CINXE.COM
Harald zur Hausen - Wikipedia
<!DOCTYPE html> <html class="client-nojs vector-feature-language-in-header-enabled vector-feature-language-in-main-page-header-disabled vector-feature-sticky-header-disabled vector-feature-page-tools-pinned-disabled vector-feature-toc-pinned-clientpref-1 vector-feature-main-menu-pinned-disabled vector-feature-limited-width-clientpref-1 vector-feature-limited-width-content-enabled vector-feature-custom-font-size-clientpref-1 vector-feature-appearance-pinned-clientpref-1 vector-feature-night-mode-enabled skin-theme-clientpref-day vector-toc-available" lang="nl" dir="ltr"> <head> <meta charset="UTF-8"> <title>Harald zur Hausen - Wikipedia</title> <script>(function(){var className="client-js vector-feature-language-in-header-enabled vector-feature-language-in-main-page-header-disabled vector-feature-sticky-header-disabled vector-feature-page-tools-pinned-disabled vector-feature-toc-pinned-clientpref-1 vector-feature-main-menu-pinned-disabled vector-feature-limited-width-clientpref-1 vector-feature-limited-width-content-enabled vector-feature-custom-font-size-clientpref-1 vector-feature-appearance-pinned-clientpref-1 vector-feature-night-mode-enabled skin-theme-clientpref-day vector-toc-available";var cookie=document.cookie.match(/(?:^|; )nlwikimwclientpreferences=([^;]+)/);if(cookie){cookie[1].split('%2C').forEach(function(pref){className=className.replace(new RegExp('(^| )'+pref.replace(/-clientpref-\w+$|[^\w-]+/g,'')+'-clientpref-\\w+( |$)'),'$1'+pref+'$2');});}document.documentElement.className=className;}());RLCONF={"wgBreakFrames":false,"wgSeparatorTransformTable":[",\t.",".\t,"],"wgDigitTransformTable":["",""], "wgDefaultDateFormat":"dmy","wgMonthNames":["","januari","februari","maart","april","mei","juni","juli","augustus","september","oktober","november","december"],"wgRequestId":"ae24d3bc-a087-42ae-a3a6-46715d251caf","wgCanonicalNamespace":"","wgCanonicalSpecialPageName":false,"wgNamespaceNumber":0,"wgPageName":"Harald_zur_Hausen","wgTitle":"Harald zur Hausen","wgCurRevisionId":64787487,"wgRevisionId":64787487,"wgArticleId":1425480,"wgIsArticle":true,"wgIsRedirect":false,"wgAction":"view","wgUserName":null,"wgUserGroups":["*"],"wgCategories":["Duits hoogleraar","Duits medicus","Duits viroloog","Winnaar van de Nobelprijs voor Fysiologie of Geneeskunde","Baarmoederhalskanker","20e-eeuws medicus"],"wgPageViewLanguage":"nl","wgPageContentLanguage":"nl","wgPageContentModel":"wikitext","wgRelevantPageName":"Harald_zur_Hausen","wgRelevantArticleId":1425480,"wgIsProbablyEditable":true,"wgRelevantPageIsProbablyEditable":true,"wgRestrictionEdit":[],"wgRestrictionMove":[],"wgNoticeProject": "wikipedia","wgCiteReferencePreviewsActive":true,"wgMediaViewerOnClick":true,"wgMediaViewerEnabledByDefault":true,"wgPopupsFlags":0,"wgVisualEditor":{"pageLanguageCode":"nl","pageLanguageDir":"ltr","pageVariantFallbacks":"nl"},"wgMFDisplayWikibaseDescriptions":{"search":true,"watchlist":true,"tagline":true,"nearby":true},"wgWMESchemaEditAttemptStepOversample":false,"wgWMEPageLength":6000,"wgRelatedArticlesCompat":[],"wgEditSubmitButtonLabelPublish":true,"wgULSPosition":"interlanguage","wgULSisCompactLinksEnabled":false,"wgVector2022LanguageInHeader":true,"wgULSisLanguageSelectorEmpty":false,"wgWikibaseItemId":"Q75847","wgCheckUserClientHintsHeadersJsApi":["brands","architecture","bitness","fullVersionList","mobile","model","platform","platformVersion"],"GEHomepageSuggestedEditsEnableTopics":true,"wgGETopicsMatchModeEnabled":false,"wgGEStructuredTaskRejectionReasonTextInputEnabled":false,"wgGELevelingUpEnabledForUser":false};RLSTATE={"ext.globalCssJs.user.styles":"ready","site.styles": "ready","user.styles":"ready","ext.globalCssJs.user":"ready","user":"ready","user.options":"loading","ext.cite.styles":"ready","skins.vector.search.codex.styles":"ready","skins.vector.styles":"ready","skins.vector.icons":"ready","jquery.makeCollapsible.styles":"ready","ext.wikimediamessages.styles":"ready","ext.visualEditor.desktopArticleTarget.noscript":"ready","ext.uls.interlanguage":"ready","wikibase.client.init":"ready","ext.wikimediaBadges":"ready"};RLPAGEMODULES=["ext.cite.ux-enhancements","site","mediawiki.page.ready","jquery.makeCollapsible","skins.vector.js","ext.centralNotice.geoIP","ext.centralNotice.startUp","ext.gadget.Direct-link-to-Commons","ext.gadget.ProtectionTemplates","ext.gadget.InterProjectLinks","ext.gadget.hoofdbetekenis-titelwijziging","ext.gadget.switcher","ext.gadget.OpenStreetMapFrame","ext.gadget.subpages","ext.urlShortener.toolbar","ext.centralauth.centralautologin","mmv.bootstrap","ext.popups","ext.visualEditor.desktopArticleTarget.init", "ext.visualEditor.targetLoader","ext.echo.centralauth","ext.eventLogging","ext.wikimediaEvents","ext.navigationTiming","ext.uls.interface","ext.cx.eventlogging.campaigns","ext.cx.uls.quick.actions","wikibase.client.vector-2022","ext.checkUser.clientHints","ext.growthExperiments.SuggestedEditSession"];</script> <script>(RLQ=window.RLQ||[]).push(function(){mw.loader.impl(function(){return["user.options@12s5i",function($,jQuery,require,module){mw.user.tokens.set({"patrolToken":"+\\","watchToken":"+\\","csrfToken":"+\\"}); }];});});</script> <link rel="stylesheet" href="/w/load.php?lang=nl&modules=ext.cite.styles%7Cext.uls.interlanguage%7Cext.visualEditor.desktopArticleTarget.noscript%7Cext.wikimediaBadges%7Cext.wikimediamessages.styles%7Cjquery.makeCollapsible.styles%7Cskins.vector.icons%2Cstyles%7Cskins.vector.search.codex.styles%7Cwikibase.client.init&only=styles&skin=vector-2022"> <script async="" src="/w/load.php?lang=nl&modules=startup&only=scripts&raw=1&skin=vector-2022"></script> <meta name="ResourceLoaderDynamicStyles" content=""> <link rel="stylesheet" href="/w/load.php?lang=nl&modules=site.styles&only=styles&skin=vector-2022"> <meta name="generator" content="MediaWiki 1.44.0-wmf.6"> <meta name="referrer" content="origin"> <meta name="referrer" content="origin-when-cross-origin"> <meta name="robots" content="max-image-preview:standard"> <meta name="format-detection" content="telephone=no"> <meta property="og:image" content="https://upload.wikimedia.org/wikipedia/commons/thumb/9/99/Harald_zur_Hausen_03.jpg/1200px-Harald_zur_Hausen_03.jpg"> <meta property="og:image:width" content="1200"> <meta property="og:image:height" content="1552"> <meta property="og:image" content="https://upload.wikimedia.org/wikipedia/commons/thumb/9/99/Harald_zur_Hausen_03.jpg/800px-Harald_zur_Hausen_03.jpg"> <meta property="og:image:width" content="800"> <meta property="og:image:height" content="1035"> <meta property="og:image" content="https://upload.wikimedia.org/wikipedia/commons/thumb/9/99/Harald_zur_Hausen_03.jpg/640px-Harald_zur_Hausen_03.jpg"> <meta property="og:image:width" content="640"> <meta property="og:image:height" content="828"> <meta name="viewport" content="width=1120"> <meta property="og:title" content="Harald zur Hausen - Wikipedia"> <meta property="og:type" content="website"> <link rel="preconnect" href="//upload.wikimedia.org"> <link rel="alternate" media="only screen and (max-width: 640px)" href="//nl.m.wikipedia.org/wiki/Harald_zur_Hausen"> <link rel="alternate" type="application/x-wiki" title="Bewerken" href="/w/index.php?title=Harald_zur_Hausen&action=edit"> <link rel="apple-touch-icon" href="/static/apple-touch/wikipedia.png"> <link rel="icon" href="/static/favicon/wikipedia.ico"> <link rel="search" type="application/opensearchdescription+xml" href="/w/rest.php/v1/search" title="Wikipedia (nl)"> <link rel="EditURI" type="application/rsd+xml" href="//nl.wikipedia.org/w/api.php?action=rsd"> <link rel="canonical" href="https://nl.wikipedia.org/wiki/Harald_zur_Hausen"> <link rel="license" href="https://creativecommons.org/licenses/by-sa/4.0/deed.nl"> <link rel="alternate" type="application/atom+xml" title="Wikipedia Atom-feed" href="/w/index.php?title=Speciaal:RecenteWijzigingen&feed=atom"> <link rel="dns-prefetch" href="//meta.wikimedia.org" /> <link rel="dns-prefetch" href="login.wikimedia.org"> </head> <body class="skin--responsive skin-vector skin-vector-search-vue mediawiki ltr sitedir-ltr mw-hide-empty-elt ns-0 ns-subject mw-editable page-Harald_zur_Hausen rootpage-Harald_zur_Hausen skin-vector-2022 action-view"><a class="mw-jump-link" href="#bodyContent">Naar inhoud springen</a> <div class="vector-header-container"> <header class="vector-header mw-header"> <div class="vector-header-start"> <nav class="vector-main-menu-landmark" aria-label="Site"> <div id="vector-main-menu-dropdown" class="vector-dropdown vector-main-menu-dropdown vector-button-flush-left vector-button-flush-right" > <input type="checkbox" id="vector-main-menu-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-main-menu-dropdown" class="vector-dropdown-checkbox " aria-label="Hoofdmenu" > <label id="vector-main-menu-dropdown-label" for="vector-main-menu-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-menu mw-ui-icon-wikimedia-menu"></span> <span class="vector-dropdown-label-text">Hoofdmenu</span> </label> <div class="vector-dropdown-content"> <div id="vector-main-menu-unpinned-container" class="vector-unpinned-container"> <div id="vector-main-menu" class="vector-main-menu vector-pinnable-element"> <div class="vector-pinnable-header vector-main-menu-pinnable-header vector-pinnable-header-unpinned" data-feature-name="main-menu-pinned" data-pinnable-element-id="vector-main-menu" data-pinned-container-id="vector-main-menu-pinned-container" data-unpinned-container-id="vector-main-menu-unpinned-container" > <div class="vector-pinnable-header-label">Hoofdmenu</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-main-menu.pin">naar zijbalk verplaatsen</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-main-menu.unpin">verbergen</button> </div> <div id="p-navigation" class="vector-menu mw-portlet mw-portlet-navigation" > <div class="vector-menu-heading"> Navigatie </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="n-mainpage" class="mw-list-item"><a href="/wiki/Hoofdpagina" title="Naar de hoofdpagina gaan [z]" accesskey="z"><span>Hoofdpagina</span></a></li><li id="n-zoekartikel" class="mw-list-item"><a href="/wiki/Portaal:Navigatie"><span>Vind een artikel</span></a></li><li id="n-today" class="mw-list-item"><a href="/wiki/11_december"><span>Vandaag</span></a></li><li id="n-Etalage" class="mw-list-item"><a href="/wiki/Wikipedia:Etalage"><span>Etalage</span></a></li><li id="n-categories" class="mw-list-item"><a href="/wiki/Categorie:Alles"><span>Categorieën</span></a></li><li id="n-recentchanges" class="mw-list-item"><a href="/wiki/Speciaal:RecenteWijzigingen" title="Een lijst met recente wijzigingen in deze wiki. [r]" accesskey="r"><span>Recente wijzigingen</span></a></li><li id="n-newpages" class="mw-list-item"><a href="/wiki/Speciaal:NieuwePaginas"><span>Nieuwe artikelen</span></a></li><li id="n-randompage" class="mw-list-item"><a href="/wiki/Speciaal:Willekeurig" title="Een willekeurige pagina bekijken [x]" accesskey="x"><span>Willekeurige pagina</span></a></li> </ul> </div> </div> <div id="p-navigation2" class="vector-menu mw-portlet mw-portlet-navigation2" > <div class="vector-menu-heading"> Informatie </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="n-portal" class="mw-list-item"><a href="/wiki/Portaal:Gebruikersportaal" title="Informatie over het project: wat u kunt doen, waar u dingen kunt vinden"><span>Gebruikersportaal</span></a></li><li id="n-Snelcursus" class="mw-list-item"><a href="/wiki/Wikipedia:Snelcursus"><span>Snelcursus</span></a></li><li id="n-help" class="mw-list-item"><a href="/wiki/Portaal:Hulp_en_beheer" title="Hulpinformatie over deze wiki"><span>Hulp en contact</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> <a href="/wiki/Hoofdpagina" class="mw-logo"> <img class="mw-logo-icon" src="/static/images/icons/wikipedia.png" alt="" aria-hidden="true" height="50" width="50"> <span class="mw-logo-container skin-invert"> <img class="mw-logo-wordmark" alt="Wikipedia" src="/static/images/mobile/copyright/wikipedia-wordmark-en.svg" style="width: 7.5em; height: 1.125em;"> <img class="mw-logo-tagline" alt="de vrije encyclopedie" src="/static/images/mobile/copyright/wikipedia-tagline-nl.svg" width="120" height="13" style="width: 7.5em; height: 0.8125em;"> </span> </a> </div> <div class="vector-header-end"> <div id="p-search" role="search" class="vector-search-box-vue