affiliation-71" ref="linksrc=author_aff"> 71 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Vasilyev+S&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Sergey Vasilyev">Sergey Vasilyev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Center of Physical Anthropology, Institute of Ethnology and Anthropology, Russian Academy of Sciences, Moscow 119991, Russia." href="#full-view-affiliation-16" ref="linksrc=author_aff"> 16 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Center for Egyptological Studies RAS, Moscow 119991, Russia." href="#full-view-affiliation-72" ref="linksrc=author_aff"> 72 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Velem%C3%ADnsk%C3%BD+P&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Petr Velemínský">Petr Velemínský</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, National Museum, Prague 115 79, Czech Republic." href="#full-view-affiliation-73" ref="linksrc=author_aff"> 73 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Voyakin+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Dmitriy Voyakin">Dmitriy Voyakin</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan." href="#full-view-affiliation-17" ref="linksrc=author_aff"> 17 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Archaeological Expertise LLP, Almaty 050060, Kazakhstan." href="#full-view-affiliation-74" ref="linksrc=author_aff"> 74 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Yermolayeva+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Antonina Yermolayeva">Antonina Yermolayeva</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan." href="#full-view-affiliation-17" ref="linksrc=author_aff"> 17 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Zahir+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Muhammad Zahir">Muhammad Zahir</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Max Planck Institute for the Science of Human History, Jena 07745, Germany." href="#full-view-affiliation-43" ref="linksrc=author_aff"> 43 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Hazara University, Mansehra 21300, Pakistan." href="#full-view-affiliation-75" ref="linksrc=author_aff"> 75 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Zubkov+VS&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Valery S Zubkov">Valery S Zubkov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="N.F. Katanov Khakassia State University, Abakan 655017, Russia." href="#full-view-affiliation-76" ref="linksrc=author_aff"> 76 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Zubova+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Alisa Zubova">Alisa Zubova</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Peter the Great Museum of Anthropology and Ethnography (Kunstkamera), Russian Academy of Science, St. Petersburg 199034, Russia." href="#full-view-affiliation-15" ref="linksrc=author_aff"> 15 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Shinde+VS&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Vasant S Shinde">Vasant S Shinde</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Deccan College Post-Graduate and Research Institute, Pune 411006, India." href="#full-view-affiliation-77" ref="linksrc=author_aff"> 77 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Lalueza-Fox+C&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Carles Lalueza-Fox">Carles Lalueza-Fox</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute of Evolutionary Biology, CSIC-Universitat Pompeu Fabra, Barcelona 08003, Spain." href="#full-view-affiliation-78" ref="linksrc=author_aff"> 78 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Meyer+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Matthias Meyer">Matthias Meyer</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Max Planck Institute for Evolutionary Anthropology, Leipzig 04103, Germany." href="#full-view-affiliation-28" ref="linksrc=author_aff"> 28 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Anthony+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="David Anthony">David Anthony</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Anthropology Department, Hartwick College, Oneonta, NY 13820, USA." href="#full-view-affiliation-79" ref="linksrc=author_aff"> 79 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Boivin+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Nicole Boivin">Nicole Boivin</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Max Planck Institute for the Science of Human History, Jena 07745, Germany." href="#full-view-affiliation-43" ref="linksrc=author_aff"> 43 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Thangaraj+K&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Kumarasamy Thangaraj">Kumarasamy Thangaraj</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="CSIR-Centre for Cellular and Molecular Biology, Hyderabad 500 007, India." href="#full-view-affiliation-32" ref="linksrc=author_aff"> 32 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Kennett+DJ&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Douglas J Kennett">Douglas J Kennett</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institutes of Energy and the Environment, Pennsylvania State University, University Park, PA 16802, USA." href="#full-view-affiliation-25" ref="linksrc=author_aff"> 25 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, Pennsylvania State University, University Park, PA 16802, USA." href="#full-view-affiliation-30" ref="linksrc=author_aff"> 30 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, University of California, Santa Barbara, CA 93106, USA." href="#full-view-affiliation-80" ref="linksrc=author_aff"> 80 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Frachetti+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Michael Frachetti">Michael Frachetti</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, Washington University in St. Louis, St. Louis, MO 63112, USA. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu." href="#full-view-affiliation-81" ref="linksrc=author_aff"> 81 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Spatial Analysis, Interpretation, and Exploration Laboratory, Washington University in St. Louis, St. Louis, MO 63112, USA." href="#full-view-affiliation-82" ref="linksrc=author_aff"> 82 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Pinhasi+R&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Ron Pinhasi">Ron Pinhasi</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu." href="#full-view-affiliation-83" ref="linksrc=author_aff"> 83 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Evolutionary Anthropology, University of Vienna, 1090 Vienna, Austria." href="#full-view-affiliation-24" ref="linksrc=author_aff"> 24 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Reich+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="David Reich">David Reich</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu." href="#full-view-affiliation-1" ref="linksrc=author_aff"> 1 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Broad Institute of Harvard and MIT, Cambridge, MA 02142, USA." href="#full-view-affiliation-7" ref="linksrc=author_aff"> 7 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#full-view-affiliation-8" ref="linksrc=author_aff"> 8 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Max Planck-Harvard Research Center for the Archaeoscience of the Ancient Mediterranean, Cambridge, MA 02138, USA." href="#full-view-affiliation-84" ref="linksrc=author_aff"> 84 </a></sup></span> </div> </div> </div> <div class="short-article-details"> Affiliations <button class="more-details" id="toggle-authors" data-alt-text="Collapse" aria-controls="expanded-authors" aria-expanded="false" data-pinger-ignore data-fetch-url="/31488661/long-authors/" data-id-prefix=""> <span class="title"> Expand </span> </button> </div> <div class="extended-article-details" id="expanded-authors" aria-hidden="true"> <div class="expanded-authors" id="full-view-expanded-authors"> <div class="affiliations"> <h3 class="title"> Affiliations </h3> <ul class="item-list"> <li data-affiliation-id="full-view-affiliation-1" id="full-view-affiliation-1" ><sup class="key">1</sup> Department of Genetics, Harvard Medical School, Boston, MA 02115, USA. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu.</li> <li data-affiliation-id="full-view-affiliation-2" id="full-view-affiliation-2" ><sup class="key">2</sup> Broad Institute of Harvard and MIT, Cambridge, MA 02142, USA. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu.</li> <li data-affiliation-id="full-view-affiliation-3" id="full-view-affiliation-3" ><sup class="key">3</sup> Radcliffe Institute for Advanced Study, Harvard University, Cambridge, MA 02138, USA.</li> <li data-affiliation-id="full-view-affiliation-4" id="full-view-affiliation-4" ><sup class="key">4</sup> Department of Molecular and Cell Biology, University of California, Berkeley, CA 94720, USA.</li> <li data-affiliation-id="full-view-affiliation-5" id="full-view-affiliation-5" ><sup class="key">5</sup> Center for Computational Biology, University of California, Berkeley, CA 94720, USA.</li> <li data-affiliation-id="full-view-affiliation-6" id="full-view-affiliation-6" ><sup class="key">6</sup> Department of Genetics, Harvard Medical School, Boston, MA 02115, USA.</li> <li data-affiliation-id="full-view-affiliation-7" id="full-view-affiliation-7" ><sup class="key">7</sup> Broad Institute of Harvard and MIT, Cambridge, MA 02142, USA.</li> <li data-affiliation-id="full-view-affiliation-8" id="full-view-affiliation-8" ><sup class="key">8</sup> Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA.</li> <li data-affiliation-id="full-view-affiliation-9" id="full-view-affiliation-9" ><sup class="key">9</sup> Harvard-MIT Division of Health Sciences and Technology, Harvard Medical School, Boston, MA 02115, USA.</li> <li data-affiliation-id="full-view-affiliation-10" id="full-view-affiliation-10" ><sup class="key">10</sup> Department of Anthropology, Harvard University, Cambridge, MA 02138, USA.</li> <li data-affiliation-id="full-view-affiliation-11" id="full-view-affiliation-11" ><sup class="key">11</sup> ISMEO - International Association of Mediterranean and Oriental Studies, Italian Archaeological Mission in Pakistan, 19200 Saidu Sharif (Swat), Pakistan.</li> <li data-affiliation-id="full-view-affiliation-12" id="full-view-affiliation-12" ><sup class="key">12</sup> Department of Environmental Biology, Sapienza University, Rome 00185, Italy.</li> <li data-affiliation-id="full-view-affiliation-13" id="full-view-affiliation-13" ><sup class="key">13</sup> Department of Cultural Heritage: Archaeology and History of Art, Cinema and Music, University of Padua, Padua 35139, Italy.</li> <li data-affiliation-id="full-view-affiliation-14" id="full-view-affiliation-14" ><sup class="key">14</sup> School of Natural and Built Environment, Queen's University Belfast, Belfast BT7 1NN, Northern Ireland, UK.</li> <li data-affiliation-id="full-view-affiliation-15" id="full-view-affiliation-15" ><sup class="key">15</sup> Peter the Great Museum of Anthropology and Ethnography (Kunstkamera), Russian Academy of Science, St. Petersburg 199034, Russia.</li> <li data-affiliation-id="full-view-affiliation-16" id="full-view-affiliation-16" ><sup class="key">16</sup> Center of Physical Anthropology, Institute of Ethnology and Anthropology, Russian Academy of Sciences, Moscow 119991, Russia.</li> <li data-affiliation-id="full-view-affiliation-17" id="full-view-affiliation-17" ><sup class="key">17</sup> A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan.</li> <li data-affiliation-id="full-view-affiliation-18" id="full-view-affiliation-18" ><sup class="key">18</sup> Al-Farabi Kazakh National University, Almaty 050040, Kazakhstan.</li> <li data-affiliation-id="full-view-affiliation-19" id="full-view-affiliation-19" ><sup class="key">19</sup> University of Pennsylvania Museum of Archaeology and Anthropology, Philadelphia, PA 19104, USA.</li> <li data-affiliation-id="full-view-affiliation-20" id="full-view-affiliation-20" ><sup class="key">20</sup> Department of Electrical Engineering and Computer Science, University of California, Berkeley, CA 94720, USA.</li> <li data-affiliation-id="full-view-affiliation-21" id="full-view-affiliation-21" ><sup class="key">21</sup> Earth Institute, University College Dublin, Dublin 4, Ireland.</li> <li data-affiliation-id="full-view-affiliation-22" id="full-view-affiliation-22" ><sup class="key">22</sup> Soprintendenza Archeologia, Belle Arti e Paesaggio per la Città Metropolitana di Cagliari e le Province di Oristano e Sud Sardegna, Cagliari 09124, Italy.</li> <li data-affiliation-id="full-view-affiliation-23" id="full-view-affiliation-23" ><sup class="key">23</sup> School of Archaeology, University College Dublin, Dublin 4, Ireland.</li> <li data-affiliation-id="full-view-affiliation-24" id="full-view-affiliation-24" ><sup class="key">24</sup> Department of Evolutionary Anthropology, University of Vienna, 1090 Vienna, Austria.</li> <li data-affiliation-id="full-view-affiliation-25" id="full-view-affiliation-25" ><sup class="key">25</sup> Institutes of Energy and the Environment, Pennsylvania State University, University Park, PA 16802, USA.</li> <li data-affiliation-id="full-view-affiliation-26" id="full-view-affiliation-26" ><sup class="key">26</sup> CIAS, Department of Life Sciences, University of Coimbra, Coimbra 3000-456, Portugal.</li> <li data-affiliation-id="full-view-affiliation-27" id="full-view-affiliation-27" ><sup class="key">27</sup> Catalan Institute of Human Paleoecology and Social Evolution (IPHES), Tarragona 43007, Spain.</li> <li data-affiliation-id="full-view-affiliation-28" id="full-view-affiliation-28" ><sup class="key">28</sup> Max Planck Institute for Evolutionary Anthropology, Leipzig 04103, Germany.</li> <li data-affiliation-id="full-view-affiliation-29" id="full-view-affiliation-29" ><sup class="key">29</sup> Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA 02138, USA.</li> <li data-affiliation-id="full-view-affiliation-30" id="full-view-affiliation-30" ><sup class="key">30</sup> Department of Anthropology, Pennsylvania State University, University Park, PA 16802, USA.</li> <li data-affiliation-id="full-view-affiliation-31" id="full-view-affiliation-31" ><sup class="key">31</sup> Institute for Anthropological Research, Zagreb 10000, Croatia.</li> <li data-affiliation-id="full-view-affiliation-32" id="full-view-affiliation-32" ><sup class="key">32</sup> CSIR-Centre for Cellular and Molecular Biology, Hyderabad 500 007, India.</li> <li data-affiliation-id="full-view-affiliation-33" id="full-view-affiliation-33" ><sup class="key">33</sup> Birbal Sahni Institute of Palaeosciences, Lucknow 226007, India.</li> <li data-affiliation-id="full-view-affiliation-34" id="full-view-affiliation-34" ><sup class="key">34</sup> Department of Anthropology, Emory University, Atlanta, GA 30322, USA.</li> <li data-affiliation-id="full-view-affiliation-35" id="full-view-affiliation-35" ><sup class="key">35</sup> Tyumen Scientific Centre SB RAS, Institute of the Problems of Northern Development, Tyumen 625003, Russia.</li> <li data-affiliation-id="full-view-affiliation-36" id="full-view-affiliation-36" ><sup class="key">36</sup> CNRS-EXT500, Directeur de la Delegation Archaologique Francaise en Afghanistan (DAFA), Embassy of France in Kabul, Afghanistan.</li> <li data-affiliation-id="full-view-affiliation-37" id="full-view-affiliation-37" ><sup class="key">37</sup> Aktobe Regional Historical Museum, Aktobe 030006, Kazakhstan.</li> <li data-affiliation-id="full-view-affiliation-38" id="full-view-affiliation-38" ><sup class="key">38</sup> Archaeology of Asia Department, ISMEO - International Association of Mediterranean and Oriental Studies, Rome RM00186, Italy.</li> <li data-affiliation-id="full-view-affiliation-39" id="full-view-affiliation-39" ><sup class="key">39</sup> Kyrgyz National University, 720033 Bishkek, Kyrgyzstan.</li> <li data-affiliation-id="full-view-affiliation-40" id="full-view-affiliation-40" ><sup class="key">40</sup> Institute of Archaeology and Ethnography, Siberian Branch, Russian Academy of Sciences, Novosibirsk 630090, Russia.</li> <li data-affiliation-id="full-view-affiliation-41" id="full-view-affiliation-41" ><sup class="key">41</sup> Department of Political History, National and State-Confessional Relations, Altai State University, Barnaul 656049, Russia.</li> <li data-affiliation-id="full-view-affiliation-42" id="full-view-affiliation-42" ><sup class="key">42</sup> Institute of Archaeology, Czech Academy of Sciences, Prague 118 01, Czech Republic.