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vector-toc-level-2"> <a class="vector-toc-link" href="#Recombination"> <div class="vector-toc-text"> <span class="vector-toc-numb">2.1</span> <span>Recombination</span> </div> </a> <ul id="toc-Recombination-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Transmission" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Transmission"> <div class="vector-toc-text"> <span class="vector-toc-numb">3</span> <span>Transmission</span> </div> </a> <ul id="toc-Transmission-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Provirus" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Provirus"> <div class="vector-toc-text"> <span class="vector-toc-numb">4</span> <span>Provirus</span> </div> </a> <ul id="toc-Provirus-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Early_evolution" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Early_evolution"> <div class="vector-toc-text"> <span class="vector-toc-numb">5</span> <span>Early evolution</span> </div> </a> <ul id="toc-Early_evolution-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Gene_therapy" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Gene_therapy"> <div class="vector-toc-text"> <span class="vector-toc-numb">6</span> <span>Gene therapy</span> </div> </a> <ul id="toc-Gene_therapy-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Cancer" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Cancer"> <div class="vector-toc-text"> <span class="vector-toc-numb">7</span> <span>Cancer</span> </div> </a> <ul id="toc-Cancer-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Classification" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Classification"> <div class="vector-toc-text"> <span class="vector-toc-numb">8</span> <span>Classification</span> </div> </a> <button aria-controls="toc-Classification-sublist" class="cdx-button cdx-button--weight-quiet cdx-button--icon-only vector-toc-toggle"> <span class="vector-icon mw-ui-icon-wikimedia-expand"></span> <span>Toggle Classification subsection</span> </button> <ul id="toc-Classification-sublist" class="vector-toc-list"> <li id="toc-Exogenous" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Exogenous"> <div class="vector-toc-text"> <span class="vector-toc-numb">8.1</span> <span>Exogenous</span> </div> </a> <ul id="toc-Exogenous-sublist" class="vector-toc-list"> <li id="toc-Group_VI_viruses" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Group_VI_viruses"> <div class="vector-toc-text"> <span class="vector-toc-numb">8.1.1</span> <span>Group VI viruses</span> </div> </a> <ul id="toc-Group_VI_viruses-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Group_VII_viruses" class="vector-toc-list-item vector-toc-level-3"> <a class="vector-toc-link" href="#Group_VII_viruses"> <div class="vector-toc-text"> <span class="vector-toc-numb">8.1.2</span> <span>Group VII viruses</span> </div> </a> <ul id="toc-Group_VII_viruses-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Endogenous" class="vector-toc-list-item vector-toc-level-2"> <a class="vector-toc-link" href="#Endogenous"> <div class="vector-toc-text"> <span class="vector-toc-numb">8.2</span> <span>Endogenous</span> </div> </a> <ul id="toc-Endogenous-sublist" class="vector-toc-list"> </ul> </li> </ul> </li> <li id="toc-Controversy" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Controversy"> <div class="vector-toc-text"> <span class="vector-toc-numb">9</span> <span>Controversy</span> </div> </a> <ul id="toc-Controversy-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Treatment" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Treatment"> <div class="vector-toc-text"> <span class="vector-toc-numb">10</span> <span>Treatment</span> </div> </a> <ul id="toc-Treatment-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Treatment_of_veterinary_retroviruses" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Treatment_of_veterinary_retroviruses"> <div class="vector-toc-text"> <span class="vector-toc-numb">11</span> <span>Treatment of veterinary retroviruses</span> </div> </a> <ul id="toc-Treatment_of_veterinary_retroviruses-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-References" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#References"> <div class="vector-toc-text"> <span class="vector-toc-numb">12</span> <span>References</span> </div> </a> <ul id="toc-References-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-Further_reading" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#Further_reading"> <div class="vector-toc-text"> <span class="vector-toc-numb">13</span> <span>Further reading</span> </div> </a> <ul id="toc-Further_reading-sublist" class="vector-toc-list"> </ul> </li> <li id="toc-External_links" class="vector-toc-list-item vector-toc-level-1 vector-toc-list-item-expanded"> <a class="vector-toc-link" href="#External_links"> <div class="vector-toc-text"> <span class="vector-toc-numb">14</span> <span>External links</span> </div> </a> <ul id="toc-External_links-sublist" class="vector-toc-list"> </ul> </li> </ul> </div> </div> </nav> </div> </div> <div class="mw-content-container"> <main id="content" class="mw-body"> <header class="mw-body-header vector-page-titlebar"> <nav aria-label="Contents" class="vector-toc-landmark"> <div id="vector-page-titlebar-toc" class="vector-dropdown vector-page-titlebar-toc vector-button-flush-left" > <input type="checkbox" id="vector-page-titlebar-toc-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-vector-page-titlebar-toc" class="vector-dropdown-checkbox " aria-label="Toggle the table of contents" > <label id="vector-page-titlebar-toc-label" for="vector-page-titlebar-toc-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--icon-only " aria-hidden="true" ><span class="vector-icon mw-ui-icon-listBullet mw-ui-icon-wikimedia-listBullet"></span> <span class="vector-dropdown-label-text">Toggle the table of contents</span> </label> <div class="vector-dropdown-content"> <div id="vector-page-titlebar-toc-unpinned-container" class="vector-unpinned-container"> </div> </div> </div> </nav> <h1 id="firstHeading" class="firstHeading mw-first-heading"><span class="mw-page-title-main">Retrovirus</span></h1> <div id="p-lang-btn" class="vector-dropdown mw-portlet mw-portlet-lang" > <input type="checkbox" id="p-lang-btn-checkbox" role="button" aria-haspopup="true" data-event-name="ui.dropdown-p-lang-btn" class="vector-dropdown-checkbox mw-interlanguage-selector" aria-label="Go to an article in another language. Available in 61 languages" > <label id="p-lang-btn-label" for="p-lang-btn-checkbox" class="vector-dropdown-label cdx-button cdx-button--fake-button cdx-button--fake-button--enabled cdx-button--weight-quiet cdx-button--action-progressive mw-portlet-lang-heading-61" aria-hidden="true" ><span class="vector-icon mw-ui-icon-language-progressive mw-ui-icon-wikimedia-language-progressive"></span> <span class="vector-dropdown-label-text">61 languages</span> </label> <div class="vector-dropdown-content"> <div class="vector-menu-content"> <ul class="vector-menu-content-list"> <li class="interlanguage-link interwiki-af mw-list-item"><a href="https://af.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Afrikaans" lang="af" hreflang="af" data-title="Retrovirus" data-language-autonym="Afrikaans" data-language-local-name="Afrikaans" class="interlanguage-link-target"><span>Afrikaans</span></a></li><li class="interlanguage-link interwiki-ar mw-list-item"><a href="https://ar.wikipedia.org/wiki/%D9%81%D9%8A%D8%B1%D9%88%D8%B3_%D8%B1%D8%A7%D8%AC%D8%B9" title="فيروس راجع – Arabic" lang="ar" hreflang="ar" data-title="فيروس راجع" data-language-autonym="العربية" data-language-local-name="Arabic" class="interlanguage-link-target"><span>العربية</span></a></li><li class="interlanguage-link interwiki-be mw-list-item"><a href="https://be.wikipedia.org/wiki/%D0%A0%D1%8D%D1%82%D1%80%D0%B0%D0%B2%D1%96%D1%80%D1%83%D1%81%D1%8B" title="Рэтравірусы – Belarusian" lang="be" hreflang="be" data-title="Рэтравірусы" data-language-autonym="Беларуская" data-language-local-name="Belarusian" class="interlanguage-link-target"><span>Беларуская</span></a></li><li class="interlanguage-link interwiki-bg mw-list-item"><a href="https://bg.wikipedia.org/wiki/%D0%A0%D0%B5%D1%82%D1%80%D0%BE%D0%B2%D0%B8%D1%80%D1%83%D1%81%D0%B8" title="Ретровируси – Bulgarian" lang="bg" hreflang="bg" data-title="Ретровируси" data-language-autonym="Български" data-language-local-name="Bulgarian" class="interlanguage-link-target"><span>Български</span></a></li><li class="interlanguage-link interwiki-bs mw-list-item"><a href="https://bs.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Bosnian" lang="bs" hreflang="bs" data-title="Retrovirus" data-language-autonym="Bosanski" data-language-local-name="Bosnian" class="interlanguage-link-target"><span>Bosanski</span></a></li><li class="interlanguage-link interwiki-ca mw-list-item"><a href="https://ca.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Catalan" lang="ca" hreflang="ca" data-title="Retrovirus" data-language-autonym="Català" data-language-local-name="Catalan" class="interlanguage-link-target"><span>Català</span></a></li><li class="interlanguage-link interwiki-cs mw-list-item"><a href="https://cs.wikipedia.org/wiki/Retroviry" title="Retroviry – Czech" lang="cs" hreflang="cs" data-title="Retroviry" data-language-autonym="Čeština" data-language-local-name="Czech" class="interlanguage-link-target"><span>Čeština</span></a></li><li class="interlanguage-link interwiki-da mw-list-item"><a href="https://da.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Danish" lang="da" hreflang="da" data-title="Retrovirus" data-language-autonym="Dansk" data-language-local-name="Danish" class="interlanguage-link-target"><span>Dansk</span></a></li><li class="interlanguage-link interwiki-de mw-list-item"><a href="https://de.wikipedia.org/wiki/Retroviren" title="Retroviren – German" lang="de" hreflang="de" data-title="Retroviren" data-language-autonym="Deutsch" data-language-local-name="German" class="interlanguage-link-target"><span>Deutsch</span></a></li><li class="interlanguage-link interwiki-et mw-list-item"><a href="https://et.wikipedia.org/wiki/Retroviirused" title="Retroviirused – Estonian" lang="et" hreflang="et" data-title="Retroviirused" data-language-autonym="Eesti" data-language-local-name="Estonian" class="interlanguage-link-target"><span>Eesti</span></a></li><li class="interlanguage-link interwiki-el mw-list-item"><a href="https://el.wikipedia.org/wiki/%CE%A1%CE%B5%CF%84%CF%81%CE%BF%CF%8A%CF%8C%CF%82" title="Ρετροϊός – Greek" lang="el" hreflang="el" data-title="Ρετροϊός" data-language-autonym="Ελληνικά" data-language-local-name="Greek" class="interlanguage-link-target"><span>Ελληνικά</span></a></li><li class="interlanguage-link interwiki-es mw-list-item"><a href="https://es.wikipedia.org/wiki/Retroviridae" title="Retroviridae – Spanish" lang="es" hreflang="es" data-title="Retroviridae" data-language-autonym="Español" data-language-local-name="Spanish" class="interlanguage-link-target"><span>Español</span></a></li><li class="interlanguage-link interwiki-eu mw-list-item"><a href="https://eu.wikipedia.org/wiki/Erretrobirus" title="Erretrobirus – Basque" lang="eu" hreflang="eu" data-title="Erretrobirus" data-language-autonym="Euskara" data-language-local-name="Basque" class="interlanguage-link-target"><span>Euskara</span></a></li><li class="interlanguage-link interwiki-fa mw-list-item"><a href="https://fa.wikipedia.org/wiki/%D8%B1%D8%AA%D8%B1%D9%88%D9%88%DB%8C%D8%B1%D9%88%D8%B3" title="رتروویروس – Persian" lang="fa" hreflang="fa" data-title="رتروویروس" data-language-autonym="فارسی" data-language-local-name="Persian" class="interlanguage-link-target"><span>فارسی</span></a></li><li class="interlanguage-link interwiki-fr mw-list-item"><a href="https://fr.wikipedia.org/wiki/Retroviridae" title="Retroviridae – French" lang="fr" hreflang="fr" data-title="Retroviridae" data-language-autonym="Français" data-language-local-name="French" class="interlanguage-link-target"><span>Français</span></a></li><li class="interlanguage-link interwiki-ga mw-list-item"><a href="https://ga.wikipedia.org/wiki/Aisv%C3%ADreas" title="Aisvíreas – Irish" lang="ga" hreflang="ga" data-title="Aisvíreas" data-language-autonym="Gaeilge" data-language-local-name="Irish" class="interlanguage-link-target"><span>Gaeilge</span></a></li><li class="interlanguage-link interwiki-gl mw-list-item"><a href="https://gl.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Galician" lang="gl" hreflang="gl" data-title="Retrovirus" data-language-autonym="Galego" data-language-local-name="Galician" class="interlanguage-link-target"><span>Galego</span></a></li><li class="interlanguage-link interwiki-ko mw-list-item"><a href="https://ko.wikipedia.org/wiki/%EB%A0%88%ED%8A%B8%EB%A1%9C%EB%B0%94%EC%9D%B4%EB%9F%AC%EC%8A%A4" title="레트로바이러스 – Korean" lang="ko" hreflang="ko" data-title="레트로바이러스" data-language-autonym="한국어" data-language-local-name="Korean" class="interlanguage-link-target"><span>한국어</span></a></li><li class="interlanguage-link interwiki-hy mw-list-item"><a href="https://hy.wikipedia.org/wiki/%D5%8C%D5%A5%D5%BF%D6%80%D5%B8%D5%BE%D5%AB%D6%80%D5%B8%D6%82%D5%BD%D5%B6%D5%A5%D6%80" title="Ռետրովիրուսներ – Armenian" lang="hy" hreflang="hy" data-title="Ռետրովիրուսներ" data-language-autonym="Հայերեն" data-language-local-name="Armenian" class="interlanguage-link-target"><span>Հայերեն</span></a></li><li class="interlanguage-link interwiki-hi mw-list-item"><a href="https://hi.wikipedia.org/wiki/%E0%A4%B0%E0%A5%87%E0%A4%9F%E0%A5%8D%E0%A4%B0%E0%A5%8B%E0%A4%B5%E0%A4%BE%E0%A4%AF%E0%A4%B0%E0%A4%B8" title="रेट्रोवायरस – Hindi" lang="hi" hreflang="hi" data-title="रेट्रोवायरस" data-language-autonym="हिन्दी" data-language-local-name="Hindi" class="interlanguage-link-target"><span>हिन्दी</span></a></li><li class="interlanguage-link interwiki-io mw-list-item"><a href="https://io.wikipedia.org/wiki/Retroviruso" title="Retroviruso – Ido" lang="io" hreflang="io" data-title="Retroviruso" data-language-autonym="Ido" data-language-local-name="Ido" class="interlanguage-link-target"><span>Ido</span></a></li><li class="interlanguage-link interwiki-id mw-list-item"><a href="https://id.wikipedia.org/wiki/Retroviridae" title="Retroviridae – Indonesian" lang="id" hreflang="id" data-title="Retroviridae" data-language-autonym="Bahasa Indonesia" data-language-local-name="Indonesian" class="interlanguage-link-target"><span>Bahasa Indonesia</span></a></li><li class="interlanguage-link interwiki-it mw-list-item"><a href="https://it.wikipedia.org/wiki/Retroviridae" title="Retroviridae – Italian" lang="it" hreflang="it" data-title="Retroviridae" data-language-autonym="Italiano" data-language-local-name="Italian" class="interlanguage-link-target"><span>Italiano</span></a></li><li class="interlanguage-link interwiki-he mw-list-item"><a href="https://he.wikipedia.org/wiki/%D7%A8%D7%98%D7%A8%D7%95-%D7%95%D7%99%D7%A8%D7%95%D7%A1" title="רטרו-וירוס – Hebrew" lang="he" hreflang="he" data-title="רטרו-וירוס" data-language-autonym="עברית" data-language-local-name="Hebrew" class="interlanguage-link-target"><span>עברית</span></a></li><li class="interlanguage-link interwiki-ka mw-list-item"><a href="https://ka.wikipedia.org/wiki/%E1%83%A0%E1%83%94%E1%83%A2%E1%83%A0%E1%83%9D%E1%83%95%E1%83%98%E1%83%A0%E1%83%A3%E1%83%A1%E1%83%94%E1%83%91%E1%83%98" title="რეტროვირუსები – Georgian" lang="ka" hreflang="ka" data-title="რეტროვირუსები" data-language-autonym="ქართული" data-language-local-name="Georgian" class="interlanguage-link-target"><span>ქართული</span></a></li><li class="interlanguage-link interwiki-kk mw-list-item"><a href="https://kk.wikipedia.org/wiki/%D0%A0%D0%B5%D1%82%D1%80%D0%BE%D0%B2%D0%B8%D1%80%D1%83%D1%81%D1%82%D0%B0%D1%80" title="Ретровирустар – Kazakh" lang="kk" hreflang="kk" data-title="Ретровирустар" data-language-autonym="Қазақша" data-language-local-name="Kazakh" class="interlanguage-link-target"><span>Қазақша</span></a></li><li class="interlanguage-link interwiki-la mw-list-item"><a href="https://la.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Latin" lang="la" hreflang="la" data-title="Retrovirus" data-language-autonym="Latina" data-language-local-name="Latin" class="interlanguage-link-target"><span>Latina</span></a></li><li class="interlanguage-link interwiki-lv mw-list-item"><a href="https://lv.wikipedia.org/wiki/Retrov%C4%ABrusi" title="Retrovīrusi – Latvian" lang="lv" hreflang="lv" data-title="Retrovīrusi" data-language-autonym="Latviešu" data-language-local-name="Latvian" class="interlanguage-link-target"><span>Latviešu</span></a></li><li class="interlanguage-link interwiki-lg mw-list-item"><a href="https://lg.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Ganda" lang="lg" hreflang="lg" data-title="Retrovirus" data-language-autonym="Luganda" data-language-local-name="Ganda" class="interlanguage-link-target"><span>Luganda</span></a></li><li class="interlanguage-link interwiki-lmo mw-list-item"><a href="https://lmo.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Lombard" lang="lmo" hreflang="lmo" data-title="Retrovirus" data-language-autonym="Lombard" data-language-local-name="Lombard" class="interlanguage-link-target"><span>Lombard</span></a></li><li class="interlanguage-link interwiki-hu mw-list-item"><a href="https://hu.wikipedia.org/wiki/Retrov%C3%ADrus" title="Retrovírus – Hungarian" lang="hu" hreflang="hu" data-title="Retrovírus" data-language-autonym="Magyar" data-language-local-name="Hungarian" class="interlanguage-link-target"><span>Magyar</span></a></li><li class="interlanguage-link interwiki-mk mw-list-item"><a href="https://mk.wikipedia.org/wiki/%D0%A0%D0%B5%D1%82%D1%80%D0%BE%D0%B2%D0%B8%D1%80%D1%83%D1%81" title="Ретровирус – Macedonian" lang="mk" hreflang="mk" data-title="Ретровирус" data-language-autonym="Македонски" data-language-local-name="Macedonian" class="interlanguage-link-target"><span>Македонски</span></a></li><li class="interlanguage-link interwiki-arz mw-list-item"><a href="https://arz.wikipedia.org/wiki/%D9%81%D9%8A%D8%B1%D9%88%D8%B3_%D8%B1%D8%A7%D8%AC%D8%B9" title="فيروس راجع – Egyptian Arabic" lang="arz" hreflang="arz" data-title="فيروس راجع" data-language-autonym="مصرى" data-language-local-name="Egyptian Arabic" class="interlanguage-link-target"><span>مصرى</span></a></li><li class="interlanguage-link interwiki-ms mw-list-item"><a href="https://ms.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Malay" lang="ms" hreflang="ms" data-title="Retrovirus" data-language-autonym="Bahasa Melayu" data-language-local-name="Malay" class="interlanguage-link-target"><span>Bahasa Melayu</span></a></li><li class="interlanguage-link interwiki-mn mw-list-item"><a href="https://mn.wikipedia.org/wiki/%D0%A0%D0%B5%D1%82%D1%80%D0%BE%D0%B2%D0%B8%D1%80%D1%83%D1%81" title="Ретровирус – Mongolian" lang="mn" hreflang="mn" data-title="Ретровирус" data-language-autonym="Монгол" data-language-local-name="Mongolian" class="interlanguage-link-target"><span>Монгол</span></a></li><li class="interlanguage-link interwiki-nl mw-list-item"><a href="https://nl.wikipedia.org/wiki/Retrovirussen" title="Retrovirussen – Dutch" lang="nl" hreflang="nl" data-title="Retrovirussen" data-language-autonym="Nederlands" data-language-local-name="Dutch" class="interlanguage-link-target"><span>Nederlands</span></a></li><li class="interlanguage-link interwiki-ja mw-list-item"><a href="https://ja.wikipedia.org/wiki/%E3%83%AC%E3%83%88%E3%83%AD%E3%82%A6%E3%82%A4%E3%83%AB%E3%82%B9%E7%A7%91" title="レトロウイルス科 – Japanese" lang="ja" hreflang="ja" data-title="レトロウイルス科" data-language-autonym="日本語" data-language-local-name="Japanese" class="interlanguage-link-target"><span>日本語</span></a></li><li class="interlanguage-link interwiki-frr mw-list-item"><a href="https://frr.wikipedia.org/wiki/Retroviridae" title="Retroviridae – Northern Frisian" lang="frr" hreflang="frr" data-title="Retroviridae" data-language-autonym="Nordfriisk" data-language-local-name="Northern Frisian" class="interlanguage-link-target"><span>Nordfriisk</span></a></li><li class="interlanguage-link interwiki-no mw-list-item"><a href="https://no.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Norwegian Bokmål" lang="nb" hreflang="nb" data-title="Retrovirus" data-language-autonym="Norsk bokmål" data-language-local-name="Norwegian Bokmål" class="interlanguage-link-target"><span>Norsk bokmål</span></a></li><li class="interlanguage-link interwiki-nn mw-list-item"><a href="https://nn.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Norwegian Nynorsk" lang="nn" hreflang="nn" data-title="Retrovirus" data-language-autonym="Norsk nynorsk" data-language-local-name="Norwegian Nynorsk" class="interlanguage-link-target"><span>Norsk nynorsk</span></a></li><li class="interlanguage-link interwiki-oc mw-list-item"><a href="https://oc.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Occitan" lang="oc" hreflang="oc" data-title="Retrovirus" data-language-autonym="Occitan" data-language-local-name="Occitan" class="interlanguage-link-target"><span>Occitan</span></a></li><li class="interlanguage-link interwiki-uz mw-list-item"><a href="https://uz.wikipedia.org/wiki/Retroviruslar" title="Retroviruslar – Uzbek" lang="uz" hreflang="uz" data-title="Retroviruslar" data-language-autonym="Oʻzbekcha / ўзбекча" data-language-local-name="Uzbek" class="interlanguage-link-target"><span>Oʻzbekcha / ўзбекча</span></a></li><li class="interlanguage-link interwiki-pl mw-list-item"><a href="https://pl.wikipedia.org/wiki/Retrowirusy" title="Retrowirusy – Polish" lang="pl" hreflang="pl" data-title="Retrowirusy" data-language-autonym="Polski" data-language-local-name="Polish" class="interlanguage-link-target"><span>Polski</span></a></li><li class="interlanguage-link interwiki-pt mw-list-item"><a href="https://pt.wikipedia.org/wiki/Retroviridae" title="Retroviridae – Portuguese" lang="pt" hreflang="pt" data-title="Retroviridae" data-language-autonym="Português" data-language-local-name="Portuguese" class="interlanguage-link-target"><span>Português</span></a></li><li class="interlanguage-link interwiki-ro mw-list-item"><a href="https://ro.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Romanian" lang="ro" hreflang="ro" data-title="Retrovirus" data-language-autonym="Română" data-language-local-name="Romanian" class="interlanguage-link-target"><span>Română</span></a></li><li class="interlanguage-link interwiki-ru mw-list-item"><a href="https://ru.wikipedia.org/wiki/%D0%A0%D0%B5%D1%82%D1%80%D0%BE%D0%B2%D0%B8%D1%80%D1%83%D1%81%D1%8B" title="Ретровирусы – Russian" lang="ru" hreflang="ru" data-title="Ретровирусы" data-language-autonym="Русский" data-language-local-name="Russian" class="interlanguage-link-target"><span>Русский</span></a></li><li class="interlanguage-link interwiki-sco mw-list-item"><a href="https://sco.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Scots" lang="sco" hreflang="sco" data-title="Retrovirus" data-language-autonym="Scots" data-language-local-name="Scots" class="interlanguage-link-target"><span>Scots</span></a></li><li class="interlanguage-link interwiki-simple mw-list-item"><a href="https://simple.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Simple English" lang="en-simple" hreflang="en-simple" data-title="Retrovirus" data-language-autonym="Simple English" data-language-local-name="Simple English" class="interlanguage-link-target"><span>Simple English</span></a></li><li class="interlanguage-link interwiki-sl mw-list-item"><a href="https://sl.wikipedia.org/wiki/Retrovirusi" title="Retrovirusi – Slovenian" lang="sl" hreflang="sl" data-title="Retrovirusi" data-language-autonym="Slovenščina" data-language-local-name="Slovenian" class="interlanguage-link-target"><span>Slovenščina</span></a></li><li class="interlanguage-link interwiki-sr mw-list-item"><a href="https://sr.wikipedia.org/wiki/%D0%A0%D0%B5%D1%82%D1%80%D0%BE%D0%B2%D0%B8%D1%80%D1%83%D1%81%D0%B8" title="Ретровируси – Serbian" lang="sr" hreflang="sr" data-title="Ретровируси" data-language-autonym="Српски / srpski" data-language-local-name="Serbian" class="interlanguage-link-target"><span>Српски / srpski</span></a></li><li class="interlanguage-link interwiki-sh mw-list-item"><a href="https://sh.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Serbo-Croatian" lang="sh" hreflang="sh" data-title="Retrovirus" data-language-autonym="Srpskohrvatski / српскохрватски" data-language-local-name="Serbo-Croatian" class="interlanguage-link-target"><span>Srpskohrvatski / српскохрватски</span></a></li><li class="interlanguage-link interwiki-fi mw-list-item"><a href="https://fi.wikipedia.org/wiki/Retrovirukset" title="Retrovirukset – Finnish" lang="fi" hreflang="fi" data-title="Retrovirukset" data-language-autonym="Suomi" data-language-local-name="Finnish" class="interlanguage-link-target"><span>Suomi</span></a></li><li class="interlanguage-link interwiki-sv mw-list-item"><a href="https://sv.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Swedish" lang="sv" hreflang="sv" data-title="Retrovirus" data-language-autonym="Svenska" data-language-local-name="Swedish" class="interlanguage-link-target"><span>Svenska</span></a></li><li class="interlanguage-link interwiki-ta mw-list-item"><a href="https://ta.wikipedia.org/wiki/%E0%AE%B0%E0%AF%86%E0%AE%9F%E0%AF%8D%E0%AE%B0%E0%AF%8B_%E0%AE%B5%E0%AF%88%E0%AE%B0%E0%AE%B8%E0%AF%8D" title="ரெட்ரோ வைரஸ் – Tamil" lang="ta" hreflang="ta" data-title="ரெட்ரோ வைரஸ்" data-language-autonym="தமிழ்" data-language-local-name="Tamil" class="interlanguage-link-target"><span>தமிழ்</span></a></li><li class="interlanguage-link interwiki-tr mw-list-item"><a href="https://tr.wikipedia.org/wiki/Retrovir%C3%BCs" title="Retrovirüs – Turkish" lang="tr" hreflang="tr" data-title="Retrovirüs" data-language-autonym="Türkçe" data-language-local-name="Turkish" class="interlanguage-link-target"><span>Türkçe</span></a></li><li class="interlanguage-link interwiki-uk mw-list-item"><a href="https://uk.wikipedia.org/wiki/%D0%A0%D0%B5%D1%82%D1%80%D0%BE%D0%B2%D1%96%D1%80%D1%83%D1%81%D0%B8" title="Ретровіруси – Ukrainian" lang="uk" hreflang="uk" data-title="Ретровіруси" data-language-autonym="Українська" data-language-local-name="Ukrainian" class="interlanguage-link-target"><span>Українська</span></a></li><li class="interlanguage-link interwiki-ur mw-list-item"><a href="https://ur.wikipedia.org/wiki/%D8%B1%DB%8C%D9%B9%D8%B1%D9%88_%D9%88%D8%A7%D8%A6%D8%B1%D8%B3" title="ریٹرو وائرس – Urdu" lang="ur" hreflang="ur" data-title="ریٹرو وائرس" data-language-autonym="اردو" data-language-local-name="Urdu" class="interlanguage-link-target"><span>اردو</span></a></li><li class="interlanguage-link interwiki-vi mw-list-item"><a href="https://vi.