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method="get"><div class='flex'> <input placeholder="Search Catalog" aria-label="Search Catalog" class="typeahead-taxa-js-hook placeholder:text-sm" type="text" name="qq" /> <input type="hidden" name="searching_from_header" value="true" autocomplete="off" /> <button type="submit" class="-ml-8 z-10"><svg xmlns="http://www.w3.org/2000/svg" fill="none" viewBox="0 0 24 24" stroke-width="2.5" stroke="currentColor" class="text-ac-blue h-5 w-5"> <path stroke-linecap="round" stroke-linejoin="round" d="M21 21l-5.197-5.197m0 0A7.5 7.5 0 105.196 5.196a7.5 7.5 0 0010.607 10.607z"></path> </svg> </button></div> </form></li> </ul> <ul class='no-bullet [&>*]:py-1'> <li><a href="/references/latest_additions">References</a></li> <li><a href="/">Catalog</a></li> <li><a href="/users/sign_in">Login</a></li> <li><a href="/users/sign_up">Sign up</a></li> <li><a href="/feedbacks/new?page=catalog%2F430164">Suggest an edit</a></li> </ul> </div> </div> <div class='clear-both'></div> <div class='pb-12 hidden lg:block'> <div class='float-left'> An Online Catalog of the Ants of the World <span class='text-sm'>by Barry Bolton</span> </div> </div> </div> </header> <section class='px-4 pt-10 pb-32 min-h-[65vh]'> <div id='fireworks'></div> <div class='layout-container'> <div class='mb-8' id='breadcrumbs'> <div class='float-left'> <a href="/">Catalog</a> &rsaquo; <a class="valid" href="/catalog/429011">Formicidae</a> &rsaquo; <a class="valid" href="/catalog/430164">Leptanillinae</a> </div> <div class='clear-both'></div> <div class='clear-both'></div> </div> <div class='row'> <div class='medium-9'>  <span class='text-lg font-bold mr-1'><a data-controller="hover-preview" data-hover-preview-url-value="/catalog/430164/hover_preview.json" class="valid" href="/catalog/430164">Leptanillinae</a></span> Emery, 1910 <span class='text-sm'> valid </span>  <div class='mb-4'> <span class='font-bold mr-1' data-testid='catalog-protonym-name'> <a data-controller="hover-preview" data-hover-preview-url-value="/protonyms/155898/hover_preview.json" class="protonym" href="/protonyms/155898">Leptanillini</a> </span> <a data-controller="hover-preview" data-hover-preview-url-value="/references/124696/hover_preview.json" class="taxt-hover-reference" href="/references/124696">Emery, 1910b</a>: 32 Type-genus: <a data-controller="hover-preview" data-hover-preview-url-value="/catalog/430169/hover_preview.json" class="valid" href="/catalog/430169"><i>Leptanilla</i></a> </div>   <div class='mb-8'> </div>     <ul class='mb-8'> <li><span class="taxt-logged-in-only-gray-bold-notice">Leptanillidae</span> as family: <a data-controller="hover-preview" data-hover-preview-url-value="/references/122634/hover_preview.json" class="taxt-hover-reference" href="/references/122634">Bernard, 1951c</a>: 1053 [<span class="taxt-logged-in-only-gray-bold-notice">Leptanillidae</span>].</li> <li><a data-controller="hover-preview" data-hover-preview-url-value="/catalog/430164/hover_preview.json" class="valid" href="/catalog/430164">Leptanillinae</a> as subfamily of <a data-controller="hover-preview" data-hover-preview-url-value="/catalog/429011/hover_preview.json" class="valid" href="/catalog/429011">Formicidae</a>: <a data-controller="hover-preview" data-hover-preview-url-value="/references/130169/hover_preview.json" class="taxt-hover-reference" href="/references/130169">Wheeler, 1923g</a>: 335; <a data-controller="hover-preview" data-hover-preview-url-value="/references/129831/hover_preview.json" class="taxt-hover-reference" href="/references/129831">Wheeler, 1928</a>: 89 (in text); <a data-controller="hover-preview" data-hover-preview-url-value="/references/129843/hover_preview.json" class="taxt-hover-reference" href="/references/129843">Wheeler & Wheeler, 1930</a>: 199; <a data-controller="hover-preview" data-hover-preview-url-value="/references/123583/hover_preview.json" class="taxt-hover-reference" href="/references/123583">Chapman & Capco, 1951</a>: 17; <a data-controller="hover-preview" data-hover-preview-url-value="/references/123635/hover_preview.json" class="taxt-hover-reference" href="/references/123635">Clark, 