</li> <li data-affiliation-id="full-view-affiliation-43" id="full-view-affiliation-43" ><sup class="key">43</sup> Department of Archaeology, Max Planck Institute for the Science of Human History, Jena 07745, Germany.</li> <li data-affiliation-id="full-view-affiliation-44" id="full-view-affiliation-44" ><sup class="key">44</sup> Oxford Radiocarbon Accelerator Unit, Research Laboratory for Archaeology and the History of Art, University of Oxford, Oxford OX1 3QY, UK.</li> <li data-affiliation-id="full-view-affiliation-45" id="full-view-affiliation-45" ><sup class="key">45</sup> Institute of History and Archaeology, Ural Branch RAS, Yekaterinburg 620990, Russia.</li> <li data-affiliation-id="full-view-affiliation-46" id="full-view-affiliation-46" ><sup class="key">46</sup> South Ural State University, Chelyabinsk 454080, Russia.</li> <li data-affiliation-id="full-view-affiliation-47" id="full-view-affiliation-47" ><sup class="key">47</sup> Department of Archaeology, Kemerovo State University, Kemerovo 650043, Russia.</li> <li data-affiliation-id="full-view-affiliation-48" id="full-view-affiliation-48" ><sup class="key">48</sup> Institute of Archaeology, University College London, London WC1H 0PY, UK.</li> <li data-affiliation-id="full-view-affiliation-49" id="full-view-affiliation-49" ><sup class="key">49</sup> School of Cultural Heritage, Northwest University, Shanxi, 710069, China.</li> <li data-affiliation-id="full-view-affiliation-50" id="full-view-affiliation-50" ><sup class="key">50</sup> Department of Archaeology, Ethnography and Museology, Altai State University, Barnaul 656049, Russia.</li> <li data-affiliation-id="full-view-affiliation-51" id="full-view-affiliation-51" ><sup class="key">51</sup> Department of Anthropology, University of Pittsburgh, Pittsburgh, PA 15260, USA.</li> <li data-affiliation-id="full-view-affiliation-52" id="full-view-affiliation-52" ><sup class="key">52</sup> Samara State University of Social Sciences and Education, Samara 443099, Russia.</li> <li data-affiliation-id="full-view-affiliation-53" id="full-view-affiliation-53" ><sup class="key">53</sup> West Kazakhstan Regional Center for History and Archaeology, Uralsk 090000, Kazakhstan.</li> <li data-affiliation-id="full-view-affiliation-54" id="full-view-affiliation-54" ><sup class="key">54</sup> Scientific and Educational Center of Study on the Problem of Nature and Man, Chelyabinsk State University, Chelyabinsk 454021, Russia.</li> <li data-affiliation-id="full-view-affiliation-55" id="full-view-affiliation-55" ><sup class="key">55</sup> Department of Cultural Heritage, University of Bologna, 48121 Ravenna, Italy.</li> <li data-affiliation-id="full-view-affiliation-56" id="full-view-affiliation-56" ><sup class="key">56</sup> Institute for Archaeological Research, Uzbekistan Academy of Sciences, Samarkand 140151, Uzbekistan.</li> <li data-affiliation-id="full-view-affiliation-57" id="full-view-affiliation-57" ><sup class="key">57</sup> Center for Research, Restoration and Protection of Historical and Cultural Heritage of Aktobe Region, Aktobe 030007, Kazakhstan.</li> <li data-affiliation-id="full-view-affiliation-58" id="full-view-affiliation-58" ><sup class="key">58</sup> Department of Anthropology, University of Winnipeg, Winnipeg, MB R3B 2E9, Canada.</li> <li data-affiliation-id="full-view-affiliation-59" id="full-view-affiliation-59" ><sup class="key">59</sup> Department of Archaeology, Simon Fraser University, Burnaby, BC V5A 1S6, Canada.</li> <li data-affiliation-id="full-view-affiliation-60" id="full-view-affiliation-60" ><sup class="key">60</sup> MiBAC - Ministero per i Beni e le Attività Culturali - Soprintendenza Archeologia, belle arti e paesaggio del Friuli Venezia Giulia, 34135 Trieste, Italy.</li> <li data-affiliation-id="full-view-affiliation-61" id="full-view-affiliation-61" ><sup class="key">61</sup> Afrosiab Museum, Samarkand 140151, Uzbekistan.</li> <li data-affiliation-id="full-view-affiliation-62" id="full-view-affiliation-62" ><sup class="key">62</sup> Department of Biological, Geological and Environmental Sciences, Alma Mater Studiorum - University of Bologna, Bologna 40126, Italy.</li> <li data-affiliation-id="full-view-affiliation-63" id="full-view-affiliation-63" ><sup class="key">63</sup> Human Origins Program, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013, USA.</li> <li data-affiliation-id="full-view-affiliation-64" id="full-view-affiliation-64" ><sup class="key">64</sup> National Research Tomsk State University, Tomsk 634050, Russia.</li> <li data-affiliation-id="full-view-affiliation-65" id="full-view-affiliation-65" ><sup class="key">65</sup> F. Voino-Yasenetsky Krasnoyarsk State Medical University, Krasnoyarsk 660022, Russia.</li> <li data-affiliation-id="full-view-affiliation-66" id="full-view-affiliation-66" ><sup class="key">66</sup> Central State Museum Republic of Kazakhstan, Samal-1 Microdistrict, Almaty 050010, Kazakhstan.</li> <li data-affiliation-id="full-view-affiliation-67" id="full-view-affiliation-67" ><sup class="key">67</sup> CHRONO Centre for Climate, the Environment, and Chronology, Queen's University of Belfast, Belfast BT7 1NN, Northern Ireland, UK.</li> <li data-affiliation-id="full-view-affiliation-68" id="full-view-affiliation-68" ><sup class="key">68</sup> Kyrgyz-Turkish Manas University, Bishkek, Kyrgyzstan.</li> <li data-affiliation-id="full-view-affiliation-69" id="full-view-affiliation-69" ><sup class="key">69</sup> Department of Anthropology, Natural History Museum Vienna, 1010 Vienna, Austria.</li> <li data-affiliation-id="full-view-affiliation-70" id="full-view-affiliation-70" ><sup class="key">70</sup> Department of Archaeology, Ethnography and Museology, The Laboratory of Interdisciplinary Studies in Archaeology of Western Siberia and Altai, Altai State University, Barnaul 656049, Russia.</li> <li data-affiliation-id="full-view-affiliation-71" id="full-view-affiliation-71" ><sup class="key">71</sup> Institute of Steppe, Ural Branch RAS, Orenburg 460000, Russia.</li> <li data-affiliation-id="full-view-affiliation-72" id="full-view-affiliation-72" ><sup class="key">72</sup> Center for Egyptological Studies RAS, Moscow 119991, Russia.</li> <li data-affiliation-id="full-view-affiliation-73" id="full-view-affiliation-73" ><sup class="key">73</sup> Department of Anthropology, National Museum, Prague 115 79, Czech Republic.</li> <li data-affiliation-id="full-view-affiliation-74" id="full-view-affiliation-74" ><sup class="key">74</sup> Archaeological Expertise LLP, Almaty 050060, Kazakhstan.</li> <li data-affiliation-id="full-view-affiliation-75" id="full-view-affiliation-75" ><sup class="key">75</sup> Department of Archaeology, Hazara University, Mansehra 21300, Pakistan.</li> <li data-affiliation-id="full-view-affiliation-76" id="full-view-affiliation-76" ><sup class="key">76</sup> N.F. Katanov Khakassia State University, Abakan 655017, Russia.</li> <li data-affiliation-id="full-view-affiliation-77" id="full-view-affiliation-77" ><sup class="key">77</sup> Department of Archaeology, Deccan College Post-Graduate and Research Institute, Pune 411006, India.</li> <li data-affiliation-id="full-view-affiliation-78" id="full-view-affiliation-78" ><sup class="key">78</sup> Institute of Evolutionary Biology, CSIC-Universitat Pompeu Fabra, Barcelona 08003, Spain.</li> <li data-affiliation-id="full-view-affiliation-79" id="full-view-affiliation-79" ><sup class="key">79</sup> Anthropology Department, Hartwick College, Oneonta, NY 13820, USA.</li> <li data-affiliation-id="full-view-affiliation-80" id="full-view-affiliation-80" ><sup class="key">80</sup> Department of Anthropology, University of California, Santa Barbara, CA 93106, USA.</li> <li data-affiliation-id="full-view-affiliation-81" id="full-view-affiliation-81" ><sup class="key">81</sup> Department of Anthropology, Washington University in St. Louis, St. Louis, MO 63112, USA. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu.</li> <li data-affiliation-id="full-view-affiliation-82" id="full-view-affiliation-82" ><sup class="key">82</sup> Spatial Analysis, Interpretation, and Exploration Laboratory, Washington University in St. Louis, St. Louis, MO 63112, USA.</li> <li data-affiliation-id="full-view-affiliation-83" id="full-view-affiliation-83" ><sup class="key">83</sup> Earth Institute, University College Dublin, Dublin 4, Ireland. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu.</li> <li data-affiliation-id="full-view-affiliation-84" id="full-view-affiliation-84" ><sup class="key">84</sup> Max Planck-Harvard Research Center for the Archaeoscience of the Ancient Mediterranean, Cambridge, MA 02138, USA.</li> </ul> </div> </div> </div> <ul class="identifiers" id="full-view-identifiers"> <li> <span class="identifier pubmed"> <span class="id-label"> PMID: </span> <strong class="current-id" title="PubMed ID">31488661</strong> </span> </li> <li> <span class="identifier pmc"> <span class="id-label"> PMCID: </span> <a class="id-link" target="_blank" rel="noopener" ref="linksrc=article_id_link&article_id=PMC6822619&id_type=PMC" href="https://pmc.ncbi.nlm.nih.gov/articles/PMC6822619/" data-ga-category="full_text" data-ga-action="PMCID" > PMC6822619 </a> </span> </li> <li> <span class="identifier doi"> <span class="id-label"> DOI: </span> <a class="id-link" target="_blank" rel="noopener" ref="linksrc=article_id_link&article_id=10.1126/science.aat7487&id_type=DOI" href="https://doi.org/10.1126/science.aat7487" data-ga-category="full_text" data-ga-action="DOI" > 10.1126/science.aat7487 </a> </span> </li> </ul> <div class="in-clipboard-label " hidden> Item in Clipboard </div> </div> <div class="short-view" id="short-view-heading"> <h1 class="heading-title"> The formation of human populations in South and Central Asia </h1> <div class="short-citation"> <span class="authors-list"> <span class="authors-list-item "><span class="full-name">Vagheesh M Narasimhan</span><span class="citation-separator"> et al.</span></span> </span> <span class="citation-journal"> Science<span class="citation-separator">.</span> </span> <span class="date"> <time class="citation-year" datetime="2019">2019</time><span class="citation-separator">.</span> </span> </div> <div class="more-article-details-section with-display-settings"> <button class="more-article-details" id="toggle-article-details" data-alt-text="Hide details" ref="linksrc=toggle_article_details" aria-controls="full-article-details" aria-expanded="false" data-fetch-url="/31488661/long-authors/" data-id-prefix=""> <span class="title">Show details</span> </button> <div class="display-options" data-format-key="format" data-default-format-value="abstract" 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href="https://www.ncbi.nlm.nih.gov/nlmcatalog?term=%22Science%22%5BTitle+Abbreviation%5D" data-ga-category="search_catalog" data-ga-action="journal_link" data-ga-label="Science"> Search in NLM Catalog </a></li><li><a class="add-to-search-link dropdown-block-link" ref ="linksrc=add_to_search_link" role="button" data-search-term='"Science"[jour]' data-ga-category="search" data-ga-action="add_to_search" data-ga-label='"Science"[jour]' href="#"> Add to Search </a></li></ul></div></div></div><span class="period">. </span><span class="cit">2019 Sep 6;365(6457):eaat7487.</span> </div> <span class="citation-doi"> doi: 10.1126/science.aat7487. </span> </div> <div class="expanded-authors" id="short-view-expanded-authors"> <div class="authors"> <h3 class="title"> Authors </h3> <div class="authors-list"> <span class="authors-list-item "><a class="full-name" href="/?term=Narasimhan+VM&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Vagheesh M Narasimhan">Vagheesh M Narasimhan</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu." href="#short-view-affiliation-1" ref="linksrc=author_aff"> 1 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Patterson+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Nick Patterson">Nick Patterson</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Broad Institute of Harvard and MIT, Cambridge, MA 02142, USA. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu." href="#short-view-affiliation-2" ref="linksrc=author_aff"> 2 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Radcliffe Institute for Advanced Study, Harvard University, Cambridge, MA 02138, USA." href="#short-view-affiliation-3" ref="linksrc=author_aff"> 3 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Moorjani+P&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Priya Moorjani">Priya Moorjani</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Molecular and Cell Biology, University of California, Berkeley, CA 94720, USA." href="#short-view-affiliation-4" ref="linksrc=author_aff"> 4 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Center for Computational Biology, University of California, Berkeley, CA 94720, USA." href="#short-view-affiliation-5" ref="linksrc=author_aff"> 5 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Rohland+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Nadin Rohland">Nadin Rohland</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Broad Institute of Harvard and MIT, Cambridge, MA 02142, USA." href="#short-view-affiliation-7" ref="linksrc=author_aff"> 7 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Bernardos+R&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Rebecca Bernardos">Rebecca Bernardos</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Mallick+S&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Swapan Mallick">Swapan Mallick</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Broad Institute of Harvard and MIT, Cambridge, MA 02142, USA." href="#short-view-affiliation-7" ref="linksrc=author_aff"> 7 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Lazaridis+I&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Iosif Lazaridis">Iosif Lazaridis</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Nakatsuka+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Nathan Nakatsuka">Nathan Nakatsuka</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Harvard-MIT Division of Health Sciences and Technology, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-9" ref="linksrc=author_aff"> 9 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Olalde+I&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Iñigo Olalde">Iñigo Olalde</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Lipson+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Mark Lipson">Mark Lipson</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Kim+AM&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Alexander M Kim">Alexander M Kim</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, Harvard University, Cambridge, MA 02138, USA." href="#short-view-affiliation-10" ref="linksrc=author_aff"> 10 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Olivieri+LM&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Luca M Olivieri">Luca M Olivieri</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="ISMEO - International Association of Mediterranean and Oriental Studies, Italian Archaeological Mission in Pakistan, 19200 Saidu Sharif (Swat), Pakistan." href="#short-view-affiliation-11" ref="linksrc=author_aff"> 11 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Coppa+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Alfredo Coppa">Alfredo Coppa</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Environmental Biology, Sapienza University, Rome 00185, Italy." href="#short-view-affiliation-12" ref="linksrc=author_aff"> 12 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Vidale+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Massimo Vidale">Massimo Vidale</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="ISMEO - International Association of Mediterranean and Oriental Studies, Italian Archaeological Mission in Pakistan, 19200 Saidu Sharif (Swat), Pakistan." href="#short-view-affiliation-11" ref="linksrc=author_aff"> 11 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Cultural Heritage: Archaeology and History of Art, Cinema and Music, University of Padua, Padua 35139, Italy." href="#short-view-affiliation-13" ref="linksrc=author_aff"> 13 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Mallory+J&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="James Mallory">James Mallory</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="School of Natural and Built Environment, Queen's University Belfast, Belfast BT7 1NN, Northern Ireland, UK." href="#short-view-affiliation-14" ref="linksrc=author_aff"> 