vector-search-box-collapses vector-search-box-show-thumbnail vector-search-box-auto-expand-width vector-search-box"> <a href="/wiki/Speciaal:Zoeken" class="cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only search-toggle" title="Doorzoek Wikipedia [f]" accesskey="f"><span class="vector-icon mw-ui-icon-search mw-ui-icon-wikimedia-search"></span> <span>Zoeken</span> </a> <div class="vector-typeahead-search-container"> <div class="cdx-typeahead-search cdx-typeahead-search--show-thumbnail cdx-typeahead-search--auto-expand-width"> <form action="/w/index.php" id="searchform" class="cdx-search-input cdx-search-input--has-end-button"> <div id="simpleSearch" class="cdx-search-input__input-wrapper" data-search-loc="header-moved"> <div class="cdx-text-input cdx-text-input--has-start-icon"> <input class="cdx-text-input__input" type="search" name="search" placeholder="Doorzoek Wikipedia" aria-label="Doorzoek Wikipedia" autocapitalize="sentences" title="Doorzoek Wikipedia [f]" accesskey="f" id="searchInput" > <span class="cdx-text-input__icon cdx-text-input__start-icon"></span> </div> <input type="hidden" name="title" value="Speciaal:Zoeken"> </div> <button class="cdx-button cdx-search-input__end-button">Zoeken</button> </form> </div> </div> </div> <nav class="vector-user-links vector-user-links-wide" aria-label="Persoonlijke hulpmiddelen"> <div class="vector-user-links-main"> <div id="p-vector-user-menu-preferences" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <div id="p-vector-user-menu-userpage" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <nav class="vector-appearance-landmark" aria-label="Uiterlijk"> <div id="vector-appearance-dropdown" class="vector-dropdown " title="De lettergrootte, breedte en kleur van de pagina wijzigen" > <input type="checkbox" id="vector-appearance-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-appearance-dropdown" class="vector-dropdown-checkbox " aria-label="Uiterlijk" > <label id="vector-appearance-dropdown-label" for="vector-appearance-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-appearance mw-ui-icon-wikimedia-appearance"></span> <span class="vector-dropdown-label-text">Uiterlijk</span> </label> <div class="vector-dropdown-content"> <div id="vector-appearance-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <div id="p-vector-user-menu-notifications" class="vector-menu mw-portlet emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> <div id="p-vector-user-menu-overflow" class="vector-menu mw-portlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-sitesupport-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="https://donate.wikimedia.org/?wmf_source=donate&wmf_medium=sidebar&wmf_campaign=nl.wikipedia.org&uselang=nl" class=""><span>Doneren</span></a> </li> <li id="pt-createaccount-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="/w/index.php?title=Speciaal:GebruikerAanmaken&returnto=Harald+zur+Hausen" title="Registreer u vooral en meld u aan. Dit is echter niet verplicht." class=""><span>Account aanmaken</span></a> </li> <li id="pt-login-2" class="user-links-collapsible-item mw-list-item user-links-collapsible-item"><a data-mw="interface" href="/w/index.php?title=Speciaal:Aanmelden&returnto=Harald+zur+Hausen" title="U wordt van harte uitgenodigd om aan te melden, maar dit is niet verplicht [o]" accesskey="o" class=""><span>Aanmelden</span></a> </li> </ul> </div> </div> </div> <div id="vector-user-links-dropdown" class="vector-dropdown vector-user-menu vector-button-flush-right vector-user-menu-logged-out" title="Meer opties" > <input type="checkbox" id="vector-user-links-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-user-links-dropdown" class="vector-dropdown-checkbox " aria-label="Persoonlijke hulpmiddelen" > <label id="vector-user-links-dropdown-label" for="vector-user-links-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-ellipsis mw-ui-icon-wikimedia-ellipsis"></span> <span class="vector-dropdown-label-text">Persoonlijke hulpmiddelen</span> </label> <div class="vector-dropdown-content"> <div id="p-personal" class="vector-menu mw-portlet mw-portlet-personal user-links-collapsible-item" title="Gebruikersmenu" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-sitesupport" class="user-links-collapsible-item mw-list-item"><a href="https://donate.wikimedia.org/?wmf_source=donate&wmf_medium=sidebar&wmf_campaign=nl.wikipedia.org&uselang=nl"><span>Doneren</span></a></li><li id="pt-createaccount" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=Speciaal:GebruikerAanmaken&returnto=Harald+zur+Hausen" title="Registreer u vooral en meld u aan. Dit is echter niet verplicht."><span class="vector-icon mw-ui-icon-userAdd mw-ui-icon-wikimedia-userAdd"></span> <span>Account aanmaken</span></a></li><li id="pt-login" class="user-links-collapsible-item mw-list-item"><a href="/w/index.php?title=Speciaal:Aanmelden&returnto=Harald+zur+Hausen" title="U wordt van harte uitgenodigd om aan te melden, maar dit is niet verplicht [o]" accesskey="o"><span class="vector-icon mw-ui-icon-logIn mw-ui-icon-wikimedia-logIn"></span> <span>Aanmelden</span></a></li> </ul> </div> </div> <div id="p-user-menu-anon-editor" class="vector-menu mw-portlet mw-portlet-user-menu-anon-editor" > <div class="vector-menu-heading"> Pagina's voor uitgelogde redacteuren <a href="/wiki/Help:Inleiding" aria-label="Meer leren over bewerken"><span>meer lezen</span></a> </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="pt-anoncontribs" class="mw-list-item"><a href="/wiki/Speciaal:MijnBijdragen" title="Bijdragen IP-adres [y]" accesskey="y"><span>Bijdragen</span></a></li><li id="pt-anontalk" class="mw-list-item"><a href="/wiki/Speciaal:MijnOverleg" title="Overlegpagina van de anonieme gebruiker van dit IP-adres [n]" accesskey="n"><span>Overleg</span></a></li> </ul> </div> </div> </div> </div> </nav> </div> </header> </div> <div class="mw-page-container"> <div class="mw-page-container-inner"> <div class="vector-sitenotice-container"> <div id="siteNotice"><!-- CentralNotice --></div> </div> <div class="vector-column-start"> <div class="vector-main-menu-container"> <div id="mw-navigation"> <nav id="mw-panel" class="vector-main-menu-landmark" aria-label="Site"> <div id="vector-main-menu-pinned-container" class="vector-pinned-container"> </div> </nav> </div> </div> <div class="vector-sticky-pinned-container"> <nav id="mw-panel-toc" aria-label="Inhoud" data-event-name="ui.sidebar-toc" class="mw-table-of-contents-container vector-toc-landmark"> <div id="vector-toc-pinned-container" class="vector-pinned-container"> <div id="vector-toc" class="vector-toc vector-pinnable-element"> <div class="vector-pinnable-header vector-toc-pinnable-header vector-pinnable-header-pinned" data-feature-name="toc-pinned" data-pinnable-element-id="vector-toc" > <h2 class="vector-pinnable-header-label">Inhoud</h2> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-toc.pin">naar zijbalk verplaatsen</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-toc.unpin">verbergen</button> </div> <ul class="vector-toc-contents" id="mw-panel-toc-list"> <li id="toc-mw-content-text" class="vector-toc-list-item vector-toc-level-1"> <a href="#" class="vector-toc-link"> <div class="vector-toc-text">Top</div> </a> </li> <li id="toc-Biografie" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Biografie"> <div class="vector-toc-text"> <span class="vector-toc-numb">1</span> <span>Biografie</span> </div> </a> <ul id="toc-Biografie-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Werk" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Werk"> <div class="vector-toc-text"> <span class="vector-toc-numb">2</span> <span>Werk</span> </div> </a> <ul id="toc-Werk-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Inhoud" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Inhoudsopgave omschakelen" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Inhoudsopgave omschakelen</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Harald zur Hausen</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Ga naar een artikel in een andere taal. Beschikbaar in 65 talen" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-65" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">65 talen</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-als mw-list-item"><a href="https://als.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Zwitserduits" lang="gsw" hreflang="gsw" data-title="Harald zur Hausen" data-language-autonym="Alemannisch" data-language-local-name="Zwitserduits" class="interlanguage-link-target"><span>Alemannisch</span></a></li><li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D9%87%D8%A7%D8%B1%D8%A7%D9%84%D8%AF_%D8%AA%D8%B3%D9%88%D8%B1_%D9%87%D8%A7%D9%88%D8%B2%D9%86" title="هارالد تسور هاوزن – Arabisch" lang="ar" hreflang="ar" data-title="هارالد تسور هاوزن" data-language-autonym="العربية" data-language-local-name="Arabisch" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D9%87%D8%A7%D8%B1%D8%A7%D9%84%D8%AF_%D8%AA%D8%B3%D9%88%D8%B1_%D9%87%D8%A7%D9%88%D8%B2%D9%86" title="هارالد تسور هاوزن – Egyptisch Arabisch" lang="arz" hreflang="arz" data-title="هارالد تسور هاوزن" data-language-autonym="مصرى" data-language-local-name="Egyptisch Arabisch" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-ast mw-list-item"><a href="https://ast.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Asturisch" lang="ast" hreflang="ast" data-title="Harald zur Hausen" data-language-autonym="Asturianu" data-language-local-name="Asturisch" class="interlanguage-link-target"><span>Asturianu</span></a></li><li class="interlanguage-link interwiki-az mw-list-item"><a href="https://az.wikipedia.org/wiki/Harald_tsur_Hauzen" title="Harald tsur Hauzen – Azerbeidzjaans" lang="az" hreflang="az" data-title="Harald tsur Hauzen" data-language-autonym="Azərbaycanca" data-language-local-name="Azerbeidzjaans" class="interlanguage-link-target"><span>Azərbaycanca</span></a></li><li class="interlanguage-link interwiki-azb mw-list-item"><a href="https://azb.wikipedia.org/wiki/%D9%87%D8%A7%D8%B1%D8%A7%D9%84%D8%AF_%D8%AA%D8%B3%D9%88%D8%B1_%D9%87%D8%A7%D9%88%D8%B2%D9%86" title="هارالد تسور هاوزن – South Azerbaijani" lang="azb" hreflang="azb" data-title="هارالد تسور هاوزن" data-language-autonym="تۆرکجه" data-language-local-name="South Azerbaijani" class="interlanguage-link-target"><span>تۆرکجه</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%A5%D0%B0%D1%80%D0%B0%D0%BB%D1%8C%D0%B4_%D1%86%D1%83%D1%80_%D0%A5%D0%B0%D1%9E%D0%B7%D0%B5%D0%BD" title="Харальд цур Хаўзен – Belarussisch" lang="be" hreflang="be" data-title="Харальд цур Хаўзен" data-language-autonym="Беларуская" data-language-local-name="Belarussisch" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%A5%D0%B0%D1%80%D0%B0%D0%BB%D0%B4_%D1%86%D1%83%D1%80_%D0%A5%D0%B0%D1%83%D0%B7%D0%B5%D0%BD" title="Харалд цур Хаузен – Bulgaars" lang="bg" hreflang="bg" data-title="Харалд цур Хаузен" data-language-autonym="Български" data-language-local-name="Bulgaars" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-bn mw-list-item"><a href="https://bn.wikipedia.org/wiki/%E0%A6%B9%E0%A6%BE%E0%A6%B0%E0%A6%BE%E0%A6%B2%E0%A7%8D%E0%A6%A1_%E0%A7%8E%E0%A6%B8%E0%A7%81%E0%A6%B0_%E0%A6%B9%E0%A6%BE%E0%A6%89%E0%A6%9C%E0%A7%87%E0%A6%A8" title="হারাল্ড ৎসুর হাউজেন – Bengaals" lang="bn" hreflang="bn" data-title="হারাল্ড ৎসুর হাউজেন" data-language-autonym="বাংলা" data-language-local-name="Bengaals" class="interlanguage-link-target"><span>বাংলা</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Catalaans" lang="ca" hreflang="ca" data-title="Harald zur Hausen" data-language-autonym="Català" data-language-local-name="Catalaans" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-ckb mw-list-item"><a href="https://ckb.wikipedia.org/wiki/%D9%87%D8%A7%D8%B1%D8%A7%DA%B5%D8%AF_%D9%87%D8%A7%D9%88%D8%B3%D9%86" title="هاراڵد هاوسن – Soranî" lang="ckb" hreflang="ckb" data-title="هاراڵد هاوسن" data-language-autonym="کوردی" data-language-local-name="Soranî" class="interlanguage-link-target"><span>کوردی</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Tsjechisch" lang="cs" hreflang="cs" data-title="Harald zur Hausen" data-language-autonym="Čeština" data-language-local-name="Tsjechisch" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Deens" lang="da" hreflang="da" data-title="Harald zur Hausen" data-language-autonym="Dansk" data-language-local-name="Deens" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Duits" lang="de" hreflang="de" data-title="Harald zur Hausen" data-language-autonym="Deutsch" data-language-local-name="Duits" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-en mw-list-item"><a href="https://en.