1951</a>: 16; <a data-controller="hover-preview" data-hover-preview-url-value="/references/123108/hover_preview.json" class="taxt-hover-reference" href="/references/123108">Brown, 1954e</a>: 28; <a data-controller="hover-preview" data-hover-preview-url-value="/references/127792/hover_preview.json" class="taxt-hover-reference" href="/references/127792">Petersen, 1968</a>: 577; <a data-controller="hover-preview" data-hover-preview-url-value="/references/129891/hover_preview.json" class="taxt-hover-reference" href="/references/129891">Wheeler & Wheeler, 1972a</a>: 37; <a data-controller="hover-preview" data-hover-preview-url-value="/references/123216/hover_preview.json" class="taxt-hover-reference" href="/references/123216">Brown, 1973b</a>: 166; <a data-controller="hover-preview" data-hover-preview-url-value="/references/122492/hover_preview.json" class="taxt-hover-reference" href="/references/122492">Baroni Urbani, 1977c</a>: 430; <a data-controller="hover-preview" data-hover-preview-url-value="/references/124165/hover_preview.json" class="taxt-hover-reference" href="/references/124165">Dlussky & Fedoseeva, 1988</a>: 79; <a data-controller="hover-preview" data-hover-preview-url-value="/references/122852/hover_preview.json" class="taxt-hover-reference" href="/references/122852">H枚lldobler & Wilson, 1990</a>: 12; <a data-controller="hover-preview" data-hover-preview-url-value="/references/122860/hover_preview.json" class="taxt-hover-reference" href="/references/122860">Bolton, 1995b</a>: 12; <a data-controller="hover-preview" data-hover-preview-url-value="/references/130789/hover_preview.json" class="taxt-hover-reference" href="/references/130789">Bolton, 2003</a>: 39, 151; <a data-controller="hover-preview" data-hover-preview-url-value="/references/132572/hover_preview.json" class="taxt-hover-reference" href="/references/132572">Ward, 2007c</a>: 555; <a data-controller="hover-preview" data-hover-preview-url-value="/references/142740/hover_preview.json" class="taxt-hover-reference" href="/references/142740">Boudinot, 2015</a>: 29; <a data-controller="hover-preview" data-hover-preview-url-value="/references/144272/hover_preview.json" class="taxt-hover-reference" href="/references/144272">Griebenow, 2024</a>: 111.</li> </ul>     <div class='mb-8'> <div> <span class='font-bold'>Tribes of Leptanillinae:</span> <a data-controller="hover-preview" data-hover-preview-url-value="/catalog/430168/hover_preview.json" class="valid" href="/catalog/430168">Leptanillini</a>, <a data-controller="hover-preview" data-hover-preview-url-value="/catalog/513819/hover_preview.json" class="valid" href="/catalog/513819">Opamyrmini</a> </div> </div>  <div> <div class='mb-4 group/reference-sections-item'> <h6 class='font-bold'>Subfamily <a data-controller="hover-preview" data-hover-preview-url-value="/catalog/430164/hover_preview.json" class="valid" href="/catalog/430164">Leptanillinae</a> catalogues</h6> <p><a data-controller="hover-preview" data-hover-preview-url-value="/references/129218/hover_preview.json" class="taxt-hover-reference" href="/references/129218">Taylor & Brown, 1985</a>: 53 (Australia); <a data-controller="hover-preview" data-hover-preview-url-value="/references/122860/hover_preview.json" class="taxt-hover-reference" href="/references/122860">Bolton, 1995b</a>: 9, 12; <a data-controller="hover-preview" data-hover-preview-url-value="/references/144355/hover_preview.json" class="taxt-hover-reference" href="/references/144355">Hamer <i>et al.</i>, 2024</a>: 109 (China checklist).