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Vietnamese" lang="vi" hreflang="vi" data-title="Retrovirus" data-language-autonym="Tiếng Việt" data-language-local-name="Vietnamese" class="interlanguage-link-target"><span>Tiếng Việt</span></a></li><li class="interlanguage-link interwiki-war mw-list-item"><a href="https://war.wikipedia.org/wiki/Retrovirus" title="Retrovirus – Waray" lang="war" hreflang="war" data-title="Retrovirus" data-language-autonym="Winaray" data-language-local-name="Waray" class="interlanguage-link-target"><span>Winaray</span></a></li><li class="interlanguage-link interwiki-zh-yue mw-list-item"><a href="https://zh-yue.wikipedia.org/wiki/%E9%80%86%E8%BD%89%E9%8C%84%E7%97%85%E6%AF%92" title="逆轉錄病毒 – Cantonese" lang="yue" hreflang="yue" data-title="逆轉錄病毒" data-language-autonym="粵語" data-language-local-name="Cantonese" class="interlanguage-link-target"><span>粵語</span></a></li><li class="interlanguage-link interwiki-zh mw-list-item"><a href="https://zh.wikipedia.org/wiki/%E9%80%86%E8%BD%AC%E5%BD%95%E7%97%85%E6%AF%92" title="逆转录病毒 – Chinese" lang="zh" hreflang="zh" data-title="逆转录病毒" data-language-autonym="中文" data-language-local-name="Chinese" class="interlanguage-link-target"><span>中文</span></a></li> </ul> <div class="after-portlet after-portlet-lang"><span class="wb-langlinks-edit wb-langlinks-link"><a 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id="siteSub" class="noprint">From Wikipedia, the free encyclopedia</div> </div> <div id="contentSub"><div id="mw-content-subtitle"></div></div> <div id="mw-content-text" class="mw-body-content"><div class="mw-content-ltr mw-parser-output" lang="en" dir="ltr"><div class="shortdescription nomobile noexcerpt noprint searchaux" style="display:none">Family of viruses</div> <style data-mw-deduplicate="TemplateStyles:r1236090951">.mw-parser-output .hatnote{font-style:italic}.mw-parser-output div.hatnote{padding-left:1.6em;margin-bottom:0.5em}.mw-parser-output .hatnote i{font-style:normal}.mw-parser-output .hatnote+link+.hatnote{margin-top:-0.5em}@media print{body.ns-0 .mw-parser-output .hatnote{display:none!important}}</style><div role="note" class="hatnote navigation-not-searchable">"Retrovirology" redirects here. For the journal, see <a href="/wiki/Retrovirology_(journal)" title="Retrovirology (journal)"><i>Retrovirology</i> (journal)</a>.</div> <p class="mw-empty-elt"> </p> <table class="infobox biota" style="text-align: left; width: 200px; font-size: 100%"> <tbody><tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(250,250,190)"><i>Retroviridae</i> </th></tr> <tr> <td colspan="2" style="text-align: center"><span class="mw-default-size" typeof="mw:File/Frameless"><a href="/wiki/File:Hiv_gross.png" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/f3/Hiv_gross.png/220px-Hiv_gross.png" decoding="async" width="220" height="189" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/f3/Hiv_gross.png/330px-Hiv_gross.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/f3/Hiv_gross.png/440px-Hiv_gross.png 2x" data-file-width="1042" data-file-height="895" /></a></span> </td></tr> <tr> <td colspan="2" style="text-align: center; font-size: 88%">HIV retrovirus schematic of cell infection, virus production and virus structure </td></tr> <tr> <th colspan="2" style="color:inherit; min-width:15em; text-align: center; background-color: rgb(250,250,190)"><a href="/wiki/Virus_classification" title="Virus classification">Virus classification</a> <span class="plainlinks taxobox-edit-taxonomy skin-invert" style="font-size:smaller; float:right; padding-right:0.4em; margin-left:-3em;"><span typeof="mw:File"><a href="/wiki/Template:Taxonomy/Retroviridae" title="Edit this classification"><img alt="Edit this classification" src="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/15px-OOjs_UI_icon_edit-ltr.svg.png" decoding="async" width="15" height="15" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/23px-OOjs_UI_icon_edit-ltr.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/8/8a/OOjs_UI_icon_edit-ltr.svg/30px-OOjs_UI_icon_edit-ltr.svg.png 2x" data-file-width="20" data-file-height="20" /></a></span></span> </th></tr> <tr> <td>(unranked): </td> <td><a href="/wiki/Virus" title="Virus">Virus</a> </td></tr> <tr> <td><i>Realm</i>: </td> <td><a href="/wiki/Riboviria" title="Riboviria"><i>Riboviria</i></a> </td></tr> <tr> <td>Kingdom: </td> <td><a href="/wiki/Revtraviricetes" title="Revtraviricetes"><i>Pararnavirae</i></a> </td></tr> <tr> <td>Phylum: </td> <td><a href="/wiki/Revtraviricetes" title="Revtraviricetes"><i>Artverviricota</i></a> </td></tr> <tr> <td>Class: </td> <td><a href="/wiki/Revtraviricetes" title="Revtraviricetes"><i>Revtraviricetes</i></a> </td></tr> <tr> <td>Order: </td> <td><a href="/wiki/Ortervirales" title="Ortervirales"><i>Ortervirales</i></a> </td></tr> <tr> <td>Family: </td> <td><b><i>Retroviridae</i></b> </td></tr> <tr> <th colspan="2" style="color:inherit; text-align: center; background-color: rgb(250,250,190)">Subfamilies and genera<sup id="cite_ref-1" class="reference"><a href="#cite_note-1"><span class="cite-bracket">[</span>1<span class="cite-bracket">]</span></a></sup> </th></tr> <tr> <td colspan="2" style="text-align: left"> <ul><li><i><a href="/wiki/Orthoretrovirinae" title="Orthoretrovirinae">Orthoretrovirinae</a></i> <ul><li><i><a href="/wiki/Alpharetrovirus" title="Alpharetrovirus">Alpharetrovirus</a></i></li> <li><i><a href="/wiki/Betaretrovirus" title="Betaretrovirus">Betaretrovirus</a></i></li> <li><i><a href="/wiki/Gammaretrovirus" title="Gammaretrovirus">Gammaretrovirus</a></i></li> <li><i><a href="/wiki/Deltaretrovirus" title="Deltaretrovirus">Deltaretrovirus</a></i></li> <li><i><a href="/wiki/Epsilonretrovirus" title="Epsilonretrovirus">Epsilonretrovirus</a></i></li> <li><i><a href="/wiki/Lentivirus" title="Lentivirus">Lentivirus</a></i></li></ul></li> <li><i><a href="/wiki/Spumaretrovirinae" title="Spumaretrovirinae">Spumaretrovirinae</a></i> <ul><li><i><a href="/w/index.php?title=Bovispumavirus&action=edit&redlink=1" class="new" title="Bovispumavirus (page does not exist)">Bovispumavirus</a></i></li> <li><i><a href="/w/index.php?title=Equispumavirus&action=edit&redlink=1" class="new" title="Equispumavirus (page does not exist)">Equispumavirus</a></i></li> <li><i><a href="/w/index.php?title=Felispumavirus&action=edit&redlink=1" class="new" title="Felispumavirus (page does not exist)">Felispumavirus</a></i></li> <li><i><a href="/w/index.php?title=Prosimiispumavirus&action=edit&redlink=1" class="new" title="Prosimiispumavirus (page does not exist)">Prosimiispumavirus</a></i></li> <li><i><a href="/w/index.php?title=Simiispumavirus&action=edit&redlink=1" class="new" title="Simiispumavirus (page does not exist)">Simiispumavirus</a></i></li></ul></li></ul> </td></tr> </tbody></table><style data-mw-deduplicate="TemplateStyles:r1238732961">@media screen{html.skin-theme-clientpref-night .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-night .mw-parser-output .infobox.biota img{background:transparent}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .infobox.biota tr{background:transparent!important}html.skin-theme-clientpref-os .mw-parser-output .infobox.biota img{background:white}}.mw-parser-output .infobox.biota .taxobox-edit-taxonomy img{background:transparent!important}body.skin-vector .mw-parser-output table.biota.infobox{margin-top:0.5em}body.skin--responsive .mw-parser-output table.biota.infobox tr.taxonrow td{padding:2px 10px}</style> <p>A <b>retrovirus</b> is a type of <a href="/wiki/Virus" title="Virus">virus</a> that inserts a DNA copy of its <a href="/wiki/RNA" title="RNA">RNA</a> genome into the <a href="/wiki/DNA" title="DNA">DNA</a> of a <a href="/wiki/Host_cell" class="mw-redirect" title="Host cell">host cell</a> that it invades, thus changing the <a href="/wiki/Genome" title="Genome">genome</a> of that cell.<sup id="cite_ref-2" class="reference"><a href="#cite_note-2"><span class="cite-bracket">[</span>2<span class="cite-bracket">]</span></a></sup> After invading a host cell's <a href="/wiki/Cytoplasm" title="Cytoplasm">cytoplasm</a>, the virus uses its own <a href="/wiki/Reverse_transcriptase" title="Reverse transcriptase">reverse transcriptase</a> enzyme to produce DNA from its RNA genome, the reverse of the usual pattern, thus <i>retro</i> (backward). The new DNA is then <a href="/wiki/Retroviral_integration" class="mw-redirect" title="Retroviral integration">incorporated</a> into the host cell genome by an <a href="/wiki/Integrase" title="Integrase">integrase</a> enzyme, at which point the retroviral DNA is referred to as a <a href="/wiki/Provirus" title="Provirus">provirus</a>. The host cell then treats the viral DNA as part of its own genome, transcribing and translating the viral genes along with the cell's own genes, producing the proteins required to assemble new copies of the virus. Many retroviruses cause serious diseases in humans, other mammals, and birds.<sup id="cite_ref-:1_3-0" class="reference"><a href="#cite_note-:1-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p><p>Retroviruses have many subfamilies in three basic groups. </p> <ul><li><a href="/wiki/Oncovirus" title="Oncovirus">Oncoretroviruses</a> (cancer-causing retroviruses) include <a href="/wiki/Human_T-lymphotropic_virus" class="mw-redirect" title="Human T-lymphotropic virus">human T-lymphotropic virus</a> (HTLV) causing a type of leukemia in humans, and murine leukemia viruses (MLVs) in mice.<sup id="cite_ref-4" class="reference"><a href="#cite_note-4"><span class="cite-bracket">[</span>4<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Lentivirus" title="Lentivirus">Lentiviruses</a> (slow viruses) include <a href="/wiki/HIV-1" class="mw-redirect" title="HIV-1">HIV-1</a> and <a href="/wiki/HIV-2" class="mw-redirect" title="HIV-2">HIV-2</a>, the cause of acquired immune deficiency syndrome (<a href="/wiki/AIDS" class="mw-redirect" title="AIDS">AIDS</a>) in humans.</li> <li><a href="/wiki/Spumavirus" class="mw-redirect" title="Spumavirus">Spumaviruses</a> (foamy viruses) are benign and not linked to any disease in humans or animals.<sup id="cite_ref-Miller,_A._D._2006_5-0" class="reference"><a href="#cite_note-Miller,_A._D._2006-5"><span class="cite-bracket">[</span>5<span class="cite-bracket">]</span></a></sup></li></ul> <p>The specialized DNA-infiltration enzymes in retroviruses make them valuable research tools in molecular biology, and they have been used successfully in gene delivery systems.<sup id="cite_ref-KurthBannert_6-0" class="reference"><a href="#cite_note-KurthBannert-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup> </p><p>Evidence from <a href="/wiki/Endogenous_retrovirus" title="Endogenous retrovirus">endogenous retroviruses</a> (inherited provirus DNA in animal genomes) suggests that retroviruses have been infecting vertebrates for at least 450 million years.<sup id="cite_ref-7" class="reference"><a href="#cite_note-7"><span class="cite-bracket">[</span>7<span class="cite-bracket">]</span></a></sup> </p> <meta property="mw:PageProp/toc" /> <div class="mw-heading mw-heading2"><h2 id="Structure">Structure</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=1" title="Edit section: Structure"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><a href="/wiki/Virion" class="mw-redirect" title="Virion">Virions</a>, viruses in the form of independent particles of retroviruses, consist of enveloped particles about 100 <a href="/wiki/Nanometre" title="Nanometre">nm</a> in diameter. The outer lipid envelope consists of glycoprotein.<sup id="cite_ref-8" class="reference"><a href="#cite_note-8"><span class="cite-bracket">[</span>8<span class="cite-bracket">]</span></a></sup> The virions also contain two identical single-stranded <a href="/wiki/RNA" title="RNA">RNA</a> molecules 7–10 <a href="/wiki/Base_pair#Length_measurements" title="Base pair">kilobases</a> in length. The two molecules are present as a dimer, formed by base pairing between complementary sequences. Interaction sites between the two RNA molecules have been identified as a "<a href="/wiki/Kissing_stem-loop" title="Kissing stem-loop">kissing stem-loop</a>".<sup id="cite_ref-:1_3-1" class="reference"><a href="#cite_note-:1-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> Although virions of different retroviruses do not have the same <a href="/wiki/Morphology_(biology)" title="Morphology (biology)">morphology</a> or biology, all the virion components are very similar.<sup id="cite_ref-she_9-0" class="reference"><a href="#cite_note-she-9"><span class="cite-bracket">[</span>9<span class="cite-bracket">]</span></a></sup> </p><p>The main virion components are: </p> <ul><li><a href="/wiki/Viral_envelope" title="Viral envelope">Envelope</a>: composed of lipids (obtained from the host <a href="/wiki/Plasma_membrane" class="mw-redirect" title="Plasma membrane">plasma membrane</a> during the <a href="/wiki/Budding" title="Budding">budding</a> process) as well as glycoprotein encoded by the <a href="/wiki/Env_(gene)" title="Env (gene)">env gene</a>. The retroviral envelope serves three distinct functions: protection from the extracellular environment via the <a href="/wiki/Lipid_bilayer" title="Lipid bilayer">lipid bilayer</a>, enabling the retrovirus to enter/exit host cells through <a href="/wiki/Endosome" title="Endosome">endosomal</a> <a href="/wiki/Membrane_vesicle_trafficking" title="Membrane vesicle trafficking">membrane trafficking</a>, and the ability to directly enter cells by fusing with their membranes.</li> <li><a href="/wiki/RNA" title="RNA">RNA</a>: consists of a <a href="/wiki/Dimer_(chemistry)" class="mw-redirect" title="Dimer (chemistry)">dimer</a> RNA. It has a cap at the <a href="/wiki/Directionality_(molecular_biology)" title="Directionality (molecular biology)">5'</a> end and a poly(A) tail at the 3' end. Genomic RNA (gRNA) is produced as a result of host RNA polymerase II (Pol II) activity and by adding a 5' methyl cap and a 3' poly-A tail is processed as a host mRNA.<sup id="cite_ref-:0_10-0" class="reference"><a href="#cite_note-:0-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> The RNA genome also has terminal noncoding regions, which are important in replication, and internal regions that encode virion proteins for <a href="/wiki/Gene_expression" title="Gene expression">gene expression</a>. The 5' end includes four regions, which are R, U5, PBS, and L. The R region is a short repeated sequence at each end of the genome used during the <a href="/wiki/Reverse_transcription" class="mw-redirect" title="Reverse transcription">reverse transcription</a> to ensure correct end-to-end transfer in the growing chain. U5, on the other hand, is a short unique sequence between R and PBS. PBS (primer binding site) consists of 18 bases complementary to 3' end of tRNA primer. L region is an untranslated leader region that gives the signal for packaging of the genome RNA. The 3' end includes three regions, which are PPT (polypurine tract), U3, and R. The PPT is a primer for plus-strand DNA synthesis during <a href="/wiki/Reverse_transcription" class="mw-redirect" title="Reverse transcription">reverse transcription</a>. U3 is a sequence between PPT and R, which serves as a signal that the provirus can use in <a href="/wiki/Transcription_(genetics)" class="mw-redirect" title="Transcription (genetics)">transcription</a>. R is the terminal repeated sequence at 3' end.</li> <li><a href="/wiki/Protein" title="Protein">Proteins</a>: consisting of gag proteins, <a href="/wiki/Protease" title="Protease">protease</a> (PR), pol proteins, and env proteins. <ul><li><a href="/wiki/Group-specific_antigen" title="Group-specific antigen">Group-specific antigen</a> (gag) proteins are major components of the viral <a href="/wiki/Capsid" title="Capsid">capsid</a>, which are about 2000–4000 copies per virion. Gag possesses two nucleic acid binding domains, including matrix (MA) and nucleocapsid (NC). Specifically recognizing, binding, and packaging the retroviral genomic RNA into assembling virions is one of the important functions of Gag protein. Gag interactions with cellular RNAs also regulate aspects of assembly.<sup id="cite_ref-11" class="reference"><a href="#cite_note-11"><span class="cite-bracket">[</span>11<span class="cite-bracket">]</span></a></sup> The expression of <i>gag</i> alone gives rise to assembly of immature virus-like particles that bud from the plasma membrane. In all retroviruses the Gag protein is the precursor to the internal structural protein.<sup id="cite_ref-12" class="reference"><a href="#cite_note-12"><span class="cite-bracket">[</span>12<span class="cite-bracket">]</span></a></sup></li> <li><a href="/wiki/Retroviral_aspartyl_protease" title="Retroviral aspartyl protease">Protease</a> (pro) is expressed differently in different viruses. It functions in proteolytic cleavages during virion maturation to make mature gag and pol proteins. Retroviral Gag proteins are responsible for coordinating many aspects of virion assembly.</li> <li><a href="/w/index.php?title=Pol_proteins&action=edit&redlink=1" class="new" title="Pol proteins (page does not exist)">Pol proteins</a> are responsible for synthesis of viral DNA and integration into host DNA after infection.</li> <li><a href="/w/index.php?title=Env_proteins&action=edit&redlink=1" class="new" title="Env proteins (page does not exist)">Env proteins</a> play a role in association and entry of virions into the host cell.<sup id="cite_ref-se_13-0" class="reference"><a href="#cite_note-se-13"><span class="cite-bracket">[</span>13<span class="cite-bracket">]</span></a></sup> Possessing a functional copy of an <a href="/wiki/Env_(gene)" title="Env (gene)">env gene</a> is what makes retroviruses distinct from <a href="/wiki/Retroelements" class="mw-redirect" title="Retroelements">retroelements</a>.<sup id="cite_ref-14" class="reference"><a href="#cite_note-14"><span class="cite-bracket">[</span>14<span class="cite-bracket">]</span></a></sup> The ability of the retrovirus to bind to its target host cell using specific cell-surface receptors is given by the surface component (SU) of the Env protein, while the ability of the retrovirus to enter the cell via <a href="/wiki/Lipid_bilayer_fusion" title="Lipid bilayer fusion">membrane fusion</a> is imparted by the membrane-anchored trans-membrane component (TM). Thus it is the Env protein that enables the retrovirus to be infectious.</li> <li>Several protein species are associated with the RNA in the retrovirus virion. Nucleocapsid (NC) protein is the most abundant protein, which coats the RNA; while other proteins, present in much smaller amounts and have enzyme activities. Some enzyme activities that are present in the retrovirus virion includes RNA-dependent DNA polymerase (reverse transcriptase; RT), DNA-dependent DNA polymerase, Ribonuclease H (RNase H) Integrase and Protease.<sup id="cite_ref-John_Wiley_&_Sons_15-0" class="reference"><a href="#cite_note-John_Wiley_&_Sons-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> The retroviral RNases H encoded by all retroviruses, including HIV have been demonstrated to show three different modes of cleavage: internal, DNA 3′ end-directed, and RNA 5′ end-directed. All three modes of cleavage constitute roles in reverse transcription. Therefore, The RNase H activity is essential in several aspects of reverse transcription. The use of an RNase H activity during retroviral replication displays a unique strategy to copy a single-stranded RNA genome into a double-stranded DNA, since the minus-strand DNA are complementary and make base pairing to retrovirus genome in the first cycle of DNA synthesis.<sup id="cite_ref-16" class="reference"><a href="#cite_note-16"><span class="cite-bracket">[</span>16<span class="cite-bracket">]</span></a></sup> The RNase H ribonuclease activity is also required in the retroviral life cycle, since it generates and removes primers essential by the Reverse Transcriptase (RT) for the initiation of DNA synthesis. Retroviruses that are lacking RNase H activity are noninfectious.<sup id="cite_ref-17" class="reference"><a href="#cite_note-17"><span class="cite-bracket">[</span>17<span class="cite-bracket">]</span></a></sup></li></ul></li></ul> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Prototypical_retrovirus_genomic_organization.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Prototypical_retrovirus_genomic_organization.svg/280px-Prototypical_retrovirus_genomic_organization.svg.png" decoding="async" width="280" height="56" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Prototypical_retrovirus_genomic_organization.svg/420px-Prototypical_retrovirus_genomic_organization.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/c/cd/Prototypical_retrovirus_genomic_organization.svg/560px-Prototypical_retrovirus_genomic_organization.svg.png 2x" data-file-width="238" data-file-height="48" /></a><figcaption>The genomic and subgenomic organization of a prototypical retrovirus. Abbreviations are explained in the file description.<sup id="cite_ref-pmid26800882_18-0" class="reference"><a href="#cite_note-pmid26800882-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup></figcaption></figure> <div class="mw-heading mw-heading3"><h3 id="Genomic_structure">Genomic structure</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=2" title="Edit section: Genomic structure"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The retroviral genome is packaged as viral particles. These viral particles are dimers of single-stranded, positive-sense, linear RNA molecules.<sup id="cite_ref-:0_10-1" class="reference"><a href="#cite_note-:0-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p><p>Retroviruses (and <a href="/wiki/Ortervirales" title="Ortervirales">orterviruses</a> in general) follow a layout of 5'–<i>gag</i>–<i>pro</i>–<i>pol</i>–<i>env</i>–3' in the RNA genome. <i>gag</i> and <i>pol</i> encode polyproteins, each managing the capsid and replication. The <i>pol</i> region encodes enzymes necessary for viral replication, such as reverse transcriptase, protease and integrase.<sup id="cite_ref-19" class="reference"><a href="#cite_note-19"><span class="cite-bracket">[</span>19<span class="cite-bracket">]</span></a></sup> Depending on the virus, the genes may overlap or fuse into larger polyprotein chains. Some viruses contain additional genes. The lentivirus genus, the spumavirus genus, the HTLV / bovine leukemia virus (BLV) genus, and a newly introduced fish virus genus are retroviruses classified as complex. These viruses have genes called accessory genes, in addition to gag, pro, pol and env genes. Accessory genes are located between pol and env, downstream from the env, including the U3 region of LTR, or in the env and overlapping portions. While accessory genes have auxiliary roles, they also coordinate and regulate viral gene expression. In addition, some retroviruses may carry genes called oncogenes or onc genes from another class. Retroviruses with these genes (also called transforming viruses) are known for their ability to quickly cause tumors in animals and transform cells in culture into an oncogenic state.<sup id="cite_ref-20" class="reference"><a href="#cite_note-20"><span class="cite-bracket">[</span>20<span class="cite-bracket">]</span></a></sup> </p><p>The polyproteins are cleaved into smaller proteins each with their own function. The nucleotides encoding them are known as <i>subgenes</i>.<sup id="cite_ref-pmid26800882_18-1" class="reference"><a href="#cite_note-pmid26800882-18"><span class="cite-bracket">[</span>18<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Multiplication">Multiplication</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=3" title="Edit section: Multiplication"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Life_Cycle_of_a_Retrovirus.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/2/2b/Life_Cycle_of_a_Retrovirus.svg/440px-Life_Cycle_of_a_Retrovirus.svg.png" decoding="async" width="440" height="173" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/2b/Life_Cycle_of_a_Retrovirus.svg/660px-Life_Cycle_of_a_Retrovirus.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/2b/Life_Cycle_of_a_Retrovirus.svg/880px-Life_Cycle_of_a_Retrovirus.svg.png 2x" data-file-width="512" data-file-height="201" /></a><figcaption>A retrovirus has a membrane containing glycoproteins, which are able to bind to a receptor protein on a host cell. There are two strands of RNA within the cell that have three enzymes: protease, reverse transcriptase, and integrase (1). The first step of replication is the binding of the glycoprotein to the receptor protein (2). Once these have been bound, the cell membrane degrades, becoming part of the host cell, and the RNA strands and enzymes enter the cell (3). Within the cell, reverse transcriptase creates a complementary strand of DNA from the retrovirus RNA and the RNA is degraded; this strand of DNA is known as cDNA (4). The cDNA is then replicated, and the two strands form a weak bond and enter the nucleus (5). Once in the nucleus, the DNA is integrated into the host cell's DNA with the help of integrase (6). This cell can either stay dormant, or RNA may be synthesized from the DNA and used to create the proteins for a new retrovirus (7). Ribosome units are used to translate the mRNA of the virus into the amino acid sequences which can be made into proteins in the rough endoplasmic reticulum. This step will also make viral enzymes and capsid proteins (8). Viral RNA will be made in the nucleus. These pieces are then gathered together and are pinched off of the cell membrane as a new retrovirus (9).</figcaption></figure> <p>When retroviruses have integrated their own genome into the <a href="/wiki/Germline" title="Germline">germ line</a>, their genome is <a href="/wiki/Inheritance" title="Inheritance">passed on</a> to a following generation. These <a href="/wiki/Endogenous_retrovirus" title="Endogenous retrovirus">endogenous retroviruses</a> (ERVs), contrasted with <a href="/wiki/Exogenous_DNA" title="Exogenous DNA">exogenous</a> ones, now make up 5–8% of the human genome.