CINXE.COM
Just a moment...
<!DOCTYPE html><html lang="en-US"><head><title>Just a moment...</title><meta http-equiv="Content-Type" content="text/html; charset=UTF-8"><meta http-equiv="X-UA-Compatible" content="IE=Edge"><meta name="robots" content="noindex,nofollow"><meta name="viewport" content="width=device-width,initial-scale=1"><style>*{box-sizing:border-box;margin:0;padding:0}html{line-height:1.15;-webkit-text-size-adjust:100%;color:#313131;font-family:system-ui,-apple-system,BlinkMacSystemFont,Segoe UI,Roboto,Helvetica Neue,Arial,Noto Sans,sans-serif,Apple Color Emoji,Segoe UI Emoji,Segoe UI Symbol,Noto Color Emoji}body{display:flex;flex-direction:column;height:100vh;min-height:100vh}.main-content{margin:8rem auto;max-width:60rem;padding-left:1.5rem}@media (width <= 720px){.main-content{margin-top:4rem}}.h2{font-size:1.5rem;font-weight:500;line-height:2.25rem}@media (width <= 720px){.h2{font-size:1.25rem;line-height:1.5rem}}#challenge-error-text{background-image:url(data:image/svg+xml;base64,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);background-repeat:no-repeat;background-size:contain;padding-left:34px}@media (prefers-color-scheme:dark){body{background-color:#222;color:#d9d9d9}}</style><meta http-equiv="refresh" content="390"></head><body class="no-js"><div class="main-wrapper" role="main"><div class="main-content"><noscript><div class="h2"><span id="challenge-error-text">Enable JavaScript and cookies to continue</span></div></noscript></div></div><script>(function(){window._cf_chl_opt={cvId: '3',cZone: "journals.biologists.com",cType: 'managed',cRay: '8eaf35017f2d4073',cH: 'o.XENlpvOSpLEGMtTYjowF2j.Fx6h7Q3CMxAEgzBoDw-1733015526-1.2.1.1-u49LUirNszbMxctV_hnYwjQVJj5DACXOj1amju8Ld8X00Q624nOsdmlmpVn4IazI',cUPMDTk: "\/dev\/article\/123\/1\/153\/39329\/Mutations-affecting-somite-formation-and?__cf_chl_tk=k0hhzrdWuIIb_FO2.l86jraSCOFldaxh0SGowYt3iSc-1733015526-1.0.1.1-mad4Jq8MDPXdnNaX6865yexwX10OuYDeko0mQCslvmQ",cFPWv: 'b',cITimeS: '1733015526',cTTimeMs: '1000',cMTimeMs: '390000',cTplC: 0,cTplV: 5,cTplB: 'cf',cK: "",fa: "\/dev\/article\/123\/1\/153\/39329\/Mutations-affecting-somite-formation-and?