</p> </div> <div class='mb-4 group/reference-sections-item'> <h6 class='font-bold'>Subfamily <a data-controller="hover-preview" data-hover-preview-url-value="/catalog/430164/hover_preview.json" class="valid" href="/catalog/430164">Leptanillinae</a> references</h6> <p><a data-controller="hover-preview" data-hover-preview-url-value="/references/124659/hover_preview.json" class="taxt-hover-reference" href="/references/124659">Emery, 1904a</a>: 107 (anatomy, affinities); <a data-controller="hover-preview" data-hover-preview-url-value="/references/130044/hover_preview.json" class="taxt-hover-reference" href="/references/130044">Wheeler, 1910a</a>: 138 (diagnosis); <a data-controller="hover-preview" data-hover-preview-url-value="/references/129831/hover_preview.json" class="taxt-hover-reference" href="/references/129831">Wheeler, 1928</a>: 85 (larva); <a data-controller="hover-preview" data-hover-preview-url-value="/references/129843/hover_preview.json" class="taxt-hover-reference" href="/references/129843">Wheeler & Wheeler, 1930</a>: 199 (diagnosis); <a data-controller="hover-preview" data-hover-preview-url-value="/references/127460/hover_preview.json" class="taxt-hover-reference" href="/references/127460">Morley, 1939</a>: 114 (phylogeny); <a data-controller="hover-preview" data-hover-preview-url-value="/references/126688/hover_preview.json" class="taxt-hover-reference" href="/references/126688">Kutter, 1948</a>: 293 (diagnosis); <a data-controller="hover-preview" data-hover-preview-url-value="/references/123108/hover_preview.json" class="taxt-hover-reference" href="/references/123108">Brown, 1954e</a>: 28 (phylogeny); <a data-controller="hover-preview" data-hover-preview-url-value="/references/122660/hover_preview.json" class="taxt-hover-reference" href="/references/122660">Bernard, 1967a</a>: 90 (diagnosis); <a data-controller="hover-preview" data-hover-preview-url-value="/references/127792/hover_preview.json" class="taxt-hover-reference" href="/references/127792">Petersen, 1968</a>: 577 (tribe, males); <a data-controller="hover-preview" data-hover-preview-url-value="/references/125611/hover_preview.json" class="taxt-hover-reference" href="/references/125611">Gotwald, 1969b</a>: 97 (mouthparts morphology); <a data-controller="hover-preview" data-hover-preview-url-value="/references/129891/hover_preview.json" class="taxt-hover-reference" href="/references/129891">Wheeler & Wheeler, 1972a</a>: 37 (diagnosis); <a data-controller="hover-preview" data-hover-preview-url-value="/references/123216/hover_preview.json" class="taxt-hover-reference" href="/references/123216">Brown, 1973b</a>: 166 (genera, distribution); <a data-controller="hover-preview" data-hover-preview-url-value="/references/129906/hover_preview.json" class="taxt-hover-reference" href="/references/129906">Wheeler & Wheeler, 1976b</a>: 46 (larvae, review and synthesis); <a data-controller="hover-preview" data-hover-preview-url-value="/references/122492/hover_preview.json" class="taxt-hover-reference" href="/references/122492">Baroni Urbani, 1977c</a>: 430 (diagnosis, all species revision); <a data-controller="hover-preview" data-hover-preview-url-value="/references/128876/hover_preview.json" class="taxt-hover-reference" href="/references/128876">Snelling, 1981</a>: 392 (synoptic classification); <a data-controller="hover-preview" data-hover-preview-url-value="/references/129918/hover_preview.json" class="taxt-hover-reference" href="/references/129918">Wheeler & Wheeler, 1985b</a>: 257 (synoptic classification); <a data-controller="hover-preview" data-hover-preview-url-value="/references/124165/hover_preview.json" class="taxt-hover-reference" href="/references/124165">Dlussky & Fedoseeva, 1988</a>: 79 (synoptic classification); <a data-controller="hover-preview" data-hover-preview-url-value="/references/122852/hover_preview.json" class="taxt-hover-reference" href="/references/122852">H枚lldobler & Wilson, 1990</a>: 9 onward (synoptic classification, genera keys); <a data-controller="hover-preview" data-hover-preview-url-value="/references/122855/hover_preview.json" class="taxt-hover-reference" href="/references/122855">Bolton, 1990b</a>: 269 (diagnosis, revision of tribes, synoptic classification, key); <a data-controller="hover-preview" data-hover-preview-url-value="/references/126562/hover_preview.json" class="taxt-hover-reference" href="/references/126562">Kugler, 1992</a>: 103 (sting structure); <a data-controller="hover-preview" data-hover-preview-url-value="/references/122513/hover_preview.json" class="taxt-hover-reference" href="/references/122513">Baroni Urbani <i>et al.