14 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Moiseyev+V&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Vyacheslav Moiseyev">Vyacheslav Moiseyev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Peter the Great Museum of Anthropology and Ethnography (Kunstkamera), Russian Academy of Science, St. Petersburg 199034, Russia." href="#short-view-affiliation-15" ref="linksrc=author_aff"> 15 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Kitov+E&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Egor Kitov">Egor Kitov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Center of Physical Anthropology, Institute of Ethnology and Anthropology, Russian Academy of Sciences, Moscow 119991, Russia." href="#short-view-affiliation-16" ref="linksrc=author_aff"> 16 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan." href="#short-view-affiliation-17" ref="linksrc=author_aff"> 17 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Al-Farabi Kazakh National University, Almaty 050040, Kazakhstan." href="#short-view-affiliation-18" ref="linksrc=author_aff"> 18 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Monge+J&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Janet Monge">Janet Monge</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="University of Pennsylvania Museum of Archaeology and Anthropology, Philadelphia, PA 19104, USA." href="#short-view-affiliation-19" ref="linksrc=author_aff"> 19 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Adamski+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Nicole Adamski">Nicole Adamski</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Alex+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Neel Alex">Neel Alex</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Electrical Engineering and Computer Science, University of California, Berkeley, CA 94720, USA." href="#short-view-affiliation-20" ref="linksrc=author_aff"> 20 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Broomandkhoshbacht+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Nasreen Broomandkhoshbacht">Nasreen Broomandkhoshbacht</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Candilio+F&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Francesca Candilio">Francesca Candilio</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland." href="#short-view-affiliation-21" ref="linksrc=author_aff"> 21 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Soprintendenza Archeologia, Belle Arti e Paesaggio per la Città Metropolitana di Cagliari e le Province di Oristano e Sud Sardegna, Cagliari 09124, Italy." href="#short-view-affiliation-22" ref="linksrc=author_aff"> 22 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Callan+K&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Kimberly Callan">Kimberly Callan</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Cheronet+O&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Olivia Cheronet">Olivia Cheronet</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland." href="#short-view-affiliation-21" ref="linksrc=author_aff"> 21 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="School of Archaeology, University College Dublin, Dublin 4, Ireland." href="#short-view-affiliation-23" ref="linksrc=author_aff"> 23 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Evolutionary Anthropology, University of Vienna, 1090 Vienna, Austria." href="#short-view-affiliation-24" ref="linksrc=author_aff"> 24 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Culleton+BJ&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Brendan J Culleton">Brendan J Culleton</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institutes of Energy and the Environment, Pennsylvania State University, University Park, PA 16802, USA." href="#short-view-affiliation-25" ref="linksrc=author_aff"> 25 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Ferry+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Matthew Ferry">Matthew Ferry</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Fernandes+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Daniel Fernandes">Daniel Fernandes</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland." href="#short-view-affiliation-21" ref="linksrc=author_aff"> 21 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="School of Archaeology, University College Dublin, Dublin 4, Ireland." href="#short-view-affiliation-23" ref="linksrc=author_aff"> 23 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Evolutionary Anthropology, University of Vienna, 1090 Vienna, Austria." href="#short-view-affiliation-24" ref="linksrc=author_aff"> 24 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="CIAS, Department of Life Sciences, University of Coimbra, Coimbra 3000-456, Portugal." href="#short-view-affiliation-26" ref="linksrc=author_aff"> 26 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Freilich+S&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Suzanne Freilich">Suzanne Freilich</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Evolutionary Anthropology, University of Vienna, 1090 Vienna, Austria." href="#short-view-affiliation-24" ref="linksrc=author_aff"> 24 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Gamarra+B&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Beatriz Gamarra">Beatriz Gamarra</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland." href="#short-view-affiliation-21" ref="linksrc=author_aff"> 21 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="School of Archaeology, University College Dublin, Dublin 4, Ireland." href="#short-view-affiliation-23" ref="linksrc=author_aff"> 23 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Catalan Institute of Human Paleoecology and Social Evolution (IPHES), Tarragona 43007, Spain." href="#short-view-affiliation-27" ref="linksrc=author_aff"> 27 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Gaudio+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Daniel Gaudio">Daniel Gaudio</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland." href="#short-view-affiliation-21" ref="linksrc=author_aff"> 21 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="School of Archaeology, University College Dublin, Dublin 4, Ireland." href="#short-view-affiliation-23" ref="linksrc=author_aff"> 23 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Hajdinjak+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Mateja Hajdinjak">Mateja Hajdinjak</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Max Planck Institute for Evolutionary Anthropology, Leipzig 04103, Germany." href="#short-view-affiliation-28" ref="linksrc=author_aff"> 28 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Harney+%C3%89&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Éadaoin Harney">Éadaoin Harney</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-8" ref="linksrc=author_aff"> 8 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA 02138, USA." href="#short-view-affiliation-29" ref="linksrc=author_aff"> 29 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Harper+TK&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Thomas K Harper">Thomas K Harper</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, Pennsylvania State University, University Park, PA 16802, USA." href="#short-view-affiliation-30" ref="linksrc=author_aff"> 30 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Keating+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Denise Keating">Denise Keating</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland." href="#short-view-affiliation-21" ref="linksrc=author_aff"> 21 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Lawson+AM&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Ann Marie Lawson">Ann Marie Lawson</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Mah+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Matthew Mah">Matthew Mah</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Broad Institute of Harvard and MIT, Cambridge, MA 02142, USA." href="#short-view-affiliation-7" ref="linksrc=author_aff"> 7 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Mandl+K&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Kirsten Mandl">Kirsten Mandl</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Evolutionary Anthropology, University of Vienna, 1090 Vienna, Austria." href="#short-view-affiliation-24" ref="linksrc=author_aff"> 24 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Michel+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Megan Michel">Megan Michel</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Novak+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Mario Novak">Mario Novak</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland." href="#short-view-affiliation-21" ref="linksrc=author_aff"> 21 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute for Anthropological Research, Zagreb 10000, Croatia." href="#short-view-affiliation-31" ref="linksrc=author_aff"> 31 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Oppenheimer+J&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Jonas Oppenheimer">Jonas Oppenheimer</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Rai+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Niraj Rai">Niraj Rai</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="CSIR-Centre for Cellular and Molecular Biology, Hyderabad 500 007, India." href="#short-view-affiliation-32" ref="linksrc=author_aff"> 32 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Birbal Sahni Institute of Palaeosciences, Lucknow 226007, India." href="#short-view-affiliation-33" ref="linksrc=author_aff"> 33 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Sirak+K&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Kendra Sirak">Kendra Sirak</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland." href="#short-view-affiliation-21" ref="linksrc=author_aff"> 21 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, Emory University, Atlanta, GA 30322, USA." href="#short-view-affiliation-34" ref="linksrc=author_aff"> 34 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Slon+V&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Viviane Slon">Viviane Slon</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Max Planck Institute for Evolutionary Anthropology, Leipzig 04103, Germany." href="#short-view-affiliation-28" ref="linksrc=author_aff"> 28 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Stewardson+K&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Kristin Stewardson">Kristin Stewardson</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Zalzala+F&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Fatma Zalzala">Fatma Zalzala</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-8" ref="linksrc=author_aff"> 8 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Zhang+Z&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Zhao Zhang">Zhao Zhang</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-6" ref="linksrc=author_aff"> 6 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Akhatov+G&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Gaziz Akhatov">Gaziz Akhatov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan." href="#short-view-affiliation-17" ref="linksrc=author_aff"> 17 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Bagashev+AN&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Anatoly N Bagashev">Anatoly N Bagashev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Tyumen Scientific Centre SB RAS, Institute of the Problems of Northern Development, Tyumen 625003, Russia." href="#short-view-affiliation-35" ref="linksrc=author_aff"> 35 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Bagnera+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Alessandra Bagnera">Alessandra Bagnera</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="ISMEO - International Association of Mediterranean and Oriental Studies, Italian Archaeological Mission in Pakistan, 19200 Saidu Sharif (Swat), Pakistan." href="#short-view-affiliation-11" ref="linksrc=author_aff"> 11 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Baitanayev+B&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Bauryzhan Baitanayev">Bauryzhan Baitanayev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan." href="#short-view-affiliation-17" ref="linksrc=author_aff"> 17 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Bendezu-Sarmiento+J&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Julio Bendezu-Sarmiento">Julio Bendezu-Sarmiento</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="CNRS-EXT500, Directeur de la Delegation Archaologique Francaise en Afghanistan (DAFA), Embassy of France in Kabul, Afghanistan." href="#short-view-affiliation-36" ref="linksrc=author_aff"> 36 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Bissembaev+AA&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Arman A Bissembaev">Arman A Bissembaev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan." href="#short-view-affiliation-17" ref="linksrc=author_aff"> 17 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Aktobe Regional Historical Museum, Aktobe 030006, Kazakhstan." href="#short-view-affiliation-37" ref="linksrc=author_aff"> 37 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Bonora+GL&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Gian Luca Bonora">Gian Luca Bonora</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Archaeology of Asia Department, ISMEO - International Association of Mediterranean and Oriental Studies, Rome RM00186, Italy." href="#short-view-affiliation-38" ref="linksrc=author_aff"> 38 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Chargynov+TT&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Temirlan T Chargynov">Temirlan T Chargynov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Kyrgyz National University, 720033 Bishkek, Kyrgyzstan." href="#short-view-affiliation-39" ref="linksrc=author_aff"> 39 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Chikisheva+T&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Tatiana Chikisheva">Tatiana Chikisheva</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute of Archaeology and Ethnography, Siberian Branch, Russian Academy of Sciences, Novosibirsk 630090, Russia." href="#short-view-affiliation-40" ref="linksrc=author_aff"> 40 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Dashkovskiy+PK&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Petr K Dashkovskiy">Petr K Dashkovskiy</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Political History, National and State-Confessional Relations, Altai State University, Barnaul 656049, Russia." href="#short-view-affiliation-41" ref="linksrc=author_aff"> 41 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Derevianko+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Anatoly Derevianko">Anatoly Derevianko</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute of Archaeology and Ethnography, Siberian Branch, Russian Academy of Sciences, Novosibirsk 630090, Russia." href="#short-view-affiliation-40" ref="linksrc=author_aff"> 40 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Dobe%C5%A1+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Miroslav Dobeš">Miroslav Dobeš</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute of Archaeology, Czech Academy of Sciences, Prague 118 01, Czech Republic." href="#short-view-affiliation-42" ref="linksrc=author_aff"> 42 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Douka+K&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Katerina Douka">Katerina Douka</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Max Planck Institute for the Science of Human History, Jena 07745, Germany." href="#short-view-affiliation-43" ref="linksrc=author_aff"> 43 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Oxford Radiocarbon Accelerator Unit, Research Laboratory for Archaeology and the History of Art, University of Oxford, Oxford OX1 3QY, UK." href="#short-view-affiliation-44" ref="linksrc=author_aff"> 44 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Dubova+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Nadezhda Dubova">Nadezhda Dubova</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Center of Physical Anthropology, Institute of Ethnology and Anthropology, Russian Academy of Sciences, Moscow 119991, Russia." href="#short-view-affiliation-16" ref="linksrc=author_aff"> 16 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Duisengali+MN&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Meiram