CINXE.COM

Cell Biology (Tetrad) (PhD) | UC San Francisco

<!doctype html> <html class="no-js" xml:lang="en" lang="en" dir="ltr"> <head> <meta http-equiv="X-UA-Compatible" content="IE=Edge" /> <title>Cell Biology (Tetrad) (PhD) | UC San Francisco</title> <meta http-equiv="Content-Type" content="text/html; charset=utf-8" /> <link rel="search" type="application/opensearchdescription+xml" href="/search/opensearch.xml" title="Catalog" /> <meta name="viewport" content="width=device-width, initial-scale=1.0, minimum-scale=1.0" /> <link href="/images/favicon.ico" rel="shortcut icon" /> <link rel="stylesheet" type="text/css" href="/css/reset.css" /> <link href="https://fonts.googleapis.com/css?family=Roboto:400,400i,500,500i,700,700i" rel="stylesheet"> <link rel="stylesheet" type="text/css" href="/fonts/font-awesome/font-awesome.min.css" /> <link rel="stylesheet" type="text/css" href="/css/courseleaf.css" /> <link rel="stylesheet" type="text/css" href="/css/screen.css?v=2024-2025" media="screen" /> <link rel="stylesheet" type="text/css" href="/css/print.css" media="print" /> <script type="text/javascript" src="/js/jquery.js"></script> <script type="text/javascript" src="/js/lfjs.js"></script> <script type="text/javascript" src="/js/lfjs_any.js"></script> <link rel="stylesheet" type="text/css" href="/js/lfjs.css" /> <script type="text/javascript" src="/js/courseleaf.js"></script> <script type="text/javascript" src="/js/custom.js?v=2024-2025"></script> </head> <body class=""> <nav aria-label="Skip content menu" class="accessible"> <div class="accessible-menu"> <ul> <li><a href="#contentarea" rel="section">Skip to Content</a></li> <li><a href="/azindex/">AZ Index</a></li> <li><a href="/">Catalog Home</a></li> </ul> </div> </nav> <header id="header"> <section class="header-universal"> <div class="wrap"> <p class="header-universal__title">University of California San Francisco</p> <a class="header-universal__give" href="https://giving.ucsf.edu/">Give to UCSF</a> </div> </section> <section class="header-main"> <div class="wrap"> <h1 id="site-title"><a href="/" rel="home">General Catalog</a></h1> <div class="mobile-toggles"> <button type="button" class="toggle-search" aria-controls="cat-search" aria-expanded="false" data-toggle="#cat-search"> <span class="sr-only">toggle search</span> <span class="toggle-search__icon" aria-hidden="true"></span> </button> <button id="hamburger" aria-controls="navigation" data-toggle="#navigation" aria-expanded="false"> <span class="sr-only">Toggle menu</span> <span class="burgertime" aria-hidden="true"> <span class="icon"></span> </span> </button> </div> <nav id="navigation" aria-label="Secondary" aria-hidden="false"> <ul id="main-nav" class="top clearfix"> <li class="isparent"> <button type="button">Courses</button> <ul> <li><a href="/course-catalog/">Course Listings</a></li> <li><a href="/course-search/">Search Courses</a></li> </ul> </li> <li><a href="/programs/">Academic Programs</a></li> <li class="isparent"> <button type="button">Schools and Programs</button> <ul> <li><a href="/schools/school-dentistry/">School of Dentistry</a></li> <li><a href="/schools/school-medicine/">School of Medicine</a></li> <li><a href="/schools/school-nursing/">School of Nursing</a></li> <li><a href="/schools/school-pharmacy/">School of Pharmacy</a></li> <li><a href="/schools/graduate-division/">Graduate Division</a></li> <li><a href="/schools/institute-global-health-sciences/">Institute for Global Health Sciences</a></li> </ul> </li> <li><a href="/university/">University Support Offices</a></li> <li><a href="/archive/">Archive</a></li> <li class="no-mobile"> <button type="button" class="toggle-search" aria-controls="cat-search" aria-expanded="false" data-toggle="#cat-search"> <span class="sr-only">open search</span> <span class="toggle-search__icon" aria-hidden="true"></span> </button> </li> </ul> <a class="header-main__give" href="https://giving.ucsf.edu/">Give to UCSF</a> </nav> <div class="search" id="cat-search"> <form action="/search/"> <label for="cat-search-term" class="sr-only">Search catalog</label> <input class="search-field" type="text" name="search" id="cat-search-term" placeholder="Search catalog..." /> <button class="search-button" type="submit"> <span class="sr-only">Submit search</span> </button> <!