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Engels" lang="en" hreflang="en" data-title="Harald zur Hausen" data-language-autonym="English" data-language-local-name="Engels" class="interlanguage-link-target"><span>English</span></a></li><li class="interlanguage-link interwiki-eo mw-list-item"><a href="https://eo.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Esperanto" lang="eo" hreflang="eo" data-title="Harald zur Hausen" data-language-autonym="Esperanto" data-language-local-name="Esperanto" class="interlanguage-link-target"><span>Esperanto</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Spaans" lang="es" hreflang="es" data-title="Harald zur Hausen" data-language-autonym="Español" data-language-local-name="Spaans" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Estisch" lang="et" hreflang="et" data-title="Harald zur Hausen" data-language-autonym="Eesti" data-language-local-name="Estisch" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Baskisch" lang="eu" hreflang="eu" data-title="Harald zur Hausen" data-language-autonym="Euskara" data-language-local-name="Baskisch" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D9%87%D8%A7%D8%B1%D8%A7%D9%84%D8%AF_%D8%AA%D8%B3%D9%88%D8%B1_%D9%87%D8%A7%D9%88%D8%B2%D9%86" title="هارالد تسور هاوزن – Perzisch" lang="fa" hreflang="fa" data-title="هارالد تسور هاوزن" data-language-autonym="فارسی" data-language-local-name="Perzisch" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Fins" lang="fi" hreflang="fi" data-title="Harald zur Hausen" data-language-autonym="Suomi" data-language-local-name="Fins" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Frans" lang="fr" hreflang="fr" data-title="Harald zur Hausen" data-language-autonym="Français" data-language-local-name="Frans" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Iers" lang="ga" hreflang="ga" data-title="Harald zur Hausen" data-language-autonym="Gaeilge" data-language-local-name="Iers" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%94%D7%A8%D7%9C%D7%93_%D7%A6%D7%95%D7%A8_%D7%94%D7%90%D7%95%D7%96%D7%9F" title="הרלד צור האוזן – Hebreeuws" lang="he" hreflang="he" data-title="הרלד צור האוזן" data-language-autonym="עברית" data-language-local-name="Hebreeuws" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-hr mw-list-item"><a href="https://hr.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Kroatisch" lang="hr" hreflang="hr" data-title="Harald zur Hausen" data-language-autonym="Hrvatski" data-language-local-name="Kroatisch" class="interlanguage-link-target"><span>Hrvatski</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Hongaars" lang="hu" hreflang="hu" data-title="Harald zur Hausen" data-language-autonym="Magyar" data-language-local-name="Hongaars" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-hy mw-list-item"><a href="https://hy.wikipedia.org/wiki/%D5%80%D5%A1%D6%80%D5%A1%D5%AC%D5%A4_%D4%B6%D5%B8%D6%82%D6%80_%D5%80%D5%A1%D5%B8%D6%82%D5%A6%D5%A5%D5%B6" title="Հարալդ Զուր Հաուզեն – Armeens" lang="hy" hreflang="hy" data-title="Հարալդ Զուր Հաուզեն" data-language-autonym="Հայերեն" data-language-local-name="Armeens" class="interlanguage-link-target"><span>Հայերեն</span></a></li><li class="interlanguage-link interwiki-iba mw-list-item"><a href="https://iba.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Iban" lang="iba" hreflang="iba" data-title="Harald zur Hausen" data-language-autonym="Jaku Iban" data-language-local-name="Iban" class="interlanguage-link-target"><span>Jaku Iban</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Indonesisch" lang="id" hreflang="id" data-title="Harald zur Hausen" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesisch" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Ido" lang="io" hreflang="io" data-title="Harald zur Hausen" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Italiaans" lang="it" hreflang="it" data-title="Harald zur Hausen" data-language-autonym="Italiano" data-language-local-name="Italiaans" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%83%8F%E3%83%A9%E3%83%AB%E3%83%89%E3%83%BB%E3%83%84%E3%82%A2%E3%83%BB%E3%83%8F%E3%82%A6%E3%82%BC%E3%83%B3" title="ハラルド・ツア・ハウゼン – Japans" lang="ja" hreflang="ja" data-title="ハラルド・ツア・ハウゼン" data-language-autonym="日本語" data-language-local-name="Japans" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%A5%D0%B0%D1%80%D0%B0%D0%BB%D1%8C%D0%B4_%D1%86%D1%83%D1%80_%D0%A5%D0%B0%D1%83%D0%B7%D0%B5%D0%BD" title="Харальд цур Хаузен – Kazachs" lang="kk" hreflang="kk" data-title="Харальд цур Хаузен" data-language-autonym="Қазақша" data-language-local-name="Kazachs" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%ED%95%98%EB%9E%84%ED%8A%B8_%EC%B6%94%EC%96%B4_%ED%95%98%EC%9A%B0%EC%A0%A0" title="하랄트 추어 하우젠 – Koreaans" lang="ko" hreflang="ko" data-title="하랄트 추어 하우젠" data-language-autonym="한국어" data-language-local-name="Koreaans" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Haroldus_zur_Hausen" title="Haroldus zur Hausen – Latijn" lang="la" hreflang="la" data-title="Haroldus zur Hausen" data-language-autonym="Latina" data-language-local-name="Latijn" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-lb mw-list-item"><a href="https://lb.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Luxemburgs" lang="lb" hreflang="lb" data-title="Harald zur Hausen" data-language-autonym="Lëtzebuergesch" data-language-local-name="Luxemburgs" class="interlanguage-link-target"><span>Lëtzebuergesch</span></a></li><li class="interlanguage-link interwiki-lt mw-list-item"><a href="https://lt.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Litouws" lang="lt" hreflang="lt" data-title="Harald zur Hausen" data-language-autonym="Lietuvių" data-language-local-name="Litouws" class="interlanguage-link-target"><span>Lietuvių</span></a></li><li class="interlanguage-link interwiki-lv mw-list-item"><a href="https://lv.wikipedia.org/wiki/Haralds_cur_Hauzens" title="Haralds cur Hauzens – Lets" lang="lv" hreflang="lv" data-title="Haralds cur Hauzens" data-language-autonym="Latviešu" data-language-local-name="Lets" class="interlanguage-link-target"><span>Latviešu</span></a></li><li class="interlanguage-link interwiki-mr mw-list-item"><a href="https://mr.wikipedia.org/wiki/%E0%A4%B9%E0%A4%B0%E0%A4%BE%E0%A4%B2%E0%A5%8D%E0%A4%A1_%E0%A4%9D%E0%A5%81%E0%A4%85%E0%A4%B0_%E0%A4%B9%E0%A4%BE%E0%A4%89%E0%A4%B8%E0%A4%A8" title="हराल्ड झुअर हाउसन – Marathi" lang="mr" hreflang="mr" data-title="हराल्ड झुअर हाउसन" data-language-autonym="मराठी" data-language-local-name="Marathi" class="interlanguage-link-target"><span>मराठी</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Maleis" lang="ms" hreflang="ms" data-title="Harald zur Hausen" data-language-autonym="Bahasa Melayu" data-language-local-name="Maleis" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-mzn mw-list-item"><a href="https://mzn.wikipedia.org/wiki/%D9%87%D8%A7%D8%B1%D8%A7%D9%84%D8%AF_%D8%AA%D8%B3%D9%88%D8%B1_%D9%87%D8%A7%D9%88%D8%B2%D9%86" title="هارالد تسور هاوزن – Mazanderani" lang="mzn" hreflang="mzn" data-title="هارالد تسور هاوزن" data-language-autonym="مازِرونی" data-language-local-name="Mazanderani" class="interlanguage-link-target"><span>مازِرونی</span></a></li><li class="interlanguage-link interwiki-nds mw-list-item"><a href="https://nds.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Nedersaksisch" lang="nds" hreflang="nds" data-title="Harald zur Hausen" data-language-autonym="Plattdüütsch" data-language-local-name="Nedersaksisch" class="interlanguage-link-target"><span>Plattdüütsch</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Noors - Bokmål" lang="nb" hreflang="nb" data-title="Harald zur Hausen" data-language-autonym="Norsk bokmål" data-language-local-name="Noors - Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Occitaans" lang="oc" hreflang="oc" data-title="Harald zur Hausen" data-language-autonym="Occitan" data-language-local-name="Occitaans" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Pools" lang="pl" hreflang="pl" data-title="Harald zur Hausen" data-language-autonym="Polski" data-language-local-name="Pools" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pnb mw-list-item"><a href="https://pnb.wikipedia.org/wiki/%DB%81%DB%8C%D8%B1%D8%A7%D9%84%DA%88_%D8%B2%D8%B1_%DB%81%D8%A7%D9%88%D8%B2%D9%86" title="ہیرالڈ زر ہاوزن – Western Punjabi" lang="pnb" hreflang="pnb" data-title="ہیرالڈ زر ہاوزن" data-language-autonym="پنجابی" data-language-local-name="Western Punjabi" class="interlanguage-link-target"><span>پنجابی</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Portugees" lang="pt" hreflang="pt" data-title="Harald zur Hausen" data-language-autonym="Português" data-language-local-name="Portugees" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Roemeens" lang="ro" hreflang="ro" data-title="Harald zur Hausen" data-language-autonym="Română" data-language-local-name="Roemeens" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%A5%D0%B0%D1%83%D0%B7%D0%B5%D0%BD,_%D0%A5%D0%B0%D1%80%D0%B0%D0%BB%D1%8C%D0%B4_%D1%86%D1%83%D1%80" title="Хаузен, Харальд цур – Russisch" lang="ru" hreflang="ru" data-title="Хаузен, Харальд цур" data-language-autonym="Русский" data-language-local-name="Russisch" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Servo-Kroatisch" lang="sh" hreflang="sh" data-title="Harald zur Hausen" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="Servo-Kroatisch" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Simple English" lang="en-simple" hreflang="en-simple" data-title="Harald zur Hausen" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sk mw-list-item"><a href="https://sk.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Slowaaks" lang="sk" hreflang="sk" data-title="Harald zur Hausen" data-language-autonym="Slovenčina" data-language-local-name="Slowaaks" class="interlanguage-link-target"><span>Slovenčina</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Sloveens" lang="sl" hreflang="sl" data-title="Harald zur Hausen" data-language-autonym="Slovenščina" data-language-local-name="Sloveens" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%A5%D0%B0%D1%80%D0%B0%D0%BB%D0%B4_%D1%86%D1%83%D1%80_%D0%A5%D0%B0%D1%83%D0%B7%D0%B5%D0%BD" title="Харалд цур Хаузен – Servisch" lang="sr" hreflang="sr" data-title="Харалд цур Хаузен" data-language-autonym="Српски / srpski" data-language-local-name="Servisch" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Zweeds" lang="sv" hreflang="sv" data-title="Harald zur Hausen" data-language-autonym="Svenska" data-language-local-name="Zweeds" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-sw mw-list-item"><a href="https://sw.