N Duisengali">Meiram N Duisengali</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Aktobe Regional Historical Museum, Aktobe 030006, Kazakhstan." href="#short-view-affiliation-37" ref="linksrc=author_aff"> 37 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Enshin+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Dmitry Enshin">Dmitry Enshin</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Tyumen Scientific Centre SB RAS, Institute of the Problems of Northern Development, Tyumen 625003, Russia." href="#short-view-affiliation-35" ref="linksrc=author_aff"> 35 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Epimakhov+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Andrey Epimakhov">Andrey Epimakhov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute of History and Archaeology, Ural Branch RAS, Yekaterinburg 620990, Russia." href="#short-view-affiliation-45" ref="linksrc=author_aff"> 45 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="South Ural State University, Chelyabinsk 454080, Russia." href="#short-view-affiliation-46" ref="linksrc=author_aff"> 46 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Fribus+AV&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Alexey V Fribus">Alexey V Fribus</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Kemerovo State University, Kemerovo 650043, Russia." href="#short-view-affiliation-47" ref="linksrc=author_aff"> 47 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Fuller+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Dorian Fuller">Dorian Fuller</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute of Archaeology, University College London, London WC1H 0PY, UK." href="#short-view-affiliation-48" ref="linksrc=author_aff"> 48 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="School of Cultural Heritage, Northwest University, Shanxi, 710069, China." href="#short-view-affiliation-49" ref="linksrc=author_aff"> 49 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Goryachev+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Alexander Goryachev">Alexander Goryachev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Tyumen Scientific Centre SB RAS, Institute of the Problems of Northern Development, Tyumen 625003, Russia." href="#short-view-affiliation-35" ref="linksrc=author_aff"> 35 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Gromov+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Andrey Gromov">Andrey Gromov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Peter the Great Museum of Anthropology and Ethnography (Kunstkamera), Russian Academy of Science, St. Petersburg 199034, Russia." href="#short-view-affiliation-15" ref="linksrc=author_aff"> 15 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Grushin+SP&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Sergey P Grushin">Sergey P Grushin</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Ethnography and Museology, Altai State University, Barnaul 656049, Russia." href="#short-view-affiliation-50" ref="linksrc=author_aff"> 50 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Hanks+B&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Bryan Hanks">Bryan Hanks</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, University of Pittsburgh, Pittsburgh, PA 15260, USA." href="#short-view-affiliation-51" ref="linksrc=author_aff"> 51 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Judd+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Margaret Judd">Margaret Judd</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, University of Pittsburgh, Pittsburgh, PA 15260, USA." href="#short-view-affiliation-51" ref="linksrc=author_aff"> 51 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Kazizov+E&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Erlan Kazizov">Erlan Kazizov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan." href="#short-view-affiliation-17" ref="linksrc=author_aff"> 17 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Khokhlov+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Aleksander Khokhlov">Aleksander Khokhlov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Samara State University of Social Sciences and Education, Samara 443099, Russia." href="#short-view-affiliation-52" ref="linksrc=author_aff"> 52 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Krygin+AP&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Aleksander P Krygin">Aleksander P Krygin</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="West Kazakhstan Regional Center for History and Archaeology, Uralsk 090000, Kazakhstan." href="#short-view-affiliation-53" ref="linksrc=author_aff"> 53 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Kupriyanova+E&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Elena Kupriyanova">Elena Kupriyanova</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Scientific and Educational Center of Study on the Problem of Nature and Man, Chelyabinsk State University, Chelyabinsk 454021, Russia." href="#short-view-affiliation-54" ref="linksrc=author_aff"> 54 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Kuznetsov+P&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Pavel Kuznetsov">Pavel Kuznetsov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Samara State University of Social Sciences and Education, Samara 443099, Russia." href="#short-view-affiliation-52" ref="linksrc=author_aff"> 52 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Luiselli+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Donata Luiselli">Donata Luiselli</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Cultural Heritage, University of Bologna, 48121 Ravenna, Italy." href="#short-view-affiliation-55" ref="linksrc=author_aff"> 55 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Maksudov+F&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Farhod Maksudov">Farhod Maksudov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute for Archaeological Research, Uzbekistan Academy of Sciences, Samarkand 140151, Uzbekistan." href="#short-view-affiliation-56" ref="linksrc=author_aff"> 56 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Mamedov+AM&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Aslan M Mamedov">Aslan M Mamedov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Center for Research, Restoration and Protection of Historical and Cultural Heritage of Aktobe Region, Aktobe 030007, Kazakhstan." href="#short-view-affiliation-57" ref="linksrc=author_aff"> 57 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Mamirov+TB&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Talgat B Mamirov">Talgat B Mamirov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan." href="#short-view-affiliation-17" ref="linksrc=author_aff"> 17 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Meiklejohn+C&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Christopher Meiklejohn">Christopher Meiklejohn</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, University of Winnipeg, Winnipeg, MB R3B 2E9, Canada." href="#short-view-affiliation-58" ref="linksrc=author_aff"> 58 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Merrett+DC&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Deborah C Merrett">Deborah C Merrett</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Simon Fraser University, Burnaby, BC V5A 1S6, Canada." href="#short-view-affiliation-59" ref="linksrc=author_aff"> 59 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Micheli+R&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Roberto Micheli">Roberto Micheli</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="ISMEO - International Association of Mediterranean and Oriental Studies, Italian Archaeological Mission in Pakistan, 19200 Saidu Sharif (Swat), Pakistan." href="#short-view-affiliation-11" ref="linksrc=author_aff"> 11 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="MiBAC - Ministero per i Beni e le Attività Culturali - Soprintendenza Archeologia, belle arti e paesaggio del Friuli Venezia Giulia, 34135 Trieste, Italy." href="#short-view-affiliation-60" ref="linksrc=author_aff"> 60 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Mochalov+O&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Oleg Mochalov">Oleg Mochalov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Samara State University of Social Sciences and Education, Samara 443099, Russia." href="#short-view-affiliation-52" ref="linksrc=author_aff"> 52 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Mustafokulov+S&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Samariddin Mustafokulov">Samariddin Mustafokulov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute for Archaeological Research, Uzbekistan Academy of Sciences, Samarkand 140151, Uzbekistan." href="#short-view-affiliation-56" ref="linksrc=author_aff"> 56 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Afrosiab Museum, Samarkand 140151, Uzbekistan." href="#short-view-affiliation-61" ref="linksrc=author_aff"> 61 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Nayak+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Ayushi Nayak">Ayushi Nayak</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Max Planck Institute for the Science of Human History, Jena 07745, Germany." href="#short-view-affiliation-43" ref="linksrc=author_aff"> 43 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Pettener+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Davide Pettener">Davide Pettener</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Biological, Geological and Environmental Sciences, Alma Mater Studiorum - University of Bologna, Bologna 40126, Italy." href="#short-view-affiliation-62" ref="linksrc=author_aff"> 62 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Potts+R&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Richard Potts">Richard Potts</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Human Origins Program, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013, USA." href="#short-view-affiliation-63" ref="linksrc=author_aff"> 63 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Razhev+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Dmitry Razhev">Dmitry Razhev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Tyumen Scientific Centre SB RAS, Institute of the Problems of Northern Development, Tyumen 625003, Russia." href="#short-view-affiliation-35" ref="linksrc=author_aff"> 35 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Rykun+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Marina Rykun">Marina Rykun</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="National Research Tomsk State University, Tomsk 634050, Russia." href="#short-view-affiliation-64" ref="linksrc=author_aff"> 64 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Sarno+S&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Stefania Sarno">Stefania Sarno</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Biological, Geological and Environmental Sciences, Alma Mater Studiorum - University of Bologna, Bologna 40126, Italy." href="#short-view-affiliation-62" ref="linksrc=author_aff"> 62 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Savenkova+TM&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Tatyana M Savenkova">Tatyana M Savenkova</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="F. Voino-Yasenetsky Krasnoyarsk State Medical University, Krasnoyarsk 660022, Russia." href="#short-view-affiliation-65" ref="linksrc=author_aff"> 65 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Sikhymbaeva+K&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Kulyan Sikhymbaeva">Kulyan Sikhymbaeva</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Central State Museum Republic of Kazakhstan, Samal-1 Microdistrict, Almaty 050010, Kazakhstan." href="#short-view-affiliation-66" ref="linksrc=author_aff"> 66 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Slepchenko+SM&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Sergey M Slepchenko">Sergey M Slepchenko</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Tyumen Scientific Centre SB RAS, Institute of the Problems of Northern Development, Tyumen 625003, Russia." href="#short-view-affiliation-35" ref="linksrc=author_aff"> 35 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Soltobaev+OA&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Oroz A Soltobaev">Oroz A Soltobaev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Kyrgyz National University, 720033 Bishkek, Kyrgyzstan." href="#short-view-affiliation-39" ref="linksrc=author_aff"> 39 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Stepanova+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Nadezhda Stepanova">Nadezhda Stepanova</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute of Archaeology and Ethnography, Siberian Branch, Russian Academy of Sciences, Novosibirsk 630090, Russia." href="#short-view-affiliation-40" ref="linksrc=author_aff"> 40 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Svyatko+S&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Svetlana Svyatko">Svetlana Svyatko</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Peter the Great Museum of Anthropology and Ethnography (Kunstkamera), Russian Academy of Science, St. Petersburg 199034, Russia." href="#short-view-affiliation-15" ref="linksrc=author_aff"> 15 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="CHRONO Centre for Climate, the Environment, and Chronology, Queen's University of Belfast, Belfast BT7 1NN, Northern Ireland, UK." href="#short-view-affiliation-67" ref="linksrc=author_aff"> 67 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Tabaldiev+K&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Kubatbek Tabaldiev">Kubatbek Tabaldiev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Kyrgyz-Turkish Manas University, Bishkek, Kyrgyzstan." href="#short-view-affiliation-68" ref="linksrc=author_aff"> 68 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Teschler-Nicola+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Maria Teschler-Nicola">Maria Teschler-Nicola</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Evolutionary Anthropology, University of Vienna, 1090 Vienna, Austria." href="#short-view-affiliation-24" ref="linksrc=author_aff"> 24 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, Natural History Museum Vienna, 1010 Vienna, Austria." href="#short-view-affiliation-69" ref="linksrc=author_aff"> 69 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Tishkin+AA&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Alexey A Tishkin">Alexey A Tishkin</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Ethnography and Museology, The Laboratory of Interdisciplinary Studies in Archaeology of Western Siberia and Altai, Altai State University, Barnaul 656049, Russia." href="#short-view-affiliation-70" ref="linksrc=author_aff"> 70 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Tkachev+VV&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Vitaly V Tkachev">Vitaly V Tkachev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute of Steppe, Ural Branch RAS, Orenburg 460000, Russia." href="#short-view-affiliation-71" ref="linksrc=author_aff"> 71 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Vasilyev+S&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Sergey Vasilyev">Sergey Vasilyev</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Center of Physical Anthropology, Institute of Ethnology and Anthropology, Russian Academy of Sciences, Moscow 119991, Russia." href="#short-view-affiliation-16" ref="linksrc=author_aff"> 16 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Center for Egyptological Studies RAS, Moscow 119991, Russia." href="#short-view-affiliation-72" ref="linksrc=author_aff"> 72 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Velem%C3%ADnsk%C3%BD+P&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Petr Velemínský">Petr Velemínský</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, National Museum, Prague 115 79, Czech Republic." href="#short-view-affiliation-73" ref="linksrc=author_aff"> 73 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Voyakin+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Dmitriy Voyakin">Dmitriy Voyakin</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan." href="#short-view-affiliation-17" ref="linksrc=author_aff"> 17 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Archaeological Expertise LLP, Almaty 050060, Kazakhstan." href="#short-view-affiliation-74" ref="linksrc=author_aff"> 74 