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Hauling-out - Wikipedia
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div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">"Haulout" redirects here. For the short documentary, see <a href="/wiki/Haulout_(film)" title="Haulout (film)">Haulout (film)</a>.</div><figure class="mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Sea_lion_group_at_haulout.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c3/Sea_lion_group_at_haulout.jpg/300px-Sea_lion_group_at_haulout.jpg" decoding="async" width="300" height="207" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c3/Sea_lion_group_at_haulout.jpg/450px-Sea_lion_group_at_haulout.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c3/Sea_lion_group_at_haulout.jpg/600px-Sea_lion_group_at_haulout.jpg 2x" data-file-width="1050" data-file-height="725" /></a><figcaption><a href="/wiki/Sea_lion" title="Sea lion">Sea lion</a> group at haulout</figcaption></figure> <figure class="mw-halign-right" typeof="mw:File/Thumb"><a href="/wiki/File:Harbor_seals_at_haulout.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/7b/Harbor_seals_at_haulout.jpg/250px-Harbor_seals_at_haulout.jpg" decoding="async" width="250" height="165" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/7b/Harbor_seals_at_haulout.jpg/375px-Harbor_seals_at_haulout.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/7b/Harbor_seals_at_haulout.jpg/500px-Harbor_seals_at_haulout.jpg 2x" data-file-width="1050" data-file-height="695" /></a><figcaption><a href="/wiki/Harbor_seals" class="mw-redirect" title="Harbor seals">Harbor seals</a> at haulout</figcaption></figure> <p><b>Hauling out</b> is a <a href="/wiki/Ethology" title="Ethology">behaviour</a> associated with <a href="/wiki/Pinniped" title="Pinniped">pinnipeds</a> (<a href="/wiki/True_seals" class="mw-redirect" title="True seals">true seals</a>, <a href="/wiki/Sea_lions" class="mw-redirect" title="Sea lions">sea lions</a>, <a href="/wiki/Fur_seals" class="mw-redirect" title="Fur seals">fur seals</a> and <a href="/wiki/Walruses" class="mw-redirect" title="Walruses">walruses</a>) temporarily leaving the water.<sup id="cite_ref-:Hoelzel_1-0" class="reference"><a href="#cite_note-:Hoelzel-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:A_2-0" class="reference"><a href="#cite_note-:A-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> Hauling-out typically occurs between periods of <a href="/wiki/Foraging" title="Foraging">foraging</a> activity.<sup id="cite_ref-:Hoelzel_1-1" class="reference"><a href="#cite_note-:Hoelzel-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:Kovacs_3-0" class="reference"><a href="#cite_note-:Kovacs-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:Gales_4-0" class="reference"><a href="#cite_note-:Gales-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> Rather than remain in the water, pinnipeds haul out onto land or sea ice for reasons such as <a href="/wiki/Reproduction" title="Reproduction">reproduction</a> and rest.<sup id="cite_ref-:Gales_4-1" class="reference"><a href="#cite_note-:Gales-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:A_2-1" class="reference"><a href="#cite_note-:A-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> Hauling out is necessary in seals for mating (with the exception of the <a href="/wiki/Baikal_seal" title="Baikal seal">Baikal seal</a><sup id="cite_ref-:Hoelzel_1-2" class="reference"><a href="#cite_note-:Hoelzel-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup>) and giving birth (although a distinction is generally made between reproductive aggregations, termed "<a href="/wiki/Rookeries" class="mw-redirect" title="Rookeries">rookeries</a>", and non-reproductive aggregations, termed "haul-outs").<sup id="cite_ref-:Gales_4-2" class="reference"><a href="#cite_note-:Gales-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:Loughlin_5-0" class="reference"><a href="#cite_note-:Loughlin-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> Other benefits of hauling out may include <a href="/wiki/Predator_avoidance" class="mw-redirect" title="Predator avoidance">predator avoidance</a>, <a href="/wiki/Thermoregulation" title="Thermoregulation">thermoregulation</a>, social activity, <a href="/wiki/Parasite" class="mw-redirect" title="Parasite">parasite</a> reduction and rest.<sup id="cite_ref-:Gales_4-3" class="reference"><a href="#cite_note-:Gales-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:A_2-2" class="reference"><a href="#cite_note-:A-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:Loughlin_5-1" class="reference"><a href="#cite_note-:Loughlin-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:3_6-0" class="reference"><a href="#cite_note-:3-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p><p>There is much variation in haul-out patterns among different seal <a href="/wiki/Species" title="Species">species</a>.