Species Diversity and Invasion Resistance in a Marine Ecosystem

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data-design="pill" data-has="right-rail" data-type="research-article" vocab="http://schema.org/" typeof="ScholarlyArticle" lang="en" dir="ltr"><header data-extent="frontmatter"><div class="core-container"><div class="mb-1_5x mt-3"> <nav class="breadcrumbs"><div class="d-none d-sm-flex align-items-center"><a href="https://web.archive.org/web/20241007153257/https://www.science.org/" title="Home" class="crumb text-uppercase prev-crumb">Home</a><i class="icon-arrow-right mx-1"></i><a href="/web/20241007153257/https://www.science.org/journal/science" title="Science" class="crumb text-uppercase prev-crumb">Science</a><i class="icon-arrow-right mx-1"></i><a href="/web/20241007153257/https://www.science.org/toc/science/286/5444" title="Vol. 286, No. 5444" class="crumb text-uppercase prev-crumb">Vol. 286, No. 5444</a><i class="icon-arrow-right mx-1"></i><span class="crumb text-uppercase current-crumb">Species Diversity and Invasion Resistance in a Marine Ecosystem</span></div><div class="d-flex 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property="familyName">Osman</span></a></span></span></span><a href="#tab-contributors" class="to-authors-affiliations" data-id="article-authors-viewall">Authors Info & Affiliations</a></div><div class="core-self-citation"><div property="isPartOf" typeof="Periodical"><span property="name">Science</span></div><div class="core-date-published"><span property="datePublished">19 Nov 1999</span></div><div class="core-enumeration"><span property="isPartOf" typeof="PublicationVolume">Vol <span property="volumeNumber">286</span></span><span class="delimiter">, </span><span property="isPartOf" typeof="PublicationIssue">Issue <span property="issueNumber">5444</span></span></div><div property="pagination">pp. <span property="pageStart">1577</span>-<span property="pageEnd">1579</span></div><div class="doi"><a href="https://web.archive.org/web/20241007153257/https://doi.org/10.1126/science.286.5444.1577" property="sameAs">DOI: 10.1126/science.286.5444.1577</a></div></div><div class="info-panel"><div 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typeof="Text" role="doc-abstract"><h2 property="name">Abstract</h2><div role="paragraph">Theory predicts that systems that are more diverse should be more resistant to exotic species, but experimental tests are needed to verify this. In experimental communities of sessile marine invertebrates, increased species richness significantly decreased invasion success, apparently because species-rich communities more completely and efficiently used available space, the limiting resource in this system. Declining biodiversity thus facilitates invasion in this system, potentially accelerating the loss of biodiversity and the homogenization of the world's biota.</div></section></div></div><div class="core-container related"> </div><section id="bodymatter" data-extent="bodymatter" property="articleBody" typeof="Text"><div class="core-container"><section aria-labelledby="denial-block-title" class="denial-block"><h2 id="denial-block-title" class="denial-block__title">Get full access to this article</h2><p class="denial-block__text">View all available purchase options and get full access to this article.</p><div class="denial-block__cta"><a href="#core-collateral-purchase-access" role="button" class="btn btn--access btn-secondary"><i class="icon-lock-open"></i><span>CHECK ACCESS</span></a></div><footer class="denial-block__footer"><span>ALREADY A SUBSCRIBER OR AAAS MEMBER? </span><span>Sign in as an </span><a href="/web/20241007153257/https://www.science.org/action/showLogin?redirectUri=%2Fdoi%2F10.1126%2Fscience.286.5444.1577">individual</a><span> or via your </span><a href="/web/20241007153257/https://www.science.org/action/ssostart?redirectUri=%2Fdoi%2F10.1126%2Fscience.286.5444.1577">institution</a></footer></section></div></section><section id="backmatter" data-extent="backmatter"><div class="core-container"><section id="bibliography" role="doc-bibliography"><h2>REFERENCES AND NOTES</h2><div role="list"><div role="listitem" data-has="label"><div class="label">1</div><div id="RF1" class="citations"><div class="citation"><div class="citation-content">Wilcove D. 