-- <input type="hidden" name="caturl" value="/programs" /> --> </form> </div> </div> </section> </header> <section id="content-container"> <div class="wrap"> <div id="col-content"> <nav id="breadcrumb" aria-label="Breadcrumbs"> <ul><li><a href="/">Home</a><span class="crumbsep">&gt;</span></li><li><a href="/programs/">Academic&nbsp;Programs</a><span class="crumbsep">&gt;</span></li><li><span class="active">Cell Biology (Tetrad) (PhD)</span></li></ul> </nav> <main id="contentarea"> <header> <h1 class="page-title">Cell Biology (Tetrad) (PhD)</h1> </header> <script type="text/javascript"> //<![CDATA[ var defshow = "text"; var bodycontainer = "#body"; var validhashes = ",text,requirementstext,"; if(window.location.hash.length) { var hash = cleanHash(window.location.hash); if(validhashes.indexOf("," + hash + ",") != -1) defshow = hash; } var ss = document.styleSheets; var workss = null; for(var i=0; i < ss.length; i++) { var currmedia = ""; if (typeof ss[i].media == "string") currmedia = ss[i].media else if (typeof ss[i].media == "object" && ss[i].media.mediaText) currmedia = ss[i].media.mediaText; if((currmedia == "screen" || currmedia == "all") && (ss[i].href.indexOf("/") == 0 || ss[i].href.indexOf(document.location.host) >= 0)) { workss = ss[i]; break; } } if(workss && workss.insertRule) { workss.insertRule(".tab_content { display: none; }", workss.cssRules.length) workss.insertRule(".sectionheader { display: none; }", workss.cssRules.length) workss.insertRule("#" + defshow + "container { display: block; }", workss.cssRules.length) } else if(workss && workss.addRule) { workss.addRule(".tab_content", "display: none"); workss.addRule(".sectionheader", "display: none"); workss.addRule("#" + defshow + "container", "display: block"); } else { if (window.console) window.console.log('screen.css <link> needs attribute media="screen"'); } //]]> </script> <nav id="tabs" aria-label="Page content tabs"> <ul class="clearfix" role="tablist"> <li id="texttab" role="presentation"> <a href="#textcontainer" role="tab" aria-controls="textcontainer" onclick="return showSection('text');">Overview</a></li> <li id="requirementstexttab" role="presentation"> <a href="#requirementstextcontainer" role="tab" aria-controls="requirementstextcontainer" onclick="return showSection('requirementstext');" >Details</a></li> </ul> </nav> <div id="textcontainer" class="page_content tab_content" role="tabpanel" aria-labelledby="texttab"> <p><a href="https://tetrad.ucsf.edu/" target="_blank" rel="noopener">Visit program website.</a></p><p><a href="https://gradapplication.ucsf.edu/" target="_blank" rel="noopener" class="applyBtn">Apply now</a></p><p><strong>Degree Offered:</strong> PhD in Cell Biology<br/> <strong>Program Leadership:</strong><br/> Natalia Jura, PhD, Director<br/> David Toczyski, PhD, Co-Director<br/> <strong>Admissions Inquiries:</strong><br/> <a href="mailto:toni.hurley@ucsf.edu">Toni Hurley</a>, Program Coordinator<br/> <a href="mailto:danny.dam@ucsf.edu">Danny Dam</a>, Student Affairs Coordinator</p> <h2>Program Description</h2> <p>The Tetrad graduate program prepares students to pose and address fundamental research problems in modern biology. The program highly values curiosity-driven research that investigates challenging questions in life sciences conducted in a collegial and scientifically rigorous manner. The program places special emphasis on modern approaches within three core and interrelated areas listed below. The research conducted under the Tetrad umbrella encompasses a wide range of structural, molecular, cellular, physiological, and pathophysiological questions.</p> <p>The Tetrad program was among the first graduate programs nationwide to break down barriers between disciplines; emphasis on interdisciplinary research remains a guiding principle of the program. Hand-in-hand with promoting interdisciplinary research is the program's emphasis on fostering a strong sense of community and interactions among students and the program's cadre of internationally recognized scientists. Program activities include an annual retreat held in Lake Tahoe, a weekly seminar program featuring top scientists from around the world, and multiple other activities that provide student-faculty interactions in both formal and informal settings.</p> <p>All Tetrad graduate students go through the same admission process, take the same classes, and have the same requirements. Depending on the specific aspects of their thesis work, their PhD degree will be in one of the areas indicated below.