<sup id="cite_ref-:Hoelzel_1-3" class="reference"><a href="#cite_note-:Hoelzel-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> Haul-out sites may be segregated by age and sex within the same species.<sup id="cite_ref-:Kovacs_3-1" class="reference"><a href="#cite_note-:Kovacs-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Many species of pinniped have only a few localized rookeries where they breed, but periodically occupy hundreds of haul-out sites throughout the range.<sup id="cite_ref-:Loughlin_5-2" class="reference"><a href="#cite_note-:Loughlin-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> For example, the <a href="/wiki/Australian_fur_seal" class="mw-redirect" title="Australian fur seal">Australian fur seals</a> breed on only nine islands in <a href="/wiki/Bass_Strait" title="Bass Strait">Bass Strait</a> but also occupy up to 50 haul-out sites in south-east Australian waters,<sup id="cite_ref-:Gales_4-4" class="reference"><a href="#cite_note-:Gales-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup> and <a href="/wiki/Steller_sea_lions" class="mw-redirect" title="Steller sea lions">Steller sea lions</a> have around 50 rookeries throughout their range, but several hundred haul-out sites.<sup id="cite_ref-:Loughlin_5-3" class="reference"><a href="#cite_note-:Loughlin-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup> Hauling-out behaviour provides numerous benefits to pinnipeds besides reproduction. This behaviour has been shown to be used for activities such as thermoregulation, predator avoidance, <a href="/wiki/Moulting" title="Moulting">moulting</a>, <a href="/wiki/Nursing" title="Nursing">nursing</a>, and resting.<sup id="cite_ref-:A_2-3" class="reference"><a href="#cite_note-:A-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> Haul-out frequency, duration, and site location (i.e. sea-ice, floating-ice, and terrestrial) are all influenced by physical constraints (i.e. air temperature, wind speed, and time of day) and biological constraints (i.e. moulting, age, and sex).<sup id="cite_ref-:A_2-4" class="reference"><a href="#cite_note-:A-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:1_7-0" class="reference"><a href="#cite_note-:1-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:3_6-1" class="reference"><a href="#cite_note-:3-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:4_8-0" class="reference"><a href="#cite_note-:4-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:7_9-0" class="reference"><a href="#cite_note-:7-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> Variations in hauling-out behaviour exist among pinnipeds for reasons such as geographical location.<sup id="cite_ref-:1_7-1" class="reference"><a href="#cite_note-:1-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Examples">Examples</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hauling-out&action=edit&section=1" title="Edit section: Examples"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <div class="mw-heading mw-heading3"><h3 id="Weddell_seals">Weddell seals</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hauling-out&action=edit&section=2" title="Edit section: Weddell seals"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Weddell_Seal_(js)1.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/7/75/Weddell_Seal_%28js%291.jpg/220px-Weddell_Seal_%28js%291.jpg" decoding="async" width="220" height="143" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/7/75/Weddell_Seal_%28js%291.jpg/330px-Weddell_Seal_%28js%291.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/7/75/Weddell_Seal_%28js%291.jpg/440px-Weddell_Seal_%28js%291.jpg 2x" data-file-width="5595" data-file-height="3633" /></a><figcaption><a href="/wiki/Weddell_seal" title="Weddell seal">Weddell seal</a> on terrestrial haul-out site.</figcaption></figure> <p>Haul-out sites of <a href="/wiki/Weddell_seal" title="Weddell seal">Weddell seals</a> are not necessarily geographically distinct from one another and vary due to physical factors (i.e. food availability) and biological factors (i.e. age).<sup id="cite_ref-:1_7-2" class="reference"><a href="#cite_note-:1-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> Weddell seals are high latitude <a href="/wiki/Antarctic" title="Antarctic">Antarctic</a> inhabitants, allowing them to haul-out onto ice as adults year round for foraging.<sup id="cite_ref-:1_7-3" class="reference"><a href="#cite_note-:1-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> Similar to other pinnipeds, Weddell seals haul-out for reasons such as feeding, rest, avoidance of predators, and thermoregulation.