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property="name">Science</span></div><div class="core-enumeration"><span property="isPartOf" typeof="PublicationVolume">Volume <span property="volumeNumber">286</span></span> | <span property="isPartOf" typeof="PublicationIssue">Issue <span property="issueNumber">5444</span></span><br/><span property="datePublished">19 November 1999</span></div></div></div></section><section class="core-history"><h4>Submission history</h4><div><b class="core-label">Received</b>: 29 June 1999</div><div><b class="core-label">Accepted</b>: 11 October 1999</div><div><b class="core-label">Published in print</b>: 19 November 1999</div></section><section class="core-permissions"><h4>Permissions</h4><div>Request permissions for this article.</div><div><a class="btn" href="https://web.archive.org/web/20241007153257/https://s100.copyright.com/AppDispatchServlet?publisherName=AAAS&publication=sci&title=Species+Diversity+and+Invasion+Resistance+in+a+Marine+Ecosystem&publicationDate=1999-11-19&author=John+J.+Stachowicz%2C+Robert+B.+Whitlatch%2C+Richard+W.+Osman&contentID=10.1126%2Fscience.286.5444.1577&volumeNum=286&issueNum=5444&startPage=1577&endPage=1579&numPages=3&orderBeanReset=true">Request Permissions</a></div></section></section><section id="tab-contributors" aria-labelledby="contributors" role="tabpanel"><h3>Authors</h3><section class="core-authors"><h4>Affiliations</h4><div id="con1" property="author" typeof="Person"><div class="heading"><span property="givenName">John J.</span> <span property="familyName">Stachowicz</span><sup class="xref"><a href="#FN150" role="doc-noteref">*</a></sup></div><div class="content"><div class="affiliations"><div property="affiliation" typeof="Organization"><span property="name">Department of Marine Sciences, University of Connecticut, 1084 Shennecossett Road, Groton, CT 06340, USA.</span></div></div><div class="core-author-link"><a href="/web/20241007153257/https://www.science.org/authored-by/Stachowicz/John+J">View all articles by this author</a></div></div></div><div id="con2" property="author" typeof="Person"><div class="heading"><span property="givenName">Robert B.</span> <span property="familyName">Whitlatch</span></div><div class="content"><div class="affiliations"><div property="affiliation" typeof="Organization"><span property="name">Department of Marine Sciences, University of Connecticut, 1084 Shennecossett Road, Groton, CT 06340, USA.</span></div></div><div class="core-author-link"><a href="/web/20241007153257/https://www.science.org/authored-by/Whitlatch/Robert+B">View all articles by this author</a></div></div></div><div id="con3" property="author" typeof="Person"><div class="heading"><span property="givenName">Richard W.</span> <span property="familyName">Osman</span></div><div class="content"><div class="affiliations"><div property="affiliation" typeof="Organization"><span property="name">Academy of Natural Sciences, Estuarine Research Center, St. Leonard, MD 20685, USA.</span></div></div><div class="core-author-link"><a href="/web/20241007153257/https://www.science.org/authored-by/Osman/Richard+W">View all articles by this author</a></div></div></div></section><section class="core-authors-notes"><h4>Notes</h4><div role="doc-footnote" data-has="label"><div class="label">*</div><div id="FN150" role="paragraph">To whom correspondence should be addressed. E-mail: <a href="/web/20241007153257/https://www.science.org/cdn-cgi/l/email-protection#4a20393e2b29220a3f292524243c27643f29252424642f2e3f"><span class="__cf_email__" data-cfemail="90fae3e4f1f3f8d0e5f3fffefee6fdbee5f3fffefebef5f4e5">[email protected]</span></a></div></div></section></section></div><div id="core-collateral-metrics" role="tabpanel"><header><h2><i class="icon-metrics" aria-hidden="true"></i>Metrics & Citations</h2></header><section id="tab-metrics-inner" aria-labelledby="metrics-inner" role="tabpanel"><h3>Metrics</h3><section> <h4 class=""> Article Usage </h4> <div class="sidebar-metric-container data-source" data-source="/pb/widgets/fullSideBarMetric/getResponse?widgetId=92466116-7ac4-46b6-9de4-96b888b4cf23&pbContext=%3Bissue%3Aissue%3Adoi%5C%3A10.1126%2Fscience.1999.286.issue-5444%3Bpage%3Astring%3AArticle%2FChapter+View%3BrequestedJournal%3Ajournal%3Ascience%3Bctype%3Astring%3AJournal+Content%3BsubPage%3Astring%3AAbstract%3Barticle%3Aarticle%3Adoi%5C%3A10.1126%2Fscience.286.5444.1577%3Bwebsite%3Awebsite%3Aaaas-site%3Bjournal%3Ajournal%3Ascience%3Bwgroup%3Astring%3APublication+Websites%3BpageGroup%3Astring%3APublication+Pages&doi=10.1126%2Fscience.286.5444.1577"></div> <div class="sidebar-metrics"> <p class="metrics-sub-title mt-3"><b>Note:</b> The article usage is presented with a three- to four-day delay and will update daily once available. 