__cf_chl_f_tk=k0hhzrdWuIIb_FO2.l86jraSCOFldaxh0SGowYt3iSc-1733015526-1.0.1.1-mad4Jq8MDPXdnNaX6865yexwX10OuYDeko0mQCslvmQ",md: "PMg8yTw5aHLQ_YeHRaa3lUyXe7qobqsvu4x7A2LOyik-1733015526-1.2.1.1-lEr28SE4HKUsc4w.Qrb4YkcGK95wx2qRosNaic3fmHiWGJkPsfFKvHpkDrIgCBUX5dsbDRkV0hLZ.9QW21_qQIChZWwurpj5VeU4gwXGvAFXdnYhtzQUO9nHrb4fJuXRHkOEr4rtH_.f4MboJeEERj4EawZNoSvDD.DP0Tqy6g3Aflo4K1cHuUf6XGVUNuUOMieT.iLuwkSSkb2TT6.4_XZ5M0mfQUxKsW5Z4Ea5OZkHmwrLlK3Su8Z37cZpde_jTiP5I4TtVurRkcheEg4cPCTaRrI43uNdeG7SZ1E1emoPHD8K_0PhoPBcIdL74JsBGlslWsHbpPYart56Crr5MV5K8W4sE0YzXDFrReZUgyxAcviwglef5BT2q7kNuCTp.WgQB5eYNnoS6WLQu3LaBkKWfFUjjCUXkkgS7qJJ2eJMiBkWbdLlaZ9ekQHQPp94kNpVv3DJgJp.SNMOqG0MeVBPKy0J1GKMXCA.M01nBw4_CiuXeANh4KROrhWQSWx_Y89rIOeqL7RBAWsNnh4u8vwaPDESItWiD6mb50Cqk5sKk.kYKg6VV7eMExdtAHjQIyNJPT6O8F5Yp6pkZ.HLoESz6wEFgtmWwb_v1uj.vX126GCDB0LxOSzhxf4pOemWsseIAuukS8tpufnU_inSYIWRJhKjc73aYdU2.whTmItcoC9Y876wJAzodsSoUQvRxUps2Ylg9XTl97U3CBL0t1pwsvk9N5CWUvvrEVsExgkCEDmNLMlSwviy1vWaqTbV9zOeHwMwrGGZkI66Gl0zeuiUwmFFI4Vl8v5ofQXb7l2TVSg9wMnJMjQt.ruqa7gNfFfMD6DhdEbzbwn9HJJ.I90bc10ZsqvSeh.8TmS6vxD9DNoX9yRUHVZMbZBDsdg2fk.HeAM29XFkWL3OkCBvialn75Mu8WEkVsr_nExaM4HBvEu94n8mVkiCKI2jQtJbYG6Pie9qfG8KZCBRIEYdUOxkLGYjET6iz8VFhqL5044lVFm0l8EpJiTSPO5zjSKCDRuR3XJPwgpQY9ZVYhH6IiSvwWShqAed3fbO4LEfkd4mSzsuGXY1qmYGDyWAgXlPiaOsFiNs_ZALkNNHth_EwsMynARKXMVac.6xm.vg8lpX9A4k9GlU8ZTAN_4TGR23bMkabHsKccU3kyFXV24ikgvPKUpzOZGknsGpG2v9m9KdiHyQOQWG4a4APB0MLxD6SjibLP_a6SX5wsxW7QHam1yVaB2YTk2fX_9TPN2tsLtyhGnYLRCZNmeZ10aUaDivsIUar0UGaGF3hcsLKeHHtQzt.wzhsZXANtq8qGEPxpzepaYREIkxNqarzeKYvtfWMEko.snSF6dYqvCNAuzkg5UxwEccHxaEjRd5GAad9V44d4F6EOIFYac2EN3MnmbCodaFp3XqRrnqNeJfahiyPTlNmYIABL0SAjIhstAeWLlnsDj4Ike5N1jtwmJQrkz2Z7sYwmi4rH98FQ6pp2AAI40_5AnS88rpbRt2VTA.gwIR3V9tgn7ZOFot14vRTbsET_7eYRGGeZnqe_H3bi_F19ZWqDZgeNa8jJ_A7PLZa84pJz1jhmexryayUPr9f3YIciTdrDzeTd9DxFNRAd1g6q9k1qRm8_hSFPsm0oPRfqjmtqOMVM5YfICSPkRwdY5gMSn6iK8Tg91OxYjEbF_smBHAq2G2i32hT7.LQF4yDbpqJGesZ0QQTr.TbMlhJqLfxhW3JYHp4Y9I6CrKRcKumJ._85HrEbY8F0_TJk3HDmzYm2P1YfGTdvJ6YT49Z4GrJ2gIo65PLOsW_QsNRjNRGU19VP6COxlqzb2Gy8LZIUyOA0duFTYsxbRVSImk2rcSQ3ZcRAlmIfjDXKKSMkURtJaNfktN77Z5S.c29.Y7RnEJ3QtODPj9Iw65CjYaRaPZoJr