</p> <h3>Faculty</h3> <p>Over 120 faculty members are associated with the Tetrad program, representing all of the basic science departments of UCSF, numerous research centers such as the Helen Diller Family Comprehensive Cancer Center, the Cardiovascular Research Institute at UCSF, and the Institute for Neurodegenerative Diseases, and numerous clinical departments.</p> <p>Tetrad is a member of the <a href="https://pibs.ucsf.edu/" target="_blank">Program in Biological Sciences</a> (PIBS) at UCSF.</p> <h3>Sub-disciplines</h3> <ul> <li>Biochemistry and Molecular Biology</li> <li>Cell Biology</li> <li>Genetics</li> </ul> <p>The Tetrad program office is located at the Mission Bay campus, though some classes and labs affiliated with the program are at the Parnassus campus as well.</p> <p>The Tetrad program is offered by the UCSF Graduate Division, administered by the UCSF Department of Biochemistry and Biophysics, and delivered by faculty members in the UCSF schools of dentistry, medicine, and pharmacy.</p> <h2>Admission Requirements</h2> <p>Because of the interdisciplinary nature of the Tetrad Graduate Program, we expect that entering graduate students will have diverse undergraduate preparation. Students with backgrounds in anatomy, biochemistry, biology, chemistry, physics and related fields are welcome to apply, providing they have demonstrated a high level of academic proficiency (generally a grade point average of 3.0 or higher in relevant science courses). Evidence of exposure to scientific research, generally as participation in a research project during at least one year (preferably two or more years), is regarded as an important attribute of the successful applicant.</p> <h2>Learning Outcomes</h2> <ol> <li><em>Establish a foundation of knowledge in the fundamental principles underlying biological processes, from molecules to organisms</em>. This objective is enabled by 1st year coursework that emphasizes basic concepts, methods of discovery, and lifelong learning, followed in later years by individual thesis projects that are motivated by addressing fundamental questions of cellular or organismal function in mechanistic depth.</li> <li><em>Learn to critically evaluate the literature and assess the significance of a given biological question</em>. This objective is enabled by in-depth paper discussions and proposal writing within courses in the 1st year, journal clubs and qualifying exam in the 2nd year, and literature study throughout the thesis project.</li> <li><em>Learn to independently make original research contributions through rigorous experimental design, data analysis and interpretation</em>. This objective is enabled through activities such as solving problem sets and formulation of research proposals on new questions in the 1st year, the NSF proposal writing workshop and qualifying exam in the 2nd year, and mentorship by thesis mentor and thesis committee during the thesis project.</li> <li><em>Learn to work across disciplines to leverage the strengths of collaborators from different scientific backgrounds</em>. This objective is enabled by the multi-disciplinary nature of the first-year course work, peer-peer learning that leverages the diversity of scientific backgrounds within each class, the multidisciplinary nature of the research programs of most faculty members, the presence of students from different graduate programs in a given thesis lab and a strong ethos of collaboration amongst the different laboratories at UCSF.</li> <li><em>Learn how to promote and support an inclusive scientific environment</em>. This objective is enabled through training activities that include diversity workshops for 1st year students, collaboration with faculty who participate in diversity and inclusion-promoting campus organizations, opportunities to mentor summer research students from under-represented backgrounds, and participation in outreach events.</li> <li><em>Establish values that drive the responsible and ethical practice of science</em>. This fundamental objective is enabled through examples of best scientific practices threaded throughout the 1st year courses, two focused courses in the Responsible Conduct of Research in the 1st and 5th years, journal clubs, seminar series, day-to-day mentor-mentee interactions and thesis committee meetings.