CINXE.COM
STAMINODE - Encyclopædia Universalis
<!DOCTYPE html> <html lang="fr-FR" xml:lang="fr" dir="ltr"> <head> <meta charset="utf-8"> <!-- This website is powered by TYPO3 - inspiring people to share! TYPO3 is a free open source Content Management Framework initially created by Kasper Skaarhoj and licensed under GNU/GPL. TYPO3 is copyright 1998-2025 of Kasper Skaarhoj. Extensions are copyright of their respective owners. Information and contribution at https://typo3.org/ --> <title>STAMINODE - Encyclopædia Universalis</title> <meta http-equiv="x-ua-compatible" content="IE=edge,chrome=1" /> <meta name="generator" content="TYPO3 CMS" /> <meta name="description" content="Nom donné, en morphologie florale, à une étamine transformée en une pièce stérile, c'est-à-dire dépourvue de sac pollinique, et qui prend souvent..." /> <meta name="robots" content="index, all, follow" /> <meta name="viewport" content="viewport-fit=cover, width=device-width, initial-scale=1.0, minimum-scale=1.0" /> <meta name="author" content="Encyclopædia Universalis" /> <meta property="og:title" content="STAMINODE" /> <meta property="og:description" content="Nom donné, en morphologie florale, à une étamine transformée en une pièce stérile, c'est-à-dire dépourvue de sac pollinique, et qui prend souvent l'apparence d'une languette. On en trouve dans la fleur de la consoude, du tilleul argenté, du cacaoyer par exemple. La fleur des cannas ne..." /> <meta property="og:url" content="https://www.universalis.fr/encyclopedie/staminode/" /> <meta property="og:locale" content="fr_FR" /> <meta property="og:site_name" content="Encyclopædia Universalis" /> <meta property="og:type" content="website" /> <meta property="og:image" content="https://www.universalis.fr/_web/static/icons/favicon.png" /> <meta property="og:image:width" content="256" /> <meta property="og:image:height" content="252" /> <meta name="twitter:title" content="STAMINODE" /> <meta name="twitter:description" content="Nom donné, en morphologie florale, à une étamine transformée en une pièce stérile, c'est-à-dire dépourvue de sac pollinique, et qui prend souvent l'apparence d'une languette. On en trouve dans la fleur de la consoude, du tilleul argenté, du cacaoyer par exemple. La fleur des cannas ne..." /> <meta name="twitter:card" content="summary" /> <meta name="twitter:site" content="@universalisfr" /> <meta name="location" content="France" /> <meta name="rating" content="General" /> <meta name="distribution" content="Global" /> <meta name="resource-type" content="document" /> <meta name="copyright" content="© Copyright 2024 - Universalis" /> <meta name="apple-mobile-web-app-title" content="Universalis" /> <meta name="apple-mobile-web-app-capable" content="yes" /> <meta name="apple-mobile-web-app-status-bar-style" content="default" /> <meta name="format-detection" content="telephone=no,date=no,address=no,email=no,url=no" /> <link rel="stylesheet" href="/_shared/pub/pub.css?1740392913" media="all"> <link rel="stylesheet" href="/_web/assets/css/home.css?1741181350" media="all"> <link rel="stylesheet" href="/_web/assets/css/classification.css?1741181350" media="all"> <link rel="canonical" href="https://www.universalis.fr/encyclopedie/staminode/" /> <script type="application/ld+json">{"@context":"https:\/\/schema.org","@type":"Article","name":"STAMINODE","headline":"STAMINODE","description":"Nom donn\u00e9, en morphologie florale, \u00e0 une \u00e9tamine transform\u00e9e en une pi\u00e8ce st\u00e9rile, c'est-\u00e0-dire d\u00e9pourvue de sac pollinique, et qui prend souvent l'apparence d'une languette. On en trouve dans la fleur de la consoude, du tilleul argent\u00e9, du cacaoyer par exemple. La fleur des cannas ne poss\u00e8de qu'une demi-\u00e9tamine produisant du pollen\u00a0; l'autre moiti\u00e9 de cette \u00e9tamine, ainsi que les cinq autres, sont transform\u00e9es en staminodes, pi\u00e8ces rouges st\u00e9riles que l'on prendrait \u00e0 premi\u00e8re vue pour les p\u00e9tales\u00a0; les p\u00e9tales proprement dits sont tr\u00e8s r\u00e9duits, les s\u00e9pales aussi\u00a0; l'un des staminodes est m\u00eame transform\u00e9 en \u00ab\u00a0labelle\u00a0\u00bb, accentuant la forte zygomorphie de cette fleur, tr\u00e8s adapt\u00e9e \u00e0 la f\u00e9condation par...","url":"https:\/\/www.universalis.fr\/encyclopedie\/staminode\/","wordCount":"133","image":{"@type":"ImageObject","url":"https:\/\/www.universalis.fr\/_web\/static\/icons\/favicon.png"},"dateModified":"2025-01-29T00:00:00+00:00","author":[{"@context":"https:\/\/schema.org","@type":"Person","name":"Jacques DAUTA","familyName":"DAUTA","givenName":"Jacques","description":"Assistant \u00e0 l'universit\u00e9 de Paris-VI.","url":"https:\/\/www.universalis.fr\/auteurs\/jacques-dauta\/"}],"publisher":{"@type":"Organization","name":"Encyclop\u00e6dia Universalis","alternateName":"Universalis","logo":{"@type":"ImageObject","url":"https:\/\/www.universalis.fr\/_web\/assets\/img\/banniere-fr.png"},"sameAs":["https:\/\/www.facebook.com\/EncyclopedieUniversalis\/","https:\/\/www.linkedin.com\/company\/encyclopaedia-universalis","https:\/\/www.twitter.com\/universalisfr","https:\/\/www.youtube.com\/channel\/UC0srTZiJaRaDw3fAvtCo4Ow"],"slogan":"Passeur de savoir"}}</script> <script type="application/ld+json">[{"@context":"https:\/\/schema.org","@type":"BreadcrumbList","itemListElement":[{"@type":"ListItem","position":1,"name":"Classification","item":"https:\/\/www.universalis.fr\/classification\/"},{"@type":"ListItem","position":2,"name":"Sciences de la vie","item":"https:\/\/www.universalis.fr\/classification\/sciences-de-la-vie\/"},{"@type":"ListItem","position":3,"name":"Botanique","item":"https:\/\/www.universalis.fr\/classification\/sciences-de-la-vie\/botanique\/"},{"@type":"ListItem","position":4,"name":"Biologie v\u00e9g\u00e9tale","item":"https:\/\/www.universalis.fr\/classification\/sciences-de-la-vie\/botanique\/biologie-vegetale\/"},{"@type":"ListItem","position":5,"name":"Anatomie v\u00e9g\u00e9tale","item":"https:\/\/www.universalis.fr\/classification\/sciences-de-la-vie\/botanique\/biologie-vegetale\/anatomie-vegetale\/"}],"numberOfItems":5,"itemListOrder":"Ascending","name":"Classification : Anatomie v\u00e9g\u00e9tale"}]</script> <meta name="apple-mobile-web-app-capable" content="yes"/> <meta name="mobile-web-app-capable" content="yes"/> <meta name="apple-mobile-web-app-status-bar-style" content="black"> <meta name="apple-mobile-web-app-title" content="Universalis"> <meta name="theme-color" content="#526D78"> <link rel="preconnect" href="https://statistiques.universalis.fr" /> <link rel="preconnect" href="https://cmp.inmobi.com" /> <link rel="search" type="application/opensearchdescription+xml" title="Universalis FR" href="/_web/static/search-fr.xml" /> <link rel="manifest" href="/_web/static/manifest.json"> <link rel="icon" type="image/png" href="/_web/static/icons/favicon.png"> <link rel="apple-touch-icon" href="/_web/static/icons/fr-apple-touch-icon.png" /> <link rel="apple-touch-icon" sizes="57x57" href="/_web/static/icons/fr-apple-touch-icon-57x57.png" /> <link rel="apple-touch-icon" sizes="72x72" href="/_web/static/icons/fr-apple-touch-icon-72x72.png" /> <link rel="apple-touch-icon" sizes="76x76" href="/_web/static/icons/fr-apple-touch-icon-76x76.png" /> <link rel="apple-touch-icon" sizes="114x114" href="/_web/static/icons/fr-apple-touch-icon-114x114.png" /> <link rel="apple-touch-icon" sizes="120x120" href="/_web/static/icons/fr-apple-touch-icon-120x120.png" /> <link rel="apple-touch-icon" sizes="144x144" href="/_web/static/icons/fr-apple-touch-icon-144x144.png" /> <link rel="apple-touch-icon" sizes="152x152" href="/_web/static/icons/fr-apple-touch-icon-152x152.png" /> <link rel="apple-touch-icon" sizes="180x180" href="/_web/static/icons/fr-apple-touch-icon-180x180.png" /> <link rel="apple-touch-startup-image" media="screen and (device-width: 834px) and (device-height: 1194px) and (-webkit-device-pixel-ratio: 2) and (orientation: portrait)" href="/_web/static/img/splash/11__iPad_Pro__10.5__iPad_Pro_portrait.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 810px) and (device-height: 1080px) and (-webkit-device-pixel-ratio: 2) and (orientation: landscape)" href="/_web/static/img/splash/10.2__iPad_landscape.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 744px) and (device-height: 1133px) and (-webkit-device-pixel-ratio: 2) and (orientation: landscape)" href="/_web/static/img/splash/8.3__iPad_Mini_landscape.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 810px) and (device-height: 1080px) and (-webkit-device-pixel-ratio: 2) and (orientation: portrait)" href="/_web/static/img/splash/10.2__iPad_portrait.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 820px) and (device-height: 1180px) and (-webkit-device-pixel-ratio: 2) and (orientation: landscape)" href="/_web/static/img/splash/10.9__iPad_Air_landscape.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 744px) and (device-height: 1133px) and (-webkit-device-pixel-ratio: 2) and (orientation: portrait)" href="/_web/static/img/splash/8.3__iPad_Mini_portrait.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 1024px) and (device-height: 1366px) and (-webkit-device-pixel-ratio: 2) and (orientation: landscape)" href="/_web/static/img/splash/12.9__iPad_Pro_landscape.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 428px) and (device-height: 926px) and (-webkit-device-pixel-ratio: 3) and (orientation: landscape)" href="/_web/static/img/splash/iPhone_14_Plus__iPhone_13_Pro_Max__iPhone_12_Pro_Max_landscape.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 414px) and (device-height: 896px) and (-webkit-device-pixel-ratio: 3) and (orientation: landscape)" href="/_web/static/img/splash/iPhone_11_Pro_Max__iPhone_XS_Max_landscape.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 1024px) and (device-height: 1366px) and (-webkit-device-pixel-ratio: 2) and (orientation: portrait)" href="/_web/static/img/splash/12.9__iPad_Pro_portrait.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 768px) and (device-height: 1024px) and (-webkit-device-pixel-ratio: 2) and (orientation: portrait)" href="/_web/static/img/splash/9.7__iPad_Pro__7.9__iPad_mini__9.7__iPad_Air__9.7__iPad_portrait.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 375px) and (device-height: 812px) and (-webkit-device-pixel-ratio: 3) and (orientation: landscape)" href="/_web/static/img/splash/iPhone_13_mini__iPhone_12_mini__iPhone_11_Pro__iPhone_XS__iPhone_X_landscape.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 820px) and (device-height: 1180px) and (-webkit-device-pixel-ratio: 2) and (orientation: portrait)" href="/_web/static/img/splash/10.9__iPad_Air_portrait.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 393px) and (device-height: 852px) and (-webkit-device-pixel-ratio: 3) and (orientation: portrait)" href="/_web/static/img/splash/iPhone_14_Pro_portrait.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 414px) and (device-height: 896px) and (-webkit-device-pixel-ratio: 3) and (orientation: portrait)" href="/_web/static/img/splash/iPhone_11_Pro_Max__iPhone_XS_Max_portrait.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 414px) and (device-height: 736px) and (-webkit-device-pixel-ratio: 3) and (orientation: portrait)" href="/_web/static/img/splash/iPhone_8_Plus__iPhone_7_Plus__iPhone_6s_Plus__iPhone_6_Plus_portrait.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 320px) and (device-height: 568px) and (-webkit-device-pixel-ratio: 2) and (orientation: portrait)" href="/_web/static/img/splash/4__iPhone_SE__iPod_touch_5th_generation_and_later_portrait.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 393px) and (device-height: 852px) and (-webkit-device-pixel-ratio: 3) and (orientation: landscape)" href="/_web/static/img/splash/iPhone_14_Pro_landscape.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 414px) and (device-height: 896px) and (-webkit-device-pixel-ratio: 2) and (orientation: landscape)" href="/_web/static/img/splash/iPhone_11__iPhone_XR_landscape.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 375px) and (device-height: 667px) and (-webkit-device-pixel-ratio: 2) and (orientation: landscape)" href="/_web/static/img/splash/iPhone_8__iPhone_7__iPhone_6s__iPhone_6__4.7__iPhone_SE_landscape.