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Swahili" lang="sw" hreflang="sw" data-title="Harald zur Hausen" data-language-autonym="Kiswahili" data-language-local-name="Swahili" class="interlanguage-link-target"><span>Kiswahili</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%B9%E0%AF%86%E0%AE%B0%E0%AE%BE%E0%AE%B2%E0%AF%8D%E0%AE%9F%E0%AF%81_%E0%AE%9A%E0%AF%82%E0%AE%B0%E0%AF%8D_%E0%AE%B9%E0%AE%BE%E0%AE%9A%E0%AF%86%E0%AE%A9%E0%AF%8D" title="ஹெரால்டு சூர் ஹாசென் – Tamil" lang="ta" hreflang="ta" data-title="ஹெரால்டு சூர் ஹாசென்" data-language-autonym="தமிழ்" data-language-local-name="Tamil" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Turks" lang="tr" hreflang="tr" data-title="Harald zur Hausen" data-language-autonym="Türkçe" data-language-local-name="Turks" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-tt mw-list-item"><a href="https://tt.wikipedia.org/wiki/%D2%BA%D0%B0%D1%80%D0%B0%D0%BB%D1%8C%D0%B4_%D1%86%D1%83%D1%80_%D2%BA%D0%B0%D1%83%D0%B7%D0%B5%D0%BD" title="Һаральд цур Һаузен – Tataars" lang="tt" hreflang="tt" data-title="Һаральд цур Һаузен" data-language-autonym="Татарча / tatarça" data-language-local-name="Tataars" class="interlanguage-link-target"><span>Татарча / tatarça</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%93%D0%B0%D1%80%D0%B0%D0%BB%D1%8C%D0%B4_%D1%86%D1%83%D1%80_%D0%93%D0%B0%D1%83%D0%B7%D0%B5%D0%BD" title="Гаральд цур Гаузен – Oekraïens" lang="uk" hreflang="uk" data-title="Гаральд цур Гаузен" data-language-autonym="Українська" data-language-local-name="Oekraïens" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%DB%81%DB%8C%D8%B1%D8%A7%D9%84%DA%88_%D8%B2%D8%B1_%DB%81%D8%A7%D9%88%D8%B3%D9%86" title="ہیرالڈ زر ہاوسن – Urdu" lang="ur" hreflang="ur" data-title="ہیرالڈ زر ہاوسن" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Vietnamees" lang="vi" hreflang="vi" data-title="Harald zur Hausen" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamees" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-wuu mw-list-item"><a href="https://wuu.wikipedia.org/wiki/%E5%93%88%E6%8B%89%E5%B0%94%E5%BE%B7%C2%B7%E6%A5%9A%E5%B0%94%C2%B7%E8%B1%AA%E6%A3%AE" title="哈拉尔德·楚尔·豪森 – Wuyu" lang="wuu" hreflang="wuu" data-title="哈拉尔德·楚尔·豪森" data-language-autonym="吴语" data-language-local-name="Wuyu" class="interlanguage-link-target"><span>吴语</span></a></li><li class="interlanguage-link interwiki-yo mw-list-item"><a href="https://yo.wikipedia.org/wiki/Harald_zur_Hausen" title="Harald zur Hausen – Yoruba" lang="yo" hreflang="yo" data-title="Harald zur Hausen" data-language-autonym="Yorùbá" data-language-local-name="Yoruba" class="interlanguage-link-target"><span>Yorùbá</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E5%93%88%E6%8B%89%E5%B0%94%E5%BE%B7%C2%B7%E6%A5%9A%E5%B0%94%C2%B7%E8%B1%AA%E6%A3%AE" title="哈拉尔德·楚尔·豪森 – Chinees" lang="zh" hreflang="zh" data-title="哈拉尔德·楚尔·豪森" data-language-autonym="中文" data-language-local-name="Chinees" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q75847#sitelinks-wikipedia" title="Taalkoppelingen bewerken" class="wbc-editpage">Koppelingen bewerken</a></span></div> </div> </div> </div> </header> <div class="vector-page-toolbar"> <div class="vector-page-toolbar-container"> <div id="left-navigation"> <nav aria-label="Naamruimten"> <div id="p-associated-pages" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-associated-pages" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-nstab-main" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Harald_zur_Hausen" title="Inhoudspagina bekijken [c]" accesskey="c"><span>Artikel</span></a></li><li id="ca-talk" class="vector-tab-noicon mw-list-item"><a href="/wiki/Overleg:Harald_zur_Hausen" rel="discussion" title="Overleg over deze pagina [t]" accesskey="t"><span>Overleg</span></a></li> </ul> </div> </div> <div id="vector-variants-dropdown" class="vector-dropdown emptyPortlet" > <input type="checkbox" id="vector-variants-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-variants-dropdown" class="vector-dropdown-checkbox " aria-label="Taalvariant wijzigen" > <label id="vector-variants-dropdown-label" for="vector-variants-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">Nederlands</span> </label> <div class="vector-dropdown-content"> <div id="p-variants" class="vector-menu mw-portlet mw-portlet-variants emptyPortlet" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> </ul> </div> </div> </div> </div> </nav> </div> <div id="right-navigation" class="vector-collapsible"> <nav aria-label="Weergaven"> <div id="p-views" class="vector-menu vector-menu-tabs mw-portlet mw-portlet-views" > <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-view" class="selected vector-tab-noicon mw-list-item"><a href="/wiki/Harald_zur_Hausen"><span>Lezen</span></a></li><li id="ca-ve-edit" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Harald_zur_Hausen&veaction=edit" title="Deze pagina bewerken [v]" accesskey="v"><span>Bewerken</span></a></li><li id="ca-edit" class="collapsible vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Harald_zur_Hausen&action=edit" title="Broncode van deze pagina bewerken [e]" accesskey="e"><span>Brontekst bewerken</span></a></li><li id="ca-history" class="vector-tab-noicon mw-list-item"><a href="/w/index.php?title=Harald_zur_Hausen&action=history" title="Eerdere versies van deze pagina [h]" accesskey="h"><span>Geschiedenis weergeven</span></a></li> </ul> </div> </div> </nav> <nav class="vector-page-tools-landmark" aria-label="Paginahulpmiddelen"> <div id="vector-page-tools-dropdown" class="vector-dropdown vector-page-tools-dropdown" > <input type="checkbox" id="vector-page-tools-dropdown-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-tools-dropdown" class="vector-dropdown-checkbox " aria-label="Hulpmiddelen" > <label id="vector-page-tools-dropdown-label" for="vector-page-tools-dropdown-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet" aria-hidden="true" ><span class="vector-dropdown-label-text">Hulpmiddelen</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-tools-unpinned-container" class="vector-unpinned-container"> <div id="vector-page-tools" class="vector-page-tools vector-pinnable-element"> <div class="vector-pinnable-header vector-page-tools-pinnable-header vector-pinnable-header-unpinned" data-feature-name="page-tools-pinned" data-pinnable-element-id="vector-page-tools" data-pinned-container-id="vector-page-tools-pinned-container" data-unpinned-container-id="vector-page-tools-unpinned-container" > <div class="vector-pinnable-header-label">Hulpmiddelen</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-page-tools.pin">naar zijbalk verplaatsen</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-page-tools.unpin">verbergen</button> </div> <div id="p-cactions" class="vector-menu mw-portlet mw-portlet-cactions emptyPortlet vector-has-collapsible-items" title="Meer opties" > <div class="vector-menu-heading"> Handelingen </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="ca-more-view" class="selected vector-more-collapsible-item mw-list-item"><a href="/wiki/Harald_zur_Hausen"><span>Lezen</span></a></li><li id="ca-more-ve-edit" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Harald_zur_Hausen&veaction=edit" title="Deze pagina bewerken [v]" accesskey="v"><span>Bewerken</span></a></li><li id="ca-more-edit" class="collapsible vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Harald_zur_Hausen&action=edit" title="Broncode van deze pagina bewerken [e]" accesskey="e"><span>Brontekst bewerken</span></a></li><li id="ca-more-history" class="vector-more-collapsible-item mw-list-item"><a href="/w/index.php?title=Harald_zur_Hausen&action=history"><span>Geschiedenis weergeven</span></a></li> </ul> </div> </div> <div id="p-tb" class="vector-menu mw-portlet mw-portlet-tb" > <div class="vector-menu-heading"> Algemeen </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="t-whatlinkshere" class="mw-list-item"><a href="/wiki/Speciaal:VerwijzingenNaarHier/Harald_zur_Hausen" title="Lijst met alle pagina's die naar deze pagina verwijzen [j]" accesskey="j"><span>Links naar deze pagina</span></a></li><li id="t-recentchangeslinked" class="mw-list-item"><a href="/wiki/Speciaal:RecenteWijzigingenGelinkt/Harald_zur_Hausen" rel="nofollow" title="Recente wijzigingen in pagina's waar deze pagina naar verwijst [k]" accesskey="k"><span>Gerelateerde wijzigingen</span></a></li><li id="t-upload" class="mw-list-item"><a href="//commons.wikimedia.org/wiki/Special:UploadWizard?uselang=nl" title="Bestanden uploaden [u]" accesskey="u"><span>Bestand uploaden</span></a></li><li id="t-specialpages" class="mw-list-item"><a href="/wiki/Speciaal:SpecialePaginas" title="Lijst met alle speciale pagina's [q]" accesskey="q"><span>Speciale pagina's</span></a></li><li id="t-permalink" class="mw-list-item"><a href="/w/index.php?title=Harald_zur_Hausen&oldid=64787487" title="Permanente koppeling naar deze versie van deze pagina"><span>Permanente koppeling</span></a></li><li id="t-info" class="mw-list-item"><a href="/w/index.php?title=Harald_zur_Hausen&action=info" title="Meer informatie over deze pagina"><span>Paginagegevens</span></a></li><li id="t-cite" class="mw-list-item"><a href="/w/index.php?title=Speciaal:Citeren&page=Harald_zur_Hausen&id=64787487&wpFormIdentifier=titleform" title="Informatie over hoe u deze pagina kunt citeren"><span>Deze pagina citeren</span></a></li><li id="t-urlshortener" class="mw-list-item"><a href="/w/index.php?title=Speciaal:UrlQ%C4%B1sald%C4%B1c%C4%B1s%C4%B1&url=https%3A%2F%2Fnl.wikipedia.org%2Fwiki%2FHarald_zur_Hausen"><span>Verkorte URL verkrijgen</span></a></li><li id="t-urlshortener-qrcode" class="mw-list-item"><a href="/w/index.php?title=Speciaal:QrKodu&url=https%3A%2F%2Fnl.wikipedia.org%2Fwiki%2FHarald_zur_Hausen"><span>QR-code downloaden</span></a></li> </ul> </div> </div> <div id="p-coll-print_export" class="vector-menu mw-portlet mw-portlet-coll-print_export" > <div class="vector-menu-heading"> Afdrukken/exporteren </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li id="coll-create_a_book" class="mw-list-item"><a href="/w/index.php?title=Speciaal:Boek&bookcmd=book_creator&referer=Harald+zur+Hausen"><span>Boek aanmaken</span></a></li><li id="coll-download-as-rl" class="mw-list-item"><a href="/w/index.php?title=Speciaal:DownloadAsPdf&page=Harald_zur_Hausen&action=show-download-screen"><span>Downloaden als PDF</span></a></li><li id="t-print" class="mw-list-item"><a href="/w/index.php?title=Harald_zur_Hausen&printable=yes" title="Printvriendelijke versie van deze pagina [p]" accesskey="p"><span>Afdrukversie</span></a></li> </ul> </div> </div> <div id="p-wikibase-otherprojects" class="vector-menu mw-portlet mw-portlet-wikibase-otherprojects" > <div class="vector-menu-heading"> In andere projecten </div> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="wb-otherproject-link wb-otherproject-commons mw-list-item"><a href="https://commons.wikimedia.org/wiki/Category:Harald_zur_Hausen" hreflang="en"><span>Wikimedia Commons</span></a></li><li id="t-wikibase" class="wb-otherproject-link wb-otherproject-wikibase-dataitem mw-list-item"><a href="https://www.wikidata.org/wiki/Special:EntityPage/Q75847" title="Koppeling naar item in verbonden gegevensrepository [g]" accesskey="g"><span>Wikidata-item</span></a></li> </ul> </div> </div> </div> </div> </div> </div> </nav> </div> </div> </div> <div class="vector-column-end"> <div class="vector-sticky-pinned-container"> <nav class="vector-page-tools-landmark" aria-label="Paginahulpmiddelen"> <div id="vector-page-tools-pinned-container" class="vector-pinned-container"> </div> </nav> <nav class="vector-appearance-landmark" aria-label="Uiterlijk"> <div id="vector-appearance-pinned-container" class="vector-pinned-container"> <div id="vector-appearance" class="vector-appearance vector-pinnable-element"> <div class="vector-pinnable-header vector-appearance-pinnable-header vector-pinnable-header-pinned" data-feature-name="appearance-pinned" data-pinnable-element-id="vector-appearance" data-pinned-container-id="vector-appearance-pinned-container" data-unpinned-container-id="vector-appearance-unpinned-container" > <div class="vector-pinnable-header-label">Uiterlijk</div> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-pin-button" data-event-name="pinnable-header.vector-appearance.pin">naar zijbalk verplaatsen</button> <button class="vector-pinnable-header-toggle-button vector-pinnable-header-unpin-button" data-event-name="pinnable-header.vector-appearance.unpin">verbergen</button> </div> </div> </div> </nav> </div> </div> <div id="bodyContent" class="vector-body" aria-labelledby="firstHeading" data-mw-ve-target-container> <div class="vector-body-before-content"> <div class="mw-indicators"> </div> <div id="siteSub" class="noprint">Uit Wikipedia, de vrije encyclopedie</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="nl" dir="ltr"><table class="infobox" cellpadding="1" cellspacing="1"> <tbody><tr> <th class="infobox-kop notheme" style="background-color:lightsteelblue; color:inherit; text-align:center; font-size:120%;" colspan="3"><span typeof="mw:File"><a href="/wiki/Bestand:Nobel_prize_medal.svg" class="mw-file-description" title="Nobelprijswinnaar"><img alt="Nobelprijswinnaar" src="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/25px-Nobel_prize_medal.svg.png" decoding="async" width="25" height="25" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/38px-Nobel_prize_medal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/32/Nobel_prize_medal.svg/50px-Nobel_prize_medal.svg.png 2x" data-file-width="150" data-file-height="150" /></a></span>  Harald zur Hausen </th></tr> <tr> <th class="infobox-kop notheme" style="background-color:lightsteelblue; color:inherit; text-align:center;" colspan="3"><a href="/wiki/11_maart" title="11 maart">11 maart</a> <a href="/wiki/1936" title="1936">1936</a> – <a href="/wiki/28_mei" title="28 mei">28 mei</a> <a href="/wiki/2023" title="2023">2023</a> </th></tr> <tr> <td style="text-align:center; padding:0;" colspan="3"><span typeof="mw:File"><a href="/wiki/Bestand:Harald_zur_Hausen_03.jpg" class="mw-file-description"><img alt="Harald zur Hausen" src="//upload.wikimedia.org/wikipedia/commons/thumb/9/99/Harald_zur_Hausen_03.jpg/266px-Harald_zur_Hausen_03.jpg" decoding="async" width="266" height="344" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/99/Harald_zur_Hausen_03.jpg/399px-Harald_zur_Hausen_03.