<sup id="cite_ref-21" class="reference"><a href="#cite_note-21"><span class="cite-bracket">[</span>21<span class="cite-bracket">]</span></a></sup> Most insertions have no known function and are often referred to as "<a href="/wiki/Junk_DNA" title="Junk DNA">junk DNA</a>". However, many endogenous retroviruses play important roles in host biology, such as control of gene transcription, cell fusion during <a href="/wiki/Placenta" title="Placenta">placental</a> development in the course of the <a href="/wiki/Germination" title="Germination">germination</a> of an <a href="/wiki/Embryo" title="Embryo">embryo</a>, and resistance to exogenous retroviral infection. Endogenous retroviruses have also received special attention in the research of <a href="/wiki/Immunology" title="Immunology">immunology</a>-related pathologies, such as <a href="/wiki/Autoimmune_disease" title="Autoimmune disease">autoimmune diseases</a> like <a href="/wiki/Multiple_sclerosis" title="Multiple sclerosis">multiple sclerosis</a>, although endogenous retroviruses have not yet been proven to play any causal role in this class of disease.<sup id="cite_ref-22" class="reference"><a href="#cite_note-22"><span class="cite-bracket">[</span>22<span class="cite-bracket">]</span></a></sup> </p><p>While transcription was classically thought to occur only from DNA to RNA, <a href="/wiki/Reverse_transcriptase" title="Reverse transcriptase">reverse transcriptase</a> transcribes RNA into DNA. The term "retro" in retrovirus refers to this reversal (making DNA from RNA) of the usual direction of transcription. It still obeys the <a href="/wiki/Central_dogma_of_molecular_biology" title="Central dogma of molecular biology">central dogma of molecular biology</a>, which states that information can be transferred from nucleic acid to nucleic acid but cannot be transferred back from protein to either protein or nucleic acid. Reverse transcriptase activity outside of retroviruses has been found in almost all <a href="/wiki/Eukaryote" title="Eukaryote">eukaryotes</a>, enabling the generation and insertion of new copies of <a href="/wiki/Retrotransposon" title="Retrotransposon">retrotransposons</a> into the host genome. These inserts are transcribed by enzymes of the host into new RNA molecules that enter the cytosol. Next, some of these RNA molecules are translated into viral proteins. The proteins encoded by the gag and pol genes are translated from genome-length mRNAs into Gag and Gag–Pol polyproteins. In example, for the <i>gag</i> gene; it is translated into molecules of the capsid protein, and for the <i>pol</i> gene; it is translated into molecules of reverse transcriptase. Retroviruses need a lot more of the Gag proteins than the Pol proteins and have developed advanced systems to synthesize the required amount of each. As an example, after Gag synthesis nearly 95 percent of the ribosomes terminate translation, while other ribosomes continue translation to synthesize Gag–Pol. In the rough endoplasmic reticulum glycosylation begins and the <i>env</i> gene is translated from spliced mRNAs in the rough endoplasmic reticulum, into molecules of the envelope protein. When the envelope protein molecules are carried to the Golgi complex, they are divided into surface glycoprotein and transmembrane glycoprotein by a host protease. These two glycoprotein products stay in close affiliation, and they are transported to the plasma membrane after further glycosylation.<sup id="cite_ref-:1_3-2" class="reference"><a href="#cite_note-:1-3"><span class="cite-bracket">[</span>3<span class="cite-bracket">]</span></a></sup> </p><p>It is important to note that a retrovirus must "bring" its own reverse transcriptase in its <a href="/wiki/Capsid" title="Capsid">capsid</a>, otherwise it is unable to utilize the enzymes of the infected cell to carry out the task, due to the unusual nature of producing DNA from RNA.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (November 2022)">citation needed</span></a></i>]</sup> </p><p>Industrial drugs that are designed as protease and <a href="/wiki/Reverse-transcriptase_inhibitor" title="Reverse-transcriptase inhibitor">reverse-transcriptase inhibitors</a> are made such that they target specific sites and sequences within their respective enzymes. However these drugs can quickly become ineffective due to the fact that the gene sequences that code for the protease and the reverse transcriptase quickly mutate. These changes in bases cause specific codons and sites with the enzymes to change and thereby avoid drug targeting by losing the sites that the drug actually targets.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (November 2022)">citation needed</span></a></i>]</sup> </p><p>Because reverse transcription lacks the usual <a href="/wiki/Proofreading_(biology)" title="Proofreading (biology)">proofreading</a> of DNA replication, a retrovirus <a href="/wiki/Mutate" class="mw-redirect" title="Mutate">mutates</a> very often. This enables the virus to grow <a href="/wiki/Immunology" title="Immunology">resistant</a> to antiviral pharmaceuticals quickly, and impedes the development of effective <a href="/wiki/Vaccines" class="mw-redirect" title="Vaccines">vaccines</a> and inhibitors for the retrovirus.<sup id="cite_ref-23" class="reference"><a href="#cite_note-23"><span class="cite-bracket">[</span>23<span class="cite-bracket">]</span></a></sup> </p><p>One difficulty faced with some retroviruses, such as the Moloney retrovirus, involves the requirement for cells to be actively dividing for transduction. As a result, cells such as neurons are very resistant to infection and transduction by retroviruses. This gives rise to a concern that insertional mutagenesis due to integration into the host genome might lead to cancer or leukemia. This is unlike <i><a href="/wiki/Lentivirus" title="Lentivirus">Lentivirus</a></i>, a genus of <i>Retroviridae</i>, which are able to integrate their RNA into the genome of non-dividing host cells.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (November 2022)">citation needed</span></a></i>]</sup> </p> <div class="mw-heading mw-heading3"><h3 id="Recombination">Recombination</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=4" title="Edit section: Recombination"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Two <a href="/wiki/RNA" title="RNA">RNA</a> <a href="/wiki/Genome" title="Genome">genomes</a> are packaged into each retrovirus particle, but, after an infection, each virus generates only one <a href="/wiki/Provirus" title="Provirus">provirus</a>.<sup id="cite_ref-Rawson2018_24-0" class="reference"><a href="#cite_note-Rawson2018-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup> After infection, <a href="/wiki/Reverse_transcriptase" title="Reverse transcriptase">reverse transcription</a> occurs and this process is accompanied by <a href="/wiki/Homologous_recombination" title="Homologous recombination">recombination</a>. Recombination involves template strand switching between the two genome copies (copy choice recombination) during reverse transcription. From 5 to 14 recombination events per genome occur at each replication cycle.<sup id="cite_ref-pmid26691546_25-0" class="reference"><a href="#cite_note-pmid26691546-25"><span class="cite-bracket">[</span>25<span class="cite-bracket">]</span></a></sup> Genetic recombination appears to be necessary for maintaining genome integrity and as a repair mechanism for salvaging damaged genomes.<sup id="cite_ref-Rawson2018_24-1" class="reference"><a href="#cite_note-Rawson2018-24"><span class="cite-bracket">[</span>24<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Transmission">Transmission</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=5" title="Edit section: Transmission"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <ul><li>Cell-to-cell<sup id="cite_ref-26" class="reference"><a href="#cite_note-26"><span class="cite-bracket">[</span>26<span class="cite-bracket">]</span></a></sup></li> <li>Fluids</li> <li>Airborne, like the <a href="/wiki/Jaagsiekte_sheep_retrovirus" title="Jaagsiekte sheep retrovirus">Jaagsiekte sheep retrovirus</a>.</li></ul> <div class="mw-heading mw-heading2"><h2 id="Provirus">Provirus</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=6" title="Edit section: Provirus"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>The DNA formed after reverse transcription (the provirus) is longer than the RNA genome because each of the terminals have the U3 - R - U5 sequences called long terminal repeat (LTR). Thus, 5' terminal has the extra U3 sequence, while the other terminal has the U5 sequence.<sup id="cite_ref-John_Wiley_&_Sons_15-1" class="reference"><a href="#cite_note-John_Wiley_&_Sons-15"><span class="cite-bracket">[</span>15<span class="cite-bracket">]</span></a></sup> LTRs are able to send signals for vital tasks to be carried out such as initiation of RNA production or management of the rate of transcription. This way, LTRs can control replication, hence, the entire progress of the viral cycle.<sup id="cite_ref-27" class="reference"><a href="#cite_note-27"><span class="cite-bracket">[</span>27<span class="cite-bracket">]</span></a></sup> Although located in the nucleus, the non-integrated retroviral cDNA is a very weak substrate for transcription. For this reason, an integrated provirus is a necessary for permanent and an effective expression of retroviral genes.<sup id="cite_ref-:0_10-2" class="reference"><a href="#cite_note-:0-10"><span class="cite-bracket">[</span>10<span class="cite-bracket">]</span></a></sup> </p><p>This DNA can be incorporated into host genome as a provirus that can be passed on to progeny cells. The retrovirus DNA is inserted at random into the host genome. Because of this, it can be inserted into <a href="/wiki/Oncogene" title="Oncogene">oncogenes</a>. In this way some retroviruses can convert normal cells into cancer cells. Some provirus remains latent in the cell for a long period of time before it is activated by the change in cell environment.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (November 2022)">citation needed</span></a></i>]</sup> </p> <div class="mw-heading mw-heading2"><h2 id="Early_evolution">Early evolution</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=7" title="Edit section: Early evolution"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Studies of retroviruses led to the first demonstrated synthesis of DNA from RNA templates, a fundamental mode for transferring genetic material that occurs in both <a href="/wiki/Eukaryote" title="Eukaryote">eukaryotes</a> and <a href="/wiki/Prokaryote" title="Prokaryote">prokaryotes</a>. It has been speculated that the RNA to DNA transcription processes used by retroviruses may have first caused DNA to be used as genetic material. In this model, the <a href="/wiki/RNA_world_hypothesis" class="mw-redirect" title="RNA world hypothesis">RNA world hypothesis</a>, cellular organisms adopted the more chemically stable DNA when retroviruses evolved to create <a href="/wiki/DNA" title="DNA">DNA</a> from the <a href="/wiki/RNA" title="RNA">RNA</a> templates.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (November 2022)">citation needed</span></a></i>]</sup> </p><p>An estimate of the date of evolution of the foamy-like endogenous retroviruses placed the time of the most recent common ancestor at > <span class="nourlexpansion plainlinks" style="white-space:nowrap;"><a rel="nofollow" class="external text" href="https://geoltime.github.io/?Ma=450">450</a> million years ago</span>.<sup id="cite_ref-Aiewsakun_28-0" class="reference"><a href="#cite_note-Aiewsakun-28"><span class="cite-bracket">[</span>28<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Gene_therapy">Gene therapy</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=8" title="Edit section: Gene therapy"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><a href="/wiki/Gammaretrovirus" title="Gammaretrovirus">Gammaretroviral</a> and <a href="/wiki/Lentiviral" class="mw-redirect" title="Lentiviral">lentiviral</a> <a href="/wiki/Vector_(molecular_biology)" title="Vector (molecular biology)">vectors</a> for <a href="/wiki/Gene_therapy" title="Gene therapy">gene therapy</a> have been developed that mediate stable genetic modification of treated cells by chromosomal integration of the transferred vector genomes. This technology is of use, not only for research purposes, but also for clinical gene therapy aiming at the long-term correction of genetic defects, e.g., in stem and progenitor cells. Retroviral vector particles with tropism for various target cells have been designed. Gammaretroviral and lentiviral vectors have so far been used in more than 300 clinical trials, addressing treatment options for various diseases.<sup id="cite_ref-KurthBannert_6-1" class="reference"><a href="#cite_note-KurthBannert-6"><span class="cite-bracket">[</span>6<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-Desportm_29-0" class="reference"><a href="#cite_note-Desportm-29"><span class="cite-bracket">[</span>29<span class="cite-bracket">]</span></a></sup> Retroviral mutations can be developed to make transgenic mouse models to study various cancers and their <a href="/wiki/Mouse_models_of_breast_cancer_metastasis" title="Mouse models of breast cancer metastasis">metastatic models</a>.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (November 2022)">citation needed</span></a></i>]</sup> </p> <div class="mw-heading mw-heading2"><h2 id="Cancer">Cancer</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=9" title="Edit section: Cancer"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Retroviruses that cause tumor growth include <i><a href="/wiki/Rous_sarcoma_virus" title="Rous sarcoma virus">Rous sarcoma virus</a></i> and <i><a href="/wiki/Mouse_mammary_tumor_virus" title="Mouse mammary tumor virus">mouse mammary tumor virus</a></i>. Cancer can be triggered by proto-oncogenes that were mistakenly incorporated into proviral DNA or by the disruption of cellular proto-oncogenes. Rous sarcoma virus contains the <a href="/wiki/Src_(gene)" class="mw-redirect" title="Src (gene)">src gene</a> that triggers tumor formation. Later it was found that a similar gene in cells is involved in cell signaling, which was most likely excised with the proviral DNA. Nontransforming viruses can randomly insert their DNA into proto-oncogenes, disrupting the expression of proteins that regulate the cell cycle. The promoter of the provirus DNA can also cause over expression of regulatory genes. Retroviruses can cause diseases such as cancer and immunodeficiency. If viral DNA is integrated into host chromosomes, it can lead to permanent infections. It is therefore important to discover the body's response to retroviruses. Exogenous retroviruses are especially associated with pathogenic diseases. For example, mice have mouse mammary tumor virus (MMTV), which is a retrovirus. This virus passes to newborn mice through mammary milk. When they are 6 months old, the mice carrying the virus get mammary cancer because of the retrovirus. In addition, leukemia virus I (HTLV-1), found in human T cell, has been found in humans for many years. It is estimated that this retrovirus causes leukemia in the ages of 40 and 50.<sup id="cite_ref-30" class="reference"><a href="#cite_note-30"><span class="cite-bracket">[</span>30<span class="cite-bracket">]</span></a></sup> It has a replicable structure that can induce cancer. In addition to the usual gene sequence of retroviruses, HTLV-1 contains a fourth region, PX. This region encodes Tax, Rex, p12, p13 and p30 regulatory proteins. The Tax protein initiates the leukemic process and organizes the transcription of all viral genes in the integrated HTLV proviral DNA. <sup id="cite_ref-31" class="reference"><a href="#cite_note-31"><span class="cite-bracket">[</span>31<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Classification">Classification</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=10" title="Edit section: Classification"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <figure class="mw-default-size" typeof="mw:File/Thumb"><a href="/wiki/File:Phylogeny_of_Retroviruses.svg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/9/97/Phylogeny_of_Retroviruses.svg/330px-Phylogeny_of_Retroviruses.svg.png" decoding="async" width="330" height="227" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/9/97/Phylogeny_of_Retroviruses.svg/495px-Phylogeny_of_Retroviruses.svg.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/9/97/Phylogeny_of_Retroviruses.svg/660px-Phylogeny_of_Retroviruses.svg.png 2x" data-file-width="570" data-file-height="392" /></a><figcaption>Phylogeny of Retroviridae</figcaption></figure> <div class="mw-heading mw-heading3"><h3 id="Exogenous">Exogenous</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=11" title="Edit section: Exogenous"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1251242444">.mw-parser-output .ambox{border:1px solid #a2a9b1;border-left:10px solid #36c;background-color:#fbfbfb;box-sizing:border-box}.mw-parser-output .ambox+link+.ambox,.mw-parser-output .ambox+link+style+.ambox,.mw-parser-output .ambox+link+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+style+.ambox,.mw-parser-output .ambox+.mw-empty-elt+link+link+.ambox{margin-top:-1px}html body.mediawiki .mw-parser-output .ambox.mbox-small-left{margin:4px 1em 4px 0;overflow:hidden;width:238px;border-collapse:collapse;font-size:88%;line-height:1.25em}.mw-parser-output .ambox-speedy{border-left:10px solid #b32424;background-color:#fee7e6}.mw-parser-output .ambox-delete{border-left:10px solid #b32424}.mw-parser-output .ambox-content{border-left:10px solid #f28500}.mw-parser-output .ambox-style{border-left:10px solid #fc3}.mw-parser-output .ambox-move{border-left:10px solid #9932cc}.mw-parser-output .ambox-protection{border-left:10px solid #a2a9b1}.mw-parser-output .ambox .mbox-text{border:none;padding:0.25em 0.5em;width:100%}.mw-parser-output .ambox .mbox-image{border:none;padding:2px 0 2px 0.5em;text-align:center}.mw-parser-output .ambox .mbox-imageright{border:none;padding:2px 0.5em 2px 0;text-align:center}.mw-parser-output .ambox .mbox-empty-cell{border:none;padding:0;width:1px}.mw-parser-output .ambox .mbox-image-div{width:52px}@media(min-width:720px){.mw-parser-output .ambox{margin:0 10%}}@media print{body.ns-0 .mw-parser-output .ambox{display:none!important}}</style><table class="box-Cleanup plainlinks metadata ambox ambox-style ambox-Cleanup" role="presentation"><tbody><tr><td class="mbox-image"><div class="mbox-image-div"><span typeof="mw:File"><span><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/f/f2/Edit-clear.svg/40px-Edit-clear.svg.png" decoding="async" width="40" height="40" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/f/f2/Edit-clear.svg/60px-Edit-clear.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/f/f2/Edit-clear.svg/80px-Edit-clear.svg.png 2x" data-file-width="48" data-file-height="48" /></span></span></div></td><td class="mbox-text"><div class="mbox-text-span">This section may <b>require <a href="/wiki/Wikipedia:Cleanup" title="Wikipedia:Cleanup">cleanup</a></b> to meet Wikipedia's <a href="/wiki/Wikipedia:Manual_of_Style" title="Wikipedia:Manual of Style">quality standards</a>. The specific problem is: <b>Not the "Retroviridae" taxon the article talks about; suggest move into <a href="/wiki/Baltimore_classification#Reverse_transcribing_viruses" title="Baltimore classification">Baltimore classification#Reverse transcribing viruses</a> or <a href="/wiki/Revtraviricetes" title="Revtraviricetes">Revtraviricetes</a>.</b><span class="hide-when-compact"> Please help <a href="/wiki/Special:EditPage/Retrovirus" title="Special:EditPage/Retrovirus">improve this section</a> if you can.</span> <span class="date-container"><i>(<span class="date">February 2021</span>)</i></span><span class="hide-when-compact"><i> (<small><a href="/wiki/Help:Maintenance_template_removal" title="Help:Maintenance template removal">Learn how and when to remove this message</a></small>)</i></span></div></td></tr></tbody></table> <p>Exogenous retroviruses are infectious RNA- or DNA-containing viruses that are transmitted from one organism to another. In the <a href="/wiki/Baltimore_classification" title="Baltimore classification">Baltimore classification</a> system, which groups viruses together based on their manner of <a href="/wiki/Messenger_RNA" title="Messenger RNA">messenger RNA</a> synthesis, they are classified into two groups: Group VI: single-stranded RNA viruses with a DNA intermediate in their life cycle, and Group VII: double-stranded DNA viruses with an RNA intermediate in their life cycle.<sup class="noprint Inline-Template Template-Fact" style="white-space:nowrap;">[<i><a href="/wiki/Wikipedia:Citation_needed" title="Wikipedia:Citation needed"><span title="This claim needs references to reliable sources. (May 2021)">citation needed</span></a></i>]</sup> </p> <div class="mw-heading mw-heading4"><h4 id="Group_VI_viruses">Group VI viruses</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=12" title="Edit section: Group VI viruses"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>All members of <a href="/wiki/Virus_classification#Baltimore_classification" title="Virus classification">Group VI</a> use virally encoded <a href="/wiki/Reverse_transcriptase" title="Reverse transcriptase">reverse transcriptase</a>, an RNA-dependent DNA polymerase, to produce DNA from the initial virion RNA genome. This DNA is often integrated into the host genome, as in the case of retroviruses and <a href="/wiki/Pseudoviridae" title="Pseudoviridae">pseudoviruses</a>, where it is replicated and <a href="/wiki/Transcription_(genetics)" class="mw-redirect" title="Transcription (genetics)">transcribed</a> by the host. </p><p>Group VI includes: </p> <ul><li>Order <a href="/wiki/Ortervirales" title="Ortervirales">Ortervirales</a> <ul><li>Family <i><a href="/wiki/Belpaoviridae" class="mw-redirect" title="Belpaoviridae">Belpaoviridae</a></i></li> <li>Family <i><a href="/wiki/Metaviridae" title="Metaviridae">Metaviridae</a></i></li> <li>Family <i><a href="/wiki/Pseudoviridae" title="Pseudoviridae">Pseudoviridae</a></i></li> <li>Family <i><a href="/wiki/Retroviridae" class="mw-redirect" title="Retroviridae">Retroviridae</a></i> – Retroviruses, e.g. <i><a href="/wiki/HIV" title="HIV">HIV</a></i></li> <li>Family <i><a href="/wiki/Caulimoviridae" title="Caulimoviridae">Caulimoviridae</a></i> – a VII group virus family (see below)</li></ul></li></ul> <p>The family <i>Retroviridae</i> was previously divided into three subfamilies (<i>Oncovirinae</i>, <i>Lentivirinae</i>, and <i>Spumavirinae</i>), but are now divided into two: <i>Orthoretrovirinae</i> and <i>Spumaretrovirinae</i>. The term <a href="/wiki/Oncovirus" title="Oncovirus">oncovirus</a> is now commonly used to describe a cancer-causing virus. This family now includes the following genera: </p> <ul><li>Subfamily <i><a href="/wiki/Orthoretrovirinae" title="Orthoretrovirinae">Orthoretrovirinae</a></i>: <ul><li>Genus <i><a href="/wiki/Alpharetrovirus" title="Alpharetrovirus">Alpharetrovirus</a></i>; including <i><a href="/wiki/Avian_leukosis_virus" class="mw-redirect" title="Avian leukosis virus">Avian leukosis virus</a></i> and <i><a href="/wiki/Rous_sarcoma_virus" title="Rous sarcoma virus">Rous sarcoma virus</a></i></li> <li>Genus <i><a href="/wiki/Betaretrovirus" title="Betaretrovirus">Betaretrovirus</a></i>; including <i><a href="/wiki/Mouse_mammary_tumour_virus" class="mw-redirect" title="Mouse mammary tumour virus">Mouse mammary tumour virus</a></i></li> <li>Genus <i><a href="/wiki/Gammaretrovirus" title="Gammaretrovirus">Gammaretrovirus</a></i>; including <i><a href="/wiki/Murine_leukemia_virus" title="Murine leukemia virus">Murine leukemia virus</a></i> and <i><a href="/wiki/Feline_leukemia_virus" title="Feline leukemia virus">Feline leukemia virus</a></i></li> <li>Genus <i><a href="/wiki/Deltaretrovirus" title="Deltaretrovirus">Deltaretrovirus</a></i>; including <i><a href="/wiki/Bovine_leukemia_virus" title="Bovine leukemia virus">Bovine leukemia virus</a></i> and the cancer-causing <i><a href="/wiki/Human_T-lymphotropic_virus" class="mw-redirect" title="Human T-lymphotropic virus">Human T-lymphotropic virus</a></i></li> <li>Genus <i><a href="/wiki/Epsilonretrovirus" title="Epsilonretrovirus">Epsilonretrovirus</a></i></li> <li>Genus <i><a href="/wiki/Lentivirus" title="Lentivirus">Lentivirus</a></i>; including <i><a href="/wiki/HIV" title="HIV">Human immunodeficiency virus 1</a></i> and <i><a href="/wiki/Simian_immunodeficiency_virus" title="Simian immunodeficiency virus">Simian</a></i> and <i><a href="/wiki/Feline_immunodeficiency_virus" title="Feline immunodeficiency virus">Feline</a></i> immunodeficiency viruses</li></ul></li> <li>Subfamily <i><a href="/wiki/Spumaviruses" class="mw-redirect" title="Spumaviruses">Spumaretrovirinae</a></i>: <ul><li>Genus <i><a href="/w/index.php?title=Bovispumavirus&action=edit&redlink=1" class="new" title="Bovispumavirus (page does not exist)">Bovispumavirus</a></i></li> <li>Genus <i><a href="/w/index.php?title=Equispumavirus&action=edit&redlink=1" class="new" title="Equispumavirus (page does not exist)">Equispumavirus</a></i></li> <li>Genus <i><a href="/w/index.php?title=Felispumavirus&action=edit&redlink=1" class="new" title="Felispumavirus (page does not exist)">Felispumavirus</a></i></li> <li>Genus <i><a href="/w/index.php?title=Prosimiispumavirus&action=edit&redlink=1" class="new" title="Prosimiispumavirus (page does not exist)">Prosimiispumavirus</a></i></li> <li>Genus <i><a href="/wiki/Simian_foamy_virus" class="mw-redirect" title="Simian foamy virus">Simiispumavirus</a></i></li></ul></li></ul> <p>Note that according to <a href="/wiki/International_Committee_on_Taxonomy_of_Viruses" title="International Committee on Taxonomy of Viruses">ICTV</a> 2017, genus <i><a href="/wiki/Spumavirus" class="mw-redirect" title="Spumavirus">Spumavirus</a></i> has been divided into five genera, and its former type species <i><a href="/wiki/Simian_foamy_virus" class="mw-redirect" title="Simian foamy virus">Simian foamy virus</a></i> is now upgraded to genus <i>Simiispumavirus</i> with not less than 14 species, including new type species <i><a href="/wiki/Eastern_chimpanzee_simian_foamy_virus" class="mw-redirect" title="Eastern chimpanzee simian foamy virus">Eastern chimpanzee simian foamy virus</a></i>.