</li> <li><em>Develop skills for effective oral and written communication of complex scientific ideas and findings</em>. This objective is enabled through the writing and oral defense of research proposals in two 1st year courses, journal club presentations, writing and oral defense of a qualifying exam proposal, writing an NSF proposal, presentations in lab meetings and thesis committee meetings, presentations at scientific meetings within and outside UCSF, and the program requirement for thesis completion of writing at least one manuscript on original work for publication in a peer-reviewed journal.</li> </ol> <h2>Additional Information</h2> <h3>Program Faculty</h3> <ul> <li><em>Find a <a href="https://tetrad.ucsf.edu/faculty" target="_blank">program faculty list</a> on the program website.</em></li> </ul> <h3>Career Outcomes</h3> <ul> <li><em>Find <a href="https://graduate.ucsf.edu/program-statistics#career" target="_blank">career outcomes and other data on PhD programs</a> on the Graduate Division website.</em></li> </ul></div><!--end #textcontainer --> <div id="requirementstextcontainer" class="page_content tab_content" role="tabpanel" aria-labelledby="requirementstexttab" aria-hidden="true"> <a name="requirementstext"></a><h2>Degree Requirements</h2> <h3>Doctoral</h3> <ul> <li>All core courses and required activities taken and passed</li> <li>Six quarters in residence including a minimum of three registered quarters after advancement to candidacy</li> <li>Pass qualifying examination</li> <li>Complete and submit the dissertation based on the current Tetrad and Graduate Division guidelines</li> <li>For additional details, please see:聽<a href="https://graduate.ucsf.edu/phd-degree" target="_blank">graduate.ucsf.edu/phd-degree</a></li> </ul> <h2>Core Courses</h2> <table pdfnorepeatheader="true" class="sc_plangrid"> <caption>Plan of Study Grid</caption><colgroup> <col pdfwidth="28" class="codecol"/> <col pdfwidth="57" class="titlecol"/> <col pdfwidth="15" class="hourscol"/> </colgroup> <tr class="plangridyear"><th id="2yFRCD_plangrid1_0" class="year" colspan="2">Year 1</th><th id="2yFRCD_plangrid1_hoursheader" scope="col" class="hourscol">Units</th></tr> <tr class="even"><td headers="2yFRCD_plangrid1_0 " class="codecol"><a href="/search/?P=BIOCHEM%20200A" title="BIOCHEM聽200A" class="bubblelink code" onclick="return showCourse(this, 'BIOCHEM 200A');">BIOCHEM聽200A</a></td> <td headers="2yFRCD_plangrid1_0 " class="titlecol">Structure of Macromolecules</td> <td headers="2yFRCD_plangrid1_0 2yFRCD_plangrid1_hoursheader" class="hourscol">3</td> </tr> <tr class="odd"><td headers="2yFRCD_plangrid1_0 " class="codecol"><a href="/search/?P=CELL%20BIOL%20245" title="CELL聽BIOL聽245" class="bubblelink code" onclick="return showCourse(this, 'CELL BIOL 245');">CELL聽BIOL聽245</a></td> <td headers="2yFRCD_plangrid1_0 " class="titlecol">Cell &amp; Developmental Biology</td> <td headers="2yFRCD_plangrid1_0 2yFRCD_plangrid1_hoursheader" class="hourscol">4</td> </tr> <tr class="even"><td headers="2yFRCD_plangrid1_0 " class="codecol"><a href="/search/?P=GRAD%20202" title="GRAD聽202" class="bubblelink code" onclick="return showCourse(this, 'GRAD 202');">GRAD聽202</a></td> <td headers="2yFRCD_plangrid1_0 " class="titlecol">Racism in Science</td> <td headers="2yFRCD_plangrid1_0 2yFRCD_plangrid1_hoursheader" class="hourscol">3</td> </tr> <tr class="odd"><td headers="2yFRCD_plangrid1_0 " class="codecol"><a href="/search/?P=BIOCHEM%20201A" title="BIOCHEM聽201A" class="bubblelink code" onclick="return showCourse(this, 'BIOCHEM 201A');">BIOCHEM聽201A</a></td> <td headers="2yFRCD_plangrid1_0 " class="titlecol">Biological Regulatory Mechanisms</td> <td headers="2yFRCD_plangrid1_0 2yFRCD_plangrid1_hoursheader" class="hourscol">4</td> </tr> <tr class="even"><td headers="2yFRCD_plangrid1_0 " class="codecol"><a href="/search/?P=GENETICS%20200A" title="GENETICS聽200A" class="bubblelink code" onclick="return showCourse(this, 'GENETICS 200A');">GENETICS聽200A</a></td> <td headers="2yFRCD_plangrid1_0 " class="titlecol">Principles of Genetics</td> <td headers="2yFRCD_plangrid1_0 2yFRCD_plangrid1_hoursheader" class="hourscol">3</td> </tr> <tr class="odd"><td headers="2yFRCD_plangrid1_0 " colspan="2" class="codecol"><span class="comment">Various: 2-3 minicourses</span></td> <td headers="2yFRCD_plangrid1_0 2yFRCD_plangrid1_hoursheader" class="hourscol">2-3</td> </tr> <tr class="even"><td headers="2yFRCD_plangrid1_0 " class="codecol"><a