1_o7TChzBFri9QEdr9xCSmncVo1U28he5._CkR5Mpz3yA6e2wZ9tYkqqOgQgyYHEyxFtgW9.afI8ytckdvmiE4ZTHkUDEXW4nrngnp9G4iIdohdZw63R7TzW6oRXJ2dxqO5umede7WHKIHNpS3sfcxe4P4s3eZeNjzAgtat6Y8KAHBsrQVRXjgrA_5j1HvXUK06.6wBTem9J31N09M_FlRSOFEN7ZWAQnZMf5ym2NAH4vinJUhH.jUkhgBHgXzvHH1ByUElU1bFeHlLO4v7G0954fIJGteiDF9u2GHQPqzQUfdiIL5xVuQLvmTSByubdhOuEWANHqKnzutnf7iFkpsYt9xRZg2bjvLPIgJf16puISbNDDw2l4mOnF2c18Bps1_dgf2ZzsX4RUAXhstNhm1Wf_QwaNwSE.r6e936GgQs3FI6tF1eICpORriX4UPWHwkTAgs1PuCfKGeomX8YqUG2kHiK6KsTqyc3n40RQOLbWNbbNQ3_9zP2_.8nHRCcs4sBYSqzKExIiVvCWJINDwxMWUBZWjNYa1ZiqmoOVY6zkdVEskwBhKqYSqDe.twPEWzoW8BuA9ad_cuCey7IlN54NTK3lohxMDZzzl._fmQj_kIwTJb52hKwpPk72WXkiM9pb6k_0b4wfVhpzbXlotq81g1AM5F2RqjmbhJxCYH_MDm.xxOjmKTW7k8U8_VW3YdqBq8pY9PxDpXcS3Sg1fDXcdzb0JX2wFB77JFYp7_QNJPVZqWeuie35mFy4tRttJFWB4sQadCRT5ag6FmmOcwdCmCslJaYfLuLAI3KbhVCBV8d1F46D7oL7kZV3_s28P2ja7DqK7LzJX61qhfwEi4jlDCcgrey1gsyN5xLpEY2cHNb9bliiGhrSvIIh5X5PbuVhC.1et05Vx6SnPzumTOIvm_zqwIfXfkKiK8Fn_h_TgG1n3kjXBfAgZPuCMC12pQGNfxMRpSbHB47Z6EFDF6bHV6rAq8cyI02I8ecA39Mhin3lU7sejXSlEbT7VYU4X7ez4.cxLmsF1eRAHvhlBvDoAnPxkpQw1R9P2miv7eAsUbl7srJsOav2Jkf3.LeYSqUxDkVG2LYug1Yx5b8IrgGlKAoadJjdZFmPOqHN.GLOSk71N5Po4cwwYS3XfzhiqySNessRoKGz..405r91Ww8a8MGVg2z2OJSHQRr8",mdrd: "j37x7j5.Mw0JxlPdx.QpFJ_5ngkrNdavN0T94zd81EE-1733015526-1.2.1.1-nf2MoaJ7LJyJL5K3U8TlJRk1oqoywrSsn59Bx0SU82i.NJO8JsYCn5NpKa_CvcKe7_DghLWawnTs7FxxfX183v0IhQY6jS5XJT9Oh1O05EGTXGdb7EthKq5J6bl8GNsoDMg8_DVQAcuKLNw4XJQwfvs2mfNiG0zDjSqWsC5GH0p.epJv805BeceOCJtoBVMX1yz.ol0_utz0lfuD5vIOdAhD7zCMOunv3gJ9s6uYOGf69nDa3vwR9ctzmPsPQiBPuhYfZaerxAabH8tdxtXOIgcZbpVthKP_qfslbCN_mwkp_soqR6RrkmB6YaTLXo_QprsxeNjxJHmsLBbJLNOr427Fc7RPazc0tUBI0pzltsJC_I6pP58ZG4AEUThqAc8dkDEvTpLH0NjIulh82Nm_OXaqAppz4iE.2XAbEZ9I9j_FpD3ZOZ4Fm3ZiIV1wPJyxVfAEhM.imWNzcJhHsKvvtkH6yZz2VC8_p.4vrd4uFMWZ9NDNj.5iYgSOcpa.1hhyJmZOkowaGi2VbmYvYdyXU1Lj5WFABC1tFlWWi31HvDB.xOAb6Wr1gQ_IFzmo1sJ8dL4XX7aCb2qtoiveHNUM6A0uwvQq9eq0rmkyED7pmeNQBN0KpwFkQg74LDpYn9G01Km1VLKAiF_oHA8.CIyHIf9j0aU5WbylpOlo4qQ17lduQigO034VWk4I0ZnAy85v8EXW2ZeW6jiBy9oOMeuMIED.A3yIaLMcyNqOD.JNWSS93RuoMt_CCCa3nJ9FXQ4X0gTbznZe6FRCK03sH3nZP9TMviyCvn_wc.m8Grv_FJtx2sy3z_buQsd8.mFqSlTtughzvohrC_lgtd0mauCNrOm5bRhZHxnIStvjkxv1a4uhBFQZoJmMFpaJRbvrgmqD5tCff.lfAU0cyrIMh1kQIXrCtF1akFQEj2A4zEapVscO_w56OMpeei4Z_uajs0SwbvBULSQH6zXclGpf9npxVkg5SVUlZB1dSEDN0kDOOK5UWnhAb5ikpfKh0vvfGsnkoseJ.7NsZbBbY6yAPRCKRDodwniffDJtMOe_ccxCxB7mHRF.X7aPiEYNdKFZpfr3Prwuu5YOWCO4dI7w3jyNqaKlhiZybZNyAu0Spn.mA8GJIEGAYP.Ub2a01httiPUtShXPyn_MeGjc6SYDhmh1LY5sG1dYiE4kBnIITugZP.x84cc_0OeJ7Qv3hw1Jayb2ihvdFq2a7jkxJT62Np93zu9U1iyDaWQES46QHmV6V8qUPdGkltE2VMe_jaZq0hArSUNBEsskci3HicrYQVy_UNDybWPsmHYcE4NFBr.zDv7GEUmcSN.WjUnWyyM5EugPI69v8XrDMm7slWpheYAD5ryVWixGqbpaXCsngaaJzQ1Eqn24S2f2ldfOfKrsUbWETR8jab2CMlQYehvdIm6LeaTVQLCPfVm7fi4uw8x60xG1o7hce3zQLMU0IrdiUSsat0ACp0_N1bS27j8ojyxNcGg5Kp0dttst..p2cu0qtYHwmIuYuiKAiOaSSdXWCz2bUXF9Kp5XU68y_H6LiHTMdWiZWIDPdrHpPecBfNk0pNLtLi.B1KUxOnutkvOBe2Mw.lebGbhSHMdhI1qDFX0owEbR2p2v2NMWnjSZRJC6fR1N2_X0RjrUGaRFlC6mGPx0K26Uc1c7HQsmmtILWLcxzqwGAl5BdLn6yUJ_wBI5yK2VALD2Ox8wbgWf9w1hZ.zIe.gpAGHt_CUk647iIzz88On48NtZrJacEu3rju1bn2HPyOSAkwnRUfK0xWllsCr6g6qbkgkYWxmD8ONoPcvWNVvASHnlON0yIjigXWTYMbrrLzvfLjX.ixR3ES97L5DCEwst9TdffNGZob8e1csXhMnQkQRMlPgxm26yq0lQRWMxNyh0.VeNWtVt7kIIF3iAldzSCHaSypmARnaZFAdZquFGMbp_2OlFXixzundxXETEhVtDQmj8mc.C6nAquHeH84WdAupxN4HRY.E__r9HjQ"};var cpo = document.createElement('script');cpo.src = '/cdn-cgi/challenge-platform/h/b/orchestrate/chl_page/v1?ray=8eaf35017f2d4073';window._cf_chl_opt.cOgUHash = location.hash === '' && location.href.indexOf('#') !== -1 ? '#' : location.hash;window._cf_chl_opt.cOgUQuery = location.search === '' && location.href.slice(0, location.href.length - window._cf_chl_opt.cOgUHash.length).indexOf('?') !== -1 ? '?' : location.search;if (window.history && window.history.replaceState) {var ogU = location.pathname + window._cf_chl_opt.cOgUQuery + window._cf_chl_opt.cOgUHash;history.replaceState(null, null, "\/dev\/article\/123\/1\/153\/39329\/Mutations-affecting-somite-formation-and?__cf_chl_rt_tk=k0hhzrdWuIIb_FO2.l86jraSCOFldaxh0SGowYt3iSc-1733015526-1.0.1.1-mad4Jq8MDPXdnNaX6865yexwX10OuYDeko0mQCslvmQ" + window._cf_chl_opt.cOgUHash);cpo.onload = function() {history.replaceState(null, null, ogU);}}document.getElementsByTagName('head')[0].appendChild(cpo);}());</script></body></html>