<sup id="cite_ref-:1_7-4" class="reference"><a href="#cite_note-:1-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:2_10-0" class="reference"><a href="#cite_note-:2-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> Seasonal variation has been indicated to influence the haul-out patterns of this species, environmental factors such as air temperature and wind speed trigger a shift from long-duration <a href="/wiki/Diurnal_cycle" title="Diurnal cycle">diurnal</a> haul-outs to short-duration <a href="/wiki/Nocturnal" class="mw-redirect" title="Nocturnal">nocturnal</a> patterns.<sup id="cite_ref-:1_7-5" class="reference"><a href="#cite_note-:1-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> Following moulting season the number of haul-outs performed increases allowing the seals to benefit from the increased air temperature and thus decreasing the energetic cost of growing new hair.<sup id="cite_ref-:1_7-6" class="reference"><a href="#cite_note-:1-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:2_10-1" class="reference"><a href="#cite_note-:2-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> The haul-out patterns of female Weddell seal are heavily influenced by the age of their pups.<sup id="cite_ref-:2_10-2" class="reference"><a href="#cite_note-:2-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:3_6-2" class="reference"><a href="#cite_note-:3-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> In the first week post <a href="/wiki/Parturition" class="mw-redirect" title="Parturition">parturition</a>, haul-out frequency is high and females remain hauled-out for longer periods prior to the pups starting to swim. Haul-out frequency decreases as the pups are <a href="/wiki/Wean" class="mw-redirect" title="Wean">weaned</a> and mating begins.<sup id="cite_ref-:1_7-7" class="reference"><a href="#cite_note-:1-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:2_10-3" class="reference"><a href="#cite_note-:2-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:3_6-3" class="reference"><a href="#cite_note-:3-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Walruses">Walruses</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hauling-out&action=edit&section=3" title="Edit section: Walruses"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Noaa-walrus9.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/f2/Noaa-walrus9.jpg/220px-Noaa-walrus9.jpg" decoding="async" width="220" height="150" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/f2/Noaa-walrus9.jpg/330px-Noaa-walrus9.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/f2/Noaa-walrus9.jpg/440px-Noaa-walrus9.jpg 2x" data-file-width="1784" data-file-height="1216" /></a><figcaption>Group of <a href="/wiki/Walrus" title="Walrus">walruses</a> on sea-ice haul-out.</figcaption></figure> <p><a href="/wiki/Walrus" title="Walrus">Walruses</a> tend to occupy both terrestrial and sea ice haul-out sites, alternating between the two depending on resource availability.<sup id="cite_ref-:4_8-1" class="reference"><a href="#cite_note-:4-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> Walruses haul-out onto land primarily for birthing, moulting, nursing, and resting, meanwhile using sea-ice haul-out sites for foraging and predator avoidance.<sup id="cite_ref-:4_8-2" class="reference"><a href="#cite_note-:4-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:5_11-0" class="reference"><a href="#cite_note-:5-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> These physiological factors are correlated with both the duration and frequency of haul-outs among walruses.<sup id="cite_ref-:4_8-3" class="reference"><a href="#cite_note-:4-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> Sea ice sites are more commonly used for shorter and more frequent haul-outs compared to terrestrial sites, which are commonly used to fulfill more time-consuming requirements (i.e. breeding and birthing).<sup id="cite_ref-:4_8-4" class="reference"><a href="#cite_note-:4-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:5_11-1" class="reference"><a href="#cite_note-:5-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> Hauling-out is also used as a method of thermoregulation, therefore it is influenced by various environmental factors such as wind speed, temperature, and even time of day.<sup id="cite_ref-:4_8-5" class="reference"><a href="#cite_note-:4-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> Accounting for these environmental factors, walruses more frequently haul-out from late morning to early evening and avoid hauling-out during weather periods of intense cold or high winds.<sup id="cite_ref-:4_8-6" class="reference"><a href="#cite_note-:4-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:5_11-2" class="reference"><a href="#cite_note-:5-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> Haul-out frequency is at a <a href="/wiki/Maxima_and_minima" class="mw-redirect" title="Maxima and minima">maximum</a> for walruses during the summer using terrestrial haul-out sites as sea ice sites are then further from foraging grounds.