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 320px) and (device-height: 568px) and (-webkit-device-pixel-ratio: 2) and (orientation: landscape)" href="/_web/static/img/splash/4__iPhone_SE__iPod_touch_5th_generation_and_later_landscape.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 390px) and (device-height: 844px) and (-webkit-device-pixel-ratio: 3) and (orientation: landscape)" href="/_web/static/img/splash/iPhone_14__iPhone_13_Pro__iPhone_13__iPhone_12_Pro__iPhone_12_landscape.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 375px) and (device-height: 812px) and (-webkit-device-pixel-ratio: 3) and (orientation: portrait)" href="/_web/static/img/splash/iPhone_13_mini__iPhone_12_mini__iPhone_11_Pro__iPhone_XS__iPhone_X_portrait.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 834px) and (device-height: 1112px) and (-webkit-device-pixel-ratio: 2) and (orientation: portrait)" href="/_web/static/img/splash/10.5__iPad_Air_portrait.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 390px) and (device-height: 844px) and (-webkit-device-pixel-ratio: 3) and (orientation: portrait)" href="/_web/static/img/splash/iPhone_14__iPhone_13_Pro__iPhone_13__iPhone_12_Pro__iPhone_12_portrait.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 768px) and (device-height: 1024px) and (-webkit-device-pixel-ratio: 2) and (orientation: landscape)" href="/_web/static/img/splash/9.7__iPad_Pro__7.9__iPad_mini__9.7__iPad_Air__9.7__iPad_landscape.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 430px) and (device-height: 932px) and (-webkit-device-pixel-ratio: 3) and (orientation: portrait)" href="/_web/static/img/splash/iPhone_14_Pro_Max_portrait.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 428px) and (device-height: 926px) and (-webkit-device-pixel-ratio: 3) and (orientation: portrait)" href="/_web/static/img/splash/iPhone_14_Plus__iPhone_13_Pro_Max__iPhone_12_Pro_Max_portrait.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 414px) and (device-height: 896px) and (-webkit-device-pixel-ratio: 2) and (orientation: portrait)" href="/_web/static/img/splash/iPhone_11__iPhone_XR_portrait.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 375px) and (device-height: 667px) and (-webkit-device-pixel-ratio: 2) and (orientation: portrait)" href="/_web/static/img/splash/iPhone_8__iPhone_7__iPhone_6s__iPhone_6__4.7__iPhone_SE_portrait.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 834px) and (device-height: 1112px) and (-webkit-device-pixel-ratio: 2) and (orientation: landscape)" href="/_web/static/img/splash/10.5__iPad_Air_landscape.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 834px) and (device-height: 1194px) and (-webkit-device-pixel-ratio: 2) and (orientation: landscape)" href="/_web/static/img/splash/11__iPad_Pro__10.5__iPad_Pro_landscape.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 414px) and (device-height: 736px) and (-webkit-device-pixel-ratio: 3) and (orientation: landscape)" href="/_web/static/img/splash/iPhone_8_Plus__iPhone_7_Plus__iPhone_6s_Plus__iPhone_6_Plus_landscape.png"> <link rel="apple-touch-startup-image" media="screen and (device-width: 430px) and (device-height: 932px) and (-webkit-device-pixel-ratio: 3) and (orientation: landscape)" href="/_web/static/img/splash/iPhone_14_Pro_Max_landscape.png"> <script>!function(a,b,c){if(c in b&&b[c]){var d,e=a.location,f=/^(a|html)$/i;a.addEventListener("click",function(a){for(d=a.target;!f.test(d.nodeName);)d=d.parentNode;"href"in d&&(d.href.indexOf("http")||~d.href.indexOf(e.host))&&(a.preventDefault(),e.href=d.href)},!1),a.querySelector("html").classList.add("standalone")}}(document,window.navigator,"standalone")</script> <style> body{ padding-bottom: 2em; } .cta{ display: block; } .folio-block { width: 100%; max-width: 550px; margin: 1em auto; border: .25em solid var(--eu-color-rgb-cta); border-radius: .375em; background: #f5f5f5; padding: .5em 1em; } .folio-block-title { font-size: 1.25em; } @media(max-width: 767px) { .folio-block-title { font-size: 1.1em; } } .folio-block ul { width: 100%; max-width: 300px; margin: 1em auto; } .folio-block ul li { font-size: .95em; } .btn-cta { background: var(--eu-color-rgb-cta) !important; color: #fff !important; font-weight: bold !important; font-size: 1.15em !important; } .btn-cta:hover { background: var(--eu-color-hover-rgb-cta) !important; color: #fff !important; text-decoration: underline; } #bandeau-footer{ display: block !important; position: fixed; bottom: 0; width: 100%; background: rgb(108,51,131); padding: .25em; z-index: 900; border-top: 1px solid rgba(166, 143, 22, 0.72); } #bandeau-footer p{ font-size: 1.375em; margin: 0; background: url('/_web/static/icons/fr-apple-touch-icon-57x57.png') no-repeat left center; background-size: 28px 28px; padding-left: 1.75em; } @media(max-width: 768px){ #bandeau-footer p{ font-size: 1.2em; margin-left: .5em; } body:not(#page_12) .login-link-container{ display: block !important; text-align: center; margin-top: -1em; } } #bandeau-footer a{ color: #fff; } .pub-container{ margin: 1em auto; display: block !important; } .pub-container .pub-placeholder{ display: flex; align-content: center; justify-content: center; align-items: center; background: #eee; color: #aaa; width: 100%; min-height: 100px; } .pub-container .pub-placeholder:before{ content: "contenu sponsorisé"; text-shadow: 0 0 1px #fff; } #ad_rightslot .pub-placeholder{ min-height: 600px; } #ad_rightslot2 .pub-placeholder{ min-height: 250px; } @media (max-width: 768px){ .pub-container{ display: none !important } } </style> <script> const getCookieValue = (name) => ( document.cookie.match('(^|;)\\s*' + name + '\\s*=\\s*([^;]+)')?.pop() || '' ); var userprofile = 'anonymous'; if (getCookieValue('user_data')) { userprofile = 'subscriber'; } var _paq = window._paq = window._paq || []; _paq.push(['setCustomDimension', 1,userprofile]); /* tracker methods like "setCustomDimension" should be called before "trackPageView" */ _paq.push(['trackPageView']); _paq.push(['enableLinkTracking']); (function() { var u="//statistiques.universalis.fr/"; _paq.push(['setTrackerUrl', u+'matomo.php']); _paq.push(['setSiteId', '1']); var d=document, g=d.createElement('script'), s=d.getElementsByTagName('script')[0]; g.async=true; g.src=u+'matomo.js'; s.parentNode.insertBefore(g,s); })(); </script> <style>.swiper-container[data-loaded=false] .swiper-wrapper .swiper-slide{display:none}.swiper-container[data-loaded=false] .swiper-wrapper .swiper-slide:first-child{display:block}img{max-width:100%;height:auto;max-height:250px}img.d-formule{max-height:none}strong{font-weight:bold !important}.dark-mode body{background:#29363a !important}.loader{display:none;height:100vh}.loader:after{display:block;margin:auto;content:"";background:url(/_web/assets/img/banniere-fr.png) no-repeat center;background-size:contain;width:50%;height:50%;position:absolute;top:50%;left:50%;transform:translate(-50%, -50%);opacity:.25}#skip{padding:5px 15px;position:absolute;background:rgba(0,0,0,.85);color:#fff;left:10%;transform:translateY(-100%);transition:transform .3s;border-bottom-left-radius:6px;border-bottom-right-radius:6px;z-index:100;text-decoration:none}#skip:focus{transform:translateY(0%);box-shadow:0 0 10px #333}.login-form-wrapper-mobile{display:none}.site-fr .article-quote{display:none !important}@media(max-width: 768px){.login-form-wrapper-mobile{display:block;text-align:center;margin-top:-1em;margin-bottom:1em}}body.loading{overflow:hidden}body.loading>*{display:none}body.loading .loader{display:block}.dropdown-menu,.d-none,#podcast audio{display:none}.list-inline-item{display:inline-block}.offcanvas{visibility:hidden}.main-navbar-wrapper .btn-universalis{background-color:#e8eced;color:#000;font-weight:normal}.offcanvas{display:none !important;height:0 !important}.offcanvas[data-loaded=true]{display:inherit !important;height:inherit !important}.clearfix{clear:both}.fr .is-edu{display:none !important}#podcast{min-height:90px}#article-medias-strip-junior{min-height:255px}#callSommaire{position:fixed;bottom:10px;left:10px;z-index:2}.edu #callSommaire{bottom:60px}#qc-cmp2-ui{padding:10px !important}.qc-cmp2-summary-buttons{justify-content:space-around !important}.qc-cmp2-summary-buttons button[mode=secondary],.qc-cmp2-footer button[mode=secondary]{border:none;box-shadow:none}.eCBsvP.eCBsvP{height:auto !important}.qc-cmp-cleanslate{background-color:rgba(33,41,52,.95) !important}.fill-universalis{fill:#1c3479}.fill-universalis-up{fill:#526c78}@media(min-width: 768px){.qc-cmp2-publisher-logo-container{text-align:center}.qc-cmp2-publisher-logo-container figure img{display:block !important}}#page_14 .filter-article-container{display:none}#newsletter_subscription .text-light{color:#fff !important}#bandeau-footer{display:none}</style> <link rel="stylesheet" id="typo-css" href="/_web/assets/css/typo.css?t=20240418-1" /><script>let cssLoaded=!1,bodyLoaded=!1;const loadedTO=setTimeout((()=>{cssLoaded=!0,bodyLoaded=!0,loaded()}),5e3),loaded=function(){cssLoaded&&bodyLoaded&&document.body.classList.contains("loading")&&(document.body.className=document.body.className.replace("loading","").trim(),clearTimeout(loadedTO))};</script> <link rel="preload" href="/_web/assets/css/lazy.css?t=20240418-1" as="style" media="all" crossorigin onload="this.rel=`stylesheet`; cssLoaded = true; loaded()" /><style>.breadcrumb{--bs-breadcrumb-padding-x: 0;--bs-breadcrumb-padding-y: 0;--bs-breadcrumb-margin-bottom: 1rem;--bs-breadcrumb-bg: ;--bs-breadcrumb-border-radius: ;--bs-breadcrumb-divider-color: #6c757d;--bs-breadcrumb-item-padding-x: 0.5rem;--bs-breadcrumb-item-active-color: #6c757d;display:flex;flex-wrap:wrap;padding:var(--bs-breadcrumb-padding-y) var(--bs-breadcrumb-padding-x);margin-bottom:var(--bs-breadcrumb-margin-bottom);font-size:var(--bs-breadcrumb-font-size);list-style:none;background-color:var(--bs-breadcrumb-bg);border-radius:var(--bs-breadcrumb-border-radius)}.breadcrumb-item+.breadcrumb-item{padding-left:var(--bs-breadcrumb-item-padding-x)}.breadcrumb-item+.breadcrumb-item::before{float:left;padding-right:var(--bs-breadcrumb-item-padding-x);color:var(--bs-breadcrumb-divider-color);content:var(--bs-breadcrumb-divider, "/") /* rtl: var(--bs-breadcrumb-divider, "/") */}.breadcrumb-item.active{color:var(--bs-breadcrumb-item-active-color)}.breadcrumb{padding:.5rem 1rem !important;background-color:#f5f5f5 !important;border-radius:.375rem !important}.breadcrumb .breadcrumb-item i.label-sub{font-size:.85em;font-weight:normal}.breadcrumb .breadcrumb-item a{text-decoration:none}.breadcrumb .breadcrumb-item a:hover{text-decoration:underline}.breadcrumb .breadcrumb-item.active{font-weight:bold}.breadcrumb .breadcrumb-item+.breadcrumb-item::before{background-image:url('data:image/svg+xml;charset=UTF-8, <svg aria-hidden="true" focusable="false" data-prefix="far" data-icon="chevron-double-right" role="img" xmlns="http://www.w3.org/2000/svg" viewBox="0 0 448 512" class="svg-inline--fa fa-chevron-double-right fa-w-14 fa-7x"><path style="fill: rgb(97, 104, 118);" d="M57.7 38.1l209.4 209.4c4.7 4.7 4.7 12.3 0 17L57.7 473.9c-4.7 4.7-12.3 4.7-17 0l-19.8-19.8c-4.7-4.7-4.7-12.3 0-17L201.6 256 20.9 74.9c-4.7-4.7-4.7-12.3 0-17l19.8-19.8c4.7-4.7 12.3-4.7 17 0zm143 0l-19.8 19.8c-4.7 4.7-4.7 12.3 0 17L361.6 256 180.9 437.1c-4.7 4.7-4.7 12.3 0 17l19.8 19.8c4.7 4.7 12.3 4.7 17 0l209.4-209.4c4.7-4.7 4.7-12.3 0-17L217.7 38.1c-4.7-4.7-12.3-4.7-17 0z" class=""></path></svg>');background-repeat:no-repeat;background-position:center;content:"";width:.5rem;height:100%;margin-right:.5rem}.list-group{--bs-list-group-color: #212529;--bs-list-group-bg: #fff;--bs-list-group-border-color: rgba(0, 0, 0, 0.125);--bs-list-group-border-width: 1px;--bs-list-group-border-radius: 0.375rem;--bs-list-group-item-padding-x: 1rem;--bs-list-group-item-padding-y: 0.5rem;--bs-list-group-action-color: #495057;--bs-list-group-action-hover-color: #495057;--bs-list-group-action-hover-bg: #f8f9fa;--bs-list-group-action-active-color: #212529;--bs-list-group-action-active-bg: #e9ecef;--bs-list-group-disabled-color: #6c757d;--bs-list-group-disabled-bg: #fff;--bs-list-group-active-color: #fff;--bs-list-group-active-bg: #526c78;--bs-list-group-active-border-color: #526c78;display:flex;flex-direction:column;padding-left:0;margin-bottom:0;border-radius:var(--bs-list-group-border-radius)}.list-group-numbered{list-style-type:none;counter-reset:section}.list-group-numbered>.list-group-item::before{content:counters(section, ".") ". ";counter-increment:section}.list-group-item-action{width:100%;color:var(--bs-list-group-action-color);text-align:inherit}.list-group-item-action:hover,.list-group-item-action:focus{z-index:1;color:var(--bs-list-group-action-hover-color);text-decoration:none;background-color:var(--bs-list-group-action-hover-bg)}.list-group-item-action:active{color:var(--bs-list-group-action-active-color);background-color:var(--bs-list-group-action-active-bg)}.list-group-item{position:relative;display:block;padding:var(--bs-list-group-item-padding-y) var(--bs-list-group-item-padding-x);color:var(--bs-list-group-color);text-decoration:none;background-color:var(--bs-list-group-bg);border:var(--bs-list-group-border-width) solid var(--bs-list-group-border-color)}.list-group-item:first-child{border-top-left-radius:inherit;border-top-right-radius:inherit}.list-group-item:last-child{border-bottom-right-radius:inherit;border-bottom-left-radius:inherit}.list-group-item.disabled,.list-group-item:disabled{color:var(--bs-list-group-disabled-color);pointer-events:none;background-color:var(--bs-list-group-disabled-bg)}.list-group-item.active{z-index:2;color:var(--bs-list-group-active-color);background-color:var(--bs-list-group-active-bg);border-color:var(--bs-list-group-active-border-color)}.list-group-item+.list-group-item{border-top-width:0}.list-group-item+.list-group-item.active{margin-top:calc(-1 * var(--bs-list-group-border-width));border-top-width:var(--bs-list-group-border-width)}.list-group-horizontal{flex-direction:row}.list-group-horizontal>.list-group-item:first-child:not(:last-child){border-bottom-left-radius:var(--bs-list-group-border-radius);border-top-right-radius:0}.list-group-horizontal>.list-group-item:last-child:not(:first-child){border-top-right-radius:var(--bs-list-group-border-radius);border-bottom-left-radius:0}.list-group-horizontal>.list-group-item.active{margin-top:0}.list-group-horizontal>.list-group-item+.list-group-item{border-top-width:var(--bs-list-group-border-width);border-left-width:0}.list-group-horizontal>.list-group-item+.list-group-item.active{margin-left:calc(-1 * var(--bs-list-group-border-width));border-left-width:var(--bs-list-group-border-width)}@media(min-width: 576px){.list-group-horizontal-sm{flex-direction:row}.list-group-horizontal-sm>.list-group-item:first-child:not(:last-child){border-bottom-left-radius:var(--bs-list-group-border-radius);border-top-right-radius:0}.list-group-horizontal-sm>.list-group-item:last-child:not(:first-child){border-top-right-radius:var(--bs-list-group-border-radius);border-bottom-left-radius:0}.list-group-horizontal-sm>.list-group-item.active{margin-top:0}.list-group-horizontal-sm>.list-group-item+.list-group-item{border-top-width:var(--bs-list-group-border-width);border-left-width:0}.list-group-horizontal-sm>.list-group-item+.list-group-item.active{margin-left:calc(-1 * var(--bs-list-group-border-width));border-left-width:var(--bs-list-group-border-width)}}@media(min-width: 768px){.list-group-horizontal-md{flex-direction:row}.list-group-horizontal-md>.list-group-item:first-child:not(:last-child){border-bottom-left-radius:var(--bs-list-group-border-radius);border-top-right-radius:0}.list-group-horizontal-md>.list-group-item:last-child:not(:first-child){border-top-right-radius:var(--bs-list-group-border-radius);border-bottom-left-radius:0}.list-group-horizontal-md>.list-group-item.active{margin-top:0}.list-group-horizontal-md>.list-group-item+.list-group-item{border-top-width:var(--bs-list-group-border-width);border-left-width:0}.list-group-horizontal-md>.list-group-item+.list-group-item.active{margin-left:calc(-1 * var(--bs-list-group-border-width));border-left-width:var(--bs-list-group-border-width)}}@media(min-width: 992px){.list-group-horizontal-lg{flex-direction:row}.list-group-horizontal-lg>.list-group-item:first-child:not(:last-child){border-bottom-left-radius:var(--bs-list-group-border-radius);border-top-right-radius:0}.list-group-horizontal-lg>.list-group-item:last-child:not(:first-child){border-top-right-radius:var(--bs-list-group-border-radius);border-bottom-left-radius:0}.list-group-horizontal-lg>.list-group-item.active{margin-top:0}.list-group-horizontal-lg>.list-group-item+.list-group-item{border-top-width:var(--bs-list-group-border-width);border-left-width:0}.list-group-horizontal-lg>.list-group-item+.list-group-item.active{margin-left:calc(-1 * var(--bs-list-group-border-width));border-left-width:var(--bs-list-group-border-width)}}@media(min-width: 1200px){.list-group-horizontal-xl{flex-direction:row}.list-group-horizontal-xl>.list-group-item:first-child:not(:last-child){border-bottom-left-radius:var(--bs-list-group-border-radius);border-top-right-radius:0}.list-group-horizontal-xl>.list-group-item:last-child:not(:first-child){border-top-right-radius:var(--bs-list-group-border-radius);border-bottom-left-radius:0}.list-group-horizontal-xl>.list-group-item.active{margin-top:0}.list-group-horizontal-xl>.list-group-item+.list-group-item{border-top-width:var(--bs-list-group-border-width);border-left-width:0}.list-group-horizontal-xl>.list-group-item+.list-group-item.active{margin-left:calc(-1 * var(--bs-list-group-border-width));border-left-width:var(--bs-list-group-border-width)}}@media(min-width: 1400px){.list-group-horizontal-xxl{flex-direction:row}.list-group-horizontal-xxl>.list-group-item:first-child:not(:last-child){border-bottom-left-radius:var(--bs-list-group-border-radius);border-top-right-radius:0}.list-group-horizontal-xxl>.list-group-item:last-child:not(:first-child){border-top-right-radius:var(--bs-list-group-border-radius);border-bottom-left-radius:0}.list-group-horizontal-xxl>.list-group-item.active{margin-top:0}.list-group-horizontal-xxl>.list-group-item+.list-group-item{border-top-width:var(--bs-list-group-border-width);border-left-width:0}.list-group-horizontal-xxl>.list-group-item+.list-group-item.active{margin-left:calc(-1 * var(--bs-list-group-border-width));border-left-width:var(--bs-list-group-border-width)}}.list-group-flush{border-radius:0}.list-group-flush>.list-group-item{border-width:0 0 var(--bs-list-group-border-width)}.list-group-flush>.list-group-item:last-child{border-bottom-width:0}.list-group-item-primary{color:#314148;background-color:#dce2e4}.list-group-item-primary.list-group-item-action:hover,.list-group-item-primary.list-group-item-action:focus{color:#314148;background-color:#c6cbcd}.list-group-item-primary.list-group-item-action.active{color:#fff;background-color:#314148;border-color:#314148}.list-group-item-secondary{color:#192023;background-color:#d4d7d8}.list-group-item-secondary.list-group-item-action:hover,.list-group-item-secondary.list-group-item-action:focus{color:#192023;background-color:#bfc2c2}.list-group-item-secondary.list-group-item-action.active{color:#fff;background-color:#192023;border-color:#192023}.list-group-item-success{color:#0f5132;background-color:#d1e7dd}.list-group-item-success.list-group-item-action:hover,.list-group-item-success.list-group-item-action:focus{color:#0f5132;background-color:#bcd0c7}.list-group-item-success.list-group-item-action.active{color:#fff;background-color:#0f5132;border-color:#0f5132}.list-group-item-info{color:#055160;background-color:#cff4fc}.list-group-item-info.list-group-item-action:hover,.list-group-item-info.list-group-item-action:focus{color:#055160;background-color:#badce3}.list-group-item-info.list-group-item-action.active{color:#fff;background-color:#055160;border-color:#055160}.list-group-item-warning{color:#664d03;background-color:#fff3cd}.list-group-item-warning.list-group-item-action:hover,.list-group-item-warning.list-group-item-action:focus{color:#664d03;background-color:#e6dbb9}.list-group-item-warning.list-group-item-action.active{color:#fff;background-color:#664d03;border-color:#664d03}.list-group-item-danger{color:#851a31;background-color:#f8d5dc}.list-group-item-danger.list-group-item-action:hover,.list-group-item-danger.list-group-item-action:focus{color:#851a31;background-color:#dfc0c6}.list-group-item-danger.list-group-item-action.active{color:#fff;background-color:#851a31;border-color:#851a31}.list-group-item-light{color:#5d5e5f;background-color:#fafbfb}.list-group-item-light.list-group-item-action:hover,.list-group-item-light.list-group-item-action:focus{color:#5d5e5f;background-color:#e1e2e2}.list-group-item-light.list-group-item-action.active{color:#fff;background-color:#5d5e5f;border-color:#5d5e5f}.list-group-item-dark{color:#192023;background-color:#d4d7d8}.list-group-item-dark.list-group-item-action:hover,.list-group-item-dark.list-group-item-action:focus{color:#192023;background-color:#bfc2c2}.list-group-item-dark.list-group-item-action.active{color:#fff;background-color:#192023;border-color:#192023}.badge{--bs-badge-padding-x: 0.65em;--bs-badge-padding-y: 0.35em;--bs-badge-font-size:0.75em;--bs-badge-font-weight: 700;--bs-badge-color: #fff;--bs-badge-border-radius: 0.375rem;display:inline-block;padding:var(--bs-badge-padding-y) var(--bs-badge-padding-x);font-size:var(--bs-badge-font-size);font-weight:var(--bs-badge-font-weight);line-height:1;color:var(--bs-badge-color);text-align:center;white-space:nowrap;vertical-align:baseline;border-radius:var(--bs-badge-border-radius)}.badge:empty{display:none}.btn .badge{position:relative;top:-1px}.btn-group,.btn-group-vertical{position:relative;display:inline-flex;vertical-align:middle}.btn-group>.btn,.btn-group-vertical>.btn{position:relative;flex:1 1 auto}.btn-group>.btn-check:checked+.btn,.btn-group>.btn-check:focus+.btn,.btn-group>.btn:hover,.btn-group>.btn:focus,.btn-group>.btn:active,.btn-group>.btn.active,.btn-group-vertical>.btn-check:checked+.btn,.btn-group-vertical>.btn-check:focus+.btn,.btn-group-vertical>.btn:hover,.btn-group-vertical>.btn:focus,.btn-group-vertical>.btn:active,.btn-group-vertical>.btn.active{z-index:1}.btn-toolbar{display:flex;flex-wrap:wrap;justify-content:flex-start}.btn-toolbar .input-group{width:auto}.btn-group{border-radius:.375rem}.btn-group>:not(.btn-check:first-child)+.btn,.btn-group>.btn-group:not(:first-child){margin-left:-1px}.btn-group>.btn:not(:last-child):not(.dropdown-toggle),.btn-group>.btn.dropdown-toggle-split:first-child,.btn-group>.btn-group:not(:last-child)>.btn{border-top-right-radius:0;border-bottom-right-radius:0}.btn-group>.btn:nth-child(n+3),.btn-group>:not(.btn-check)+.btn,.btn-group>.btn-group:not(:first-child)>.btn{border-top-left-radius:0;border-bottom-left-radius:0}.dropdown-toggle-split{padding-right:.5625rem;padding-left:.5625rem}.dropdown-toggle-split::after,.dropup .dropdown-toggle-split::after,.dropend .dropdown-toggle-split::after{margin-left:0}.dropstart .dropdown-toggle-split::before{margin-right:0}.btn-sm+.dropdown-toggle-split,.btn-group-sm>.btn+.dropdown-toggle-split{padding-right:.375rem;padding-left:.375rem}.btn-lg+.dropdown-toggle-split,.btn-group-lg>.btn+.dropdown-toggle-split{padding-right:.75rem;padding-left:.75rem}.btn-group-vertical{flex-direction:column;align-items:flex-start;justify-content:center}.btn-group-vertical>.btn,.btn-group-vertical>.btn-group{width:100%}.btn-group-vertical>.btn:not(:first-child),.btn-group-vertical>.btn-group:not(:first-child){margin-top:-1px}.btn-group-vertical>.btn:not(:last-child):not(.dropdown-toggle),.btn-group-vertical>.btn-group:not(:last-child)>.btn{border-bottom-right-radius:0;border-bottom-left-radius:0}.btn-group-vertical>.btn~.btn,.btn-group-vertical>.btn-group:not(:first-child)>.btn{border-top-left-radius:0;border-top-right-radius:0}.pagination{--bs-pagination-padding-x: 0.75rem;--bs-pagination-padding-y: 0.375rem;--bs-pagination-font-size:0.95rem;--bs-pagination-color: var(--bs-link-color);--bs-pagination-bg: #fff;--bs-pagination-border-width: 1px;--bs-pagination-border-color: #dee2e6;--bs-pagination-border-radius: 0.375rem;--bs-pagination-hover-color: var(--bs-link-hover-color);--bs-pagination-hover-bg: #e9ecef;--bs-pagination-hover-border-color: #dee2e6;--bs-pagination-focus-color: var(--bs-link-hover-color);--bs-pagination-focus-bg: #e9ecef;--bs-pagination-focus-box-shadow: 0 0 