CINXE.COM
Just a moment...
<!DOCTYPE html><html lang="en-US"><head><title>Just a moment...</title><meta http-equiv="Content-Type" content="text/html; charset=UTF-8"><meta http-equiv="X-UA-Compatible" content="IE=Edge"><meta name="robots" content="noindex,nofollow"><meta name="viewport" content="width=device-width,initial-scale=1"><style>*{box-sizing:border-box;margin:0;padding:0}html{line-height:1.15;-webkit-text-size-adjust:100%;color:#313131;font-family:system-ui,-apple-system,BlinkMacSystemFont,Segoe UI,Roboto,Helvetica Neue,Arial,Noto Sans,sans-serif,Apple Color Emoji,Segoe UI Emoji,Segoe UI Symbol,Noto Color Emoji}body{display:flex;flex-direction:column;height:100vh;min-height:100vh}.main-content{margin:8rem auto;max-width:60rem;padding-left:1.5rem}@media (width <= 720px){.main-content{margin-top:4rem}}.h2{font-size:1.5rem;font-weight:500;line-height:2.25rem}@media (width <= 720px){.h2{font-size:1.25rem;line-height:1.5rem}}#challenge-error-text{background-image:url(data:image/svg+xml;base64,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);background-repeat:no-repeat;background-size:contain;padding-left:34px}@media (prefers-color-scheme:dark){body{background-color:#222;color:#d9d9d9}}</style><meta http-equiv="refresh" content="390"></head><body class="no-js"><div class="main-wrapper" role="main"><div class="main-content"><noscript><div class="h2"><span id="challenge-error-text">Enable JavaScript and cookies to continue</span></div></noscript></div></div><script>(function(){window._cf_chl_opt={cvId: '3',cZone: "ift.onlinelibrary.wiley.com",cType: 'managed',cRay: '8ea9ae7f1e5dcde6',cH: 'Qf0Yswqz3Uv7VI_u4JGUqaG.0V4i_o..3eDC_a7O0Vc-1732957588-1.2.1.1-J0w.WhHfGUiYfjevul1apkQICzmTMeOpQ8ShEll2J0ToMKnRB7P20N30_xathrUg',cUPMDTk: "\/doi\/10.1111\/j.1365-2621.1963.tb00247.x?__cf_chl_tk=CGgb6ZPwPHVjMaJiuUxtmAsJTB8vNxh9opQb7mYIvIQ-1732957588-1.0.1.1-EGJ7vu6gcfQJkf4k4iGjEuh4.yp5rfQziNEfsFOnASI",cFPWv: 'b',cITimeS: '1732957588',cTTimeMs: '1000',cMTimeMs: '390000',cTplC: 0,cTplV: 5,cTplB: 'cf',cK: "",fa: "\/doi\/10.1111\/j.1365-2621.1963.tb00247.x?