</i>, 1992</a>: 316, 317 (phylogeny); <a data-controller="hover-preview" data-hover-preview-url-value="/references/122834/hover_preview.json" class="taxt-hover-reference" href="/references/122834">Bolton, 1994</a>: 69 (diagnosis, synoptic classification, genera key); <a data-controller="hover-preview" data-hover-preview-url-value="/references/122859/hover_preview.json" class="taxt-hover-reference" href="/references/122859">Bolton, 1995a</a>: 1040 (census); <a data-controller="hover-preview" data-hover-preview-url-value="/references/127591/hover_preview.json" class="taxt-hover-reference" href="/references/127591">Ogata <i>et al.</i>, 1995</a>: 32 (genera, classification); <a data-controller="hover-preview" data-hover-preview-url-value="/references/132308/hover_preview.json" class="taxt-hover-reference" href="/references/132308">Shattuck, 1999</a>: 117 (Australia, synopsis); <a data-controller="hover-preview" data-hover-preview-url-value="/references/130789/hover_preview.json" class="taxt-hover-reference" href="/references/130789">Bolton, 2003</a>: 39, 151 (diagnosis, synopsis); <a data-controller="hover-preview" data-hover-preview-url-value="/references/131969/hover_preview.json" class="taxt-hover-reference" href="/references/131969">Ouellette <i>et al.</i>, 2006</a>: 359 (phylogeny); <a data-controller="hover-preview" data-hover-preview-url-value="/references/132431/hover_preview.json" class="taxt-hover-reference" href="/references/132431">Terayama, 2009</a>: 125 (Taiwan genera key); <a data-controller="hover-preview" data-hover-preview-url-value="/references/132809/hover_preview.json" class="taxt-hover-reference" href="/references/132809">Borowiec <i>et al.</i>, 2011</a>: 13 (genera key); <a data-controller="hover-preview" data-hover-preview-url-value="/references/132879/hover_preview.json" class="taxt-hover-reference" href="/references/132879">Keller, 2011</a>: 1 (morphology, phylogeny); <a data-controller="hover-preview" data-hover-preview-url-value="/references/133060/hover_preview.json" class="taxt-hover-reference" href="/references/133060">General & Alpert, 2012</a>: 73 (Philippines genera key); <a data-controller="hover-preview" data-hover-preview-url-value="/references/134418/hover_preview.json" class="taxt-hover-reference" href="/references/134418">Xu, 2012c</a>: 479 (genera key); <a data-controller="hover-preview" data-hover-preview-url-value="/references/142647/hover_preview.json" class="taxt-hover-reference" href="/references/142647">Eguchi <i>et al.</i>, 2014</a>: 3 (Vietnam genera key); <a data-controller="hover-preview" data-hover-preview-url-value="/references/142740/hover_preview.json" class="taxt-hover-reference" href="/references/142740">Boudinot, 2015</a>: 29 (males diagnosis); <a data-controller="hover-preview" data-hover-preview-url-value="/references/142963/hover_preview.json" class="taxt-hover-reference" href="/references/142963">Fisher & Bolton, 2016</a>: 49 (diagnosis); <a data-controller="hover-preview" data-hover-preview-url-value="/references/143787/hover_preview.json" class="taxt-hover-reference" href="/references/143787">Griebenow, 2020</a>: 235 (phylogeny, tribal revision), 237 (males diagnosis), 241 (key to known males); <a data-controller="hover-preview" data-hover-preview-url-value="/references/144154/hover_preview.json" class="taxt-hover-reference" href="/references/144154">Griebenow <i>et al.</i>, 2022</a>: 1130 (genera key); <a data-controller="hover-preview" data-hover-preview-url-value="/references/144260/hover_preview.json" class="taxt-hover-reference" href="/references/144260">Griebenow <i>et al.</i>, 2023</a>: 945 (male genital skeletomusculature, evolution); <a data-controller="hover-preview" data-hover-preview-url-value="/references/144272/hover_preview.json" class="taxt-hover-reference" href="/references/144272">Griebenow, 2024</a>: 83 (systematic revision), 111 (worker diagnosis), 112 (gyne diagnosis), 113 (male diagnosis), 114 (larva diagnosis), 148 (worker-based keys), 155 (male-based key to major subclades), 176 (biogeography and ecology); <a data-controller="hover-preview" data-hover-preview-url-value="/references/144355/hover_preview.json" class="taxt-hover-reference" href="/references/144355">Hamer <i>et al.