0 0.25rem rgba(82, 108, 120, 0.25);--bs-pagination-active-color: #fff;--bs-pagination-active-bg: #526c78;--bs-pagination-active-border-color: #526c78;--bs-pagination-disabled-color: #6c757d;--bs-pagination-disabled-bg: #fff;--bs-pagination-disabled-border-color: #dee2e6;display:flex;padding-left:0;list-style:none}.page-link{position:relative;display:block;padding:var(--bs-pagination-padding-y) var(--bs-pagination-padding-x);font-size:var(--bs-pagination-font-size);color:var(--bs-pagination-color);text-decoration:none;background-color:var(--bs-pagination-bg);border:var(--bs-pagination-border-width) solid var(--bs-pagination-border-color);transition:color .15s ease-in-out,background-color .15s ease-in-out,border-color .15s ease-in-out,box-shadow .15s ease-in-out}@media(prefers-reduced-motion: reduce){.page-link{transition:none}}.page-link:hover{z-index:2;color:var(--bs-pagination-hover-color);background-color:var(--bs-pagination-hover-bg);border-color:var(--bs-pagination-hover-border-color)}.page-link:focus{z-index:3;color:var(--bs-pagination-focus-color);background-color:var(--bs-pagination-focus-bg);outline:0;box-shadow:var(--bs-pagination-focus-box-shadow)}.page-link.active,.active>.page-link{z-index:3;color:var(--bs-pagination-active-color);background-color:var(--bs-pagination-active-bg);border-color:var(--bs-pagination-active-border-color)}.page-link.disabled,.disabled>.page-link{color:var(--bs-pagination-disabled-color);pointer-events:none;background-color:var(--bs-pagination-disabled-bg);border-color:var(--bs-pagination-disabled-border-color)}.page-item:not(:first-child) .page-link{margin-left:-1px}.page-item:first-child .page-link{border-top-left-radius:var(--bs-pagination-border-radius);border-bottom-left-radius:var(--bs-pagination-border-radius)}.page-item:last-child .page-link{border-top-right-radius:var(--bs-pagination-border-radius);border-bottom-right-radius:var(--bs-pagination-border-radius)}.pagination-lg{--bs-pagination-padding-x: 1.5rem;--bs-pagination-padding-y: 0.75rem;--bs-pagination-font-size:1.1875rem;--bs-pagination-border-radius: 0.5rem}.pagination-sm{--bs-pagination-padding-x: 0.5rem;--bs-pagination-padding-y: 0.25rem;--bs-pagination-font-size:0.83125rem;--bs-pagination-border-radius: 0.25rem}.list-group a.list-group-item{border-color:#f8fafc}.list-group a.list-group-item.active{background-color:#526c78}.list-group a.list-group-item h3{margin:0;line-height:1.5;font-size:.95rem;font-family:system-ui,-apple-system,"Segoe UI",Roboto,"Helvetica Neue","Noto Sans","Liberation Sans",Arial,sans-serif,"Apple Color Emoji","Segoe UI Emoji","Segoe UI Symbol","Noto Color Emoji";padding-bottom:0}.list-group a.list-group-item h3:hover{color:#fff}.list-group a.list-group-item h3 .badge{line-height:1.25}.list-group a.list-group-item:hover{background-color:#526c78;color:#fff}.list-group a.list-group-item:hover h3{color:inherit}i.anchor{display:block;position:relative;visibility:hidden;top:calc(-25vh + 15px)}.article__props{font-size:.8rem;color:#000;font-style:italic;padding:.2rem 0;margin:0 auto;display:block}.content h6,.h6,h5,.h5,h4,.h4,h3,.h3,h2,.h2,h1,.h1{margin-bottom:0;padding-bottom:.5rem}titre_main+titre_subs{margin-left:.25em}pc,.pc{text-transform:uppercase;font-size:.8em}h2 .pc,h3 .pc,h4 .pc,h5 .pc,h6 .pc{font-size:inherit !important}.sup-sub{position:relative;margin-right:1rem}.sup-sub sup,.sup-sub sub,.sup-sub inf{position:absolute;display:block;left:0}.sup-sub sup{top:.25rem}.sup-sub sub,.sup-sub inf{bottom:.25rem}p:empty{display:none !important}.content.whole .read-more{display:none}blockquote{position:relative;text-align:center;padding:1rem 1.4rem;width:80%;margin:1rem auto 2rem;font-family:Times;font-style:italic;box-shadow:0 .5rem 1rem rgba(0,0,0,.15);border-radius:.375rem}blockquote:before{position:absolute;color:#526c78;font-size:3rem;content:"«";top:-22px;margin-right:-28px;right:100%}blockquote:after{position:absolute;color:#526c78;font-size:3rem;content:"»";margin-left:-28px;left:100%;top:auto;bottom:-12px}.offcanvas,.offcanvas-xxl,.offcanvas-xl,.offcanvas-lg,.offcanvas-md,.offcanvas-sm{--bs-offcanvas-zindex: 1045;--bs-offcanvas-width: 400px;--bs-offcanvas-height: 30vh;--bs-offcanvas-padding-x: 1rem;--bs-offcanvas-padding-y: 1rem;--bs-offcanvas-color: ;--bs-offcanvas-bg: #fff;--bs-offcanvas-border-width: 1px;--bs-offcanvas-border-color: var(--bs-border-color-translucent);--bs-offcanvas-box-shadow: 0 0.125rem 0.25rem rgba(0, 0, 0, 0.075)}@media(max-width: 575.98px){.offcanvas-sm{position:fixed;bottom:0;z-index:var(--bs-offcanvas-zindex);display:flex;flex-direction:column;max-width:100%;color:var(--bs-offcanvas-color);visibility:hidden;background-color:var(--bs-offcanvas-bg);background-clip:padding-box;outline:0;transition:transform .3s ease-in-out}}@media(max-width: 575.98px)and (prefers-reduced-motion: reduce){.offcanvas-sm{transition:none}}@media(max-width: 575.98px){.offcanvas-sm.offcanvas-start{top:0;left:0;width:var(--bs-offcanvas-width);border-right:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateX(-100%)}}@media(max-width: 575.98px){.offcanvas-sm.offcanvas-end{top:0;right:0;width:var(--bs-offcanvas-width);border-left:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateX(100%)}}@media(max-width: 575.98px){.offcanvas-sm.offcanvas-top{top:0;right:0;left:0;height:var(--bs-offcanvas-height);max-height:100%;border-bottom:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateY(-100%)}}@media(max-width: 575.98px){.offcanvas-sm.offcanvas-bottom{right:0;left:0;height:var(--bs-offcanvas-height);max-height:100%;border-top:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateY(100%)}}@media(max-width: 575.98px){.offcanvas-sm.showing,.offcanvas-sm.show:not(.hiding){transform:none}}@media(max-width: 575.98px){.offcanvas-sm.showing,.offcanvas-sm.hiding,.offcanvas-sm.show{visibility:visible}}@media(min-width: 576px){.offcanvas-sm{--bs-offcanvas-height: auto;--bs-offcanvas-border-width: 0;background-color:transparent !important}.offcanvas-sm .offcanvas-header{display:none}.offcanvas-sm .offcanvas-body{display:flex;flex-grow:0;padding:0;overflow-y:visible;background-color:transparent !important}}@media(max-width: 767.98px){.offcanvas-md{position:fixed;bottom:0;z-index:var(--bs-offcanvas-zindex);display:flex;flex-direction:column;max-width:100%;color:var(--bs-offcanvas-color);visibility:hidden;background-color:var(--bs-offcanvas-bg);background-clip:padding-box;outline:0;transition:transform .3s ease-in-out}}@media(max-width: 767.98px)and (prefers-reduced-motion: reduce){.offcanvas-md{transition:none}}@media(max-width: 767.98px){.offcanvas-md.offcanvas-start{top:0;left:0;width:var(--bs-offcanvas-width);border-right:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateX(-100%)}}@media(max-width: 767.98px){.offcanvas-md.offcanvas-end{top:0;right:0;width:var(--bs-offcanvas-width);border-left:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateX(100%)}}@media(max-width: 767.98px){.offcanvas-md.offcanvas-top{top:0;right:0;left:0;height:var(--bs-offcanvas-height);max-height:100%;border-bottom:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateY(-100%)}}@media(max-width: 767.98px){.offcanvas-md.offcanvas-bottom{right:0;left:0;height:var(--bs-offcanvas-height);max-height:100%;border-top:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateY(100%)}}@media(max-width: 767.98px){.offcanvas-md.showing,.offcanvas-md.show:not(.hiding){transform:none}}@media(max-width: 767.98px){.offcanvas-md.showing,.offcanvas-md.hiding,.offcanvas-md.show{visibility:visible}}@media(min-width: 768px){.offcanvas-md{--bs-offcanvas-height: auto;--bs-offcanvas-border-width: 0;background-color:transparent !important}.offcanvas-md .offcanvas-header{display:none}.offcanvas-md .offcanvas-body{display:flex;flex-grow:0;padding:0;overflow-y:visible;background-color:transparent !important}}@media(max-width: 991.98px){.offcanvas-lg{position:fixed;bottom:0;z-index:var(--bs-offcanvas-zindex);display:flex;flex-direction:column;max-width:100%;color:var(--bs-offcanvas-color);visibility:hidden;background-color:var(--bs-offcanvas-bg);background-clip:padding-box;outline:0;transition:transform .3s ease-in-out}}@media(max-width: 991.98px)and (prefers-reduced-motion: reduce){.offcanvas-lg{transition:none}}@media(max-width: 991.98px){.offcanvas-lg.offcanvas-start{top:0;left:0;width:var(--bs-offcanvas-width);border-right:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateX(-100%)}}@media(max-width: 991.98px){.offcanvas-lg.offcanvas-end{top:0;right:0;width:var(--bs-offcanvas-width);border-left:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateX(100%)}}@media(max-width: 991.98px){.offcanvas-lg.offcanvas-top{top:0;right:0;left:0;height:var(--bs-offcanvas-height);max-height:100%;border-bottom:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateY(-100%)}}@media(max-width: 991.98px){.offcanvas-lg.offcanvas-bottom{right:0;left:0;height:var(--bs-offcanvas-height);max-height:100%;border-top:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateY(100%)}}@media(max-width: 991.98px){.offcanvas-lg.showing,.offcanvas-lg.show:not(.hiding){transform:none}}@media(max-width: 991.98px){.offcanvas-lg.showing,.offcanvas-lg.hiding,.offcanvas-lg.show{visibility:visible}}@media(min-width: 992px){.offcanvas-lg{--bs-offcanvas-height: auto;--bs-offcanvas-border-width: 0;background-color:transparent !important}.offcanvas-lg .offcanvas-header{display:none}.offcanvas-lg .offcanvas-body{display:flex;flex-grow:0;padding:0;overflow-y:visible;background-color:transparent !important}}@media(max-width: 1199.98px){.offcanvas-xl{position:fixed;bottom:0;z-index:var(--bs-offcanvas-zindex);display:flex;flex-direction:column;max-width:100%;color:var(--bs-offcanvas-color);visibility:hidden;background-color:var(--bs-offcanvas-bg);background-clip:padding-box;outline:0;transition:transform .3s ease-in-out}}@media(max-width: 1199.98px)and (prefers-reduced-motion: reduce){.offcanvas-xl{transition:none}}@media(max-width: 1199.98px){.offcanvas-xl.offcanvas-start{top:0;left:0;width:var(--bs-offcanvas-width);border-right:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateX(-100%)}}@media(max-width: 1199.98px){.offcanvas-xl.offcanvas-end{top:0;right:0;width:var(--bs-offcanvas-width);border-left:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateX(100%)}}@media(max-width: 1199.98px){.offcanvas-xl.offcanvas-top{top:0;right:0;left:0;height:var(--bs-offcanvas-height);max-height:100%;border-bottom:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateY(-100%)}}@media(max-width: 1199.98px){.offcanvas-xl.offcanvas-bottom{right:0;left:0;height:var(--bs-offcanvas-height);max-height:100%;border-top:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateY(100%)}}@media(max-width: 1199.98px){.offcanvas-xl.showing,.offcanvas-xl.show:not(.hiding){transform:none}}@media(max-width: 1199.98px){.offcanvas-xl.showing,.offcanvas-xl.hiding,.offcanvas-xl.show{visibility:visible}}@media(min-width: 1200px){.offcanvas-xl{--bs-offcanvas-height: auto;--bs-offcanvas-border-width: 0;background-color:transparent !important}.offcanvas-xl .offcanvas-header{display:none}.offcanvas-xl .offcanvas-body{display:flex;flex-grow:0;padding:0;overflow-y:visible;background-color:transparent !important}}@media(max-width: 1399.98px){.offcanvas-xxl{position:fixed;bottom:0;z-index:var(--bs-offcanvas-zindex);display:flex;flex-direction:column;max-width:100%;color:var(--bs-offcanvas-color);visibility:hidden;background-color:var(--bs-offcanvas-bg);background-clip:padding-box;outline:0;transition:transform .3s ease-in-out}}@media(max-width: 1399.98px)and (prefers-reduced-motion: reduce){.offcanvas-xxl{transition:none}}@media(max-width: 1399.98px){.offcanvas-xxl.offcanvas-start{top:0;left:0;width:var(--bs-offcanvas-width);border-right:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateX(-100%)}}@media(max-width: 1399.98px){.offcanvas-xxl.offcanvas-end{top:0;right:0;width:var(--bs-offcanvas-width);border-left:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateX(100%)}}@media(max-width: 1399.98px){.offcanvas-xxl.offcanvas-top{top:0;right:0;left:0;height:var(--bs-offcanvas-height);max-height:100%;border-bottom:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateY(-100%)}}@media(max-width: 