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Stachowicz <em>et al.</em></span></li> </ul> <span class="ml-n1">,</span></div><span class="ml-1">Species Diversity and Invasion Resistance in a Marine Ecosystem.</span><span class="ml-1"><i>Science</i></span><span class="ml-1"><b>286</b>,</span><span class="ml-1">1577-1579</span><span class="ml-1">(1999).</span><span class="ml-1">DOI:<a href="https://web.archive.org/web/20241007153257/https://doi.org/10.1126/science.286.5444.1577" class="ml-1">10.1126/science.286.5444.1577</a></span></div> <div class="pill__item"> <h4 class="">Export citation</h4> <p>Select the format you want to export the citation of this publication.</p> <form action="/web/20241007153257/https://www.science.org/action/downloadCitation" name="frmCitmgr" class="citation-form" method="post" target="_self"><input type="hidden" name="doi" value="10.1126/science.286.5444.1577"><input type="hidden" name="downloadFileName" value="csp_286_"><input type="hidden" name="include" value="abs"> <fieldset class="format-select"> <div class="select-container"> <select id="slct_format" name="format" class="js__slcInclude" style=" padding: 7px; color: #595959; border: 1px solid #7f7f7f; "> <option value="" selected="selected">Please select one from the list</option> <option value="ris">RIS (ProCite, Reference Manager)</option> <option value="endnote">EndNote</option> <option value="bibtex">BibTex</option> <option value="medlars">Medlars</option> <option value="refworks">RefWorks</option> </select> </div> <label class="label-direct" for="direct"> <input id="direct" type="checkbox" name="direct" value="" checked="checked" style=" margin-top: 5px; margin-right: 7px; "> <span class="round-check"><span class="check"></span></span> Direct import </label> </fieldset> <footer class="form-footer"> <input onclick="onCitMgrSubmit()" class="btn btn-outline-dark btn-sm collapsed text-no-transform" type="submit" name="submit" value="EXPORT CITATION"> </footer> </form> </div> </section><section> <div data-widget-def="UX3CitedByWidget" data-widget-id="7dad93bb-d32a-4ca1-a6e0-6b0a63c85850" class="pill__item"> <h4 class=""> Cited by </h4> <section id="cited-by" class="cited-by"><div class="cited-by__wrapper"><div id="cited-by__content" class="cited-by__content"><ol data-total="4" data-source="" data-lazyload="false" class="cited-by__list list-unstyled"><li class="cited-by__entry pb-3 mb-4 border-bottom"><ul class="cited-by__entry__authors list-inline"><li class="list-inline-item cited-by__entry__author">Gail V. 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Rocha, </li><li class="list-inline-item cited-by__entry__author">Carlos Rumbold, </li><li class="list-inline-item cited-by__entry__author">Carlos Sánchez, </li><li class="list-inline-item cited-by__entry__author">Adolfo Sanjuan-Muñoz, </li><li class="list-inline-item cited-by__entry__author">Carmen Schlöder, </li><li class="list-inline-item cited-by__entry__author">Evangelina Schwindt, </li><li class="list-inline-item cited-by__entry__author">Janina Seemann, </li><li class="list-inline-item cited-by__entry__author">Alan Shanks, </li><li class="list-inline-item cited-by__entry__author">Nuno Simoes, </li><li class="list-inline-item cited-by__entry__author">Luis Skinner, </li><li class="list-inline-item cited-by__entry__author">Nancy Yolimar Suárez-Mozo, </li><li class="list-inline-item cited-by__entry__author">Martin Thiel, </li><li class="list-inline-item cited-by__entry__author">Nelson Valdivia, </li><li class="list-inline-item cited-by__entry__author">Ximena Velez-Zuazo, </li><li class="list-inline-item cited-by__entry__author">Edson A. Vieira, </li><li class="list-inline-item cited-by__entry__author">Bruno Vildoso, </li><li class="list-inline-item cited-by__entry__author">Ingo S. Wehrtmann, </li><li class="list-inline-item cited-by__entry__author">Matt Whalen, </li><li class="list-inline-item cited-by__entry__author">Lynn Wilbur, </li><li class="list-inline-item cited-by__entry__author">Gregory M. Ruiz, </li></ul><span class="cited-by__entry__title">Predator control of marine communities increases with temperature across 115 degrees of latitude, </span><span class="cited-by__entry__series-title">Science, </span><span class="cited-by__entry__volume"><strong>376</strong>, </span><span class="cited-by__entry__issue">6598, </span><span class="cited-by__entry__page-range">(1215-1219), </span><span class="cited-by__entry__pub-date">(2022).