<sup id="cite_ref-32" class="reference"><a href="#cite_note-32"><span class="cite-bracket">[</span>32<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading4"><h4 id="Group_VII_viruses">Group VII viruses</h4><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=13" title="Edit section: Group VII viruses"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/DsDNA-RT_virus" class="mw-redirect" title="DsDNA-RT virus">dsDNA-RT virus</a></div> <p>Both families in <a href="/wiki/Virus_classification#Baltimore_classification" title="Virus classification">Group VII</a> have DNA genomes contained within the invading virus particles. The DNA genome is transcribed into both mRNA, for use as a transcript in protein synthesis, and pre-genomic RNA, for use as the template during genome replication. Virally encoded <a href="/wiki/Reverse_transcriptase" title="Reverse transcriptase">reverse transcriptase</a> uses the pre-genomic RNA as a template for the creation of genomic DNA. </p><p>Group VII includes: </p> <ul><li>Family <i><a href="/wiki/Caulimoviridae" title="Caulimoviridae">Caulimoviridae</a></i> — e.g. <i><a href="/wiki/Cauliflower_mosaic_virus" title="Cauliflower mosaic virus">Cauliflower mosaic virus</a></i></li> <li>Family <i><a href="/wiki/Hepadnaviridae" title="Hepadnaviridae">Hepadnaviridae</a></i> — e.g. <i><a href="/wiki/Hepatitis_B_virus" title="Hepatitis B virus">Hepatitis B virus</a></i></li></ul> <p>The latter family is closely related to the newly proposed </p> <ul><li>Family <i><a href="/wiki/Nackednaviridae" class="mw-redirect" title="Nackednaviridae">Nackednaviridae</a></i> — e.g. <i><a href="/w/index.php?title=African_cichlid_nackednavirus&action=edit&redlink=1" class="new" title="African cichlid nackednavirus (page does not exist)">African cichlid nackednavirus</a></i> (ACNDV), formerly named <i>African cichlid hepatitis B virus</i> (ACHBV).<sup id="cite_ref-Lauber2017_33-0" class="reference"><a href="#cite_note-Lauber2017-33"><span class="cite-bracket">[</span>33<span class="cite-bracket">]</span></a></sup></li></ul> <p>whilst families <i><a href="/wiki/Belpaoviridae" class="mw-redirect" title="Belpaoviridae">Belpaoviridae</a></i>, <i><a href="/wiki/Metaviridae" title="Metaviridae">Metaviridae</a></i>, <i><a href="/wiki/Pseudoviridae" title="Pseudoviridae">Pseudoviridae</a></i>, <i><a href="/wiki/Retroviridae" class="mw-redirect" title="Retroviridae">Retroviridae</a></i>, and <i><a href="/wiki/Caulimoviridae" title="Caulimoviridae">Caulimoviridae</a></i> constitute the order <i><a href="/wiki/Ortervirales" title="Ortervirales">Ortervirales</a></i>.<sup id="cite_ref-34" class="reference"><a href="#cite_note-34"><span class="cite-bracket">[</span>34<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading3"><h3 id="Endogenous">Endogenous</h3><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=14" title="Edit section: Endogenous"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236090951"><div role="note" class="hatnote navigation-not-searchable">Main article: <a href="/wiki/Endogenous_retrovirus" title="Endogenous retrovirus">Endogenous retrovirus</a></div> <p>Endogenous retroviruses are not formally included in this classification system, and are broadly classified into three classes, on the basis of relatedness to exogenous genera: </p> <ul><li>Class I are most similar to the gammaretroviruses</li> <li>Class II are most similar to the betaretroviruses and alpharetroviruses</li> <li>Class III are most similar to the spumaviruses.</li></ul> <div class="mw-heading mw-heading2"><h2 id="Controversy">Controversy</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=15" title="Edit section: Controversy"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p>Retroviruses have been the focus of several recent claims and assertions which have been largely discredited by the science community. An initial study in 2009 seemed to make new findings which might change some of the established knowledge on this topic. However, although later research disproved some of the claims made about retroviruses, there are several controversial figures who continue to make claims which overall are considered to not have any valid basis or consensus in support of these claims.<sup id="cite_ref-35" class="reference"><a href="#cite_note-35"><span class="cite-bracket">[</span>35<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-36" class="reference"><a href="#cite_note-36"><span class="cite-bracket">[</span>36<span class="cite-bracket">]</span></a></sup><sup id="cite_ref-37" class="reference"><a href="#cite_note-37"><span class="cite-bracket">[</span>37<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Treatment">Treatment</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=16" title="Edit section: Treatment"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><a href="/wiki/Antiretroviral_drug" class="mw-redirect" title="Antiretroviral drug">Antiretroviral drugs</a> are medications for the treatment of infection by retroviruses, primarily <a href="/wiki/HIV" title="HIV">HIV</a>. Different classes of antiretroviral drugs act on different stages of the HIV <a href="/wiki/Biological_life_cycle" title="Biological life cycle">life cycle</a>. Combination of several (typically three or four) antiretroviral drugs is known as highly active antiretroviral therapy (HAART).<sup id="cite_ref-38" class="reference"><a href="#cite_note-38"><span class="cite-bracket">[</span>38<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="Treatment_of_veterinary_retroviruses">Treatment of veterinary retroviruses</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=17" title="Edit section: Treatment of veterinary retroviruses"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <p><i><a href="/wiki/Feline_leukemia_virus" title="Feline leukemia virus">Feline leukemia virus</a></i> and <i><a href="/wiki/Feline_immunodeficiency_virus" title="Feline immunodeficiency virus">Feline immunodeficiency virus</a></i> infections are treated with <a href="/wiki/Biologic_medical_product" class="mw-redirect" title="Biologic medical product">biologics</a>, including the only immunomodulator currently licensed for sale in the United States, <a href="/wiki/Lymphocyte_T-Cell_Immune_Modulator" class="mw-redirect" title="Lymphocyte T-Cell Immune Modulator">Lymphocyte T-Cell Immune Modulator</a> (LTCI).<sup id="cite_ref-39" class="reference"><a href="#cite_note-39"><span class="cite-bracket">[</span>39<span class="cite-bracket">]</span></a></sup> </p> <div class="mw-heading mw-heading2"><h2 id="References">References</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=18" title="Edit section: References"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239543626">.mw-parser-output .reflist{margin-bottom:0.5em;list-style-type:decimal}@media screen{.mw-parser-output .reflist{font-size:90%}}.mw-parser-output .reflist .references{font-size:100%;margin-bottom:0;list-style-type:inherit}.mw-parser-output .reflist-columns-2{column-width:30em}.mw-parser-output .reflist-columns-3{column-width:25em}.mw-parser-output .reflist-columns{margin-top:0.3em}.mw-parser-output .reflist-columns ol{margin-top:0}.mw-parser-output .reflist-columns li{page-break-inside:avoid;break-inside:avoid-column}.mw-parser-output .reflist-upper-alpha{list-style-type:upper-alpha}.mw-parser-output .reflist-upper-roman{list-style-type:upper-roman}.mw-parser-output .reflist-lower-alpha{list-style-type:lower-alpha}.mw-parser-output .reflist-lower-greek{list-style-type:lower-greek}.mw-parser-output .reflist-lower-roman{list-style-type:lower-roman}</style><div class="reflist"> <div class="mw-references-wrap mw-references-columns"><ol class="references"> <li id="cite_note-1"><span class="mw-cite-backlink"><b><a href="#cite_ref-1">^</a></b></span> <span class="reference-text"><style data-mw-deduplicate="TemplateStyles:r1238218222">.mw-parser-output cite.citation{font-style:inherit;word-wrap:break-word}.mw-parser-output .citation q{quotes:"\"""\"""'""'"}.mw-parser-output .citation:target{background-color:rgba(0,127,255,0.133)}.mw-parser-output .id-lock-free.id-lock-free a{background:url("//upload.wikimedia.org/wikipedia/commons/6/65/Lock-green.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-limited.id-lock-limited a,.mw-parser-output .id-lock-registration.id-lock-registration a{background:url("//upload.wikimedia.org/wikipedia/commons/d/d6/Lock-gray-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .id-lock-subscription.id-lock-subscription a{background:url("//upload.wikimedia.org/wikipedia/commons/a/aa/Lock-red-alt-2.svg")right 0.1em center/9px no-repeat}.mw-parser-output .cs1-ws-icon a{background:url("//upload.wikimedia.org/wikipedia/commons/4/4c/Wikisource-logo.svg")right 0.1em center/12px no-repeat}body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-free a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-limited a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-registration a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .id-lock-subscription a,body:not(.skin-timeless):not(.skin-minerva) .mw-parser-output .cs1-ws-icon a{background-size:contain;padding:0 1em 0 0}.mw-parser-output .cs1-code{color:inherit;background:inherit;border:none;padding:inherit}.mw-parser-output .cs1-hidden-error{display:none;color:var(--color-error,#d33)}.mw-parser-output .cs1-visible-error{color:var(--color-error,#d33)}.mw-parser-output .cs1-maint{display:none;color:#085;margin-left:0.3em}.mw-parser-output .cs1-kern-left{padding-left:0.2em}.mw-parser-output .cs1-kern-right{padding-right:0.2em}.mw-parser-output .citation .mw-selflink{font-weight:inherit}@media screen{.mw-parser-output .cs1-format{font-size:95%}html.skin-theme-clientpref-night .mw-parser-output .cs1-maint{color:#18911f}}@media screen and (prefers-color-scheme:dark){html.skin-theme-clientpref-os .mw-parser-output .cs1-maint{color:#18911f}}</style><cite class="citation web cs1"><a rel="nofollow" class="external text" href="https://ictv.global/taxonomy">"Virus Taxonomy: 2018b Release"</a>. <i>International Committee on Taxonomy of Viruses (ICTV)</i>. March 2019<span class="reference-accessdate">. Retrieved <span class="nowrap">16 March</span> 2019</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=unknown&rft.jtitle=International+Committee+on+Taxonomy+of+Viruses+%28ICTV%29&rft.atitle=Virus+Taxonomy%3A+2018b+Release&rft.date=2019-03&rft_id=https%3A%2F%2Fictv.global%2Ftaxonomy&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-2"><span class="mw-cite-backlink"><b><a href="#cite_ref-2">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite class="citation encyclopaedia cs1"><a rel="nofollow" class="external text" href="https://web.archive.org/web/20180926051822/https://en.oxforddictionaries.com/definition/retrovirus">"retrovirus"</a>. <i>Oxford English Dictionary</i>. Archived from <a rel="nofollow" class="external text" href="https://en.oxforddictionaries.com/definition/retrovirus">the original</a> on 26 September 2018<span class="reference-accessdate">. Retrieved <span class="nowrap">25 September</span> 2018</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=retrovirus&rft.btitle=Oxford+English+Dictionary&rft_id=https%3A%2F%2Fen.oxforddictionaries.com%2Fdefinition%2Fretrovirus&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-:1-3"><span class="mw-cite-backlink">^ <a href="#cite_ref-:1_3-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-:1_3-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-:1_3-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCarterSaunders2007" class="citation book cs1">Carter JB, Saunders VA (2007). <i>Virology: principles and applications</i> (1st ed.). Chichester, England: John Wiley & Sons. p. 191. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-470-02386-0" title="Special:BookSources/978-0-470-02386-0"><bdi>978-0-470-02386-0</bdi></a>. <a href="/wiki/OCLC_(identifier)" class="mw-redirect" title="OCLC (identifier)">OCLC</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/oclc/124160564">124160564</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Virology%3A+principles+and+applications&rft.place=Chichester%2C+England&rft.pages=191&rft.edition=1st&rft.pub=John+Wiley+%26+Sons&rft.date=2007&rft_id=info%3Aoclcnum%2F124160564&rft.isbn=978-0-470-02386-0&rft.aulast=Carter&rft.aufirst=John+B&rft.au=Saunders%2C+Venetia+A.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-4"><span class="mw-cite-backlink"><b><a href="#cite_ref-4">^</a></b></span> <span class="reference-text">Coffin JM, Hughes SH, Varmus HE, eds. (1997). <a href="/wiki/Retroviruses" class="mw-redirect" title="Retroviruses">Retroviruses</a>. Cold Spring Harbor Laboratory. <link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-87969-571-2" title="Special:BookSources/978-0-87969-571-2">978-0-87969-571-2</a>.</span> </li> <li id="cite_note-Miller,_A._D._2006-5"><span class="mw-cite-backlink"><b><a href="#cite_ref-Miller,_A._D._2006_5-0">^</a></b></span> <span class="reference-text">{Miller, A. D. (2006). Retroviral Vectors in Gene Therapy. Encyclopedia of Life Sciences. doi:10.1038/npg.els.0005741}</span> </li> <li id="cite_note-KurthBannert-6"><span class="mw-cite-backlink">^ <a href="#cite_ref-KurthBannert_6-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-KurthBannert_6-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFKurthBannert2010" class="citation book cs1">Kurth R, Bannert N, eds. (2010). <a rel="nofollow" class="external text" href="https://books.google.com/books?id=Vx3HbMRDj1IC"><i>Retroviruses: Molecular Biology, Genomics and Pathogenesis</i></a>. Horizon Scientific. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-1-904455-55-4" title="Special:BookSources/978-1-904455-55-4"><bdi>978-1-904455-55-4</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Retroviruses%3A+Molecular+Biology%2C+Genomics+and+Pathogenesis&rft.pub=Horizon+Scientific&rft.date=2010&rft.isbn=978-1-904455-55-4&rft_id=https%3A%2F%2Fbooks.google.com%2Fbooks%3Fid%3DVx3HbMRDj1IC&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-7"><span class="mw-cite-backlink"><b><a href="#cite_ref-7">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFZhengWeiHan2022" class="citation journal cs1">Zheng, Jialu; Wei, Yutong; Han, Guan-Zhu (1 February 2022). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC8922424">"The diversity and evolution of retroviruses: Perspectives from viral "fossils"<span class="cs1-kern-right"></span>"</a>. <i>Virologica Sinica</i>. <b>37</b> (1): 11–18. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.virs.2022.01.019">10.1016/j.virs.2022.01.019</a>. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/1995-820X">1995-820X</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC8922424">8922424</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/35234634">35234634</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Virologica+Sinica&rft.atitle=The+diversity+and+evolution+of+retroviruses%3A+Perspectives+from+viral+%22fossils%22&rft.volume=37&rft.issue=1&rft.pages=11-18&rft.date=2022-02-01&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC8922424%23id-name%3DPMC&rft.issn=1995-820X&rft_id=info%3Apmid%2F35234634&rft_id=info%3Adoi%2F10.1016%2Fj.virs.2022.01.019&rft.aulast=Zheng&rft.aufirst=Jialu&rft.au=Wei%2C+Yutong&rft.au=Han%2C+Guan-Zhu&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC8922424&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-8"><span class="mw-cite-backlink"><b><a href="#cite_ref-8">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCoffinHughesVarmus1997" class="citation book cs1">Coffin, John M.; Hughes, Stephen H.; Varmus, Harold E. (1997). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/books/NBK19382/"><i>The Place of Retroviruses in Biology</i></a>. Cold Spring Harbor Laboratory Press.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=The+Place+of+Retroviruses+in+Biology&rft.pub=Cold+Spring+Harbor+Laboratory+Press&rft.date=1997&rft.aulast=Coffin&rft.aufirst=John+M.&rft.au=Hughes%2C+Stephen+H.&rft.au=Varmus%2C+Harold+E.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fbooks%2FNBK19382%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-she-9"><span class="mw-cite-backlink"><b><a href="#cite_ref-she_9-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCoffin1992" class="citation book cs1">Coffin JM (1992). "Structure and Classification of Retroviruses". In Levy JA (ed.). <i>The Retroviridae</i>. Vol. 1 (1st ed.). New York: Plenum. p. 20. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-306-44074-8" title="Special:BookSources/978-0-306-44074-8"><bdi>978-0-306-44074-8</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=Structure+and+Classification+of+Retroviruses&rft.btitle=The+Retroviridae&rft.place=New+York&rft.pages=20&rft.edition=1st&rft.pub=Plenum&rft.date=1992&rft.isbn=978-0-306-44074-8&rft.aulast=Coffin&rft.aufirst=John+M.&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-:0-10"><span class="mw-cite-backlink">^ <a href="#cite_ref-:0_10-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-:0_10-1"><sup><i><b>b</b></i></sup></a> <a href="#cite_ref-:0_10-2"><sup><i><b>c</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFPainterCollins2019" class="citation cs2">Painter, Mark M.; Collins, Kathleen L. (1 January 2019), <a rel="nofollow" class="external text" href="http://www.sciencedirect.com/science/article/pii/B9780128012383662025">"HIV and Retroviruses"</a>, in Schmidt, Thomas M. (ed.), <i>Encyclopedia of Microbiology (Fourth Edition)</i>, Academic Press, pp. 613–628, <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fb978-0-12-801238-3.66202-5">10.1016/b978-0-12-801238-3.66202-5</a>, <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-12-811737-8" title="Special:BookSources/978-0-12-811737-8"><bdi>978-0-12-811737-8</bdi></a>, <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:188750910">188750910</a><span class="reference-accessdate">, retrieved <span class="nowrap">3 May</span> 2020</span></cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=HIV+and+Retroviruses&rft.btitle=Encyclopedia+of+Microbiology+%28Fourth+Edition%29&rft.pages=613-628&rft.pub=Academic+Press&rft.date=2019-01-01&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A188750910%23id-name%3DS2CID&rft_id=info%3Adoi%2F10.1016%2Fb978-0-12-801238-3.66202-5&rft.isbn=978-0-12-811737-8&rft.aulast=Painter&rft.aufirst=Mark+M.&rft.au=Collins%2C+Kathleen+L.&rft_id=http%3A%2F%2Fwww.sciencedirect.com%2Fscience%2Farticle%2Fpii%2FB9780128012383662025&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-11"><span class="mw-cite-backlink"><b><a href="#cite_ref-11">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFOlsonMusier-Forsyth2019" class="citation journal cs1">Olson ED, Musier-Forsyth K (February 2019). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6167211">"Retroviral Gag protein-RNA interactions: Implications for specific genomic RNA packaging and virion assembly"</a>. <i>Seminars in Cell & Developmental Biology</i>. SI: Human dendritic cells. <b>86</b>: 129–139. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.semcdb.2018.03.015">10.1016/j.semcdb.2018.03.015</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6167211">6167211</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/29580971">29580971</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Seminars+in+Cell+%26+Developmental+Biology&rft.atitle=Retroviral+Gag+protein-RNA+interactions%3A+Implications+for+specific+genomic+RNA+packaging+and+virion+assembly&rft.volume=86&rft.pages=129-139&rft.date=2019-02&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC6167211%23id-name%3DPMC&rft_id=info%3Apmid%2F29580971&rft_id=info%3Adoi%2F10.1016%2Fj.semcdb.2018.03.015&rft.aulast=Olson&rft.aufirst=ED&rft.au=Musier-Forsyth%2C+K&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC6167211&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-12"><span class="mw-cite-backlink"><b><a href="#cite_ref-12">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCoffinHughesVarmus1997" class="citation book cs1">Coffin JM, Hughes SH, Varmus HE (1997). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/books/NBK19464/"><i>Virion Proteins</i></a>. Cold Spring Harbor Laboratory Press. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-87969-571-2" title="Special:BookSources/978-0-87969-571-2"><bdi>978-0-87969-571-2</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Virion+Proteins&rft.pub=Cold+Spring+Harbor+Laboratory+Press&rft.date=1997&rft.isbn=978-0-87969-571-2&rft.aulast=Coffin&rft.aufirst=John+M.&rft.au=Hughes%2C+Stephen+H.&rft.au=Varmus%2C+Harold+E.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fbooks%2FNBK19464%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-se-13"><span class="mw-cite-backlink"><b><a href="#cite_ref-se_13-0">^</a></b></span> <span class="reference-text"><a href="#CITEREFCoffin1992">Coffin 1992</a>, pp. 26–34</span> </li> <li id="cite_note-14"><span class="mw-cite-backlink"><b><a href="#cite_ref-14">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFKimBattiniManelSitbon2004" class="citation journal cs1">Kim FJ, Battini JL, Manel N, Sitbon M (January 2004). <a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.virol.2003.09.026">"Emergence of vertebrate retroviruses and envelope capture"</a>. <i>Virology</i>. <b>318</b> (1): 183–91. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.virol.2003.09.026">10.1016/j.virol.2003.09.026</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/14972546">14972546</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Virology&rft.atitle=Emergence+of+vertebrate+retroviruses+and+envelope+capture&rft.volume=318&rft.issue=1&rft.pages=183-91&rft.date=2004-01&rft_id=info%3Adoi%2F10.1016%2Fj.virol.2003.09.026&rft_id=info%3Apmid%2F14972546&rft.aulast=Kim&rft.aufirst=FJ&rft.au=Battini%2C+JL&rft.au=Manel%2C+N&rft.au=Sitbon%2C+M&rft_id=https%3A%2F%2Fdoi.org%2F10.1016%252Fj.virol.2003.09.026&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-John_Wiley_&_Sons-15"><span class="mw-cite-backlink">^ <a href="#cite_ref-John_Wiley_&_Sons_15-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-John_Wiley_&_Sons_15-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCarterSaunders2007" class="citation book cs1">Carter JB, Saunders VA (2007). <i>Virology : principles and applications</i>. Chichester, England: John Wiley & Sons. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-470-02386-0" title="Special:BookSources/978-0-470-02386-0"><bdi>978-0-470-02386-0</bdi></a>. <a href="/wiki/OCLC_(identifier)" class="mw-redirect" title="OCLC (identifier)">OCLC</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/oclc/124160564">124160564</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Virology+%3A+principles+and+applications&rft.place=Chichester%2C+England&rft.pub=John+Wiley+%26+Sons&rft.date=2007&rft_id=info%3Aoclcnum%2F124160564&rft.isbn=978-0-470-02386-0&rft.aulast=Carter&rft.aufirst=John+B&rft.au=Saunders%2C+Venetia+A&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-16"><span class="mw-cite-backlink"><b><a href="#cite_ref-16">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFChampouxSchultz2009" class="citation journal cs1">Champoux JJ, Schultz SJ (June 2009). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2464458">"RNase H Activity: Structure, Specificity, and Function in Reverse Transcription"</a>. <i>The FEBS Journal</i>. <b>134</b> (1–2): 86–103. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.virusres.2007.12.007">10.1016/j.virusres.2007.12.007</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC2464458">2464458</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/18261820">18261820</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+FEBS+Journal&rft.atitle=RNase+H+Activity%3A+Structure%2C+Specificity%2C+and+Function+in+Reverse+Transcription&rft.volume=134&rft.issue=1%E2%80%932&rft.pages=86-103&rft.date=2009-06&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC2464458%23id-name%3DPMC&rft_id=info%3Apmid%2F18261820&rft_id=info%3Adoi%2F10.1016%2Fj.virusres.2007.12.007&rft.aulast=Champoux&rft.aufirst=JJ&rft.au=Schultz%2C+SJ&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC2464458&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-17"><span class="mw-cite-backlink"><b><a href="#cite_ref-17">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMoellingBroeckerKerrigan2014" class="citation book cs1">Moelling K, Broecker F, Kerrigan JE (2014). "RNase H: specificity, mechanisms of action, and antiviral target". <i>Human Retroviruses</i>. Methods in Molecular Biology. Vol. 1087. pp. 71–84. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1007%2F978-1-62703-670-2_7">10.1007/978-1-62703-670-2_7</a>. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-1-62703-669-6" title="Special:BookSources/978-1-62703-669-6"><bdi>978-1-62703-669-6</bdi></a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/24158815">24158815</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=RNase+H%3A+specificity%2C+mechanisms+of+action%2C+and+antiviral+target&rft.btitle=Human+Retroviruses&rft.series=Methods+in+Molecular+Biology&rft.pages=71-84&rft.date=2014&rft_id=info%3Apmid%2F24158815&rft_id=info%3Adoi%2F10.1007%2F978-1-62703-670-2_7&rft.isbn=978-1-62703-669-6&rft.aulast=Moelling&rft.aufirst=K&rft.au=Broecker%2C+F&rft.au=Kerrigan%2C+JE&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-pmid26800882-18"><span class="mw-cite-backlink">^ <a href="#cite_ref-pmid26800882_18-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-pmid26800882_18-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFVargiuRodriguez-ToméSperberCadeddu2016" class="citation journal cs1">Vargiu L, Rodriguez-Tomé P, Sperber GO, Cadeddu M, Grandi N, Blikstad V, et al. (January 2016). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4724089">"Classification and characterization of human endogenous retroviruses; mosaic forms are common"</a>. <i>Retrovirology</i>. <b>13</b>: 7. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1186%2Fs12977-015-0232-y">10.1186/s12977-015-0232-y</a></span>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4724089">4724089</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/26800882">26800882</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Retrovirology&rft.atitle=Classification+and+characterization+of+human+endogenous+retroviruses%3B+mosaic+forms+are+common&rft.volume=13&rft.pages=7&rft.date=2016-01&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC4724089%23id-name%3DPMC&rft_id=info%3Apmid%2F26800882&rft_id=info%3Adoi%2F10.1186%2Fs12977-015-0232-y&rft.aulast=Vargiu&rft.aufirst=L&rft.au=Rodriguez-Tom%C3%A9%2C+P&rft.au=Sperber%2C+GO&rft.au=Cadeddu%2C+M&rft.au=Grandi%2C+N&rft.au=Blikstad%2C+V&rft.au=Tramontano%2C+E&rft.au=Blomberg%2C+J&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC4724089&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-19"><span class="mw-cite-backlink"><b><a href="#cite_ref-19">^</a></b></span> <span class="reference-text">Peters, P. J., Marston, B. J., Weidle, P. J., & Brooks, J. T. (2013). Human Immunodeficiency Virus Infection. Hunter's Tropical Medicine and Emerging Infectious Disease, 217–247. doi:10.1016/b978-1-4160-4390-4.00027-8</span> </li> <li id="cite_note-20"><span class="mw-cite-backlink"><b><a href="#cite_ref-20">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCoffinHughesVarmus1997" class="citation book cs1">Coffin JM, Hughes SH, Varmus HE (1997). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/books/NBK19370/">"Genetic Organization"</a>. <i>Retroviruses</i>. Cold Spring Harbor Laboratory Press. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-87969-571-2" title="Special:BookSources/978-0-87969-571-2"><bdi>978-0-87969-571-2</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=bookitem&rft.atitle=Genetic+Organization&rft.btitle=Retroviruses&rft.pub=Cold+Spring+Harbor+Laboratory+Press&rft.date=1997&rft.isbn=978-0-87969-571-2&rft.aulast=Coffin&rft.aufirst=John+M.&rft.au=Hughes%2C+Stephen+H.&rft.au=Varmus%2C+Harold+E.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fbooks%2FNBK19370%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-21"><span class="mw-cite-backlink"><b><a href="#cite_ref-21">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFBelshawPereiraKatzourakisTalbot2004" class="citation journal cs1">Belshaw R, Pereira V, Katzourakis A, Talbot G, Paces J, Burt A, Tristem M (April 2004). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC387345">"Long-term reinfection of the human genome by endogenous retroviruses"</a>. <i>Proceedings of the National Academy of Sciences of the United States of America</i>. <b>101</b> (14): 4894–9. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2004PNAS..101.4894B">2004PNAS..101.4894B</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1073%2Fpnas.0307800101">10.1073/pnas.0307800101</a></span>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC387345">387345</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/15044706">15044706</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Proceedings+of+the+National+Academy+of+Sciences+of+the+United+States+of+America&rft.atitle=Long-term+reinfection+of+the+human+genome+by+endogenous+retroviruses&rft.volume=101&rft.issue=14&rft.pages=4894-9&rft.date=2004-04&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC387345%23id-name%3DPMC&rft_id=info%3Apmid%2F15044706&rft_id=info%3Adoi%2F10.1073%2Fpnas.0307800101&rft_id=info%3Abibcode%2F2004PNAS..101.4894B&rft.aulast=Belshaw&rft.aufirst=R&rft.au=Pereira%2C+V&rft.au=Katzourakis%2C+A&rft.au=Talbot%2C+G&rft.au=Paces%2C+J&rft.au=Burt%2C+A&rft.au=Tristem%2C+M&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC387345&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-22"><span class="mw-cite-backlink"><b><a href="#cite_ref-22">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMedstrandvan_de_LagemaatDunnLandry2005" class="citation journal cs1">Medstrand P, van de Lagemaat LN, Dunn CA, Landry JR, Svenback D, Mager DL (2005). "Impact of transposable elements on the evolution of mammalian gene regulation". <i>Cytogenetic and Genome Research</i>. <b>110</b> (1–4): 342–52. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1159%2F000084966">10.1159/000084966</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/16093686">16093686</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:25307890">25307890</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Cytogenetic+and+Genome+Research&rft.atitle=Impact+of+transposable+elements+on+the+evolution+of+mammalian+gene+regulation&rft.volume=110&rft.issue=1%E2%80%934&rft.pages=342-52&rft.date=2005&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A25307890%23id-name%3DS2CID&rft_id=info%3Apmid%2F16093686&rft_id=info%3Adoi%2F10.1159%2F000084966&rft.aulast=Medstrand&rft.aufirst=P&rft.au=van+de+Lagemaat%2C+LN&rft.au=Dunn%2C+CA&rft.au=Landry%2C+JR&rft.au=Svenback%2C+D&rft.au=Mager%2C+DL&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-23"><span class="mw-cite-backlink"><b><a href="#cite_ref-23">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFSvarovskaiaCheslockZhangHu2003" class="citation journal cs1">Svarovskaia ES, Cheslock SR, Zhang WH, Hu WS, Pathak VK (January 2003). <a rel="nofollow" class="external text" href="https://doi.org/10.2741%2F957">"Retroviral mutation rates and reverse transcriptase fidelity"</a>. <i>Frontiers in Bioscience</i>. <b>8</b> (1–3): d117–34. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.2741%2F957">10.2741/957</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/12456349">12456349</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Frontiers+in+Bioscience&rft.atitle=Retroviral+mutation+rates+and+reverse+transcriptase+fidelity&rft.volume=8&rft.issue=1%E2%80%933&rft.pages=d117-34&rft.date=2003-01&rft_id=info%3Adoi%2F10.2741%2F957&rft_id=info%3Apmid%2F12456349&rft.aulast=Svarovskaia&rft.aufirst=ES&rft.au=Cheslock%2C+SR&rft.au=Zhang%2C+WH&rft.au=Hu%2C+WS&rft.au=Pathak%2C+VK&rft_id=https%3A%2F%2Fdoi.org%2F10.2741%252F957&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-Rawson2018-24"><span class="mw-cite-backlink">^ <a href="#cite_ref-Rawson2018_24-0"><sup><i><b>a</b></i></sup></a> <a href="#cite_ref-Rawson2018_24-1"><sup><i><b>b</b></i></sup></a></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFRawsonNikolaitchikKeelePathak2018" class="citation journal cs1">Rawson JM, Nikolaitchik OA, Keele BF, Pathak VK, Hu WS (November 2018). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6237782">"Recombination is required for efficient HIV-1 replication and the maintenance of viral genome integrity"</a>. <i>Nucleic Acids Research</i>. <b>46</b> (20): 10535–10545. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1093%2Fnar%2Fgky910">10.1093/nar/gky910</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6237782">6237782</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/30307534">30307534</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nucleic+Acids+Research&rft.atitle=Recombination+is+required+for+efficient+HIV-1+replication+and+the+maintenance+of+viral+genome+integrity&rft.volume=46&rft.issue=20&rft.pages=10535-10545&rft.date=2018-11&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC6237782%23id-name%3DPMC&rft_id=info%3Apmid%2F30307534&rft_id=info%3Adoi%2F10.1093%2Fnar%2Fgky910&rft.aulast=Rawson&rft.aufirst=JM&rft.au=Nikolaitchik%2C+OA&rft.au=Keele%2C+BF&rft.au=Pathak%2C+VK&rft.au=Hu%2C+WS&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC6237782&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-pmid26691546-25"><span class="mw-cite-backlink"><b><a href="#cite_ref-pmid26691546_25-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCromerGrimmSchlubMak2016" class="citation journal cs1">Cromer D, Grimm AJ, Schlub TE, Mak J, Davenport MP (January 2016). <a rel="nofollow" class="external text" href="https://doi.org/10.1097%2FQAD.0000000000000936">"Estimating the in-vivo HIV template switching and recombination rate"</a>. <i>AIDS</i>. <b>30</b> (2): 185–92. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.1097%2FQAD.0000000000000936">10.1097/QAD.0000000000000936</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/26691546">26691546</a>. <a href="/wiki/S2CID_(identifier)" class="mw-redirect" title="S2CID (identifier)">S2CID</a> <a rel="nofollow" class="external text" href="https://api.semanticscholar.org/CorpusID:20086739">20086739</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=AIDS&rft.atitle=Estimating+the+in-vivo+HIV+template+switching+and+recombination+rate&rft.volume=30&rft.issue=2&rft.pages=185-92&rft.date=2016-01&rft_id=https%3A%2F%2Fapi.semanticscholar.org%2FCorpusID%3A20086739%23id-name%3DS2CID&rft_id=info%3Apmid%2F26691546&rft_id=info%3Adoi%2F10.1097%2FQAD.0000000000000936&rft.aulast=Cromer&rft.aufirst=D&rft.au=Grimm%2C+AJ&rft.au=Schlub%2C+TE&rft.au=Mak%2C+J&rft.au=Davenport%2C+MP&rft_id=https%3A%2F%2Fdoi.org%2F10.1097%252FQAD.0000000000000936&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-26"><span class="mw-cite-backlink"><b><a href="#cite_ref-26">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFJolly2011" class="citation journal cs1">Jolly C (March 2011). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3053447">"Cell-to-cell transmission of retroviruses: Innate immunity and interferon-induced restriction factors"</a>. <i>Virology</i>. <b>411</b> (2): 251–9. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.virol.2010.12.031">10.1016/j.virol.2010.12.031</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3053447">3053447</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/21247613">21247613</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Virology&rft.atitle=Cell-to-cell+transmission+of+retroviruses%3A+Innate+immunity+and+interferon-induced+restriction+factors&rft.volume=411&rft.issue=2&rft.pages=251-9&rft.date=2011-03&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3053447%23id-name%3DPMC&rft_id=info%3Apmid%2F21247613&rft_id=info%3Adoi%2F10.1016%2Fj.virol.2010.12.031&rft.aulast=Jolly&rft.aufirst=C&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC3053447&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-27"><span class="mw-cite-backlink"><b><a href="#cite_ref-27">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFMacLachlanDubovi2011" class="citation book cs1">MacLachlan, N. James; Dubovi, Edward J. (2011). <a rel="nofollow" class="external text" href="https://www.sciencedirect.com/science/article/pii/B9780123751584000146"><i>Fenner's Veterinary Virology</i></a> (Fourth ed.). Academic Press. p. 250. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-12-375159-1" title="Special:BookSources/978-0-12-375159-1"><bdi>978-0-12-375159-1</bdi></a><span class="reference-accessdate">. Retrieved <span class="nowrap">6 May</span> 2020</span>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Fenner%27s+Veterinary+Virology&rft.pages=250&rft.edition=Fourth&rft.pub=Academic+Press&rft.date=2011&rft.isbn=978-0-12-375159-1&rft.aulast=MacLachlan&rft.aufirst=N.+James&rft.au=Dubovi%2C+Edward+J.&rft_id=https%3A%2F%2Fwww.sciencedirect.com%2Fscience%2Farticle%2Fpii%2FB9780123751584000146&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-Aiewsakun-28"><span class="mw-cite-backlink"><b><a href="#cite_ref-Aiewsakun_28-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFAiewsakunKatzourakis2017" class="citation journal cs1">Aiewsakun P, Katzourakis A (January 2017). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5512871">"Marine origin of retroviruses in the early Palaeozoic Era"</a>. <i>Nature Communications</i>. <b>8</b>: 13954. <a href="/wiki/Bibcode_(identifier)" class="mw-redirect" title="Bibcode (identifier)">Bibcode</a>:<a rel="nofollow" class="external text" href="https://ui.adsabs.harvard.edu/abs/2017NatCo...813954A">2017NatCo...813954A</a>. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1038%2Fncomms13954">10.1038/ncomms13954</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5512871">5512871</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/28071651">28071651</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Nature+Communications&rft.atitle=Marine+origin+of+retroviruses+in+the+early+Palaeozoic+Era&rft.volume=8&rft.pages=13954&rft.date=2017-01&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5512871%23id-name%3DPMC&rft_id=info%3Apmid%2F28071651&rft_id=info%3Adoi%2F10.1038%2Fncomms13954&rft_id=info%3Abibcode%2F2017NatCo...813954A&rft.aulast=Aiewsakun&rft.aufirst=P&rft.au=Katzourakis%2C+A&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5512871&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-Desportm-29"><span class="mw-cite-backlink"><b><a href="#cite_ref-Desportm_29-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFDesport2010" class="citation book cs1">Desport M, ed. (2010). <i>Lentiviruses and Macrophages: Molecular and Cellular Interactions</i>. Caister Academic. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-1-904455-60-8" title="Special:BookSources/978-1-904455-60-8"><bdi>978-1-904455-60-8</bdi></a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Lentiviruses+and+Macrophages%3A+Molecular+and+Cellular+Interactions&rft.pub=Caister+Academic&rft.date=2010&rft.isbn=978-1-904455-60-8&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-30"><span class="mw-cite-backlink"><b><a href="#cite_ref-30">^</a></b></span> <span class="reference-text">Ross, S. R. (2018). Cellular Immune Responses to Retroviruses. In Retrovirus-Cell Interactions (pp. 401–420). Elsevier. <a rel="nofollow" class="external free" href="https://doi.org/10.1016/B978-0-12-811185-7.00011-X">https://doi.org/10.1016/B978-0-12-811185-7.00011-X</a></span> </li> <li id="cite_note-31"><span class="mw-cite-backlink"><b><a href="#cite_ref-31">^</a></b></span> <span class="reference-text">Burrell, C. J., Howard, C. R., & Murphy, F. A. (2017). Retroviruses. In Fenner and White's Medical Virology (pp. 317–344). Elsevier. <a rel="nofollow" class="external free" href="https://doi.org/10.1016/b978-0-12-375156-0.00023-0">https://doi.org/10.1016/b978-0-12-375156-0.00023-0</a></span> </li> <li id="cite_note-32"><span class="mw-cite-backlink"><b><a href="#cite_ref-32">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://ictv.global/taxonomy">ICTV Taxonomy</a> Browser</span> </li> <li id="cite_note-Lauber2017-33"><span class="mw-cite-backlink"><b><a href="#cite_ref-Lauber2017_33-0">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFLauberSeitzMatteiSuh2017" class="citation journal cs1">Lauber C, Seitz S, Mattei S, Suh A, Beck J, Herstein J, et al. (September 2017). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5604429">"Deciphering the Origin and Evolution of Hepatitis B Viruses by Means of a Family of Non-enveloped Fish Viruses"</a>. <i>Cell Host & Microbe</i>. <b>22</b> (3): 387–399.e6. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1016%2Fj.chom.2017.07.019">10.1016/j.chom.2017.07.019</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5604429">5604429</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/28867387">28867387</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Cell+Host+%26+Microbe&rft.atitle=Deciphering+the+Origin+and+Evolution+of+Hepatitis+B+Viruses+by+Means+of+a+Family+of+Non-enveloped+Fish+Viruses&rft.volume=22&rft.issue=3&rft.pages=387-399.e6&rft.date=2017-09&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5604429%23id-name%3DPMC&rft_id=info%3Apmid%2F28867387&rft_id=info%3Adoi%2F10.1016%2Fj.chom.2017.07.019&rft.aulast=Lauber&rft.aufirst=C&rft.au=Seitz%2C+S&rft.au=Mattei%2C+S&rft.au=Suh%2C+A&rft.au=Beck%2C+J&rft.au=Herstein%2C+J&rft.au=B%C3%B6rold%2C+J&rft.au=Salzburger%2C+W&rft.au=Kaderali%2C+L&rft.au=Briggs%2C+JA&rft.au=Bartenschlager%2C+R&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5604429&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span> and <a rel="nofollow" class="external text" href="https://www.cell.com/cms/attachment/2119344051/2092493318/mmc1.pdf">PDF</a></span> </li> <li id="cite_note-34"><span class="mw-cite-backlink"><b><a href="#cite_ref-34">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFKrupovicBlombergCoffinDasgupta2018" class="citation journal cs1">Krupovic M, Blomberg J, Coffin JM, Dasgupta I, Fan H, Geering AD, et al. (June 2018). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5974489">"Ortervirales: New Virus Order Unifying Five Families of Reverse-Transcribing Viruses"</a>. <i>Journal of Virology</i>. <b>92</b> (12). <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1128%2FJVI.00515-18">10.1128/JVI.00515-18</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5974489">5974489</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/29618642">29618642</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Journal+of+Virology&rft.atitle=Ortervirales%3A+New+Virus+Order+Unifying+Five+Families+of+Reverse-Transcribing+Viruses&rft.volume=92&rft.issue=12&rft.date=2018-06&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5974489%23id-name%3DPMC&rft_id=info%3Apmid%2F29618642&rft_id=info%3Adoi%2F10.1128%2FJVI.00515-18&rft.aulast=Krupovic&rft.aufirst=M&rft.au=Blomberg%2C+J&rft.au=Coffin%2C+JM&rft.au=Dasgupta%2C+I&rft.au=Fan%2C+H&rft.au=Geering%2C+AD&rft.au=Gifford%2C+R&rft.au=Harrach%2C+B&rft.au=Hull%2C+R&rft.au=Johnson%2C+W&rft.au=Kreuze%2C+JF&rft.au=Lindemann%2C+D&rft.au=Llorens%2C+C&rft.au=Lockhart%2C+B&rft.au=Mayer%2C+J&rft.au=Muller%2C+E&rft.au=Olszewski%2C+NE&rft.au=Pappu%2C+HR&rft.au=Pooggin%2C+MM&rft.au=Richert-P%C3%B6ggeler%2C+KR&rft.au=Sabanadzovic%2C+S&rft.au=Sanfa%C3%A7on%2C+H&rft.au=Schoelz%2C+JE&rft.au=Seal%2C+S&rft.au=Stavolone%2C+L&rft.au=Stoye%2C+JP&rft.au=Teycheney%2C+PY&rft.au=Tristem%2C+M&rft.au=Koonin%2C+EV&rft.au=Kuhn%2C+JH&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC5974489&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-35"><span class="mw-cite-backlink"><b><a href="#cite_ref-35">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://www.science.org/content/article/fact-checking-judy-mikovits-controversial-virologist-attacking-anthony-fauci-viral">Fact-checking Judy Mikovits, the controversial virologist attacking Anthony Fauci in a viral conspiracy video</a>, By Martin Enserink, Jon Cohen, May 8, 2020, accessed June 17, 2022, science.org website.</span> </li> <li id="cite_note-36"><span class="mw-cite-backlink"><b><a href="#cite_ref-36">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFNeilCampbell2020" class="citation journal cs1">Neil, Stuart J.D.; Campbell, Edward M. (2020). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7398426">"Fake Science: XMRV, COVID-19, and the Toxic Legacy of Dr. Judy Mikovits"</a>. <i>AIDS Research and Human Retroviruses</i>. <b>36</b> (7): 545–549. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1089%2Faid.2020.0095">10.1089/aid.2020.0095</a>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7398426">7398426</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/32414291">32414291</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=AIDS+Research+and+Human+Retroviruses&rft.atitle=Fake+Science%3A+XMRV%2C+COVID-19%2C+and+the+Toxic+Legacy+of+Dr.+Judy+Mikovits&rft.volume=36&rft.issue=7&rft.pages=545-549&rft.date=2020&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC7398426%23id-name%3DPMC&rft_id=info%3Apmid%2F32414291&rft_id=info%3Adoi%2F10.1089%2Faid.2020.0095&rft.aulast=Neil&rft.aufirst=Stuart+J.D.&rft.au=Campbell%2C+Edward+M.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC7398426&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-37"><span class="mw-cite-backlink"><b><a href="#cite_ref-37">^</a></b></span> <span class="reference-text"><a rel="nofollow" class="external text" href="https://www.nytimes.com/2020/05/09/technology/plandemic-judy-mikovitz-coronavirus-disinformation.html">Virus Conspiracists Elevate a New Champion</a>, by <a href="/wiki/Davey_Alba" title="Davey Alba">Davey Alba</a>, May 9, 2020, nytimes.com</span> </li> <li id="cite_note-38"><span class="mw-cite-backlink"><b><a href="#cite_ref-38">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFRutherfordSanganiKennedy2003" class="citation journal cs1">Rutherford GW, Sangani PR, Kennedy GE (2003). "Three- or four- versus two-drug antiretroviral maintenance regimens for HIV infection". <i>The Cochrane Database of Systematic Reviews</i> (4): CD002037. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<a rel="nofollow" class="external text" href="https://doi.org/10.1002%2F14651858.CD002037">10.1002/14651858.CD002037</a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/14583945">14583945</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+Cochrane+Database+of+Systematic+Reviews&rft.atitle=Three-+or+four-+versus+two-drug+antiretroviral+maintenance+regimens+for+HIV+infection&rft.issue=4&rft.pages=CD002037&rft.date=2003&rft_id=info%3Adoi%2F10.1002%2F14651858.CD002037&rft_id=info%3Apmid%2F14583945&rft.aulast=Rutherford&rft.aufirst=GW&rft.au=Sangani%2C+PR&rft.au=Kennedy%2C+GE&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> <li id="cite_note-39"><span class="mw-cite-backlink"><b><a href="#cite_ref-39">^</a></b></span> <span class="reference-text"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFGingerich2008" class="citation journal cs1">Gingerich DA (2008). <a rel="nofollow" class="external text" href="http://jarvm.com/articles/Vol6Iss2/Vol6Iss2Gingerich61-68.pdf">"Lymphocyte T-cell immunomodulator (LTCI): Review of the immunopharmacology of a new biologic"</a> <span class="cs1-format">(PDF)</span>. <i>International Journal of Applied Research in Veterinary Medicine</i>. <b>6</b> (2): 61–68. <a href="/wiki/ISSN_(identifier)" class="mw-redirect" title="ISSN (identifier)">ISSN</a> <a rel="nofollow" class="external text" href="https://search.worldcat.org/issn/1559-470X">1559-470X</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=International+Journal+of+Applied+Research+in+Veterinary+Medicine&rft.atitle=Lymphocyte+T-cell+immunomodulator+%28LTCI%29%3A+Review+of+the+immunopharmacology+of+a+new+biologic&rft.volume=6&rft.issue=2&rft.pages=61-68&rft.date=2008&rft.issn=1559-470X&rft.aulast=Gingerich&rft.aufirst=Daniel+A.&rft_id=http%3A%2F%2Fjarvm.com%2Farticles%2FVol6Iss2%2FVol6Iss2Gingerich61-68.pdf&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></span> </li> </ol></div></div> <div class="mw-heading mw-heading2"><h2 id="Further_reading">Further reading</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=19" title="Edit section: Further reading"><span>edit</span></a><span class="mw-editsection-bracket">]</span></span></div> <style data-mw-deduplicate="TemplateStyles:r1239549316">.mw-parser-output .refbegin{margin-bottom:0.5em}.mw-parser-output .refbegin-hanging-indents>ul{margin-left:0}.mw-parser-output .refbegin-hanging-indents>ul>li{margin-left:0;padding-left:3.2em;text-indent:-3.2em}.mw-parser-output .refbegin-hanging-indents ul,.mw-parser-output .refbegin-hanging-indents ul li{list-style:none}@media(max-width:720px){.mw-parser-output .refbegin-hanging-indents>ul>li{padding-left:1.6em;text-indent:-1.6em}}.mw-parser-output .refbegin-columns{margin-top:0.3em}.mw-parser-output .refbegin-columns ul{margin-top:0}.mw-parser-output .refbegin-columns li{page-break-inside:avoid;break-inside:avoid-column}@media screen{.mw-parser-output .refbegin{font-size:90%}}</style><div class="refbegin" style=""> <ul><li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFCoffinHughesVarmus1997" class="citation book cs1">Coffin JM, Hughes SH, Varmus HE (1997). Coffin JM, Hughes SH, Varmus HE (eds.). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/books/NBK19376/"><i>Retroviruses</i></a>. Cold Spring Harbor Laboratory. <a href="/wiki/ISBN_(identifier)" class="mw-redirect" title="ISBN (identifier)">ISBN</a> <a href="/wiki/Special:BookSources/978-0-87969-571-2" title="Special:BookSources/978-0-87969-571-2"><bdi>978-0-87969-571-2</bdi></a>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/21433340">21433340</a>. NBK19376.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Abook&rft.genre=book&rft.btitle=Retroviruses&rft.pub=Cold+Spring+Harbor+Laboratory&rft.date=1997&rft_id=info%3Apmid%2F21433340&rft.isbn=978-0-87969-571-2&rft.aulast=Coffin&rft.aufirst=J.+M.&rft.au=Hughes%2C+S.+H.&rft.au=Varmus%2C+H.+E.&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fbooks%2FNBK19376%2F&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFSpecter2007" class="citation news cs1">Specter M (3 December 2007). <a rel="nofollow" class="external text" href="http://www.newyorker.com/reporting/2007/12/03/071203fa_fact_specter">"Annals of Science: Darwin's Surprise"</a>. <i><a href="/wiki/The_New_Yorker" title="The New Yorker">The New Yorker</a></i>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=The+New+Yorker&rft.atitle=Annals+of+Science%3A+Darwin%27s+Surprise&rft.date=2007-12-03&rft.aulast=Specter&rft.aufirst=Michael&rft_id=http%3A%2F%2Fwww.newyorker.com%2Freporting%2F2007%2F12%2F03%2F071203fa_fact_specter&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></li> <li><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1238218222"><cite id="CITEREFDostálkováVokatáKaufmanUlbrich2021" class="citation journal cs1">Dostálková, A; Vokatá, B; Kaufman, F; Ulbrich, P; Ruml, T; Rumlová, M (2021). <a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7831069">"Effect of Small Polyanions on In Vitro Assembly of Selected Members of Alpha-, Beta- and Gammaretroviruses"</a>. <i>Viruses</i>. <b>13</b> (1): 129. <a href="/wiki/Doi_(identifier)" class="mw-redirect" title="Doi (identifier)">doi</a>:<span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://doi.org/10.3390%2Fv13010129">10.3390/v13010129</a></span>. <a href="/wiki/PMC_(identifier)" class="mw-redirect" title="PMC (identifier)">PMC</a> <span class="id-lock-free" title="Freely accessible"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7831069">7831069</a></span>. <a href="/wiki/PMID_(identifier)" class="mw-redirect" title="PMID (identifier)">PMID</a> <a rel="nofollow" class="external text" href="https://pubmed.ncbi.nlm.nih.gov/33477490">33477490</a>.