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Yermolayeva+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Antonina Yermolayeva">Antonina Yermolayeva</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan." href="#short-view-affiliation-17" ref="linksrc=author_aff"> 17 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Zahir+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Muhammad Zahir">Muhammad Zahir</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Max Planck Institute for the Science of Human History, Jena 07745, Germany." href="#short-view-affiliation-43" ref="linksrc=author_aff"> 43 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Hazara University, Mansehra 21300, Pakistan." href="#short-view-affiliation-75" ref="linksrc=author_aff"> 75 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Zubkov+VS&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Valery S Zubkov">Valery S Zubkov</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="N.F. Katanov Khakassia State University, Abakan 655017, Russia." href="#short-view-affiliation-76" ref="linksrc=author_aff"> 76 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Zubova+A&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Alisa Zubova">Alisa Zubova</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Peter the Great Museum of Anthropology and Ethnography (Kunstkamera), Russian Academy of Science, St. Petersburg 199034, Russia." href="#short-view-affiliation-15" ref="linksrc=author_aff"> 15 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Shinde+VS&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Vasant S Shinde">Vasant S Shinde</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Deccan College Post-Graduate and Research Institute, Pune 411006, India." href="#short-view-affiliation-77" ref="linksrc=author_aff"> 77 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Lalueza-Fox+C&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Carles Lalueza-Fox">Carles Lalueza-Fox</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institute of Evolutionary Biology, CSIC-Universitat Pompeu Fabra, Barcelona 08003, Spain." href="#short-view-affiliation-78" ref="linksrc=author_aff"> 78 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Meyer+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Matthias Meyer">Matthias Meyer</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Max Planck Institute for Evolutionary Anthropology, Leipzig 04103, Germany." href="#short-view-affiliation-28" ref="linksrc=author_aff"> 28 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Anthony+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="David Anthony">David Anthony</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Anthropology Department, Hartwick College, Oneonta, NY 13820, USA." href="#short-view-affiliation-79" ref="linksrc=author_aff"> 79 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Boivin+N&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Nicole Boivin">Nicole Boivin</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Archaeology, Max Planck Institute for the Science of Human History, Jena 07745, Germany." href="#short-view-affiliation-43" ref="linksrc=author_aff"> 43 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Thangaraj+K&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Kumarasamy Thangaraj">Kumarasamy Thangaraj</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="CSIR-Centre for Cellular and Molecular Biology, Hyderabad 500 007, India." href="#short-view-affiliation-32" ref="linksrc=author_aff"> 32 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Kennett+DJ&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Douglas J Kennett">Douglas J Kennett</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Institutes of Energy and the Environment, Pennsylvania State University, University Park, PA 16802, USA." href="#short-view-affiliation-25" ref="linksrc=author_aff"> 25 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, Pennsylvania State University, University Park, PA 16802, USA." href="#short-view-affiliation-30" ref="linksrc=author_aff"> 30 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, University of California, Santa Barbara, CA 93106, USA." href="#short-view-affiliation-80" ref="linksrc=author_aff"> 80 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Frachetti+M&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Michael Frachetti">Michael Frachetti</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Anthropology, Washington University in St. Louis, St. Louis, MO 63112, USA. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu." href="#short-view-affiliation-81" ref="linksrc=author_aff"> 81 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Spatial Analysis, Interpretation, and Exploration Laboratory, Washington University in St. Louis, St. Louis, MO 63112, USA." href="#short-view-affiliation-82" ref="linksrc=author_aff"> 82 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Pinhasi+R&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="Ron Pinhasi">Ron Pinhasi</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Earth Institute, University College Dublin, Dublin 4, Ireland. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu." href="#short-view-affiliation-83" ref="linksrc=author_aff"> 83 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Evolutionary Anthropology, University of Vienna, 1090 Vienna, Austria." href="#short-view-affiliation-24" ref="linksrc=author_aff"> 24 </a></sup><span class="comma">, </span></span><span class="authors-list-item "><a class="full-name" href="/?term=Reich+D&cauthor_id=31488661" ref="linksrc=author_name_link" data-ga-category="search" data-ga-action="author_link" data-ga-label="David Reich">David Reich</a><sup class="affiliation-links"><span class="author-sup-separator"> </span><a class="affiliation-link" title="Department of Genetics, Harvard Medical School, Boston, MA 02115, USA. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu." href="#short-view-affiliation-1" ref="linksrc=author_aff"> 1 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Broad Institute of Harvard and MIT, Cambridge, MA 02142, USA." href="#short-view-affiliation-7" ref="linksrc=author_aff"> 7 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA." href="#short-view-affiliation-8" ref="linksrc=author_aff"> 8 </a><span class="author-sup-separator"> </span><a class="affiliation-link" title="Max Planck-Harvard Research Center for the Archaeoscience of the Ancient Mediterranean, Cambridge, MA 02138, USA." href="#short-view-affiliation-84" ref="linksrc=author_aff"> 84 </a></sup></span> </div> </div> <div class="affiliations"> <h3 class="title"> Affiliations </h3> <ul class="item-list"> <li data-affiliation-id="short-view-affiliation-1" id="short-view-affiliation-1" ><sup class="key">1</sup> Department of Genetics, Harvard Medical School, Boston, MA 02115, USA. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu.</li> <li data-affiliation-id="short-view-affiliation-2" id="short-view-affiliation-2" ><sup class="key">2</sup> Broad Institute of Harvard and MIT, Cambridge, MA 02142, USA. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu.</li> <li data-affiliation-id="short-view-affiliation-3" id="short-view-affiliation-3" ><sup class="key">3</sup> Radcliffe Institute for Advanced Study, Harvard University, Cambridge, MA 02138, USA.</li> <li data-affiliation-id="short-view-affiliation-4" id="short-view-affiliation-4" ><sup class="key">4</sup> Department of Molecular and Cell Biology, University of California, Berkeley, CA 94720, USA.</li> <li data-affiliation-id="short-view-affiliation-5" id="short-view-affiliation-5" ><sup class="key">5</sup> Center for Computational Biology, University of California, Berkeley, CA 94720, USA.</li> <li data-affiliation-id="short-view-affiliation-6" id="short-view-affiliation-6" ><sup class="key">6</sup> Department of Genetics, Harvard Medical School, Boston, MA 02115, USA.</li> <li data-affiliation-id="short-view-affiliation-7" id="short-view-affiliation-7" ><sup class="key">7</sup> Broad Institute of Harvard and MIT, Cambridge, MA 02142, USA.</li> <li data-affiliation-id="short-view-affiliation-8" id="short-view-affiliation-8" ><sup class="key">8</sup> Howard Hughes Medical Institute, Harvard Medical School, Boston, MA 02115, USA.</li> <li data-affiliation-id="short-view-affiliation-9" id="short-view-affiliation-9" ><sup class="key">9</sup> Harvard-MIT Division of Health Sciences and Technology, Harvard Medical School, Boston, MA 02115, USA.</li> <li data-affiliation-id="short-view-affiliation-10" id="short-view-affiliation-10" ><sup class="key">10</sup> Department of Anthropology, Harvard University, Cambridge, MA 02138, USA.</li> <li data-affiliation-id="short-view-affiliation-11" id="short-view-affiliation-11" ><sup class="key">11</sup> ISMEO - International Association of Mediterranean and Oriental Studies, Italian Archaeological Mission in Pakistan, 19200 Saidu Sharif (Swat), Pakistan.</li> <li data-affiliation-id="short-view-affiliation-12" id="short-view-affiliation-12" ><sup class="key">12</sup> Department of Environmental Biology, Sapienza University, Rome 00185, Italy.</li> <li data-affiliation-id="short-view-affiliation-13" id="short-view-affiliation-13" ><sup class="key">13</sup> Department of Cultural Heritage: Archaeology and History of Art, Cinema and Music, University of Padua, Padua 35139, Italy.</li> <li data-affiliation-id="short-view-affiliation-14" id="short-view-affiliation-14" ><sup class="key">14</sup> School of Natural and Built Environment, Queen's University Belfast, Belfast BT7 1NN, Northern Ireland, UK.</li> <li data-affiliation-id="short-view-affiliation-15" id="short-view-affiliation-15" ><sup class="key">15</sup> Peter the Great Museum of Anthropology and Ethnography (Kunstkamera), Russian Academy of Science, St. Petersburg 199034, Russia.</li> <li data-affiliation-id="short-view-affiliation-16" id="short-view-affiliation-16" ><sup class="key">16</sup> Center of Physical Anthropology, Institute of Ethnology and Anthropology, Russian Academy of Sciences, Moscow 119991, Russia.</li> <li data-affiliation-id="short-view-affiliation-17" id="short-view-affiliation-17" ><sup class="key">17</sup> A.Kh. Margulan Institute of Archaeology, Almaty 050010, Kazakhstan.</li> <li data-affiliation-id="short-view-affiliation-18" id="short-view-affiliation-18" ><sup class="key">18</sup> Al-Farabi Kazakh National University, Almaty 050040, Kazakhstan.</li> <li data-affiliation-id="short-view-affiliation-19" id="short-view-affiliation-19" ><sup class="key">19</sup> University of Pennsylvania Museum of Archaeology and Anthropology, Philadelphia, PA 19104, USA.</li> <li data-affiliation-id="short-view-affiliation-20" id="short-view-affiliation-20" ><sup class="key">20</sup> Department of Electrical Engineering and Computer Science, University of California, Berkeley, CA 94720, USA.</li> <li data-affiliation-id="short-view-affiliation-21" id="short-view-affiliation-21" ><sup class="key">21</sup> Earth Institute, University College Dublin, Dublin 4, Ireland.</li> <li data-affiliation-id="short-view-affiliation-22" id="short-view-affiliation-22" ><sup class="key">22</sup> Soprintendenza Archeologia, Belle Arti e Paesaggio per la Città Metropolitana di Cagliari e le Province di Oristano e Sud Sardegna, Cagliari 09124, Italy.</li> <li data-affiliation-id="short-view-affiliation-23" id="short-view-affiliation-23" ><sup class="key">23</sup> School of Archaeology, University College Dublin, Dublin 4, Ireland.</li> <li data-affiliation-id="short-view-affiliation-24" id="short-view-affiliation-24" ><sup class="key">24</sup> Department of Evolutionary Anthropology, University of Vienna, 1090 Vienna, Austria.</li> <li data-affiliation-id="short-view-affiliation-25" id="short-view-affiliation-25" ><sup class="key">25</sup> Institutes of Energy and the Environment, Pennsylvania State University, University Park, PA 16802, USA.</li> <li data-affiliation-id="short-view-affiliation-26" id="short-view-affiliation-26" ><sup class="key">26</sup> CIAS, Department of Life Sciences, University of Coimbra, Coimbra 3000-456, Portugal.</li> <li data-affiliation-id="short-view-affiliation-27" id="short-view-affiliation-27" ><sup class="key">27</sup> Catalan Institute of Human Paleoecology and Social Evolution (IPHES), Tarragona 43007, Spain.</li> <li data-affiliation-id="short-view-affiliation-28" id="short-view-affiliation-28" ><sup class="key">28</sup> Max Planck Institute for Evolutionary Anthropology, Leipzig 04103, Germany.</li> <li data-affiliation-id="short-view-affiliation-29" id="short-view-affiliation-29" ><sup class="key">29</sup> Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA 02138, USA.</li> <li data-affiliation-id="short-view-affiliation-30" id="short-view-affiliation-30" ><sup class="key">30</sup> Department of Anthropology, Pennsylvania State University, University Park, PA 16802, USA.</li> <li data-affiliation-id="short-view-affiliation-31" id="short-view-affiliation-31" ><sup class="key">31</sup> Institute for Anthropological Research, Zagreb 10000, Croatia.</li> <li data-affiliation-id="short-view-affiliation-32" id="short-view-affiliation-32" ><sup class="key">32</sup> CSIR-Centre for Cellular and Molecular Biology, Hyderabad 500 007, India.</li> <li data-affiliation-id="short-view-affiliation-33" id="short-view-affiliation-33" ><sup class="key">33</sup> Birbal Sahni Institute of Palaeosciences, Lucknow 226007, India.</li> <li data-affiliation-id="short-view-affiliation-34" id="short-view-affiliation-34" ><sup class="key">34</sup> Department of Anthropology, Emory University, Atlanta, GA 30322, USA.</li> <li data-affiliation-id="short-view-affiliation-35" id="short-view-affiliation-35" ><sup class="key">35</sup> Tyumen Scientific Centre SB RAS, Institute of the Problems of Northern Development, Tyumen 625003, Russia.</li> <li data-affiliation-id="short-view-affiliation-36" id="short-view-affiliation-36" ><sup class="key">36</sup> CNRS-EXT500, Directeur de la Delegation Archaologique Francaise en Afghanistan (DAFA), Embassy of France in Kabul, Afghanistan.</li> <li data-affiliation-id="short-view-affiliation-37" id="short-view-affiliation-37" ><sup class="key">37</sup> Aktobe Regional Historical Museum, Aktobe 030006, Kazakhstan.</li> <li data-affiliation-id="short-view-affiliation-38" id="short-view-affiliation-38" ><sup class="key">38</sup> Archaeology of Asia Department, ISMEO - International Association of Mediterranean and Oriental Studies, Rome RM00186, Italy.</li> <li data-affiliation-id="short-view-affiliation-39" id="short-view-affiliation-39" ><sup class="key">39</sup> Kyrgyz National University, 720033 Bishkek, Kyrgyzstan.</li> <li data-affiliation-id="short-view-affiliation-40" id="short-view-affiliation-40" ><sup class="key">40</sup> Institute of Archaeology and Ethnography, Siberian Branch, Russian Academy of Sciences, Novosibirsk 630090, Russia.</li> <li data-affiliation-id="short-view-affiliation-41" id="short-view-affiliation-41" ><sup class="key">41</sup> Department of Political History, National and State-Confessional Relations, Altai State University, Barnaul 656049, Russia.</li> <li data-affiliation-id="short-view-affiliation-42" id="short-view-affiliation-42" ><sup class="key">42</sup> Institute of Archaeology, Czech Academy of Sciences, Prague 118 01, Czech Republic.</li> <li data-affiliation-id="short-view-affiliation-43" id="short-view-affiliation-43" ><sup class="key">43</sup> Department of Archaeology, Max Planck Institute for the Science of Human History, Jena 07745, Germany.</li> <li data-affiliation-id="short-view-affiliation-44" id="short-view-affiliation-44" ><sup class="key">44</sup> Oxford Radiocarbon Accelerator Unit, Research Laboratory for Archaeology and the History of Art, University of Oxford, Oxford OX1 3QY, UK.</li> <li data-affiliation-id="short-view-affiliation-45" id="short-view-affiliation-45" ><sup class="key">45</sup> Institute of History and Archaeology, Ural Branch RAS, Yekaterinburg 620990, Russia.</li> <li data-affiliation-id="short-view-affiliation-46" id="short-view-affiliation-46" ><sup class="key">46</sup> South Ural State University, Chelyabinsk 454080, Russia.</li> <li data-affiliation-id="short-view-affiliation-47" id="short-view-affiliation-47" ><sup class="key">47</sup> Department of Archaeology, Kemerovo State University, Kemerovo 650043, Russia.</li> <li data-affiliation-id="short-view-affiliation-48" id="short-view-affiliation-48" ><sup class="key">48</sup> Institute of Archaeology, University College London, London WC1H 0PY, UK.