</span><a href="/web/20241007153257/https://www.science.org/doi/10.1126/science.abc4916" title="Abstract" class="cited-by__entry__doi">/doi/10.1126/science.abc4916</a><div class="cited-by__entry__extra-links"><a href="/web/20241007153257/https://www.science.org/doi/10.1126/science.abc4916" class="cited-by__entry__visitable">Abstract</a></div></li><li class="cited-by__entry pb-3 mb-4 border-bottom"><ul class="cited-by__entry__authors list-inline"><li class="list-inline-item cited-by__entry__author">Steven Murawski, </li><li class="list-inline-item cited-by__entry__author">Richard Methot, </li><li class="list-inline-item cited-by__entry__author">Galen Tromble, </li></ul><span class="cited-by__entry__title">Biodiversity Loss in the Ocean: How Bad Is It?, </span><span class="cited-by__entry__series-title">Science, </span><span class="cited-by__entry__volume"><strong>316</strong>, </span><span class="cited-by__entry__issue">5829, </span><span class="cited-by__entry__page-range">(1281-1284), </span><span class="cited-by__entry__pub-date">(2021).</span><a href="/web/20241007153257/https://www.science.org/doi/10.1126/science.316.5829.1281b" title="Abstract" class="cited-by__entry__doi">/doi/10.1126/science.316.5829.1281b</a><div class="cited-by__entry__extra-links"><a href="/web/20241007153257/https://www.science.org/doi/10.1126/science.316.5829.1281b" class="cited-by__entry__visitable">Abstract</a></div></li><li class="cited-by__entry pb-3 mb-4 border-bottom"><ul class="cited-by__entry__authors list-inline"><li class="list-inline-item cited-by__entry__author">Boris Worm, </li><li class="list-inline-item cited-by__entry__author">Edward B. Barbier, </li><li class="list-inline-item cited-by__entry__author">Nicola Beaumont, </li><li class="list-inline-item cited-by__entry__author">J. Emmett Duffy, </li><li class="list-inline-item cited-by__entry__author">Carl Folke, </li><li class="list-inline-item cited-by__entry__author">Benjamin S. Halpern, </li><li class="list-inline-item cited-by__entry__author">Jeremy B. C. Jackson, </li><li class="list-inline-item cited-by__entry__author">Heike K. Lotze, </li><li class="list-inline-item cited-by__entry__author">Fiorenza Micheli, </li><li class="list-inline-item cited-by__entry__author">Stephen R. Palumbi, </li><li class="list-inline-item cited-by__entry__author">Enric Sala, </li><li class="list-inline-item cited-by__entry__author">Kimberley A. Selkoe, </li><li class="list-inline-item cited-by__entry__author">John J. Stachowicz, </li><li class="list-inline-item cited-by__entry__author">Reg Watson, </li></ul><span class="cited-by__entry__title">Impacts of Biodiversity Loss on Ocean Ecosystem Services, </span><span class="cited-by__entry__series-title">Science, </span><span class="cited-by__entry__volume"><strong>314</strong>, </span><span class="cited-by__entry__issue">5800, </span><span class="cited-by__entry__page-range">(787-790), </span><span class="cited-by__entry__pub-date">(2021).</span><a href="/web/20241007153257/https://www.science.org/doi/10.1126/science.1132294" title="Abstract" class="cited-by__entry__doi">/doi/10.1126/science.1132294</a><div class="cited-by__entry__extra-links"><a href="/web/20241007153257/https://www.science.org/doi/10.1126/science.1132294" class="cited-by__entry__visitable">Abstract</a></div></li><li class="cited-by__entry pb-3 mb-4 border-bottom"><ul class="cited-by__entry__authors list-inline"><li class="list-inline-item cited-by__entry__author">Jeremy B. C. Jackson, </li><li class="list-inline-item cited-by__entry__author">Michael X. Kirby, </li><li class="list-inline-item cited-by__entry__author">Wolfgang H. Berger, </li><li class="list-inline-item cited-by__entry__author">Karen A. Bjorndal, </li><li class="list-inline-item cited-by__entry__author">Louis W. Botsford, </li><li class="list-inline-item cited-by__entry__author">Bruce J. Bourque, </li><li class="list-inline-item cited-by__entry__author">Roger H. Bradbury, </li><li class="list-inline-item cited-by__entry__author">Richard Cooke, </li><li class="list-inline-item cited-by__entry__author">Jon Erlandson, </li><li