1399.98px){.offcanvas-xxl.offcanvas-bottom{right:0;left:0;height:var(--bs-offcanvas-height);max-height:100%;border-top:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateY(100%)}}@media(max-width: 1399.98px){.offcanvas-xxl.showing,.offcanvas-xxl.show:not(.hiding){transform:none}}@media(max-width: 1399.98px){.offcanvas-xxl.showing,.offcanvas-xxl.hiding,.offcanvas-xxl.show{visibility:visible}}@media(min-width: 1400px){.offcanvas-xxl{--bs-offcanvas-height: auto;--bs-offcanvas-border-width: 0;background-color:transparent !important}.offcanvas-xxl .offcanvas-header{display:none}.offcanvas-xxl .offcanvas-body{display:flex;flex-grow:0;padding:0;overflow-y:visible;background-color:transparent !important}}.offcanvas{position:fixed;bottom:0;z-index:var(--bs-offcanvas-zindex);display:flex;flex-direction:column;max-width:100%;color:var(--bs-offcanvas-color);visibility:hidden;background-color:var(--bs-offcanvas-bg);background-clip:padding-box;outline:0;transition:transform .3s ease-in-out}@media(prefers-reduced-motion: reduce){.offcanvas{transition:none}}.offcanvas.offcanvas-start{top:0;left:0;width:var(--bs-offcanvas-width);border-right:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateX(-100%)}.offcanvas.offcanvas-end{top:0;right:0;width:var(--bs-offcanvas-width);border-left:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateX(100%)}.offcanvas.offcanvas-top{top:0;right:0;left:0;height:var(--bs-offcanvas-height);max-height:100%;border-bottom:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateY(-100%)}.offcanvas.offcanvas-bottom{right:0;left:0;height:var(--bs-offcanvas-height);max-height:100%;border-top:var(--bs-offcanvas-border-width) solid var(--bs-offcanvas-border-color);transform:translateY(100%)}.offcanvas.showing,.offcanvas.show:not(.hiding){transform:none}.offcanvas.showing,.offcanvas.hiding,.offcanvas.show{visibility:visible}.offcanvas-backdrop{position:fixed;top:0;left:0;z-index:1040;width:100vw;height:100vh;background-color:#000}.offcanvas-backdrop.fade{opacity:0}.offcanvas-backdrop.show{opacity:.5}.offcanvas-header{display:flex;align-items:center;justify-content:space-between;padding:var(--bs-offcanvas-padding-y) var(--bs-offcanvas-padding-x)}.offcanvas-header .btn-close{padding:calc(var(--bs-offcanvas-padding-y) * .5) calc(var(--bs-offcanvas-padding-x) * .5);margin-top:calc(-.5 * var(--bs-offcanvas-padding-y));margin-right:calc(-.5 * var(--bs-offcanvas-padding-x));margin-bottom:calc(-.5 * var(--bs-offcanvas-padding-y))}.offcanvas-title{margin-bottom:0;line-height:1.5}.offcanvas-body{flex-grow:1;padding:var(--bs-offcanvas-padding-y) var(--bs-offcanvas-padding-x);overflow-y:auto}.article__single__toc .float-end{float:right !important}.article__single__toc .list-group-item{border:none;border-radius:0}.article__single__toc .list-group-item i{transition:transform .2s;position:absolute;left:.25rem;margin-top:.25rem}.article__single__toc .list-group-item.expanded i{transform:rotateZ(90deg)}.article__single__toc>.list-group{background:transparent}.article__single__toc>.list-group>.list-group{margin-left:.5em}.article__single__toc>.list-group>.list-group>.list-group{margin-left:.75em;border-left:1px solid #ccc}.article__single__toc>.list-group>.list-group>.list-group>.list-group-item{padding:.5em}#callSommaire{position:fixed;bottom:10px;left:10px;z-index:2}.free-toc-container{position:fixed;bottom:3em;left:.5em;max-width:300px;z-index:10}@media(max-width: 768px){.free-toc-container{bottom:5em}}#free-toc{max-height:calc(100vh - 275px);overflow:auto}.free-toc-container .list-group a{text-decoration:none}.free-toc-container .list-group li:not(.active):hover{background-color:#f5f5f5;color:#fff}.free-toc-container .list-group .active a{color:#fff;pointer-events:none;font-weight:bold}.pagination{--bs-pagination-padding-x: 0.75rem;--bs-pagination-padding-y: 0.375rem;--bs-pagination-font-size:0.95rem;--bs-pagination-color: var(--bs-link-color);--bs-pagination-bg: #fff;--bs-pagination-border-width: 1px;--bs-pagination-border-color: #dee2e6;--bs-pagination-border-radius: 0.375rem;--bs-pagination-hover-color: var(--bs-link-hover-color);--bs-pagination-hover-bg: #e9ecef;--bs-pagination-hover-border-color: #dee2e6;--bs-pagination-focus-color: var(--bs-link-hover-color);--bs-pagination-focus-bg: #e9ecef;--bs-pagination-focus-box-shadow: 0 0 0 0.25rem rgba(82, 108, 120, 0.25);--bs-pagination-active-color: #fff;--bs-pagination-active-bg: #526c78;--bs-pagination-active-border-color: #526c78;--bs-pagination-disabled-color: #6c757d;--bs-pagination-disabled-bg: #fff;--bs-pagination-disabled-border-color: #dee2e6;display:flex;padding-left:0;list-style:none}.page-link{position:relative;display:block;padding:var(--bs-pagination-padding-y) var(--bs-pagination-padding-x);font-size:var(--bs-pagination-font-size);color:var(--bs-pagination-color);text-decoration:none;background-color:var(--bs-pagination-bg);border:var(--bs-pagination-border-width) solid var(--bs-pagination-border-color);transition:color .15s ease-in-out,background-color .15s ease-in-out,border-color .15s ease-in-out,box-shadow .15s ease-in-out}@media(prefers-reduced-motion: reduce){.page-link{transition:none}}.page-link:hover{z-index:2;color:var(--bs-pagination-hover-color);background-color:var(--bs-pagination-hover-bg);border-color:var(--bs-pagination-hover-border-color)}.page-link:focus{z-index:3;color:var(--bs-pagination-focus-color);background-color:var(--bs-pagination-focus-bg);outline:0;box-shadow:var(--bs-pagination-focus-box-shadow)}.page-link.active,.active>.page-link{z-index:3;color:var(--bs-pagination-active-color);background-color:var(--bs-pagination-active-bg);border-color:var(--bs-pagination-active-border-color)}.page-link.disabled,.disabled>.page-link{color:var(--bs-pagination-disabled-color);pointer-events:none;background-color:var(--bs-pagination-disabled-bg);border-color:var(--bs-pagination-disabled-border-color)}.page-item:not(:first-child) .page-link{margin-left:-1px}.page-item:first-child .page-link{border-top-left-radius:var(--bs-pagination-border-radius);border-bottom-left-radius:var(--bs-pagination-border-radius)}.page-item:last-child .page-link{border-top-right-radius:var(--bs-pagination-border-radius);border-bottom-right-radius:var(--bs-pagination-border-radius)}.pagination-lg{--bs-pagination-padding-x: 1.5rem;--bs-pagination-padding-y: 0.75rem;--bs-pagination-font-size:1.1875rem;--bs-pagination-border-radius: 0.5rem}.pagination-sm{--bs-pagination-padding-x: 0.5rem;--bs-pagination-padding-y: 0.25rem;--bs-pagination-font-size:0.83125rem;--bs-pagination-border-radius: 0.25rem}.table{--bs-table-color: var(--bs-body-color);--bs-table-bg: transparent;--bs-table-border-color: var(--bs-border-color);--bs-table-accent-bg: transparent;--bs-table-striped-color: var(--bs-body-color);--bs-table-striped-bg: rgba(0, 0, 0, 0.05);--bs-table-active-color: var(--bs-body-color);--bs-table-active-bg: rgba(0, 0, 0, 0.1);--bs-table-hover-color: var(--bs-body-color);--bs-table-hover-bg: rgba(0, 0, 0, 0.075);width:100%;margin-bottom:1rem;color:var(--bs-table-color);vertical-align:top;border-color:var(--bs-table-border-color)}.table>:not(caption)>*>*{padding:.5rem .5rem;background-color:var(--bs-table-bg);border-bottom-width:1px;box-shadow:inset 0 0 0 9999px var(--bs-table-accent-bg)}.table>tbody{vertical-align:inherit}.table>thead{vertical-align:bottom}.table-group-divider{border-top:2px solid currentcolor}.caption-top{caption-side:top}.table-sm>:not(caption)>*>*{padding:.25rem .25rem}.table-bordered>:not(caption)>*{border-width:1px 0}.table-bordered>:not(caption)>*>*{border-width:0 1px}.table-borderless>:not(caption)>*>*{border-bottom-width:0}.table-borderless>:not(:first-child){border-top-width:0}.table-striped>tbody>tr:nth-of-type(odd)>*{--bs-table-accent-bg: var(--bs-table-striped-bg);color:var(--bs-table-striped-color)}.table-striped-columns>:not(caption)>tr>:nth-child(even){--bs-table-accent-bg: var(--bs-table-striped-bg);color:var(--bs-table-striped-color)}.table-active{--bs-table-accent-bg: var(--bs-table-active-bg);color:var(--bs-table-active-color)}.table-hover>tbody>tr:hover>*{--bs-table-accent-bg: var(--bs-table-hover-bg);color:var(--bs-table-hover-color)}.table-primary{--bs-table-color: #000;--bs-table-bg: #dce2e4;--bs-table-border-color: #c6cbcd;--bs-table-striped-bg: #d1d7d9;--bs-table-striped-color: #000;--bs-table-active-bg: #c6cbcd;--bs-table-active-color: #000;--bs-table-hover-bg: #ccd1d3;--bs-table-hover-color: #000;color:var(--bs-table-color);border-color:var(--bs-table-border-color)}.table-secondary{--bs-table-color: #000;--bs-table-bg: #e2e3e5;--bs-table-border-color: #cbccce;--bs-table-striped-bg: #d7d8da;--bs-table-striped-color: #000;--bs-table-active-bg: #cbccce;--bs-table-active-color: #000;--bs-table-hover-bg: #d1d2d4;--bs-table-hover-color: #000;color:var(--bs-table-color);border-color:var(--bs-table-border-color)}.table-success{--bs-table-color: #000;--bs-table-bg: #d1e7dd;--bs-table-border-color: #bcd0c7;--bs-table-striped-bg: #c7dbd2;--bs-table-striped-color: #000;--bs-table-active-bg: #bcd0c7;--bs-table-active-color: #000;--bs-table-hover-bg: #c1d6cc;--bs-table-hover-color: #000;color:var(--bs-table-color);border-color:var(--bs-table-border-color)}.table-info{--bs-table-color: #000;--bs-table-bg: #cff4fc;--bs-table-border-color: #badce3;--bs-table-striped-bg: #c5e8ef;--bs-table-striped-color: #000;--bs-table-active-bg: #badce3;--bs-table-active-color: #000;--bs-table-hover-bg: #bfe2e9;--bs-table-hover-color: #000;color:var(--bs-table-color);border-color:var(--bs-table-border-color)}.table-warning{--bs-table-color: #000;--bs-table-bg: #fff3cd;--bs-table-border-color: #e6dbb9;--bs-table-striped-bg: #f2e7c3;--bs-table-striped-color: #000;--bs-table-active-bg: #e6dbb9;--bs-table-active-color: #000;--bs-table-hover-bg: #ece1be;--bs-table-hover-color: #000;color:var(--bs-table-color);border-color:var(--bs-table-border-color)}.table-danger{--bs-table-color: #000;--bs-table-bg: #f8d5dc;--bs-table-border-color: #dfc0c6;--bs-table-striped-bg: #eccad1;--bs-table-striped-color: #000;--bs-table-active-bg: #dfc0c6;--bs-table-active-color: #000;--bs-table-hover-bg: #e5c5cc;--bs-table-hover-color: #000;color:var(--bs-table-color);border-color:var(--bs-table-border-color)}.table-light{--bs-table-color: #000;--bs-table-bg: #e8eced;--bs-table-border-color: #d1d4d5;--bs-table-striped-bg: #dce0e1;--bs-table-striped-color: #000;--bs-table-active-bg: #d1d4d5;--bs-table-active-color: #000;--bs-table-hover-bg: #d7dadb;--bs-table-hover-color: #000;color:var(--bs-table-color);border-color:var(--bs-table-border-color)}.table-dark{--bs-table-color: #fff;--bs-table-bg: #29363a;--bs-table-border-color: #3e4a4e;--bs-table-striped-bg: #344044;--bs-table-striped-color: #fff;--bs-table-active-bg: #3e4a4e;--bs-table-active-color: #fff;--bs-table-hover-bg: #394549;--bs-table-hover-color: #fff;color:var(--bs-table-color);border-color:var(--bs-table-border-color)}.table-responsive{overflow-x:auto;-webkit-overflow-scrolling:touch}@media(max-width: 575.98px){.table-responsive-sm{overflow-x:auto;-webkit-overflow-scrolling:touch}}@media(max-width: 767.98px){.table-responsive-md{overflow-x:auto;-webkit-overflow-scrolling:touch}}@media(max-width: 991.98px){.table-responsive-lg{overflow-x:auto;-webkit-overflow-scrolling:touch}}@media(max-width: 1199.98px){.table-responsive-xl{overflow-x:auto;-webkit-overflow-scrolling:touch}}@media(max-width: 1399.98px){.table-responsive-xxl{overflow-x:auto;-webkit-overflow-scrolling:touch}}span.rg{font-weight:bold}.lined-bg{background:repeating-linear-gradient(45deg, #606dbc, #606dbc 10px, #465298 10px, #465298 20px)}@media(min-width: 768px){.geography-list-group{grid-template-columns:repeat(2, 1fr);display:grid;grid-gap:.5rem}.geography-list-group .list-group-item+.list-group-item{border-top-width:1px}}@media(min-width: 1200px){.geography-list-group{grid-template-columns:repeat(3, 1fr);display:grid;grid-gap:.5rem}.geography-list-group .list-group-item+.list-group-item{border-top-width:1px}}table .note{font-size:.8rem;line-height:1.1;position:relative;padding-left:.8rem;display:block;margin-bottom:.5rem;font-style:italic}table .note b{position:absolute;font-style:normal;left:0;top:0}h1 em.label-sub{font-style:normal}.event-date{font-size:1rem;font-style:italic;color:#526c78}.event-label{font-weight:bold;margin-bottom:0}.event-label .event-date{font-size:.8em}.badge{background-color:#e8eced !important;border-radius:0;color:#526c78}.badge.text-bg-primary{color:#fff !important;background-color:#0d6efd !important}.badge.text-bg-success{color:#fff !important;background-color:#198754 !important}.badge .bad a{text-decoration:underline}.event-single .badge{font-size:.85rem;font-weight:normal;margin-top:-0.25rem;padding-left:1rem !important;padding-right:1rem !important}.figure{display:flex;flex-direction:column;justify-content:center}@media(max-width: 768px){.figure{background:#f5f5f5;box-shadow:0 .125rem .25rem rgba(0,0,0,.075);margin-bottom:1em !important;padding-top:.5em !important}.figure .popover{background:transparent}.figure .popover .popover-header{color:#222}}img.media_resized{max-width:100%;height:auto;width:auto;margin:0 auto;max-height:200px}.protected{position:relative}.protected:after{position:absolute;content:"";width:100%;height:100%;top:0;left:0;background:transparent}@media(min-width: 992px){.inline-media-group{display:flex;justify-content:space-around}}@media screen and (max-width: 480px){.nav.nav-tabs{display:block}.nav.nav-tabs .nav-item{display:block !important}.nav.nav-tabs .nav-link{border:none;width:100%;margin:0}.nav.nav-tabs .nav-link.active{background-color:#e8eced;margin:0}}.index-list{border:1px solid #dfdfdf;background-color:#fafafa;border-radius:.375em;box-shadow:0 .125rem .25rem rgba(0,0,0,.075) !important;margin-bottom:1rem !important;margin-right:.25rem !important;margin-left:.25rem !important;padding:1rem}.index-list a{padding:.125em .25em;border-radius:.375em;display:inline-block}.index-list a.active{background:#526c78;color:#fff;text-decoration:none}.index-list a.text-muted{color:#a4a9ae !important}.card{--bs-card-spacer-y: 1rem;--bs-card-spacer-x: 1rem;--bs-card-title-spacer-y: 0.5rem;--bs-card-border-width: 1px;--bs-card-border-color: var(--bs-border-color-translucent);--bs-card-border-radius: 0.375rem;--bs-card-box-shadow: ;--bs-card-inner-border-radius: calc(0.375rem - 1px);--bs-card-cap-padding-y: 0.5rem;--bs-card-cap-padding-x: 1rem;--bs-card-cap-bg: rgba(0, 0, 0, 0.03);--bs-card-cap-color: ;--bs-card-height: ;--bs-card-color: ;--bs-card-bg: #fff;--bs-card-img-overlay-padding: 1rem;--bs-card-group-margin: 0.75rem;position:relative;display:flex;flex-direction:column;min-width:0;height:var(--bs-card-height);word-wrap:break-word;background-color:var(--bs-card-bg);background-clip:border-box;border:var(--bs-card-border-width) solid var(--bs-card-border-color);border-radius:var(--bs-card-border-radius)}.card>hr{margin-right:0;margin-left:0}.card>.list-group{border-top:inherit;border-bottom:inherit}.card>.list-group:first-child{border-top-width:0;border-top-left-radius:var(--bs-card-inner-border-radius);border-top-right-radius:var(--bs-card-inner-border-radius)}.card>.list-group:last-child{border-bottom-width:0;border-bottom-right-radius:var(--bs-card-inner-border-radius);border-bottom-left-radius:var(--bs-card-inner-border-radius)}.card>.card-header+.list-group,.card>.list-group+.card-footer{border-top:0}.card-body{flex:1 1 auto;padding:var(--bs-card-spacer-y) var(--bs-card-spacer-x);color:var(--bs-card-color)}.card-title{margin-bottom:var(--bs-card-title-spacer-y)}.card-subtitle{margin-top:calc(-.5 * var(--bs-card-title-spacer-y));margin-bottom:0}.card-text:last-child{margin-bottom:0}.card-link+.card-link{margin-left:var(--bs-card-spacer-x)}.card-header{padding:var(--bs-card-cap-padding-y) var(--bs-card-cap-padding-x);margin-bottom:0;color:var(--bs-card-cap-color);background-color:var(--bs-card-cap-bg);border-bottom:var(--bs-card-border-width) solid var(--bs-card-border-color)}.card-header:first-child{border-radius:var(--bs-card-inner-border-radius) var(--bs-card-inner-border-radius) 0 0}.card-footer{padding:var(--bs-card-cap-padding-y) var(--bs-card-cap-padding-x);color:var(--bs-card-cap-color);background-color:var(--bs-card-cap-bg);border-top:var(--bs-card-border-width) solid var(--bs-card-border-color)}.card-footer:last-child{border-radius:0 0 var(--bs-card-inner-border-radius) var(--bs-card-inner-border-radius)}.card-header-tabs{margin-right:calc(-.5 * var(--bs-card-cap-padding-x));margin-bottom:calc(-1 * var(--bs-card-cap-padding-y));margin-left:calc(-.5 * var(--bs-card-cap-padding-x));border-bottom:0}.card-header-tabs .nav-link.active{background-color:var(--bs-card-bg);border-bottom-color:var(--bs-card-bg)}.card-header-pills{margin-right:calc(-.5 * var(--bs-card-cap-padding-x));margin-left:calc(-.5 * var(--bs-card-cap-padding-x))}.card-img-overlay{position:absolute;top:0;right:0;bottom:0;left:0;padding:var(--bs-card-img-overlay-padding);border-radius:var(--bs-card-inner-border-radius)}.card-img,.card-img-top,.card-img-bottom{width:100%}.card-img,.card-img-top{border-top-left-radius:var(--bs-card-inner-border-radius);border-top-right-radius:var(--bs-card-inner-border-radius)}.card-img,.card-img-bottom{border-bottom-right-radius:var(--bs-card-inner-border-radius);border-bottom-left-radius:var(--bs-card-inner-border-radius)}.card-group>.card{margin-bottom:var(--bs-card-group-margin)}@media(min-width: 576px){.card-group{display:flex;flex-flow:row wrap}.card-group>.card{flex:1 0 0%;margin-bottom:0}.card-group>.card+.card{margin-left:0;border-left:0}.card-group>.card:not(:last-child){border-top-right-radius:0;border-bottom-right-radius:0}.card-group>.card:not(:last-child) .card-img-top,.card-group>.card:not(:last-child) .card-header{border-top-right-radius:0}.card-group>.card:not(:last-child) .card-img-bottom,.card-group>.card:not(:last-child) .card-footer{border-bottom-right-radius:0}.card-group>.card:not(:first-child){border-top-left-radius:0;border-bottom-left-radius:0}.card-group>.card:not(:first-child) .card-img-top,.card-group>.card:not(:first-child) .card-header{border-top-left-radius:0}.card-group>.card:not(:first-child) .card-img-bottom,.card-group>.card:not(:first-child) .card-footer{border-bottom-left-radius:0}}.alert{--bs-alert-bg: transparent;--bs-alert-padding-x: 1rem;--bs-alert-padding-y: 1rem;--bs-alert-margin-bottom: 1rem;--bs-alert-color: inherit;--bs-alert-border-color: transparent;--bs-alert-border: 1px solid var(--bs-alert-border-color);--bs-alert-border-radius: 0.375rem;position:relative;padding:var(--bs-alert-padding-y) var(--bs-alert-padding-x);margin-bottom:var(--bs-alert-margin-bottom);color:var(--bs-alert-color);background-color:var(--bs-alert-bg);border:var(--bs-alert-border);border-radius:var(--bs-alert-border-radius)}.alert-heading{color:inherit}.alert-link{font-weight:700}.alert-dismissible{padding-right:3rem}.alert-dismissible .btn-close{position:absolute;top:0;right:0;z-index:2;padding:1.25rem 1rem}.alert-primary{--bs-alert-color: #314148;--bs-alert-bg: #dce2e4;--bs-alert-border-color: #cbd3d7}.alert-primary .alert-link{color:#27343a}.alert-secondary{--bs-alert-color: #192023;--bs-alert-bg: #d4d7d8;--bs-alert-border-color: #bfc3c4}.alert-secondary .alert-link{color:#141a1c}.alert-success{--bs-alert-color: #0f5132;--bs-alert-bg: #d1e7dd;--bs-alert-border-color: #badbcc}.alert-success .alert-link{color:#0c4128}.alert-info{--bs-alert-color: #055160;--bs-alert-bg: #cff4fc;--bs-alert-border-color: #b6effb}.alert-info .alert-link{color:#04414d}.alert-warning{--bs-alert-color: #664d03;--bs-alert-bg: #fff3cd;--bs-alert-border-color: #ffecb5}.alert-warning .alert-link{color:#523e02}.alert-danger{--bs-alert-color: #851a31;--bs-alert-bg: #f8d5dc;--bs-alert-border-color: #f5bfcb}.alert-danger .alert-link{color:#6a1527}.alert-light{--bs-alert-color: #5d5e5f;--bs-alert-bg: #fafbfb;--bs-alert-border-color: #f8f9fa}.alert-light .alert-link{color:#4a4b4c}.alert-dark{--bs-alert-color: #192023;--bs-alert-bg: #d4d7d8;--bs-alert-border-color: #bfc3c4}.alert-dark .alert-link{color:#141a1c}.media-landing .media-reference-copy{display:block;position:relative !important;top:auto;left:auto;width:auto;height:auto}.media-landing .media-reference-copy:after{content:"Afficher le média";background:rgba(255,255,255,.1);position:absolute;top:0;left:0;width:100%;height:100%;display:flex;align-items:center;justify-content:center;color:#222;font-size:2em;font-weight:bold;text-shadow:0 0 5px #fff;backdrop-filter:blur(5px)}.media-landing .media-reference-copy img{width:auto !important;height:auto !important}</style><script type="application/ld+json"> { "@context": "https://schema.org", "@type": "WebSite", "name": "Encyclopædia Universalis", "alternateName": "Universalis", "url": "https://www.universalis.fr", "sameAs": [ "https://www.facebook.com/EncyclopedieUniversalis/", "https://www.linkedin.com/company/encyclopaedia-universalis", "https://www.twitter.com/universalisfr", "https://www.youtube.com/channel/UC0srTZiJaRaDw3fAvtCo4Ow" ] } </script> </head> <body id="page_3" class="edu SENIOR loading" > <div class="loader"></div> <a href="#main-content" id="skip"> <i aria-hidden="true" class="fa fa-fw fa-arrow-down"></i> Accéder au contenu </a> <header class="d-none d-md-block d-print-none"> <div class="container row-between"> <a class="header-logo mr-3" title="Retour à la page d'accueil" href="/"> <svg viewBox="0 0 370 76.44" height="70" width="277" id="logo-main"> <text class="fill-universalis text-darkblue" transform="translate(74.25 37.1)" font-family="Arial" font-size="43.37"> <tspan x="0" y="0">UNIVERSALIS</tspan> </text> <text class="text-darkblue subtitle" transform="translate(76.75 66.74)" font-family="Arial" font-size="35.8"> <tspan class="fill-universalis-up" x="0" y="0" font-size=".85em">.fr</tspan> </text> <g> <path class="fill-universalis-up" d="M57,6.36H4.25C1.9,6.36,0,8.27,0,10.62V62.51c0,2.35,1.9,4.25,4.25,4.25H57c2.35,0,4.25-1.9,4.25-4.25V10.62c0-2.35-1.9-4.25-4.25-4.25ZM29.63,31.87h7.33v8.06c-.77-1.77-1.61-2.99-2.96-4.38-1.41-1.44-2.42-2.35-4.36-3.29v-.39Zm2.37,29.2c-.85,.68-1.75,1.28-2.71,1.75-.11,.05-.22,.1-.33,.15H13.39c-1.67-.77-3.19-1.8-4.53-3.14-3.13-3.13-4.69-7.06-4.69-11.77s1.56-8.84,4.67-12.04c3.12-3.19,7.02-4.79,11.72-4.79,2.71,0,5.14,.49,7.3,1.43,1.81,.79,3.44,1.9,4.87,3.36,1.35,1.38,2.4,2.97,3.17,4.74,1,2.31,1.52,4.94,1.52,7.9l-.03,1.51H11.86c.37,2.25,1.36,4.04,2.95,5.37,1.6,1.33,3.64,1.99,6.13,1.99,2.79,0,4.96-.67,6.99-2.49l6.74,3.26c-.84,1.12-1.61,1.9-2.67,2.75Zm24.74-15.61c0,5.53-.44,9.31-1.31,11.35-1.07,2.49-2.64,4.39-4.72,5.72-.24,.15-.49,.3-.75,.44h-14.33c-.44-.23-.87-.49-1.26-.77-.26-.18-.43-.29-.66-.49,.29-.3,2.13-2.54,3.11-3.92,0,0-7.13-3.32-7.3-3.39-.22-1.04-.28-1.34-.44-2.83h7.98c.1,1.11,.27,1.26,.5,1.93,.41,1.15,1.07,2.05,1.97,2.69,.9,.64,2.02,.96,3.34,.96s2.45-.32,3.37-.95c.92-.63,1.61-1.55,2.06-2.78,.33-.91,.5-2.86,.5-5.85v-15.51h7.95v13.42Z"/> <path class="fill-universalis-up" d="M29.44,43.92c-.32-1.09-.87-2.04-1.58-2.88-.44-.52-.93-1-1.52-1.43-1.54-1.11-3.34-1.66-5.38-1.66-2.22,0-4.16,.62-5.83,1.86-1.05,.78-2.02,2.14-2.92,4.11H29.44Z"/> </g> </svg> </a> <div class="p-1 d-flex justify-content-end flex-column" style="height: 80px"> <div class="user-welcome"></div> <div class="login-form-wrapper" style="align-items: flex-end; display: flex; flex-direction: column" data-identification_uri="/identification/"> <div class="login-form"> <div class="btn btn-sm btn-primary keep-label-space-top"> <i aria-hidden="true" class="fa fa-user"></i> Mon Compte </div> </div> </div> <div id="toast-manager"></div> </div> </div> </header> <div class="d-print-none sticky-top"> <div role="navigation" aria-label="Navigation mobile" class="main-navbar-wrapper shadow"> <a class="navbar-brand-md d-none d-md-block" href="/"> <img alt="Universalis" src=" /typo3temp/assets/_processed_/e/b/csm_eu-blanc-2x_ca998e8b78.webp " width="32" height="32" alt=""> </a> <div class="container"> <nav class="navbar navbar-expand-lg navbar-dark"> <a class="navbar-brand d-md-none" href="/"> <img alt="Universalis" src=" /typo3temp/assets/_processed_/e/b/csm_eu-blanc-2x_ca998e8b78.webp " width="36" height="36" alt=""> </a> <div id="navbar-collapse-grid" class="navbar-collapse collapse pt-2 pt-md-0"> <ul class="navbar-nav me-auto"> <li><a href="/atlas/" title="Atlas" class="nav-link">Atlas</a></li><li><a href="/classification/" title="Classification" class="nav-link">Classification</a></li><li><a href="/dictionnaire/" title="Dictionnaire français Universalis : retrouvez toutes les