</li> <li data-affiliation-id="short-view-affiliation-49" id="short-view-affiliation-49" ><sup class="key">49</sup> School of Cultural Heritage, Northwest University, Shanxi, 710069, China.</li> <li data-affiliation-id="short-view-affiliation-50" id="short-view-affiliation-50" ><sup class="key">50</sup> Department of Archaeology, Ethnography and Museology, Altai State University, Barnaul 656049, Russia.</li> <li data-affiliation-id="short-view-affiliation-51" id="short-view-affiliation-51" ><sup class="key">51</sup> Department of Anthropology, University of Pittsburgh, Pittsburgh, PA 15260, USA.</li> <li data-affiliation-id="short-view-affiliation-52" id="short-view-affiliation-52" ><sup class="key">52</sup> Samara State University of Social Sciences and Education, Samara 443099, Russia.</li> <li data-affiliation-id="short-view-affiliation-53" id="short-view-affiliation-53" ><sup class="key">53</sup> West Kazakhstan Regional Center for History and Archaeology, Uralsk 090000, Kazakhstan.</li> <li data-affiliation-id="short-view-affiliation-54" id="short-view-affiliation-54" ><sup class="key">54</sup> Scientific and Educational Center of Study on the Problem of Nature and Man, Chelyabinsk State University, Chelyabinsk 454021, Russia.</li> <li data-affiliation-id="short-view-affiliation-55" id="short-view-affiliation-55" ><sup class="key">55</sup> Department of Cultural Heritage, University of Bologna, 48121 Ravenna, Italy.</li> <li data-affiliation-id="short-view-affiliation-56" id="short-view-affiliation-56" ><sup class="key">56</sup> Institute for Archaeological Research, Uzbekistan Academy of Sciences, Samarkand 140151, Uzbekistan.</li> <li data-affiliation-id="short-view-affiliation-57" id="short-view-affiliation-57" ><sup class="key">57</sup> Center for Research, Restoration and Protection of Historical and Cultural Heritage of Aktobe Region, Aktobe 030007, Kazakhstan.</li> <li data-affiliation-id="short-view-affiliation-58" id="short-view-affiliation-58" ><sup class="key">58</sup> Department of Anthropology, University of Winnipeg, Winnipeg, MB R3B 2E9, Canada.</li> <li data-affiliation-id="short-view-affiliation-59" id="short-view-affiliation-59" ><sup class="key">59</sup> Department of Archaeology, Simon Fraser University, Burnaby, BC V5A 1S6, Canada.</li> <li data-affiliation-id="short-view-affiliation-60" id="short-view-affiliation-60" ><sup class="key">60</sup> MiBAC - Ministero per i Beni e le Attività Culturali - Soprintendenza Archeologia, belle arti e paesaggio del Friuli Venezia Giulia, 34135 Trieste, Italy.</li> <li data-affiliation-id="short-view-affiliation-61" id="short-view-affiliation-61" ><sup class="key">61</sup> Afrosiab Museum, Samarkand 140151, Uzbekistan.</li> <li data-affiliation-id="short-view-affiliation-62" id="short-view-affiliation-62" ><sup class="key">62</sup> Department of Biological, Geological and Environmental Sciences, Alma Mater Studiorum - University of Bologna, Bologna 40126, Italy.</li> <li data-affiliation-id="short-view-affiliation-63" id="short-view-affiliation-63" ><sup class="key">63</sup> Human Origins Program, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013, USA.</li> <li data-affiliation-id="short-view-affiliation-64" id="short-view-affiliation-64" ><sup class="key">64</sup> National Research Tomsk State University, Tomsk 634050, Russia.</li> <li data-affiliation-id="short-view-affiliation-65" id="short-view-affiliation-65" ><sup class="key">65</sup> F. Voino-Yasenetsky Krasnoyarsk State Medical University, Krasnoyarsk 660022, Russia.</li> <li data-affiliation-id="short-view-affiliation-66" id="short-view-affiliation-66" ><sup class="key">66</sup> Central State Museum Republic of Kazakhstan, Samal-1 Microdistrict, Almaty 050010, Kazakhstan.</li> <li data-affiliation-id="short-view-affiliation-67" id="short-view-affiliation-67" ><sup class="key">67</sup> CHRONO Centre for Climate, the Environment, and Chronology, Queen's University of Belfast, Belfast BT7 1NN, Northern Ireland, UK.</li> <li data-affiliation-id="short-view-affiliation-68" id="short-view-affiliation-68" ><sup class="key">68</sup> Kyrgyz-Turkish Manas University, Bishkek, Kyrgyzstan.</li> <li data-affiliation-id="short-view-affiliation-69" id="short-view-affiliation-69" ><sup class="key">69</sup> Department of Anthropology, Natural History Museum Vienna, 1010 Vienna, Austria.</li> <li data-affiliation-id="short-view-affiliation-70" id="short-view-affiliation-70" ><sup class="key">70</sup> Department of Archaeology, Ethnography and Museology, The Laboratory of Interdisciplinary Studies in Archaeology of Western Siberia and Altai, Altai State University, Barnaul 656049, Russia.</li> <li data-affiliation-id="short-view-affiliation-71" id="short-view-affiliation-71" ><sup class="key">71</sup> Institute of Steppe, Ural Branch RAS, Orenburg 460000, Russia.</li> <li data-affiliation-id="short-view-affiliation-72" id="short-view-affiliation-72" ><sup class="key">72</sup> Center for Egyptological Studies RAS, Moscow 119991, Russia.</li> <li data-affiliation-id="short-view-affiliation-73" id="short-view-affiliation-73" ><sup class="key">73</sup> Department of Anthropology, National Museum, Prague 115 79, Czech Republic.</li> <li data-affiliation-id="short-view-affiliation-74" id="short-view-affiliation-74" ><sup class="key">74</sup> Archaeological Expertise LLP, Almaty 050060, Kazakhstan.</li> <li data-affiliation-id="short-view-affiliation-75" id="short-view-affiliation-75" ><sup class="key">75</sup> Department of Archaeology, Hazara University, Mansehra 21300, Pakistan.</li> <li data-affiliation-id="short-view-affiliation-76" id="short-view-affiliation-76" ><sup class="key">76</sup> N.F. Katanov Khakassia State University, Abakan 655017, Russia.</li> <li data-affiliation-id="short-view-affiliation-77" id="short-view-affiliation-77" ><sup class="key">77</sup> Department of Archaeology, Deccan College Post-Graduate and Research Institute, Pune 411006, India.</li> <li data-affiliation-id="short-view-affiliation-78" id="short-view-affiliation-78" ><sup class="key">78</sup> Institute of Evolutionary Biology, CSIC-Universitat Pompeu Fabra, Barcelona 08003, Spain.</li> <li data-affiliation-id="short-view-affiliation-79" id="short-view-affiliation-79" ><sup class="key">79</sup> Anthropology Department, Hartwick College, Oneonta, NY 13820, USA.</li> <li data-affiliation-id="short-view-affiliation-80" id="short-view-affiliation-80" ><sup class="key">80</sup> Department of Anthropology, University of California, Santa Barbara, CA 93106, USA.</li> <li data-affiliation-id="short-view-affiliation-81" id="short-view-affiliation-81" ><sup class="key">81</sup> Department of Anthropology, Washington University in St. Louis, St. Louis, MO 63112, USA. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu.</li> <li data-affiliation-id="short-view-affiliation-82" id="short-view-affiliation-82" ><sup class="key">82</sup> Spatial Analysis, Interpretation, and Exploration Laboratory, Washington University in St. Louis, St. Louis, MO 63112, USA.</li> <li data-affiliation-id="short-view-affiliation-83" id="short-view-affiliation-83" ><sup class="key">83</sup> Earth Institute, University College Dublin, Dublin 4, Ireland. vagheesh@mail.harvard.edu nickp@broadinstitute.org frachetti@wustl.edu ron.pinhasi@univie.ac.at reich@genetics.med.harvard.edu.</li> <li data-affiliation-id="short-view-affiliation-84" id="short-view-affiliation-84" ><sup class="key">84</sup> Max Planck-Harvard Research Center for the Archaeoscience of the Ancient Mediterranean, Cambridge, MA 02138, USA.</li> </ul> </div> </div> <ul class="identifiers" id="short-view-identifiers"> <li> <span class="identifier pubmed"> <span class="id-label"> PMID: </span> <strong class="current-id" title="PubMed ID">31488661</strong> </span> </li> <li> <span class="identifier pmc"> <span class="id-label"> PMCID: </span> <a class="id-link" target="_blank" rel="noopener" ref="linksrc=article_id_link&article_id=PMC6822619&id_type=PMC" href="https://pmc.ncbi.nlm.nih.gov/articles/PMC6822619/" data-ga-category="full_text" data-ga-action="PMCID" > PMC6822619 </a> </span> </li> <li> <span class="identifier doi"> <span class="id-label"> DOI: </span> <a class="id-link" target="_blank" rel="noopener" ref="linksrc=article_id_link&article_id=10.1126/science.aat7487&id_type=DOI" href="https://doi.org/10.1126/science.aat7487" data-ga-category="full_text" data-ga-action="DOI" > 10.1126/science.aat7487 </a> </span> </li> </ul> <div class="in-clipboard-label " hidden> Item in Clipboard </div> </div> </div> </header> <div class="actions-buttons inline"><div class="inner-wrap"><button class="full-text-links-button full-text-links-dialog-trigger" title="Open dialog with full text links" ref="linksrc=full_text_links"> Full text links </button><button class="citation-button citation-dialog-trigger" aria-label="Open dialog with citation text in different styles" data-ga-category="save_share" data-ga-action="cite" data-ga-label="open" data-all-citations-url="/31488661/citations/" data-citation-style="nlm" data-pubmed-format-link="/31488661/export/"><span class="button-label">Cite</span></button><button class="more-actions-button more-actions-dialog-trigger" title="Open dialog with more actions to take" ref="linksrc=more_actions_btn"></button><div class="display-options" data-format-key="format" data-default-format-value="abstract" data-current-format-value="abstract" role="region"><button class="trigger usa-button" title="Change format" data-ga-category="display_options" data-ga-action="display_options_dialog"><span class="button-label">Display options</span></button><div class="dropdown" hidden><div class="title"> Display options </div><div class="content"><div class="format-container"><span class="option-label"> Format </span><select name="format" data-initial-value="abstract" aria-label="Display format" id="inline-actions-display-format"><option value="abstract">Abstract</option><option value="pubmed">PubMed</option><option value="pmid">PMID</option></select></div></div></div></div></div></div> <div class="abstract" id="abstract"> <h2 class="title"> Abstract </h2> <div class="abstract-content selected" id="eng-abstract"> <p> By sequencing 523 ancient humans, we show that the primary source of ancestry in modern South Asians is a prehistoric genetic gradient between people related to early hunter-gatherers of Iran and Southeast Asia. After the Indus Valley Civilization's decline, its people mixed with individuals in the southeast to form one of the two main ancestral populations of South Asia, whose direct descendants live in southern India. Simultaneously, they mixed with descendants of Steppe pastoralists who, starting around 4000 years ago, spread via Central Asia to form the other main ancestral population. The Steppe ancestry in South Asia has the same profile as that in Bronze Age Eastern Europe, tracking a movement of people that affected both regions and that likely spread the distinctive features shared between Indo-Iranian and Balto-Slavic languages. </p> </div> </div> <p class="copyright" id="copyright"> Copyright © 2019 The Authors, some rights reserved; exclusive licensee American Association for the Advancement of Science. No claim to original U.S. Government Works. </p> <p class="disclaimer" id="disclaimer"> <a href="/disclaimer/" target="_blank" ga_category="literature_resources" ga_action="disclaimer_link" ga_label="under_abstract">PubMed Disclaimer</a> </p> <div class="conflict-of-interest" id="conflict-of-interest"> <h2 class="title"> Conflict of interest statement </h2> <div class="statement"> <p xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:p1="http://pubmed.gov/pub-one"><bold>Competing interests:</bold> The authors declare no competing interests.</p> </div> </div> <div class="figures" id="figures"> <h2 class="title"> Figures </h2> <div class="figures-list" id="slides-container" itemscope itemtype="http://schema.org/ImageGallery"> <figure class="figure-item " itemscope itemtype="http://schema.org/ImageObject" itemprop="associatedMedia" data-slide-index="0" data-label-slug="fig-1"> <a class="figure-link" href="https://cdn.ncbi.nlm.nih.gov/pmc/blobs/d74d/6822619/b308eecc98f0/nihms-1053677-f0002.jpg" itemprop="contentUrl" aria-describedby="figure-caption-0" role="button" data-image-width="800" data-image-height="584" data-image-alt="Fig. 1" data-pmc-id="PMC6822619" data-figure-id="F1"> <img class="figure-thumb" itemprop="thumbnail" id="article-image-0" src="https://cdn.ncbi.nlm.nih.gov/pmc/blobs/d74d/6822619/011aa57317c4/nihms-1053677-f0002.gif" alt="Fig. 1" /> </a> <meta itemprop="width" itemtype="http://schema.org/ImageObject" content="800"> <meta itemprop="height" itemtype="http://schema.org/ImageObject" content="584"> <div class="figure-caption figure-caption-text" itemtype="http://schema.org/ImageObject" itemprop="caption" aria-hidden="true"> <div class="caption-wrap"> <strong class="figure-label"> <p> Fig. 1. Overview of ancient DNA data. </p> </strong> </div> </div> <div class="figure-caption-medium figure-caption-text" aria-hidden="true"> <strong class="figure-label"> <p> Fig. 1. Overview of ancient DNA data. </p> </strong> <div class="figure-caption-contents"><p> ( <b> A </b> ) Distribution of sites and associated… </p></div> </div> <figcaption id="figure-caption-0" class="figure-caption-full figure-caption-text" itemtype="http://schema.org/ImageObject" itemprop="description"> <strong class="figure-label"> Fig. 1. Overview of ancient DNA data. </strong> <div class="figure-caption-contents">(<b>A</b>) Distribution of sites and associated archeological or radiocarbon dates along with the number of individuals meeting our analysis thresholds from each site. (<b>B</b>) Locations of ancient individuals for whom we generated ancient DNA that passed our analysis thresholds along with the locations of individuals from 140 groups from present-day South Asia that we analyzed as forming the <i>Modern Indian Cline</i>. Shapes distinguish the individuals from different sites. Data from 106 South Asian groups that do not fit along the <i>Modern Indian Cline</i> as well as <i>AHG</i> are not shown. <b>(C)</b> PCA analysis of ancient and modern individuals projected onto a basis formed by 1,340 present day Eurasians reflects clustering of individuals that mirrors their geographical relationships. An interactive version of this figure is presented in the Online Data Visualizer.</div> </figcaption> </figure> <figure class="figure-item " itemscope itemtype="http://schema.org/ImageObject" itemprop="associatedMedia" data-slide-index="1" data-label-slug="fig-2"> <a class="figure-link" href="https://cdn.ncbi.nlm.nih.gov/pmc/blobs/d74d/6822619/c705f8c5411f/nihms-1053677-f0003.jpg" itemprop="contentUrl" aria-describedby="figure-caption-1" role="button" data-image-width="750" data-image-height="733" data-image-alt="Fig. 2." data-pmc-id="PMC6822619" data-figure-id="F2"> <img class="figure-thumb" itemprop="thumbnail" id="article-image-1" src="https://cdn.ncbi.nlm.nih.gov/pmc/blobs/d74d/6822619/506e79d437e8/nihms-1053677-f0003.gif" alt="Fig. 2." /> </a> <meta itemprop="width" itemtype="http://schema.org/ImageObject" content="750"> <meta itemprop="height" itemtype="http://schema.org/ImageObject" content="733"> <div class="figure-caption figure-caption-text" itemtype="http://schema.org/ImageObject" itemprop="caption" aria-hidden="true"> <div class="caption-wrap"> <strong class="figure-label"> <p> Fig. 2.. Outlier analysis reveals ancient contacts… </p> </strong> </div> </div> <div class="figure-caption-medium figure-caption-text" aria-hidden="true"> <strong class="figure-label"> <p> Fig. 2.. Outlier analysis reveals ancient contacts between sites. </p> </strong> <div class="figure-caption-contents"><p> We plot the average of Principal… </p></div> </div> <figcaption id="figure-caption-1" class="figure-caption-full figure-caption-text" itemtype="http://schema.org/ImageObject" itemprop="description"> <strong class="figure-label"> Fig. 2.. Outlier analysis reveals ancient contacts between sites. </strong> <div class="figure-caption-contents">We plot the average of Principal Component 1 (x-axis) and Principal Component 2 (y-axis) for the West Eurasian and All Eurasian PCA plots, as we found that this aids visual separation of the ancestry profiles. (<b>A</b>) In the Middle to Late Bronze Age Steppe, we observe in addition to the <i>Western_Steppe_MLBA</i> and <i>Central_Steppe_MLBA</i> clusters (indistinguishable in this projection), outliers admixed with other ancestries. The <i>BMAC</i>-related admixture in Kazakhstan documents northward gene flow onto the Steppe and confirms the Inner Asian Mountain Corridor as a conduit for movement of people. <b>(B)</b> At Shahr-i-Sokhta in eastern Iran, there are two primary groupings: one with ~20% Anatolian farmer-related ancestry and no detectable <i>AHG-</i>related ancestry, and the other with ~0% Anatolian farmer-related ancestry and substantial <i>AHG-</i>related ancestry (<i>Indus Periphery Cline</i>). (<b>C</b>) In individuals of the BMAC and successor sites, we observe a main cluster as well as numerous outliers: outliers >2000 BCE with admixture related to <i>WSHG,</i> outliers >2000 BCE on the <i>Indus Periphery Cline</i> (with an ancestral similar similar to the outliers at Shahr-i-Sokhta), and outliers after 2000 BCE that reveal how <i>Central_Steppe_MLBA</i> ancestry had arrived. <b>(D)</b> In the Late Bronze Age and Iron Age of northernmost South Asia, we observe a main cluster consistent with admixture between peoples of the <i>Indus Periphery Cline</i> and <i>Central_Steppe_MLBA</i>, and variable Steppe pastoralist-related admixture.