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/99/Harald_zur_Hausen_03.jpg/532px-Harald_zur_Hausen_03.jpg 2x" data-file-width="1583" data-file-height="2048" /></a></span> </td></tr> <tr> <td style="text-align:center;" colspan="3"><div style="font-size:90%">Harald zur Hausen</div> </td></tr> <tr> <th>Geboorteland </th> <td colspan="2"><span class="mw-image-border" typeof="mw:File"><a href="/wiki/Bestand:Flag_of_Germany_(1933%E2%80%931935).svg" class="mw-file-description" title="Vlag van het Derde Rijk"><img alt="Vlag van het Derde Rijk" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/84/Flag_of_Germany_%281933%E2%80%931935%29.svg/20px-Flag_of_Germany_%281933%E2%80%931935%29.svg.png" decoding="async" width="20" height="12" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/84/Flag_of_Germany_%281933%E2%80%931935%29.svg/30px-Flag_of_Germany_%281933%E2%80%931935%29.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/84/Flag_of_Germany_%281933%E2%80%931935%29.svg/40px-Flag_of_Germany_%281933%E2%80%931935%29.svg.png 2x" data-file-width="512" data-file-height="307" /></a></span> <a href="/wiki/Nazi-Duitsland" title="Nazi-Duitsland">Grootduitse Rijk</a> </td></tr> <tr> <th>Geboorteplaats </th> <td colspan="2"><a href="/wiki/Gelsenkirchen-Buer" title="Gelsenkirchen-Buer">Gelsenkirchen-Buer</a> </td></tr> <tr> <th>Overlijdensplaats </th> <td colspan="2"><a href="/wiki/Heidelberg_(Duitsland)" title="Heidelberg (Duitsland)">Heidelberg</a>, <span class="mw-image-border noviewer" typeof="mw:File"><a href="/wiki/Bestand:Flag_of_Germany.svg" class="mw-file-description" title="Vlag van Duitsland"><img alt="Vlag van Duitsland" src="//upload.wikimedia.org/wikipedia/commons/thumb/b/ba/Flag_of_Germany.svg/20px-Flag_of_Germany.svg.png" decoding="async" width="20" height="12" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/b/ba/Flag_of_Germany.svg/30px-Flag_of_Germany.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/b/ba/Flag_of_Germany.svg/40px-Flag_of_Germany.svg.png 2x" data-file-width="1000" data-file-height="600" /></a></span> <a href="/wiki/Duitsland_(hoofdbetekenis)" class="mw-redirect" title="Duitsland (hoofdbetekenis)">Duitsland</a> </td></tr> <tr> <th>Nobelprijs </th> <td colspan="2"><a href="/wiki/Nobelprijs_voor_de_Fysiologie_of_Geneeskunde" class="mw-redirect" title="Nobelprijs voor de Fysiologie of Geneeskunde">Fysiologie of Geneeskunde</a> </td></tr> <tr> <th>Jaar </th> <td colspan="2"><a href="/wiki/2008" title="2008">2008</a> </td></tr> <tr> <th>Reden </th> <td colspan="2">Voor zijn ontdekking van humane papillomavirussen (HPV) die baarmoederhalskanker kunnen veroorzaken </td></tr> <tr> <th>Gedeeld met </th> <td colspan="2"><a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Luc Montagnier</a><br /><a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Françoise Barré-Sinoussi</a> </td></tr> <tr> <th><span class="nowrap">Voorganger(s)</span> </th> <td colspan="2"><a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Oliver Smithies</a><br /><a href="/wiki/Martin_Evans" title="Martin Evans">Martin Evans</a><br /><a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Mario Capecchi</a> </td></tr> <tr> <th>Opvolger(s) </th> <td colspan="2"><a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Elizabeth Blackburn</a><br /><a href="/wiki/Carol_Greider" title="Carol Greider">Carol Greider</a><br /><a href="/wiki/Jack_Szostak" title="Jack Szostak">Jack Szostak</a> </td></tr> <tr> <td class="infobox-kop notheme center" style="background-color:lightsteelblue; color:inherit;" colspan="3"> <table cellspacing="0" cellpadding="0"> <tbody><tr> <td style="vertical-align:middle;"><b>Portaal</b>  <span class="noviewer" typeof="mw:File"><a href="/wiki/Bestand:Portal.svg" class="mw-file-description" title="Portaalicoon"><img alt="Portaalicoon" src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/22px-Portal.svg.png" decoding="async" width="22" height="20" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/33px-Portal.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c9/Portal.svg/44px-Portal.svg.png 2x" data-file-width="36" data-file-height="32" /></a></span>   </td> <td class="ta-left"><b><a href="/wiki/Portaal:Geneeskunde" title="Portaal:Geneeskunde">Geneeskunde</a> </b> </td></tr></tbody></table> </td></tr> </tbody></table> <p><b>Harald zur Hausen</b> (<a href="/wiki/Gelsenkirchen-Buer" title="Gelsenkirchen-Buer">Gelsenkirchen-Buer</a>, <a href="/wiki/11_maart" title="11 maart">11 maart</a> <a href="/wiki/1936" title="1936">1936</a> – <a href="/wiki/Heidelberg_(Duitsland)" title="Heidelberg (Duitsland)">Heidelberg</a>, <a href="/wiki/28_mei" title="28 mei">28 mei</a> <a href="/wiki/2023" title="2023">2023</a>) was een <a href="/wiki/Duitsland" title="Duitsland">Duitse</a> <a href="/wiki/Arts" title="Arts">arts</a> en <a href="/wiki/Viroloog" class="mw-redirect" title="Viroloog">viroloog</a>. Hij kreeg de <a href="/wiki/Nobelprijs_voor_de_Fysiologie_of_Geneeskunde" class="mw-redirect" title="Nobelprijs voor de Fysiologie of Geneeskunde">Nobelprijs voor de Fysiologie of Geneeskunde</a> van <a href="/wiki/2008" title="2008">2008</a><sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> voor zijn ontdekking van <a href="/wiki/Humaan_papillomavirus" title="Humaan papillomavirus">humane papillomavirussen</a> (HPV) die <a href="/wiki/Baarmoederhalskanker" title="Baarmoederhalskanker">baarmoederhalskanker</a> kunnen veroorzaken. De Nobelprijs moest hij delen met de Franse virologen <a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Françoise Barré-Sinoussi</a> en <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Luc Montagnier</a> voor hun ontdekking van het menselijk immuundeficiëntievirus (<a href="/wiki/Hiv" title="Hiv">hiv</a>). </p> <div class="mw-heading mw-heading2"><h2 id="Biografie">Biografie</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Harald_zur_Hausen&veaction=edit&section=1" title="Bewerk dit kopje: Biografie" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Harald_zur_Hausen&action=edit&section=1" title="De broncode bewerken van de sectie: Biografie"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Zur Hausen studeerde geneeskunde aan de universiteiten van Bonn, Hamburg en Düsseldorf waar hij in 1960 zijn medische graad behaalde. Na twee jaar gewerkt te hebben als medisch assistent ging hij als onderzoeker werken bij het Instituut voor Microbiologie aan de <a href="/wiki/Heinrich_Heine-Universiteit" title="Heinrich Heine-Universiteit">Heinrich Heine-Universiteit</a> te <a href="/wiki/D%C3%BCsseldorf" title="Düsseldorf">Düsseldorf</a>. </p><p>Vanaf 1966 was hij werkzaam op het viruslaboratorium van het kinderziekenhuis van <a href="/wiki/Philadelphia_(Pennsylvania)" title="Philadelphia (Pennsylvania)">Philadelphia</a> (onder de Duitse biologen <a href="/w/index.php?title=Werner_Henle&action=edit&redlink=1" class="new" title="Werner Henle (de pagina bestaat niet)">Werner</a> en <a href="/w/index.php?title=Gertrude_Henle&action=edit&redlink=1" class="new" title="Gertrude Henle (de pagina bestaat niet)">Gertrude Henle</a>) Later was hij ook docent virologie aan de <a href="/wiki/Universiteit_van_Pennsylvania" title="Universiteit van Pennsylvania">Universiteit van Pennsylvania</a>, waarna hij in 1969 terugkeerde naar Duitsland. Achtereenvolgens was hij hoogleraar virologie aan de <a href="/wiki/Julius_Maximilians-Universiteit" title="Julius Maximilians-Universiteit">Julius Maximilians-Universiteit</a> in Würzburg (1969-1972), de <a href="/wiki/Friedrich-Alexander-Universit%C3%A4t_Erlangen-N%C3%BCrnberg" title="Friedrich-Alexander-Universität Erlangen-Nürnberg">Friedrich-Alexander-Universität Erlangen-Nürnberg</a> (1972-1977) en de <a href="/wiki/Universiteit_van_Freiburg" title="Universiteit van Freiburg">Universiteit van Freiburg</a> (1977-1983). Het laatste deel van zijn carrière was hij wetenschappelijk directeur van het Duitse Kankeronderzoekscentrum in Heidelberg. Daarnaast was hij hoofdredacteur van het <i>International Journal of Cancer</i>. </p><p>Zur Hausen overleed op 87-jarige leeftijd.<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-3" class="reference"><a href="#cite_note-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Werk">Werk</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Harald_zur_Hausen&veaction=edit&section=2" title="Bewerk dit kopje: Werk" class="mw-editsection-visualeditor"><span>bewerken</span></a><span class="mw-editsection-divider"> | </span><a href="/w/index.php?title=Harald_zur_Hausen&action=edit&section=2" title="De broncode bewerken van de sectie: Werk"><span>brontekst bewerken</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Begin jaren 1970 vermoedde Zur Hausen dat vrouwen die besmet waren met <a href="/wiki/Genitale_wrat" title="Genitale wrat">genitale wratten</a> een verhoogde kans hadden op <a href="/wiki/Baarmoederhalskanker" title="Baarmoederhalskanker">baarmoederhalskanker</a>. Genitale wratten ontstaan door (seksuele) besmetting met het humane papillomavirus (HPV) en als zijn theorie juist was zouden fragmenten van het virus-DNA terug te vinden zijn in de tumorcellen. Zijn hypothese publiceerde hij in het vakblad Cancer Research.<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Onderzoek naar welk virustype hiervoor verantwoordelijk was bleek niet eenvoudig, maar na meer dan tien jaar slaagde Zur Hausen in zijn opzet. In 1983 respectievelijk 1984 vond en identificeerde hij de belangrijkste genotypes van het virus, de types 16 en 18. Deze twee virustypes zijn verantwoordelijk voor 70% van het aantal gevallen van deze vorm van kanker.<sup id="cite_ref-5" class="reference"><a href="#cite_note-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> </p><p>Hoewel zijn bevindingen aanvankelijk met enige scepsis werd ontvangen door de medische wetenschap, leidde zijn onderzoeksresultaten tot de ontwikkeling van <a href="/wiki/Gardasil" title="Gardasil">Gardasil</a>, een vaccin tegen baarmoederhalskanker. </p> <div class="toccolours appendix" role="presentation" style="font-size:90%; margin:1em 0 -0.5em; clear:both;"> <div><span style="font-weight:bold">Bronnen, noten en/of referenties</span></div> <ul><li><style data-mw-deduplicate="TemplateStyles:r67679320">.mw-parser-output .taalaanduiding{font-family:sans-serif;font-size:85%;cursor:help;color:var(--color-subtle,#555)}.mw-parser-output .taalaanduiding span{border-bottom:1px dotted var(--color-subtle,#555)}</style><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span><a rel="nofollow" class="external text" href="https://www.nobelprize.org/nobel_prizes/medicine/laureates/2008/hausen-bio.html">Autobiografie Harold zur Hausen</a> op Nobelprize.org</li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67679320"><span class="taalaanduiding" title="Taal: Engels">(<span>en</span>) </span><a rel="nofollow" class="external text" href="https://web.archive.org/web/20100103192040/http://www.monstersandcritics.com/news/europe/news/article_1435030.php/PROFILE_Harald_zur_Hausen_-_Nobel_honour_for_cancer_research">Profiel: Harald zur Hausen - Nobel honour for cancer research</a></li></ul> <hr /> <div class="reflist" style="list-style-type: decimal;"><div class="mw-references-wrap"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><a href="#cite_ref-1">↑</a></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://www.nu.nl/wetenschap/1776610/nobelprijs-geneeskunde-naar-duitser-en-twee-fransen-video.html">Nobelprijs Geneeskunde naar Duitser en twee Fransen</a>, <a href="/wiki/NU.nl" title="NU.nl">NU.nl</a>, 6 oktober 2008. <a rel="nofollow" class="external text" href="https://web.archive.org/web/20230602024222/https://www.nu.nl/wetenschap/1776610/nobelprijs-geneeskunde-naar-duitser-en-twee-fransen-video.html">Gearchiveerd</a> op 2 juni 2023.