href="/search/?P=GRAD%20214" title="GRAD聽214" class="bubblelink code" onclick="return showCourse(this, 'GRAD 214');">GRAD聽214</a></td> <td headers="2yFRCD_plangrid1_0 " class="titlecol">Responsible Conduct of Research and Rigor &amp; Reproducibility</td> <td headers="2yFRCD_plangrid1_0 2yFRCD_plangrid1_hoursheader" class="hourscol">1.5</td> </tr> <tr class="odd"><td headers="2yFRCD_plangrid1_0 " class="codecol"><a href="/search/?P=BIOCHEM%20215" title="BIOCHEM聽215" class="bubblelink code" onclick="return showCourse(this, 'BIOCHEM 215');">BIOCHEM聽215</a></td> <td headers="2yFRCD_plangrid1_0 " class="titlecol">Laboratory Rotation (<span class="comment">Taken all terms</span>)</td> <td headers="2yFRCD_plangrid1_0 2yFRCD_plangrid1_hoursheader" class="hourscol">9</td> </tr> <tr class="even"><td headers="2yFRCD_plangrid1_0 " class="codecol"><a href="/search/?P=BIOCHEM%20220" title="BIOCHEM聽220" class="bubblelink code" onclick="return showCourse(this, 'BIOCHEM 220');">BIOCHEM聽220</a></td> <td headers="2yFRCD_plangrid1_0 " class="titlecol">Biochemistry Basic Science Seminar Series</td> <td headers="2yFRCD_plangrid1_0 2yFRCD_plangrid1_hoursheader" class="hourscol">1</td> </tr> <tr class="odd"><td headers="2yFRCD_plangrid1_0 " class="codecol"><a href="/search/?P=BIOCHEM%20221" title="BIOCHEM聽221" class="bubblelink code" onclick="return showCourse(this, 'BIOCHEM 221');">BIOCHEM聽221</a></td> <td headers="2yFRCD_plangrid1_0 " class="titlecol">Selected Topics (<span class="comment">Journal Club, taken all terms</span>)</td> <td headers="2yFRCD_plangrid1_0 2yFRCD_plangrid1_hoursheader" class="hourscol">3</td> </tr> <tr class="plangridsum even"><td>聽</td><td headers="2yFRCD_plangrid1_0 " class="titlecol">Units</td><td headers="2yFRCD_plangrid1_0 2yFRCD_plangrid1_hoursheader" class="hourscol">33.5-34.5</td></tr> <tr class="plangridyear"><th id="2yFRCD_plangrid1_1" class="year" colspan="3">Year 2</th></tr> <tr class="odd"><td headers=" " class="codecol"><a href="/search/?P=BIOCHEM%20221" title="BIOCHEM聽221" class="bubblelink code" onclick="return showCourse(this, 'BIOCHEM 221');">BIOCHEM聽221</a></td> <td headers=" " class="titlecol">Selected Topics (<span class="comment">Journal Club, taken all terms</span>)</td> <td headers=" 2yFRCD_plangrid1_hoursheader" class="hourscol">3</td> </tr> <tr class="even"><td headers=" " class="codecol"><a href="/search/?P=CELL%20BIOL%20250" title="CELL聽BIOL聽250" class="bubblelink code" onclick="return showCourse(this, 'CELL BIOL 250');">CELL聽BIOL聽250</a></td> <td headers=" " class="titlecol">Research (<span class="comment">Taken all terms</span>)</td> <td headers=" 2yFRCD_plangrid1_hoursheader" class="hourscol">24</td> </tr> <tr class="odd"><td headers=" " colspan="2" class="codecol"><span class="comment">Any approved electives</span></td> <td headers=" 2yFRCD_plangrid1_hoursheader" class="hourscol">聽</td> </tr> <tr class="plangridsum even"><td>聽</td><td headers=" " class="titlecol">Units</td><td headers=" 2yFRCD_plangrid1_hoursheader" class="hourscol">27</td></tr> <tr class="plangridyear"><th id="2yFRCD_plangrid1_2" class="year" colspan="3">Year 3 and Beyond</th></tr> <tr class="odd"><td headers=" " colspan="2" class="codecol"><span class="comment">Any approved electives</span></td> <td headers=" 2yFRCD_plangrid1_hoursheader" class="hourscol">聽</td> </tr> <tr class="even"><td headers=" " class="codecol"><a href="/search/?P=CELL%20BIOL%20250" title="CELL聽BIOL聽250" class="bubblelink code" onclick="return showCourse(this, 'CELL BIOL 250');">CELL聽BIOL聽250</a></td> <td headers=" " class="titlecol">Research (<span class="comment">Taken all terms</span>)</td> <td headers=" 2yFRCD_plangrid1_hoursheader" class="hourscol">24</td> </tr> <tr class="plangridsum odd"><td>聽</td><td headers=" " class="titlecol">Units</td><td headers=" 2yFRCD_plangrid1_hoursheader" class="hourscol">24</td></tr> <tr class="plangridtotal lastrow even"><td>聽</td><td>Total Units</td><td headers="2yFRCD_plangrid1_hoursheader" class="hourscol">84.5-85.5</td></tr> </table> <h3>Approved Electives</h3> <table class="sc_courselist"> <caption class="hidden noscript">Course List</caption> <colgroup> <col class="codecol"/> <col class="titlecol"/> <col align="char" char="." class="hourscol"/> </colgroup> <thead> <tr class="hidden noscript"> <th scope="col">Code</th> <th scope="col">Title</th> <th scope="col" class="hourscol">Units</th> </tr> </thead> <tbody> <tr