</div> </figcaption> </figure> <figure class="figure-item " itemscope itemtype="http://schema.org/ImageObject" itemprop="associatedMedia" data-slide-index="2" data-label-slug="fig-3"> <a class="figure-link" href="https://cdn.ncbi.nlm.nih.gov/pmc/blobs/d74d/6822619/9d5f0f39fcf7/nihms-1053677-f0004.jpg" itemprop="contentUrl" aria-describedby="figure-caption-2" role="button" data-image-width="800" data-image-height="882" data-image-alt="Fig. 3." data-pmc-id="PMC6822619" data-figure-id="F3"> <img class="figure-thumb" itemprop="thumbnail" id="article-image-2" src="https://cdn.ncbi.nlm.nih.gov/pmc/blobs/d74d/6822619/7845e3ad48a5/nihms-1053677-f0004.gif" alt="Fig. 3." /> </a> <meta itemprop="width" itemtype="http://schema.org/ImageObject" content="800"> <meta itemprop="height" itemtype="http://schema.org/ImageObject" content="882"> <div class="figure-caption figure-caption-text" itemtype="http://schema.org/ImageObject" itemprop="caption" aria-hidden="true"> <div class="caption-wrap"> <strong class="figure-label"> <p> Fig. 3.. Ancestry Transformations in Holocene Eurasia. </p> </strong> </div> </div> <div class="figure-caption-medium figure-caption-text" aria-hidden="true"> <strong class="figure-label"> <p> Fig. 3.. Ancestry Transformations in Holocene Eurasia. </p> </strong> <div class="figure-caption-contents"><b> (A) </b> Ancestry clines before and after the advent…</div> </div> <figcaption id="figure-caption-2" class="figure-caption-full figure-caption-text" itemtype="http://schema.org/ImageObject" itemprop="description"> <strong class="figure-label"> Fig. 3.. Ancestry Transformations in Holocene Eurasia. </strong> <div class="figure-caption-contents"><b>(A)</b> Ancestry clines before and after the advent of farming. We document a <i>South Eurasian Early Holocene Cline</i> of increasing Iranian farmer- and West Siberian hunter-gatherer related ancestry moving west-to-east from Anatolia to Iran, and a <i>North Eurasian Early Holocene Cline</i> of increasing relatedness to East Asians moving west-to-east from Europe to Siberia. Mixtures of peoples along these two clines following the spread of farming formed five later gradients (shaded): moving west-to-east: the <i>European Cline,</i> the <i>Caucasus Cline</i> from which the Yamnaya formed, the <i>Central Asian Cline</i> which characterized much of Central Asia in the Copper and Bronze Ages, the <i>Southwest Asian Cline</i> established by spreads of farmers in multiple directions from several loci of domestication, and the <i>Indus Periphery Cline</i>. <b>(B)</b> Following the appearance of the Yamnaya Steppe pastoralists, <i>Western_Steppe_EMBA</i> (<i>Yamnaya</i>-like) ancestry then spread across this vast region. We use arrows to show plausible directions of spread of increasingly diluted ancestry (the arrows are not meant as exact routes which we do not have enough sampling to determine at present). Rough estimates of the timing of the arrival of this ancestry and estimated ancestry proportions are shown.</div> </figcaption> </figure> <figure class="figure-item " itemscope itemtype="http://schema.org/ImageObject" itemprop="associatedMedia" data-slide-index="3" data-label-slug="fig-4"> <a class="figure-link" href="https://cdn.ncbi.nlm.nih.gov/pmc/blobs/d74d/6822619/ff17677b794d/nihms-1053677-f0005.jpg" itemprop="contentUrl" aria-describedby="figure-caption-3" role="button" data-image-width="800" data-image-height="731" data-image-alt="Fig. 4." data-pmc-id="PMC6822619" data-figure-id="F4"> <img class="figure-thumb" itemprop="thumbnail" id="article-image-3" src="https://cdn.ncbi.nlm.nih.gov/pmc/blobs/d74d/6822619/6f016162a42f/nihms-1053677-f0005.gif" alt="Fig. 4." /> </a> <meta itemprop="width" itemtype="http://schema.org/ImageObject" content="800"> <meta itemprop="height" itemtype="http://schema.org/ImageObject" content="731"> <div class="figure-caption figure-caption-text" itemtype="http://schema.org/ImageObject" itemprop="caption" aria-hidden="true"> <div class="caption-wrap"> <strong class="figure-label"> <p> Fig. 4.. The Genomic Formation of South… </p> </strong> </div> </div> <div class="figure-caption-medium figure-caption-text" aria-hidden="true"> <strong class="figure-label"> <p> Fig. 4.. The Genomic Formation of South Asia. </p> </strong> <div class="figure-caption-contents"><b> (A) </b> The degree of allele sharing with…</div> </div> <figcaption id="figure-caption-3" class="figure-caption-full figure-caption-text" itemtype="http://schema.org/ImageObject" itemprop="description"> <strong class="figure-label"> Fig. 4.. The Genomic Formation of South Asia. </strong> <div class="figure-caption-contents"><b>(A)</b> The degree of allele sharing with southern Asian hunter-gatherers (<i>AASI</i>) measured by <i>f</i><sub><i>4</i></sub><i>(Ethiopia_4500BP, X; Ganj_Dareh_N</i>, <i>AHG</i>) and with Steppe pastoralists measured by <i>f</i><sub><i>4</i></sub><i>(Ethiopia_4500BP, X; Central_Steppe_MLBA, Ganj_Dareh_N</i>) reveals three ancestry clines that succeeded each other in time: the <i>Indus Periphery Cline</i> prior to ~2000 BCE, the <i>Steppe Cline</i> represented by northern South Asian individuals after ~2000 BCE, and the <i>Modern Indian Cline.</i> <b>(B)</b> Modeling South Asians as a mixture of <i>Central_Steppe_MLBA, AHG</i> (as a proxy for <i>AASI</i>), and <i>Indus_Periphery_West</i> (the individual from the <i>Indus Periphery Cline</i> with the least <i>AASI</i> ancestry). Groups along the edges of the triangle fit a two-way model, and in the interior only fit a three-way model. The 140 present-day South Asian groups on the <i>Modern Indian Cline</i> are shown as small dots. (<b>C</b>) Groups that traditionally view themselves as being of priestly status in this and the preceding panel are shown in red (“Brahmin,” “Pandit,” and “Bhumihar” but excluding “Catholic Brahmins”), and tend to have a significantly higher ratio of <i>Central_Steppe_MLBA</i> to <i>Indus_Periphery_Cline</i> ancestry than other groups. (<b>D</b>) Plot of the proportion of <i>Central_Steppe_MLBA</i> ancestry on the autosomes (x-axis) and the Y chromosome (y-axis) shows that the source of this ancestry is primarily from females in Late Bronze Age and Iron Age individuals from the Swat District of northernmost South Asia, and primarily from males in most present-day South Asians.</div> </figcaption> </figure> <figure class="figure-item " itemscope itemtype="http://schema.org/ImageObject" itemprop="associatedMedia" data-slide-index="4" data-label-slug="fig-5"> <a class="figure-link" href="https://cdn.ncbi.nlm.nih.gov/pmc/blobs/d74d/6822619/f3d6c5c929c1/nihms-1053677-f0006.jpg" itemprop="contentUrl" aria-describedby="figure-caption-4" role="button" data-image-width="800" data-image-height="569" data-image-alt="Fig. 5." data-pmc-id="PMC6822619" data-figure-id="F5"> <img class="figure-thumb" itemprop="thumbnail" id="article-image-4" src="https://cdn.ncbi.nlm.nih.gov/pmc/blobs/d74d/6822619/f638ffb61967/nihms-1053677-f0006.gif" alt="Fig. 5." /> </a> <meta itemprop="width" itemtype="http://schema.org/ImageObject" content="800"> <meta itemprop="height" itemtype="http://schema.org/ImageObject" content="569"> <div class="figure-caption figure-caption-text" itemtype="http://schema.org/ImageObject" itemprop="caption" aria-hidden="true"> <div class="caption-wrap"> <strong class="figure-label"> <p> Fig. 5.. Admixture Graph Model. </p> </strong> <div class="figure-caption-contents"><p> The largest… </p></div> </div> </div> <div class="figure-caption-medium figure-caption-text" aria-hidden="true"> <strong class="figure-label"> <p> Fig. 5.. Admixture Graph Model. </p> </strong> <div class="figure-caption-contents"><p> The largest deviation between empirical and theoretical <i> f </i> -statistics is… </p></div> </div> <figcaption id="figure-caption-4" class="figure-caption-full figure-caption-text" itemtype="http://schema.org/ImageObject" itemprop="description"> <strong class="figure-label"> Fig. 5.. Admixture Graph Model. </strong> <div class="figure-caption-contents">The largest deviation between empirical and theoretical <i>f</i>-statistics is |Z|=2.9, indicating a good fit considering the large number of <i>f-</i>statistics analyzed. Admixture events are shown as dotted lines labeled by proportions, with the minor ancestry in gray. The present-day groups are shown in orange ovals, the ancient ones in blue, and unsampled groups in white. (The ovals and admixture events are positioned according to guesses about their relative dates to help in visualization, although the dates are in no way meant to be exact.) In this graph we do not attempt to model the contribution of <i>WSHG</i> and Anatolian farmer-related ancestry, and thus cannot model <i>Central_Steppe_EMBA,</i> the proximal source of Steppe ancestry in South Asia (instead we model the Steppe ancestry in South Asia through the more distally related <i>Yamnaya</i>). However, the admixture graph does highlight several key findings of the study, including the deep separation of the <i>AASI</i> from other Eurasian lineages, and the fact that some Austroasiatic-speaking groups in South Asia (e.g. Juang) harbor ancestry from a South Asian group with a higher ratio of <i>AASI-</i>related to Iranian farmer-related ancestry than any groups on the <i>Modern Indian Cline</i>, thus revealing that groups with substantial Iranian farmer-related ancestry were not ubiquitous in peninsular South Asia in the 3<sup>rd</sup> millennium BCE when Austroasiatic languages likely spread across the subcontinent.</div> </figcaption> </figure> </div>  <div class="pswp figures-dialog" id="figures-dialog" tabindex="-1" role="dialog" aria-hidden="true" data-image-gallery-uid="article-figures" aria-label="Figures Dialog">  <div class="pswp__bg"></div>  <div class="pswp__scroll-wrap">  <div class="pswp__container"> <div class="pswp__item"></div> <div class="pswp__item"></div> <div class="pswp__item"></div> </div>  <div class="pswp__ui pswp__ui--hidden"> <div class="pswp__top-bar">  <div class="pswp__counter"></div> <button class="pswp__button pswp__button--close" title="Close (Esc)"></button> <button class="pswp__button pswp__button--share" title="Share"></button> <button class="pswp__button pswp__button--fs" title="Toggle fullscreen"></button> <button class="pswp__button pswp__button--zoom" title="Zoom in/out"></button>   <div class="pswp__preloader"> <div class="pswp__preloader__icn"> <div class="pswp__preloader__cut"> <div class="pswp__preloader__donut"></div> </div> </div> </div> </div> <div class="pswp__share-modal pswp__share-modal--hidden pswp__single-tap"> <div class="pswp__share-tooltip"></div> </div> <button class="pswp__button pswp__button--arrow--left" title="Previous (arrow left)"> </button> <button class="pswp__button pswp__button--arrow--right" title="Next (arrow right)"> </button> <div class="pswp__caption"> <div class="pswp__caption__center"></div> </div> </div> </div> </div> <div class="figure-dialog-caption"> <div class="figure-caption-link-container"> <a class="figure-caption-link" href="#">See this image and copyright information in PMC</a> </div> <div class="figure-caption-center" data-expand-title="click to show full caption" data-collapse-title="click to hide full caption"> </div> <button class="figure-caption-toggle-btn" aria-label="show full caption" data-expand-text="show full caption" data-collapse-text="hide full caption">  </button> </div> </div> <div class="linked-articles" id="linked-commentary"> <h2 class="title"> Comment in </h2> <ul class="item-list usa-unstyled-list"> <li class="full-docsum"> <div class="docsum-content" style="display:inline;"> <a class="docsum-title" href="/31488673/" ref="article_id=31488673&linksrc=comments_link" data-ga-category="comment_correction" data-ga-action="abstract_page_link" data-ga-label="linked-commentary"> New middle chapter in the story of human evolution. </a> <div class="docsum-citation full-citation"> <span class="docsum-authors full-authors">Schaefer NK, Shapiro B.</span> <span class="docsum-authors short-authors">Schaefer NK, et al.</span> <span class="docsum-journal-citation full-journal-citation">Science. 2019 Sep 6;365(6457):981-982. doi: 10.1126/science.aay3550.</span> <span class="docsum-journal-citation short-journal-citation">Science. 2019.</span> <span class="citation-part">PMID: <span class="docsum-pmid">31488673</span></span> <span class="no-abstract spaced-citation-item citation-part">No abstract available.</span> </div> </div> </li> </ul> </div> <div class="similar-articles" id="similar"> <h2 class="title"> Similar articles </h2> <ul class="articles-list" id="similar-articles-list"> <li class="full-docsum"> <div class="docsum-content" style="display:inline;"> <a class="docsum-title" href="/31495572/" ref="article_id=31495572&linksrc=similar_articles_link&ordinalpos=1" data-ga-category="similar_article" data-ga-action="31495572" data-ga-label=""> An Ancient Harappan Genome Lacks Ancestry from Steppe Pastoralists or Iranian Farmers. </a> <div class="docsum-citation full-citation"> <span class="docsum-authors full-authors">Shinde V, Narasimhan VM, Rohland N, Mallick S, Mah M, Lipson M, Nakatsuka N, Adamski N, Broomandkhoshbacht N, Ferry M, Lawson AM, Michel M, Oppenheimer J, Stewardson K, Jadhav N, Kim YJ, Chatterjee M, Munshi A, Panyam A, Waghmare P, Yadav Y, Patel H, Kaushik A, Thangaraj K, Meyer M, Patterson N, Rai N, Reich D.</span> <span class="docsum-authors short-authors">Shinde V, et al.</span> <span class="docsum-journal-citation full-journal-citation">Cell. 2019 Oct 17;179(3):729-735.e10. doi: 10.1016/j.cell.2019.08.048. 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(Cambridge University Press, Cambridge, 2017), pp. 304–331. </li> </ol> </li> <li> <ol class="references-and-notes-list"> <li class="skip-numbering" value="1"> Stevens CJ et al., Between China and South Asia: A Middle Asian corridor of crop dispersal and agricultural innovation in the Bronze Age. The Holocene. 