</span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><a href="#cite_ref-2">↑</a></span> <span class="reference-text"><span class="citation" id="CITEREF2023"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67679320"><span class="taalaanduiding" title="Taal: Duits">(<span>de</span>) </span><a rel="nofollow" class="external text" href="https://www.zdf.de/uri/256c6e60-9cca-4619-a8b9-236e1f0718b0">Medizin-Nobelpreisträger Harald zur Hausen gestorben</a>. <i>ZDF</i> (<span class="mw-formatted-date" title="2023-05-29">29 mei 2023</span>). </span></span> </li> <li id="cite_note-3"><span class="mw-cite-backlink"><a href="#cite_ref-3">↑</a></span> <span class="reference-text"><span class="citation" id="CITEREF2023"><a rel="nofollow" class="external text" href="https://www.standaard.be/cnt/dmf20230529_96445417">Nobelprijswinnaar Harald zur Hausen overleden</a>. <i>De Standaard</i> (<span class="mw-formatted-date" title="2023-05-29">29 mei 2023</span>). <a rel="nofollow" class="external text" href="https://web.archive.org/web/20230608092113/https://www.standaard.be/cnt/dmf20230529_96445417">Gearchiveerd</a> op <span class="mw-formatted-date" title="2023-06-08">8 juni 2023</span>. </span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><a href="#cite_ref-4">↑</a></span> <span class="reference-text"><span class="citation" id="CITEREFHarald_zur_Hausen1976"><span style="font-variant:small-caps;">Harald zur Hausen</span> (1976). <cite><a rel="nofollow" class="external text" href="https://archive.org/details/sim_cancer-research_1976-02_36_2/page/794">Condylomata Acuminata and Human Genital Cancer</a></cite>. <i>Cancer Research</i> <b>36</b>: 794. <a href="/wiki/PubMed" title="PubMed">PMID</a> <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pubmed/175942">175942</a>. </span></span> </li> <li id="cite_note-5"><span class="mw-cite-backlink"><a href="#cite_ref-5">↑</a></span> <span class="reference-text">Zie <a href="/wiki/Baarmoederhalskanker" title="Baarmoederhalskanker">baarmoederhalskanker</a> op Wikipedia</span> </li> </ol></div></div> </div> <style data-mw-deduplicate="TemplateStyles:r67837862">.mw-parser-output .navigatie{position:relative;clear:both;overflow:auto;margin:1em auto -0.5em;padding:2px;background-color:var(--background-color-neutral-subtle,#f8f9fa);border:1px solid var(--border-color-base,#a2a9b1);text-align:center;font-size:87%}.mw-parser-output .navigatie-bewerken{margin-left:0.5em}.mw-parser-output .navigatie-bewerken .mw-ui-icon::before{background-size:0.9em}.mw-parser-output .navigatie-afb-links,.mw-parser-output .navigatie-afb-rechts{position:absolute}.mw-parser-output .navigatie-afb-rechts{right:2px}.mw-parser-output .navigatie-afb-groot{float:right;padding-left:0.5em}.mw-parser-output .navigatie-titel{background-color:#ddeef8;padding:2px 5.5em;font-weight:bold}.mw-parser-output .navigatie-inhoud{padding:0.5em}.mw-parser-output .navigatie-inhoud p:first-child{margin:0}.mw-parser-output .navigatie div[style*="background-color"],.mw-parser-output .navigatie div[style*="background"]{color:inherit}@media screen{html.skin-theme-clientpref-night .mw-parser-output .navigatie-titel{background-color:var(--background-color-interactive,#eaecf0)!important}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .navigatie-titel{background-color:var(--background-color-interactive,#eaecf0)!important}}</style><div class="navigatie mw-collapsible mw-collapsed" role="navigation" aria-labelledby="Bibliografische informatie" data-collapsetext="Inklappen" data-expandtext="Uitklappen"><div id="Bibliografische_informatie" class="navigatie-titel"><a href="/wiki/Wikipedia:Bibliografische_informatie" title="Wikipedia:Bibliografische informatie">Bibliografische informatie</a></div><div class="navigatie-inhoud mw-collapsible-content plainlist ta-left"><ul style="column-width:36em"><li><b><a href="/wiki/Bibsys" class="mw-redirect" title="Bibsys">Bibsys</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://authority.bibsys.no/authority/rest/authorities/html/90933354">90933354</a></span></li><li><b><a href="/wiki/Biblioteca_Nacional_de_Espa%C3%B1a" title="Biblioteca Nacional de España">Biblioteca Nacional de España</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="http://catalogo.bne.es/uhtbin/authoritybrowse.cgi?action=display&authority_id=XX5065098">XX5065098</a></span></li><li><b><a href="/wiki/Biblioth%C3%A8que_nationale_de_France" title="Bibliothèque nationale de France">Bibliothèque nationale de France</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://catalogue.bnf.fr/ark:/12148/cb12279302x">cb12279302x</a> <a rel="nofollow" class="external text" href="https://data.bnf.fr/ark:/12148/cb12279302x">(data)</a></span></li><li><b><a href="/wiki/CiNii" title="CiNii">CiNii</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://ci.nii.ac.jp/author/DA00595349?l=en">DA00595349</a></span></li><li><b><a href="/wiki/Gemeinsame_Normdatei" title="Gemeinsame Normdatei">Gemeinsame Normdatei</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://d-nb.info/gnd/121279731">121279731</a></span></li><li><b><a href="/wiki/International_Standard_Name_Identifier" title="International Standard Name Identifier">International Standard Name Identifier</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="http://isni.org/isni/0000000109206695">0000 0001 0920 6695</a></span></li><li><b><a href="/wiki/Library_of_Congress_Control_Number" title="Library of Congress Control Number">Library of Congress Control Number</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://id.loc.gov/authorities/names/n80083202">n80083202</a></span></li><li><b><a href="/wiki/Nationale_Bibliotheek_van_Letland" title="Nationale Bibliotheek van Letland">Nationale Bibliotheek van Letland</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://kopkatalogs.lv/F?func=direct&local_base=lnc10&doc_number=000258358&P_CON_LNG=ENG">000258358</a></span></li><li><b><a href="/wiki/Nationale_Bibliotheek_van_Tsjechi%C3%AB" title="Nationale Bibliotheek van Tsjechië">Nationale Bibliotheek van Tsjechië</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://aleph.nkp.cz/F/?func=find-c&local_base=aut&ccl_term=ica=mzk2005294787&CON_LNG=ENG">mzk2005294787</a></span></li><li><b><a href="/wiki/Nationale_Bibliotheek_van_Isra%C3%ABl" title="Nationale Bibliotheek van Israël">Nationale Bibliotheek van Israël</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://www.nli.org.il/en/authorities/987007435260505171">987007435260505171</a></span></li><li><b><a href="/wiki/Nationale_Bibliotheek_van_Polen" title="Nationale Bibliotheek van Polen">Nationale Bibliotheek van Polen</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://dbn.bn.org.pl/descriptor-details/a0000003135568">a0000003135568</a></span></li><li><b><a href="https://www.wikidata.org/wiki/Property:P1006" class="extiw" title="d:Property:P1006">Nederlandse Thesaurus van Auteursnamen</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="http://data.bibliotheken.nl/id/thes/p068242530">068242530</a></span></li><li><b><a href="/wiki/ORCID" title="ORCID">ORCID</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://orcid.org/0000-0003-1103-0347">0000-0003-1103-0347</a></span></li><li><b><a href="/wiki/R%C3%89RO" title="RÉRO">Réseau des bibliothèques de Suisse occidentale</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="http://data.rero.ch/02-A004003349">02-A004003349</a></span></li><li><b><a href="/wiki/LIBRIS" title="LIBRIS">LIBRIS</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://libris.kb.se/auth/319933">319933</a></span></li><li><b><a href="/wiki/Syst%C3%A8me_universitaire_de_documentation" title="Système universitaire de documentation">Système universitaire de documentation</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://www.idref.fr/031611699">031611699</a></span></li><li><b><a href="/wiki/Virtual_International_Authority_File" title="Virtual International Authority File">Virtual International Authority File</a>:</b> <span class="uid"><a rel="nofollow" class="external text" href="https://viaf.org/viaf/84630562">84630562</a></span></li><li><b></b> <b><a href="/wiki/WorldCat" title="WorldCat">WorldCat</a></b> (via VIAF): <a rel="nofollow" class="external text" href="https://www.worldcat.org/identities/containsVIAFID/84630562">84630562</a></li></ul></div></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r67837862"> <div class="navigatie mw-collapsible mw-collapsed" role="navigation" aria-labelledby="Winnaars_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" data-expandtext="Uitklappen" data-collapsetext="Inklappen"> <div class="navigatie-afb-links noviewer"><div class="navigatie-bewerken show-autoconfirmed nomobile"><span class="nowrap skin-invert-image"><span typeof="mw:File"><a href="/wiki/Sjabloon:Navigatie_winnaars_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Sjabloon:Navigatie winnaars van de Nobelprijs voor Fysiologie of Geneeskunde"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c4/OOjs_UI_icon_article-ltr.svg/16px-OOjs_UI_icon_article-ltr.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c4/OOjs_UI_icon_article-ltr.svg/24px-OOjs_UI_icon_article-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c4/OOjs_UI_icon_article-ltr.svg/32px-OOjs_UI_icon_article-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span> · <span class="ui-state-disabled" typeof="mw:File"><span title="Overleg sjabloon (de pagina bestaat niet)"><img alt="Overleg sjabloon (de pagina bestaat niet)" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/40/OOjs_UI_icon_speechBubbles-ltr.svg/16px-OOjs_UI_icon_speechBubbles-ltr.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/40/OOjs_UI_icon_speechBubbles-ltr.svg/24px-OOjs_UI_icon_speechBubbles-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/40/OOjs_UI_icon_speechBubbles-ltr.svg/32px-OOjs_UI_icon_speechBubbles-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></span></span> · <span typeof="mw:File"><a href="//nl.wikipedia.org/w/index.php?title=Sjabloon:Navigatie_winnaars_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde&action=edit" title="Sjabloon bewerken"><img alt="Sjabloon bewerken" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/16px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/24px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/32px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span></div></div> <div id="Winnaars_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" class="navigatie-titel">Winnaars van de <a href="/wiki/Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Nobelprijs voor Fysiologie of Geneeskunde">Nobelprijs voor Fysiologie of Geneeskunde</a></div> <div class="navigatie-inhoud mw-collapsible-content"> <p>1901: <a href="/wiki/Emil_Adolf_von_Behring" title="Emil Adolf von Behring">Behring</a> · 1902: <a href="/wiki/Ronald_Ross" title="Ronald Ross">Ross</a> · 1903: <a href="/wiki/Niels_Ryberg_Finsen" title="Niels Ryberg Finsen">Finsen</a> · 1904: <a href="/wiki/Ivan_Pavlov" title="Ivan Pavlov">Pavlov</a> · 1905: <a href="/wiki/Robert_Koch" title="Robert Koch">Koch</a> · 1906: <a href="/wiki/Camillo_Golgi" title="Camillo Golgi">Golgi</a>, <a href="/wiki/Santiago_Ram%C3%B3n_y_Cajal" title="Santiago Ramón y Cajal">Ramón y Cajal</a> · 1907: <a href="/wiki/Charles_Louis_Alphonse_Laveran" title="Charles Louis Alphonse Laveran">Laveran</a> · 1908: <a href="/wiki/Ilja_Iljitsj_Metsjnikov" title="Ilja Iljitsj Metsjnikov">Mechnikov</a>, <a href="/wiki/Paul_Ehrlich" title="Paul Ehrlich">Ehrlich</a> · 1909: <a href="/wiki/Emil_Theodor_Kocher" title="Emil Theodor Kocher">Kocher</a> · 1910: <a href="/wiki/Albrecht_Kossel" title="Albrecht Kossel">Kossel</a> · 1911: <a href="/wiki/Allvar_Gullstrand" title="Allvar Gullstrand">Gullstrand</a> · 1912: <a href="/wiki/Alexis_Carrel" title="Alexis Carrel">Carrel</a> · 1913: <a href="/wiki/Charles_Richet" title="Charles Richet">Richet</a> · 1914: <a href="/wiki/Robert_B%C3%A1r%C3%A1ny" title="Robert Bárány">Bárány</a> · 1919: <a href="/wiki/Jules_Bordet" title="Jules Bordet">Bordet</a> · 1920: <a href="/wiki/August_Krogh" title="August Krogh">Krogh</a> · 1922: <a href="/wiki/Archibald_Hill" title="Archibald Hill">Hill</a>, <a href="/wiki/Otto_Fritz_Meyerhof" title="Otto