class="even firstrow"><td class="codecol"><a href="/search/?P=BIOCHEM%20210" title="BIOCHEM聽210" class="bubblelink code" onclick="return showCourse(this, 'BIOCHEM 210');">BIOCHEM聽210</a></td><td>Special Topics</td><td class="hourscol">3</td></tr> <tr class="odd"><td class="codecol"><a href="/search/?P=BIOCHEM%20241" title="BIOCHEM聽241" class="bubblelink code" onclick="return showCourse(this, 'BIOCHEM 241');">BIOCHEM聽241</a></td><td>Startup 101</td><td class="hourscol">3</td></tr> <tr class="even"><td class="codecol"><a href="/search/?P=BIOMED%20SCI%20270" title="BIOMED聽SCI聽270" class="bubblelink code" onclick="return showCourse(this, 'BIOMED SCI 270');">BIOMED聽SCI聽270</a></td><td>Special Topics in Biomedical Sciences</td><td class="hourscol">3</td></tr> <tr class="odd"><td class="codecol"><a href="/search/?P=BIO%20MD%20INF%20219" title="BIO聽MD聽INF聽219" class="bubblelink code" onclick="return showCourse(this, 'BIO MD INF 219');">BIO聽MD聽INF聽219</a></td><td>Special Topics in Bioinformatics</td><td class="hourscol">3</td></tr> <tr class="even"><td class="codecol"><a href="/search/?P=BIOPHYSICS%20219" title="BIOPHYSICS聽219" class="bubblelink code" onclick="return showCourse(this, 'BIOPHYSICS 219');">BIOPHYSICS聽219</a></td><td>Special Topics in Biophysics</td><td class="hourscol">3</td></tr> <tr class="odd"><td class="codecol"><a href="/search/?P=CHEMISTRY%20219" title="CHEMISTRY聽219" class="bubblelink code" onclick="return showCourse(this, 'CHEMISTRY 219');">CHEMISTRY聽219</a></td><td>Special Topics in Basic and Translational Chemical Biology</td><td class="hourscol">3</td></tr> <tr class="even"><td class="codecol"><a href="/search/?P=GRAD%20210" title="GRAD聽210" class="bubblelink code" onclick="return showCourse(this, 'GRAD 210');">GRAD聽210</a></td><td>Justice, Equity, Diversity and Inclusion Academic Leadership (with approval from the Graduate Division)</td><td class="hourscol">4</td></tr> <tr class="odd"><td class="codecol"><a href="/search/?P=DEV%20STMCEL%20270" title="DEV聽STMCEL聽270" class="bubblelink code" onclick="return showCourse(this, 'DEV STMCEL 270');">DEV聽STMCEL聽270</a></td><td>Special Topics in Developmental &amp; Stem Cell Biology</td><td class="hourscol">3</td></tr> <tr class="even"><td class="codecol"><a href="/search/?P=GRAD%20213" title="GRAD聽213" class="bubblelink code" onclick="return showCourse(this, 'GRAD 213');">GRAD聽213</a></td><td>Motivating INformed Decisions (MIND) Catalytic Course</td><td class="hourscol">2</td></tr> <tr class="odd"><td class="codecol"><a href="/search/?P=GRAD%20286" title="GRAD聽286" class="bubblelink code" onclick="return showCourse(this, 'GRAD 286');">GRAD聽286</a></td><td>GSICE Curricular Practicum</td><td class="hourscol">1</td></tr> <tr class="even"><td class="codecol"><a href="/search/?P=GRAD%20219A" title="GRAD聽219A" class="bubblelink code" onclick="return showCourse(this, 'GRAD 219A');">GRAD聽219A</a></td><td>Special topics in racism and social justice in science</td><td class="hourscol">3</td></tr> <tr class="odd"><td class="codecol"><a href="/search/?P=GRAD%20219B" title="GRAD聽219B" class="bubblelink code" onclick="return showCourse(this, 'GRAD 219B');">GRAD聽219B</a></td><td>Special topics in racism and social justice in science</td><td class="hourscol">3</td></tr> <tr class="even"><td class="codecol"><a href="/search/?P=GRAD%20219C" title="GRAD聽219C" class="bubblelink code" onclick="return showCourse(this, 'GRAD 219C');">GRAD聽219C</a></td><td>Special topics in racism and social justice in science</td><td class="hourscol">3</td></tr> <tr class="odd"><td class="codecol"><a href="/search/?P=NEUROSCI%20219" title="NEUROSCI聽219" class="bubblelink code" onclick="return showCourse(this, 'NEUROSCI 219');">NEUROSCI聽219</a></td><td>Special Topics in Basic and Translational Neuroscience</td><td class="hourscol">3</td></tr> <tr class="even lastrow"><td colspan="2"><span class="courselistcomment">Other electives may be approved by program on a case-by-case basis</span></td><td class="hourscol"></td></tr> </tbody> </table> <h2>Non-course Core Requirements</h2> <ol> <li>Attendance and participation in Boot Camp: <ol> <li>We realize that our incoming students come from a wide variety of experimental backgrounds, so the first year begins with a week-long series of intensive hands-on courses on cutting-edge concepts and experimental techniques that will be useful in a modern biomedical research lab.