définitions, étymologies, expressions, synonymes, exemples, et flexions de plus de 120 000 mots de la langue française." class="nav-link">Dictionnaire</a></li> <li class="nav-item dropdown"> <a class="nav-link dropdown-toggle" href="#" role="button" data-bs-toggle="dropdown" aria-expanded="false"> Données Pays </a> <ul class="dropdown-menu"> <li> <a class="dropdown-item" href="https://www.universalis.fr/donnees-pays/"><i class="fa fa-wand-magic-sparkles" aria-hidden="true"></i> Comparateur</a> </li> <li><hr class="dropdown-divider"></li> <li> <a class="dropdown-item" href="/donnees-pays/indicateur/liste/"><i class="fa fa-chart-line" aria-hidden="true"></i> Données</a> </li> <li> <a class="dropdown-item" href="/donnees-pays/pays/liste/"><i class="fa fa-globe" aria-hidden="true"></i> Pays</a> </li> <li><hr class="dropdown-divider"/></li> <li><button class="dropdown-item" data-bs-toggle="modal" data-bs-target="#wd-help"><i class="fa fa-circle-info" aria-hidden="true"></i> Aide</button></li> </ul> </li> <li class="nav-item"> <a href="https://boutique.universalis.fr" class="nav-link">Boutique</a> </li> </ul> </div> <form method="get" name="searchQuery" class="flex-grow-1 my-1" id="search-form" autocomplete="off" action="/recherche/redirect/"> <div> <input type="hidden" name="tx_solrdata_pi1[__referrer][@extension]" value="SolrData" /> <input type="hidden" name="tx_solrdata_pi1[__referrer][@controller]" value="SearchForm" /> <input type="hidden" name="tx_solrdata_pi1[__referrer][@action]" value="showFr" /> <input type="hidden" name="tx_solrdata_pi1[__referrer][arguments]" value="YTowOnt9789d077876a7a0adeb4a9a4c4853f315af9bb201" /> <input type="hidden" name="tx_solrdata_pi1[__referrer][@request]" value="{"@extension":"SolrData","@controller":"SearchForm","@action":"showFr"}db953a457671272659a1e7710096726899d321a7" /> <input type="hidden" name="tx_solrdata_pi1[__trustedProperties]" value="{"data-action":1,"searchQuery":{"resultType":1,"searchedString":1}}36e15bfcc110b23d6ba91a6810381c8ec6b87146" /> </div> <input type="hidden" name="tx_solrdata_pi1[data-action]" value="https://www.universalis.fr/recherche/___searchedString___/article/1/" /> <input type="hidden" name="tx_solrdata_pi1[searchQuery][resultType]" value="ARTICLE" /> <div class="input-wrapper input-group"> <input placeholder="Rechercher sur Universalis" class="form-control" id="searchedString" autocomplete="off" type="text" name="tx_solrdata_pi1[searchQuery][searchedString]" value="" /> <button type="submit" class="btn btn-sm btn-light"> <span class="d-none d-md-block">Recherche</span> <span class="visually-hidden">Recherche</span> <span class="d-md-none"> <i aria-hidden="true" class="fa fa-fw fa-magnifying-glass"></i></span> </button> </div> </form> <button title="Afficher / masquer le menu du site" class="navbar-toggler ms-1" type="button" data-bs-toggle="collapse" data-bs-target="#navbar-collapse-grid"> <span class="navbar-toggler-icon"></span> </button> </nav> </div> </div> </div> <main class="site-fr container main rounded-bottom shadow mb-3"> <div class="user-welcome"></div> <div class="login-form-wrapper-mobile"> <div class="login-form"> <div class="btn btn-sm btn-primary keep-label-space-top"> <i aria-hidden="true" class="fa fa-user"></i> Mon Compte </div> </div> </div> <div class="d-lg-none text-center mb-3"> <a href="/abonnement" title="Je m'abonne" data-matomo-category="CTA" data-matomo-name="Clic" data-matomo-value="Top (mobile)"> <img src="/typo3temp/assets/_processed_/1/6/csm_banner-cta_431c3102be.webp" width="728" height="90" alt="Abonnez-vous à Universalis pour 1 euro" class="img-max shadow-sm"/> </a> </div> <div class="row"> <div id="main-content" class="col-lg-8 col-xl-9 pb-4"> <h1 class="mb-1"> <strong>STAMINODE</strong> </h1> <ul class="pagination justify-content-center public-links "><li class="page-item active"><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 1" class="page-link" title="Abonnez-vous" data-loaded="true">1</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 2" class="page-link" title="Abonnez-vous" data-loaded="true">2</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 3" class="page-link" title="Abonnez-vous" data-loaded="true">3</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 4" class="page-link" title="Abonnez-vous" data-loaded="true">4</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 5" class="page-link" title="Abonnez-vous" data-loaded="true">5</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 6" class="page-link" title="Abonnez-vous" data-loaded="true">6</a></li></ul><!-- f:if condition=""><i aria-hidden="true" id="chiffres"></i><div class="card factbox my-3 mx-auto shadow-sm"><div class="card-body p-0"><table class="table table-sm table-striped mb-0"><tbody></tbody></table><nav class="d-flex justify-content-center d-print-none"><a href=" / " class="btn btn-sm btn-primary flex-grow-1 m-1"><i aria-hidden="true" class="fa fa-fw fa-globe"></i> Cartes </a></nav></div></div></f:if --><p class="text-sm text-muted mb-1">Article modifié le <time datetime="2025-01-29"> 29/01/2025 </time></p><div class="content whole type-article" ><div class="block_1" data-ucode="yT322135_0"><i class="anchor" id="yT322135_0"></i><p> Nom donné, en morphologie florale, à une étamine transformée en une pièce stérile, c'est-à-dire dépourvue de sac pollinique, et qui prend souvent l'apparence d'une languette. On en trouve dans la <a data-bs-toggle="popover" data-bs-content="La fleur n'est pas un organe morphologiquement défini. Chez les Angiospermes les moins évoluées, la fleur est évidemment, comme Goethe lui-même l'avait découvert, un rameau feuillé (système d'axes foliarisés) très contracté, dans le prolongement de dispositifs déjà réalisés chez des Végétaux vasculaires archaïques (Coniférophytes, Cycadophytes et même Cryptogames vasculai..." title="Article : FLEUR" href="/encyclopedie/fleur/">fleur</a> de la consoude, du tilleul argenté, du cacaoyer par exemple. La fleur des cannas ne possède qu'une demi-étamine produisant du pollen ; l'autre moitié de cette étamine, ainsi que les cinq autres, sont transformées en staminodes, pièces rouges stériles que l'on prendrait à première vue pour les pétales ; les pétales proprement dits sont très réduits, les sépales aussi ; l'un des staminodes est même transformé en « labelle », accentuant la forte <a data-bs-toggle="popover" data-bs-content="Type de symétrie florale dans lequel les pièces sont symétriquement disposées par rapport à un plan. Les fleurs sont dites alors zygomorphes ou irrégulières par opposition aux fleurs actinomorphes. Ce plan de symétrie bilatérale peut contenir l'axe de l'inflorescence : la moitié droite de la fleur est symétrique de la moitié gauche (zygomorphie médiane ; exemples : le pois, la ..." title="Article : ZYGOMORPHIE" href="/encyclopedie/zygomorphie/">zygomorphie</a> de cette fleur, très adaptée à la fécondation par les insectes.</p><div class="clearfix"></div><p class="author"> — Jacques DAUTA</p><div class="clearfix"></div></div><div class="clearfix"></div></div><ul class="pagination justify-content-center public-links "><li class="page-item active"><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 1" class="page-link" title="Abonnez-vous" data-loaded="true">1</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 2" class="page-link" title="Abonnez-vous" data-loaded="true">2</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 3" class="page-link" title="Abonnez-vous" data-loaded="true">3</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 4" class="page-link" title="Abonnez-vous" data-loaded="true">4</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 5" class="page-link" title="Abonnez-vous" data-loaded="true">5</a></li><li class="page-item "><a href="/abonnement" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="page 6" class="page-link" title="Abonnez-vous" data-loaded="true">6</a></li></ul><div class="folio-block"><p class="folio-block-title text-center text-uc"><strong>La suite de cet article est accessible aux abonnés</strong></p><ul class="list-unstyled"><li><i class="fa fa-fw fa-check-circle"></i> Des contenus variés, complets et fiables</li><li><i class="fa fa-fw fa-check-circle"></i> Accessible sur tous les écrans</li><li><i class="fa fa-fw fa-check-circle"></i> Pas de publicité</li></ul><p class="text-center"><a href="/abonnement/" class="btn btn-cta text-uc" id="folio-button" data-matomo-category="Article" data-matomo-name="Folio" data-matomo-value="Block bottom">Découvrez nos offres </a></p><p class="text-center text-sm"> Déjà abonné ? <a href="/identification/" id="folio-login">Se connecter</a></p></div><div id="ad_mpuslot" class="pub-container text-center"><div class="pub-placeholder"></div></div><section class="mt-3 mb-5"><i class="anchor" id="T322135_AUTHORS"></i><p class="title-minor">Écrit par</p><ul class="list-group"><li class="list-group-item" data-dc-author="DAUTA, Jacques"><a title="Auteur : Jacques DAUTA" rel="author" href="/auteurs/jacques-dauta/"><strong>Jacques DAUTA</strong></a> : assistant à l'université de Paris-VI </li></ul></section><section class="mb-5"><div class="text-center mb-3 d-print-none"><a class="d-inline-flex flex-row justify-content-center align-items-center btn btn-light btn-large text-start" title="Carte mentale : STAMINODE" href="/carte-mentale/staminode/"><i class="fa fa-sitemap fa-rotate-270 me-1 fs-1 text-universalis"></i><p class="m-0"><strong class="fs-4">Carte mentale</strong><br /> Élargissez votre recherche dans Universalis </p></a></div></section> <section class="mb-5 article-classification" data-print-label="la classification" data-print-target=".article-classification"> <i class="anchor" id="yT322135_ROOT_LINES"></i> <p class="title-minor">Classification</p> <nav aria-label="Classification 1"> <ol class="breadcrumb"> <li class="breadcrumb-item"> <a title="Classification : Sciences de la vie" href="/classification/sciences-de-la-vie/"> Sciences de la vie </a> </li> <li class="breadcrumb-item"> <a title="Classification : Botanique" href="/classification/sciences-de-la-vie/botanique/"> Botanique </a> </li> <li class="breadcrumb-item"> <a title="Classification : Biologie végétale" href="/classification/sciences-de-la-vie/botanique/biologie-vegetale/"> Biologie végétale </a> </li> <li class="breadcrumb-item active" data-dc-subject="Anatomie végétale"> <a title="Classification : Anatomie végétale" href="/classification/sciences-de-la-vie/botanique/biologie-vegetale/anatomie-vegetale/"> Anatomie végétale </a> </li> </ol> </nav> </section> <section class="mb-5 d-print-none learn-more"> <i class="anchor" id="yT322135_REFERENCES"></i> <ul class="list-group"> </ul> </section> <section class="mb-5 d-print-none article-indexes learn-more"> <i class="anchor" id="yT322135_INDEXES"></i> <p class="title-minor">Voir aussi</p> <ul class="list-inline list-inline-bullet"> <li class="list-inline-item m-1"> <a title="Index : ÉTAMINE" href="/index/etamine/"> <span class="rg">ÉTAMINE</span> </a> </li> <li class="list-inline-item m-1"> <a title="Index : PÉTALE" href="/index/petale/"> <span class="rg">PÉTALE</span> </a> </li> <li class="list-inline-item m-1"> <a title="Index : ANATOMIE ET MORPHOLOGIE VÉGÉTALE" href="/index/anatomie-vegetale/"> <span class="rg">ANATOMIE ET MORPHOLOGIE VÉGÉTALE</span> </a> </li> </ul> </section> </div> <div class="col-lg-4 col-xl-3 px-0 d-none d-lg-block side-bar position-relative"> <div id="ad_rightslot" class="pub-container text-center"> <div class="pub-placeholder"></div> </div> <div id="ad_rightslot2" class="pub-container text-center"> <div class="pub-placeholder"></div> </div> <a class="wrapper cta" href="/abonnement" title="Abonnez-vous à Universalis pour 1 euro" data-matomo-category="CTA" data-matomo-name="Clic" data-matomo-value="Col right"> <img src="/typo3temp/assets/_processed_/b/6/csm_abo-3_fd5f912304.webp" class="full-height" loading="lazy" width="290" height="476" alt="Abonnez-vous à Universalis pour 1 euro"> </a> <div id="newsletter_subscription" class="wrapper p-3"> <p class="fs-3 fw-bold text-center text-light mb-1">Rejoignez-nous</p> <p class="text-center"> Inscrivez-vous à notre <strong>newsletter</strong></p> <div class="input-group mb-3"> <input type="text" class="form-control" placeholder="votre email"> <div class="input-group-append"> <button id="newsletter_subscription_button" class="btn btn-primary text-uppercase" type="button"> <strong>S'inscrire</strong></button> </div> </div> </div> </div> </div> </main> <div id="bandeau-footer"> <div class="container"> <p><a href="/abonnement" title="Abonnez-vous à Universalis pour 1 euro">Accédez à l'intégralité d'Universalis.fr <strong>sans publicité</strong></a></p> </div> </div> <footer class="container footer text-center d-print-none"> <ul class="list-inline list-inline-bullet mb-2"> <li class="list-inline-item"> <a href="http://www.encyclopaedia-universalis.fr/" title="Encyclopædia Universalis" rel="noopener" target="_blank">Encyclopædia Universalis </a> </li> <li class="list-inline-item"> <a href="https://www.encyclopaedia-universalis.fr/nous-contacter/infos/" title="Contactez-nous" rel="noopener nofollow" target="_blank">Contact</a></li> <li class="list-inline-item"> <a title="Mentions légales" rel="nofollow" href="/mentions-legales/">Mentions légales et politique de confidentialité</a> </li> <li class="list-inline-item"> <a title="Crédits images Accueil" rel="nofollow" href="/credits-illustrations/">Crédits images</a> </li> <li class="list-inline-item"> <a href="https://abonnement.universalis.fr/typo3conf/ext/mailer/Resources/Public/Pdf/cgv.pdf" rel="noopener nofollow" target="_blank" title="Conditions générales de vente">CGV</a> </li> <li class="list-inline-item"> <a href="javascript:" id="rgpd" rel="nofollow" title="Consentement RGPD">RGPD</a> </li> </ul> <p class="mb-2">Consulter le <a title="Dictionnaire de l'Encyclopædia Universalis" href="/dictionnaire/"> dictionnaire </a> de l'Encyclopædia Universalis </p> <p class="mb-3">© 2025 Encyclopædia Universalis France. Tous droits de propriété industrielle et intellectuelle réservés.