Fritz Meyerhof">Meyerhof</a> · 1923: <a href="/wiki/Frederick_Banting" title="Frederick Banting">Banting</a>, <a href="/wiki/John_Macleod_(medicus)" title="John Macleod (medicus)">Macleod</a> · 1924: <a href="/wiki/Willem_Einthoven" title="Willem Einthoven">Einthoven</a> · 1926: <a href="/wiki/Johannes_Andreas_Grib_Fibiger" title="Johannes Andreas Grib Fibiger">Fibiger</a> · 1927: <a href="/wiki/Julius_Wagner-Jauregg" title="Julius Wagner-Jauregg">Wagner-Jauregg</a> · 1928: <a href="/wiki/Charles_Nicolle" title="Charles Nicolle">Nicolle</a> · 1929: <a href="/wiki/Christiaan_Eijkman" title="Christiaan Eijkman">Eijkman</a>, <a href="/wiki/Frederick_Gowland_Hopkins" title="Frederick Gowland Hopkins">Hopkins</a> · 1930: <a href="/wiki/Karl_Landsteiner" title="Karl Landsteiner">Landsteiner</a> · 1931: <a href="/wiki/Otto_Heinrich_Warburg" title="Otto Heinrich Warburg">Warburg</a> · 1932: <a href="/wiki/Charles_Scott_Sherrington" title="Charles Scott Sherrington">Sherrington</a>, <a href="/wiki/Edgar_Douglas_Adrian" title="Edgar Douglas Adrian">Adrian</a> · 1933: <a href="/wiki/Thomas_Hunt_Morgan" title="Thomas Hunt Morgan">Morgan</a> · 1934: <a href="/wiki/George_Whipple" title="George Whipple">Whipple</a>, <a href="/wiki/George_Minot" title="George Minot">Minot</a>, <a href="/wiki/William_Murphy" title="William Murphy">Murphy</a> · 1935: <a href="/wiki/Hans_Spemann" title="Hans Spemann">Spemann</a> · 1936: <a href="/wiki/Henry_Hallett_Dale" title="Henry Hallett Dale">Dale</a>, <a href="/wiki/Otto_Loewi" title="Otto Loewi">Loewi</a> · 1937: <a href="/wiki/Albert_Szent-Gy%C3%B6rgyi" title="Albert Szent-Györgyi">Szent-Györgyi</a> · 1938: <a href="/wiki/Corneel_Heymans" title="Corneel Heymans">Heymans</a> · 1939: <a href="/wiki/Gerhard_Domagk" title="Gerhard Domagk">Domagk</a> · 1943: <a href="/wiki/Henrik_Dam" title="Henrik Dam">Dam</a>, <a href="/wiki/Edward_Adelbert_Doisy" title="Edward Adelbert Doisy">Doisy</a> · 1944: <a href="/wiki/Joseph_Erlanger" title="Joseph Erlanger">Erlanger</a>, <a href="/wiki/Herbert_Spencer_Gasser" title="Herbert Spencer Gasser">Gasser</a> · 1945: <a href="/wiki/Alexander_Fleming" title="Alexander Fleming">Fleming</a>, <a href="/wiki/Ernst_Boris_Chain" title="Ernst Boris Chain">Chain</a>, <a href="/wiki/Howard_Florey" title="Howard Florey">Florey</a> · 1946: <a href="/wiki/Hermann_Joseph_Muller" title="Hermann Joseph Muller">Muller</a> · 1947: <span class="nowrap"><a href="/wiki/Carl_Ferdinand_Cori" title="Carl Ferdinand Cori">C. Cori</a></span>, <span class="nowrap"><a href="/wiki/Gerty_Cori" title="Gerty Cori">G. Cori</a></span>, <a href="/wiki/Bernardo_Houssay" title="Bernardo Houssay">Houssay</a> · 1948: <a href="/wiki/Paul_Hermann_M%C3%BCller" title="Paul Hermann Müller">Müller</a> · 1949: <a href="/wiki/Walter_Rudolf_Hess" title="Walter Rudolf Hess">Hess</a>, <a href="/wiki/Ant%C3%B3nio_Egas_Moniz" title="António Egas Moniz">Moniz</a> · 1950: <a href="/wiki/Edward_Calvin_Kendall" title="Edward Calvin Kendall">Kendall</a>, <a href="/wiki/Tadeus_Reichstein" title="Tadeus Reichstein">Reichstein</a>, <a href="/wiki/Philip_Showalter_Hench" title="Philip Showalter Hench">Hench</a> · 1951: <a href="/wiki/Max_Theiler" title="Max Theiler">Theiler</a> · 1952: <a href="/wiki/Selman_Waksman" title="Selman Waksman">Waksman</a> · 1953: <a href="/wiki/Hans_Krebs_(arts)" title="Hans Krebs (arts)">Krebs</a>,<a href="/wiki/Fritz_Albert_Lipmann" title="Fritz Albert Lipmann">Lipmann</a> · 1954: <a href="/wiki/John_Franklin_Enders" title="John Franklin Enders">Enders</a>, <a href="/wiki/Thomas_Huckle_Weller" title="Thomas Huckle Weller">Weller</a>, <a href="/wiki/Frederick_Chapman_Robbins" title="Frederick Chapman Robbins">Robbins</a> · 1955: <a href="/wiki/Hugo_Theorell" title="Hugo Theorell">Theorell</a> · 1956: <a href="/wiki/Andr%C3%A9_Fr%C3%A9d%C3%A9ric_Cournand" title="André Frédéric Cournand">Cournand</a>, <a href="/wiki/Werner_Forssmann" title="Werner Forssmann">Forssmann</a>, <a href="/wiki/Dickinson_W._Richards" title="Dickinson W. Richards">Richards</a> · 1957: <a href="/wiki/Daniel_Bovet" title="Daniel Bovet">Bovet</a> · 1958: <a href="/wiki/George_Wells_Beadle" title="George Wells Beadle">Beadle</a>, <a href="/wiki/Edward_Lawrie_Tatum" title="Edward Lawrie Tatum">Tatum</a>, <a href="/wiki/Joshua_Lederberg" title="Joshua Lederberg">Lederberg</a> · 1959: <a href="/wiki/Severo_Ochoa" title="Severo Ochoa">Ochoa</a>, <a href="/wiki/Arthur_Kornberg" title="Arthur Kornberg">Kornberg</a> · 1960: <a href="/wiki/Frank_Macfarlane_Burnet" title="Frank Macfarlane Burnet">Burnet</a>, <a href="/wiki/Peter_Medawar" title="Peter Medawar">Medawar</a> · 1961: <a href="/wiki/Georg_von_B%C3%A9k%C3%A9sy" title="Georg von Békésy">Békésy</a> · 1962: <a href="/wiki/Francis_Crick" title="Francis Crick">Crick</a>, <a href="/wiki/James_Watson" title="James Watson">Watson</a>, <a href="/wiki/Maurice_Wilkins" title="Maurice Wilkins">Wilkins</a> · 1963: <a href="/wiki/John_Eccles" title="John Eccles">Eccles</a>, <a href="/wiki/Alan_Lloyd_Hodgkin" title="Alan Lloyd Hodgkin">Hodgkin</a>, <a href="/wiki/Andrew_Huxley" title="Andrew Huxley">Huxley</a> · 1964: <a href="/wiki/Konrad_Emil_Bloch" title="Konrad Emil Bloch">Bloch</a>, <a href="/wiki/Feodor_Felix_Konrad_Lynen" title="Feodor Felix Konrad Lynen">Lynen</a> · 1965: <a href="/wiki/Fran%C3%A7ois_Jacob" title="François Jacob">Jacob</a>, <a href="/wiki/Andr%C3%A9_Michael_Lwoff" title="André Michael Lwoff">Lwoff</a>, <a href="/wiki/Jacques_Monod" title="Jacques Monod">Monod</a> · 1966: <a href="/wiki/Francis_Peyton_Rous" title="Francis Peyton Rous">Rous</a>, <a href="/wiki/Charles_Brenton_Huggins" title="Charles Brenton Huggins">Huggins</a> · 1967: <a href="/wiki/Ragnar_Granit" title="Ragnar Granit">Granit</a>, <a href="/wiki/Haldan_Keffer_Hartline" title="Haldan Keffer Hartline">Hartline</a>, <a href="/wiki/George_Wald" title="George Wald">Wald</a> · 1968: <a href="/wiki/Robert_W._Holley" title="Robert W. Holley">Holley</a>, <a href="/wiki/Har_Gobind_Khorana" title="Har Gobind Khorana">Khorana</a>, <a href="/wiki/Marshall_Warren_Nirenberg" title="Marshall Warren Nirenberg">Nirenberg</a> · 1969: <a href="/wiki/Max_Delbr%C3%BCck" title="Max Delbrück">Delbrück</a>, <a href="/wiki/Alfred_Hershey" title="Alfred Hershey">Hershey</a>, <a href="/wiki/Salvador_Luria" title="Salvador Luria">Luria</a> · 1970: <a href="/wiki/Bernard_Katz" title="Bernard Katz">Katz</a>, <a href="/wiki/Ulf_Svante_von_Euler" title="Ulf Svante von Euler">Euler</a>, <a href="/wiki/Julius_Axelrod" title="Julius Axelrod">Axelrod</a> · 1971: <a href="/wiki/Earl_Wilbur_Sutherland_jr." title="Earl Wilbur Sutherland jr.">Sutherland</a> · 1972: <a href="/wiki/Gerald_Edelman" title="Gerald Edelman">Edelman</a>, <a href="/wiki/Rodney_Porter" title="Rodney Porter">Porter</a> · 1973: <a href="/wiki/Karl_von_Frisch" title="Karl von Frisch">Frisch</a>, <a href="/wiki/Konrad_Lorenz" title="Konrad Lorenz">Lorenz</a>, <a href="/wiki/Niko_Tinbergen" title="Niko Tinbergen">Tinbergen</a> · 1974: <a href="/wiki/Albert_Claude" title="Albert Claude">Claude</a>, <a href="/wiki/Christian_de_Duve" title="Christian de Duve">De Duve</a>, <a href="/wiki/George_Emil_Palade" title="George Emil Palade">Palade</a> · 1975: <a href="/wiki/David_Baltimore" title="David Baltimore">Baltimore</a>, <a href="/wiki/Renato_Dulbecco" title="Renato Dulbecco">Dulbecco</a>, <a href="/wiki/Howard_Martin_Temin" title="Howard Martin Temin">Temin</a> · 1976: <a href="/wiki/Baruch_Samuel_Blumberg" title="Baruch Samuel Blumberg">Blumberg</a>, <a href="/wiki/Daniel_Carleton_Gajdusek" title="Daniel Carleton Gajdusek">Gajdusek</a> · 1977: <a href="/wiki/Roger_Guillemin" title="Roger Guillemin">Guillemin</a>, <a href="/wiki/Andrew_Schally" title="Andrew Schally">Schally</a>, <a href="/wiki/Rosalyn_Sussman_Yalow" title="Rosalyn Sussman Yalow">Yalow</a> · 1978: <a href="/wiki/Werner_Arber" title="Werner Arber">Arber</a>, <a href="/wiki/Daniel_Nathans" title="Daniel Nathans">Nathans</a>, <a href="/wiki/Hamilton_Othanel_Smith" title="Hamilton Othanel Smith">Smith</a> · 1979: <a href="/wiki/Allan_McLeod_Cormack" title="Allan McLeod Cormack">Cormack</a>, <a href="/wiki/Godfrey_Hounsfield" title="Godfrey Hounsfield">Hounsfield</a> · 1980: <a href="/wiki/Baruj_Benacerraf" title="Baruj Benacerraf">Benacerraf</a>, <a href="/wiki/Jean_Dausset" title="Jean Dausset">Dausset</a>, <a href="/wiki/George_Davis_Snell" title="George Davis Snell">Snell</a> · 1981: <a href="/wiki/Roger_Sperry" title="Roger Sperry">Sperry</a>, <a href="/wiki/David_H._Hubel" title="David H. Hubel">Hubel</a>, <a href="/wiki/Torsten_Wiesel" title="Torsten Wiesel">Wiesel</a> · 1982: <a href="/wiki/Sune_Karl_Bergstr%C3%B6m" title="Sune Karl Bergström">Bergström</a>, <a href="/wiki/Bengt_Ingemar_Samuelsson" title="Bengt Ingemar Samuelsson">Samuelsson</a>, <a href="/wiki/John_Robert_Vane" title="John Robert Vane">Vane</a> · 1983: <a href="/wiki/Barbara_McClintock" title="Barbara McClintock">McClintock</a> · 1984: <a href="/wiki/Niels_Kaj_Jerne" title="Niels Kaj Jerne">Jerne</a>, <a href="/wiki/Georges_J.F._K%C3%B6hler" title="Georges J.F. Köhler">Köhler</a>, <a href="/wiki/C%C3%A9sar_Milstein" title="César Milstein">Milstein</a> · 1985: <a href="/wiki/Michael_Stuart_Brown" title="Michael Stuart Brown">Brown</a>, <a href="/wiki/Joseph_Goldstein" title="Joseph Goldstein">Goldstein</a> · 1986: <a href="/wiki/Stanley_Cohen_(biochemicus)" title="Stanley Cohen (biochemicus)">Cohen</a>, <span class="nowrap"><a href="/wiki/Rita_Levi-Montalcini" title="Rita Levi-Montalcini">Levi-Montalcini</a></span> · 1987: <a href="/wiki/Susumu_Tonegawa" title="Susumu Tonegawa">Tonegawa</a> · 1988: <a href="/wiki/James_Whyte_Black" title="James Whyte Black">Black</a>, <a href="/wiki/Gertrude_Elion" title="Gertrude Elion">Elion</a>, <a href="/wiki/George_H._Hitchings" title="George H. Hitchings">Hitchings</a> · 1989: <a href="/wiki/John_Michael_Bishop" title="John Michael Bishop">Bishop</a>, <a href="/wiki/Harold_E._Varmus" title="Harold E. Varmus">Varmus</a> · 1990: <a href="/wiki/Joseph_Murray_(chirurg)" title="Joseph Murray (chirurg)">Murray</a>, <a href="/wiki/Edward_Donnall_Thomas" title="Edward Donnall Thomas">Thomas</a> · 1991: <a href="/wiki/Erwin_Neher" title="Erwin Neher">Neher</a>, <a href="/wiki/Bert_Sakmann" title="Bert Sakmann">Sakmann</a> · 1992: <a href="/wiki/Edmond_H._Fischer" title="Edmond H. Fischer">Fischer</a>, <a href="/wiki/Edwin_Gerhard_Krebs" title="Edwin Gerhard Krebs">Krebs</a> · 1993: <a href="/wiki/Richard_John_Roberts" title="Richard John Roberts">Roberts</a>, <a href="/wiki/Phillip_Allen_Sharp" title="Phillip Allen Sharp">Sharp</a> · 1994: <a href="/wiki/Alfred_Goodman_Gilman" title="Alfred Goodman Gilman">Gilman</a>, <a href="/wiki/Martin_Rodbell" title="Martin Rodbell">Rodbell</a> · 1995: <a href="/wiki/Edward_B._Lewis" title="Edward B. Lewis">Lewis</a>, <span class="nowrap"><a href="/wiki/Christiane_N%C3%BCsslein-Volhard" title="Christiane Nüsslein-Volhard">Nüsslein-Volhard</a></span>, <a href="/wiki/Eric_F._Wieschaus" title="Eric F. Wieschaus">Wieschaus</a> · 1996: <a href="/wiki/Peter_C._Doherty" title="Peter C. Doherty">Doherty</a>, <a href="/wiki/Rolf_M._Zinkernagel" title="Rolf M. Zinkernagel">Zinkernagel</a> · 1997: <a href="/wiki/Stanley_B._Prusiner" title="Stanley B. Prusiner">Prusiner</a> · 1998: <a href="/wiki/Robert_F._Furchgott" title="Robert F. Furchgott">Furchgott</a>, <a href="/wiki/Louis_Ignarro" title="Louis Ignarro">Ignarro</a>, <a href="/wiki/Ferid_Murad" title="Ferid Murad">Murad</a> · 1999: <a href="/wiki/G%C3%BCnter_Blobel" title="Günter Blobel">Blobel</a> · 2000: <a href="/wiki/Arvid_Carlsson" title="Arvid Carlsson">Carlsson</a>, <a href="/wiki/Paul_Greengard" title="Paul Greengard">Greengard</a>, <a href="/wiki/Eric_Kandel" title="Eric Kandel">Kandel</a> · 2001: <a href="/wiki/Leland_H._Hartwell" title="Leland H. Hartwell">Hartwell</a>, <a href="/wiki/Tim_Hunt" title="Tim Hunt">Hunt</a>, <a href="/wiki/Paul_Nurse" title="Paul Nurse">Nurse</a> · 2002: <a href="/wiki/Sydney_Brenner" title="Sydney Brenner">Brenner</a>, <a href="/wiki/H._Robert_Horvitz" title="H. Robert Horvitz">Horvitz</a>, <a href="/wiki/John_Edward_Sulston" title="John Edward Sulston">Sulston</a> · 2003: <a href="/wiki/Paul_Lauterbur" title="Paul Lauterbur">Lauterbur</a>, <a