26, 1541–1555 (2016). - <a class="reference-link" href="https://pmc.ncbi.nlm.nih.gov/articles/PMC5125436/" ref="linksrc=references_link&ordinalpos=1" data-ga-category="reference" data-ga-action="PMC5125436" > PMC </a> - <a class="reference-link" href="/27942165/" ref="linksrc=references_link&ordinalpos=2" data-ga-category="reference" data-ga-action="27942165" > PubMed </a> </li> </ol> </li> <li> <ol class="references-and-notes-list"> <li class="skip-numbering" value="1"> Allaby RG, Stevens C, Lucas L, Maeda O, Fuller DQ, Geographic mosaics and changing rates of cereal domestication. Philos. Trans. R. Soc. B Biol. Sci 372, 20160429 (2017). - <a class="reference-link" href="https://pmc.ncbi.nlm.nih.gov/articles/PMC5665816/" ref="linksrc=references_link&ordinalpos=1" data-ga-category="reference" data-ga-action="PMC5665816" > PMC </a> - <a class="reference-link" href="/29061901/" ref="linksrc=references_link&ordinalpos=2" data-ga-category="reference" data-ga-action="29061901" > PubMed </a> </li> </ol> </li> <li> <ol class="references-and-notes-list"> <li class="skip-numbering" value="1"> Dani AH et al., History of Civilizations of Central Asia: The Development of Sedentary and Nomadic Civilizations, 700 B. C. to A (UNESCO Publishing, 1994). </li> </ol> </li> </ol> <div class="actions-bar"> <button class="show-all" ref="linksrc=show_all_references_link" aria-controls="top-references-list-1" data-next-page-url="/31488661/references/" data-ga-category="reference" data-ga-action="show_more" > Show all 228 references </button> </div> </div> </div> </div> <div id="publication-types" class="publication-types keywords-section"> <h2 class="title"> Publication types </h2> <ul class="keywords-list"><li><div class="keyword-actions dropdown-block"><button class="keyword-actions-trigger trigger keyword-link" aria-label="Toggle dropdown menu for keyword Historical Article" tabindex="0" aria-controls="keyword-actions-publication-types-0" aria-expanded="false" data-pinger-ignore> Historical Article </button><div id="keyword-actions-publication-types-0" class="keyword-actions-dropdown dropdown dropdown-container" hidden aria-label="Dropdown menu for keyword Historical Article"><div 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id="keyword-actions-mesh-terms-0" class="keyword-actions-dropdown dropdown dropdown-container" hidden aria-label="Dropdown menu for keyword Asia, Central"><div class="title">Actions</div><div class="content"><ul class="keyword-actions-links"><li><a class="search-in-pubmed-link dropdown-block-link" href="/?term=%22Asia%2C+Central%22%5BMeSH%5D&sort=date&sort_order=desc" data-href="/?term=%22Asia%2C+Central%22%5BMeSH%5D&sort=date&sort_order=desc" ref="linksrc=search_in_pubmed_keyword_link&mesh="Asia, Central"[MeSH]" data-ga-category="search" data-ga-action="mesh_term" data-ga-label="Asia, Central"> Search in PubMed </a></li><li><a class="search-in-mesh-link dropdown-block-link" ref="linksrc=search_in_mesh_link" href="https://www.ncbi.nlm.nih.gov/mesh?term=Asia%2C+Central" data-ga-category="search_mesh" data-ga-action="mesh_term_link" data-ga-label="Asia, Central"> Search in MeSH </a></li><li><a class="add-to-search-link dropdown-block-link" ref="linksrc=add_to_search_link" role="button" data-search-term=""Asia, Central"[MeSH]" data-ga-category="search" data-ga-action="add_to_search" data-ga-label=""Asia, Central"[MeSH]" href="#"> Add to Search </a></li></ul></div></div></div></li><li><div class="keyword-actions dropdown-block"><button class="keyword-actions-trigger trigger keyword-link" aria-label="Toggle dropdown menu for keyword Asia, Southeastern" tabindex="0" aria-controls="keyword-actions-mesh-terms-1" aria-expanded="false" data-pinger-ignore> Asia, Southeastern </button><div id="keyword-actions-mesh-terms-1" class="keyword-actions-dropdown dropdown dropdown-container" hidden aria-label="Dropdown menu for keyword Asia, Southeastern"><div class="title">Actions</div><div class="content"><ul class="keyword-actions-links"><li><a class="search-in-pubmed-link dropdown-block-link" href="/?term=%22Asia%2C+Southeastern%22%5BMeSH%5D&sort=date&sort_order=desc" data-href="/?term=%22Asia%2C+Southeastern%22%5BMeSH%5D&sort=date&sort_order=desc" ref="linksrc=search_in_pubmed_keyword_link&mesh="Asia, Southeastern"[MeSH]" data-ga-category="search" data-ga-action="mesh_term" data-ga-label="Asia, Southeastern"> Search in PubMed </a></li><li><a class="search-in-mesh-link dropdown-block-link" ref="linksrc=search_in_mesh_link" href="https://www.ncbi.nlm.nih.gov/mesh?term=Asia%2C+Southeastern" data-ga-category="search_mesh" data-ga-action="mesh_term_link" data-ga-label="Asia, Southeastern"> Search in MeSH </a></li><li><a class="add-to-search-link dropdown-block-link" ref="linksrc=add_to_search_link" role="button" data-search-term=""Asia, Southeastern"[MeSH]" data-ga-category="search" data-ga-action="add_to_search" data-ga-label=""Asia, Southeastern"[MeSH]" href="#"> Add to Search </a></li></ul></div></div></div></li><li><div class="keyword-actions dropdown-block"><button class="keyword-actions-trigger trigger keyword-link major" aria-label="Toggle dropdown menu for keyword Asian People / genetics*" tabindex="0" aria-controls="keyword-actions-mesh-terms-major-2" aria-expanded="false" data-pinger-ignore> Asian People / genetics* </button><div id="keyword-actions-mesh-terms-major-2" class="keyword-actions-dropdown dropdown dropdown-container" hidden aria-label="Dropdown menu for keyword Asian People / genetics*"><div class="title">Actions</div><div class="content"><ul class="keyword-actions-links"><li><a class="search-in-pubmed-link dropdown-block-link" href="/?term=%22Asian+People%2Fgenetics%22%5BMAJR%5D&sort=date&sort_order=desc" data-href="/?term=%22Asian+People%2Fgenetics%22%5BMAJR%5D&sort=date&sort_order=desc" ref="linksrc=search_in_pubmed_keyword_link&mesh="Asian People/genetics"[MAJR]" data-ga-category="search" data-ga-action="mesh_term" data-ga-label="Asian People / genetics*"> Search in PubMed </a></li><li><a class="search-in-mesh-link dropdown-block-link" ref="linksrc=search_in_mesh_link" href="https://www.ncbi.nlm.nih.gov/mesh?term=Asian+People" data-ga-category="search_mesh" data-ga-action="mesh_term_link" data-ga-label="Asian People / genetics*"> Search in MeSH </a></li><li><a class="add-to-search-link dropdown-block-link" ref="linksrc=add_to_search_link" role="button" data-search-term=""Asian People/genetics"[MAJR]" data-ga-category="search" data-ga-action="add_to_search" data-ga-label=""Asian People/genetics"[MAJR]" href="#"> Add to Search </a></li></ul></div></div></div></li><li><div class="keyword-actions dropdown-block"><button class="keyword-actions-trigger trigger keyword-link major" aria-label="Toggle dropdown menu for keyword Farms / history*" tabindex="0" aria-controls="keyword-actions-mesh-terms-major-3" aria-expanded="false" data-pinger-ignore> Farms / history* </button><div id="keyword-actions-mesh-terms-major-3" class="keyword-actions-dropdown dropdown dropdown-container" hidden aria-label="Dropdown menu for keyword Farms / history*"><div class="title">Actions</div><div class="content"><ul class="keyword-actions-links"><li><a class="search-in-pubmed-link dropdown-block-link" href="/?term=%22Farms%2Fhistory%22%5BMAJR%5D&sort=date&sort_order=desc" data-href="/?term=%22Farms%2Fhistory%22%5BMAJR%5D&sort=date&sort_order=desc" ref="linksrc=search_in_pubmed_keyword_link&mesh="Farms/history"[MAJR]" data-ga-category="search" data-ga-action="mesh_term" data-ga-label="Farms / history*"> Search in PubMed </a></li><li><a class="search-in-mesh-link dropdown-block-link" ref="linksrc=search_in_mesh_link" href="https://www.ncbi.nlm.nih.gov/mesh?term=Farms" data-ga-category="search_mesh" data-ga-action="mesh_term_link" data-ga-label="Farms / history*"> Search in MeSH </a></li><li><a class="add-to-search-link dropdown-block-link" ref="linksrc=add_to_search_link" role="button" data-search-term=""Farms/history"[MAJR]" data-ga-category="search" data-ga-action="add_to_search" data-ga-label=""Farms/history"[MAJR]" href="#"> Add to Search </a></li></ul></div></div></div></li><li><div class="keyword-actions dropdown-block"><button class="keyword-actions-trigger trigger keyword-link" aria-label="Toggle dropdown menu for keyword Gene Flow" tabindex="0" aria-controls="keyword-actions-mesh-terms-4" aria-expanded="false" data-pinger-ignore> Gene Flow </button><div id="keyword-actions-mesh-terms-4" class="keyword-actions-dropdown dropdown dropdown-container" hidden aria-label="Dropdown menu for keyword Gene Flow"><div class="title">Actions</div><div class="content"><ul class="keyword-actions-links"><li><a class="search-in-pubmed-link dropdown-block-link" href="/?term=%22Gene+Flow%22%5BMeSH%5D&sort=date&sort_order=desc" data-href="/?term=%22Gene+Flow%22%5BMeSH%5D&sort=date&sort_order=desc" ref="linksrc=search_in_pubmed_keyword_link&mesh="Gene Flow"[MeSH]" data-ga-category="search" data-ga-action="mesh_term" data-ga-label="Gene Flow"> Search in PubMed </a></li><li><a class="search-in-mesh-link dropdown-block-link" ref="linksrc=search_in_mesh_link" href="https://www.ncbi.nlm.nih.gov/mesh?term=Gene+Flow" data-ga-category="search_mesh" data-ga-action="mesh_term_link" data-ga-label="Gene Flow"> Search in MeSH </a></li><li><a class="add-to-search-link dropdown-block-link" ref="linksrc=add_to_search_link" role="button" data-search-term=""Gene Flow"[MeSH]" data-ga-category="search" data-ga-action="add_to_search" data-ga-label=""Gene Flow"[MeSH]" href="#"> Add to Search </a></li></ul></div></div></div></li><li><div class="keyword-actions dropdown-block"><button class="keyword-actions-trigger trigger keyword-link" aria-label="Toggle dropdown menu for keyword History, Ancient" tabindex="0" aria-controls="keyword-actions-mesh-terms-5" aria-expanded="false" data-pinger-ignore> History, Ancient </button><div id="keyword-actions-mesh-terms-5" class="keyword-actions-dropdown dropdown dropdown-container" hidden aria-label="Dropdown menu for keyword History, Ancient"><div class="title">Actions</div><div class="content"><ul class="keyword-actions-links"><li><a class="search-in-pubmed-link dropdown-block-link" href="/?term=%22History%2C+Ancient%22%5BMeSH%5D&sort=date&sort_order=desc" data-href="/?term=%22History%2C+Ancient%22%5BMeSH%5D&sort=date&sort_order=desc" ref="linksrc=search_in_pubmed_keyword_link&mesh="History, Ancient"[MeSH]" data-ga-category="search" data-ga-action="mesh_term" data-ga-label="History, Ancient"> Search in PubMed </a></li><li><a class="search-in-mesh-link dropdown-block-link" ref="linksrc=search_in_mesh_link" href="https://www.ncbi.nlm.nih.gov/mesh?term=History%2C+Ancient" data-ga-category="search_mesh" data-ga-action="mesh_term_link" data-ga-label="History, Ancient"> Search in MeSH </a></li><li><a class="add-to-search-link dropdown-block-link" ref="linksrc=add_to_search_link" role="button" data-search-term=""History, Ancient"[MeSH]" data-ga-category="search" data-ga-action="add_to_search" data-ga-label=""History, Ancient"[MeSH]" href="#"> Add to Search </a></li></ul></div></div></div></li><li><div class="keyword-actions dropdown-block"><button class="keyword-actions-trigger trigger keyword-link major" aria-label="Toggle dropdown menu for keyword Human Migration / history*" tabindex="0" aria-controls="keyword-actions-mesh-terms-major-6" aria-expanded="false" data-pinger-ignore> Human Migration / history* </button><div id="keyword-actions-mesh-terms-major-6" class="keyword-actions-dropdown dropdown dropdown-container" hidden aria-label="Dropdown menu for keyword Human Migration / history*"><div class="title">Actions</div><div class="content"><ul class="keyword-actions-links"><li><a class="search-in-pubmed-link dropdown-block-link" href="/?term=%22Human+Migration%2Fhistory%22%5BMAJR%5D&sort=date&sort_order=desc" data-href="/?term=%22Human+Migration%2Fhistory%22%5BMAJR%5D&sort=date&sort_order=desc" ref="linksrc=search_in_pubmed_keyword_link&mesh="Human Migration/history"[MAJR]" data-ga-category="search" data-ga-action="mesh_term" data-ga-label="Human Migration / history*"> Search in PubMed </a></li><li><a class="search-in-mesh-link dropdown-block-link" ref="linksrc=search_in_mesh_link" href="https://www.ncbi.nlm.nih.gov/mesh?term=Human+Migration" data-ga-category="search_mesh" data-ga-action="mesh_term_link" data-ga-label="Human Migration / history*"> Search in MeSH </a></li><li><a class="add-to-search-link dropdown-block-link" ref="linksrc=add_to_search_link" role="button" data-search-term=""Human Migration/history"[MAJR]" data-ga-category="search" data-ga-action="add_to_search" data-ga-label=""Human Migration/history"[MAJR]" href="#"> Add to Search </a></li></ul></div></div></div></li><li><div class="keyword-actions dropdown-block"><button class="keyword-actions-trigger trigger keyword-link" aria-label="Toggle dropdown menu for keyword Humans" tabindex="0" aria-controls="keyword-actions-mesh-terms-7" aria-expanded="false" data-pinger-ignore> Humans </button><div id="keyword-actions-mesh-terms-7" class="keyword-actions-dropdown dropdown dropdown-container" hidden aria-label="Dropdown menu for keyword Humans"><div class="title">Actions</div><div class="content"><ul class="keyword-actions-links"><li><a class="search-in-pubmed-link dropdown-block-link" href="/?term=%22Humans%22%5BMeSH%5D&sort=date&sort_order=desc" data-href="/?term=%22Humans%22%5BMeSH%5D&sort=date&sort_order=desc" ref="linksrc=search_in_pubmed_keyword_link&mesh="Humans"[MeSH]" data-ga-category="search" data-ga-action="mesh_term" data-ga-label="Humans"> Search in PubMed </a></li><li><a class="search-in-mesh-link dropdown-block-link" ref="linksrc=search_in_mesh_link" href="https://www.ncbi.nlm.nih.gov/mesh?term=Humans" data-ga-category="search_mesh" data-ga-action="mesh_term_link" data-ga-label="Humans"> Search in MeSH </a></li><li><a class="add-to-search-link dropdown-block-link" ref="linksrc=add_to_search_link" role="button" data-search-term=""Humans"[MeSH]" data-ga-category="search" data-ga-action="add_to_search" data-ga-label=""Humans"[MeSH]" href="#"> Add to Search </a></li></ul></div></div></div></li><li><div class="keyword-actions dropdown-block"><button class="keyword-actions-trigger trigger keyword-link" aria-label="Toggle dropdown menu for keyword Iran" tabindex="0" aria-controls="keyword-actions-mesh-terms-8" aria-expanded="false" data-pinger-ignore> Iran </button><div id="keyword-actions-mesh-terms-8" class="keyword-actions-dropdown dropdown dropdown-container" hidden aria-label="Dropdown menu for keyword Iran"><div class="title">Actions</div><div class="content"><ul class="keyword-actions-links"><li><a class="search-in-pubmed-link dropdown-block-link" href="/?term=%22Iran%22%5BMeSH%5D&sort=date&sort_order=desc" data-href="/?term=%22Iran%22%5BMeSH%5D&sort=date&sort_order=desc" ref="linksrc=search_in_pubmed_keyword_link&mesh="Iran"[MeSH]" data-ga-category="search" data-ga-action="mesh_term" data-ga-label="Iran"> Search in PubMed </a></li><li><a class="search-in-mesh-link dropdown-block-link" ref="linksrc=search_in_mesh_link" href="https://www.ncbi.nlm.nih.gov/mesh?term=Iran" data-ga-category="search_mesh" data-ga-action="mesh_term_link" data-ga-label="Iran"> Search in MeSH </a></li><li><a class="add-to-search-link dropdown-block-link" ref="linksrc=add_to_search_link" role="button" data-search-term=""Iran"[MeSH]" data-ga-category="search" data-ga-action="add_to_search" data-ga-label=""Iran"[MeSH]" href="#"> Add to Search </a></li></ul></div></div></div></li><li><div class="keyword-actions dropdown-block"><button class="keyword-actions-trigger trigger keyword-link major" aria-label="Toggle dropdown menu for keyword Population / genetics*" tabindex="0" aria-controls="keyword-actions-mesh-terms-major-9" aria-expanded="false" data-pinger-ignore> Population / genetics* </button><div id="keyword-actions-mesh-terms-major-9" class="keyword-actions-dropdown dropdown dropdown-container" hidden aria-label="Dropdown menu for keyword Population / genetics*"><div class="title">Actions</div><div class="content"><ul class="keyword-actions-links"><li><a class="search-in-pubmed-link dropdown-block-link" href="/?term=%22Population%2Fgenetics%22%5BMAJR%5D&sort=date&sort_order=desc" data-href="/?term=%22Population%2Fgenetics%22%5BMAJR%5D&sort=date&sort_order=desc" ref="linksrc=search_in_pubmed_keyword_link&mesh="Population/genetics"[MAJR]" data-ga-category="search" data-ga-action="mesh_term" data-ga-label="Population / genetics*"> Search in PubMed </a></li><li><a class="search-in-mesh-link dropdown-block-link" ref="linksrc=search_in_mesh_link" href="https://www.ncbi.nlm.nih.gov/mesh?term=Population" data-ga-category="search_mesh" data-ga-action="mesh_term_link" data-ga-label="Population / genetics*"> Search in MeSH </a></li><li><a class="add-to-search-link dropdown-block-link" ref="linksrc=add_to_search_link" role="button" data-search-term=""Population/genetics"[MAJR]" data-ga-category="search" data-ga-action="add_to_search" data-ga-label=""Population/genetics"[MAJR]" href="#"> Add to Search 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DNA"> Search in PubMed </a></li><li><a class="search-in-mesh-link dropdown-block-link" ref="linksrc=search_in_mesh_link" href="https://www.ncbi.nlm.nih.gov/mesh?term=Sequence+Analysis%2C+DNA" data-ga-category="search_mesh" data-ga-action="mesh_term_link" data-ga-label="Sequence Analysis, DNA"> Search in MeSH </a></li><li><a class="add-to-search-link dropdown-block-link" ref="linksrc=add_to_search_link" role="button" data-search-term=""Sequence Analysis, DNA"[MeSH]" data-ga-category="search" data-ga-action="add_to_search" data-ga-label=""Sequence Analysis, DNA"[MeSH]" href="#"> Add to Search </a></li></ul></div></div></div></li></ul> </div> <div id="related-links" class="related-links"> <h2 class="title"> Related information </h2> <ul class="related-links-list"><li><a title="PubMed links associated with Books" href="https://www.ncbi.nlm.nih.gov/books?linkname=pubmed_books_refs&from_uid=31488661" ref="log$=recordlinks&logdbfrom=pubmed&ncbi_uid=31488661&ordinalpos=1&linkpos=1"> Cited in 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