</li> </ol> </li> <li>Passing qualifying exam</li> <li>Teaching Assistantship in year 2</li> <li>Submission/publication of first-author research paper</li> <li>Presentation of thesis talk</li> </ol> </div> </main> </div> <!-- end col-content --> <div id="col-nav"> <aside id="sidebar"> <div class="sidebar-item"> <h2 id="edition" class="sidebar-header"><a href="/">2024-2025 Edition</a></h2> <nav id="cl-menu" aria-label="Primary"> <ul class="nav levelzero" id="/"> <li><a href="/welcome/">Welcome from the Chancellor</a></li> <li class="isparent"><a href="/schools/">Schools and Programs</a></li> <li class="active isparent"><a href="/programs/">Academic Programs</a> <ul class="nav levelone" id="/programs/"> <li><a href="/programs/atcr-certificate/">Advanced Training in Clinical Research (Certificate)</a></li> <li><a href="/programs/ai-comp-drug-discovery-development/">Artificial Intelligence and Computational Drug Discovery and Development (MS)</a></li> <li><a href="/programs/biochem-molec-bio-tetrad/">Biochemistry and Molecular Biology (Tetrad) (PhD)</a></li> <li><a href="/programs/bioengineering/">Bioengineering (PhD)</a></li> <li><a href="/programs/biological-medical-informatics/">Biological and Medical Informatics (PhD)</a></li> <li><a href="/programs/biomedical-imaging/">Biomedical Imaging (MS)</a></li> <li><a href="/programs/biomedical-sciences/">Biomedical Sciences (PhD)</a></li> <li><a href="/programs/biophysics/">Biophysics (PhD)</a></li> <li class="active self"><a href="#" onclick="return false;">Cell Biology (Tetrad) (PhD)</a></li> <li><a href="/programs/chemistry-chemical-biology/">Chemistry and Chemical Biology (PhD)</a></li> <li><a href="/programs/clinical-epidemiologic-research-mas/">Clinical and Epidemiologic Research (MAS)</a></li> <li><a href="/programs/computational-precision-health/">Computational Precision Health (PhD)</a></li> <li><a href="/programs/dentistry-post-graduate-certificate/">Dentistry Postgraduate Program -&#8203; Certificate</a></li> <li><a href="/programs/dscb/">Developmental and Stem Cell Biology (PhD)</a></li> <li><a href="/programs/doctor-dental-surgery/">Doctor of Dental Surgery (DDS)</a></li> <li><a href="/programs/medicine-md/">Doctor of Medicine (MD)</a></li> <li><a href="/programs/pharmd/">Doctor of Pharmacy (PharmD)</a></li> <li><a href="/programs/physical-therapy-dpt/">Doctor of Physical Therapy (DPT)</a></li> <li><a href="/programs/epidemiology-translational-science/">Epidemiology and Translational Science (PhD)</a></li> <li><a href="/programs/equity-in-brain-health-certificate/">Equity in Brain Health (Certificate)</a></li> <li><a href="/programs/genetic-counseling-ms/">Genetic Counseling (MS)</a></li> <li><a href="/programs/genetics-tetrad/">Genetics (Tetrad) (PhD)</a></li> <li><a href="/programs/global-health-sciences-ms/">Global Health Sciences (MS)</a></li> <li><a href="/programs/global-health-sciences/">Global Health Sciences (PhD)</a></li> <li><a href="/programs/health-data-sci-certificate/">Health Data Science (Certificate)</a></li> <li><a href="/programs/health-data-sci-ms/">Health Data Science (MS)</a></li> <li><a href="/programs/health-policy-and-law-ms/">Health Policy and Law (MS)</a></li> <li><a href="/programs/healthcare-administration-interprofessional-leadership-ms/">Healthcare Administration and Interprofessional Leadership (MS)</a></li> <li><a href="/programs/history-health-sciences-ma/">History of Health Sciences (MA)</a></li> <li><a href="/programs/history-health-sciences/">History of Health Sciences (PhD)</a></li> <li><a href="/programs/international-dentist-pathway-dds/">International Dentist Pathway (DDS)</a></li> <li><a href="/programs/interprofessional-health-postbac-dentistry/">Interprofessional Health Post-&#8203;Baccalaureate Certificate Program in Dentistry</a></li> <li><a href="/programs/interprofessional-health-postbac-medicine/">Interprofessional Health Post-&#8203;Baccalaureate Certificate Program in Medicine</a></li> <li><a href="/programs/interprofessional-health-postbac-pharmacy/">Interprofessional Health Post-&#8203;Baccalaureate Certificate Program in Pharmacy</a></li> <li><a href="/programs/medical-anthropology/">Medical Anthropology (PhD)</a></li> <li><a href="/programs/mstp/">Medical Scientist Training Program</a></li> <li><a href="/programs/neuroscience/">Neuroscience (PhD)</a></li> <li><a href="/programs/doctor-nursing-practice-dnp/">Nursing (DNP)</a></li> <li><a href="/programs/nursing-ms/">Nursing (MS)</a></li> <li><a href="/programs/phd-nursing/">Nursing (PhD)</a></li> <li><a href="/programs/nursing-mepn/">Nursing Master鈥檚 Entry Program</a></li> <li><a href="/programs/nursing-post-masters-certificate/">Nursing Post-&#8203;Master's Certificate Program</a></li> <li><a href="/programs/doctor-nursing-practice-post-bsn/">Nursing Practice (Post-&#8203;BSN)</a></li> <li><a href="/programs/oral-craniofacial-sciences-ms/">Oral and Craniofacial Sciences (MS)</a></li> <li><a href="/programs/oral-craniofacial-sciences-graduate/">Oral and Craniofacial Sciences (PhD)</a></li> <li><a href="/programs/oral-craniofacial-sciences-dds-phd/">Oral and Craniofacial Sciences DDS/&#8203;PhD Program</a></li> <li><a href="/programs/pharmaceutical-sciences-pharmacogenomics/">Pharmaceutical Sciences and Pharmacogenomics (PhD)</a></li> <li><a href="/programs/postgraduate-program-dental-public-health-certificate/">Postgraduate Program in Dental Public Health -&#8203; Certificate</a></li> <li><a href="/programs/postgraduate-program-oral-medicine-certificate/">Postgraduate Program in Oral Medicine -&#8203; Certificate</a></li> <li><a href="/programs/postgraduate-program-orthodontics-certificate/">Postgraduate Program in Orthodontics -&#8203; Certificate</a></li> <li><a href="/programs/pediatric-dentistry-certificate/">Postgraduate Program in Pediatric Dentistry -&#8203; Certificate</a></li> <li><a href="/programs/postgraduate-program-periodontology-certificate/">Postgraduate Program in Periodontology -&#8203; Certificate</a></li> <li><a href="/programs/rehabilitation-science-phd/">Rehabilitation Science (PhD)</a></li> <li><a href="/programs/sociology/">Sociology (PhD)</a></li> <li><a href="/programs/translational-medicine-mtm/">Translational Medicine (MTM)</a></li> <li><a href="/programs/ucsf-ucb-jmp/">UCSF/&#8203;UC Berkeley Joint Medical Program</a></li> </ul> </li> <li class="isparent"><a href="/residency-fellowship/">Residency and Fellowship Programs</a></li> <li class="isparent"><a href="/university/">University Support Offices</a></li> <li><a href="/academic-administrative-calendar/">Academic and Administrative Calendar</a></li> <li><a href="/accreditation/">Accreditation</a></li> <li class="isparent"><a href="/course-catalog/">Course Catalog</a></li> <li class="isparent"><a href="/course-search/">Search Courses</a></li> <li><a href="/catalogcontents/">Catalog Contents</a></li> <li><a href="/azindex/">Catalog A-&#8203;Z Index</a></li> </ul> </nav> </div> <div class="sidebar-item"> <button id="print-btn" href="#print-dialog" onclick="showPrintDialog(); return false;"> <span class="fa fa-print" aria-hidden="true"></span> Print Options </button> </div> </aside> </div> <!-- end col-nav --> </div> </section> <footer id="footer"> <div class="footer__bottom wrap"> <nav aria-label="footer" class="footer__nav"> <ul> <li><a href="//www.ucsf.edu/contact-us">Contact Us</a></li> <li><a href="https://websites.ucsf.edu/digital-accessibility">Accessibility</a></li> <li><a href="https://www.ucsf.edu/website-privacy-policy">Privacy Policy</a></li> <li><a href="https://websites.ucsf.edu/website-terms-use">Terms of Use </a></li> <li><a href="https://websites.ucsf.edu/azlist">A-Z Website List</a></li> </ul> </nav> <p class="footer__copyright">&copy; 2024-2025 The Regents of the University of California</p> </div> </footer> <aside id="print-dialog" aria-labelledby="dialog-title" class="screen" role="dialog"> <div class="print-header"> <button onclick="hidePrintDialog(); return false;" aria-controls="#print-dialog"> <span class="sr-only">Close this window</span> </button> <h2 id="dialog-title">Print Options</h2> </div> <div class="print-body"> <ul> <li> <p><a class="option-name" href="#" role="button" onclick="hidePrintDialog();window.print();return false">Send Page to Printer</a></p> <p class="option-desc">Print this page.</p> </li> <li><p><a class="option-name" href="/programs/cell-bio-tetrad/cell-bio-tetrad.pdf" role="button">Download Page (PDF)</a></p><p class="option-desc">The PDF will include all information unique to this page.</p></li><li><p><a class="option-name" href="/pdf/2024-25_UCSF_General_Catalog.pdf" target="_blank">Download Full 2024-25 Catalog</a></p><p class="option-desc">A PDF of the entire catalog.</p></li> </ul> </div> </aside> </body> </html>

Pages: 1 2 3 4 5 6 7 8 9 10