href="/wiki/Peter_Mansfield" title="Peter Mansfield">Mansfield</a> · 2004: <a href="/wiki/Richard_Axel" title="Richard Axel">Axel</a>, <a href="/wiki/Linda_B._Buck" title="Linda B. Buck">Buck</a> · 2005: <a href="/wiki/Barry_Marshall" title="Barry Marshall">Marshall</a>, <a href="/wiki/Robin_Warren" title="Robin Warren">Warren</a> · 2006: <a href="/wiki/Andrew_Fire" title="Andrew Fire">Fire</a>, <a href="/wiki/Craig_Mello" title="Craig Mello">Mello</a> · 2007: <a href="/wiki/Mario_Capecchi" title="Mario Capecchi">Capecchi</a>, <a href="/wiki/Oliver_Smithies" title="Oliver Smithies">Smithies</a>, <a href="/wiki/Martin_Evans" title="Martin Evans">Evans</a> · 2008: <a class="mw-selflink selflink">Zur Hausen</a>, <span class="nowrap"><a href="/wiki/Fran%C3%A7oise_Barr%C3%A9-Sinoussi" title="Françoise Barré-Sinoussi">Barré–Sinoussi</a>,</span> <a href="/wiki/Luc_Montagnier" title="Luc Montagnier">Montagnier</a> · 2009: <a href="/wiki/Elizabeth_Blackburn" title="Elizabeth Blackburn">Blackburn</a>, <a href="/wiki/Carol_Greider" title="Carol Greider">Greider</a>, <a href="/wiki/Jack_Szostak" title="Jack Szostak">Szostak</a> · 2010: <a href="/wiki/Robert_Edwards_(fysioloog)" title="Robert Edwards (fysioloog)">Edwards</a> · 2011: <a href="/wiki/Bruce_Beutler" title="Bruce Beutler">Beutler</a>, <a href="/wiki/Jules_Hoffmann" title="Jules Hoffmann">Hoffmann</a>, <a href="/wiki/Ralph_Steinman" title="Ralph Steinman">Steinman</a> · 2012: <a href="/wiki/John_Gurdon" title="John Gurdon">Gurdon</a>, <a href="/wiki/Shinya_Yamanaka" title="Shinya Yamanaka">Yamanaka</a> · 2013: <a href="/wiki/James_Rothman" title="James Rothman">Rothman</a>, <a href="/wiki/Randy_Schekman" title="Randy Schekman">Schekman</a>, <a href="/wiki/Thomas_S%C3%BCdhof" title="Thomas Südhof">Südhof</a> · 2014: <a href="/wiki/John_O%27Keefe" title="John O'Keefe">O'Keefe</a>, <a href="/wiki/May-Britt_Moser" title="May-Britt Moser">Moser</a>, <a href="/wiki/Edvard_Moser" title="Edvard Moser">Moser</a> · 2015: <a href="/wiki/William_C._Campbell_(wetenschapper)" title="William C. Campbell (wetenschapper)">Campbell</a>, <a href="/wiki/Satoshi_Omura" title="Satoshi Omura">Omura</a>, <a href="/wiki/Youyou_Tu" title="Youyou Tu">Tu</a> · 2016: <a href="/wiki/Yoshinori_Osumi" title="Yoshinori Osumi">Osumi</a> · 2017: <a href="/wiki/Jeffrey_C._Hall" title="Jeffrey C. Hall">Hall</a>, <a href="/wiki/Michael_Rosbash" title="Michael Rosbash">Rosbash</a>, <a href="/wiki/Michael_W._Young" title="Michael W. Young">Young</a> · 2018: <a href="/wiki/James_Allison" title="James Allison">Allison</a>, <a href="/wiki/Tasuku_Honjo" title="Tasuku Honjo">Honjo</a> · 2019: <a href="/wiki/William_Kaelin_jr." title="William Kaelin jr.">Kaelin</a>, <a href="/wiki/Peter_J._Ratcliffe" title="Peter J. Ratcliffe">Ratcliffe</a>, <a href="/wiki/Gregg_L._Semenza" title="Gregg L. Semenza">Semenza</a> · 2020: <a href="/wiki/Harvey_J._Alter" title="Harvey J. Alter">Alter</a>, <a href="/wiki/Michael_Houghton" title="Michael Houghton">Houghton</a>, <a href="/wiki/Charles_M._Rice" title="Charles M. Rice">Rice</a> · 2021: <a href="/wiki/David_Julius" title="David Julius">Julius</a>, <a href="/wiki/Ardem_Patapoutian" title="Ardem Patapoutian">Patapoutian</a> · 2022: <a href="/wiki/Svante_P%C3%A4%C3%A4bo" title="Svante Pääbo">Pääbo</a> · 2023: <a href="/wiki/Katalin_Karik%C3%B3" title="Katalin Karikó">Karikó</a>, <a href="/wiki/Drew_Weissman" title="Drew Weissman">Weissman</a> · 2024: <a href="/wiki/Victor_Ambros" title="Victor Ambros">Ambros</a>, <a href="/wiki/Gary_Ruvkun" title="Gary Ruvkun">Ruvkun</a> </p> </div></div> <div class="interProject commons mw-list-item" style="display:none;"><a href="https://commons.wikimedia.org/wiki/Category:Harald_zur_Hausen#mw-subcategories" class="extiw" title="commons:Category:Harald zur Hausen">Mediabestanden</a></div> <div class="interProjectTemplate interProject-groot toccolours" style="display:flex; gap:1em; align-items:center; clear:both; margin:1em 0 -0.5em 0;"> <div style="min-width:max-content;"><span class="noviewer noresize" typeof="mw:File"><a href="/wiki/Bestand:Commons-logo.svg" class="mw-file-description"><img alt="" src="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/25px-Commons-logo.svg.png" decoding="async" width="25" height="34" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/38px-Commons-logo.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/4/4a/Commons-logo.svg/50px-Commons-logo.svg.png 2x" data-file-width="1024" data-file-height="1376" /></a></span></div> <div>Zie de categorie <i><b><a href="https://commons.wikimedia.org/wiki/Category:Harald_zur_Hausen#mw-subcategories" class="extiw" title="commons:Category:Harald zur Hausen">Harald zur Hausen</a></b></i> van <a href="/wiki/Wikimedia_Commons" title="Wikimedia Commons">Wikimedia Commons</a> voor mediabestanden over dit onderwerp.</div> </div> <!-- NewPP limit report Parsed by mw‐web.eqiad.main‐74878d4b‐59nwf Cached time: 20241204205759 Cache expiry: 2592000 Reduced expiry: false Complications: [] CPU time usage: 0.200 seconds Real time usage: 0.313 seconds Preprocessor visited node count: 2175/1000000 Post‐expand include size: 43962/2097152 bytes Template argument size: 15396/2097152 bytes Highest expansion depth: 18/100 Expensive parser function count: 2/500 Unstrip recursion depth: 1/20 Unstrip post‐expand size: 7131/5000000 bytes Lua time usage: 0.029/10.000 seconds Lua memory usage: 1005256/52428800 bytes Number of Wikibase entities loaded: 1/400 --> <!-- Transclusion expansion time report (%,ms,calls,template) 100.00% 221.619 1 -total 35.41% 78.479 1 Sjabloon:Appendix 31.54% 69.909 1 Sjabloon:Infobox_Nobelprijswinnaar 29.95% 66.370 1 Sjabloon:Infobox_generiek 21.82% 48.350 1 Sjabloon:References 14.93% 33.078 1 Sjabloon:Bibliografische_informatie 13.31% 29.499 1 Sjabloon:Wikidata_afbeelding 9.48% 21.001 1 Sjabloon:Commonscat 9.12% 20.215 2 Sjabloon:Citeer_web 7.17% 15.886 1 Sjabloon:Zusterproject_box --> <!-- Saved in parser cache with key nlwiki:pcache:1425480:|#|:idhash:dateformat=default and timestamp 20241204205759 and revision id 64787487. Rendering was triggered because: page-view --> </div><!--esi <esi:include src="/esitest-fa8a495983347898/content" /> --><noscript><img src="https://login.wikimedia.org/wiki/Special:CentralAutoLogin/start?useformat=desktop&type=1x1&usesul3=0" alt="" width="1" height="1" style="border: none; position: absolute;"></noscript> <div class="printfooter" data-nosnippet="">Overgenomen van "<a dir="ltr" href="https://nl.wikipedia.org/w/index.php?title=Harald_zur_Hausen&oldid=64787487">https://nl.wikipedia.org/w/index.php?title=Harald_zur_Hausen&oldid=64787487</a>"</div></div> <div id="catlinks" class="catlinks" data-mw="interface"><div id="mw-normal-catlinks" class="mw-normal-catlinks"><a href="/wiki/Categorie:Alles" title="Categorie:Alles">Categorieën</a>: <ul><li><a href="/wiki/Categorie:Duits_hoogleraar" title="Categorie:Duits hoogleraar">Duits hoogleraar</a></li><li><a href="/wiki/Categorie:Duits_medicus" title="Categorie:Duits medicus">Duits medicus</a></li><li><a href="/wiki/Categorie:Duits_viroloog" title="Categorie:Duits viroloog">Duits viroloog</a></li><li><a href="/wiki/Categorie:Winnaar_van_de_Nobelprijs_voor_Fysiologie_of_Geneeskunde" title="Categorie:Winnaar van de Nobelprijs voor Fysiologie of Geneeskunde">Winnaar van de Nobelprijs voor Fysiologie of Geneeskunde</a></li><li><a href="/wiki/Categorie:Baarmoederhalskanker" title="Categorie:Baarmoederhalskanker">Baarmoederhalskanker</a></li><li><a href="/wiki/Categorie:20e-eeuws_medicus" title="Categorie:20e-eeuws medicus">20e-eeuws medicus</a></li></ul></div></div> </div> </main> </div> <div class="mw-footer-container"> <footer id="footer" class="mw-footer" > <ul id="footer-info"> <li id="footer-info-lastmod"> Deze pagina is voor het laatst bewerkt op 25 jul 2023 om 22:02.</li> <li id="footer-info-copyright">De tekst is beschikbaar onder de licentie <a rel="nofollow" class="external text" href="//creativecommons.org/licenses/by-sa/4.0/deed.nl">Creative Commons Naamsvermelding/Gelijk delen</a>, er kunnen aanvullende voorwaarden van toepassing zijn. Zie de <a class="external text" href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Terms_of_Use/nl">gebruiksvoorwaarden</a> voor meer informatie.<br /> Wikipedia® is een geregistreerd handelsmerk van de <a rel="nofollow" class="external text" href="https://www.wikimediafoundation.org">Wikimedia Foundation, Inc.</a>, een organisatie zonder winstoogmerk.</li> </ul> <ul id="footer-places"> <li id="footer-places-privacy"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Privacy_policy">Privacybeleid</a></li> <li id="footer-places-about"><a href="/wiki/Wikipedia">Over Wikipedia</a></li> <li id="footer-places-disclaimers"><a href="/wiki/Wikipedia:Algemeen_voorbehoud">Disclaimers</a></li> <li id="footer-places-wm-codeofconduct"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Universal_Code_of_Conduct">Gedragscode</a></li> <li id="footer-places-developers"><a href="https://developer.wikimedia.org">Ontwikkelaars</a></li> <li id="footer-places-statslink"><a href="https://stats.wikimedia.org/#/nl.wikipedia.org">Statistieken</a></li> <li id="footer-places-cookiestatement"><a href="https://foundation.wikimedia.org/wiki/Special:MyLanguage/Policy:Cookie_statement">Cookieverklaring</a></li> <li id="footer-places-mobileview"><a href="//nl.m.wikipedia.org/w/index.php?title=Harald_zur_Hausen&mobileaction=toggle_view_mobile" class="noprint stopMobileRedirectToggle">Mobiele weergave</a></li> </ul> <ul id="footer-icons" class="noprint"> <li id="footer-copyrightico"><a href="https://wikimediafoundation.org/" class="cdx-button cdx-button--fake-button cdx-button--size-large cdx-button--fake-button--enabled"><img src="/static/images/footer/wikimedia-button.svg" width="84" height="29" alt="Wikimedia Foundation" loading="lazy"></a></li> <li id="footer-poweredbyico"><a href="https://www.mediawiki.org/" class="cdx-button cdx-button--fake-button cdx-button--size-large cdx-button--fake-button--enabled"><img src="/w/resources/assets/poweredby_mediawiki.svg" alt="Powered by MediaWiki" width="88" height="31" loading="lazy"></a></li> </ul> </footer> </div> </div> </div> <div class="vector-settings" id="p-dock-bottom"> <ul></ul> </div><script>(RLQ=window.RLQ||[]).push(function(){mw.config.set({"wgHostname":"mw-web.codfw.main-5ccf8d5c58-fzhd7","wgBackendResponseTime":141,"wgPageParseReport":{"limitreport":{"cputime":"0.200","walltime":"0.313","ppvisitednodes":{"value":2175,"limit":1000000},"postexpandincludesize":{"value":43962,"limit":2097152},"templateargumentsize":{"value":15396,"limit":2097152},"expansiondepth":{"value":18,"limit":100},"expensivefunctioncount":{"value":2,"limit":500},"unstrip-depth":{"value":1,"limit":20},"unstrip-size":{"value":7131,"limit":5000000},"entityaccesscount":{"value":1,"limit":400},"timingprofile":["100.00% 221.619 1 -total"," 35.41% 78.479 1 Sjabloon:Appendix"," 31.54% 69.909 1 Sjabloon:Infobox_Nobelprijswinnaar"," 29.95% 66.370 1 Sjabloon:Infobox_generiek"," 21.82% 48.350 1 Sjabloon:References"," 14.93% 33.078 1 Sjabloon:Bibliografische_informatie"," 13.31% 29.499 1 Sjabloon:Wikidata_afbeelding"," 9.48% 21.001 1 Sjabloon:Commonscat"," 9.12% 20.215 2 Sjabloon:Citeer_web"," 7.17% 15.886 1 Sjabloon:Zusterproject_box"]},"scribunto":{"limitreport-timeusage":{"value":"0.029","limit":"10.000"},"limitreport-memusage":{"value":1005256,"limit":52428800}},"cachereport":{"origin":"mw-web.eqiad.main-74878d4b-59nwf","timestamp":"20241204205759","ttl":2592000,"transientcontent":false}}});});</script> <script type="application/ld+json">{"@context":"https:\/\/schema.org","@type":"Article","name":"Harald zur Hausen","url":"https:\/\/nl.wikipedia.org\/wiki\/Harald_zur_Hausen","sameAs":"http:\/\/www.wikidata.org\/entity\/Q75847","mainEntity":"http:\/\/www.wikidata.org\/entity\/Q75847","author":{"@type":"Organization","name":"Bijdragers aan Wikimedia-projecten"},"publisher":{"@type":"Organization","name":"Wikimedia Foundation, Inc.","logo":{"@type":"ImageObject","url":"https:\/\/www.wikimedia.org\/static\/images\/wmf-hor-googpub.png"}},"datePublished":"2008-10-06T15:36:00Z","dateModified":"2023-07-25T21:02:42Z","image":"https:\/\/upload.wikimedia.org\/wikipedia\/commons\/9\/99\/Harald_zur_Hausen_03.jpg","headline":"Duits viroloog (1936\u20132023)"}</script> </body> </html>