__cf_chl_f_tk=CGgb6ZPwPHVjMaJiuUxtmAsJTB8vNxh9opQb7mYIvIQ-1732957588-1.0.1.1-EGJ7vu6gcfQJkf4k4iGjEuh4.yp5rfQziNEfsFOnASI",md: "RbtkVc9pWky98KSANNMZwQodIOw2yenHCWr_KrJFNaE-1732957588-1.2.1.1-KVGOTalopkeeDiSeq1I_2YbA2DeGc_oZp3qDAi9hNYBJt43TV.BUInFCgyzULkmCVCE_XIsjM0MEVwRxyMBDF43r8pBs__mw9yHb.GidT4P33LeImKhyi7SD2d7OKg5GDnjuwjNZlCfA_TsG88rwXfRrDdnN3tlKOosm1940wFCPqLRPoy0Gl6Iklp.7pNcPLBRmbyHn890siI2pNuzTi45kBGP2ZKoQBFwiXX48FklivFBR8sHuK.FDLu9QR3lHPYJcfRd7zqrpRmNko8KYrHw3po6xvPXFUMSs7rlynf4YfvDol6S66S8v_q68I8JXKS003X7fnN4GlwW7s71Q_OCpPdxjJeST6kZbqTV7Mp8sWXk0cU.uw2Kgklid9frfAZVJmEk0yrIfw7JdSUFCMmZulwIMiIQpfodSkYN6w4AzR9bjN9oihJw9Pth9htA_oYMof_RoT.gnCS82nsrH4KkS1BRKnwMSMJieGGnUlG3EgULtplfPQbhl9uIRn9.4jX7gVvzOw8F71WLdUL3a7WkJYiXXVnS.2Ab4uQ_c6N6uaB0HvLiHRIOxdBkO5np5o2lSWEWYnxb1sPJCI3SYwBkF2Qj.kfzvrlxMoKum2vN75PFLNG5qN.RbSPDfXE8TYe0ZsEKbBVAzJ8Km7MB1HVdRT8LqNWsiSwPq8VUM5USjm4MdYHWEjTdzOdNe_LMdcDi0COArB.qPyTO4UzbFbEW9gXaJXR6FowNBibcAIaho55Y_e872qNr12jqLPXi9Xti0BL7DucCgOLYXxFU7eIYjFp2.KxxpVLhdCt_W7twVrDUh8RVIHEKC4GoEkv6psdB52f2vi8IbFgK4vpvxinifm.KmrBINx_bVVGwL4YEyYOcMCgKyje7igdLbZpK2LWoBh8ZZgH8zWOPH9jxjKgaeaUE4_vMzUpuCwq3FXywxESzHRvi5Fu2IAswquRauGX5IcluYr1PF_p0gH9.a06Rwwz5Wy9CX9gcB6Kooa0RLOr.gaGQeXzbOshobCH7T.Xuu6NWWctpXFxBJUeBpGgF5yxToaGKT5zk245OT_dYf8.ARW8fA04wWBy_o20mqVk2HdBH9jz2BUtboqfkxvDBRtMDcEyv3BADbBK35MIPXFyRd9TnYWiKeptInaQxFpbN6sjUzKUKh8EI_MgGQVtWPiLeFxW7WMof7flicV99oc0FKKbKGEmEEcInJBuIgRsrAZ8_C4B42yyeqlXGAlMthxigm74fREGbe4D5F_tsz2e.4HDU3q1jDRtgE7xtDb9UT3sT0jAxyZS1CkNdwdApyiBAs9ogSi.DvYILhKl0Nwv.I49udosrBOAnTZZai4yYwyN0xzGQRIxq_WzgSJo8ynqnnocrA4UMgiud9lMxIeOBtpnUCa77JNqbNDn71vsNzp5HD9Q1nSqD8K4BiZEc1xDHEz6_bNWNc5J00Ctb_vLkyruNT543YcW_3H24FzzBZ90xrBzvr5_901VtpKbzpNI0ZPI1v5uYbtdgbs__tcl5iYS2r2i7JnUvA46roVzPjMqVpmQnCM5MpXKHj52.glIb0eeT5DREeWVoxwEO.xP_tMNVehzi7WI3QOEji_ylu5bD8BUPzRYdl1CabD_9wQDHr3M9jz6H6nBg2M6ADVTPIbToo8Nff.cSpFgcKMD4DnTc.bisTaNps6LIPauJoUy0klYwrkMDbkCWsiRHtI7zc4qV8oh0dy6FzCJKSIPWSEZYcC2iTJDmgqmHmt_97eJK6KoXdpgE_J0xDmngAN8fxFUfBUjnlo6Nt4T9lxkSCaMGataTm0N_0zLkGBROgG0las6xWD9xGfrHguJnSlO9HmI_2U9uXjkAV2479YgvIL0InRMV0eG3_DcFoLXPW9vfxiFdM_BTVHq0Kw0nh8tHNo6d_M8S70iJX7daIq7Uc6SmEaKo_9H.F1wCHHCigHWdXMG8NsITELsUHWSVMBxYXTDFjqCbD6SHn_wVcs_Tk8dWy2c1K1y7hSu00bESX94Bmw51HLGLsj96Tk_NNSqGc71qf.i63VDMI.3t7ipI8DU_hSoS7sIbSf4Civ48_SsJWqWE_b0ufEG6lH24.2lF_LOodyPsIKKeMAbfzZ3rQD4yj7yqloLRTsMViWo75mOxWboy.dV.uqVpSkR3csNaLKc4sOzqOemHA28S.nnyE_Ll19Muf24P6b9GYwS74KOgRsZZ0_FU4XDxcbVUqtHxV5QhwIqdRRrxgoPyYOUlGMKwLKg9yHDniyV5xV_BAd5R8pyNEooUP._TOFpvtzeKTN2_qi1CXrYqU0zgvBPdy8w4vmoniPKUGIE6s4k6hyaSaDChdXFXsa3wUVWgGmnKT0BQhTA9CPAPMqWnndi5t31dgDKqkL6fcnVauclPx6cC1vmaZxHzav5tHn6OW0ZKTGmd6XI7cG49MvluOWExT.5EIFapek7WB28c7Q9Jl2nTG1ggE920Kd.wCzsCZacEMSvs2QtJdxneYnSbnkkmUyJVs6zSqU_aQZUA.uj3JTww7fSLTDXJRrmzQaCpvehpgTk.j_G5JS1ds73Jmg5mbgeSRV.xWL8nRP4KQ80ViyIflEoiOprdFnteq8vz0YBCQDVRuh7.jkqFDhV7NJNvWYrOtY4AtRhLKMy_zwI9tftgMrgYZpGqqOP3DEgyJyarxGRpO2IA2lRHuf0AQLJaz5E4Mz8bBGt37wBUl5EFL819b3VRDWuysMEdqcavj_CXbqy2Kelec1.SHVz88FDhkk4xiTNxuyuZexB_wDyG5veBhRdRiRntaVGHM1R5IfNzbrClOQ6KcHJ0qgrT98iGuwLwGHqNirVh1Pc5U3LYkaU7tJ2RRDK8nFWUPNqFaCFQIRdwEfelmmQIPeQKtpHtx7poUBT4lxHSI5Cq0zMHdA19IswsuOROeyFzTLGF1sMwzh3PBpAy0jGMqQaFtF_4rt1OVk83JpILnyvTNzlyhWPmUd4EctnSOaXcZSrh0mqEsWTngPqrOIHv3XQ7aAqvqNhvxxRXp6dUDPgNK6A",mdrd: "uGSzsPVCFndcIoR5xiU.hXOcIdWsEBCzLqnMhj3Fzao-1732957588-1.2.1.1-B3Ly5byGt3RFS0bTQetjm5SHBQRzHWDl8F8btYU5HnuoTTR3.DzAqr68WMDSpvj3Bm8_GmpFr6xoZODYee_O6PVNuZKIo.oXcnPy27DNUXvCp6yVA5HBMpGkOZHM6kaSgP0kk2utqkMNtDAcjseRRuzopFLzfJTdZ.PJPEuhETVMK9Jtsjsu8x1FISKJ7YkXzIWHxPnZHlgM3e9pSy6FtTPvqBtpnPT_djpSmpeKEjcdWl9OrZVXiM83VWlXwFY5pyL.FncyjMlbUnYnEpJija_nwqcXsK7yBLZbpZWJP3wh17M.uYe6Mokg9byDvXm.ZInZHIglwamZLmGWYUSjcdl21thR2D24Z08FkreL_0HPw0KTokwf.98LKD9zcK8kg_5xIcCnTrMKOE9qMQRbZcuN0PaiyzUaEZX19kKPwzcSHhd1dzwUV8vrFXF0ZtIaawn27ILwBIDEfahe2vz998ZuTESuoaglluU4CGIpU.w3_vYP7QpvXbW.w1ml9oqszzYAbBMGTuu_cur3y58i8an2rQiNZxasDZ5DqHSzcaVVTs9GWq6bE6SlfFuJVn3e4q63njAwx4PR8VKcfax9WpkwYjO8A_iFLcOlWekd3ALw8Mqgu8CR_73q9GNvasdZbSOAm7LKUG_k.aVKp9T7lXjjb52PkSStgw04VXtjW0SwRfu17Pz_X7AeW.GRnsyxQmbTS79DijhdOb5v9OdHdJGaA8Y1HZneCtIhu10zpVNnH44Rb4GiDwhYFzeAdiiCL3DCxxmsqMOordBsqcZ3qR0qepAYmcMNHB.Mrvp.My.4UE0.cMTdDSlPLl7pKFAAn_JGCudnb5kxdxctj8WHv9Gc4_.DYhhCMyy4QR9ZzlApVTtWf615uVlwrPmWOuAGpbggq6szcB2Cmxx7TT2raOd0zdNKQTe.U45Lq.RqZn5ihaTuOKFG363UBfWTK90c35605KzFUGfuhVeqiM5IYY5LhNzXi8.LTykGJynzrGRMS3__VcydboFT_U5Q9ysgniopQ4bYyXZwQtRhAXW4kEwLl90ATPxjxgB8S4lU3IIpjAN1FlIjSqT3E0lqF_55ocqvfg6jTsfsXjMdV0a2CFcJMLlDvWeV3LW3rhXC8TkR6xr6h6imiC7G_3W87nzJ5ob5F8BJYQj4D.qK8bA897k7YtQSGOnH7caNgYAFI4NrvK8LSUfgTQO9OPNZCR6NKgeWO4EjdNys.7T7MMS9ngJuNr0IggoqY4S.tR4qhLqMLEc0FoaFXKMd.SYN6DGS3aj4k8ROwOcgpCbFYiPsWpWooDrpZwxihAlPSgbr1oLDD2BhP8ACYe4YnEIv0QTCIS67sNw8tcN7xHIfYMKW0BAoSaD1C9nQZIbCP5U1SeHZNjkFx6VoqRZNMkfakI_nHlOunjTZeKaIoKnXBym3zc9snqNfTki6IZdw8t5x4W2X9hYYpvKUNe5RtlT4DAbHFkuitQwmKZwI653ZFPNamTMSfhPVR_o_zPjJ7Os7uOw8Slj2xQ9lYbe3VQKkQdxlR0JsO5q2BDHolRrxTFHJl9b7zkkOjK_.n5CNH6ZAC3Je2OAZz5I3rjR2YjMezKeVtyzPVpSIcXxj0pJThIACzbFQWlT6a4ykojZUx.0xG2gPFpgoYsmoRlCJ8LenEVuzE_y6HTrZ2kr9gLsefLBQhuLMm_xDwNdDEnrQzVK2Zdi7f16oEdCI1gKalJ5wnjDIjhYppSWr0aP74MRMD9k_RrAsPpqxwExpZFXibgMdl6tLkTpbtylKx9wu6JhZFfcqc6GPWb4pKR89hMIqm18v0US2ypXPkfIM0o2srl.R_IM4kjAsFZTJZ7fEL.5lceh_8KWEi3C1Xqx9Jeu_6DuCPGh4UXhyXUhaOCtWLno.D5i9Pm87y5Sbi8QxkfQ6hfMFbrT91iL26BkNqy63wl_KuKRgBJ9mD2OsPULDST7KfFJ5CaKIC7GEYYU0P66eH6I8D3XcWYwuC8RZDHltiNgdwg"};var cpo = document.createElement('script');cpo.src = '/cdn-cgi/challenge-platform/h/b/orchestrate/chl_page/v1?ray=8ea9ae7f1e5dcde6';window._cf_chl_opt.cOgUHash = location.hash === '' && location.href.indexOf('#') !== -1 ? '#' : location.hash;window._cf_chl_opt.cOgUQuery = location.search === '' && location.href.slice(0, location.href.length - window._cf_chl_opt.cOgUHash.length).indexOf('?') !== -1 ? '?' : location.search;if (window.history && window.history.replaceState) {var ogU = location.pathname + window._cf_chl_opt.cOgUQuery + window._cf_chl_opt.cOgUHash;history.replaceState(null, null, "\/doi\/10.1111\/j.1365-2621.1963.tb00247.x?__cf_chl_rt_tk=CGgb6ZPwPHVjMaJiuUxtmAsJTB8vNxh9opQb7mYIvIQ-1732957588-1.0.1.1-EGJ7vu6gcfQJkf4k4iGjEuh4.yp5rfQziNEfsFOnASI" + window._cf_chl_opt.cOgUHash);cpo.onload = function() {history.replaceState(null, null, ogU);}}document.getElementsByTagName('head')[0].appendChild(cpo);}());</script></body></html>