</i>, 2024</a>: 99 (diversity, distribution in China).</p> </div> </div>  <div class='mb-12'></div> <h5 class='font-bold'>Citations</h5> <ul> <li class='group/citations-reference [&_a]:break-words'> <a href="/authors/6918">Baroni Urbani, C.</a> 1977c. <a href="/references/122492">Materiali per una revisione della sottofamiglia Leptanillinae Emery (Hymenoptera: Formicidae).</a> Entomologica Basiliensia 2:427-488. <a class="pdf-link" rel="nofollow" href="https://antcat.org/documents/164/6319.pdf">PDF</a> </li> <li class='group/citations-reference [&_a]:break-words'> <a href="/authors/6918">Baroni Urbani, C.</a>; <a href="/authors/6762">Bolton, B.</a>; <a href="/authors/6927">Ward, P. 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M.</a> 1923g. <a href="/references/130169">Social life among the insects.</a> New York: Harcourt, Brace and Co., vii + 375 pp. </li> <li class='group/citations-reference [&_a]:break-words'> <a href="/authors/9382">Xu, Z.-H.</a> 2012c. <a href="/references/134418"><i>Furcotanilla</i>, a new genus of the ant subfamily Leptanillinae from China with descriptions of two new species of <i>Protanilla</i> and <i>P. rafflesi</i> Taylor (Hymenoptera: Formicidae).</a> Sociobiology 59:477-491. <a class="pdf-link" rel="nofollow" href="https://antcat.org/documents/5724/xu_2012_sociobiology-furcotanilla.pdf">PDF</a> </li> </ul> </div> <div class='medium-3'> <div class='callout-white mb-4'> <p>Extant: 2 valid tribes, 3 valid genera, 82 valid species</p> <a class="btn-default" href="/catalog/430164?include_full_statistics=true">More statistics</a> </div> <div class='callout-white'> <a class="external-link" href="https://www.antweb.org/description.do?rank=subfamily&subfamily=leptanillinae&project=worldants">AntWeb</a> <a class="external-link" href="https://www.antwiki.org/wiki/Leptanillinae">AntWiki</a> <a class="external-link" href="//scholar.google.com/scholar?q=Leptanillinae+Emery%2C+1910">Google Scholar</a> </div> </div> </div> </div> </section> <div class='sticky color-coded-catalog-links nottiny:bottom-0' id='taxon-browser'> <div class='bg-white overflow-hidden h-auto' id='taxon-browser-css-tabs'> <div class='hidden px-4' data-toggler-name='taxon_key'> <div class='flex justify-between gap-1 flex-wrap'> <div>&dagger;fossil</div> <div class='valid'>valid</div> <div class='synonym'>synonym</div> <div class='unidentifiable'>unidentifiable/excluded from Formicidae</div> <div class='homonym'>homonym</div> <div class='unavailable'>unavailable/nomen nudum</div> <div class='obsolete-combination'>obsolete combination</div> </div> </div> <input class='taxon-browser-tab-input hidden' id='taxon-browser-tab-0' name='taxon_browser_tab' type='radio' value='selected tab'> <label class='text-ac-blue-text py-1 px-4 !m-0' for='taxon-browser-tab-0'>Formicidae subfamilies</label> <input class='taxon-browser-tab-input hidden' id='taxon-browser-tab-1' name='taxon_browser_tab' type='radio' value='selected tab'> <label class='text-ac-blue-text py-1 px-4 !m-0' for='taxon-browser-tab-1'>Leptanillinae tribes</label> <input checked='checked' class='taxon-browser-tab-input hidden' id='taxon-browser-tab-2' name='taxon_browser_tab' type='radio' value='selected tab'> <label class='text-ac-blue-text py-1 px-4 !m-0' for='taxon-browser-tab-2'>All Leptanillinae genera</label> <div class='float-right hidden sm:block'> <button class='btn-neutral' data-action='click->toggler#toggle' data-toggler-target='taxon_key'>toggle legend</button> <span class='base__btn [&_a]:text-white bg-gray-500 hover:bg-gray-600 border-gray-500'> <a rel="nofollow" data-method="put" href="/catalog/toggle_view?show=valid_and_invalid">show invalid<i class="antcat_icon refresh"></i></a> </span> </div> <div class='hidden overflow-y-auto clear-right p-4 pt-0 min-h-[10rem] max-h-[24vh]' id='taxon-browser-tab-0-content'> <ul class='no-bullet py-2 columns-[9rem]'> <li class='uppercase