</cite><span title="ctx_ver=Z39.88-2004&rft_val_fmt=info%3Aofi%2Ffmt%3Akev%3Amtx%3Ajournal&rft.genre=article&rft.jtitle=Viruses&rft.atitle=Effect+of+Small+Polyanions+on+In+Vitro+Assembly+of+Selected+Members+of+Alpha-%2C+Beta-+and+Gammaretroviruses&rft.volume=13&rft.issue=1&rft.pages=129&rft.date=2021&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC7831069%23id-name%3DPMC&rft_id=info%3Apmid%2F33477490&rft_id=info%3Adoi%2F10.3390%2Fv13010129&rft.aulast=Dost%C3%A1lkov%C3%A1&rft.aufirst=A&rft.au=Vokat%C3%A1%2C+B&rft.au=Kaufman%2C+F&rft.au=Ulbrich%2C+P&rft.au=Ruml%2C+T&rft.au=Rumlov%C3%A1%2C+M&rft_id=https%3A%2F%2Fwww.ncbi.nlm.nih.gov%2Fpmc%2Farticles%2FPMC7831069&rfr_id=info%3Asid%2Fen.wikipedia.org%3ARetrovirus" class="Z3988"></span></li></ul> </div> <div class="mw-heading mw-heading2"><h2 id="External_links">External links</h2><span class="mw-editsection"><span class="mw-editsection-bracket">[</span><a href="/w/index.php?title=Retrovirus&action=edit&section=20" title="Edit section: 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viruses"</i>)</li> <li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20090405172319/http://www.1lec.com/Microbiology/Replication%20Cycle%20of%20a%20Retrovirus/index.html">Retrovirus Animation</a> (Flash Required)</li> <li><a rel="nofollow" class="external text" href="https://retrovirology.biomedcentral.com/">Retrovirology</a> Scientific journal</li> <li><a rel="nofollow" class="external text" href="https://web.archive.org/web/20130329004127/http://users.rcn.com/jkimball.ma.ultranet/BiologyPages/R/Retroviruses.html">Retrovirus life cycle chapter</a> From Kimball's <i>Biology</i> (online biology textbook pages)</li></ul> <div class="navbox-styles"><style data-mw-deduplicate="TemplateStyles:r1129693374">.mw-parser-output .hlist dl,.mw-parser-output .hlist ol,.mw-parser-output .hlist ul{margin:0;padding:0}.mw-parser-output .hlist dd,.mw-parser-output .hlist dt,.mw-parser-output .hlist li{margin:0;display:inline}.mw-parser-output .hlist.inline,.mw-parser-output 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<li><a href="/wiki/Virology" title="Virology">Virology</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Components</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Capsid" title="Capsid">Capsid</a></li> <li><a href="/wiki/Viral_envelope" title="Viral envelope">Viral envelope</a></li> <li><a href="/wiki/Viral_protein" title="Viral protein">Viral protein</a></li></ul> </div></td><td class="noviewer navbox-image" rowspan="5" style="width:1px;padding:0 0 0 2px"><div><span typeof="mw:File"><a href="/wiki/File:Rotavirus_Reconstruction.jpg" class="mw-file-description"><img src="//upload.wikimedia.org/wikipedia/commons/thumb/f/fa/Rotavirus_Reconstruction.jpg/100px-Rotavirus_Reconstruction.jpg" decoding="async" width="100" height="88" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/f/fa/Rotavirus_Reconstruction.jpg/150px-Rotavirus_Reconstruction.jpg 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/f/fa/Rotavirus_Reconstruction.jpg/200px-Rotavirus_Reconstruction.jpg 2x" data-file-width="1986" data-file-height="1739" /></a></span></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Viral_life_cycle" title="Viral life cycle">Viral life cycle</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Viral_entry" title="Viral entry">Viral entry</a></li> <li><a href="/wiki/Viral_replication" title="Viral replication">Viral replication</a></li> <li><a href="/wiki/Viral_shedding" title="Viral shedding">Viral shedding</a></li> <li><a href="/wiki/Viroplasm" title="Viroplasm">Viroplasm</a></li> <li><a href="/wiki/Virus_latency" title="Virus latency">Virus latency</a></li> <li><a href="/wiki/Lytic_cycle" title="Lytic cycle">Lytic cycle</a></li> <li><a href="/wiki/Lysogenic_cycle" title="Lysogenic cycle">Lysogenic cycle</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Genetics</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Antigenic_drift" title="Antigenic drift">Antigenic drift</a></li> <li><a href="/wiki/Antigenic_shift" title="Antigenic shift">Antigenic shift</a></li> <li><a href="/wiki/Phenotype_mixing" title="Phenotype mixing">Phenotype mixing</a></li> <li><a href="/wiki/Reassortment" title="Reassortment">Reassortment</a></li> <li><a href="/wiki/Viral_evolution" title="Viral evolution">Viral evolution</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">By host</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Animal_virus" title="Animal virus">Animal virus</a></li> <li><a href="/wiki/Bacteriophage" title="Bacteriophage">Bacteriophage</a></li> <li><a href="/wiki/Virophage" title="Virophage">Virophage</a></li> <li><a href="/wiki/Human_virome" title="Human virome">Human virome</a></li> <li><a href="/wiki/Mycovirus" title="Mycovirus">Mycovirus</a></li> <li><a href="/wiki/Plant_virus" title="Plant virus">Plant virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Other</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Antiviral_drug" title="Antiviral drug">Antiviral drug</a></li> <li><a href="/wiki/Giant_virus" title="Giant virus">Giant virus</a></li> <li><a href="/wiki/Helper_virus" title="Helper virus">Helper virus</a></li> <li><a href="/wiki/Viral_vector" title="Viral vector">Viral vector</a> <ul><li><a href="/wiki/Helper_dependent_virus" title="Helper dependent virus">Helper dependent virus</a></li></ul></li> <li><a href="/wiki/Laboratory_diagnosis_of_viral_infections" title="Laboratory diagnosis of viral infections">Laboratory diagnosis of viral infections</a></li> <li><a href="/wiki/Marine_viruses" title="Marine viruses">Marine viruses</a></li> <li><a href="/wiki/Neurotropic_virus" title="Neurotropic virus">Neurotropic virus</a></li> <li><a href="/wiki/Oncovirus" title="Oncovirus">Oncovirus</a></li> <li><a href="/wiki/Satellite_(biology)" title="Satellite (biology)">Satellites</a></li> <li><a href="/wiki/Viral_disease" title="Viral disease">Viral disease</a></li> <li><a href="/wiki/Viral_load" title="Viral load">Viral load</a></li> <li><a href="/wiki/Virus-like_particle" title="Virus-like particle">Virus-like particle</a></li> <li><a href="/wiki/Virus_classification" title="Virus classification">Virus classification</a></li> <li><a href="/wiki/Virus_quantification" title="Virus quantification">Virus quantification</a></li> <li><a href="/wiki/Virome" title="Virome">Virome</a></li> <li><a href="/wiki/Virosphere" title="Virosphere">Virosphere</a></li></ul> </div></td></tr><tr><td class="navbox-abovebelow" colspan="3"><div> <ul><li><span class="noviewer" typeof="mw:File"><a href="/wiki/File:Symbol_portal_class.svg" class="mw-file-description" title="Portal"><img alt="" src="//upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/16px-Symbol_portal_class.svg.png" decoding="async" width="16" height="16" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/23px-Symbol_portal_class.svg.png 1.5x, //upload.wikimedia.org/wikipedia/en/thumb/e/e2/Symbol_portal_class.svg/31px-Symbol_portal_class.svg.png 2x" data-file-width="180" data-file-height="185" /></a></span> <b><a href="/wiki/Portal:Viruses" title="Portal:Viruses">Portal</a></b></li> <li><span class="noviewer" typeof="mw:File"><span 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rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Virus:_Reverse_transcribing_viruses_(Revtraviricetes)" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Retroviruses" title="Template:Retroviruses"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Retroviruses" title="Template talk:Retroviruses"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Retroviruses" title="Special:EditPage/Template:Retroviruses"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Virus:_Reverse_transcribing_viruses_(Revtraviricetes)" style="font-size:114%;margin:0 4em"><a href="/wiki/Virus" title="Virus">Virus</a>: <a href="/wiki/Reverse_transcriptase" title="Reverse transcriptase">Reverse transcribing</a> viruses (<i><a href="/wiki/Revtraviricetes" title="Revtraviricetes">Revtraviricetes</a></i>)</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Blubervirales" class="mw-redirect" title="Blubervirales">Blubervirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Hepadnaviridae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Hepadnaviridae" title="Hepadnaviridae">Hepadnaviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Avihepadnavirus" title="Avihepadnavirus">Avihepadnavirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Duck_hepatitis_B_virus" title="Duck hepatitis B virus">Duck hepatitis B virus</a></li> <li><a href="/wiki/Heron_hepatitis_B_virus" class="mw-redirect" title="Heron hepatitis B virus">Heron hepatitis B virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/w/index.php?title=Herpetohepadnavirus&action=edit&redlink=1" class="new" title="Herpetohepadnavirus (page does not exist)">Herpetohepadnavirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li class="mw-empty-elt"></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/w/index.php?title=Metahepadnavirus&action=edit&redlink=1" class="new" title="Metahepadnavirus (page does not exist)">Metahepadnavirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li class="mw-empty-elt"></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Orthohepadnavirus" title="Orthohepadnavirus">Orthohepadnavirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Ground_squirrel_hepatitis_virus" title="Ground squirrel hepatitis virus">Ground squirrel hepatitis virus</a></li> <li><a href="/wiki/Hepatitis_B_virus" title="Hepatitis B virus">Hepatitis B virus</a></li> <li><a href="/wiki/Woodchuck_hepatitis_virus" title="Woodchuck hepatitis virus">Woodchuck hepatitis virus</a></li> <li><a href="/wiki/Woolly_monkey_hepatitis_B_virus" title="Woolly monkey hepatitis B virus">Woolly monkey hepatitis B virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/w/index.php?title=Parahepadnavirus&action=edit&redlink=1" class="new" title="Parahepadnavirus (page does not exist)">Parahepadnavirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li class="mw-empty-elt"></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ortervirales" title="Ortervirales">Ortervirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%">(<a href="/wiki/SsRNA-RT_virus" class="mw-redirect" title="SsRNA-RT virus">ssRNA-RT viruses</a>)</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Belpaoviridae" class="mw-redirect" title="Belpaoviridae">Belpaoviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Semotivirus" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Semotivirus" title="Semotivirus">Semotivirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Caenorhabditis_elegans_Cer13_virus" title="Caenorhabditis elegans Cer13 virus">Caenorhabditis elegans Cer13 virus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Metaviridae" title="Metaviridae">Metaviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/w/index.php?title=Errantivirus&action=edit&redlink=1" class="new" title="Errantivirus (page does not exist)">Errantivirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li class="mw-empty-elt"></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Metavirus" title="Metavirus">Metavirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Caenorhabditis_elegans_Cer1_virus" title="Caenorhabditis elegans Cer1 virus">Caenorhabditis elegans Cer1 virus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pseudoviridae" title="Pseudoviridae">Pseudoviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/w/index.php?title=Hemivirus&action=edit&redlink=1" class="new" title="Hemivirus (page does not exist)">Hemivirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li class="mw-empty-elt"></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pseudovirus_(genus)" class="mw-redirect" title="Pseudovirus (genus)">Pseudovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li class="mw-empty-elt"></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/w/index.php?title=Sirevirus&action=edit&redlink=1" class="new" title="Sirevirus (page does not exist)">Sirevirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li class="mw-empty-elt"></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a class="mw-selflink selflink">Retroviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Orthoretrovirinae" title="Orthoretrovirinae">Orthoretrovirinae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Alpharetrovirus" title="Alpharetrovirus">Alpharetrovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Avian_sarcoma_leukosis_virus" title="Avian sarcoma leukosis virus">Avian sarcoma leukosis virus</a></li> <li><a href="/wiki/Rous_sarcoma_virus" title="Rous sarcoma virus">Rous sarcoma virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Betaretrovirus" title="Betaretrovirus">Betaretrovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Mouse_mammary_tumor_virus" title="Mouse mammary tumor virus">Mouse mammary tumor virus</a></li> <li><a href="/wiki/Jaagsiekte_sheep_retrovirus" title="Jaagsiekte sheep retrovirus">Jaagsiekte sheep retrovirus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Deltaretrovirus" title="Deltaretrovirus">Deltaretrovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Human_T-lymphotropic_virus" class="mw-redirect" title="Human T-lymphotropic virus">Human T-lymphotropic virus</a> (<a href="/wiki/Human_T-lymphotropic_virus_1" title="Human T-lymphotropic virus 1">HTLV-1</a></li> <li><a href="/wiki/Human_T-lymphotropic_virus_2" title="Human T-lymphotropic virus 2">HTLV-2</a>, 3, 4)</li> <li><a href="/wiki/Simian-T-lymphotropic_virus" class="mw-redirect" title="Simian-T-lymphotropic virus">Simian-T-lymphotropic virus</a> (types 1-4)</li> <li><a href="/wiki/Bovine_leukemia_virus" title="Bovine leukemia virus">Bovine leukemia virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Epsilonretrovirus" title="Epsilonretrovirus">Epsilonretrovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Walleye_epidermal_hyperplasia_virus" title="Walleye epidermal hyperplasia virus">Walleye epidermal hyperplasia virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Gammaretrovirus" title="Gammaretrovirus">Gammaretrovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Murine_leukemia_virus" title="Murine leukemia virus">Murine leukemia virus</a></li> <li><a href="/wiki/Abelson_murine_leukemia_virus" title="Abelson murine leukemia virus">Abelson murine leukemia virus</a></li> <li><a href="/wiki/Friend_virus" title="Friend virus">Friend virus</a></li> <li><a href="/wiki/Feline_leukemia_virus" title="Feline leukemia virus">Feline leukemia virus</a></li> <li><a href="/wiki/Koala_retrovirus" title="Koala retrovirus">Koala retrovirus</a> (KoRV)</li> <li><a href="/wiki/Xenotropic_murine_leukemia_virus-related_virus" class="mw-redirect" title="Xenotropic murine leukemia virus-related virus">Xenotropic murine leukemia virus-related virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Lentivirus" title="Lentivirus">Lentivirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/HIV" title="HIV">HIV</a> (human immunodeficiency viruses)</li> <li><a href="/wiki/Simian_immunodeficiency_virus" title="Simian immunodeficiency virus">Simian immunodeficiency viruses</a> (SIV)</li> <li><a href="/wiki/Feline_immunodeficiency_virus" title="Feline immunodeficiency virus">Feline immunodeficiency virus</a> (FIV)</li> <li><a href="/wiki/Puma_lentivirus" title="Puma lentivirus">Puma lentivirus</a> (PLV)</li> <li><a href="/wiki/Equine_infectious_anemia" title="Equine infectious anemia">Equine infectious anemia virus</a> (EIAV)</li> <li><a href="/wiki/Bovine_immunodeficiency_virus" title="Bovine immunodeficiency virus">Bovine immunodeficiency virus</a> (BIV)</li> <li><a href="/wiki/Caprine_arthritis_encephalitis_virus" title="Caprine arthritis encephalitis virus">Caprine arthritis encephalitis virus</a></li> <li><a href="/wiki/Visna-maedi_virus" title="Visna-maedi virus">Visna-maedi virus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Spumaretrovirinae" title="Spumaretrovirinae">Spumaretrovirinae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/w/index.php?title=Bovispumavirus&action=edit&redlink=1" class="new" title="Bovispumavirus (page does not exist)">Bovispumavirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li class="mw-empty-elt"></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/w/index.php?title=Equispumavirus&action=edit&redlink=1" class="new" title="Equispumavirus (page does not exist)">Equispumavirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li class="mw-empty-elt"></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/w/index.php?title=Felispumavirus&action=edit&redlink=1" class="new" title="Felispumavirus (page does not exist)">Felispumavirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Feline_foamy_virus" title="Feline foamy virus">Feline foamy virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/w/index.php?title=Prosimiispumavirus&action=edit&redlink=1" class="new" title="Prosimiispumavirus (page does not exist)">Prosimiispumavirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li class="mw-empty-elt"></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/w/index.php?title=Simiispumavirus&action=edit&redlink=1" class="new" title="Simiispumavirus (page does not exist)">Simiispumavirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Simian_foamy_virus" class="mw-redirect" title="Simian foamy virus">Simian foamy virus</a></li> <li><a href="/wiki/Human_foamy_virus" title="Human foamy virus">Human foamy virus</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">(<a href="/wiki/DsDNA-RT_virus" class="mw-redirect" title="DsDNA-RT virus">dsDNA-RT viruses</a>)</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Caulimoviridae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Caulimoviridae" title="Caulimoviridae">Caulimoviridae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Badnavirus" title="Badnavirus">Badnavirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Banana_streak_MY_virus" title="Banana streak MY virus">Banana streak MY virus</a></li> <li><a href="/wiki/Cacao_swollen_shoot_virus" title="Cacao swollen shoot virus">Cacao swollen shoot virus</a></li> <li><a href="/wiki/Kalanchoe_top-spotting_virus" title="Kalanchoe top-spotting virus">Kalanchoe top-spotting virus</a></li> <li><a href="/wiki/Rubus_yellow_net_virus" title="Rubus yellow net virus">Rubus yellow net virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Caulimovirus" title="Caulimovirus">Caulimovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Carnation_etched_ring_virus" title="Carnation etched ring virus">Carnation etched ring virus</a></li> <li><a href="/wiki/Cauliflower_mosaic_virus" title="Cauliflower mosaic virus">Cauliflower mosaic virus</a></li> <li><a href="/wiki/Strawberry_vein_banding_virus" title="Strawberry vein banding virus">Strawberry vein banding virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cavemovirus" title="Cavemovirus">Cavemovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Cassava_vein_mosaic_virus" title="Cassava vein mosaic virus">Cassava vein mosaic virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/w/index.php?title=Dioscovirus&action=edit&redlink=1" class="new" title="Dioscovirus (page does not exist)">Dioscovirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li 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navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li class="mw-empty-elt"></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/w/index.php?title=Vaccinivirus&action=edit&redlink=1" class="new" title="Vaccinivirus (page does not exist)">Vaccinivirus</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li class="mw-empty-elt"></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Endogenous_retrovirus" title="Endogenous retrovirus">Endogenous</a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/ERV-Fc" title="ERV-Fc">ERV-Fc</a></li> <li><a href="/wiki/Syncytin-1" title="Syncytin-1">ERVWE1</a></li> <li><a href="/wiki/HCP5" title="HCP5">HCP5</a></li> <li><a href="/wiki/Human_teratocarcinoma-derived_virus" class="mw-redirect" title="Human teratocarcinoma-derived virus">Human teratocarcinoma-derived virus</a></li> <li><a href="/wiki/Human_Endogenous_Retrovirus-W" class="mw-redirect" title="Human Endogenous Retrovirus-W">Human Endogenous Retrovirus-W</a></li></ul> </div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Baltimore_(virus_classification)" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><link rel="mw-deduplicated-inline-style" 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classification">virus classification</a>)</div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/DNA_virus" title="DNA virus">DNA</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="I:_dsDNA_viruses" style="font-size:114%;margin:0 4em">I: <a href="/wiki/DsDNA_virus" class="mw-redirect" title="DsDNA virus">dsDNA viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Adnaviria" title="Adnaviria">Adnaviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Zilligvirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Zilligvirae" class="mw-redirect" title="Zilligvirae">Zilligvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Taleaviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Taleaviricota" class="mw-redirect" title="Taleaviricota">Taleaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Tokiviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tokiviricetes" class="mw-redirect" title="Tokiviricetes">Tokiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ligamenvirales" title="Ligamenvirales">Ligamenvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Lipothrixviridae" title="Lipothrixviridae">Lipothrixviridae</a></i></li> <li><i><a href="/wiki/Rudiviridae" title="Rudiviridae">Rudiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Primavirales" class="mw-redirect" title="Primavirales">Primavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Tristromaviridae" title="Tristromaviridae">Tristromaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Duplodnaviria" title="Duplodnaviria">Duplodnaviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Heunggongvirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Heunggongvirae" class="mw-redirect" title="Heunggongvirae">Heunggongvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Peploviricota" class="mw-redirect" title="Peploviricota">Peploviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Herviviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Herviviricetes" class="mw-redirect" title="Herviviricetes">Herviviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Herpesvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Herpesvirales" title="Herpesvirales">Herpesvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alloherpesviridae" title="Alloherpesviridae">Alloherpesviridae</a></i></li> <li><i><a href="/wiki/Herpesviridae" title="Herpesviridae">Herpesviridae</a></i></li> <li><i><a href="/wiki/Malacoherpesviridae" title="Malacoherpesviridae">Malacoherpesviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Uroviricota" class="mw-redirect" title="Uroviricota">Uroviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Caudoviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Caudoviricetes" title="Caudoviricetes">Caudoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Caudovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Caudovirales" class="mw-redirect" title="Caudovirales">Caudovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Ackermannviridae" title="Ackermannviridae">Ackermannviridae</a></i></li> <li><i><a href="/wiki/Autographiviridae" title="Autographiviridae">Autographiviridae</a></i></li> <li><i><a href="/w/index.php?title=Chaseviridae&action=edit&redlink=1" class="new" title="Chaseviridae (page does not exist)">Chaseviridae</a></i></li> <li><i><a href="/w/index.php?title=Demerecviridae&action=edit&redlink=1" class="new" title="Demerecviridae (page does not exist)">Demerecviridae</a></i></li> <li><i><a href="/w/index.php?title=Drexlerviridae&action=edit&redlink=1" class="new" title="Drexlerviridae (page does not exist)">Drexlerviridae</a></i></li> <li><i><a href="/w/index.php?title=Guelinviridae&action=edit&redlink=1" class="new" title="Guelinviridae (page does not exist)">Guelinviridae</a></i></li> <li><i><a href="/wiki/Herelleviridae" title="Herelleviridae">Herelleviridae</a></i></li> <li><i><a href="/wiki/Myoviridae" title="Myoviridae">Myoviridae</a></i></li> <li><i><a href="/wiki/Podoviridae" title="Podoviridae">Podoviridae</a></i></li> <li><i><a href="/w/index.php?title=Rountreeviridae&action=edit&redlink=1" class="new" title="Rountreeviridae (page does not exist)">Rountreeviridae</a></i></li> <li><i><a href="/w/index.php?title=Salasmaviridae&action=edit&redlink=1" class="new" title="Salasmaviridae (page does not exist)">Salasmaviridae</a></i></li> <li><i><a href="/w/index.php?title=Schitoviridae&action=edit&redlink=1" class="new" title="Schitoviridae (page does not exist)">Schitoviridae</a></i></li> <li><i><a href="/wiki/Siphoviridae" title="Siphoviridae">Siphoviridae</a></i></li> <li><i><a href="/w/index.php?title=Zobellviridae&action=edit&redlink=1" class="new" title="Zobellviridae (page does not exist)">Zobellviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Monodnaviria" title="Monodnaviria">Monodnaviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Shotokuvirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Shotokuvirae" title="Shotokuvirae">Shotokuvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Cossaviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cossaviricota" title="Cossaviricota">Cossaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Papovaviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Papovaviricetes" title="Papovaviricetes">Papovaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Sepolyvirales" class="mw-redirect" title="Sepolyvirales">Sepolyvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Polyomaviridae" title="Polyomaviridae">Polyomaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Zurhausenvirales" class="mw-redirect" title="Zurhausenvirales">Zurhausenvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Papillomaviridae" title="Papillomaviridae">Papillomaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Varidnaviria" title="Varidnaviria">Varidnaviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Bamfordvirae" title="Bamfordvirae">Bamfordvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Nucleocytoviricota" title="Nucleocytoviricota">Nucleocytoviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pokkesviricetes" title="Pokkesviricetes">Pokkesviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Asfuvirales" class="mw-redirect" title="Asfuvirales">Asfuvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Asfarviridae" title="Asfarviridae">Asfarviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Chitovirales" class="mw-redirect" title="Chitovirales">Chitovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Poxviridae" title="Poxviridae">Poxviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Megaviricetes" title="Megaviricetes">Megaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Algavirales" class="mw-redirect" title="Algavirales">Algavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Phycodnaviridae" title="Phycodnaviridae">Phycodnaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Imitervirales" class="mw-redirect" title="Imitervirales">Imitervirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Mimiviridae" title="Mimiviridae">Mimiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pimascovirales" title="Pimascovirales">Pimascovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Ascoviridae" title="Ascoviridae">Ascoviridae</a></i></li> <li><i><a