</p> </footer> <a href="javascript:" class="btn btn-primary" id="back-to-top" title="Retour en haut de page"><i aria-hidden="true" class="far fa-arrow-up"></i></a> <script> // noinspection ES6ConvertVarToLetConst,JSUnusedGlobalSymbols var subscription_uri = 'https://www.universalis.fr/abonnement/' bodyLoaded = true; loaded(); </script> <div class="modal" id="wd-help" tabindex="-1" style="display:none"> <div class="modal-dialog modal-xl"> <div class="modal-content"> <div class="modal-header"> <h5 class="modal-title">Aide</h5> <button type="button" class="btn-close" data-bs-dismiss="modal" aria-label="Close"></button> </div> <div class="modal-body"> <p>L'outil Données Pays met à disposition un très grand nombre de données. Aussi, pour des raisons de performances et pour conserver une bonne expérience utilisateur, des règles de comparaison en nombre et en durée ont été mises en place :</p> <table class="table table-vcenter table-bordered table-sm"> <thead> <tr> <th>Pays</th> <th>Indicateurs</th> <th>Années</th> <th>Graphe</th> </tr> </thead> <tbody> <tr> <td>Jusqu'à 12 pays</td> <td>Toutes les données</td> <td>De 1960 à nos jours</td> <td> <ul class="list-unstyled m-0"> <li> <i class="fa fa-check text-success" aria-hidden="true"></i> <span class="visually-hidden">Affiché</span> en courbe </li> <li> <i class="fa fa-check text-success" aria-hidden="true"></i> <span class="visually-hidden">Affiché</span> en barre </li> </ul> </td> </tr> <tr> <td>De 13 à 50 pays</td> <td>5 données simultanées</td> <td>De 1960 à nos jours</td> <td> <ul class="list-unstyled m-0"> <li> <i class="fa fa-times text-danger" aria-hidden="true"></i> <span class="visually-hidden">Masqué</span> en courbe </li> <li> <i class="fa fa-check text-success" aria-hidden="true"></i> <span class="visually-hidden">Affiché</span> en barre </li> </ul> </td> </tr> <tr> <td>51 pays et plus</td> <td>1 seule donnée</td> <td>Sur une durée de 25 ans maximum</td> <td> <ul class="list-unstyled m-0"> <li> <i class="fa fa-times text-danger" aria-hidden="true"></i> <span class="visually-hidden">Masqué</span> en courbe </li> <li> <i class="fa fa-times text-danger" aria-hidden="true"></i> <span class="visually-hidden">Masqué</span> en barre </li> </ul> </td> </tr> </tbody> </table> <p>Lorsque la recherche porte sur <strong>une seule année</strong>, les résultats s’affichent sous forme de <strong>graphe en barre</strong>.</p> <p>Lorsque la recherche porte sur <strong>une période</strong>, les résultats s’affichent sous forme de <strong>graphe en courbe</strong>.</p> <p>Lorsque la recherche contient uniquement des données <strong>Informations générales</strong>, le champ <strong>Année ou période est désactivé</strong>, ces données ne possédant pas d'historique.</p> <p class="mb-0 text-sm text-muted text-end">v2.0</p> </div> </div> </div> </div> <script>(()=>{var e,a,r,_,c,d={},b={};function __webpack_require__(e){var a=b[e];if(void 0!==a)return a.exports;var r=b[e]={id:e,loaded:!1,exports:{}};return d[e].call(r.exports,r,r.exports,__webpack_require__),r.loaded=!0,r.exports}__webpack_require__.m=d,__webpack_require__.n=e=>{var a=e&&e.__esModule?()=>e.default:()=>e;return __webpack_require__.d(a,),a},__webpack_require__.d=(e,a)=>{for(var r in a)__webpack_require__.o(a,r)&&!__webpack_require__.o(e,r)&&Object.defineProperty(e,r,{enumerable:!0,get:a[r]})},__webpack_require__.f={},__webpack_require__.e=e=>Promise.all(Object.keys(__webpack_require__.f).reduce(((a,r)=>(__webpack_require__.f[r](e,a),a)),[])),__webpack_require__.u=e=>"js/"+e+"."+{177:"333ddd8e201baf8b0ef8",190:"a2441ee3e33c1b0dfb46",227:"981873bb3ea9a3e2e3f4",305:"c88c90542337f7fdd2de",346:"9ae5854f9802be2b3f01",477:"d09be99baa6950c38795",741:"a93fdc8c8a0167d66f8f",780:"9353cebb460c5e2269f8",786:"138eb76e7e39d1ce2ef6",789:"7efe182d60e54790dce7",830:"3ed10eef4abea47b3440",840:"42480cfc5e9984993429",849:"e05b8d506ec312e78384",882:"da3184782d3541205108",883:"b504c671ee56b3f5edf8",1031:"71fa0551886a59bcf1ca",1120:"7324fe97941817a45d24",1142:"fc1a3a90c8a7cd0d7807",1222:"77e66626c184b7d31555",1480:"fef98c6a57f99f65fda8",1552:"a2d448d7d22d351a95bd",1641:"20b900e0adda4c80db1d",1731:"7271fc6a9e929960b210",1752:"eddbaf2c7e344a179c08",1800:"1379f1df3ace760a2e7c",1926:"7f59c7a726bec3d92f10",2003:"f48ccb771cd49cb7cf66",2136:"13dd4555663ea6eb4188",2307:"be05dd39a77150ede1ff",2363:"05f46879c7f3e969d3e5",2385:"c3da4f2c357252c9df08",2528:"35cf15469387068e6a85",2895:"a8f0413fcf2fee539813",3015:"e1084ababa33d08a81f0",3138:"94e862fda8508652322f",3343:"95ea7335a61f647d7e16",3507:"a636b4602eb43840133a",3630:"d0cc0fd84b36f88e15e6",3706:"15030bd90e02872595a8",3758:"957c19e64d42e47c33c7",4010:"bf993b78bd832234df6e",4047:"da30a71b441aa46bb8e5",4116:"94b769711c17c522d8c9",4280:"a995ffa72fa330a58630",4304:"d36dfb828762700c9765",4354:"4e0a70f1a566added829",4415:"7b18d362eeeac2a0f13a",4441:"dca649f001491a2bd607",4465:"308ce3c81e5393f25865",4595:"c37517f72707c874a2b1",4692:"b98cac1fb67e4e908d64",4709:"b47e5b787871f5b71d2b",4766:"8ca54e680606959b402d",4769:"934ef9da0386bf0b5e50",4793:"801bcc6ee25931aea123",4800:"976c343f52e42fa93fdc",4882:"ac4bf9b5216b41e62db4",5037:"45597a02f6e5efbbe8ab",5097:"70b5339794033b90dc7f",5107:"55931eae2cd4b39a0de0",5156:"dfe562b76cf0ff9f24e0",5263:"88eb3ab427a4cbc934b9",5324:"87f3337881bf106bccdd",5366:"046ebce9f739eb2e0c89",5423:"c13018ad8a1f937349ca",5486:"1fed20421da47434c298",5506:"aa3617dbc66db7f1c9a1",5614:"c9a8ac19be4023b7b250",5738:"6466f626e725a42b3698",5975:"88ee9cdd28f0245914f8",6036:"057d4e35fe7d6aedfd70",6037:"e7cdaffc1d87265d5627",6111:"c25dd0a941839ee93c38",6251:"36a514e5dc40273fc7e1",6423:"a8e0bb7db6d3fded7dda",6600:"f619d552df8bcc9fecb5",7033:"6a366be1fc4535fb0efb",7096:"25a463e58dd1075911f1",7204:"3e5234153f1f490db272",7278:"ab7009f38bbb5732c15d",7319:"c14f96c5a30814af47fa",7555:"a5f570d29bfea1ef7506",7604:"530b168f12bee7c9f974",7686:"0c417333213158f7d38b",7700:"e6a6a14ecf4012ebdafc",7766:"6f944a74026e1fe1264f",7900:"05c142408eaaa8435ffd",8025:"13ae5cfe5d2bbbd536b9",8113:"1a49932d3f9f58a0e19b",8141:"0226899ac199d372352a",8264:"55490fb5d4a9513d0175",8515:"eedd52d13507ac3fc95b",8615:"b508710cbd8a37db7ec5",8642:"1b6778bd5cc7a3c2bc58",8776:"8830e16a52d7de41cbc0",8871:"37df2861ac3d33fe4501",9112:"8b5424231c22fe32757a",9207:"d925c96ca941b509bb11",9416:"6ee4ffb0e861caab07f2",9424:"6538e064c5cb624ae4c6",9428:"6764efb670f6393e3957",9759:"a6a9959235213314d1b6",9797:"d76c4091e0c7d0b6b3f9",9845:"8ce2f2d84affc2a07c6a",9871:"f06db479356ec8fadfeb",9957:"75378c34d0a170e87382"}[e]+".bundle.js",__webpack_require__.miniCssF=e=>4826===e?"css/index.css":"css/"+e+".css",__webpack_require__.g=function(){if("object"==typeof globalThis)return globalThis;try{return this||new Function("return this")()}catch(e){if("object"==typeof window)return window}}(),__webpack_require__.o=(e,a)=>Object.prototype.hasOwnProperty.call(e,a),e={},a="@universalis/web_fr:",__webpack_require__.l=(r,_,c,d)=>{if(e[r])e[r].push(_);else{var b,f;if(void 0!==c)for(var t=document.getElementsByTagName("script"),i=0;i<t.length;i++){var n=t[i];if(n.getAttribute("src")==r||n.getAttribute("data-webpack")==a+c){b=n;break}}b||(f=!0,(b=document.createElement("script")).charset="utf-8",b.timeout=120,__webpack_require__.nc&&b.setAttribute("nonce",__webpack_require__.nc),b.setAttribute("data-webpack",a+c),b.src=r),e[r]=[_];var onScriptComplete=(a,_)=>{b.onerror=b.onload=null,clearTimeout(u);var c=e[r];if(delete e[r],b.parentNode&&b.parentNode.removeChild(b),c&&c.forEach((e=>e(_))),a)return a(_)},u=setTimeout(onScriptComplete.bind(null,void 0,{type:"timeout",target:b}),12e4);b.onerror=onScriptComplete.bind(null,b.onerror),b.onload=onScriptComplete.bind(null,b.onload),f&&document.head.appendChild(b)}},__webpack_require__.r=e=>{"undefined"!=typeof Symbol&&Symbol.toStringTag&&Object.defineProperty(e,Symbol.toStringTag,{value:"Module"}),Object.defineProperty(e,"__esModule",{value:!0})},__webpack_require__.nmd=e=>(e.paths=[],e.children||(e.children=[]),e),__webpack_require__.p="/_web/assets/",r=(e,a,r)=>{var _=document.createElement("link");return _.rel="stylesheet",_.type="text/css",_.onload=a,_.onerror=function(a){var c=a&&a.target&&a.target.src||e,d=new Error("Loading CSS chunk "+chunkId+" failed.\n("+c+")");d.code="CSS_CHUNK_LOAD_FAILED",d.request=c,_.parentNode.removeChild(_),r(d)},_.href=e,document.getElementsByTagName("head")[0].appendChild(_),_},_=e=>new Promise(((a,_)=>{var c=__webpack_require__.miniCssF(e),d=__webpack_require__.p+c;if(((e,a)=>{for(var r=document.getElementsByTagName("link"),_=0;_<r.length;_++){var c=(b=r[_]).getAttribute("data-href")||b.getAttribute("href");if("stylesheet"===b.rel&&(c===e||c===a))return b}var d=document.getElementsByTagName("style");for(_=0;_<d.length;_++){var b;if((c=(b=d[_]).getAttribute("data-href"))===e||c===a)return b}})(c,d))return a();r(d,a,_)})),c={4826:0},__webpack_require__.f.miniCss=(e,a)=>{c[e]?a.push(c[e]):0!==c[e]&&{2528:1,3343:1,3507:1,3630:1,3706:1,4769:1,5366:1,6036:1,9428:1,9845:1}[e]&&a.push(c[e]=_(e).then((()=>{c[e]=0}),(a=>{throw delete c[e],a})))},(()=>{var e={4826:0};__webpack_require__.f.j=(a,r)=>{var _=__webpack_require__.o(e,a)?e[a]:void 0;if(0!==_)if(_)r.push(_[2]);else{var c=new Promise(((r,c)=>_=e[a]=[r,c]));r.push(_[2]=c);var d=__webpack_require__.p+__webpack_require__.u(a),b=new Error;__webpack_require__.l(d,(r=>{if(__webpack_require__.o(e,a)&&(0!==(_=e[a])&&(e[a]=void 0),_)){var c=r&&("load"===r.type?"missing":r.type),d=r&&r.target&&r.target.src;b.message="Loading chunk "+a+" failed.\n("+c+": "+d+")",b.name="ChunkLoadError",b.type=c,b.request=d,_[1](b)}}),"chunk-"+a,a)}};var webpackJsonpCallback=(a,r)=>{var _,c,[d,b,f]=r,t=0;if(d.some((a=>0!==e[a]))){for(_ in b)__webpack_require__.o(b,_)&&(__webpack_require__.m[_]=b[_]);if(f)f(__webpack_require__)}for(a&&a(r);t<d.length;t++)c=d[t],__webpack_require__.o(e,c)&&e[c]&&e[c][0](),e[c]=0},a=self.webpackChunk_universalis_web_fr=self.webpackChunk_universalis_web_fr||[];a.forEach(webpackJsonpCallback.bind(null,0)),a.push=webpackJsonpCallback.bind(null,a.push.bind(a))})(),__webpack_require__.nc=void 0,document.addEventListener("DOMContentLoaded",(function(){__webpack_require__.e(3138).then(__webpack_require__.bind(__webpack_require__,73138)).then((function(){Promise.all([__webpack_require__.e(7686),__webpack_require__.e(786)]).then(__webpack_require__.bind(__webpack_require__,786)).then((function(e){e.default()}))}))}))})();</script> <script> let eu_article_ucode = "T322135"</script> </body> </html>