whitespace-nowrap'><span class="label-white-smaller"><a href="/catalog/429011?view=all_genera_in_family">All genera</a></span></li> <li class='uppercase whitespace-nowrap'><span class="label-white-smaller"><a href="/catalog/429011?view=incertae_sedis_in_family">Incertae sedis</a></span></li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/430014">Agroecomyrmecinae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/430020">Amblyoponinae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/429060">Aneuretinae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/507398">Apomyrminae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/430175">鈥燗rmaniinae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/430196">鈥燘rownimeciinae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/429071">Dolichoderinae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/429481">Dorylinae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/429497">Ectatomminae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/430199">鈥燜ormiciinae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/429149">Formicinae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/508390">鈥燞aidomyrmecinae</a> </li> <li class='whitespace-nowrap selected'> <a class="valid subfamily" href="/catalog/430164">Leptanillinae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/430173">Martialinae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/429387">Myrmeciinae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/429529">Myrmicinae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/430147">Paraponerinae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/430052">Ponerinae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/430150">Proceratiinae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/429404">Pseudomyrmecinae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/430185">鈥燬phecomyrminae</a> </li> <li class='whitespace-nowrap'> <a class="valid subfamily" href="/catalog/511088">鈥燴igrasimeciinae</a> </li> </ul> </div> <div class='hidden overflow-y-auto clear-right p-4 pt-0 min-h-[10rem] max-h-[24vh]' id='taxon-browser-tab-1-content'> <ul class='no-bullet py-2 columns-[9rem]'> <li class='uppercase whitespace-nowrap'><span class="selected"><a href="/catalog/430164?view=all_genera_in_subfamily">All genera</a></span></li> <li class='uppercase whitespace-nowrap'><span class="label-white-smaller"><a href="/catalog/430164?view=without_tribe">Without tribe</a></span></li> <li class='whitespace-nowrap'> <a class="valid tribe" href="/catalog/430168">Leptanillini</a> </li> <li class='whitespace-nowrap'> <a class="valid tribe" href="/catalog/513819">Opamyrmini</a> </li> </ul> </div> <div class='hidden overflow-y-auto clear-right p-4 pt-0 min-h-[10rem] max-h-[24vh]' id='taxon-browser-tab-2-content'> <ul class='no-bullet py-2 columns-[9rem]'> <li class='whitespace-nowrap'> <a class="valid genus" href="/catalog/430169">Leptanilla</a> </li> <li class='whitespace-nowrap'> <a class="valid genus" href="/catalog/430041">Opamyrma</a> </li> <li class='whitespace-nowrap'> <a class="valid genus" href="/catalog/430167">Protanilla</a> </li> </ul> </div> </div> </div> <footer class='bg-ac-blue text-white [&_a]:text-white [&_a:hover]:font-bold'> <div class='layout-container text-center text-sm pt-12 pb-8'> <div class='w-full flex flex-wrap'> <div class='w-full flex items-center justify-around lg:w-6/12'> <ul class='no-bullet'> <li><img alt="AntCat" class="inline-block mb-4" src="/assets/logos/antcat_head_white_27x89-cb88c6c1f42f938e3b6c9fb7f5742e78e70c88d5dc3664585e277b54dadae324.png" /></li> <li><a href="/about">About</a></li> <li><a href="/users">List of users</a></li> <li><a href="mailto:bfisher@calacademy.org?subject=AntCat">Contact us </a></li> </ul> <ul class='no-bullet'> <li><a href="/activities">Activity feed</a></li> <li class='whitespace-nowrap'> <a href="/contribute">Contribute <i class="antcat_icon star"></i> <i class="antcat_icon star"></i> <i class="antcat_icon star"></i> </a></li> <li><a href="/catalog/random">Random 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