href="/wiki/Iridoviridae" title="Iridoviridae">Iridoviridae</a></i></li> <li><i><a href="/wiki/Marseilleviridae" title="Marseilleviridae">Marseilleviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Preplasmiviricota" title="Preplasmiviricota">Preplasmiviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Maveriviricetes" class="mw-redirect" title="Maveriviricetes">Maveriviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Priklausovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Priklausovirales" class="mw-redirect" title="Priklausovirales">Priklausovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Lavidaviridae" class="mw-redirect" title="Lavidaviridae">Lavidaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Polintoviricetes" class="mw-redirect" title="Polintoviricetes">Polintoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Orthopolintovirales" class="mw-redirect" title="Orthopolintovirales">Orthopolintovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Adintoviridae" title="Adintoviridae">Adintoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tectiliviricetes" title="Tectiliviricetes">Tectiliviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Belfryvirales" class="mw-redirect" title="Belfryvirales">Belfryvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Turriviridae" title="Turriviridae">Turriviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Kalamavirales" class="mw-redirect" title="Kalamavirales">Kalamavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Tectiviridae" class="mw-redirect" title="Tectiviridae">Tectiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Rowavirales" class="mw-redirect" title="Rowavirales">Rowavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Adenoviridae" title="Adenoviridae">Adenoviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Vinavirales" class="mw-redirect" title="Vinavirales">Vinavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Corticoviridae" class="mw-redirect" title="Corticoviridae">Corticoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Helvetiavirae" class="mw-redirect" title="Helvetiavirae">Helvetiavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Dividoviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Dividoviricota" class="mw-redirect" title="Dividoviricota">Dividoviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Laserviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Laserviricetes" class="mw-redirect" title="Laserviricetes">Laserviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Halopanivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Halopanivirales" title="Halopanivirales">Halopanivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Matshushitaviridae" class="mw-redirect" title="Matshushitaviridae">Matshushitaviridae</a></i></li> <li><i><a href="/wiki/Simuloviridae" class="mw-redirect" title="Simuloviridae">Simuloviridae</a></i></li> <li><i><a href="/wiki/Sphaerolipoviridae" class="mw-redirect" title="Sphaerolipoviridae">Sphaerolipoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Naldaviricetes" title="Naldaviricetes">Naldaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Lefavirales" title="Lefavirales">Lefavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Baculoviridae" title="Baculoviridae">Baculoviridae</a></i></li> <li><i><a href="/wiki/Hytrosaviridae" title="Hytrosaviridae">Hytrosaviridae</a></i></li> <li><i><a href="/wiki/Nudiviridae" class="mw-redirect" title="Nudiviridae">Nudiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Nimaviridae" class="mw-redirect" title="Nimaviridae">Nimaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><b>Families</b>: <i><a href="/wiki/Ampullaviridae" title="Ampullaviridae">Ampullaviridae</a></i></li> <li><i><a href="/wiki/Bicaudaviridae" title="Bicaudaviridae">Bicaudaviridae</a></i></li> <li><i><a href="/wiki/Clavaviridae" title="Clavaviridae">Clavaviridae</a></i></li> <li><i><a href="/wiki/Fuselloviridae" title="Fuselloviridae">Fuselloviridae</a></i></li> <li><i><a href="/wiki/Globuloviridae" title="Globuloviridae">Globuloviridae</a></i></li> <li><i><a href="/wiki/Guttaviridae" title="Guttaviridae">Guttaviridae</a></i></li> <li><i><a href="/wiki/Halspiviridae" title="Halspiviridae">Halspiviridae</a></i></li> <li><i><a href="/wiki/Ovaliviridae" title="Ovaliviridae">Ovaliviridae</a></i></li> <li><i><a href="/wiki/Plasmaviridae" title="Plasmaviridae">Plasmaviridae</a></i></li> <li><i><a href="/wiki/Polydnaviridae" class="mw-redirect" title="Polydnaviridae">Polydnaviridae</a></i></li> <li><i><a href="/wiki/Portogloboviridae" title="Portogloboviridae">Portogloboviridae</a></i></li> <li><i><a href="/wiki/Thaspiviridae" title="Thaspiviridae">Thaspiviridae</a></i></li> <li><b>Genera</b>: <i><a href="/wiki/Dinodnavirus" title="Dinodnavirus">Dinodnavirus</a></i></li> <li><i><a href="/wiki/Rhizidiovirus" title="Rhizidiovirus">Rhizidiovirus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="II:_ssDNA_viruses" style="font-size:114%;margin:0 4em">II: <a href="/wiki/SsDNA_virus" class="mw-redirect" title="SsDNA virus">ssDNA viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Monodnaviria" title="Monodnaviria">Monodnaviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Loebvirae" class="mw-redirect" title="Loebvirae">Loebvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Hofneiviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Hofneiviricota" class="mw-redirect" title="Hofneiviricota">Hofneiviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Faserviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Faserviricetes" class="mw-redirect" title="Faserviricetes">Faserviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Tubulavirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tubulavirales" title="Tubulavirales">Tubulavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Inoviridae" class="mw-redirect" title="Inoviridae">Inoviridae</a></i></li> <li><i><a href="/w/index.php?title=Paulinoviridae&action=edit&redlink=1" class="new" title="Paulinoviridae (page does not exist)">Paulinoviridae</a></i></li> <li><i><a href="/w/index.php?title=Plectroviridae&action=edit&redlink=1" class="new" title="Plectroviridae (page does not exist)">Plectroviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Sangervirae" class="mw-redirect" title="Sangervirae">Sangervirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Phixviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Phixviricota" class="mw-redirect" title="Phixviricota">Phixviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Malgrandaviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Malgrandaviricetes" class="mw-redirect" title="Malgrandaviricetes">Malgrandaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Petitvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Petitvirales" class="mw-redirect" title="Petitvirales">Petitvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Microviridae" title="Microviridae">Microviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Shotokuvirae" title="Shotokuvirae">Shotokuvirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cossaviricota" title="Cossaviricota">Cossaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Mouviricetes" class="mw-redirect" title="Mouviricetes">Mouviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Polivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Polivirales" class="mw-redirect" title="Polivirales">Polivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Bidnaviridae" class="mw-redirect" title="Bidnaviridae">Bidnaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Quintoviricetes" class="mw-redirect" title="Quintoviricetes">Quintoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Piccovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Piccovirales" class="mw-redirect" title="Piccovirales">Piccovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Parvoviridae" title="Parvoviridae">Parvoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cressdnaviricota" title="Cressdnaviricota">Cressdnaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Arfiviricetes" title="Arfiviricetes">Arfiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Baphyvirales" class="mw-redirect" title="Baphyvirales">Baphyvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Bacilladnaviridae" title="Bacilladnaviridae">Bacilladnaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cirlivirales" class="mw-redirect" title="Cirlivirales">Cirlivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Circoviridae" title="Circoviridae">Circoviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cremevirales" class="mw-redirect" title="Cremevirales">Cremevirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Smacoviridae" title="Smacoviridae">Smacoviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Mulpavirales" title="Mulpavirales">Mulpavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Metaxyviridae&action=edit&redlink=1" class="new" title="Metaxyviridae (page does not exist)">Metaxyviridae</a></i></li> <li><i><a href="/wiki/Nanoviridae" title="Nanoviridae">Nanoviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Recrevirales" class="mw-redirect" title="Recrevirales">Recrevirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Redondoviridae" title="Redondoviridae">Redondoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Repensiviricetes" class="mw-redirect" title="Repensiviricetes">Repensiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Geplafuvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Geplafuvirales" title="Geplafuvirales">Geplafuvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Geminiviridae" title="Geminiviridae">Geminiviridae</a></i></li> <li><i><a href="/wiki/Genomoviridae" title="Genomoviridae">Genomoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Trapavirae" class="mw-redirect" title="Trapavirae">Trapavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Saleviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Saleviricota" class="mw-redirect" title="Saleviricota">Saleviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Huolimaviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Huolimaviricetes" class="mw-redirect" title="Huolimaviricetes">Huolimaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Haloruvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Haloruvirales" class="mw-redirect" title="Haloruvirales">Haloruvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Pleolipoviridae" title="Pleolipoviridae">Pleolipoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><b>Families</b>: <i><a href="/wiki/Anelloviridae" title="Anelloviridae">Anelloviridae</a></i></li> <li><i><a href="/wiki/Finnlakeviridae" title="Finnlakeviridae">Finnlakeviridae</a></i></li> <li><i><a href="/wiki/Spiraviridae" title="Spiraviridae">Spiraviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/RNA_virus" title="RNA virus">RNA</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="III:_dsRNA_viruses" style="font-size:114%;margin:0 4em">III: <a href="/wiki/Double-stranded_RNA_viruses" title="Double-stranded RNA viruses">dsRNA viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Riboviria" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Riboviria" title="Riboviria">Riboviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Orthornavirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Orthornavirae" title="Orthornavirae">Orthornavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Duplornaviricota" title="Duplornaviricota">Duplornaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Chrymotiviricetes" class="mw-redirect" title="Chrymotiviricetes">Chrymotiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Ghabrivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ghabrivirales" title="Ghabrivirales">Ghabrivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Chrysoviridae" title="Chrysoviridae">Chrysoviridae</a></i></li> <li><i><a href="/wiki/Megabirnaviridae" title="Megabirnaviridae">Megabirnaviridae</a></i></li> <li><i><a href="/wiki/Quadriviridae" title="Quadriviridae">Quadriviridae</a></i></li> <li><i><a href="/wiki/Totiviridae" title="Totiviridae">Totiviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Resentoviricetes" class="mw-redirect" title="Resentoviricetes">Resentoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Reovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Reovirales" class="mw-redirect" title="Reovirales">Reovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Reoviridae" class="mw-redirect" title="Reoviridae">Reoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Vidaverviricetes" class="mw-redirect" title="Vidaverviricetes">Vidaverviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Mindivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Mindivirales" class="mw-redirect" title="Mindivirales">Mindivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Cystoviridae" class="mw-redirect" title="Cystoviridae">Cystoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pisuviricota" title="Pisuviricota">Pisuviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Duplopiviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Duplopiviricetes" class="mw-redirect" title="Duplopiviricetes">Duplopiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Durnavirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Durnavirales" title="Durnavirales">Durnavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Amalgaviridae" title="Amalgaviridae">Amalgaviridae</a></i></li> <li><i><a href="/w/index.php?title=Curvulaviridae&action=edit&redlink=1" class="new" title="Curvulaviridae (page does not exist)">Curvulaviridae</a></i></li> <li><i><a href="/wiki/Hypoviridae" class="mw-redirect" title="Hypoviridae">Hypoviridae</a></i></li> <li><i><a href="/wiki/Picobirnaviridae" class="mw-redirect" title="Picobirnaviridae">Picobirnaviridae</a></i></li> <li><i><a href="/wiki/Partitiviridae" title="Partitiviridae">Partitiviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><b>Families</b>: <i><a href="/wiki/Birnaviridae" title="Birnaviridae">Birnaviridae</a></i></li> <li><i><a href="/w/index.php?title=Polymycoviridae&action=edit&redlink=1" class="new" title="Polymycoviridae (page does not exist)">Polymycoviridae</a></i></li> <li><b>Genera</b>: <i><a href="/w/index.php?title=Botybirnavirus&action=edit&redlink=1" class="new" title="Botybirnavirus (page does not exist)">Botybirnavirus</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="IV:_(+)ssRNA_viruses" style="font-size:114%;margin:0 4em">IV: <a href="/wiki/Positive-sense_ssRNA_virus" class="mw-redirect" title="Positive-sense ssRNA virus">(+)ssRNA viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Riboviria" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Riboviria" title="Riboviria">Riboviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Orthornavirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Orthornavirae" title="Orthornavirae">Orthornavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Kitrinoviricota" title="Kitrinoviricota">Kitrinoviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Alsuviricetes" title="Alsuviricetes">Alsuviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Hepelivirales" title="Hepelivirales">Hepelivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alphatetraviridae" title="Alphatetraviridae">Alphatetraviridae</a></i></li> <li><i><a href="/wiki/Benyviridae" class="mw-redirect" title="Benyviridae">Benyviridae</a></i></li> <li><i><a href="/wiki/Hepeviridae" title="Hepeviridae">Hepeviridae</a></i></li> <li><i><a href="/wiki/Matonaviridae" class="mw-redirect" title="Matonaviridae">Matonaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Martellivirales" title="Martellivirales">Martellivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Bromoviridae" title="Bromoviridae">Bromoviridae</a></i></li> <li><i><a href="/wiki/Closteroviridae" title="Closteroviridae">Closteroviridae</a></i></li> <li><i><a href="/wiki/Endornaviridae" title="Endornaviridae">Endornaviridae</a></i></li> <li><i><a href="/w/index.php?title=Kitaviridae&action=edit&redlink=1" class="new" title="Kitaviridae (page does not exist)">Kitaviridae</a></i></li> <li><i><a href="/w/index.php?title=Mayoviridae&action=edit&redlink=1" class="new" title="Mayoviridae (page does not exist)">Mayoviridae</a></i></li> <li><i><a href="/wiki/Togaviridae" class="mw-redirect" title="Togaviridae">Togaviridae</a></i></li> <li><i><a href="/wiki/Virgaviridae" title="Virgaviridae">Virgaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tymovirales" title="Tymovirales">Tymovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alphaflexiviridae" title="Alphaflexiviridae">Alphaflexiviridae</a></i></li> <li><i><a href="/wiki/Betaflexiviridae" title="Betaflexiviridae">Betaflexiviridae</a></i></li> <li><i><a href="/w/index.php?title=Deltaflexiviridae&action=edit&redlink=1" class="new" title="Deltaflexiviridae (page does not exist)">Deltaflexiviridae</a></i></li> <li><i><a href="/wiki/Gammaflexiviridae" title="Gammaflexiviridae">Gammaflexiviridae</a></i></li> <li><i><a href="/wiki/Tymoviridae" title="Tymoviridae">Tymoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Flasuviricetes" class="mw-redirect" title="Flasuviricetes">Flasuviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Amarillovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Amarillovirales" class="mw-redirect" title="Amarillovirales">Amarillovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Flaviviridae" title="Flaviviridae">Flaviviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Magsaviricetes" class="mw-redirect" title="Magsaviricetes">Magsaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Nodamuvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Nodamuvirales" title="Nodamuvirales">Nodamuvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Nodaviridae" title="Nodaviridae">Nodaviridae</a></i></li> <li><i><a href="/w/index.php?title=Sinhaliviridae&action=edit&redlink=1" class="new" title="Sinhaliviridae (page does not exist)">Sinhaliviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tolucaviricetes" class="mw-redirect" title="Tolucaviricetes">Tolucaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Tolivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Tolivirales" title="Tolivirales">Tolivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Carmotetraviridae" title="Carmotetraviridae">Carmotetraviridae</a></i></li> <li><i><a href="/wiki/Luteoviridae" title="Luteoviridae">Luteoviridae</a></i></li> <li><i><a href="/wiki/Tombusviridae" title="Tombusviridae">Tombusviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Lenarviricota" title="Lenarviricota">Lenarviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Leviviricetes" title="Leviviricetes">Leviviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Norzivirales" title="Norzivirales">Norzivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Atkinsviridae" title="Atkinsviridae">Atkinsviridae</a></i></li> <li><i><a href="/wiki/Duinviridae" title="Duinviridae">Duinviridae</a></i></li> <li><i><a href="/wiki/Fiersviridae" title="Fiersviridae">Fiersviridae</a></i></li> <li><i><a href="/wiki/Solspiviridae" title="Solspiviridae">Solspiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Timlovirales" title="Timlovirales">Timlovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Blumeviridae" title="Blumeviridae">Blumeviridae</a></i></li> <li><i><a href="/wiki/Steitzviridae" title="Steitzviridae">Steitzviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Amabiliviricetes" class="mw-redirect" title="Amabiliviricetes">Amabiliviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Wolframvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Wolframvirales" class="mw-redirect" title="Wolframvirales">Wolframvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Narnaviridae" class="mw-redirect" title="Narnaviridae">Narnaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Howeltoviricetes" class="mw-redirect" title="Howeltoviricetes">Howeltoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Cryppavirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Cryppavirales" class="mw-redirect" title="Cryppavirales">Cryppavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Mitoviridae" class="mw-redirect" title="Mitoviridae">Mitoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Miaviricetes" class="mw-redirect" title="Miaviricetes">Miaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Ourlivirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ourlivirales" class="mw-redirect" title="Ourlivirales">Ourlivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Botourmiaviridae" title="Botourmiaviridae">Botourmiaviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pisuviricota" title="Pisuviricota">Pisuviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pisoniviricetes" title="Pisoniviricetes">Pisoniviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Nidovirales" title="Nidovirales">Nidovirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Abyssoviridae" class="mw-redirect" title="Abyssoviridae">Abyssoviridae</a></i></li> <li><i><a href="/wiki/Arteriviridae" title="Arteriviridae">Arteriviridae</a></i></li> <li><i><a href="/w/index.php?title=Cremegaviridae&action=edit&redlink=1" class="new" title="Cremegaviridae (page does not exist)">Cremegaviridae</a></i></li> <li><i><a href="/wiki/Coronaviridae" title="Coronaviridae">Coronaviridae</a></i></li> <li><i><a href="/w/index.php?title=Euroniviridae&action=edit&redlink=1" class="new" title="Euroniviridae (page does not exist)">Euroniviridae</a></i></li> <li><i><a href="/w/index.php?title=Gresnaviridae&action=edit&redlink=1" class="new" title="Gresnaviridae (page does not exist)">Gresnaviridae</a></i></li> <li><i><a href="/w/index.php?title=Medioniviridae&action=edit&redlink=1" class="new" title="Medioniviridae (page does not exist)">Medioniviridae</a></i></li> <li><i><a href="/wiki/Mesoniviridae" title="Mesoniviridae">Mesoniviridae</a></i></li> <li><i><a href="/wiki/Mononiviridae" class="mw-redirect" title="Mononiviridae">Mononiviridae</a></i></li> <li><i><a href="/w/index.php?title=Nanghoshaviridae&action=edit&redlink=1" class="new" title="Nanghoshaviridae (page does not exist)">Nanghoshaviridae</a></i></li> <li><i><a href="/w/index.php?title=Nanhypoviridae&action=edit&redlink=1" class="new" title="Nanhypoviridae (page does not exist)">Nanhypoviridae</a></i></li> <li><i><a href="/w/index.php?title=Olifoviridae&action=edit&redlink=1" class="new" title="Olifoviridae (page does not exist)">Olifoviridae</a></i></li> <li><i><a href="/wiki/Roniviridae" class="mw-redirect" title="Roniviridae">Roniviridae</a></i></li> <li><i><a href="/wiki/Tobaniviridae" title="Tobaniviridae">Tobaniviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Picornavirales" title="Picornavirales">Picornavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Picornaviridae" class="mw-redirect" title="Picornaviridae">Picornaviridae</a></i></li> <li><i><a href="/wiki/Marnaviridae" title="Marnaviridae">Marnaviridae</a></i></li> <li><i><a href="/w/index.php?title=Solinviviridae&action=edit&redlink=1" class="new" title="Solinviviridae (page does not exist)">Solinviviridae</a></i></li> <li><i><a href="/wiki/Caliciviridae" title="Caliciviridae">Caliciviridae</a></i></li> <li><i><a href="/wiki/Iflaviridae" title="Iflaviridae">Iflaviridae</a></i></li> <li><i><a href="/wiki/Secoviridae" title="Secoviridae">Secoviridae</a></i></li> <li><i><a href="/wiki/Dicistroviridae" title="Dicistroviridae">Dicistroviridae</a></i></li> <li><i><a href="/w/index.php?title=Polycipiviridae&action=edit&redlink=1" class="new" title="Polycipiviridae (page does not exist)">Polycipiviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Sobelivirales" title="Sobelivirales">Sobelivirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Alvernaviridae" title="Alvernaviridae">Alvernaviridae</a></i></li> <li><i><a href="/wiki/Barnaviridae" title="Barnaviridae">Barnaviridae</a></i></li> <li><i><a href="/wiki/Solemoviridae" title="Solemoviridae">Solemoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Stelpaviricetes" title="Stelpaviricetes">Stelpaviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Patatavirales" class="mw-redirect" title="Patatavirales">Patatavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Potyviridae" title="Potyviridae">Potyviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Stellavirales" class="mw-redirect" title="Stellavirales">Stellavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Astroviridae" class="mw-redirect" title="Astroviridae">Astroviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%">Unassigned</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><b>Families</b>: <i><a href="/wiki/Permutotetraviridae" title="Permutotetraviridae">Permutotetraviridae</a></i></li> <li><i><a href="/wiki/Sarthroviridae" class="mw-redirect" title="Sarthroviridae">Sarthroviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="V:_(–)ssRNA_viruses" style="font-size:114%;margin:0 4em">V: <a href="/wiki/Negative-sense_ssRNA_virus" class="mw-redirect" title="Negative-sense ssRNA virus">(–)ssRNA viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Riboviria" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Riboviria" title="Riboviria">Riboviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Orthornavirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Orthornavirae" title="Orthornavirae">Orthornavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Negarnaviricota" class="mw-redirect" title="Negarnaviricota">Negarnaviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Chunqiuviricetes" class="mw-redirect" title="Chunqiuviricetes">Chunqiuviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Muvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Muvirales" class="mw-redirect" title="Muvirales">Muvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Qinviridae" class="mw-redirect" title="Qinviridae">Qinviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ellioviricetes" class="mw-redirect" title="Ellioviricetes">Ellioviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Bunyavirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Bunyavirales" title="Bunyavirales">Bunyavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Cruliviridae" title="Cruliviridae">Cruliviridae</a></i></li> <li><i><a