class="list-inline-item cited-by__entry__author">James A. Estes, </li><li class="list-inline-item cited-by__entry__author">Terence P. Hughes, </li><li class="list-inline-item cited-by__entry__author">Susan Kidwell, </li><li class="list-inline-item cited-by__entry__author">Carina B. Lange, </li><li class="list-inline-item cited-by__entry__author">Hunter S. Lenihan, </li><li class="list-inline-item cited-by__entry__author">John M. Pandolfi, </li><li class="list-inline-item cited-by__entry__author">Charles H. Peterson, </li><li class="list-inline-item cited-by__entry__author">Robert S. Steneck, </li><li class="list-inline-item cited-by__entry__author">Mia J. Tegner, </li><li class="list-inline-item cited-by__entry__author">Robert R. Warner, </li></ul><span class="cited-by__entry__title">Historical Overfishing and the Recent Collapse of Coastal Ecosystems, </span><span class="cited-by__entry__series-title">Science, </span><span class="cited-by__entry__volume"><strong>293</strong>, </span><span class="cited-by__entry__issue">5530, </span><span class="cited-by__entry__page-range">(629-637), </span><span class="cited-by__entry__pub-date">(2021).</span><a href="/web/20241007153257/https://www.science.org/doi/10.1126/science.1059199" title="Abstract" class="cited-by__entry__doi">/doi/10.1126/science.1059199</a><div class="cited-by__entry__extra-links"><a href="/web/20241007153257/https://www.science.org/doi/10.1126/science.1059199" class="cited-by__entry__visitable">Abstract</a></div></li></ol></div><div class="cited-by__spinner justify-content-center align-items-center d-none"><div role="status" class="spinner-border text-gray"><span class="sr-only">Loading...</span></div></div></div></section> </div> <div class="sidebar-metrics"> <p class="metrics-sub-title mt-3">Citation information is sourced from <a href="https://web.archive.org/web/20241007153257/https://www.crossref.org/services/cited-by/" class="external-link" target="_blank" rel="nofollow noopener" title="Follow link">Crossref Cited-by</a> service.</p> </div> </section></section></div><div id="core-collateral-fulltext-options" role="tabpanel"><header><h2><i class="icon-eye" aria-hidden="true"></i>View Options</h2></header><div tabindex="0" class="section--wrapper"><h3>Check Access</h3><section> <div class="pill__item pill__item--border-bottom"><h4>Log in to view the full text</h4>  <p><a type="button" class="btn btn-outline-primary btn-sm mt-3" href="/web/20241007153257/https://www.science.org/action/ssostart?redirectUri="><span>AAAS ID LOGIN</span></a></p> <div class="seamlessAccessDenial"><div class="seamlessAccessDenial__wrapper"><div role="button" class="seamlessAccessDenial__button"><span class="icon"><div class="js--loader"></div></span><span class="text"><b class="text-truncate">Loading institution options</b></span></div><div class="extra-login"><a href="/web/20241007153257/https://www.science.org/action/ssostart?redirectUri=" class="extra-login__link"><i class="icon-edit"></i>Select another institution</a></div></div></div> <p>AAAS login provides access to Science for AAAS Members, and access to other journals in the Science family to users who have purchased individual subscriptions.</p> <ul class="list-unstyled"> <li class="mb-2"><a href="https://web.archive.org/web/20241007153257/http://www.aaas.org/join/?CTC=SMAAJN">Become a AAAS Member</a></li> <li class="mb-2"><a href="https://web.archive.org/web/20241007153257/https://account.aaas.org/Identity/Lookup">Activate your AAAS ID</a></li> <li class="mb-2"><a href="/web/20241007153257/https://www.science.org/content/page/purchase-journal-access">Purchase Access to Other Journals in the Science Family</a></li> <li><a href="/web/20241007153257/https://www.science.org/content/page/access-and-subscriptions">Account Help</a></li> </ul> </div> <div class="pill__item pill__item--border-bottom"><h4>More options</h4><p><a href="https://web.archive.org/web/20241007153257/https://purchase.aaas.org/order/startnew/580?&returnUrl=https://www.science.org/doi/10.1126/science.286.5444.1577&ctc=SPREGFRE" class="animation-icon-shift d-flex align-items-center"><span class="text-uppercase">Register for free to read this article</span><i aria-hidden="true" class="icon-arrow-right ml-1 text-xl"></i></a></p><p class="mb-1_5x">As a service to the community, this article is available for free. 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