<sup id="cite_ref-:4_8-7" class="reference"><a href="#cite_note-:4-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:5_11-3" class="reference"><a href="#cite_note-:5-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> As female walruses haul-out for parturition, the males are <a href="/wiki/Territorial" class="mw-redirect" title="Territorial">territorial</a> of the haul-out site surrounding the female <a href="/wiki/Herd" title="Herd">herd</a>.<sup id="cite_ref-:5_11-4" class="reference"><a href="#cite_note-:5-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> In these instances, hauling-out provides an opportunity for more <a href="/wiki/Aggressive" class="mw-redirect" title="Aggressive">aggressive</a> and territorial males to mate.<sup id="cite_ref-:6_12-0" class="reference"><a href="#cite_note-:6-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Ringed_seal">Ringed seal</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hauling-out&action=edit&section=4" title="Edit section: Ringed seal"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Pusa_hispida_hispida_NOAA_1.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/3/33/Pusa_hispida_hispida_NOAA_1.jpg/220px-Pusa_hispida_hispida_NOAA_1.jpg" decoding="async" width="220" height="146" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/3/33/Pusa_hispida_hispida_NOAA_1.jpg/330px-Pusa_hispida_hispida_NOAA_1.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/3/33/Pusa_hispida_hispida_NOAA_1.jpg/440px-Pusa_hispida_hispida_NOAA_1.jpg 2x" data-file-width="700" data-file-height="465" /></a><figcaption><a href="/wiki/Ringed_seal" title="Ringed seal">Ringed seal</a> on sea-ice haul-out.</figcaption></figure> <p><a href="/wiki/Ringed_seal" title="Ringed seal">Ringed seal</a> hauling-out occurs throughout any point in the year, however it reaches a maximum during the spring.<sup id="cite_ref-:7_9-1" class="reference"><a href="#cite_note-:7-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> In comparison to other pinniped species, ringed seals haul-out with a shorter duration year round.<sup id="cite_ref-:7_9-2" class="reference"><a href="#cite_note-:7-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> Ringed seals have a <a href="/wiki/Diel" class="mw-redirect" title="Diel">diel</a> haul-out pattern in which they spend more time hauled-out during the night, an uncommon feature among <a href="/wiki/Pinniped" title="Pinniped">pinnipeds</a>.<sup id="cite_ref-:7_9-3" class="reference"><a href="#cite_note-:7-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:10_13-0" class="reference"><a href="#cite_note-:10-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> Hauling-out spikes an increase in the herding behaviour of ringed seals, particularly in the Ladoga subspecies.<sup id="cite_ref-:8_14-0" class="reference"><a href="#cite_note-:8-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> Subspecies of the ringed seal prefer different haul-out sites depending on their geographical location and environmental constraints.<sup id="cite_ref-:9_15-0" class="reference"><a href="#cite_note-:9-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> For example, 5 subspecies of ringed seals prefer hauling-out onto land-fast ice, however <i>Phoca hispida ochotensis</i> prefers drifting pack ice, meanwhile <i>Phoca hispida hispida</i> occupies both land-fast ice and far offshore areas of relatively stable ice.<sup id="cite_ref-:9_15-1" class="reference"><a href="#cite_note-:9-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> The majority of ringed seals however use terrestrial haul-out sites to create birth layers in the snow for newborn seal pups.<sup id="cite_ref-:9_15-2" class="reference"><a href="#cite_note-:9-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Harbour_seal">Harbour seal</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hauling-out&action=edit&section=5" title="Edit section: Harbour seal"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Harbor_seal_rocks.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/c8/Harbor_seal_rocks.jpg/220px-Harbor_seal_rocks.jpg" decoding="async" width="220" height="181" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/c8/Harbor_seal_rocks.jpg/330px-Harbor_seal_rocks.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/c8/Harbor_seal_rocks.jpg/440px-Harbor_seal_rocks.jpg 2x" data-file-width="2747" data-file-height="2263" /></a><figcaption><a href="/wiki/Harbour_seal" class="mw-redirect" title="Harbour seal">Harbour seals</a> on <a href="/wiki/Intertidal" class="mw-redirect" title="Intertidal">intertidal</a> site</figcaption></figure> <p><a href="/wiki/Harbour_seal" class="mw-redirect" title="Harbour seal">Harbour seals</a> are the most abundant pinniped in the eastern <a href="/wiki/North_Pacific_Ocean" class="mw-redirect" title="North Pacific Ocean">North Pacific Ocean</a>.