href="/wiki/Arenaviridae" class="mw-redirect" title="Arenaviridae">Arenaviridae</a></i></li> <li><i><a href="/wiki/Fimoviridae" class="mw-redirect" title="Fimoviridae">Fimoviridae</a></i></li> <li><i><a href="/wiki/Hantaviridae" title="Hantaviridae">Hantaviridae</a></i></li> <li><i><a href="/w/index.php?title=Leishbuviridae&action=edit&redlink=1" class="new" title="Leishbuviridae (page does not exist)">Leishbuviridae</a></i></li> <li><i><a href="/w/index.php?title=Mypoviridae&action=edit&redlink=1" class="new" title="Mypoviridae (page does not exist)">Mypoviridae</a></i></li> <li><i><a href="/wiki/Nairoviridae" title="Nairoviridae">Nairoviridae</a></i></li> <li><i><a href="/wiki/Peribunyaviridae" title="Peribunyaviridae">Peribunyaviridae</a></i></li> <li><i><a href="/wiki/Phasmaviridae" title="Phasmaviridae">Phasmaviridae</a></i></li> <li><i><a href="/wiki/Phenuiviridae" title="Phenuiviridae">Phenuiviridae</a></i></li> <li><i><a href="/wiki/Tospoviridae" class="mw-redirect" title="Tospoviridae">Tospoviridae</a></i></li> <li><i><a href="/w/index.php?title=Wupedeviridae&action=edit&redlink=1" class="new" title="Wupedeviridae (page does not exist)">Wupedeviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Insthoviricetes" class="mw-redirect" title="Insthoviricetes">Insthoviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Articulavirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Articulavirales" title="Articulavirales">Articulavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Amnoonviridae" class="mw-redirect" title="Amnoonviridae">Amnoonviridae</a></i></li> <li><i><a href="/wiki/Orthomyxoviridae" title="Orthomyxoviridae">Orthomyxoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Milneviricetes" class="mw-redirect" title="Milneviricetes">Milneviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Serpentovirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Serpentovirales" class="mw-redirect" title="Serpentovirales">Serpentovirales </a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Aspiviridae" title="Aspiviridae">Aspiviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Monjiviricetes" title="Monjiviricetes">Monjiviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Jingchuvirales" title="Jingchuvirales">Jingchuvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Aliusviridae&action=edit&redlink=1" class="new" title="Aliusviridae (page does not exist)">Aliusviridae</a></i></li> <li><i><a href="/wiki/Chuviridae" title="Chuviridae">Chuviridae</a></i></li> <li><i><a href="/w/index.php?title=Crepuscuviridae&action=edit&redlink=1" class="new" title="Crepuscuviridae (page does not exist)">Crepuscuviridae</a></i></li> <li><i><a href="/w/index.php?title=Myriaviridae&action=edit&redlink=1" class="new" title="Myriaviridae (page does not exist)">Myriaviridae</a></i></li> <li><i><a href="/w/index.php?title=Natareviridae&action=edit&redlink=1" class="new" title="Natareviridae (page does not exist)">Natareviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Mononegavirales" title="Mononegavirales">Mononegavirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Artoviridae" title="Artoviridae">Artoviridae</a></i></li> <li><i><a href="/wiki/Bornaviridae" title="Bornaviridae">Bornaviridae</a></i></li> <li><i><a href="/wiki/Filoviridae" title="Filoviridae">Filoviridae</a></i></li> <li><i><a href="/w/index.php?title=Lispiviridae&action=edit&redlink=1" class="new" title="Lispiviridae (page does not exist)">Lispiviridae</a></i></li> <li><i><a href="/wiki/Mymonaviridae" title="Mymonaviridae">Mymonaviridae</a></i></li> <li><i><a href="/wiki/Nyamiviridae" title="Nyamiviridae">Nyamiviridae</a></i></li> <li><i><a href="/wiki/Paramyxoviridae" title="Paramyxoviridae">Paramyxoviridae</a></i></li> <li><i><a href="/wiki/Pneumoviridae" title="Pneumoviridae">Pneumoviridae</a></i></li> <li><i><a href="/wiki/Rhabdoviridae" title="Rhabdoviridae">Rhabdoviridae</a></i></li> <li><i><a href="/wiki/Sunviridae" title="Sunviridae">Sunviridae</a></i></li> <li><i><a href="/w/index.php?title=Xinmoviridae&action=edit&redlink=1" class="new" title="Xinmoviridae (page does not exist)">Xinmoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Yunchangviricetes" class="mw-redirect" title="Yunchangviricetes">Yunchangviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Goujianvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Goujianvirales" class="mw-redirect" title="Goujianvirales">Goujianvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Yueviridae" class="mw-redirect" title="Yueviridae">Yueviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ambiviricota" title="Ambiviricota">Ambiviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Suforviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Suforviricetes" class="mw-redirect" title="Suforviricetes">Suforviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Crytulvirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Crytulvirales" class="mw-redirect" title="Crytulvirales">Crytulvirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/w/index.php?title=Dumbiviridae&action=edit&redlink=1" class="new" title="Dumbiviridae (page does not exist)">Dumbiviridae</a></i></li> <li><i><a href="/w/index.php?title=Quambiviridae&action=edit&redlink=1" class="new" title="Quambiviridae (page does not exist)">Quambiviridae</a></i></li> <li><i><a href="/w/index.php?title=Trimbiviridae&action=edit&redlink=1" class="new" title="Trimbiviridae (page does not exist)">Trimbiviridae</a></i></li> <li><i><a href="/w/index.php?title=Unambiviridae&action=edit&redlink=1" class="new" title="Unambiviridae (page does not exist)">Unambiviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><a href="/wiki/Reverse_transcribing_virus" title="Reverse transcribing virus">RT</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="VI:_ssRNA-RT_viruses" style="font-size:114%;margin:0 4em">VI: <a href="/wiki/SsRNA-RT_virus" class="mw-redirect" title="SsRNA-RT virus">ssRNA-RT viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Riboviria" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Riboviria" title="Riboviria">Riboviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Pararnavirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pararnavirae" class="mw-redirect" title="Pararnavirae">Pararnavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Artverviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Artverviricota" class="mw-redirect" title="Artverviricota">Artverviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Revtraviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Revtraviricetes" title="Revtraviricetes">Revtraviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Ortervirales" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ortervirales" title="Ortervirales">Ortervirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Belpaoviridae" class="mw-redirect" title="Belpaoviridae">Belpaoviridae</a></i></li> <li><i><a href="/wiki/Metaviridae" title="Metaviridae">Metaviridae</a></i></li> <li><i><a href="/wiki/Pseudoviridae" title="Pseudoviridae">Pseudoviridae</a></i></li> <li><i><a href="/wiki/Retroviridae" class="mw-redirect" title="Retroviridae">Retroviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks mw-collapsible mw-collapsed navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="VII:_dsDNA-RT_viruses" style="font-size:114%;margin:0 4em">VII: <a href="/wiki/DsDNA-RT_virus" class="mw-redirect" title="DsDNA-RT virus">dsDNA-RT viruses</a></div></th></tr><tr><td colspan="2" class="navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Riboviria" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Riboviria" title="Riboviria">Riboviria</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Pararnavirae" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Pararnavirae" class="mw-redirect" title="Pararnavirae">Pararnavirae</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Artverviricota" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Artverviricota" class="mw-redirect" title="Artverviricota">Artverviricota</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th id="Revtraviricetes" scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Revtraviricetes" title="Revtraviricetes">Revtraviricetes</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Blubervirales" class="mw-redirect" title="Blubervirales">Blubervirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Hepadnaviridae" title="Hepadnaviridae">Hepadnaviridae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%"><i><a href="/wiki/Ortervirales" title="Ortervirales">Ortervirales</a></i></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Caulimoviridae" title="Caulimoviridae">Caulimoviridae</a></i></li></ul> </div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table><div></div></td></tr></tbody></table></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Self-replicating_organic_structures" style="padding:3px"><table class="nowraplinks mw-collapsible autocollapse navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2" style="text-align: center;"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1239400231"><div class="navbar plainlinks hlist navbar-mini"><ul><li class="nv-view"><a href="/wiki/Template:Self-replicating_organic_structures" title="Template:Self-replicating organic structures"><abbr title="View this template">v</abbr></a></li><li class="nv-talk"><a href="/wiki/Template_talk:Self-replicating_organic_structures" title="Template talk:Self-replicating organic structures"><abbr title="Discuss this template">t</abbr></a></li><li class="nv-edit"><a href="/wiki/Special:EditPage/Template:Self-replicating_organic_structures" title="Special:EditPage/Template:Self-replicating organic structures"><abbr title="Edit this template">e</abbr></a></li></ul></div><div id="Self-replicating_organic_structures" style="font-size:114%;margin:0 4em"><a href="/wiki/Self-replication" title="Self-replication">Self-replicating</a> organic structures</div></th></tr><tr><th scope="row" class="navbox-group" style="text-align: center;;width:1%"><a href="/wiki/Life" title="Life">Cellular life</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Bacteria" title="Bacteria">Bacteria</a></li> <li><a href="/wiki/Archaea" title="Archaea">Archaea</a></li> <li><a href="/wiki/Eukaryote" title="Eukaryote">Eukaryota</a> <ul><li><a href="/wiki/Animal" title="Animal">Animalia</a></li> <li><a href="/wiki/Fungus" title="Fungus">Fungi</a></li> <li><a href="/wiki/Plant" title="Plant">Plantae</a></li> <li><a href="/wiki/Protist" title="Protist">Protista</a></li></ul></li> <li><i><a href="/wiki/Incertae_sedis" title="Incertae sedis">Incertae sedis</a></i> <ul><li><i><a href="/wiki/Parakaryon" title="Parakaryon">Parakaryon</a></i></li> <li><a href="/wiki/Biological_dark_matter" title="Biological dark matter">Biological dark matter</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="text-align: center;;width:1%;background:#CEDAF2;"><a href="/wiki/Virus" title="Virus">Virus</a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/DNA_virus#Group_I:_dsDNA_viruses" title="DNA virus">dsDNA virus</a> <ul><li><a href="/wiki/Giant_virus" title="Giant virus">Giant virus</a></li></ul></li> <li><a href="/wiki/DNA_virus#Group_II:_ssDNA_viruses" title="DNA virus">ssDNA virus</a></li> <li><a href="/wiki/Double-stranded_RNA_viruses" title="Double-stranded RNA viruses">dsRNA virus</a></li> <li><a href="/wiki/RNA_virus#Group_IV—positive-sense_ssRNA_viruses" title="RNA virus">(+)ssRNA virus</a></li> <li><a href="/wiki/RNA_virus#Group_V—negative-sense_ssRNA_viruses" title="RNA virus">(−)ssRNA virus</a></li> <li><a class="mw-selflink selflink">ssRNA-RT virus</a></li> <li><a href="/wiki/DsDNA-RT_virus" class="mw-redirect" title="DsDNA-RT virus">dsDNA-RT virus</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="text-align: center;;width:1%;background:#CEDAF2;"><a href="/wiki/Subviral_agents" class="mw-redirect" title="Subviral agents">Subviral<br />agents</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:7em;text-align: center;;background:#CEDAF2;"><a href="/wiki/Viroid" title="Viroid">Viroid</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><i><a href="/wiki/Pospiviroidae" title="Pospiviroidae">Pospiviroidae</a></i></li> <li><i><a href="/wiki/Avsunviroidae" title="Avsunviroidae">Avsunviroidae</a></i></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7em;text-align: center;;background:#CEDAF2;"><a href="/wiki/Helper_virus" title="Helper virus">Helper-virus<br />dependent</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:1%;text-align: center;;background:#CEDAF2;"><a href="/wiki/Satellite_(biology)" title="Satellite (biology)">Satellite</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li>ssRNA satellite virus</li> <li>dsDNA satellite virus (<a href="/wiki/Virophage" title="Virophage">Virophage</a>)</li> <li>ssDNA satellite virus</li> <li>ssDNA satellite</li> <li>dsRNA satellite</li> <li>ssRNA satellite (<a href="/wiki/Virusoid" title="Virusoid">Virusoid</a>)</li> <li>Satellite-like nucleic acids <ul><li>RNA</li> <li>DNA</li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: center;;background:#CEDAF2;">Other</th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Defective_interfering_particle" title="Defective interfering particle">Defective interfering particle</a> <ul><li>RNA</li> <li>DNA</li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="width:7em;text-align: center;"><a href="/wiki/Prion" title="Prion">Prion</a></th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Prion" title="Prion">Mammalian prion</a></li> <li><a href="/wiki/Fungal_prion" title="Fungal prion">Fungal prion</a></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="text-align: center;;width:1%"><a href="/wiki/Nucleic_acid" title="Nucleic acid">Nucleic acid</a><br />self-replication</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"></div><table class="nowraplinks navbox-subgroup" style="border-spacing:0"><tbody><tr><th scope="row" class="navbox-group" style="width:7em;text-align: center;"><a href="/wiki/Mobile_genetic_elements" title="Mobile genetic elements">Mobile genetic<br />elements</a></th><td class="navbox-list-with-group navbox-list navbox-odd" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Mobilome" title="Mobilome">Mobilome</a> <ul><li><a href="/wiki/Horizontal_gene_transfer" title="Horizontal gene transfer">Horizontal gene transfer</a></li> <li><a href="/wiki/Genomic_island" title="Genomic island">Genomic island</a></li></ul></li> <li><a href="/wiki/Transposable_element" title="Transposable element">Transposable element</a> <ul><li><a href="/wiki/Retrotransposon" title="Retrotransposon">Class I or retrotransposon</a></li> <li><a href="/wiki/DNA_transposon" title="DNA transposon">Class II or DNA transposon</a></li></ul></li> <li><a href="/wiki/Plasmid" title="Plasmid">Plasmid</a> <ul><li><a href="/wiki/Fertility_factor_(bacteria)" class="mw-redirect" title="Fertility factor (bacteria)">Fertility</a></li> <li><a href="/wiki/R-factor" class="mw-redirect" title="R-factor">Resistance</a></li> <li><a href="/wiki/Colicin" title="Colicin">Col</a></li> <li>Degradative</li> <li><a href="/wiki/Virulence_factor" title="Virulence factor">Virulence</a>/<a href="/wiki/Ti_plasmid" title="Ti plasmid">Ti</a></li> <li>Cryptic</li></ul></li> <li><a href="/wiki/Cosmid" title="Cosmid">Cosmid</a> <ul><li><a href="/wiki/Fosmid" title="Fosmid">Fosmid</a></li></ul></li> <li><a href="/wiki/Phagemid" title="Phagemid">Phagemid</a></li> <li><a href="/wiki/Group_I_catalytic_intron" title="Group I catalytic intron">Group I intron</a></li> <li><a href="/wiki/Group_II_intron" title="Group II intron">Group II intron</a></li> <li><a href="/wiki/Retrozyme" title="Retrozyme">Retrozyme</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="width:7em;text-align: center;">Other aspects</th><td class="navbox-list-with-group navbox-list navbox-even" style="padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/DNA_replication" title="DNA replication">DNA replication</a> <ul><li><a href="/wiki/RNA-dependent_RNA_polymerase" title="RNA-dependent RNA polymerase">RNA replication</a></li></ul></li> <li><a href="/wiki/Chromosome" title="Chromosome">Chromosome</a> <ul><li><a href="/wiki/Linear_chromosome" title="Linear chromosome">Linear</a></li> <li><a href="/wiki/Circular_chromosome" title="Circular chromosome">Circular</a></li> <li><a href="/wiki/Extrachromosomal_DNA" title="Extrachromosomal DNA">Extrachromosomal DNA</a></li> <li><a href="/wiki/Secondary_chromosome" title="Secondary chromosome">Secondary chromosome</a></li></ul></li> <li><a href="/wiki/Genome" title="Genome">Genome</a> <ul><li><a href="/wiki/Gene" title="Gene">Gene</a></li> <li><a href="/wiki/Gene_duplication" title="Gene duplication">Gene duplication</a></li> <li><a href="/wiki/Non-coding_DNA" title="Non-coding DNA">Non-coding DNA</a></li></ul></li> <li><a href="/wiki/Origin_of_replication" title="Origin of replication">Origin of replication</a> <ul><li><a href="/wiki/Replicon_(genetics)" title="Replicon (genetics)">Replicon</a></li></ul></li> <li><a href="/wiki/Endogenous_viral_element" title="Endogenous viral element">Endogenous viral element</a> <ul><li><a href="/wiki/Provirus" title="Provirus">Provirus</a></li> <li><a href="/wiki/Prophage" title="Prophage">Prophage</a></li> <li><a href="/wiki/Endogenous_retrovirus" title="Endogenous retrovirus">Endogenous retrovirus</a></li> <li><a href="/wiki/Transpoviron" title="Transpoviron">Transpoviron</a></li></ul></li> <li><a href="/wiki/Repeated_sequence_(DNA)" title="Repeated sequence (DNA)">Repeated sequences in DNA</a> <ul><li><a href="/wiki/Tandem_repeat" title="Tandem repeat">Tandem repeat</a></li> <li><a href="/wiki/Interspersed_repeat" title="Interspersed repeat">Interspersed repeat</a></li></ul></li></ul> </div></td></tr></tbody></table><div></div></td></tr><tr><th scope="row" class="navbox-group" style="text-align: center;;width:1%"><a href="/wiki/Endosymbiont" title="Endosymbiont">Endosymbiosis</a></th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Mitochondrion" title="Mitochondrion">Mitochondrion</a> <ul><li><a href="/wiki/Mitosome" title="Mitosome">Mitosome</a></li> <li><a href="/wiki/Hydrogenosome" title="Hydrogenosome">Hydrogenosome</a></li></ul></li> <li><a href="/wiki/Plastid" title="Plastid">Plastid</a> <ul><li><a href="/wiki/Chloroplast" title="Chloroplast">Chloroplast</a></li> <li><a href="/wiki/Chromoplast" title="Chromoplast">Chromoplast</a></li> <li><a href="/wiki/Gerontoplast" title="Gerontoplast">Gerontoplast</a></li> <li><a href="/wiki/Leucoplast" title="Leucoplast">Leucoplast</a></li> <li><a href="/wiki/Apicoplast" title="Apicoplast">Apicoplast</a></li></ul></li> <li><a href="/wiki/Kappa_organism" title="Kappa organism">Kappa organism</a></li> <li>Organs <ul><li><a href="/wiki/Bacteriome" title="Bacteriome">Bacteriome</a></li> <li><a href="/wiki/Trophosome" title="Trophosome">Trophosome</a></li></ul></li> <li><a href="/wiki/Nitroplast" title="Nitroplast">Nitroplast</a></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="text-align: center;;width:1%"><a href="/wiki/Abiogenesis" title="Abiogenesis">Abiogenesis</a></th><td class="navbox-list-with-group navbox-list navbox-even hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Last_universal_common_ancestor" title="Last universal common ancestor">Last universal common ancestor</a></li> <li><a href="/wiki/Earliest_known_life_forms" title="Earliest known life forms">Earliest known life forms</a></li> <li>?<a href="/wiki/RNA_world" title="RNA world">RNA life</a> <ul><li><a href="/wiki/Ribozyme" title="Ribozyme">Ribozyme</a></li></ul></li> <li>†<a href="/wiki/Protocell" title="Protocell">Protocell</a></li> <li><a href="/wiki/Coacervate" title="Coacervate">Coacervate</a></li> <li><a href="/wiki/Proteinoid" title="Proteinoid">Proteinoid</a></li> <li><a href="/wiki/Sulphobes" title="Sulphobes">Sulphobe</a></li> <li>Research <ul><li><a href="/wiki/Model_lipid_bilayer" title="Model lipid bilayer">Model lipid bilayer</a></li> <li><a href="/wiki/Jeewanu" title="Jeewanu">Jeewanu</a></li></ul></li></ul> </div></td></tr><tr><th scope="row" class="navbox-group" style="text-align: center;;width:1%">See also</th><td class="navbox-list-with-group navbox-list navbox-odd hlist" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><a href="/wiki/Organism" title="Organism">Organism</a></li> <li><a href="/wiki/Cell_(biology)" title="Cell (biology)">Cell</a> <ul><li><a href="/wiki/Cell_division" title="Cell division">Cell division</a></li> <li><a href="/wiki/Artificial_cell" title="Artificial cell">Artificial cell</a></li></ul></li> <li><a href="/wiki/Non-cellular_life" title="Non-cellular life">Non-cellular life</a></li> <li><a href="/wiki/Synthetic_virology" title="Synthetic virology">Synthetic virus</a> <ul><li><a href="/wiki/Viral_vector" title="Viral vector">Viral vector</a></li> <li><a href="/wiki/Helper_dependent_virus" title="Helper dependent virus">Helper dependent virus</a></li></ul></li> <li>?<a href="/wiki/Nanobacterium" title="Nanobacterium">Nanobacterium</a></li> <li>?<a href="/wiki/Nanobe" title="Nanobe">Nanobe</a></li> <li><a href="/wiki/Cancer_cell" title="Cancer cell">Cancer cell</a> <ul><li><a href="/wiki/HeLa" title="HeLa">HeLa</a></li> <li><a href="/wiki/Clonally_transmissible_cancer" title="Clonally transmissible cancer">Clonally transmissible cancer</a></li></ul></li> <li><a href="/wiki/Virome" title="Virome">Virome</a></li></ul> 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.portal-bar{font-size:88%;font-weight:bold;display:flex;flex-flow:column wrap;align-items:baseline}.mw-parser-output .portal-bar-header{text-align:center;flex:0;padding-left:0.5em;margin:0 auto}.mw-parser-output .portal-bar-related{font-size:100%;align-items:flex-start}.mw-parser-output .portal-bar-content{display:flex;flex-flow:row wrap;align-items:center;flex:0;column-gap:1em;border-top:1px solid #a2a9b1;margin:0 auto;list-style:none}.mw-parser-output .portal-bar-content-related{border-top:none;margin:0;list-style:none}}.mw-parser-output .navbox+link+.portal-bar,.mw-parser-output .navbox+style+.portal-bar,.mw-parser-output .navbox+link+.portal-bar-bordered,.mw-parser-output .navbox+style+.portal-bar-bordered,.mw-parser-output .sister-bar+link+.portal-bar,.mw-parser-output .sister-bar+style+.portal-bar,.mw-parser-output .portal-bar+.navbox-styles+.navbox,.mw-parser-output .portal-bar+.navbox-styles+.sister-bar{margin-top:-1px}</style><div class="portal-bar noprint metadata noviewer portal-bar-bordered" role="navigation" aria-label="Portals"><span class="portal-bar-header"><a href="/wiki/Wikipedia:Contents/Portals" title="Wikipedia:Contents/Portals">Portal</a>:</span><ul class="portal-bar-content"><li class="portal-bar-item"><span class="nowrap"><span typeof="mw:File"><a href="/wiki/File:Sida-aids.png" class="mw-file-description"><img alt="icon" src="//upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Sida-aids.png/19px-Sida-aids.png" decoding="async" width="19" height="19" class="mw-file-element" srcset="//upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Sida-aids.png/29px-Sida-aids.png 1.5x, //upload.wikimedia.org/wikipedia/commons/thumb/2/2f/Sida-aids.png/38px-Sida-aids.png 2x" data-file-width="290" data-file-height="290" /></a></span> </span><a href="/wiki/Portal:Viruses" title="Portal:Viruses">Viruses</a></li></ul></div> <div class="navbox-styles"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1129693374"><link rel="mw-deduplicated-inline-style" href="mw-data:TemplateStyles:r1236075235"></div><div role="navigation" class="navbox" aria-labelledby="Taxon_identifiers" style="padding:3px"><table class="nowraplinks hlist navbox-inner" style="border-spacing:0;background:transparent;color:inherit"><tbody><tr><th scope="col" class="navbox-title" colspan="2"><div id="Taxon_identifiers" style="font-size:114%;margin:0 4em"><a href="/wiki/Help:Taxon_identifiers" title="Help:Taxon identifiers">Taxon identifiers</a></div></th></tr><tr><th scope="row" class="navbox-group" style="width:1%;text-align: left;"><i>Retroviridae</i></th><td class="navbox-list-with-group navbox-list navbox-odd" style="width:100%;padding:0"><div style="padding:0 0.25em"> <ul><li><span style="white-space:nowrap;"><a href="/wiki/Wikidata" title="Wikidata">Wikidata</a>: <span class="uid"><span class="external"><a href="https://www.wikidata.org/wiki/Q189308" class="extiw" title="wikidata:Q189308">Q189308</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Wikispecies" title="Wikispecies">Wikispecies</a>: <span class="uid"><span class="external"><a href="https://species.wikimedia.org/wiki/Retroviridae" class="extiw" title="wikispecies:Retroviridae">Retroviridae</a></span></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Catalogue_of_Life" title="Catalogue of Life">CoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.catalogueoflife.org/data/taxon/FM2">FM2</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Encyclopedia_of_Life" title="Encyclopedia of Life">EoL</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://eol.org/pages/8053">8053</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Global_Biodiversity_Information_Facility" title="Global Biodiversity Information Facility">GBIF</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.gbif.org/species/1940">1940</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/Interim_Register_of_Marine_and_Nonmarine_Genera" title="Interim Register of Marine and Nonmarine Genera">IRMNG</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.irmng.org/aphia.php?p=taxdetails&id=113829">113829</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/National_Center_for_Biotechnology_Information" title="National Center for Biotechnology Information">NCBI</a>: <span class="uid"><a rel="nofollow" class="external text" href="https://www.ncbi.nlm.nih.gov/Taxonomy/Browser/wwwtax.cgi?mode=Info&id=11632">11632</a></span></span></li> <li><span style="white-space:nowrap;"><a href="/wiki/World_Register_of_Marine_Species" title="World Register of Marine Species">WoRMS</a>: <span class="uid"><a rel="nofollow" class="external text" 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