<sup id="cite_ref-:A_2-5" class="reference"><a href="#cite_note-:A-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> Much like other pinnipeds, harbour seals haul-out for reasons such as thermoregulation, breeding, mating, moulting, resting, and foraging.<sup id="cite_ref-:11_16-0" class="reference"><a href="#cite_note-:11-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:A_2-6" class="reference"><a href="#cite_note-:A-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> They commonly haul-out onto <a href="/wiki/Intertidal" class="mw-redirect" title="Intertidal">intertidal</a> ledges, <a href="/wiki/Mudflat" title="Mudflat">mudflats</a>, <a href="/wiki/Beaches" class="mw-redirect" title="Beaches">beaches</a>, and ice floes year round.<sup id="cite_ref-:11_16-1" class="reference"><a href="#cite_note-:11-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> Haul-out sites are often revisited on a regular basis by the same herd<sup id="cite_ref-:A_2-7" class="reference"><a href="#cite_note-:A-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> and are heavily affected by tide height.<sup id="cite_ref-:A_2-8" class="reference"><a href="#cite_note-:A-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:11_16-2" class="reference"><a href="#cite_note-:11-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> Harbour seals are likely to move haul-out sites in response to inclement weather conditions (i.e. wind chill and wave size) to more favourable sites in rocky reefs, mudflats, and beaches that are exposed during lower tides.<sup id="cite_ref-:A_2-9" class="reference"><a href="#cite_note-:A-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:11_16-3" class="reference"><a href="#cite_note-:11-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> Frequency and duration of the behaviour is at a maxima during early afternoon when lower tides and higher air temperatures are prevalent.<sup id="cite_ref-:A_2-10" class="reference"><a href="#cite_note-:A-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:11_16-4" class="reference"><a href="#cite_note-:11-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> During parturition and weaning, females spend more time hauled-out ashore until their pups begin to swim, meanwhile males spend less time hauled-out and maintain aquatic <a href="/wiki/Territories" class="mw-redirect" title="Territories">territories</a> instead.<sup id="cite_ref-:A_2-11" class="reference"><a href="#cite_note-:A-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> Moulting and predation risk also increase the time spent hauled-out.<sup id="cite_ref-:A_2-12" class="reference"><a href="#cite_note-:A-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> Despite the increased time ashore for females and decreased time ashore or males during birthing and weaning, biological constraints such as age and sex have not been shown to effect harbour seal haul-outs.<sup id="cite_ref-:A_2-13" class="reference"><a href="#cite_note-:A-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> Both male and female harbour seals of all ages are consistent with time spent hauled-out.<sup id="cite_ref-:A_2-14" class="reference"><a href="#cite_note-:A-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-:11_16-5" class="reference"><a href="#cite_note-:11-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> Harbour seals commonly inhabit regions susceptible to human <a href="/wiki/Disturbance_(ecology)" title="Disturbance (ecology)">disturbances</a> (i.e. industries such as the <a href="/wiki/Fishery" title="Fishery">fishery</a>), a factor that has been studied and shown to alter haul-out patterns.<sup id="cite_ref-:12_17-0" class="reference"><a href="#cite_note-:12-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> Human disturbances negatively influence the duration and frequency of harbour seal haul-outs, decreasing the occurrence of this behaviour as human interference increases.<sup id="cite_ref-:12_17-1" class="reference"><a href="#cite_note-:12-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Hauling-out&action=edit&section=6" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-:Hoelzel-1"><span class="mw-cite-backlink">^ <a href="#cite_ref-:Hoelzel_1-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-:Hoelzel_1-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-:Hoelzel_1-2"><sup><i><b>c</b></i></sup></a> <a href="#cite_ref-:Hoelzel_1-3"><sup><i><b>d</b></i></sup></a></span> <span class="reference-text">Hoelzel, A. Rus. (2002). <i>Marine Mammal Biology: An Evolutionary Approach</i>. 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